treatments-xml/data/03/FE/72/03FE7253F514FFEBFF18FA39FA8DFCF2.xml

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<mods:title id="8C8C0DAFE6DE6FD1AD73CB2867147FD3">Guide To The Aquatic Heteroptera Of Singapore And Peninsular Malaysia. Xi. Infraorder Nepomorpha- Families Naucoridae And Aphelocheiridae</mods:title>
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<taxonomicName id="4C57B8C6F514FFE8FF18FA39FD81FA51" ID-CoL="3K4T4" authority="Montandon, 1897" authorityName="Montandon" authorityYear="1897" box="[230,636,1437,1462]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="678" phylum="Arthropoda" rank="species" species="ovatus">
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<emphasis id="B9231F57F514FFE8FF18FA39FE56FA51" bold="true" box="[230,427,1437,1462]" italics="true" pageId="13" pageNumber="678">Heleocoris ovatus</emphasis>
Montandon, 1897
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(
<figureCitation id="136CDFC0F514FFE8FE8EFA1AFE37FA31" box="[368,458,1470,1494]" captionStart="Figs" captionStartId="10.[136,180,1861,1883]" captionTargetBox="[139,775,714,1834]" captionTargetId="figure-8@10.[117,790,706,1848]" captionTargetPageId="10" captionText="Figs. 24–28. Heleocoris species, male phallotheca. 24. Heleocoris malayensis, new species, specimen from Malaysia, Terengganu, Sungai Brang. 25. Heleocoris montandoni Lundblad, specimen from Singapore, Nee Soon swamp forest. 26. Heleocoris nebulosus Montandon, specimen from Indonesia, Bali, Melangit River. 27. Heleocoris ovatus Montandon, specimen from Vietnam, Lam Dong Prov., Pongour Falls, CL 3091. 28. Heleocoris strabus Montandon, specimen from Thailand, Chiang Mai Prov., Huay Hia Creek, CL 2198." figureDoi="http://doi.org/10.5281/zenodo.5352718" httpUri="https://zenodo.org/record/5352718/files/figure.png" pageId="13" pageNumber="678">Figs. 27</figureCitation>
, 
<figureCitation id="136CDFC0F514FFE8FE29FA1AFE0EFA31" box="[471,499,1470,1494]" captionStart="Figs" captionStartId="13.[812,856,2002,2024]" captionTargetBox="[892,1329,785,1964]" captionTargetId="figure-226@13.[872,1393,776,1974]" captionTargetPageId="13" captionText="Figs. 34, 35. Heleocoris species, male left abdominal paratergites. 34. Heleocoris malayensis, new species, specimen from Malaysia, Terengganu, Sungai Brang. 35. Heleocoris ovatus Montandon, specimen from Vietnam, Lam Dong Prov., Pongour Falls." figureDoi="http://doi.org/10.5281/zenodo.5352726" httpUri="https://zenodo.org/record/5352726/files/figure.png" pageId="13" pageNumber="678">35</figureCitation>
)
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<taxonomicName id="4C57B8C6F514FFE8FF8FFA5AFDECF9F4" ID-CoL="3K4T4" authority="Montandon, 1897 b: 451" authorityName="Montandon" authorityYear="1897" box="[113,529,1533,1555]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="678" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="B9231F57F514FFE8FF8FFA5AFEDCF9F5" box="[113,289,1534,1554]" italics="true" pageId="13" pageNumber="678">Heleocoris ovatus</emphasis>
<bibRefCitation id="EFC6BEB4F514FFE8FED6FA59FDECF9F4" author="Montandon, A" box="[296,529,1533,1555]" pageId="13" pageNumber="678" pagination="435 - 454" refId="ref18667" refString="Montandon, A. L., 1897 b. Hemiptera cryptocerata. Fam. Naucoridae. - Sous. - fam. Laccocorinae. Verhhandlungen der kaiserlich-koniglichen zoologisch-botanischen Gesellschaft in Wien, 47: 435 - 454." type="journal article" year="1897">Montandon, 1897b: 451</bibRefCitation>
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<emphasis id="B9231F57F514FFE8FF8FF991FE4DF9AB" bold="true" box="[113,432,1589,1612]" pageId="13" pageNumber="678">
<emphasis id="B9231F57F514FFE8FF8FF991FE57F9AC" bold="true" box="[113,426,1589,1611]" italics="true" pageId="13" pageNumber="678">Extralimital material examined</emphasis>
.
</emphasis>
— 
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<collectingCountry id="F34083D5F514FFE8FE28F993FDBAF9AB" box="[470,583,1591,1612]" name="Vietnam" pageId="13" pageNumber="678">VIETNAM</collectingCountry>
, 
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<collectingRegion id="49930DA7F514FFE8FDACF992FD17F9AB" box="[594,746,1590,1612]" country="Vietnam" name="Nghe An" pageId="13" pageNumber="678">Nghê An Prov.</collectingRegion>
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: 
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, 
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, SW of 
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, 
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stream, 
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, water temp. 22°C., 
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<collectingDate id="EFAD1C6DF514FFE8FDE1F9D4FD70F962" box="[543,653,1648,1669]" pageId="13" pageNumber="678" value="2000-04-01">1 Apr.2000</collectingDate>
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, CL 4382, coll. J. 
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</materialsCitation>
. 
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&amp; 
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(
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, 
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)
</materialsCitation>
. 
<materialsCitation id="3B3FC918F514FFE8FD81F929FE15F91E" collectingDate="1998-05-13" collectingDateMax="1998-05-16" collectingDateMin="1998-05-13" collectionCode="T" country="Vietnam" elevation="825" latitude="11.688611" location="Pongour Falls" longLatPrecision="21" longitude="108.265274" pageId="13" pageNumber="678" specimenCount="8" specimenCount-female="3" specimenCount-male="5" stateProvince="Lam Dong">
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<collectingRegion id="49930DA7F514FFE8FD81F929FF5AF958" country="Vietnam" name="Lam Dong" pageId="13" pageNumber="678">Lam Dong Prov.</collectingRegion>
</emphasis>
: 
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, 
<specimenCount id="9D5108CCF514FFE8FEE8F90EFE84F927" box="[278,377,1706,1728]" pageId="13" pageNumber="678" type="female">3 females</specimenCount>
, 
<location id="8E88959EF514FFE8FE78F90EFDE9F927" LSID="urn:lsid:plazi:treatment:03FE7253F514FFEBFF18FA39FA8DFCF2:8E88959EF514FFE8FE78F90EFDE9F927" box="[390,532,1706,1728]" country="Vietnam" latitude="11.688611" longLatPrecision="21" longitude="108.265274" name="Pongour Falls" pageId="13" pageNumber="678" stateProvince="Lam Dong">Pongour Falls</location>
, 
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<quantity id="4CAF6EA0F514FFE8FDDFF90EFD9FF927" box="[545,610,1706,1728]" metricMagnitude="4" metricUnit="m" metricValue="4.7" pageId="13" pageNumber="678" unit="km" value="47.0">47 km</quantity>
SW of Dalat
</locationDeviation>
, 
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<elevation id="007A2476F514FFE8FF8FF962FF4FF93B" box="[113,178,1734,1756]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="13" pageNumber="678" unit="m" value="825.0">825 m</elevation>
</quantity>
, 
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, 
<geoCoordinate id="EE63A582F514FFE8FEC5F962FE45F93B" box="[315,440,1734,1756]" degrees="108" direction="east" minutes="15" orientation="longitude" pageId="13" pageNumber="678" precision="15" seconds="55" value="108.265274">108°15'55&quot;E</geoCoordinate>
, water temp. 25.5°C, 
<date id="FFE9E585F514FFE8FD71F963FF2BF91F" pageId="13" pageNumber="678" value="1998-05-13" valueMax="1998-05-16" valueMin="1998-05-13">
<collectingDate id="EFAD1C6DF514FFE8FD71F963FF2BF91F" pageId="13" pageNumber="678" value="1998-05-13" valueMax="1998-05-16" valueMin="1998-05-13">13 and 16 May 1998</collectingDate>
</date>
, CL 3091, coll. D. A. &amp; J. 
<collectionCode id="ED465B80F514FFE8FE26F940FE15F91E" box="[472,488,1764,1785]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="678">T</collectionCode>
</materialsCitation>
. 
<materialsCitation id="3B3FC918F514FFE8FE0FF947FD10F91E" box="[497,749,1763,1785]" collectionCode="JTPC, BPBM" country="Vietnam" location="Polhemus" pageId="13" pageNumber="678" specimenCount="1" stateProvince="Lam Dong">
<location id="8E88959EF514FFE8FE0FF947FDADF91E" LSID="urn:lsid:plazi:treatment:03FE7253F514FFEBFF18FA39FA8DFCF2:8E88959EF514FFE8FE0FF947FDADF91E" box="[497,592,1763,1785]" country="Vietnam" name="Polhemus" pageId="13" pageNumber="678" stateProvince="Lam Dong">Polhemus</location>
(
<collectionCode id="ED465B80F514FFE8FDA3F947FD6BF91F" box="[605,662,1763,1784]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33646" lsid="urn:lsid:biocol.org:col:33646" name="Colorado Entomological Museum (formerly John T. Polhemus collection)" pageId="13" pageNumber="678">JTPC</collectionCode>
, 
<collectionCode id="ED465B80F514FFE8FD61F940FD1BF91E" box="[671,742,1764,1785]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="13" pageNumber="678" type="Museum">BPBM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B3FC918F514FFE8FF8FF8A4FEF6F8A8" collectingDate="2001-03-27" collectionCode="T" country="Vietnam" elevation="825" latitude="11.604167" location="Di Linh" longLatPrecision="21" longitude="108.32139" municipality="Lan Hanh" pageId="13" pageNumber="678" specimenCount="1" specimenCount-female="1" stateProvince="Lam Dong">
<specimenCount id="9D5108CCF514FFE8FF8FF8A4FF37F8F1" box="[113,202,1792,1814]" pageId="13" pageNumber="678" type="female">1 female</specimenCount>
, small stream nr. 
<collectingMunicipality id="6B8C593FF514FFE8FE83F8A4FE1CF8F1" box="[381,481,1792,1814]" pageId="13" pageNumber="678">Lan Hanh</collectingMunicipality>
, 
<quantity id="4CAF6EA0F514FFE8FE10F8A4FDD3F8F1" box="[494,558,1792,1814]" metricMagnitude="4" metricUnit="m" metricValue="3.1" pageId="13" pageNumber="678" unit="km" value="31.0">31 km</quantity>
E. of 
<location id="8E88959EF514FFE8FD8EF8A4FD3CF8F1" LSID="urn:lsid:plazi:treatment:03FE7253F514FFEBFF18FA39FA8DFCF2:8E88959EF514FFE8FD8EF8A4FD3CF8F1" box="[624,705,1792,1814]" country="Vietnam" latitude="11.604167" longLatPrecision="21" longitude="108.32139" municipality="Lan Hanh" name="Di Linh" pageId="13" pageNumber="678" stateProvince="Lam Dong">Di Linh</location>
, 
<quantity id="4CAF6EA0F514FFE8FD33F8A4FF7BF8D5" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="13" pageNumber="678" unit="m" value="825.0">
<elevation id="007A2476F514FFE8FD33F8A4FF7BF8D5" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="13" pageNumber="678" unit="m" value="825.0">825 m</elevation>
</quantity>
, 
<geoCoordinate id="EE63A582F514FFE8FF6FF8B9FEFFF8D4" box="[145,258,1821,1843]" degrees="11" direction="north" minutes="36" orientation="latitude" pageId="13" pageNumber="678" precision="15" seconds="15" value="11.604167">11°36'15&quot;N</geoCoordinate>
, 
<geoCoordinate id="EE63A582F514FFE8FEF3F8B9FE77F8D5" box="[269,394,1821,1842]" degrees="108" direction="east" minutes="19" orientation="longitude" pageId="13" pageNumber="678" precision="15" seconds="17" value="108.32139">108°19'17&quot;E</geoCoordinate>
, water temp. 24°C, 
<date id="FFE9E585F514FFE8FDB5F8B9FD34F8D5" box="[587,713,1821,1842]" pageId="13" pageNumber="678" value="2001-03-27">
<collectingDate id="EFAD1C6DF514FFE8FDB5F8B9FD34F8D5" box="[587,713,1821,1842]" pageId="13" pageNumber="678" value="2001-03-27">27 Mar.2001</collectingDate>
</date>
, CL 3094, coll. J. 
<collectionCode id="ED465B80F514FFE8FF05F89EFEF6F8A8" box="[251,267,1850,1871]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="678">T</collectionCode>
</materialsCitation>
. 
<materialsCitation id="3B3FC918F514FFE8FEE9F89EFE25F8B7" box="[279,472,1850,1872]" collectionCode="USNM" country="Vietnam" location="Polhemus" pageId="13" pageNumber="678" specimenCount="1" stateProvince="Lam Dong">
<location id="8E88959EF514FFE8FEE9F89EFE8BF8B7" LSID="urn:lsid:plazi:treatment:03FE7253F514FFEBFF18FA39FA8DFCF2:8E88959EF514FFE8FEE9F89EFE8BF8B7" box="[279,374,1850,1872]" country="Vietnam" name="Polhemus" pageId="13" pageNumber="678" stateProvince="Lam Dong">Polhemus</location>
(
<collectionCode id="ED465B80F514FFE8FE78F89EFE2CF8B7" box="[390,465,1850,1872]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="678" type="Museum">USNM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B3FC918F514FFE8FE1AF89EFEDBF842" collectingDate="2001-03-08" collectionCode="T" country="Vietnam" elevation="500" latitude="14.200556" location="Ialy" longLatPrecision="21" longitude="107.81167" municipality="Ialing Rapids" pageId="13" pageNumber="678" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Kon Tum">
<collectingRegion id="49930DA7F514FFE8FE1AF89EFD84F8A8" box="[484,633,1850,1871]" country="Vietnam" name="Kon Tum" pageId="13" pageNumber="678">
<emphasis id="B9231F57F514FFE8FE1AF89EFD8EF8A8" bold="true" box="[484,627,1850,1871]" pageId="13" pageNumber="678">Kontum Prov</emphasis>
.
</collectingRegion>
: 
<specimenCount id="9D5108CCF514FFE8FD77F89EFD25F8B7" box="[649,728,1850,1872]" pageId="13" pageNumber="678" type="male">2 males</specimenCount>
, 
<specimenCount id="9D5108CCF514FFE8FD1BF89EFF43F88B" pageId="13" pageNumber="678" type="female">2 females</specimenCount>
, 
<collectingMunicipality id="6B8C593FF514FFE8FF35F8F2FEADF88B" box="[203,336,1878,1900]" pageId="13" pageNumber="678">Ialing Rapids</collectingMunicipality>
, 
<locationDeviation id="9A5ADA67F514FFE8FEA3F8F2FDC2F88B" box="[349,575,1878,1900]" location="Kontum" pageId="13" pageNumber="678">
<quantity id="4CAF6EA0F514FFE8FEA3F8F2FE60F88B" box="[349,413,1878,1900]" metricMagnitude="4" metricUnit="m" metricValue="6.4" pageId="13" pageNumber="678" unit="km" value="64.0">64 km</quantity>
SW of 
<collectingRegion id="49930DA7F514FFE8FE10F8F3FDC2F88B" box="[494,575,1879,1900]" country="Vietnam" name="Kon Tum" pageId="13" pageNumber="678">Kontum</collectingRegion>
</locationDeviation>
, 
<quantity id="4CAF6EA0F514FFE8FDB5F8F3FD82F88B" box="[587,639,1878,1900]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="678" unit="km" value="4.0">4 km</quantity>
W. of 
<location id="8E88959EF514FFE8FD39F8F2FD10F88B" LSID="urn:lsid:plazi:treatment:03FE7253F514FFEBFF18FA39FA8DFCF2:8E88959EF514FFE8FD39F8F2FD10F88B" box="[711,749,1878,1900]" country="Vietnam" latitude="14.200556" longLatPrecision="21" longitude="107.81167" municipality="Ialing Rapids" name="Ialy" pageId="13" pageNumber="678" stateProvince="Kon Tum">Ialy</location>
, 
<quantity id="4CAF6EA0F514FFE8FF8FF8D7FF4FF86E" box="[113,178,1907,1929]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="678" unit="m" value="500.0">
<elevation id="007A2476F514FFE8FF8FF8D7FF4FF86E" box="[113,178,1907,1929]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="678" unit="m" value="500.0">500 m</elevation>
</quantity>
, 
<geoCoordinate id="EE63A582F514FFE8FF42F8D7FECDF86F" box="[188,304,1907,1928]" degrees="14" direction="north" minutes="12" orientation="latitude" pageId="13" pageNumber="678" precision="15" seconds="02" value="14.200556">14°12'02&quot;N</geoCoordinate>
, 
<geoCoordinate id="EE63A582F514FFE8FEC5F8D7FE4AF86F" box="[315,439,1907,1928]" degrees="107" direction="east" minutes="48" orientation="longitude" pageId="13" pageNumber="678" precision="15" seconds="42" value="107.81167">107°48'42&quot;E</geoCoordinate>
, water temp. 24°C, 
<date id="FFE9E585F514FFE8FD85F8D7FD10F86F" box="[635,749,1907,1928]" pageId="13" pageNumber="678" value="2001-03-08">
<collectingDate id="EFAD1C6DF514FFE8FD85F8D7FD10F86F" box="[635,749,1907,1928]" pageId="13" pageNumber="678" value="2001-03-08">8 Mar.2001</collectingDate>
</date>
, CL 4285, coll. J. 
<collectionCode id="ED465B80F514FFE8FEE8F834FEDBF842" box="[278,294,1936,1957]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="678">T</collectionCode>
</materialsCitation>
. 
<materialsCitation id="3B3FC918F514FFE8FED0F834FD90F841" box="[302,621,1936,1958]" collectionCode="USNM" collectorName="P. Nguyen" country="Vietnam" location="Polhemus" pageId="13" pageNumber="678" specimenCount="1" stateProvince="Kon Tum">
<location id="8E88959EF514FFE8FED0F834FE70F841" LSID="urn:lsid:plazi:treatment:03FE7253F514FFEBFF18FA39FA8DFCF2:8E88959EF514FFE8FED0F834FE70F841" box="[302,397,1936,1958]" country="Vietnam" name="Polhemus" pageId="13" pageNumber="678" stateProvince="Kon Tum">Polhemus</location>
&amp; 
<collectorName id="26A2A693F514FFE8FE52F834FDF3F842" box="[428,526,1936,1957]" pageId="13" pageNumber="678">P. Nguyen</collectorName>
(
<collectionCode id="ED465B80F514FFE8FDE5F834FD9BF841" box="[539,614,1936,1958]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="678" type="Museum">USNM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B3FC918F514FFE8FD88F834FD96F81B" collectingDate="2001-03-18" collectionCode="T" collectorName="D. A. Polhemus" country="Vietnam" elevation="24120" latitude="14.719444" location="Hwy." longLatPrecision="21" longitude="108.59667" municipality="Nuoc Xi" pageId="13" pageNumber="678" specimenCount="20" specimenCount-female="10" specimenCount-male="10" stateProvince="Quang Ngai">
<collectingRegion id="49930DA7F514FFE8FD88F834FF5BF825" country="Vietnam" name="Quang Ngai" pageId="13" pageNumber="678">
<emphasis id="B9231F57F514FFE8FD88F834FF5CF825" bold="true" pageId="13" pageNumber="678">Quang Ngai Prov</emphasis>
.
</collectingRegion>
, 
<specimenCount id="9D5108CCF514FFE8FF4CF809FEFAF824" box="[178,263,1965,1987]" pageId="13" pageNumber="678" type="male">10 males</specimenCount>
, 
<specimenCount id="9D5108CCF514FFE8FEF1F809FE8AF824" box="[271,375,1965,1987]" pageId="13" pageNumber="678" type="female">10 females</specimenCount>
, 
<collectingMunicipality id="6B8C593FF514FFE8FE81F809FE32F824" box="[383,463,1965,1987]" pageId="13" pageNumber="678">Nuoc Xi</collectingMunicipality>
stream, 
<locationDeviation id="9A5ADA67F514FFE8FDE2F809FD0CF825" box="[540,753,1965,1987]" location="Kontum" pageId="13" pageNumber="678">
<quantity id="4CAF6EA0F514FFE8FDE2F809FD9FF824" box="[540,610,1965,1987]" metricMagnitude="5" metricUnit="m" metricValue="1.23" pageId="13" pageNumber="678" unit="km" value="123.0">123 km</quantity>
NE of 
<collectingRegion id="49930DA7F514FFE8FD5DF809FD0CF825" box="[675,753,1965,1986]" country="Vietnam" name="Kon Tum" pageId="13" pageNumber="678">Kontum</collectingRegion>
</locationDeviation>
on 
<location id="8E88959EF514FFE8FF6EF86EFF38F838" LSID="urn:lsid:plazi:treatment:03FE7253F514FFEBFF18FA39FA8DFCF2:8E88959EF514FFE8FF6EF86EFF38F838" box="[144,197,1994,2015]" country="Vietnam" latitude="14.719444" longLatPrecision="21" longitude="108.59667" municipality="Nuoc Xi" name="Hwy." pageId="13" pageNumber="678" stateProvince="Quang Ngai">Hwy.</location>
<quantity id="4CAF6EA0F514FFE8FF35F86EFECDF838" box="[203,304,1994,2015]" metricMagnitude="4" metricUnit="m" metricValue="2.412" pageId="13" pageNumber="678" unit="m" value="24120.0">
<elevation id="007A2476F514FFE8FF35F86EFECDF838" box="[203,304,1994,2015]" metricMagnitude="4" metricUnit="m" metricValue="2.412" pageId="13" pageNumber="678" unit="m" value="24120.0">24, 120 m</elevation>
</quantity>
, 
<geoCoordinate id="EE63A582F514FFE8FEC4F86EFE50F838" box="[314,429,1994,2015]" degrees="14" direction="north" minutes="43" orientation="latitude" pageId="13" pageNumber="678" precision="15" seconds="10" value="14.719444">14°43'10&quot;N</geoCoordinate>
, 
<geoCoordinate id="EE63A582F514FFE8FE46F86DFDC9F838" box="[440,564,1993,2015]" degrees="108" direction="east" minutes="35" orientation="longitude" pageId="13" pageNumber="678" precision="15" seconds="48" value="108.59667">108°35'48&quot;E</geoCoordinate>
, water temp. 26°C, 
<date id="FFE9E585F514FFE8FF8FF843FF0CF81B" box="[113,241,2023,2044]" pageId="13" pageNumber="678" value="2001-03-18">
<collectingDate id="EFAD1C6DF514FFE8FF8FF843FF0CF81B" box="[113,241,2023,2044]" pageId="13" pageNumber="678" value="2001-03-18">18 Mar.2001</collectingDate>
</date>
, CL 4299, coll. 
<collectorName id="26A2A693F514FFE8FE68F843FDCBF81B" box="[406,566,2022,2044]" pageId="13" pageNumber="678">D. A. Polhemus</collectorName>
, J. 
<collectionCode id="ED465B80F514FFE8FDA5F843FD96F81B" box="[603,619,2023,2044]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="678">T</collectionCode>
</materialsCitation>
. 
<materialsCitation id="3B3FC918F514FFE8FD89F842FEC5F7FE" collectionCode="USNM" collectorName="P. Nguyen" country="Vietnam" location="Polhemus" pageId="13" pageNumber="678" specimenCount="1" stateProvince="Quang Ngai">
<location id="8E88959EF514FFE8FD89F842FD2BF81B" LSID="urn:lsid:plazi:treatment:03FE7253F514FFEBFF18FA39FA8DFCF2:8E88959EF514FFE8FD89F842FD2BF81B" box="[631,726,2022,2044]" country="Vietnam" name="Polhemus" pageId="13" pageNumber="678" stateProvince="Quang Ngai">Polhemus</location>
&amp; 
<collectorName id="26A2A693F514FFE8FF8FF7A0FF2AF7FE" box="[113,215,2052,2073]" pageId="13" pageNumber="678">P. Nguyen</collectorName>
(
<collectionCode id="ED465B80F514FFE8FF18F7A7FECCF7FE" box="[230,305,2051,2073]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="678" type="Museum">USNM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B3FC918F514FFE8FEBDF7A7FD0CF7D2" collectorName="An Khe Pass &amp; An Khe" country="Vietnam" location="Binh Dinh Prov." pageId="13" pageNumber="678" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Binh Dinh">
<collectingRegion id="49930DA7F514FFE8FEBDF7A7FE12F7FE" box="[323,495,2051,2073]" country="Vietnam" name="Binh Dinh" pageId="13" pageNumber="678">
<emphasis id="B9231F57F514FFE8FEBDF7A7FE17F7FF" bold="true" box="[323,490,2051,2073]" pageId="13" pageNumber="678">Binh Dinh Prov</emphasis>
.
</collectingRegion>
, 
<specimenCount id="9D5108CCF514FFE8FE00F7A7FDB9F7FE" box="[510,580,2051,2073]" pageId="13" pageNumber="678" type="male">1 male</specimenCount>
, 
<specimenCount id="9D5108CCF514FFE8FDAEF7A7FD54F7FE" box="[592,681,2051,2073]" pageId="13" pageNumber="678" type="female">1 female</specimenCount>
, spring fed stream on E. side of 
<collectorName id="26A2A693F514FFE8FE98F784FE15F7D2" box="[358,488,2080,2102]" pageId="13" pageNumber="678">An Khe Pass</collectorName>
, 
<quantity id="4CAF6EA0F514FFE8FE0AF784FDB8F7D1" box="[500,581,2080,2102]" metricMagnitude="4" metricUnit="m" metricValue="1.55" pageId="13" pageNumber="678" unit="km" value="15.5">15.5 km</quantity>
E. of 
<collectorName id="26A2A693F514FFE8FD7AF784FD2CF7D1" box="[644,721,2080,2102]" pageId="13" pageNumber="678">An Khe</collectorName>
on
</materialsCitation>
</paragraph>
<paragraph id="8BE8C345F514FFE8FADBFB0EFAB2FB2C" blockId="13.[1317,1359,1194,1227]" box="[1317,1359,1194,1227]" pageId="13" pageNumber="678">34</paragraph>
<caption id="DF2893CDF514FFE8FCD2F876FAA3F7D9" ID-DOI="http://doi.org/10.5281/zenodo.5352726" ID-Zenodo-Dep="5352726" httpUri="https://zenodo.org/record/5352726/files/figure.png" pageId="13" pageNumber="678" startId="13.[812,856,2002,2024]" targetBox="[892,1329,785,1964]" targetPageId="13">
<paragraph id="8BE8C345F514FFE8FCD2F876FAA3F7D9" blockId="13.[812,1453,2001,2110]" pageId="13" pageNumber="678">
Figs. 34, 35. 
<taxonomicName id="4C57B8C6F514FFE8FC55F877FB9EF80F" box="[939,1123,2002,2024]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="678" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9231F57F514FFE8FC55F877FBE9F800" box="[939,1044,2003,2023]" italics="true" pageId="13" pageNumber="678">Heleocoris</emphasis>
species
</taxonomicName>
, male left abdominal paratergites. 34. 
<taxonomicName id="4C57B8C6F514FFE8FCAEF854FBD6F7E3" authority="Polhemus &amp; Polhemus, 2013" authorityName="Polhemus &amp; Polhemus" authorityYear="2013" box="[848,1067,2032,2052]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="678" phylum="Arthropoda" rank="species" species="malayensis" status="sp. nov.">
<emphasis id="B9231F57F514FFE8FCAEF854FBD6F7E3" box="[848,1067,2032,2052]" italics="true" pageId="13" pageNumber="678">Heleocoris malayensis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A210A22CF514FFE8FBC9F84BFB50F7E3" box="[1079,1197,2030,2052]" pageId="13" pageNumber="678" rank="species">new species</taxonomicNameLabel>
, specimen from Malaysia, Terengganu, Sungai Brang. 35. 
<taxonomicName id="4C57B8C6F514FFE8FB8DF7A8FA5AF7C6" authority="Montandon" authorityName="Montandon" authorityYear="1897" box="[1139,1447,2059,2081]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="678" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="B9231F57F514FFE8FB8DF7A8FAD4F7C7" box="[1139,1321,2060,2080]" italics="true" pageId="13" pageNumber="678">Heleocoris ovatus</emphasis>
Montandon
</taxonomicName>
, specimen from Vietnam, Lam Dong Prov., Pongour Falls.
</paragraph>
</caption>
<paragraph id="8BE8C345F517FFEBFF76FF1AFDF7FE11" blockId="14.[136,776,189,502]" pageId="14" pageNumber="679">
apex dark yellow; hemelytra dark blackish brown with tiny pale asperities, outer half of embolium dark yellow, wing membrane in submacropterous forms poorly defined, dark brown; abdominal laterotergites dark yellow, extreme posterior margins dark brown. This species may recognised by its moderate size for the genus, the predominantly dark hemelytra which lack yellow markings except on the outer half of the corium, the projecting tab on the lateral margin of male abdominal left paratergite 
<collectionCode id="ED465B80F517FFEBFE30FE1AFE1CFE32" box="[462,481,446,469]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="14" pageNumber="679" type="Museum">V</collectionCode>
(
<figureCitation id="136CDFC0F517FFEBFE10FE19FDC7FE32" box="[494,570,445,469]" captionStart="Figs" captionStartId="13.[812,856,2002,2024]" captionTargetBox="[892,1329,785,1964]" captionTargetId="figure-226@13.[872,1393,776,1974]" captionTargetPageId="13" captionText="Figs. 34, 35. Heleocoris species, male left abdominal paratergites. 34. Heleocoris malayensis, new species, specimen from Malaysia, Terengganu, Sungai Brang. 35. Heleocoris ovatus Montandon, specimen from Vietnam, Lam Dong Prov., Pongour Falls." figureDoi="http://doi.org/10.5281/zenodo.5352726" httpUri="https://zenodo.org/record/5352726/files/figure.png" pageId="14" pageNumber="679">Fig. 35</figureCitation>
), and the distinctive male genitalic strucutures (
<figureCitation id="136CDFC0F517FFEBFE53FE79FE03FE11" box="[429,510,477,502]" captionStart="Figs" captionStartId="10.[136,180,1861,1883]" captionTargetBox="[139,775,714,1834]" captionTargetId="figure-8@10.[117,790,706,1848]" captionTargetPageId="10" captionText="Figs. 24–28. Heleocoris species, male phallotheca. 24. Heleocoris malayensis, new species, specimen from Malaysia, Terengganu, Sungai Brang. 25. Heleocoris montandoni Lundblad, specimen from Singapore, Nee Soon swamp forest. 26. Heleocoris nebulosus Montandon, specimen from Indonesia, Bali, Melangit River. 27. Heleocoris ovatus Montandon, specimen from Vietnam, Lam Dong Prov., Pongour Falls, CL 3091. 28. Heleocoris strabus Montandon, specimen from Thailand, Chiang Mai Prov., Huay Hia Creek, CL 2198." figureDoi="http://doi.org/10.5281/zenodo.5352718" httpUri="https://zenodo.org/record/5352718/files/figure.png" pageId="14" pageNumber="679">Fig. 27</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34D90CEF517FFEBFF76FDB8FE1DFB51" pageId="14" pageNumber="679" type="distribution">
<paragraph id="8BE8C345F517FFEBFF76FDB8FE1DFB51" blockId="14.[136,776,540,1206]" pageId="14" pageNumber="679">
<emphasis id="B9231F57F517FFEBFF76FDB8FEEBFDD1" bold="true" box="[136,278,540,566]" pageId="14" pageNumber="679">
<emphasis id="B9231F57F517FFEBFF76FDB8FEF2FDD2" bold="true" box="[136,271,540,565]" italics="true" pageId="14" pageNumber="679">Distribution</emphasis>
.
</emphasis>
— 
<bibRefCitation id="EFC6BEB4F517FFEBFEB8FDB9FDDDFDD1" author="Montandon, A" box="[326,544,541,566]" pageId="14" pageNumber="679" pagination="365 - 377" refId="ref18620" refString="Montandon, A. L., 1897 a. Viaggio di Leonardo Fea in Birmania e regioni vicine. LXXV. Hemiptera Cryptocerata. Annali del Museo Civico di Storia Naturale di Genova, Serie 2, 17 (37): 365 - 377." type="journal article" year="1897">Montandon (1897a)</bibRefCitation>
described 
<taxonomicName id="4C57B8C6F517FFEBFD63FDBBFCF5FDD1" authorityName="Montandon" authorityYear="1897" box="[669,776,542,566]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="B9231F57F517FFEBFD63FDBBFD4DFDD1" box="[669,688,543,566]" italics="true" pageId="14" pageNumber="679">H</emphasis>
. 
<emphasis id="B9231F57F517FFEBFD3FFDBBFCF5FDD1" box="[705,776,543,566]" italics="true" pageId="14" pageNumber="679">ovatus</emphasis>
</taxonomicName>
from a single specimen taken in the vicinity of Lakhon, in northern 
<collectingCountry id="F34083D5F517FFEBFF14FDFAFEDDFD92" box="[234,288,606,629]" name="Laos" pageId="14" pageNumber="679">Laos</collectingCountry>
, by François Jules Harmand, a French doctor who from 1877–1878 made a traverse of the country lying between Lakhon, on the Mekong River, and 
<collectingRegion id="49930DA7F517FFEBFD8AFD3AFD18FD52" box="[628,741,669,694]" country="Vietnam" name="Quang Tri" pageId="14" pageNumber="679">Quang Tri</collectingRegion>
, in modern 
<collectingCountry id="F34083D5F517FFEBFF1EFD19FEC3FD32" box="[224,318,701,725]" name="Vietnam" pageId="14" pageNumber="679">Vietnam</collectingCountry>
. Montandon did not indicate the sex of his Laotian 
<typeStatus id="54EC7DE7F517FFEBFF1CFD79FEBFFD12" box="[226,322,733,757]" pageId="14" pageNumber="679" type="holotype">holotype</typeStatus>
, and although this specimen was recently re-examined and discussed by Sites &amp; Vitheepradit (2011), these latter authors did not indicate the sex in their publication either. We have contacted Eric Guilbert of the Muséum National d’Histoire Naturelle in Paris, where the 
<typeStatus id="54EC7DE7F517FFEBFD54FCF9FCFAFC92" box="[682,775,861,885]" pageId="14" pageNumber="679" type="holotype">holotype</typeStatus>
is held, who has confirmed that this specimen is a female, and has provided useful notes on its morphology. As noted by Sites &amp; Vitheepradit (2011), 
<taxonomicName id="4C57B8C6F517FFEBFE06FC1BFD9EFC31" authorityName="Montandon" authorityYear="1897" box="[504,611,958,982]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="B9231F57F517FFEBFE06FC1BFDF6FC31" box="[504,523,959,982]" italics="true" pageId="14" pageNumber="679">H</emphasis>
. 
<emphasis id="B9231F57F517FFEBFDE2FC1BFD9EFC31" box="[540,611,959,982]" italics="true" pageId="14" pageNumber="679">ovatus</emphasis>
</taxonomicName>
is widespread in Indochina, with records provided by these authors for 
<collectingCountry id="F34083D5F517FFEBFF76FC59FF35FBF2" box="[136,200,1021,1045]" name="China" pageId="14" pageNumber="679">China</collectingCountry>
(
<collectingRegion id="49930DA7F517FFEBFF24FC59FED0FBF2" box="[218,301,1021,1045]" country="China" name="Hainan" pageId="14" pageNumber="679">Hainan</collectingRegion>
), 
<collectingCountry id="F34083D5F517FFEBFEBCFC5AFE73FBF2" box="[322,398,1022,1045]" name="Myanmar" pageId="14" pageNumber="679">Burma</collectingCountry>
, 
<collectingCountry id="F34083D5F517FFEBFE65FC59FE00FBF2" box="[411,509,1021,1045]" name="Thailand" pageId="14" pageNumber="679">Thailand</collectingCountry>
, and 
<collectingCountry id="F34083D5F517FFEBFDC2FC59FD67FBF2" box="[572,666,1021,1045]" name="Vietnam" pageId="14" pageNumber="679">Vietnam</collectingCountry>
, to which we can now add a new record from 
<collectingCountry id="F34083D5F517FFEBFDD8FBBAFD55FBD1" box="[550,680,1054,1078]" name="Hong Kong" pageId="14" pageNumber="679">Hong Kong</collectingCountry>
as well. As explained below, records of 
<taxonomicName id="4C57B8C6F517FFEBFE25FB9BFDBFFBB1" authorityName="Montandon" authorityYear="1897" box="[475,578,1087,1110]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="B9231F57F517FFEBFE25FB9BFDBFFBB1" box="[475,578,1087,1110]" italics="true" pageId="14" pageNumber="679">H. ovatus</emphasis>
</taxonomicName>
for 
<collectingCountry id="F34083D5F517FFEBFD91FB99FD2CFBB2" box="[623,721,1085,1109]" name="Malaysia" pageId="14" pageNumber="679">Malaysia</collectingCountry>
(and perhaps elsewhere in Indochina) listed by these latter authors are misidentifications, and refer instead to the new species 
<taxonomicName id="4C57B8C6F517FFEBFF76FB3BFEDCFB51" authorityName="Polhemus &amp; Polhemus" authorityYear="2013" box="[136,289,1182,1206]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="B9231F57F517FFEBFF76FB3BFF66FB51" box="[136,155,1183,1206]" italics="true" pageId="14" pageNumber="679">H</emphasis>
. 
<emphasis id="B9231F57F517FFEBFF54FB3BFEDCFB51" box="[170,289,1183,1206]" italics="true" pageId="14" pageNumber="679">malayensis</emphasis>
</taxonomicName>
described herein.
</paragraph>
</subSubSection>
<subSubSection id="C34D90CEF517FFEBFF76FB79FA8DFCF2" pageId="14" pageNumber="679" type="discussion">
<paragraph id="8BE8C345F517FFEBFF76FB79FD50F8F2" blockId="14.[136,776,1245,1814]" pageId="14" pageNumber="679">
<emphasis id="B9231F57F517FFEBFF76FB79FEFDFB12" bold="true" box="[136,256,1245,1269]" italics="true" pageId="14" pageNumber="679">Discussion</emphasis>
. — Males of 
<taxonomicName id="4C57B8C6F517FFEBFE68FB7BFE00FB11" authorityName="Montandon" authorityYear="1897" box="[406,509,1246,1270]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="B9231F57F517FFEBFE68FB7BFE54FB11" box="[406,425,1247,1270]" italics="true" pageId="14" pageNumber="679">H</emphasis>
. 
<emphasis id="B9231F57F517FFEBFE49FB7BFE00FB11" box="[439,509,1247,1270]" italics="true" pageId="14" pageNumber="679">ovatus</emphasis>
</taxonomicName>
are easily recognised by the blunt, anteriorly projecting tab on the lateral margin of male left abdominal paratergite VI (
<figureCitation id="136CDFC0F517FFEBFDF1FAB9FDA2FAD2" box="[527,607,1309,1333]" captionStart="Figs" captionStartId="13.[812,856,2002,2024]" captionTargetBox="[892,1329,785,1964]" captionTargetId="figure-226@13.[872,1393,776,1974]" captionTargetPageId="13" captionText="Figs. 34, 35. Heleocoris species, male left abdominal paratergites. 34. Heleocoris malayensis, new species, specimen from Malaysia, Terengganu, Sungai Brang. 35. Heleocoris ovatus Montandon, specimen from Vietnam, Lam Dong Prov., Pongour Falls." figureDoi="http://doi.org/10.5281/zenodo.5352726" httpUri="https://zenodo.org/record/5352726/files/figure.png" pageId="14" pageNumber="679">Fig. 35</figureCitation>
), which is very different in form from the hooked projection on the left abdominal paratergite VI in males of 
<taxonomicName id="4C57B8C6F517FFEBFDF2FAFBFD5CFA91" authorityName="Polhemus &amp; Polhemus" authorityYear="2013" box="[524,673,1374,1398]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="B9231F57F517FFEBFDF2FAFBFDE2FA91" box="[524,543,1375,1398]" italics="true" pageId="14" pageNumber="679">H</emphasis>
. 
<emphasis id="B9231F57F517FFEBFDD5FAFBFD5CFA91" box="[555,673,1375,1398]" italics="true" pageId="14" pageNumber="679">malayensis</emphasis>
</taxonomicName>
(
<figureCitation id="136CDFC0F517FFEBFD53FAF9FD01FA91" box="[685,764,1373,1398]" captionStart="Figs" captionStartId="13.[812,856,2002,2024]" captionTargetBox="[892,1329,785,1964]" captionTargetId="figure-226@13.[872,1393,776,1974]" captionTargetPageId="13" captionText="Figs. 34, 35. Heleocoris species, male left abdominal paratergites. 34. Heleocoris malayensis, new species, specimen from Malaysia, Terengganu, Sungai Brang. 35. Heleocoris ovatus Montandon, specimen from Vietnam, Lam Dong Prov., Pongour Falls." figureDoi="http://doi.org/10.5281/zenodo.5352726" httpUri="https://zenodo.org/record/5352726/files/figure.png" pageId="14" pageNumber="679">Fig. 34</figureCitation>
). No other species of 
<taxonomicName id="4C57B8C6F517FFEBFEA5FADBFE2DFA71" authorityName="Stal" authorityYear="1876" box="[347,464,1407,1430]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="genus">
<emphasis id="B9231F57F517FFEBFEA5FADBFE2DFA71" box="[347,464,1407,1430]" italics="true" pageId="14" pageNumber="679">Heleocoris</emphasis>
</taxonomicName>
so far known from Southeast Asia possess similar projections on male left paratergite VI. The male phallotheca also differs in shape between the two species, with the tip being more produced and evenly rounded in 
<taxonomicName id="4C57B8C6F517FFEBFEFAFA5BFE60F9F1" authorityName="Polhemus &amp; Polhemus" authorityYear="2013" box="[260,413,1534,1558]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="B9231F57F517FFEBFEFAFA5BFEEAF9F1" box="[260,279,1535,1558]" italics="true" pageId="14" pageNumber="679">H</emphasis>
. 
<emphasis id="B9231F57F517FFEBFED8FA5BFE60F9F1" box="[294,413,1535,1558]" italics="true" pageId="14" pageNumber="679">malayensis</emphasis>
</taxonomicName>
, and the distal internal sclerite of a different shape (compare 
<figureCitation id="136CDFC0F517FFEBFE55F9B9FDAEF9D1" box="[427,595,1565,1590]" captionStart="Figs" captionStartId="10.[136,180,1861,1883]" captionTargetBox="[139,775,714,1834]" captionTargetId="figure-8@10.[117,790,706,1848]" captionTargetPageId="10" captionText="Figs. 24–28. Heleocoris species, male phallotheca. 24. Heleocoris malayensis, new species, specimen from Malaysia, Terengganu, Sungai Brang. 25. Heleocoris montandoni Lundblad, specimen from Singapore, Nee Soon swamp forest. 26. Heleocoris nebulosus Montandon, specimen from Indonesia, Bali, Melangit River. 27. Heleocoris ovatus Montandon, specimen from Vietnam, Lam Dong Prov., Pongour Falls, CL 3091. 28. Heleocoris strabus Montandon, specimen from Thailand, Chiang Mai Prov., Huay Hia Creek, CL 2198." figureDoi="http://doi.org/10.5281/zenodo.5352718" httpUri="https://zenodo.org/record/5352718/files/figure.png" pageId="14" pageNumber="679">Figs. 24 and 27</figureCitation>
). Females are by contrast far more similar, with the posterior margin of the subgenital plate being broadly concave in 
<taxonomicName id="4C57B8C6F517FFEBFDC2F9FBFD2DF991" authorityName="Polhemus &amp; Polhemus" authorityYear="2013" box="[572,720,1631,1654]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="B9231F57F517FFEBFDC2F9FBFD2DF991" box="[572,720,1631,1654]" italics="true" pageId="14" pageNumber="679">H. malayensis</emphasis>
</taxonomicName>
(
<figureCitation id="136CDFC0F517FFEBFD23F9F9FF5BF971" captionStart="Figs" captionStartId="12.[835,879,1091,1113]" captionTargetBox="[834,1469,233,1050]" captionTargetId="figure-8@12.[819,1486,216,1072]" captionTargetPageId="12" captionText="Figs. 31, 32. Heleocoris species, female terminal abdomen, ventral view, showing shape of subgenital plate. 31. Heleocoris montandoni Lundblad, specimen from Vietnam, Lam Dong Prov., CL 3094. 32. Heleocoris malayensis, new species, specimen from Malaysia, Perak, Kerunai River, CL 2078." figureDoi="http://doi.org/10.5281/zenodo.5352722" httpUri="https://zenodo.org/record/5352722/files/figure.png" pageId="14" pageNumber="679">Fig. 32</figureCitation>
), rather than bearing a broad, V-shaped incision as in 
<taxonomicName id="4C57B8C6F517FFEBFD10F9DBFF33F951" authorityName="Montandon" authorityYear="1897" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="B9231F57F517FFEBFD10F9DBFF33F951" italics="true" pageId="14" pageNumber="679">H. ovatus</emphasis>
</taxonomicName>
; the latter character state has been confirmed on the basis of an examination of the 
<typeStatus id="54EC7DE7F517FFEBFE29F919FDC9F932" box="[471,564,1725,1749]" pageId="14" pageNumber="679" type="holotype">holotype</typeStatus>
female by Guilbert in relation to illustrations of the subgenital plates of both 
<taxonomicName id="4C57B8C6F517FFEBFD10F97BFF33F8F1" authorityName="Montandon" authorityYear="1897" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="B9231F57F517FFEBFD10F97BFF33F8F1" italics="true" pageId="14" pageNumber="679">H. ovatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C57B8C6F517FFEBFEFAF95BFE60F8F1" authorityName="Polhemus &amp; Polhemus" authorityYear="2013" box="[260,413,1791,1814]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="B9231F57F517FFEBFEFAF95BFE60F8F1" box="[260,413,1791,1814]" italics="true" pageId="14" pageNumber="679">H. malayensis</emphasis>
</taxonomicName>
provided by the authors.
</paragraph>
<paragraph id="8BE8C345F517FFEBFF76F899FA8DFCF2" blockId="14.[136,776,1853,2102]" lastBlockId="14.[834,1475,189,790]" pageId="14" pageNumber="679">
Although Sites &amp; Vitheepradit (2011) provided records of 
<taxonomicName id="4C57B8C6F517FFEBFF76F8FBFF0CF891" authorityName="Montandon" authorityYear="1897" box="[136,241,1886,1910]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="B9231F57F517FFEBFF76F8FBFF66F891" box="[136,155,1887,1910]" italics="true" pageId="14" pageNumber="679">H</emphasis>
. 
<emphasis id="B9231F57F517FFEBFF55F8FBFF0CF891" box="[171,241,1887,1910]" italics="true" pageId="14" pageNumber="679">ovatus</emphasis>
</taxonomicName>
from the Peninsular Malaysian states of 
<collectingRegion id="49930DA7F517FFEBFD4CF8F9FCFEF892" box="[690,771,1885,1909]" country="Malaysia" name="Pahang" pageId="14" pageNumber="679">Pahang</collectingRegion>
, 
<collectingRegion id="49930DA7F517FFEBFF76F8DAFEE8F871" box="[136,277,1918,1942]" country="Malaysia" name="Terengganu" pageId="14" pageNumber="679">Terengganu</collectingRegion>
, and 
<collectingRegion id="49930DA7F517FFEBFEA5F8D9FE38F872" box="[347,453,1917,1941]" country="Malaysia" name="Selangor" pageId="14" pageNumber="679">Selangor</collectingRegion>
, we have re-examined the 
<collectingRegion id="49930DA7F517FFEBFF76F83AFEF8F851" box="[136,261,1950,1974]" country="Malaysia" name="Terengganu" pageId="14" pageNumber="679">Terengganu</collectingRegion>
series and determined that it is in fact composed of specimens of 
<taxonomicName id="4C57B8C6F517FFEBFEAEF81BFE0BF831" authorityName="Polhemus &amp; Polhemus" authorityYear="2013" box="[336,502,1982,2006]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="B9231F57F517FFEBFEAEF81BFE9EF831" box="[336,355,1983,2006]" italics="true" pageId="14" pageNumber="679">H</emphasis>
. 
<emphasis id="B9231F57F517FFEBFE89F81BFE0BF831" box="[375,502,1983,2006]" italics="true" pageId="14" pageNumber="679">malayensis</emphasis>
</taxonomicName>
(see 
<typeStatus id="54EC7DE7F517FFEBFDC5F81AFD60F831" box="[571,669,1982,2006]" pageId="14" pageNumber="679" type="paratype">paratype</typeStatus>
material listed under that species). Given that all other specimens of 
<taxonomicName id="4C57B8C6F517FFEBFF56F85BFEE3F7F1" authorityName="Stal" authorityYear="1876" box="[168,286,2047,2070]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="genus">
<emphasis id="B9231F57F517FFEBFF56F85BFEE3F7F1" box="[168,286,2047,2070]" italics="true" pageId="14" pageNumber="679">Heleocoris</emphasis>
</taxonomicName>
of appropriate size and colouration that we have examined from Peninsular 
<collectingCountry id="F34083D5F517FFEBFE19F7B9FDB4F7D2" box="[487,585,2077,2101]" name="Malaysia" pageId="14" pageNumber="679">Malaysia</collectingCountry>
also represent 
<taxonomicName id="4C57B8C6F517FFEBFD10F7BBFC44FF31" authorityName="Polhemus &amp; Polhemus" authorityYear="2013" box="[750,953,191,2102]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="B9231F57F517FFEBFD10F7BBFCFCF7D1" box="[750,769,2079,2102]" italics="true" pageId="14" pageNumber="679">H</emphasis>
. 
<emphasis id="B9231F57F517FFEBFCBCFF1BFC44FF31" box="[834,953,191,214]" italics="true" pageId="14" pageNumber="679">malayensis</emphasis>
</taxonomicName>
rather than 
<taxonomicName id="4C57B8C6F517FFEBFBBDFF1BFB50FF31" authorityName="Montandon" authorityYear="1897" box="[1091,1197,190,214]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="B9231F57F517FFEBFBBDFF1BFBABFF31" box="[1091,1110,191,214]" italics="true" pageId="14" pageNumber="679">H</emphasis>
. 
<emphasis id="B9231F57F517FFEBFB99FF1BFB50FF31" box="[1127,1197,191,214]" italics="true" pageId="14" pageNumber="679">ovatus</emphasis>
</taxonomicName>
, we have concluded that the Malaysian records of the latter species listed by Sites &amp; Vitheepradit (2011) are probably all misidentifications. In addition, because the above authors did not utilise male paratergite or genitalic characters in their taxonomic analysis, and therefore did not realise that two species were co-mingled under their concept of 
<taxonomicName id="4C57B8C6F517FFEBFBBCFEDBFB51FE71" authorityName="Montandon" authorityYear="1897" box="[1090,1196,382,406]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="B9231F57F517FFEBFBBCFEDBFBA8FE71" box="[1090,1109,383,406]" italics="true" pageId="14" pageNumber="679">H</emphasis>
. 
<emphasis id="B9231F57F517FFEBFB98FEDBFB51FE71" box="[1126,1196,383,406]" italics="true" pageId="14" pageNumber="679">ovatus</emphasis>
</taxonomicName>
, we consider it probable that at least some of the specimens of “ 
<taxonomicName id="4C57B8C6F517FFEBFB14FE3BFAAFFE51" authorityName="Montandon" authorityYear="1897" box="[1258,1362,414,438]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="B9231F57F517FFEBFB14FE3BFB00FE51" box="[1258,1277,415,438]" italics="true" pageId="14" pageNumber="679">H</emphasis>
. 
<emphasis id="B9231F57F517FFEBFAF2FE3BFAAFFE51" box="[1292,1362,415,438]" italics="true" pageId="14" pageNumber="679">ovatus</emphasis>
</taxonomicName>
” that they list from southern peninsular 
<collectingCountry id="F34083D5F517FFEBFB6AFE19FB08FE32" box="[1172,1269,445,469]" name="Thailand" pageId="14" pageNumber="679">Thailand</collectingCountry>
may represent 
<taxonomicName id="4C57B8C6F517FFEBFA56FE1BFC44FE11" authorityName="Polhemus &amp; Polhemus" authorityYear="2013" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="B9231F57F517FFEBFA56FE1BFA46FE31" box="[1448,1467,447,470]" italics="true" pageId="14" pageNumber="679">H</emphasis>
. 
<emphasis id="B9231F57F517FFEBFCBCFE7BFC44FE11" box="[834,953,479,502]" italics="true" pageId="14" pageNumber="679">malayensis</emphasis>
</taxonomicName>
as well, since it appears that 
<taxonomicName id="4C57B8C6F517FFEBFB0BFE7BFA70FE11" authorityName="Polhemus &amp; Polhemus" authorityYear="2013" box="[1269,1421,478,502]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="B9231F57F517FFEBFB0BFE7BFAF5FE11" box="[1269,1288,479,502]" italics="true" pageId="14" pageNumber="679">H</emphasis>
. 
<emphasis id="B9231F57F517FFEBFAE8FE7BFA70FE11" box="[1302,1421,479,502]" italics="true" pageId="14" pageNumber="679">malayensis</emphasis>
</taxonomicName>
may occur as far north as 
<collectingCountry id="F34083D5F517FFEBFBDEFE5AFBA9FDF2" box="[1056,1108,510,533]" name="Laos" pageId="14" pageNumber="679">Laos</collectingCountry>
(Zettel, in litt.). As such, all of the extensive material listed as 
<taxonomicName id="4C57B8C6F517FFEBFB9DFDBBFB3AFDD1" authorityName="Montandon" authorityYear="1897" box="[1123,1223,542,566]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="B9231F57F517FFEBFB9DFDBBFB8BFDD1" box="[1123,1142,543,566]" italics="true" pageId="14" pageNumber="679">H</emphasis>
. 
<emphasis id="B9231F57F517FFEBFB7CFDBBFB3AFDD1" box="[1154,1223,543,566]" italics="true" pageId="14" pageNumber="679">ovatus</emphasis>
</taxonomicName>
by Sites &amp; Vitheepradit (2011) will need to be critically re-examined on the basis of the characters discussed above in order to determine which of these two species was represented at any given locality, and the distribution map for this species given in their 
<figureCitation id="136CDFC0F517FFEBFAABFD39FA6FFD51" box="[1365,1426,669,694]" captionStart="Figs" captionStartId="2.[136,180,2004,2026]" captionTargetBox="[219,1386,187,1958]" captionTargetId="figure-8@2.[199,1408,156,1984]" captionTargetPageId="2" captionText="Figs. 2–10. Naucoris species, structural details of male genitalia. 2–4. N. sigaloeis La Rivers, specimen from NE Thailand. 2. Left paramere. 3. Right paramere. 4. Phallotheca. 5, 6. N. scutellaris Stål, specimen from Singapore, Lower Fort Peirce Forest. 5. Left paramere. 6. Right paramere. 7–10. N. minutus, new species, specimen from Singapore, Chestnut Drive. 7. Left paramere. 8. Right paramere. 9. Phallotheca. 10. Proctiger." figureDoi="http://doi.org/10.5281/zenodo.5352704" httpUri="https://zenodo.org/record/5352704/files/figure.png" pageId="14" pageNumber="679">Fig. 9</figureCitation>
will need to be revised for the southern half of the distribution in question, given that some of the symbols likely depict populations of 
<taxonomicName id="4C57B8C6F517FFEBFC1BFD5BFB83FCF1" authorityName="Polhemus &amp; Polhemus" authorityYear="2013" box="[997,1150,767,790]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="B9231F57F517FFEBFC1BFD5BFB83FCF1" box="[997,1150,767,790]" italics="true" pageId="14" pageNumber="679">H. malayensis</emphasis>
</taxonomicName>
rather than 
<taxonomicName id="4C57B8C6F517FFEBFAFCFD5BFA97FCF1" authorityName="Montandon" authorityYear="1897" box="[1282,1386,767,790]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="679" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="B9231F57F517FFEBFAFCFD5BFA97FCF1" box="[1282,1386,767,790]" italics="true" pageId="14" pageNumber="679">H. ovatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>