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<document id="4A2B04255ECED895D286D3890770C1BE" ID-CLB-Dataset="263342" ID-DOI="10.11646/zootaxa.5286.1.1" ID-GBIF-Dataset="67e7a105-1f62-414a-83d9-f758039eb726" ID-ISSN="1175-5326" ID-Zenodo-Dep="7958937" ID-ZooBank="F9834684-24D3-4795-B5EB-77B451DF856D" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1684793492880" checkinUser="plazi" docAuthor="Shi, Hongliang &amp; Liang, Hongbin" docDate="2023" docId="038776236203FFA42DEFB367FC055C8B" docLanguage="en" docName="zootaxa.5286.1.1.pdf" docOrigin="Zootaxa 5286 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5286.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Parena (Bothynoptera) shapingensis Xie &amp; Yu. B 1993" docType="treatment" docVersion="8" lastPageNumber="72" masterDocId="FFBE0E5B6246FFE32D78B27BFFE75D5A" masterDocTitle="Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)" masterLastPageNumber="144" masterPageNumber="1" pageNumber="70" updateTime="1699506634024" updateUser="plazi">
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<mods:title id="E2B8E375E3E6FE63EE276E0C7CF37BA7">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
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<subSubSection id="C33494BE6203FFA62DEFB367FCA85C6C" box="[151,847,283,310]" pageId="69" pageNumber="70" type="nomenclature">
<paragraph id="8B91C7356203FFA62DEFB367FCA85C6C" blockId="69.[151,847,283,346]" box="[151,847,283,310]" pageId="69" pageNumber="70">
<emphasis id="B95A1B276203FFA62DEFB367FCA85C6C" bold="true" box="[151,847,283,310]" pageId="69" pageNumber="70">
[17]
<taxonomicName id="4C2EBCB66203FFA62DB7B367FCA85C6C" ID-CoL="4DKQJ" authority="Xie &amp; Yu, 1993" authorityName="Xie &amp; Yu. B" authorityYear="1993" box="[207,847,283,310]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="shapingensis" subGenus="Bothynoptera">
<emphasis id="B95A1B276203FFA62DB7B367FEC05C6C" bold="true" box="[207,295,284,310]" italics="true" pageId="69" pageNumber="70">Parena</emphasis>
(
<emphasis id="B95A1B276203FFA62C40B360FE3A5C6C" bold="true" box="[312,477,283,310]" italics="true" pageId="69" pageNumber="70">Bothynoptera</emphasis>
)
<emphasis id="B95A1B276203FFA62C96B360FD6D5C6C" bold="true" box="[494,650,283,310]" italics="true" pageId="69" pageNumber="70">shapingensis</emphasis>
<bibRefCitation id="EFBFBAC46203FFA62FEAB367FCA85C6C" author="Xie, W. P. &amp; Yu, P. Y." box="[658,847,284,310]" pageId="69" pageNumber="70" pagination="185 - 195" refId="ref113326" refString="Xie, W. P. &amp; Yu, P. Y. (1993) On the Chinese species of Parena Motschulsky (Coleoptera: Carabidae). Sinozoologia, 10, 185 - 195." type="journal article" year="1993">Xie &amp; Yu, 1993</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6203FFA62DEFB33BFD495C00" box="[151,686,320,346]" pageId="69" pageNumber="70" type="description">
<paragraph id="8B91C7356203FFA62DEFB33BFD495C00" blockId="69.[151,847,283,346]" box="[151,686,320,346]" pageId="69" pageNumber="70">
Habitus:
<figureCitation id="1315DBB06203FFA62D85B33BFE425C00" box="[253,421,320,346]" captionStart="FIGURE 40" captionStartId="69.[151,250,1854,1878]" captionTargetBox="[155,1433,502,1823]" captionTargetId="figure-95@69.[151,1436,498,1831]" captionTargetPageId="69" captionText="FIGURE 40. Habitus of Parena (Bothynoptera) shapingensis Xie &amp; Yu and Parena (Bothynoptera) monostigma (Bates). A Holotype of Parena shapingensis Xie &amp; Yu. B P. shapingensis Xie &amp; Yu, male from Yunnan. C Lectotype of Parena monostigma (Bates). D Holotype of Parena japoincia Jedlička [=Parena monostigma (Bates)]. E Holotype of Parena koreana Kirschenhofer [=Parena monostigma (Bates)]. F P. monostigma (Bates), male from Zhejiang. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959019" httpUri="https://zenodo.org/record/7959019/files/figure.png" pageId="69" pageNumber="70">Figs 40A, 40B</figureCitation>
. Male genitalia:
<figureCitation id="1315DBB06203FFA62F27B33BFD495C00" box="[607,686,320,346]" captionStart="FIGURE 41" captionStartId="70.[151,250,1651,1675]" captionTargetBox="[162,1420,826,1618]" captionTargetId="figure-315@70.[151,1436,813,1628]" captionTargetPageId="70" captionText="FIGURE 41. Median lobe of male genitalia for Parena (Bothynoptera) shapingensis Xie &amp; Yu, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959021" httpUri="https://zenodo.org/record/7959021/files/figure.png" pageId="69" pageNumber="70">Fig. 41</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6203FFA62DEFB3FFFCC65C80" pageId="69" pageNumber="70" type="reference_group">
<paragraph id="8B91C7356203FFA62DEFB3FFFBCF5CC7" blockId="69.[151,1437,388,474]" box="[151,1064,388,413]" pageId="69" pageNumber="70">
<bibRefCitation id="EFBFBAC46203FFA62DEFB3FFFE8D5CC7" author="Xie, W. P. &amp; Yu, P. Y." box="[151,362,388,413]" pageId="69" pageNumber="70" pagination="185 - 195" refId="ref113326" refString="Xie, W. P. &amp; Yu, P. Y. (1993) On the Chinese species of Parena Motschulsky (Coleoptera: Carabidae). Sinozoologia, 10, 185 - 195." type="journal article" year="1993">Xie &amp; Yu, 1993: 192</bibRefCitation>
(type locality: Shaping;
<typeStatus id="549579976203FFA62F1DB3FFFD595CC6" box="[613,702,388,412]" pageId="69" pageNumber="70" type="holotype">holotype</typeStatus>
in IZAS);
<bibRefCitation id="EFBFBAC46203FFA62E54B3FFFBC35CC7" author="Kirschenhofer, E." box="[812,1060,388,413]" pageId="69" pageNumber="70" pagination="87 - 103" refId="ref111612" refString="Kirschenhofer, E. (2006) Neue Arten der Gattung Parena Motschulsky aus Japan, China Und Indonesien (Sulawesi) (Coleoptera: Carabidae). Mitteilungen des Internationalen Entomologischen Vereins E. V. Frankfurt, 31 (3 - 4), 87 - 103." type="journal article" year="2006">Kirschenhofer, 2006: 89</bibRefCitation>
.
</paragraph>
<paragraph id="8B91C7356203FFA62DEFB3D8FCC65C80" blockId="69.[151,1437,388,474]" pageId="69" pageNumber="70">
<taxonomicName id="4C2EBCB66203FFA62DEFB3D8FD815CE1" ID-CoL="4DKQX" authority="Kirschenhofer, 1994: 1019" authorityName="Kirschenhofer" authorityPageNumber="1019" authorityYear="1994" box="[151,614,419,443]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="yunnana">
<emphasis id="B95A1B276203FFA62DEFB3D8FEAE5CE0" bold="true" box="[151,329,419,442]" italics="true" pageId="69" pageNumber="70">Parena yunnana</emphasis>
<materialsCitation id="3B46CD686203FFA62C2AB3D8FD815CE1" ID-GBIF-Occurrence="4115871360" box="[338,614,419,443]" country="Austria" location="Naturhistorisches Museum" pageId="69" pageNumber="70" specimenCount="1" stateProvince="Wien" typeStatus="holotype">
<bibRefCitation id="EFBFBAC46203FFA62C2AB3D8FD815CE1" author="Kirschenhofer, E." box="[338,614,419,443]" pageId="69" pageNumber="70" pagination="999 - 1067" refId="ref111570" refString="Kirschenhofer, E. (1994) Neue und wenig bekannte Carabidae aus der palaarktischen und orientalischen Region (Col. Carabidae, Lebiinae, Odacanthinae, Brachininae, Panagaeinae). Linzer Biologische Beitrage, 26, 999 - 1067." type="journal article" year="1994">Kirschenhofer, 1994: 1019</bibRefCitation>
</materialsCitation>
</taxonomicName>
(type locality: Yunnan: Lijiang;
<typeStatus id="549579976203FFA62EC4B3D8FBF25CE1" box="[956,1045,419,443]" pageId="69" pageNumber="70" type="holotype">holotype</typeStatus>
in NMW:
<location id="8EF191EE6203FFA629F2B3D8FA7A5CE1" LSID="urn:lsid:plazi:treatment:038776236203FFA42DEFB367FC055C8B:8EF191EE6203FFA629F2B3D8FA7A5CE1" box="[1162,1437,419,443]" country="Austria" name="Naturhistorisches Museum" pageId="69" pageNumber="70" stateProvince="Wien">Naturhistorisches Museum</location>
<collectingRegion id="49EA09D76203FFA62DBFB3B9FEE75C80" box="[199,256,450,474]" country="Austria" name="Wien" pageId="69" pageNumber="70">Wien</collectingRegion>
,
<collectingRegion id="49EA09D76203FFA62C71B3B9FEB45C80" box="[265,339,450,474]" country="Austria" name="Wien" pageId="69" pageNumber="70">Vienne</collectingRegion>
,
<collectingCountry id="F33987A56203FFA62C24B3B9FE4C5C80" box="[348,427,450,474]" name="Austria" pageId="69" pageNumber="70">Austria</collectingCountry>
);
<bibRefCitation id="EFBFBAC46203FFA62CC4B3B9FD525C80" author="Kirschenhofer, E." box="[444,693,450,474]" pageId="69" pageNumber="70" pagination="87 - 103" refId="ref111612" refString="Kirschenhofer, E. (2006) Neue Arten der Gattung Parena Motschulsky aus Japan, China Und Indonesien (Sulawesi) (Coleoptera: Carabidae). Mitteilungen des Internationalen Entomologischen Vereins E. V. Frankfurt, 31 (3 - 4), 87 - 103." type="journal article" year="2006">Kirschenhofer, 2006: 89</bibRefCitation>
.
<emphasis id="B95A1B276203FFA62FC7B3BAFCC65C80" bold="true" box="[703,801,449,474]" pageId="69" pageNumber="70">Syn. nov.</emphasis>
</paragraph>
</subSubSection>
<caption id="DF5197BD6203FFA62DEFB545FB985A98" ID-DOI="http://doi.org/10.5281/zenodo.7959019" ID-Zenodo-Dep="7959019" httpUri="https://zenodo.org/record/7959019/files/figure.png" pageId="69" pageNumber="70" startId="69.[151,250,1854,1878]" targetBox="[155,1433,502,1823]" targetPageId="69" targetType="figure">
<paragraph id="8B91C7356203FFA62DEFB545FB985A98" blockId="69.[151,1436,1854,1987]" pageId="69" pageNumber="70">
<emphasis id="B95A1B276203FFA62DEFB545FEFF5A0C" bold="true" box="[151,280,1854,1878]" pageId="69" pageNumber="70">FIGURE 40</emphasis>
. Habitus of
<taxonomicName id="4C2EBCB66203FFA62CEBB544FC8F5A0C" authority="Xie &amp; Yu" authorityName="Xie &amp; Yu. B" authorityYear="1993" box="[403,872,1854,1879]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="shapingensis" subGenus="Bothynoptera">
<emphasis id="B95A1B276203FFA62CEBB544FE3A5A0C" box="[403,477,1855,1878]" italics="true" pageId="69" pageNumber="70">Parena</emphasis>
(
<emphasis id="B95A1B276203FFA62C92B545FD935A0C" box="[490,628,1854,1878]" italics="true" pageId="69" pageNumber="70">Bothynoptera</emphasis>
)
<emphasis id="B95A1B276203FFA62FFAB545FCE25A0C" box="[642,773,1854,1878]" italics="true" pageId="69" pageNumber="70">shapingensis</emphasis>
Xie &amp; Yu
</taxonomicName>
and
<taxonomicName id="4C2EBCB66203FFA62EE0B544FAB35A0D" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[920,1364,1854,1879]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="monostigma" subGenus="Bothynoptera">
<emphasis id="B95A1B276203FFA62EE0B544FC055A0C" box="[920,994,1855,1878]" italics="true" pageId="69" pageNumber="70">Parena</emphasis>
(
<emphasis id="B95A1B276203FFA62E97B545FB9E5A0C" box="[1007,1145,1854,1878]" italics="true" pageId="69" pageNumber="70">Bothynoptera</emphasis>
)
<emphasis id="B95A1B276203FFA629FFB544FAE45A0C" box="[1159,1283,1855,1878]" italics="true" pageId="69" pageNumber="70">monostigma</emphasis>
(Bates)
</taxonomicName>
.
<emphasis id="B95A1B276203FFA62825B545FA885A0C" bold="true" box="[1373,1391,1854,1878]" pageId="69" pageNumber="70">A</emphasis>
Holotype of
<taxonomicName id="4C2EBCB66203FFA62D81B518FDB65A21" authority="Xie &amp; Yu. B" authorityName="Xie &amp; Yu. B" authorityYear="1993" box="[249,593,1890,1915]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="shapingensis">
<emphasis id="B95A1B276203FFA62D81B518FE2B5A20" box="[249,460,1890,1914]" italics="true" pageId="69" pageNumber="70">Parena shapingensis</emphasis>
Xie &amp; Yu.
<emphasis id="B95A1B276203FFA62F38B518FDB65A21" bold="true" box="[576,593,1891,1915]" pageId="69" pageNumber="70">B</emphasis>
</taxonomicName>
<taxonomicName id="4C2EBCB66203FFA62F2FB518FCBA5A20" authority="Xie &amp; Yu" authorityName="Xie &amp; Yu. B" authorityYear="1993" box="[599,861,1890,1915]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="shapingensis">
<emphasis id="B95A1B276203FFA62F2FB518FD145A20" box="[599,755,1890,1914]" italics="true" pageId="69" pageNumber="70">P. shapingensis</emphasis>
Xie &amp; Yu
</taxonomicName>
, male from Yunnan.
<emphasis id="B95A1B276203FFA62948B519FBA55A20" bold="true" box="[1072,1090,1890,1914]" pageId="69" pageNumber="70">C</emphasis>
Lectotype of
<taxonomicName id="4C2EBCB66203FFA629A8B518FF045AC5" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1873" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="monostigma">
<emphasis id="B95A1B276203FFA629A8B518FA7B5A20" box="[1232,1436,1891,1914]" italics="true" pageId="69" pageNumber="70">Parena monostigma</emphasis>
(Bates)
</taxonomicName>
.
<emphasis id="B95A1B276203FFA62D94B5FCFF195AC5" bold="true" box="[236,254,1927,1951]" pageId="69" pageNumber="70">D</emphasis>
Holotype of
<taxonomicName id="4C2EBCB66203FFA62CF9B5FCFD6F5AC4" authority="Jedlicka" authorityName="Jedlicka" authorityYear="1946" box="[385,648,1926,1950]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="japoincia">
<emphasis id="B95A1B276203FFA62CF9B5FCFDD75AC4" box="[385,560,1927,1950]" italics="true" pageId="69" pageNumber="70">Parena japoincia</emphasis>
Jedlička
</taxonomicName>
[=
<taxonomicName id="4C2EBCB66203FFA62FDCB5FCFC245AC5" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[676,963,1927,1951]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="monostigma">
<emphasis id="B95A1B276203FFA62FDCB5FCFC885AC4" box="[676,879,1927,1950]" italics="true" pageId="69" pageNumber="70">Parena monostigma</emphasis>
(Bates)
</taxonomicName>
].
<emphasis id="B95A1B276203FFA62EA9B5FCFC055AC5" bold="true" box="[977,994,1927,1951]" pageId="69" pageNumber="70">E</emphasis>
Holotype of
<taxonomicName id="4C2EBCB66203FFA6291DB5FCFA7B5AC4" authority="Kirschenhofer" authorityName="Kirschenhofer" authorityYear="1994" box="[1125,1436,1926,1950]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="koreana">
<emphasis id="B95A1B276203FFA6291DB5FCFAE15AC4" box="[1125,1286,1926,1950]" italics="true" pageId="69" pageNumber="70">Parena koreana</emphasis>
Kirschenhofer
</taxonomicName>
[=
<taxonomicName id="4C2EBCB66203FFA62DD6B5D0FE285A99" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[174,463,1963,1987]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="monostigma">
<emphasis id="B95A1B276203FFA62DD6B5D0FE9D5A98" box="[174,378,1963,1986]" italics="true" pageId="69" pageNumber="70">Parena monostigma</emphasis>
(Bates)
</taxonomicName>
].
<emphasis id="B95A1B276203FFA62CA6B5D0FE0A5A99" bold="true" box="[478,493,1963,1987]" pageId="69" pageNumber="70">F</emphasis>
<taxonomicName id="4C2EBCB66203FFA62C8CB5D0FD3D5A99" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[500,730,1963,1987]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="monostigma">
<emphasis id="B95A1B276203FFA62C8CB5D0FD6E5A98" box="[500,649,1963,1986]" italics="true" pageId="69" pageNumber="70">P. monostigma</emphasis>
(Bates)
</taxonomicName>
, male from Zhejiang. Scale bars = 2 mm.
</paragraph>
</caption>
<subSubSection id="C33494BE6200FFA52DEFB2ECFC425DA3" pageId="70" pageNumber="71" type="materials_examined">
<paragraph id="8B91C7356200FFA52DEFB2ECFC425DA3" blockId="70.[151,1437,150,753]" pageId="70" pageNumber="71">
<emphasis id="B95A1B276200FFA52DEFB2ECFE5D5DEA" bold="true" box="[151,442,150,177]" pageId="70" pageNumber="71">Type material examined</emphasis>
.
<taxonomicName id="4C2EBCB66200FFA52CB1B2ECFCD45DEB" authority="Xie &amp; Yu" authorityName="Xie &amp; Yu. B" authorityYear="1993" box="[457,819,150,177]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="shapingensis">
<emphasis id="B95A1B276200FFA52CB1B2ECFCD45DEB" bold="true" box="[457,819,150,177]" pageId="70" pageNumber="71">
<emphasis id="B95A1B276200FFA52CB1B2ECFD5C5DEB" bold="true" box="[457,699,150,177]" italics="true" pageId="70" pageNumber="71">Parena shapingensis</emphasis>
Xie &amp; Yu
</emphasis>
</taxonomicName>
:
<emphasis id="B95A1B276200FFA52E3BB2ECFC575DEB" bold="true" box="[835,944,151,177]" pageId="70" pageNumber="71">
<typeStatus id="549579976200FFA52E3BB2ECFC575DEB" box="[835,944,151,177]" pageId="70" pageNumber="71" type="holotype">Holotype</typeStatus>
</emphasis>
(IZAS,
<figureCitation id="1315DBB06200FFA52977B2ECFB9E5DEB" box="[1039,1145,151,177]" captionStart="FIGURE 40" captionStartId="69.[151,250,1854,1878]" captionTargetBox="[155,1433,502,1823]" captionTargetId="figure-95@69.[151,1436,498,1831]" captionTargetPageId="69" captionText="FIGURE 40. Habitus of Parena (Bothynoptera) shapingensis Xie &amp; Yu and Parena (Bothynoptera) monostigma (Bates). A Holotype of Parena shapingensis Xie &amp; Yu. B P. shapingensis Xie &amp; Yu, male from Yunnan. C Lectotype of Parena monostigma (Bates). D Holotype of Parena japoincia Jedlička [=Parena monostigma (Bates)]. E Holotype of Parena koreana Kirschenhofer [=Parena monostigma (Bates)]. F P. monostigma (Bates), male from Zhejiang. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959019" httpUri="https://zenodo.org/record/7959019/files/figure.png" pageId="70" pageNumber="71">Fig. 40A</figureCitation>
): male, body length =
<quantity id="4CD66AD06200FFA52801B2EDFF225D8F" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="70" pageNumber="71" unit="mm" value="6.5">6.5 mm</quantity>
, board mounted, &quot;
<emphasis id="B95A1B276200FFA52CE8B2C0FE0B5D8F" box="[400,492,187,213]" italics="true" pageId="70" pageNumber="71">Shaping</emphasis>
/
<emphasis id="B95A1B276200FFA52F78B2C0FD9B5D8F" box="[512,636,187,213]" italics="true" pageId="70" pageNumber="71">
<date id="FF90E1F56200FFA52F78B2C0FD9B5D8F" box="[512,636,187,213]" pageId="70" pageNumber="71" value="1943-09-27">27.IX.1943</date>
</emphasis>
&quot; [in Chinese], &quot;
<typeStatus id="549579976200FFA52E54B2C0FC2F5D8F" box="[812,968,187,213]" pageId="70" pageNumber="71" type="holotype">HOLOTYPE</typeStatus>
&quot; [red label], &quot;
<taxonomicName id="4C2EBCB66200FFA52918B2C0FAAE5D8F" authorityName="Xie &amp; Yu. B" authorityYear="1993" box="[1120,1353,187,213]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="shapingensis">
<emphasis id="B95A1B276200FFA52918B2C0FAAE5D8F" box="[1120,1353,187,213]" italics="true" pageId="70" pageNumber="71">Parena shapingensis</emphasis>
</taxonomicName>
/
<emphasis id="B95A1B276200FFA52825B2C0FF555DA2" italics="true" pageId="70" pageNumber="71">Xie et Yu</emphasis>
/ Xie Weiping det. 1991&quot;, &quot;IOZ(E) 211403&quot; [blue label] &lt;
<figureCitation id="1315DBB06200FFA52E47B2A4FC775DA3" box="[831,912,223,249]" captionStart="FIGURE 41" captionStartId="70.[151,250,1651,1675]" captionTargetBox="[162,1420,826,1618]" captionTargetId="figure-315@70.[151,1436,813,1628]" captionTargetPageId="70" captionText="FIGURE 41. Median lobe of male genitalia for Parena (Bothynoptera) shapingensis Xie &amp; Yu, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959021" httpUri="https://zenodo.org/record/7959021/files/figure.png" pageId="70" pageNumber="71">Fig. 41</figureCitation>
&gt;.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6200FFA52DBFB378FA685F3B" pageId="70" pageNumber="71" type="discussion">
<paragraph id="8B91C7356200FFA52DBFB378FD285CAF" blockId="70.[151,1437,150,753]" pageId="70" pageNumber="71">
<emphasis id="B95A1B276200FFA52DBFB378FDC45C47" bold="true" box="[199,547,259,285]" pageId="70" pageNumber="71">Notes on types and synonyms</emphasis>
.
<taxonomicName id="4C2EBCB66200FFA52F49B378FC735C47" authority="Xie &amp; Yu" authorityName="Xie &amp; Yu. B" authorityYear="1993" box="[561,916,258,285]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="shapingensis">
<emphasis id="B95A1B276200FFA52F49B378FC735C47" bold="true" box="[561,916,258,285]" pageId="70" pageNumber="71">
<emphasis id="B95A1B276200FFA52F49B378FCC75C47" bold="true" box="[561,800,258,285]" italics="true" pageId="70" pageNumber="71">Parena shapingensis</emphasis>
Xie &amp; Yu
</emphasis>
</taxonomicName>
: The
<typeStatus id="549579976200FFA52EACB378FBD15C47" box="[980,1078,259,285]" pageId="70" pageNumber="71" type="holotype">holotype</typeStatus>
of this species is briefly labeled as &quot;Shaping,
<date id="FF90E1F56200FFA52C53B35CFE495C1B" box="[299,430,295,321]" pageId="70" pageNumber="71" value="1943-09-27">
<collectingDate id="EFD4181D6200FFA52C53B35CFE495C1B" box="[299,430,295,321]" pageId="70" pageNumber="71" value="1943-09-27">27.IX.1943</collectingDate>
</date>
&quot; in handwritten
<collectingCounty id="62F0BFB96200FFA52F1EB35CFD275C1B" box="[614,704,295,321]" pageId="70" pageNumber="71">Chinese</collectingCounty>
without other explanation of this locality.
<collectingMunicipality id="6BF55D4F6200FFA529E4B35CFB545C1B" box="[1180,1203,295,321]" pageId="70" pageNumber="71">In</collectingMunicipality>
the original description, the type locality was explained as
<collectingRegion id="49EA09D76200FFA52F2AB330FD325C3F" box="[594,725,331,357]" country="China" name="Guangdong" pageId="70" pageNumber="71">Guangdong</collectingRegion>
(Shaping), referring to
<location id="8EF191EE6200FFA52EA6B330FB9E5C3F" LSID="urn:lsid:plazi:treatment:038776236203FFA42DEFB367FC055C8B:8EF191EE6200FFA52EA6B330FB9E5C3F" box="[990,1145,331,357]" county="Chinese" latitude="29.533333" longLatPrecision="1226" longitude="106.45" municipality="In" name="Shaping town" pageId="70" pageNumber="71" stateProvince="Guangdong">Shaping town</location>
of
<location id="8EF191EE6200FFA529E6B330FB155C3F" LSID="urn:lsid:plazi:treatment:038776236203FFA42DEFB367FC055C8B:8EF191EE6200FFA529E6B330FB155C3F" box="[1182,1266,331,357]" county="Chinese" latitude="29.533333" longLatPrecision="1226" longitude="106.45" municipality="In" name="Heshan" pageId="70" pageNumber="71" stateProvince="Guangdong">Heshan</location>
city.
<location id="8EF191EE6200FFA52848B330FA7F5C3F" LSID="urn:lsid:plazi:treatment:038776236203FFA42DEFB367FC055C8B:8EF191EE6200FFA52848B330FA7F5C3F" box="[1328,1432,331,357]" county="Chinese" latitude="29.533333" longLatPrecision="1226" longitude="106.45" municipality="In" name="However" pageId="70" pageNumber="71" stateProvince="Guangdong">However</location>
, this speculation could be wrong because
<collectingRegion id="49EA09D76200FFA52F0EB314FD1E5CD3" box="[630,761,367,393]" country="China" name="Guangdong" pageId="70" pageNumber="71">Guangdong</collectingRegion>
is far from all other known localities of this species.
<location id="8EF191EE6200FFA52809B314FA7A5CD3" LSID="urn:lsid:plazi:treatment:038776236203FFA42DEFB367FC055C8B:8EF191EE6200FFA52809B314FA7A5CD3" box="[1393,1437,367,393]" county="Chinese" latitude="29.533333" longLatPrecision="1226" longitude="106.45" municipality="In" name="The" pageId="70" pageNumber="71" stateProvince="Guangdong">The</location>
handwriting of the locality label is identified that of a famous
<location id="8EF191EE6200FFA52E34B3E8FC415CF7" LSID="urn:lsid:plazi:treatment:038776236203FFA42DEFB367FC055C8B:8EF191EE6200FFA52E34B3E8FC415CF7" box="[844,934,403,429]" county="Chinese" latitude="29.533333" longLatPrecision="1226" longitude="106.45" municipality="In" name="Chinese" pageId="70" pageNumber="71" stateProvince="Guangdong">Chinese</location>
entomologist, Prof. Sicien Chen. He worked in
<location id="8EF191EE6200FFA52DCDB3CCFE925C8B" LSID="urn:lsid:plazi:treatment:038776236203FFA42DEFB367FC055C8B:8EF191EE6200FFA52DCDB3CCFE925C8B" box="[181,373,439,465]" county="Chinese" latitude="29.533333" longLatPrecision="1226" longitude="106.45" municipality="In" name="Academia Sinica" pageId="70" pageNumber="71" stateProvince="Guangdong">Academia Sinica</location>
in
<collectingRegion id="49EA09D76200FFA52CE5B3CCFDFE5C8B" box="[413,537,439,465]" country="China" name="Chongqing" pageId="70" pageNumber="71">Chongqing</collectingRegion>
from 1939 to 1946.
<location id="8EF191EE6200FFA52E73B3CCFCA45C8B" LSID="urn:lsid:plazi:treatment:038776236203FFA42DEFB367FC055C8B:8EF191EE6200FFA52E73B3CCFCA45C8B" box="[779,835,439,465]" county="Chinese" latitude="29.533333" longLatPrecision="1226" longitude="106.45" municipality="In" name="Thus" pageId="70" pageNumber="71" stateProvince="Guangdong">Thus</location>
the type locality &quot;Shaping&quot; is very likely to refer to
<location id="8EF191EE6200FFA52DEFB3A0FEEA5CAF" LSID="urn:lsid:plazi:treatment:038776236203FFA42DEFB367FC055C8B:8EF191EE6200FFA52DEFB3A0FEEA5CAF" box="[151,269,475,501]" county="Chinese" latitude="29.533333" longLatPrecision="1226" longitude="106.45" municipality="In" name="Shapingba" pageId="70" pageNumber="71" stateProvince="Guangdong">Shapingba</location>
(
<geoCoordinate id="EE1AA1F26200FFA52C65B3A0FE9C5CAF" box="[285,379,475,501]" degrees="29" direction="north" minutes="32" orientation="latitude" pageId="70" pageNumber="71" precision="925" value="29.533333">N29°32'</geoCoordinate>
<geoCoordinate id="EE1AA1F26200FFA52CF9B3A0FE0C5CAF" box="[385,491,475,501]" degrees="106" direction="east" minutes="27" orientation="longitude" pageId="70" pageNumber="71" precision="925" value="106.45">E106°27'</geoCoordinate>
) in
<collectingRegion id="49EA09D76200FFA52F6EB3A0FD755CAF" box="[534,658,475,501]" country="China" name="Chongqing" pageId="70" pageNumber="71">Chongqing</collectingRegion>
<location id="8EF191EE6200FFA52FE1B3A0FD2C5CAF" LSID="urn:lsid:plazi:treatment:038776236203FFA42DEFB367FC055C8B:8EF191EE6200FFA52FE1B3A0FD2C5CAF" box="[665,715,475,501]" county="Chinese" latitude="29.533333" longLatPrecision="1226" longitude="106.45" municipality="In" name="City" pageId="70" pageNumber="71" stateProvince="Guangdong">City</location>
.
</paragraph>
<paragraph id="8B91C7356200FFA52DBFB384FA685F3B" blockId="70.[151,1437,150,753]" pageId="70" pageNumber="71">
<taxonomicName id="4C2EBCB66200FFA52DBFB384FDA65F42" authority="Kirschenhofer" authorityName="Kirschenhofer" authorityYear="1994" box="[199,577,510,537]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="yunnana">
<emphasis id="B95A1B276200FFA52DBFB384FDA65F42" bold="true" box="[199,577,510,537]" pageId="70" pageNumber="71">
<emphasis id="B95A1B276200FFA52DBFB384FE6C5F43" bold="true" box="[199,395,511,537]" italics="true" pageId="70" pageNumber="71">Parena yunnana</emphasis>
Kirschenhofer
</emphasis>
</taxonomicName>
: We did not examine the
<typeStatus id="549579976200FFA52E13B384FC7C5F43" box="[875,923,511,537]" pageId="70" pageNumber="71">type</typeStatus>
of this species, but the specimens from very close to its
<typeStatus id="549579976200FFA52C6EB058FEA15F67" box="[278,326,547,573]" pageId="70" pageNumber="71">type</typeStatus>
locality that we examined are perfectly in accord with the original description and illustration of
<taxonomicName id="4C2EBCB66200FFA528F0B058FF105F3A" authorityName="Kirschenhofer" authorityYear="1994" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="yunnana">
<emphasis id="B95A1B276200FFA528F0B058FF105F3A" italics="true" pageId="70" pageNumber="71">P. yunnana</emphasis>
</taxonomicName>
. So, we confirmed that
<taxonomicName id="4C2EBCB66200FFA52F7CB03CFD985F3A" authorityName="Kirschenhofer" authorityYear="1994" box="[516,639,583,608]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="yunnana">
<emphasis id="B95A1B276200FFA52F7CB03CFD985F3A" box="[516,639,583,608]" italics="true" pageId="70" pageNumber="71">P. yunnana</emphasis>
</taxonomicName>
is identical to
<taxonomicName id="4C2EBCB66200FFA52E5DB03CFC365F3B" authorityName="Xie &amp; Yu. B" authorityYear="1993" box="[805,977,583,609]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="shapingensis">
<emphasis id="B95A1B276200FFA52E5DB03CFC365F3B" box="[805,977,583,609]" italics="true" pageId="70" pageNumber="71">P. shapingensis</emphasis>
</taxonomicName>
in external and male genital characters.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6200FFA52DBFB010FC135FAB" pageId="70" pageNumber="71" type="materials_examined">
<paragraph id="8B91C7356200FFA52DBFB010FC135FAB" blockId="70.[151,1437,150,753]" pageId="70" pageNumber="71">
<emphasis id="B95A1B276200FFA52DBFB010FDF95FDE" bold="true" box="[199,542,618,645]" pageId="70" pageNumber="71">Non-type material examined</emphasis>
.
<materialsCitation id="3B46CD686200FFA52F56B010FECA5FF3" ID-GBIF-Occurrence="4115871343" collectingDate="2009-07-24" collectingDateMax="2009-07-25" collectingDateMin="2009-07-24" location="Shangmeng" municipality="Lixian" pageId="70" pageNumber="71" specimenCount="1" specimenCount-female="1" stateProvince="Sichuan">
<specimenCount id="9D280CBC6200FFA52F56B010FD775FDF" box="[558,656,619,645]" pageId="70" pageNumber="71" type="female">1 female</specimenCount>
(HBUM), &quot;
<collectingRegion id="49EA09D76200FFA52E6FB010FC905FDF" box="[791,887,619,645]" country="China" name="Sichuan" pageId="70" pageNumber="71">Sichuan</collectingRegion>
,
<collectingMunicipality id="6BF55D4F6200FFA52EFCB010FC375FDF" box="[900,976,619,645]" pageId="70" pageNumber="71">Lixian</collectingMunicipality>
,
<location id="8EF191EE6200FFA52EA4B010FB845FDF" LSID="urn:lsid:plazi:treatment:038776236203FFA42DEFB367FC055C8B:8EF191EE6200FFA52EA4B010FB845FDF" box="[988,1123,619,645]" municipality="Lixian" name="Shangmeng" pageId="70" pageNumber="71" stateProvince="Sichuan">Shangmeng</location>
,
<date id="FF90E1F56200FFA52908B010FAC25FDF" box="[1136,1317,618,645]" pageId="70" pageNumber="71" value="2009-07-24" valueMax="2009-07-25" valueMin="2009-07-24">
<collectingDate id="EFD4181D6200FFA52908B010FAC25FDF" box="[1136,1317,618,645]" pageId="70" pageNumber="71" value="2009-07-24" valueMax="2009-07-25" valueMin="2009-07-24">2009.VII.24–25</collectingDate>
</date>
, Gao ZH., Niu YP. lgt.&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD686200FFA52C43B0F4FA965FF3" ID-GBIF-Occurrence="4115871416" box="[315,1393,655,681]" collectingDate="1979-08-12" collectorName="Zou Huanguang" location="Yulongshan" municipality="Lijiang" pageId="70" pageNumber="71" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="9D280CBC6200FFA52C43B0F4FE6F5FF3" box="[315,392,655,681]" pageId="70" pageNumber="71" type="male">1 male</specimenCount>
(IZAS), &quot;
<collectingRegion id="49EA09D76200FFA52C82B0F4FDB15FF3" box="[506,598,655,681]" country="China" name="Yunnan" pageId="70" pageNumber="71">Yunnan</collectingRegion>
,
<collectingMunicipality id="6BF55D4F6200FFA52F1CB0F4FD5F5FF3" box="[612,696,655,681]" pageId="70" pageNumber="71">Lijiang</collectingMunicipality>
,
<location id="8EF191EE6200FFA52FBCB0F4FCAC5FF3" LSID="urn:lsid:plazi:treatment:038776236203FFA42DEFB367FC055C8B:8EF191EE6200FFA52FBCB0F4FCAC5FF3" box="[708,843,655,681]" municipality="Lijiang" name="Yulongshan" pageId="70" pageNumber="71" stateProvince="Yunnan">Yulongshan</location>
,
<date id="FF90E1F56200FFA52E21B0F4FC0A5FF3" box="[857,1005,655,681]" pageId="70" pageNumber="71" value="1979-08-12">
<collectingDate id="EFD4181D6200FFA52E21B0F4FC0A5FF3" box="[857,1005,655,681]" pageId="70" pageNumber="71" value="1979-08-12">1979.VIII.12</collectingDate>
</date>
,
<collectorName id="26DBA2E36200FFA52E83B0F4FB545FF3" box="[1019,1203,655,681]" pageId="70" pageNumber="71">Zou Huanguang</collectorName>
leg.&quot; &lt;
<figureCitation id="1315DBB06200FFA52872B0F4FA965FF3" box="[1290,1393,655,681]" captionStart="FIGURE 40" captionStartId="69.[151,250,1854,1878]" captionTargetBox="[155,1433,502,1823]" captionTargetId="figure-95@69.[151,1436,498,1831]" captionTargetPageId="69" captionText="FIGURE 40. Habitus of Parena (Bothynoptera) shapingensis Xie &amp; Yu and Parena (Bothynoptera) monostigma (Bates). A Holotype of Parena shapingensis Xie &amp; Yu. B P. shapingensis Xie &amp; Yu, male from Yunnan. C Lectotype of Parena monostigma (Bates). D Holotype of Parena japoincia Jedlička [=Parena monostigma (Bates)]. E Holotype of Parena koreana Kirschenhofer [=Parena monostigma (Bates)]. F P. monostigma (Bates), male from Zhejiang. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959019" httpUri="https://zenodo.org/record/7959019/files/figure.png" pageId="70" pageNumber="71">Fig. 40B</figureCitation>
</materialsCitation>
&gt;.
<materialsCitation id="3B46CD686200FFA528F6B0F4FB065F97" ID-GBIF-Occurrence="4115871524" collectorName="S. Becvar" country="China" elevation="1800" latitude="26.53" location="Lijiang" longitude="100.18" pageId="70" pageNumber="71" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="9D280CBC6200FFA528F6B0F4FF2A5F97" pageId="70" pageNumber="71" type="male">1 male</specimenCount>
(CRS), &quot;
<collectingCountry id="F33987A56200FFA52C4EB0C8FE9A5F97" box="[310,381,691,717]" name="China" pageId="70" pageNumber="71">China</collectingCountry>
<collectingRegion id="49EA09D76200FFA52CFCB0C8FE3A5F97" box="[388,477,691,717]" country="China" name="Yunnan" pageId="70" pageNumber="71">Yunnan</collectingRegion>
,
<quantity id="4CD66AD06200FFA52C92B0C8FDA45F97" box="[490,579,691,717]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="70" pageNumber="71" unit="m" value="1800.0">
<elevation id="000320066200FFA52C92B0C8FDA45F97" box="[490,579,691,717]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="70" pageNumber="71" unit="m" value="1800.0">1800 m</elevation>
</quantity>
,
<location id="8EF191EE6200FFA52F29B0C8FD455F97" LSID="urn:lsid:plazi:treatment:038776236203FFA42DEFB367FC055C8B:8EF191EE6200FFA52F29B0C8FD455F97" box="[593,674,691,717]" country="China" latitude="100.18" name="Lijiang" pageId="70" pageNumber="71" stateProvince="Yunnan">Lijiang</location>
23.6–21.7,
<geoCoordinate id="EE1AA1F26200FFA52E53B0C9FC0F5F97" box="[811,1000,690,717]" degrees="100.18" direction="north" orientation="latitude" pageId="70" pageNumber="71" precision="555" value="100.18">26.53N 100.18E</geoCoordinate>
, lgt.
<collectorName id="26DBA2E36200FFA5295BB0C8FB765F97" box="[1059,1169,691,717]" pageId="70" pageNumber="71">S. Becvar</collectorName>
1992&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD686200FFA52996B0C8FC085FAB" ID-GBIF-Occurrence="4115871509" collectingDate="2004-06-27" country="China" location="World Horticultural Expo Garden" municipality="Kunming" pageId="70" pageNumber="71" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="9D280CBC6200FFA52996B0C8FADC5F97" box="[1262,1339,691,717]" pageId="70" pageNumber="71" type="male">1 male</specimenCount>
(IZAS), &quot;
<collectingMunicipality id="6BF55D4F6200FFA52DE7B0ACFEE85FAB" box="[159,271,727,753]" pageId="70" pageNumber="71">Kunming</collectingMunicipality>
,
<location id="8EF191EE6200FFA52C62B0ACFD725FAB" LSID="urn:lsid:plazi:treatment:038776236203FFA42DEFB367FC055C8B:8EF191EE6200FFA52C62B0ACFD725FAB" box="[282,661,727,753]" country="China" municipality="Kunming" name="World Horticultural Expo Garden" pageId="70" pageNumber="71" stateProvince="Yunnan">World Horticultural Expo Garden</location>
,
<date id="FF90E1F56200FFA52FD8B0ACFCF45FAB" box="[672,787,727,753]" pageId="70" pageNumber="71" value="2004-06-27">
<collectingDate id="EFD4181D6200FFA52FD8B0ACFCF45FAB" box="[672,787,727,753]" pageId="70" pageNumber="71" value="2004-06-27">2004.6.27</collectingDate>
</date>
, Ou Xiaohong lgt.&quot;
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF5197BD6200FFA52DEFB408FCDE5BF5" ID-DOI="http://doi.org/10.5281/zenodo.7959021" ID-Zenodo-Dep="7959021" httpUri="https://zenodo.org/record/7959021/files/figure.png" pageId="70" pageNumber="71" startId="70.[151,250,1651,1675]" targetBox="[162,1420,826,1618]" targetPageId="70" targetType="figure">
<paragraph id="8B91C7356200FFA52DEFB408FCDE5BF5" blockId="70.[151,1436,1651,1711]" pageId="70" pageNumber="71">
<emphasis id="B95A1B276200FFA52DEFB408FEFC5BD1" bold="true" box="[151,283,1651,1675]" pageId="70" pageNumber="71">FIGURE 41</emphasis>
. Median lobe of male genitalia for
<taxonomicName id="4C2EBCB66200FFA52FF7B40FFB915BD1" authority="Xie &amp; Yu" authorityName="Xie &amp; Yu. B" authorityYear="1993" box="[655,1142,1651,1675]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="shapingensis" subGenus="Bothynoptera">
<emphasis id="B95A1B276200FFA52FF7B40FFD3E5BD1" box="[655,729,1652,1675]" italics="true" pageId="70" pageNumber="71">Parena</emphasis>
(
<emphasis id="B95A1B276200FFA52F92B408FC935BD1" box="[746,884,1651,1675]" italics="true" pageId="70" pageNumber="71">Bothynoptera</emphasis>
)
<emphasis id="B95A1B276200FFA52EFCB408FBE05BD1" box="[900,1031,1651,1675]" italics="true" pageId="70" pageNumber="71">shapingensis</emphasis>
Xie &amp; Yu
</taxonomicName>
, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, holotype. Scale bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C33494BE6200FFA52DBFB4A6FE1B5A39" pageId="70" pageNumber="71" type="diagnosis">
<paragraph id="8B91C7356200FFA52DBFB4A6FE1B5A39" blockId="70.[151,1437,1757,2036]" pageId="70" pageNumber="71">
<emphasis id="B95A1B276200FFA52DBFB4A6FE815BAD" bold="true" box="[199,358,1757,1783]" pageId="70" pageNumber="71">Comparisons</emphasis>
. This species is easily distinguished from all other species of subgenus
<taxonomicName id="4C2EBCB66200FFA52900B4A6FAF75BAD" authority="Schaum, 1863" box="[1144,1296,1757,1783]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B276200FFA52900B4A6FAF75BAD" box="[1144,1296,1757,1783]" italics="true" pageId="70" pageNumber="71">Bothynoptera</emphasis>
</taxonomicName>
by the small size and elytra light yellowish brown with an indistinct large pale patch. The male genitalia of this species are very similar to those of
<taxonomicName id="4C2EBCB66200FFA52C15B55DFDE25A65" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[365,517,1830,1855]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="tripunctata">
<emphasis id="B95A1B276200FFA52C15B55DFDE25A65" box="[365,517,1830,1855]" italics="true" pageId="70" pageNumber="71">P. tripunctata</emphasis>
</taxonomicName>
, but differ in the ventral margin of median lobe, which is nearly straight, and the apical lamella not bent dorsally.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6200FFA42DBFB516FD705CF7" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71" type="description">
<paragraph id="8B91C7356200FFA42DBFB516FD705CF7" blockId="70.[151,1437,1757,2036]" lastBlockId="71.[151,1437,151,465]" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71">
<emphasis id="B95A1B276200FFA52DBFB516FEB65ADD" bold="true" box="[199,337,1901,1927]" pageId="70" pageNumber="71">Description</emphasis>
. Body length
<quantity id="4CD66AD06200FFA52C94B516FD925AD2" box="[492,629,1901,1928]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" metricValueMax="6.7" metricValueMin="6.5" pageId="70" pageNumber="71" unit="mm" value="6.6" valueMax="6.7" valueMin="6.5">6.5–6.7 mm</quantity>
. Head and pronotum yellowish red; elytra gradually graded from yellowish brown basally to pale yellow subapically, apical eighth of elytra yellowish red, forming a vague, large pale patch on the middle of elytra; legs, antennae, and mouthparts yellowish brown; venter yellowish brown. Labrum slightly dilated to apex, apex very weakly emarginate. Pronotum nearly cordate, PW/PL = 1.26–1.36, slightly narrower than head, PW/HW = 0.93–0.97; lateral explanations narrow; widest at anterior third, well rounded at anterior half, and then strongly sinuate before posterior angles; posterior angles nearly rectangular with distinct apex, slightly pointed outward. Elytra slightly dilated to apex, surface without microsculpture; striae shallowly incised, with fine puncture rows; intervals with very sparse punctures. Males with biseriate adhesive setae near apex of mesotarsomere 2 and apical two-thirds of mesotarsomere 3. Median lobe of aedeagus very stout, in dorsal view apex distinctly bent to right; in lateral view ventral margin nearly straight before middle; apical lamella thick and short, LL much smaller than LW, apex rounded-truncate; in lateral view apical lamella not bent dorsally. Endophallus with squamate sac partly divided. Female ovipositor not studied.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6201FFA42DBFB3CDFC055C8B" box="[199,994,438,465]" pageId="71" pageNumber="72" type="distribution">
<paragraph id="8B91C7356201FFA42DBFB3CDFC055C8B" blockId="71.[151,1437,151,465]" box="[199,994,438,465]" pageId="71" pageNumber="72">
<emphasis id="B95A1B276201FFA42DBFB3CDFEBE5C8A" bold="true" box="[199,345,438,464]" pageId="71" pageNumber="72">Distribution</emphasis>
(
<figureCitation id="1315DBB06201FFA42C10B3CCFE545C8B" box="[360,435,439,465]" captionStart="MAP 8" captionStartId="65.[152,211,1703,1727]" captionTargetBox="[180,1408,802,1680]" captionTargetId="figure-242@65.[180,1408,802,1680]" captionText="MAP 8. Distributions for species in the Parena (Bothynoptera) tripunctata group." figureDoi="http://doi.org/10.5281/zenodo.7963675" httpUri="https://zenodo.org/record/7963675/files/figure.png" pageId="71" pageNumber="72">Map 8</figureCitation>
, magenta).
<collectingCountry id="F33987A56201FFA42F4EB3CCFD9E5C8B" box="[566,633,439,465]" name="China" pageId="71" pageNumber="72">China</collectingCountry>
(
<collectingRegion id="49EA09D76201FFA42FF0B3CCFCEE5C8B" box="[648,777,439,465]" country="China" name="Chongqing" pageId="71" pageNumber="72">Chongqing</collectingRegion>
,
<collectingRegion id="49EA09D76201FFA42E6CB3CCFC965C8B" box="[788,881,439,465]" country="China" name="Sichuan" pageId="71" pageNumber="72">Sichuan</collectingRegion>
,
<collectingRegion id="49EA09D76201FFA42E03B3CCFC315C8B" box="[891,982,439,465]" country="China" name="Yunnan" pageId="71" pageNumber="72">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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