treatments-xml/data/03/FE/4B/03FE4B3B7B4CFF86FF0355CCFE4B7868.xml
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<document id="E564CD5C6D2851FD00580BD85B67ECB5" ID-DOI="10.11646/zootaxa.5446.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="11101578" ID-ZooBank="B1B73538-4AD4-4E0B-843F-74411F6E9AB3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1714634582387" checkinUser="plazi" docAuthor="Samoh, Abdulloh &amp; Grootaert, Patrick" docDate="2024" docId="03FE4B3B7B4CFF86FF0355CCFE4B7868" docLanguage="en" docName="zootaxa.5446.2.2.pdf" docOrigin="Zootaxa 5446 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5446.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hercostomus squamatus Samoh &amp; Grootaert 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="189" masterDocId="FFC733437B49FF8CFF94533BFFE77B51" masterDocTitle="New species and records of the genus Hercostomus Loew (Diptera: Dolichopodidae) from Thailand mangroves, with notes on the Hercostomus fauna of Singapore mangroves" masterLastPageNumber="204" masterPageNumber="179" pageNumber="184" updateTime="1714635081008" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="39E15C29EC85A876578AAC76BF39F20A">New species and records of the genus Hercostomus Loew (Diptera: Dolichopodidae) from Thailand mangroves, with notes on the Hercostomus fauna of Singapore mangroves</mods:title>
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<mods:name id="CD350BAC05A01822750CAEA4B90FBA26" type="personal">
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<mods:namePart id="0158106C31C0C7B416EE1712D04FFE23">Samoh, Abdulloh</mods:namePart>
<mods:affiliation id="1F93F720E1D79245A46551F90950B8C3">Princess Maha Chakri Sirindhorn Natural History Museum (PSUNHM), Prince of Songkla University, Kho Hong, Hat Yai, Songkhla, Thailand, 90110</mods:affiliation>
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<mods:namePart id="5E1232DDE0B5270BC9DF40934872265D">Grootaert, Patrick</mods:namePart>
<mods:affiliation id="74C505DC1131EDE5A5B24002F1DB2BC7">National Biodiversity Centre, NParks, Singapore; Lee Kong Chian Natural History Museum, National University of Singapore; Department of Entomology, Royal Belgian Institute of Natural Sciences, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; Princess Maha Chakri Sirindhorn Natural History Museum (PSUNHM), Prince of Songkla University, Kho Hong, Hat Yai, Songkhla, Thailand, 90110</mods:affiliation>
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<mods:date id="55B62E6B66F2F008F60AAF9A58A10BAC">2024</mods:date>
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<subSubSection id="C34DA9A67B4CFF89FF0355CCFDC17C41" box="[151,550,1782,1809]" pageId="5" pageNumber="184" type="nomenclature">
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<emphasis id="B923263F7B4CFF89FF0355CCFDC17C41" bold="true" box="[151,550,1782,1809]" pageId="5" pageNumber="184">
<taxonomicName id="4C5781AE7B4CFF89FF0355CCFE277C40" authority="Samoh &amp; Grootaert, 2024" authorityName="Samoh &amp; Grootaert" authorityYear="2024" box="[151,448,1783,1809]" class="Insecta" family="Dolichopodidae" genus="Hercostomus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="184" phylum="Arthropoda" rank="species" species="squamatus" status="sp. nov.">
<emphasis id="B923263F7B4CFF89FF0355CCFE277C40" bold="true" box="[151,448,1783,1809]" italics="true" pageId="5" pageNumber="184">Hercostomus squamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2109B447B4CFF89FE5C55CDFDC17C41" box="[456,550,1782,1808]" pageId="5" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C34DA9A67B4CFF89FF035421FEEC7C65" box="[151,267,1818,1844]" pageId="5" pageNumber="184" type="description">
<paragraph id="8BE8FA2D7B4CFF89FF035421FEEC7C65" blockId="5.[151,550,1782,1844]" box="[151,267,1818,1844]" pageId="5" pageNumber="184">
(
<figureCitation id="136CE6A87B4CFF89FF0B5421FEE47C65" box="[159,259,1818,1844]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="6.[151,250,1943,1967]" captionStartId-1="7.[151,250,1944,1968]" captionStartId-2="8.[151,250,965,989]" captionTargetBox-0="[220,1350,201,1898]" captionTargetBox-1="[193,1363,214,1879]" captionTargetBox-2="[168,1420,198,924]" captionTargetId-0="figure-21@6.[214,1373,181,1919]" captionTargetId-1="figure-18@7.[193,1395,181,1920]" captionTargetId-2="figure-21@8.[151,1436,181,941]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURE 4. Hercostomus squamatus sp. nov. Male habitus (photo: Abdulloh Samoh)." captionText-1="FIGURE 5. Hercostomus squamatus sp. nov. Female habitus (photo: Abdulloh Samoh)." captionText-2="FIGURE 6. Hercostomus squamatus sp. nov. Male terminalia (Singapore, Semakau, ZRCBDP00002799). A. Male terminalia lateral; B. ventral border of epandrium with apicoventral epandrial lobe; C. Tip of aedeagus and hypandrium; D. Lateral view of cercus and surstylus. Scale 0.1 mm (image P. Grootaert)." figureDoi-0="http://doi.org/10.5281/zenodo.11101586" figureDoi-1="http://doi.org/10.5281/zenodo.11101588" figureDoi-2="http://doi.org/10.5281/zenodo.11101590" httpUri-0="https://zenodo.org/record/11101586/files/figure.png" httpUri-1="https://zenodo.org/record/11101588/files/figure.png" httpUri-2="https://zenodo.org/record/11101590/files/figure.png" pageId="5" pageNumber="184">Figs 46</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34DA9A67B4CFF84FF035459FD727E14" lastPageId="8" lastPageNumber="187" pageId="5" pageNumber="184" type="materials_examined">
<paragraph id="8BE8FA2D7B4CFF89FF035459FA7D7CB9" blockId="5.[151,1437,1890,2025]" pageId="5" pageNumber="184">
<emphasis id="B923263F7B4CFF89FF035459FEA27C2D" bold="true" box="[151,325,1890,1916]" pageId="5" pageNumber="184">Type material.</emphasis>
<materialsCitation id="3B3FF0707B4CFF89FEDA5459FB0F7CF1" collectingDate="2015-04-29" collectorName="A. Samoh" country="Thailand" county="Langu District" latitude="6.841778" location="Ban Pakbara" longLatPrecision="1" longitude="99.77581" municipality="Langu Subdistrict" pageId="5" pageNumber="184" specimenCount="1" specimenCount-male="1" stateProvince="Satun" typeStatus="holotype">
<emphasis id="B923263F7B4CFF89FEDA5459FE0C7C2D" bold="true" box="[334,491,1890,1916]" pageId="5" pageNumber="184">
<typeStatus id="54EC448F7B4CFF89FEDA5459FE0C7C2D" box="[334,491,1890,1916]" pageId="5" pageNumber="184" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
<specimenCount id="9D5131A47B4CFF89FE675458FDED7C2C" box="[499,522,1891,1917]" count="1" pageId="5" pageNumber="184" type="male"></specimenCount>
,
<emphasis id="B923263F7B4CFF89FD8C5459FC927C2D" bold="true" box="[536,885,1890,1916]" pageId="5" pageNumber="184">
<collectingCountry id="F340BABD7B4CFF89FD8C5459FD557C2D" box="[536,690,1890,1916]" name="Thailand" pageId="5" pageNumber="184">THAILAND</collectingCountry>
:
<collectingRegion id="499334CF7B4CFF89FD2B5459FC927C2D" box="[703,885,1890,1916]" country="Thailand" name="Satun" pageId="5" pageNumber="184">Satun Province</collectingRegion>
</emphasis>
,
<collectingCounty id="628982A17B4CFF89FC115458FBCD7C2D" box="[901,1066,1890,1917]" pageId="5" pageNumber="184">Langu District</collectingCounty>
,
<collectingMunicipality id="6B8C60577B4CFF89FBA35458FAE67C2D" box="[1079,1281,1890,1917]" pageId="5" pageNumber="184">Langu Subdistrict</collectingMunicipality>
,
<location id="8E88ACF67B4CFF89FA9A5458FA7B7C2D" LSID="urn:lsid:plazi:treatment:03FE4B3B7B4CFF86FF0355CCFE4B7868:8E88ACF67B4CFF89FA9A5458FA7B7C2D" box="[1294,1436,1890,1917]" country="Thailand" county="Langu District" latitude="6.841778" longLatPrecision="1" longitude="99.77581" municipality="Langu Subdistrict" name="Ban Pakbara" pageId="5" pageNumber="184" stateProvince="Satun">Ban Pakbara</location>
(
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), sweep netting,
<date id="FFE9DCED7B4CFF89FD0754BDFCF77CF0" box="[659,784,1926,1953]" pageId="5" pageNumber="184" value="2015-04-29">
<collectingDate id="EFAD25057B4CFF89FD0754BDFCF77CF0" box="[659,784,1926,1953]" pageId="5" pageNumber="184" value="2015-04-29">29.IV.2015</collectingDate>
</date>
,
<collectorName id="26A29FFB7B4CFF89FC8F54BCFC6C7CF1" box="[795,907,1926,1953]" pageId="5" pageNumber="184">A. Samoh</collectorName>
leg., PSUNHM, PSUZCEN11
</materialsCitation>
.
<materialsCitation id="3B3FF0707B4CFF89FB6754BDFD177CB9" collectingDate="2015-04-29" collectorName="A. Samoh" country="Thailand" county="Langu District" latitude="6.841778" location="Ban Pakbara" longLatPrecision="1" longitude="99.77581" municipality="Langu Subdistrict" pageId="5" pageNumber="184" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Satun" typeStatus="paratype">
<emphasis id="B923263F7B4CFF89FB6754BDFE0D7C95" bold="true" pageId="5" pageNumber="184">
<typeStatus id="54EC448F7B4CFF89FB6754BDFA707CF1" box="[1267,1431,1926,1952]" pageId="5" pageNumber="184" type="paratype">PARATYPES</typeStatus>
.
<collectingCountry id="F340BABD7B4CFF89FF035491FED67C95" box="[151,305,1962,1988]" name="Thailand" pageId="5" pageNumber="184">THAILAND</collectingCountry>
:
<collectingRegion id="499334CF7B4CFF89FEAD5491FE0D7C95" box="[313,490,1962,1988]" country="Thailand" name="Satun" pageId="5" pageNumber="184">Satun Province</collectingRegion>
</emphasis>
,
<specimenCount id="9D5131A47B4CFF89FE615491FDFA7C94" box="[501,541,1962,1989]" count="3" pageId="5" pageNumber="184" type="male">3♂</specimenCount>
,
<specimenCount id="9D5131A47B4CFF89FDB15491FDAC7C94" box="[549,587,1962,1989]" count="2" pageId="5" pageNumber="184" type="female">2♀</specimenCount>
,
<collectingCounty id="628982A17B4CFF89FDC25490FD117C95" box="[598,758,1962,1989]" pageId="5" pageNumber="184">Langu District</collectingCounty>
,
<collectingMunicipality id="6B8C60577B4CFF89FD695490FC247C95" box="[765,963,1962,1989]" pageId="5" pageNumber="184">Langu Subdistrict</collectingMunicipality>
,
<location id="8E88ACF67B4CFF89FC5E5490FBB47C95" LSID="urn:lsid:plazi:treatment:03FE4B3B7B4CFF86FF0355CCFE4B7868:8E88ACF67B4CFF89FC5E5490FBB47C95" box="[970,1107,1962,1989]" country="Thailand" county="Langu District" latitude="6.841778" longLatPrecision="1" longitude="99.77581" municipality="Langu Subdistrict" name="Ban Pakbara" pageId="5" pageNumber="184" stateProvince="Satun">Ban Pakbara</location>
(
<geoCoordinate id="EE639CEA7B4CFF89FBCB5491FB087C94" box="[1119,1263,1962,1989]" degrees="6" direction="north" minutes="50" orientation="latitude" pageId="5" pageNumber="184" precision="1" seconds="30.4" value="6.841778">6°5030.4”N</geoCoordinate>
<geoCoordinate id="EE639CEA7B4CFF89FB675491FA777C95" box="[1267,1424,1962,1988]" degrees="99" direction="east" minutes="46" orientation="longitude" pageId="5" pageNumber="184" precision="1" seconds="32.9" value="99.77581">99°4632.9”E</geoCoordinate>
), sweep netting,
<date id="FFE9DCED7B4CFF89FEAA54F5FE277CB8" box="[318,448,1998,2025]" pageId="5" pageNumber="184" value="2015-04-29">
<collectingDate id="EFAD25057B4CFF89FEAA54F5FE277CB8" box="[318,448,1998,2025]" pageId="5" pageNumber="184" value="2015-04-29">29.IV.2015</collectingDate>
</date>
,
<collectorName id="26A29FFB7B4CFF89FE5E54F5FDDD7CB9" box="[458,570,1998,2024]" pageId="5" pageNumber="184">A. Samoh</collectorName>
leg., PSUNHM
</materialsCitation>
.
<materialsCitation id="3B3FF0707B4CFF89FD6E54F5FA7D7CB9" box="[762,1434,1998,2025]" collectorName="Thai" country="Thailand" location="Thai Mueang district" pageId="5" pageNumber="184" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Phangnga" typeStatus="paratype">
<emphasis id="B923263F7B4CFF89FD6E54F5FC097CB9" bold="true" box="[762,1006,1998,2024]" pageId="5" pageNumber="184">
<collectingRegion id="499334CF7B4CFF89FD6E54F5FC097CB9" box="[762,1006,1998,2024]" country="Thailand" name="Phangnga" pageId="5" pageNumber="184">Phang Nga Province</collectingRegion>
</emphasis>
,
<specimenCount id="9D5131A47B4CFF89FC6854F5FBCF7CB8" box="[1020,1064,1998,2025]" count="1" pageId="5" pageNumber="184" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D5131A47B4CFF89FBA054F5FBB97CB8" box="[1076,1118,1998,2025]" count="1" pageId="5" pageNumber="184" type="female">1 ♀</specimenCount>
,
<location id="8E88ACF67B4CFF89FBF854F5FABB7CB9" LSID="urn:lsid:plazi:treatment:03FE4B3B7B4CFF86FF0355CCFE4B7868:8E88ACF67B4CFF89FBF854F5FABB7CB9" box="[1132,1372,1998,2025]" country="Thailand" name="Thai Mueang district" pageId="5" pageNumber="184" stateProvince="Phangnga">Thai Mueang district</location>
,
<collectorName id="26A29FFB7B4CFF89FAF254F5FA7D7CB9" box="[1382,1434,1998,2024]" pageId="5" pageNumber="184">Thai</collectorName>
</materialsCitation>
</paragraph>
<caption id="DF28AAA57B4FFF8AFF0354ACFBE47CE1" ID-DOI="http://doi.org/10.5281/zenodo.11101586" ID-Zenodo-Dep="11101586" box="[151,1027,1943,1968]" httpUri="https://zenodo.org/record/11101586/files/figure.png" pageId="6" pageNumber="185" startId="6.[151,250,1943,1967]" targetBox="[220,1350,201,1898]" targetPageId="6" targetType="figure">
<paragraph id="8BE8FA2D7B4FFF8AFF0354ACFBE47CE1" blockId="6.[151,1027,1943,1968]" box="[151,1027,1943,1968]" pageId="6" pageNumber="185">
<emphasis id="B923263F7B4FFF8AFF0354ACFEF47CFE" bold="true" box="[151,275,1943,1967]" pageId="6" pageNumber="185">FIGURE 4.</emphasis>
<taxonomicName id="4C5781AE7B4FFF8AFE8D54A3FDF77CFE" authority="Samoh &amp; Grootaert, 2024" authorityName="Samoh &amp; Grootaert" authorityYear="2024" box="[281,528,1944,1967]" class="Insecta" family="Dolichopodidae" genus="Hercostomus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="185" phylum="Arthropoda" rank="species" species="squamatus" status="sp. nov.">
<emphasis id="B923263F7B4FFF8AFE8D54A3FDF77CFE" box="[281,528,1944,1967]" italics="true" pageId="6" pageNumber="185">Hercostomus squamatus</emphasis>
</taxonomicName>
<emphasis id="B923263F7B4FFF8AFD8354A3FD8F7CE1" bold="true" box="[535,616,1944,1968]" pageId="6" pageNumber="185">
<taxonomicNameLabel id="A2109B447B4FFF8AFD8354A3FD8F7CE1" box="[535,616,1944,1968]" pageId="6" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Male habitus (photo: Abdulloh Samoh).
</paragraph>
</caption>
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<paragraph id="8BE8FA2D7B4EFF8BFF0354A3FBFE7CE1" blockId="7.[151,1049,1944,1968]" box="[151,1049,1944,1968]" pageId="7" pageNumber="186">
<emphasis id="B923263F7B4EFF8BFF0354A3FEF47CE1" bold="true" box="[151,275,1944,1968]" pageId="7" pageNumber="186">FIGURE 5.</emphasis>
<taxonomicName id="4C5781AE7B4EFF8BFE8D54A2FDF77CE1" authority="Samoh &amp; Grootaert, 2024" authorityName="Samoh &amp; Grootaert" authorityYear="2024" box="[281,528,1945,1968]" class="Insecta" family="Dolichopodidae" genus="Hercostomus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="186" phylum="Arthropoda" rank="species" species="squamatus" status="sp. nov.">
<emphasis id="B923263F7B4EFF8BFE8D54A2FDF77CE1" box="[281,528,1945,1968]" italics="true" pageId="7" pageNumber="186">Hercostomus squamatus</emphasis>
</taxonomicName>
<emphasis id="B923263F7B4EFF8BFD8354A3FD8F7CE1" bold="true" box="[535,616,1944,1968]" pageId="7" pageNumber="186">
<taxonomicNameLabel id="A2109B447B4EFF8BFD8354A3FD8F7CE1" box="[535,616,1944,1968]" pageId="7" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Female habitus (photo: Abdulloh Samoh).
</paragraph>
</caption>
<caption id="DF28AAA57B41FF84FF0350FEFD157F74" ID-DOI="http://doi.org/10.5281/zenodo.11101590" ID-Zenodo-Dep="11101590" httpUri="https://zenodo.org/record/11101590/files/figure.png" pageId="8" pageNumber="187" startId="8.[151,250,965,989]" targetBox="[168,1420,198,924]" targetPageId="8" targetType="figure">
<paragraph id="8BE8FA2D7B41FF84FF0350FEFD157F74" blockId="8.[151,1437,965,1061]" pageId="8" pageNumber="187">
<emphasis id="B923263F7B41FF84FF0350FEFEF4788C" bold="true" box="[151,275,965,989]" pageId="8" pageNumber="187">FIGURE 6.</emphasis>
<taxonomicName id="4C5781AE7B41FF84FE8E50FDFDF6788C" authority="Samoh &amp; Grootaert, 2024" authorityName="Samoh &amp; Grootaert" authorityYear="2024" box="[282,529,966,989]" class="Insecta" family="Dolichopodidae" genus="Hercostomus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="squamatus" status="sp. nov.">
<emphasis id="B923263F7B41FF84FE8E50FDFDF6788C" box="[282,529,966,989]" italics="true" pageId="8" pageNumber="187">Hercostomus squamatus</emphasis>
</taxonomicName>
<emphasis id="B923263F7B41FF84FD8C50FEFD8E788C" bold="true" box="[536,617,965,989]" pageId="8" pageNumber="187">
<taxonomicNameLabel id="A2109B447B41FF84FD8C50FEFD8E788C" box="[536,617,965,989]" pageId="8" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Male terminalia (Singapore, Semakau, ZRCBDP00002799).
<emphasis id="B923263F7B41FF84FB4850FEFB09788C" bold="true" box="[1244,1262,965,989]" pageId="8" pageNumber="187">A</emphasis>
. Male terminalia lateral;
<emphasis id="B923263F7B41FF84FF7150D2FF1B7F50" bold="true" box="[229,252,1001,1025]" pageId="8" pageNumber="187">B.</emphasis>
ventral border of epandrium with apicoventral epandrial lobe;
<emphasis id="B923263F7B41FF84FC1450D2FC7F7F50" bold="true" box="[896,920,1001,1025]" pageId="8" pageNumber="187">C.</emphasis>
Tip of aedeagus and hypandrium;
<emphasis id="B923263F7B41FF84FB6950D2FAF27F50" bold="true" box="[1277,1301,1001,1025]" pageId="8" pageNumber="187">D.</emphasis>
Lateral view of cercus and surstylus. Scale 0.1 mm (image P. Grootaert).
</paragraph>
</caption>
<paragraph id="8BE8FA2D7B41FF84FF035743FD647FAC" blockId="8.[151,1437,1143,2040]" pageId="8" pageNumber="187">
<materialsCitation id="3B3FF0707B41FF84FF035743FAF37FE4" collectingDate="2021-04-04" collectionCode="SMO" collectorName="A. Samoh" country="Singapore" county="Semakau Island" latitude="8.482944" location="Mueang" longLatPrecision="1" longitude="98.25486" municipality="Mangrove creek" pageId="8" pageNumber="187" specimenCode="SMO3" specimenCount="1" specimenCount-male="1">
<location id="8E88ACF67B41FF84FF035743FF157FC3" LSID="urn:lsid:plazi:treatment:03FE4B3B7B4CFF86FF0355CCFE4B7868:8E88ACF67B41FF84FF035743FF157FC3" box="[151,242,1144,1170]" country="Singapore" county="Semakau Island" latitude="8.482944" longLatPrecision="1" longitude="98.25486" municipality="Mangrove creek" name="Mueang" pageId="8" pageNumber="187">Mueang</location>
subdistrict,
<location id="8E88ACF67B41FF84FE165743FDC47FC0" LSID="urn:lsid:plazi:treatment:03FE4B3B7B4CFF86FF0355CCFE4B7868:8E88ACF67B41FF84FE165743FDC47FC0" box="[386,547,1143,1170]" country="Singapore" county="Semakau Island" latitude="8.482944" longLatPrecision="1" longitude="98.25486" municipality="Mangrove creek" name="Ban Thap Pla" pageId="8" pageNumber="187">Ban Thap Pla</location>
,
<collectingMunicipality id="6B8C60577B41FF84FDA65743FD0C7FC0" box="[562,747,1143,1170]" pageId="8" pageNumber="187">Mangrove creek</collectingMunicipality>
and local fishing port (
<geoCoordinate id="EE639CEA7B41FF84FB95574CFB747FC3" box="[1025,1171,1143,1170]" degrees="8" direction="north" minutes="28" orientation="latitude" pageId="8" pageNumber="187" precision="1" seconds="58.6" value="8.482944">8°2858.6”N</geoCoordinate>
,
<geoCoordinate id="EE639CEA7B41FF84FB36574CFAD87FC3" box="[1186,1343,1143,1170]" degrees="98" direction="east" minutes="15" orientation="longitude" pageId="8" pageNumber="187" precision="1" seconds="17.5" value="98.25486">98°1517.5”E</geoCoordinate>
), sweep netting,
<date id="FFE9DCED7B41FF84FF6857A0FE8B7FE4" box="[252,364,1179,1206]" pageId="8" pageNumber="187" value="2021-04-04">
<collectingDate id="EFAD25057B41FF84FF6857A0FE8B7FE4" box="[252,364,1179,1206]" pageId="8" pageNumber="187" value="2021-04-04">4.IV.2021</collectingDate>
</date>
,
<collectorName id="26A29FFB7B41FF84FE1457A0FE1C7FE4" box="[384,507,1179,1205]" pageId="8" pageNumber="187">A. Samoh</collectorName>
leg., PSUNHM.
<emphasis id="B923263F7B41FF84FD4E57A0FC6E7FE4" bold="true" box="[730,905,1179,1205]" pageId="8" pageNumber="187">
<collectingCountry id="F340BABD7B41FF84FD4E57A0FC677FE4" box="[730,896,1179,1205]" name="Singapore" pageId="8" pageNumber="187">SINGAPORE</collectingCountry>
:
</emphasis>
<specimenCount id="9D5131A47B41FF84FC0F57A0FC367FE7" box="[923,977,1179,1206]" count="1" pageId="8" pageNumber="187" type="male">1 ♂</specimenCount>
,
<collectingCounty id="628982A17B41FF84FC7357A0FB447FE4" box="[999,1187,1179,1205]" pageId="8" pageNumber="187">Semakau Island</collectingCounty>
(
<specimenCode id="DBF152567B41FF84FB2857A0FAEB7FE4" box="[1212,1292,1179,1205]" collectionCode="SMO" pageId="8" pageNumber="187">SMO3</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B3FF0707B41FF84FABE57A0FD647FAC" collectingDate="2012-05-24" collectionCode="LKCNHM" country="Singapore" location="Semakau Island" pageId="8" pageNumber="187" specimenCode="ZRCBDP00002799, SMO3, ZRCBDP00007660" specimenCount="1" specimenCount-female="1">
<date id="FFE9DCED7B41FF84FABE57A0FA7B7FE4" box="[1322,1436,1179,1205]" pageId="8" pageNumber="187" value="2012-05-24">
<collectingDate id="EFAD25057B41FF84FABE57A0FA7B7FE4" box="[1322,1436,1179,1205]" pageId="8" pageNumber="187" value="2012-05-24">24.V.2012</collectingDate>
</date>
(
<specimenCode id="DBF152567B41FF84FF0B5784FE657F88" box="[159,386,1215,1241]" collectionCode="ZRCBDP" pageId="8" pageNumber="187">ZRCBDP00002799</specimenCode>
: barcoded, dissected and figured (
<collectionCode id="ED4662E87B41FF84FC8D5784FC7E7F88" box="[793,921,1215,1241]" pageId="8" pageNumber="187">LKCNHM</collectionCode>
);
<specimenCount id="9D5131A47B41FF84FC265784FC067F8B" box="[946,993,1215,1242]" count="1" pageId="8" pageNumber="187" type="female">1 ♀</specimenCount>
,
<location id="8E88ACF67B41FF84FC615784FB4B7F88" LSID="urn:lsid:plazi:treatment:03FE4B3B7B4CFF86FF0355CCFE4B7868:8E88ACF67B41FF84FC615784FB4B7F88" box="[1013,1196,1215,1241]" country="Singapore" name="Semakau Island" pageId="8" pageNumber="187">Semakau Island</location>
(
<specimenCode id="DBF152567B41FF84FB555784FAF67F88" box="[1217,1297,1215,1241]" collectionCode="SMO" pageId="8" pageNumber="187">SMO3</specimenCode>
),
<date id="FFE9DCED7B41FF84FABE5784FA7B7F88" box="[1322,1436,1215,1241]" pageId="8" pageNumber="187" value="2012-05-24">
<collectingDate id="EFAD25057B41FF84FABE5784FA7B7F88" box="[1322,1436,1215,1241]" pageId="8" pageNumber="187" value="2012-05-24">24.V.2012</collectingDate>
</date>
(
<specimenCode id="DBF152567B41FF84FF0B57D8FE657FAC" box="[159,386,1251,1277]" collectionCode="ZRCBDP" pageId="8" pageNumber="187">ZRCBDP00007660</specimenCode>
: barcoded;
<collectionCode id="ED4662E87B41FF84FD9657D8FD987FAC" box="[514,639,1251,1277]" pageId="8" pageNumber="187">LKCNHM</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE8FA2D7B41FF84FF53563CFD727E14" blockId="8.[151,1437,1143,2040]" pageId="8" pageNumber="187">
<emphasis id="B923263F7B41FF84FF53563CFE197E70" bold="true" box="[199,510,1287,1313]" pageId="8" pageNumber="187">Other material examined.</emphasis>
About
<specimenCount id="9D5131A47B41FF84FDC7563CFD1D7E70" box="[595,762,1287,1313]" count="180" pageId="8" pageNumber="187" type="generic">180 specimens</specimenCount>
of
<taxonomicName id="4C5781AE7B41FF84FCB65633FC5A7E70" authority="Samoh &amp; Grootaert, 2024" authorityName="Samoh &amp; Grootaert" authorityYear="2024" box="[802,957,1288,1313]" class="Insecta" family="Dolichopodidae" genus="Hercostomus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="squamatus" status="sp. nov.">
<emphasis id="B923263F7B41FF84FCB65633FC5A7E70" box="[802,957,1288,1313]" italics="true" pageId="8" pageNumber="187">H. squamatus</emphasis>
</taxonomicName>
<emphasis id="B923263F7B41FF84FC51563CFBC77E70" bold="true" box="[965,1056,1287,1313]" pageId="8" pageNumber="187">
<taxonomicNameLabel id="A2109B447B41FF84FC51563CFBC77E70" box="[965,1056,1287,1313]" pageId="8" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from
<collectingCountry id="F340BABD7B41FF84FBF3563CFB3E7E70" box="[1127,1241,1287,1313]" name="Singapore" pageId="8" pageNumber="187">Singapore</collectingCountry>
have been NGSbarcoded and are all conserved at
<collectionCode id="ED4662E87B41FF84FD805610FD767E14" box="[532,657,1323,1349]" pageId="8" pageNumber="187">LKCNHM</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C34DA9A67B41FF84FF535674FE7A7EA8" pageId="8" pageNumber="187" type="diagnosis">
<paragraph id="8BE8FA2D7B41FF84FF535674FE7A7EA8" blockId="8.[151,1437,1143,2040]" pageId="8" pageNumber="187">
<emphasis id="B923263F7B41FF84FF535674FEA57E38" bold="true" box="[199,322,1359,1385]" pageId="8" pageNumber="187">Diagnosis.</emphasis>
The first flagellomere is elongated, three times as long as wide. All coxae and femora are black, with the exception of the tips of the fore and mid femora. The hind femur lacks long ventral setae at the base. Hind tibia with a very long ventral seta on apical third, being at least 3 times as long as tibia is deep. The male genitalia are notably long, extending to the thorax, and the cercus is triangular-shaped, equipped with seven strong, black bristles along the outer margin.
</paragraph>
</subSubSection>
<subSubSection id="C34DA9A67B41FF86FF535538FB327B84" lastPageId="10" lastPageNumber="189" pageId="8" pageNumber="187" type="description">
<paragraph id="8BE8FA2D7B41FF85FF535538FEBB7A4C" blockId="8.[151,1437,1143,2040]" lastBlockId="9.[151,1437,150,285]" lastPageId="9" lastPageNumber="188" pageId="8" pageNumber="187">
<emphasis id="B923263F7B41FF84FF535538FE787D4C" bold="true" box="[199,415,1539,1565]" pageId="8" pageNumber="187">Description. Male</emphasis>
(
<figureCitation id="136CE6A87B41FF84FE255538FE1E7D4C" box="[433,505,1539,1565]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[220,1350,201,1898]" captionTargetId="figure-21@6.[214,1373,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Hercostomus squamatus sp. nov. Male habitus (photo: Abdulloh Samoh)." figureDoi="http://doi.org/10.5281/zenodo.11101586" httpUri="https://zenodo.org/record/11101586/files/figure.png" pageId="8" pageNumber="187">Fig. 4</figureCitation>
). Small to medium species (body length
<quantity id="4CAF57C87B41FF84FC435538FB857D4F" box="[983,1122,1539,1566]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.6" metricValueMin="3.2" pageId="8" pageNumber="187" unit="mm" value="3.4" valueMax="3.6" valueMin="3.2">3.23.6 mm</quantity>
, wing length
<quantity id="4CAF57C87B41FF84FA965538FA777D4C" box="[1282,1424,1539,1566]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" metricValueMax="3.5" metricValueMin="3.2" pageId="8" pageNumber="187" unit="mm" value="3.35" valueMax="3.5" valueMin="3.2">3.23.5 mm</quantity>
).
<emphasis id="B923263F7B41FF84FF03551CFF327D13" bold="true" box="[151,213,1575,1602]" italics="true" pageId="8" pageNumber="187">Head</emphasis>
metallic green with pale grey pollen. Face narrowing downward, distinctly narrower than first flagellomere. Hairs and bristles black. Postocular bristles black. Antenna blackish brown, first flagellomere elongated, 3 times as long as wide; arista apicodorsal, minutely pubescent with basal segment 0.3 times as long as apical segment. Proboscis blackish brown with black hairs, palpus black with black hairs and black apical bristles.
<emphasis id="B923263F7B41FF84FB7255A8FADB7DFF" bold="true" box="[1254,1340,1683,1710]" italics="true" pageId="8" pageNumber="187">Thorax</emphasis>
metallic green with pale grey pollen. Hairs and bristles black. Five strong dc, 6 pairs of acr; scutellum with 2 pairs of bristles (basal pair short and weak). Propleuron with 1 black bristle on lower part. Legs almost yellow; all coxae dark brown to black; femora also dark brown to black except tip of fore and mid femora. Mid and hind tarsi from tip of tarsomere 1 onward brownish. Hairs and bristles black. Mid and hind coxae each with 1 exterior bristle. Mid and hind femora each with 1 preapical bristle. Hind femur with 3 brown vs. near base (slightly shorter than width of femur). Fore tibia with 2 ad and 1 weak pd, apically with 3 long bristles, longest 0.8 times as long as tarsomere 1; mid tibia with 3 ad, 2 pd and 1 av, apically with 4 bristles; hind tibia with 3 ad, 3 pd and very long ventral on apical third, apically with 3 bristles. Fore tarsomere 1 with 2 vs.; hind tarsomere 1 with 1 vs at base.
<emphasis id="B923263F7B41FF84FB6B5488FADB7C9C" bold="true" box="[1279,1340,1971,1997]" italics="true" pageId="8" pageNumber="187">Wing</emphasis>
hyaline, tinged with greyish; veins brown, R
<subScript id="17D3F8687B41FF84FDBF54D2FDA27CA6" attach="left" box="[555,581,2025,2039]" fontSize="6" pageId="8" pageNumber="187">4+5</subScript>
and M
<subScript id="17D3F8687B41FF84FD0054D1FD497CA9" attach="left" box="[660,686,2026,2040]" fontSize="6" pageId="8" pageNumber="187">1+2</subScript>
parallel apically, CuAx ratio 0.7. Squama yellow with black hairs. Halter pale yellow.
<emphasis id="B923263F7B40FF85FEEC53ADFE017BE0" bold="true" box="[376,486,150,177]" italics="true" pageId="9" pageNumber="188">Abdomen</emphasis>
metallic green with pale grey pollen. Hairs and bristles black. Male genitalia long, reaching thorax. Epandrium about 3x as long as wide (
<figureCitation id="136CE6A87B40FF85FD675380FCAB7B84" box="[755,844,187,213]" captionStart="FIGURE 6" captionStartId="8.[151,250,965,989]" captionTargetBox="[168,1420,198,924]" captionTargetId="figure-21@8.[151,1436,181,941]" captionTargetPageId="8" captionText="FIGURE 6. Hercostomus squamatus sp. nov. Male terminalia (Singapore, Semakau, ZRCBDP00002799). A. Male terminalia lateral; B. ventral border of epandrium with apicoventral epandrial lobe; C. Tip of aedeagus and hypandrium; D. Lateral view of cercus and surstylus. Scale 0.1 mm (image P. Grootaert)." figureDoi="http://doi.org/10.5281/zenodo.11101590" httpUri="https://zenodo.org/record/11101590/files/figure.png" pageId="9" pageNumber="188">Fig. 6A</figureCitation>
), ventral border of epandrium with narrow digitiform apicoventral epandrial lobe with 3 bristles (
<figureCitation id="136CE6A87B40FF85FD1953E4FCEB7BA8" box="[653,780,223,249]" captionStart="FIGURE 6" captionStartId="8.[151,250,965,989]" captionTargetBox="[168,1420,198,924]" captionTargetId="figure-21@8.[151,1436,181,941]" captionTargetPageId="8" captionText="FIGURE 6. Hercostomus squamatus sp. nov. Male terminalia (Singapore, Semakau, ZRCBDP00002799). A. Male terminalia lateral; B. ventral border of epandrium with apicoventral epandrial lobe; C. Tip of aedeagus and hypandrium; D. Lateral view of cercus and surstylus. Scale 0.1 mm (image P. Grootaert)." figureDoi="http://doi.org/10.5281/zenodo.11101590" httpUri="https://zenodo.org/record/11101590/files/figure.png" pageId="9" pageNumber="188">Fig. 6A, B</figureCitation>
). Cercus triangular with 7 strong, black bristles on outer margin (
<figureCitation id="136CE6A87B40FF85FF615238FEB77A4C" box="[245,336,259,285]" captionStart="FIGURE 6" captionStartId="8.[151,250,965,989]" captionTargetBox="[168,1420,198,924]" captionTargetId="figure-21@8.[151,1436,181,941]" captionTargetPageId="8" captionText="FIGURE 6. Hercostomus squamatus sp. nov. Male terminalia (Singapore, Semakau, ZRCBDP00002799). A. Male terminalia lateral; B. ventral border of epandrium with apicoventral epandrial lobe; C. Tip of aedeagus and hypandrium; D. Lateral view of cercus and surstylus. Scale 0.1 mm (image P. Grootaert)." figureDoi="http://doi.org/10.5281/zenodo.11101590" httpUri="https://zenodo.org/record/11101590/files/figure.png" pageId="9" pageNumber="188">Fig. 6D</figureCitation>
).
</paragraph>
<caption id="DF28AAA57B40FF85FF035456FBBF7CD4" ID-DOI="http://doi.org/10.5281/zenodo.11101592" ID-Zenodo-Dep="11101592" box="[151,1112,1901,1926]" httpUri="https://zenodo.org/record/11101592/files/figure.png" pageId="9" pageNumber="188" startId="9.[151,250,1901,1925]" targetBox="[162,1425,320,1867]" targetPageId="9" targetType="figure">
<paragraph id="8BE8FA2D7B40FF85FF035456FBBF7CD4" blockId="9.[151,1112,1901,1926]" box="[151,1112,1901,1926]" pageId="9" pageNumber="188">
<emphasis id="B923263F7B40FF85FF035456FEF47CD7" bold="true" box="[151,275,1901,1926]" pageId="9" pageNumber="188">FIGURE 7.</emphasis>
Distribution map of the mangrove associated species of
<taxonomicName id="4C5781AE7B40FF85FCC65455FC307CD4" authorityName="Loew" authorityYear="1857" box="[850,983,1902,1925]" class="Insecta" family="Dolichopodidae" genus="Hercostomus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B923263F7B40FF85FCC65455FC307CD4" box="[850,983,1902,1925]" italics="true" pageId="9" pageNumber="188">Hercostomus</emphasis>
</taxonomicName>
in Thailand.
</paragraph>
</caption>
<paragraph id="8BE8FA2D7B43FF86FF5353ACFB327B84" blockId="10.[151,1437,150,825]" pageId="10" pageNumber="189">
<emphasis id="B923263F7B43FF86FF5353ACFEF97BE0" bold="true" box="[199,286,151,177]" pageId="10" pageNumber="189">Female</emphasis>
(
<figureCitation id="136CE6A87B43FF86FEBA53ACFE957BE0" box="[302,370,150,177]" captionStart="FIGURE 5" captionStartId="7.[151,250,1944,1968]" captionTargetBox="[193,1363,214,1879]" captionTargetId="figure-18@7.[193,1395,181,1920]" captionTargetPageId="7" captionText="FIGURE 5. Hercostomus squamatus sp. nov. Female habitus (photo: Abdulloh Samoh)." figureDoi="http://doi.org/10.5281/zenodo.11101588" httpUri="https://zenodo.org/record/11101588/files/figure.png" pageId="10" pageNumber="189">Fig. 5</figureCitation>
). Body length
<quantity id="4CAF57C87B43FF86FD8D53ACFD457BE0" box="[537,674,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="10" pageNumber="189" unit="mm" value="2.75" valueMax="2.8" valueMin="2.7">2.72.8 mm</quantity>
, wing length
<quantity id="4CAF57C87B43FF86FCAE53ADFC237BE0" box="[826,964,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="10" pageNumber="189" unit="mm" value="2.45" valueMax="2.5" valueMin="2.4">2.42.5 mm</quantity>
. Similar to male, except first flagellomere as long as wide; hind femur without vs. at base and fore tibia without long apical ventral bristles.
</paragraph>
</subSubSection>
<subSubSection id="C34DA9A67B43FF86FF5353E4FC487A34" pageId="10" pageNumber="189" type="etymology">
<paragraph id="8BE8FA2D7B43FF86FF5353E4FEC97A4C" blockId="10.[151,1437,150,825]" pageId="10" pageNumber="189">
<emphasis id="B923263F7B43FF86FF5353E4FEA97BA8" bold="true" box="[199,334,223,249]" pageId="10" pageNumber="189">Etymology.</emphasis>
The specific epithet,
<taxonomicName id="4C5781AE7B43FF86FDD253E4FD597BA9" authorityName="Samoh &amp; Grootaert" authorityYear="2024" box="[582,702,223,248]" class="Insecta" family="Dolichopodidae" genus="Hercostomus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="squamatus">
<emphasis id="B923263F7B43FF86FDD253E4FD597BA9" box="[582,702,223,248]" italics="true" pageId="10" pageNumber="189">squamatus</emphasis>
</taxonomicName>
refers to the numerous squamiform bristles on the apical border of the cercus.
</paragraph>
<paragraph id="8BE8FA2D7B43FF86FF53521DFC487A34" blockId="10.[151,1437,150,825]" pageId="10" pageNumber="189">
<emphasis id="B923263F7B43FF86FF53521DFEAC7A11" bold="true" box="[199,331,294,320]" pageId="10" pageNumber="189">Bionomics.</emphasis>
This species was collected from shaded areas of mangrove creeks and back mangroves, which had an abundance of small pebbles, sand, and muddy soils in their vicinity.
</paragraph>
</subSubSection>
<subSubSection id="C34DA9A67B43FF86FF535255FF0B7AFC" pageId="10" pageNumber="189" type="distribution">
<paragraph id="8BE8FA2D7B43FF86FF535255FF0B7AFC" blockId="10.[151,1437,150,825]" pageId="10" pageNumber="189">
<emphasis id="B923263F7B43FF86FF535255FE867AD9" bold="true" box="[199,353,366,392]" pageId="10" pageNumber="189">Distribution.</emphasis>
<collectingCountry id="F340BABD7B43FF86FEF35254FE1B7AD8" box="[359,508,367,393]" name="Thailand" pageId="10" pageNumber="189">THAILAND</collectingCountry>
:
<collectingRegion id="499334CF7B43FF86FD915254FD677AD8" box="[517,640,367,393]" country="Thailand" name="Phangnga" pageId="10" pageNumber="189">Phang Nga</collectingRegion>
and
<collectingRegion id="499334CF7B43FF86FD215254FD127AD8" box="[693,757,367,393]" country="Thailand" name="Satun" pageId="10" pageNumber="189">Satun</collectingRegion>
provinces (
<figureCitation id="136CE6A87B43FF86FCE25254FC5D7AD8" box="[886,954,367,393]" captionStart="FIGURE 7" captionStartId="9.[151,250,1901,1925]" captionTargetBox="[162,1425,320,1867]" captionTargetId="figure-74@9.[151,1436,309,1878]" captionTargetPageId="9" captionText="FIGURE 7. Distribution map of the mangrove associated species of Hercostomus in Thailand." figureDoi="http://doi.org/10.5281/zenodo.11101592" httpUri="https://zenodo.org/record/11101592/files/figure.png" pageId="10" pageNumber="189">Fig. 7</figureCitation>
),
<collectingCountry id="F340BABD7B43FF86FC5A5254FB977AD8" box="[974,1136,367,393]" name="Singapore" pageId="10" pageNumber="189">SINGAPORE</collectingCountry>
: Pulau Semakau (Semakau Island).
</paragraph>
</subSubSection>
<subSubSection id="C34DA9A67B43FF86FF53528CFE4B7868" pageId="10" pageNumber="189" type="discussion">
<paragraph id="8BE8FA2D7B43FF86FF53528CFE4B7868" blockId="10.[151,1437,150,825]" pageId="10" pageNumber="189">
<emphasis id="B923263F7B43FF86FF53528CFEDC7A80" bold="true" box="[199,315,439,465]" pageId="10" pageNumber="189">Remarks.</emphasis>
The newly discovered species bears a resemblance to
<taxonomicName id="4C5781AE7B43FF86FC03528CFBAB7A80" authorityName="Zhang, Yang &amp; Grootaert" authorityYear="2008" box="[919,1100,439,465]" class="Insecta" family="Dolichopodidae" genus="Hercostomus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="brevidigitalis">
<emphasis id="B923263F7B43FF86FC03528CFBAB7A80" box="[919,1100,439,465]" italics="true" pageId="10" pageNumber="189">H. brevidigitalis</emphasis>
</taxonomicName>
, particularly in the elongation of the first flagellomere, which is 3 times as long as wide. However, there are notable differences between the two species. In the new species, the hind femur lacks long ventral setae at the base. The hind tibia bears a very long ventral seta on apical third, being at least 3 times as long as tibia is deep. Furthermore, the cercus of the new species is triangular-shaped and characterised by the presence of seven robust, black squamiform bristles along its outer margin. In contrast, in
<taxonomicName id="4C5781AE7B43FF86FE415150FD6C79D4" authorityName="Zhang, Yang &amp; Grootaert" authorityYear="2008" box="[469,651,619,645]" class="Insecta" family="Dolichopodidae" genus="Hercostomus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="brevidigitalis">
<emphasis id="B923263F7B43FF86FE415150FD6C79D4" box="[469,651,619,645]" italics="true" pageId="10" pageNumber="189">H. brevidigitalis</emphasis>
</taxonomicName>
two long ventral setae are present at the base of the hind femur nearly as long as femur is deep, the hind tibia bears a much shorter ventral seta on the apical third that is about twice as long as tibia is deep and the cercus bears only three strong, black squamiform bristles on the outer margin (
<figureCitation id="136CE6A87B43FF86FAFB5188FF5179A0" captionStart="FIGURE 12" captionStartId="15.[151,250,1707,1731]" captionTargetBox="[169,1418,537,1666]" captionTargetId="figure-137@15.[151,1436,525,1684]" captionTargetPageId="15" captionText="FIGURE 12. Hercostomus brevidigitalis Zhang, Yang &amp; Grootaert, 2008. Male terminalia (Singapore, Pulau Ubin Ma10627). A. Tip of terminalia with cercus in lateral view; B. Apex of aedeagus and hypandrium in lateral view; C. Apicoventral lobe of epandrium; D. Cerci and surstyli in ventral view; E. Surstyli in lateral view. Scale 0.1 mm (image P. Grootaert)." figureDoi="http://doi.org/10.5281/zenodo.11101610" httpUri="https://zenodo.org/record/11101610/files/figure.png" pageId="10" pageNumber="189">Fig. 12</figureCitation>
). The genetic difference of
<taxonomicName id="4C5781AE7B43FF86FE6751ECFD6879A1" authority="Samoh &amp; Grootaert, 2024" authorityName="Samoh &amp; Grootaert" authorityYear="2024" box="[499,655,727,752]" class="Insecta" family="Dolichopodidae" genus="Hercostomus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="squamatus" status="sp. nov.">
<emphasis id="B923263F7B43FF86FE6751ECFD6879A1" box="[499,655,727,752]" italics="true" pageId="10" pageNumber="189">H. squamatus</emphasis>
</taxonomicName>
<emphasis id="B923263F7B43FF86FD0C51ECFD1479A0" bold="true" box="[664,755,727,753]" pageId="10" pageNumber="189">
<taxonomicNameLabel id="A2109B447B43FF86FD0C51ECFD1479A0" box="[664,755,727,753]" pageId="10" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C5781AE7B43FF86FCB951ECFC0379A0" authorityName="Zhang, Yang &amp; Grootaert" authorityYear="2008" box="[813,996,727,753]" class="Insecta" family="Dolichopodidae" genus="Hercostomus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="brevidigitalis">
<emphasis id="B923263F7B43FF86FCB951ECFC0379A0" box="[813,996,727,753]" italics="true" pageId="10" pageNumber="189">H. brevidigitalis</emphasis>
</taxonomicName>
is about 10%. The genetic distance of
<taxonomicName id="4C5781AE7B43FF86FF0351C0FED67845" authority="Samoh &amp; Grootaert, 2024" authorityName="Samoh &amp; Grootaert" authorityYear="2024" box="[151,305,763,788]" class="Insecta" family="Dolichopodidae" genus="Hercostomus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="squamatus" status="sp. nov.">
<emphasis id="B923263F7B43FF86FF0351C0FED67845" box="[151,305,763,788]" italics="true" pageId="10" pageNumber="189">H. squamatus</emphasis>
</taxonomicName>
<emphasis id="B923263F7B43FF86FEAC51C0FE757844" bold="true" box="[312,402,763,789]" pageId="10" pageNumber="189">
<taxonomicNameLabel id="A2109B447B43FF86FEAC51C0FE757844" box="[312,402,763,789]" pageId="10" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
between South-Thailand and
<collectingCountry id="F340BABD7B43FF86FD7651C0FCB37844" box="[738,852,763,789]" name="Singapore" pageId="10" pageNumber="189">Singapore</collectingCountry>
is between 12%, but no morphological differences have been distinguished.
</paragraph>
</subSubSection>
</treatment>
</document>