243 lines
35 KiB
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243 lines
35 KiB
XML
<document id="BC979B44493824910305A2A134E4C81F" ID-DOI="10.17109/AZH.64.4.277.2018" ID-ISSN="2064-2474" ID-Zenodo-Dep="12626311" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638277157792" checkinUser="felipe" docAuthor="Nabozhenko, Maxim & Ando, Kiyoshi" docDate="2018" docId="03BBAE26FFEAFFE7FE1CA5EC57E93CFF" docLanguage="en" docName="ActaZoolAcadSciH.64.4.277-327.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 64 (4)" docSource="http://dx.doi.org/10.17109/azh.64.4.277.2018" docStyle="DocumentStyle:183DC955716D8EEDE76F04077FE39880.8:ActaZoolAcadSciH.2015-2023.journal_article" docStyleId="183DC955716D8EEDE76F04077FE39880" docStyleName="ActaZoolAcadSciH.2015-2023.journal_article" docStyleVersion="8" docTitle="Nalassus Mulsant 1854" docType="treatment" docVersion="2" lastPageNumber="295" masterDocId="FF82D65EFFFAFFF5FFCCA16D56513801" masterDocTitle="Subtribal, Generic And Subgeneric Composition Of Darkling Beetles Of The Tribe Helopini (Coleoptera: Tenebrionidae) In The Eastern Palaearctic Region" masterLastPageNumber="327" masterPageNumber="277" pageNumber="293" updateTime="1719929912661" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="BF71A0CBB0780BE69C2781F64A6D4056">Subtribal, Generic And Subgeneric Composition Of Darkling Beetles Of The Tribe Helopini (Coleoptera: Tenebrionidae) In The Eastern Palaearctic Region</mods:title>
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<mods:namePart id="02FB1BF785A10B0753E692B148899874">Nabozhenko, Maxim</mods:namePart>
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<mods:namePart id="CF607B6515536409691C1A2959AE614F">Ando, Kiyoshi</mods:namePart>
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<mods:title id="5A5B8D42E6BC05030D78AA4714CBAE2F">Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
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<mods:date id="2D4BEB850DC0BF2CB10C5FF022133375">2018</mods:date>
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<heading id="D0E5A85CFFEAFFE5FE1CA5EC54A83C9D" box="[464,761,1153,1180]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="293" reason="2">
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<taxonomicName id="4C1264B3FFEAFFE5FE1CA5EC54A83C9D" authority="Mulsant, 1854" authorityName="Mulsant" authorityYear="1854" box="[464,761,1153,1180]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="293" phylum="Arthropoda" rank="genus">
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<emphasis id="B966C322FFEAFFE5FE1CA5EC546D3C9A" box="[464,572,1153,1179]" italics="true" pageId="16" pageNumber="293">Nalassus</emphasis>
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Mulsant, 1854
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3084CBBFFEAFFE7FDEEA5C857E93CFF" lastPageId="18" lastPageNumber="295" pageId="16" pageNumber="293" type="description">
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<paragraph id="8BAD1F30FFEAFFE5FDEEA5C854F73CC1" blockId="16.[546,678,1189,1216]" box="[546,678,1189,1216]" pageId="16" pageNumber="293">
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<heading id="D0E5A85CFFEAFFE5FDEEA5C854F73CC1" box="[546,678,1189,1216]" centered="true" fontSize="11" level="3" pageId="16" pageNumber="293" reason="3">
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(
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<figureCitation id="132903B5FFEAFFE5FDE6A5C854CF3CC1" box="[554,670,1189,1216]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStart-7="Fig" captionStart-8="Fig" captionStart-9="Fig" captionStartId-0="17.[174,210,1457,1479]" captionStartId-1="20.[136,172,1483,1505]" captionStartId-2="21.[174,210,1525,1547]" captionStartId-3="23.[174,210,1462,1484]" captionStartId-4="24.[137,173,1473,1495]" captionStartId-5="26.[138,174,1501,1523]" captionStartId-6="29.[175,211,1499,1521]" captionStartId-7="30.[136,172,1528,1550]" captionStartId-8="31.[174,210,1550,1572]" captionStartId-9="32.[136,172,1496,1518]" captionTargetBox-0="[190,1108,185,1431]" captionTargetBox-1="[136,1088,741,1458]" captionTargetBox-2="[177,1126,798,1501]" captionTargetBox-3="[175,1124,193,1429]" captionTargetBox-4="[140,1081,642,1444]" captionTargetBox-5="[138,1088,854,1477]" captionTargetBox-6="[181,1121,518,1478]" captionTargetBox-7="[151,1074,797,1499]" captionTargetBox-8="[178,1120,1043,1522]" captionTargetBox-9="[136,1088,459,1472]" captionTargetId-0="figure-17@17.[188,1111,182,1432]" captionTargetId-1="figure-209@20.[136,1088,741,1458]" captionTargetId-2="figure-168@21.[174,1126,798,1504]" captionTargetId-3="figure-17@23.[174,1124,189,1429]" captionTargetId-4="figure-134@24.[136,1088,638,1444]" captionTargetId-5="figure-184@26.[136,1088,850,1480]" captionTargetId-6="figure-88@29.[180,1121,518,1478]" captionTargetId-7="figure-172@30.[151,1074,797,1499]" captionTargetId-8="figure-259@31.[174,1126,1037,1529]" captionTargetId-9="figure-84@32.[136,1088,459,1472]" captionTargetPageId-0="17" captionTargetPageId-1="20" captionTargetPageId-2="21" captionTargetPageId-3="23" captionTargetPageId-4="24" captionTargetPageId-5="26" captionTargetPageId-6="29" captionTargetPageId-7="30" captionTargetPageId-8="31" captionTargetPageId-9="32" captionText-0="Fig. 7. The genus Nalassus, genital structure: N. (s. str.) pilushenmuus (A–D), N. (s. str.) zoltani (E–H), N. (Nipponalassus) andoi (I–M), N. (Nipponalassus) lewisi (O–R): A, E, I, O = aedeagus, ventral view, B, F, J, P = aedeagus, lateral view, C, G, K, Q = male inner sternite VIII, D, H, L, R = gastral spicula, M = female genital tubes (ag – accessory gland of spermatheca," captionText-1="Fig. 8. Nalassus (s. str.) brunneus brunneus, details of structure: A = head and prothorax, ventral view, B = head, ventral view (arrow shows weakly developed posterior grooves beginning at lower aspect of eyes), C = prosternum, D = prosternal process, procoxae and" captionText-2="Fig. 9. Nalassus brunneus konoi, holotype, female: A = habitus, B = labels, C = protibia, D = mesotibia" captionText-3="Fig. 10. Nalassus (s. str.) akitai, holotype and paratype: A = male, holotype, habitus, B = fe-" captionText-4="Fig. 11. Nalassus (s. str.) kawabatai, holotype and paratype. A = male, holotype, habitus, B = aedeagus, dorsal view, C = female paratype, habitus, D = male protibia, E = male mesotibia, F = male metatibia, G = male profemur, inner side, H = male prothorax, ventral view, I =" captionText-5="Fig. 12. Nalassus (s. str.) from Taiwan, habitus. A = N. formosanus, female, B = N. zoltani, male, C = N. pilushenmuus, male" captionText-6="Fig. 13. Nalassus (Nipponalassus) andoi, holotype and paratype. A = male, holotype, habitus, B = female paratype, habitus, C = male fore leg, dorsal margin, D = male profemur, inner side, E = aedeagus, dorsal view, F = prothorax, ventral view, G = abdominal ventrite 5, H = labels of the holotype" captionText-7="Fig. 14. Nalassus (Nipponalassus) andoi, details of structure. A = Head, ventral view, B = prothorax and base of fore legs, ventral side, C = male meso- and metaventrite, middle legs" captionText-8="Fig. 15. Nalassus (Nipponalassus) lewisi, holotype and paratype.A = male, holotype, habitus, B = female paratype, habitus, C = labels of the holotype, D = aedeagus, dorsal view" captionText-9="Fig. 16. Nalassus (Nipponalassus) lewisi (A, C–F) and N. (Nipponalassus) andoi (B), details of structure: A = head, ventral view, B = male abdominal ventrite 5, C = male meso- and metaventrite, middle legs basally, D = male abdominal ventrites, E = male profemur, inner side, F = male profemur dorsally" figureDoi-0="http://doi.org/10.5281/zenodo.12626333" figureDoi-1="http://doi.org/10.5281/zenodo.12626335" figureDoi-2="http://doi.org/10.5281/zenodo.12626337" figureDoi-3="http://doi.org/10.5281/zenodo.12626339" figureDoi-4="http://doi.org/10.5281/zenodo.12626341" figureDoi-5="http://doi.org/10.5281/zenodo.12626343" figureDoi-6="http://doi.org/10.5281/zenodo.12626345" figureDoi-7="http://doi.org/10.5281/zenodo.12626347" figureDoi-8="http://doi.org/10.5281/zenodo.12626349" figureDoi-9="http://doi.org/10.5281/zenodo.12626353" httpUri-0="https://zenodo.org/record/12626333/files/figure.png" httpUri-1="https://zenodo.org/record/12626335/files/figure.png" httpUri-2="https://zenodo.org/record/12626337/files/figure.png" httpUri-3="https://zenodo.org/record/12626339/files/figure.png" httpUri-4="https://zenodo.org/record/12626341/files/figure.png" httpUri-5="https://zenodo.org/record/12626343/files/figure.png" httpUri-6="https://zenodo.org/record/12626345/files/figure.png" httpUri-7="https://zenodo.org/record/12626347/files/figure.png" httpUri-8="https://zenodo.org/record/12626349/files/figure.png" httpUri-9="https://zenodo.org/record/12626353/files/figure.png" pageId="16" pageNumber="293">Figs 7–16</figureCitation>
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)
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</heading>
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</paragraph>
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<paragraph id="8BAD1F30FFEAFFE5FF44A5B057F13D11" blockId="16.[136,1089,1244,1296]" pageId="16" pageNumber="293">
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<typeStatus id="54A9A192FFEAFFE5FF44A5B056EF3CF3" box="[136,190,1245,1266]" pageId="16" pageNumber="293">Type</typeStatus>
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species:
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<taxonomicName id="4C1264B3FFEAFFE5FEEAA5B0578D3CF3" authority="" box="[294,476,1245,1266]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="293" phylum="Arthropoda" rank="subGenus" subGenus="Nalassus">
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<emphasis id="B966C322FFEAFFE5FEEAA5B057383CF3" box="[294,361,1245,1266]" italics="true" pageId="16" pageNumber="293">Helops</emphasis>
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(
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<emphasis id="B966C322FFEAFFE5FEB0A5B057853CF3" box="[380,468,1245,1266]" italics="true" pageId="16" pageNumber="293">Nalassus</emphasis>
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)
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</taxonomicName>
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<emphasis id="B966C322FFEAFFE5FE2BA5B054343CF3" box="[487,613,1245,1266]" italics="true" pageId="16" pageNumber="293">dryadophilus</emphasis>
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Mulsant, 1854, by subsequent designation (
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<bibRefCitation id="EF8362C1FFEAFFE5FF00A59457C43D11" author="NABOZHENKO, M. V." box="[204,405,1273,1296]" pageId="16" pageNumber="293" pagination="909 - 942" refId="ref19247" refString="NABOZHENKO, M. V. (2001): On the classification of the tenebrionid tribe Helopini, with a review of the genera Nalassus Mulsant and Odocnemis Allard (Coleoptera, Tenebrionidae) of the European part of CIS and the Caucasus. - Entomologicheskoe Obozrenie 80: 627 - 668. [In Russian; English translation: Entomological Review 81: 909 - 942.]" type="journal article" year="2001">Nabozhenko, 2001</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BAD1F30FFEAFFE4FF09A45E550F3E3B" blockId="16.[136,1089,1331,1584]" lastBlockId="17.[174,1127,1457,1594]" lastPageId="17" lastPageNumber="294" pageId="16" pageNumber="293">
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General morphology of adults. Body medial in size (length
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<quantity id="4CEAB2D5FFEAFFE5FC92A45E55943D49" box="[862,965,1331,1353]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="16" pageNumber="293" unit="mm" value="8.5" valueMax="12.0" valueMin="5.0">5–12 mm</quantity>
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), brown or rarely black, usually shiny, rarely with metallic luster, but sometimes dull, moderately flattened. Head with posterior grooves beginning at lower aspect of eyes. This character is, sometimes, not clear and head with posterior longitudinal depression instead of the grooves. East Asian species mainly without these grooves. Eyes moderately large or small (subgenus
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<taxonomicName id="4C1264B3FFEAFFE5FE8EA4AE54CC3DD8" authority="Nabozhenko, 2001" authorityName="Nabozhenko" authorityYear="2001" box="[322,669,1475,1497]" class="Insecta" family="Tenebrionidae" genus="Caucasonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="293" phylum="Arthropoda" rank="genus">
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<emphasis id="B966C322FFEAFFE5FE8EA4AE57983DD9" box="[322,457,1475,1496]" italics="true" pageId="16" pageNumber="293">Caucasonotus</emphasis>
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<bibRefCitation id="EF8362C1FFEAFFE5FE1EA4AE54CC3DD8" author="NABOZHENKO, M. V." box="[466,669,1475,1497]" pageId="16" pageNumber="293" pagination="909 - 942" refId="ref19247" refString="NABOZHENKO, M. V. (2001): On the classification of the tenebrionid tribe Helopini, with a review of the genera Nalassus Mulsant and Odocnemis Allard (Coleoptera, Tenebrionidae) of the European part of CIS and the Caucasus. - Entomologicheskoe Obozrenie 80: 627 - 668. [In Russian; English translation: Entomological Review 81: 909 - 942.]" type="journal article" year="2001">Nabozhenko, 2001</bibRefCitation>
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</taxonomicName>
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), not strongly transverse. Antennae of male often with weakly or strongly thickened in each antennomere, rarely (in Japanese and Taiwanese species) male and female antennomeres not thickened. Anterior margin of frontoclypeus straight (excluding deeply emarginate in
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<emphasis id="B966C322FFEAFFE5FCCDA77755DB3E2F" box="[769,906,1561,1583]" italics="true" pageId="16" pageNumber="293">N. melchiades</emphasis>
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(
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<bibRefCitation id="EF8362C1FFEAFFE5FC57A777527F3E31" author="REITTER, E." box="[923,1070,1562,1584]" pageId="16" pageNumber="293" pagination="113 - 171" refId="ref20299" refString="REITTER, E. (1922): Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). - Wiener Entomologische Zeitung 39: 1 - 44, 113 - 171. https: // doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">REITTER, 1922</bibRefCitation>
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)).
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<emphasis id="B966C322FFEBFFE4FE7AA74857933E3B" box="[438,450,1573,1594]" italics="true" pageId="17" pageNumber="294">v</emphasis>
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– vagina,
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<emphasis id="B966C322FFEBFFE4FDE2A74854663E3B" box="[558,567,1573,1594]" italics="true" pageId="17" pageNumber="294">s</emphasis>
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– spermatheca,
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<emphasis id="B966C322FFEBFFE4FD2EA74854BC3E3B" box="[738,749,1573,1594]" italics="true" pageId="17" pageNumber="294">o</emphasis>
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– oviduct)
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</paragraph>
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<caption id="DF6D4FB8FFEBFFE4FF62A4DC52373E1C" ID-DOI="http://doi.org/10.5281/zenodo.12626333" ID-Zenodo-Dep="12626333" httpUri="https://zenodo.org/record/12626333/files/figure.png" pageId="17" pageNumber="294" startId="17.[174,210,1457,1479]" targetBox="[190,1108,185,1431]" targetPageId="17">
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||
<paragraph id="8BAD1F30FFEBFFE4FF62A4DC52363E01" blockId="17.[174,1127,1457,1594]" pageId="17" pageNumber="294">
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<emphasis id="B966C322FFEBFFE4FF62A4DC56BE3DC7" bold="true" box="[174,239,1457,1479]" pageId="17" pageNumber="294">Fig. 7.</emphasis>
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The genus
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<taxonomicName id="4C1264B3FFEBFFE4FEA6A4DC57933DC7" authorityName="Mulsant" authorityYear="1854" box="[362,450,1457,1478]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="294" phylum="Arthropoda" rank="genus">
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<emphasis id="B966C322FFEBFFE4FEA6A4DC57933DC7" box="[362,450,1457,1478]" italics="true" pageId="17" pageNumber="294">Nalassus</emphasis>
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</taxonomicName>
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, genital structure:
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<emphasis id="B966C322FFEBFFE4FD46A4DF54F53DC6" box="[650,676,1458,1479]" italics="true" pageId="17" pageNumber="294">N.</emphasis>
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(s. str.)
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<emphasis id="B966C322FFEBFFE4FD34A4DC55D03DC7" box="[760,897,1457,1478]" italics="true" pageId="17" pageNumber="294">pilushenmuus</emphasis>
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(A–D),
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<taxonomicName id="4C1264B3FFEBFFE4FC19A4DF56843DE2" baseAuthorityName="Masumoto" baseAuthorityYear="1981" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="294" phylum="Arthropoda" rank="species" species="zoltani">
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<emphasis id="B966C322FFEBFFE4FC19A4DF55BE3DC6" box="[981,1007,1458,1479]" italics="true" pageId="17" pageNumber="294">N.</emphasis>
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(s. str.)
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<emphasis id="B966C322FFEBFFE4FB8FA4DC56843DE2" italics="true" pageId="17" pageNumber="294">zoltani</emphasis>
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</taxonomicName>
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(E–H),
|
||
<taxonomicName id="4C1264B3FFEBFFE4FEEEA4A3544B3DE2" authorityName="Masumoto" authorityYear="1993" box="[290,538,1486,1507]" class="Insecta" family="Tenebrionidae" genus="Tarpela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="294" phylum="Arthropoda" rank="species" species="andoi" subGenus="Nipponalassus">
|
||
<emphasis id="B966C322FFEBFFE4FEEEA4A3576D3DE2" box="[290,316,1486,1507]" italics="true" pageId="17" pageNumber="294">N.</emphasis>
|
||
(
|
||
<emphasis id="B966C322FFEBFFE4FE85A4A357893DE2" box="[329,472,1486,1507]" italics="true" pageId="17" pageNumber="294">Nipponalassus</emphasis>
|
||
)
|
||
<emphasis id="B966C322FFEBFFE4FE29A4A3544B3DE2" box="[485,538,1486,1507]" italics="true" pageId="17" pageNumber="294">andoi</emphasis>
|
||
</taxonomicName>
|
||
(I–M),
|
||
<taxonomicName id="4C1264B3FFEBFFE4FDA8A4A3550E3DE2" baseAuthorityName="Masumoto" baseAuthorityYear="1993" box="[612,863,1486,1507]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="294" phylum="Arthropoda" rank="species" species="lewisi" subGenus="Nipponalassus">
|
||
<emphasis id="B966C322FFEBFFE4FDA8A4A3542F3DE2" box="[612,638,1486,1507]" italics="true" pageId="17" pageNumber="294">N.</emphasis>
|
||
(
|
||
<emphasis id="B966C322FFEBFFE4FD47A4A3554B3DE2" box="[651,794,1486,1507]" italics="true" pageId="17" pageNumber="294">Nipponalassus</emphasis>
|
||
)
|
||
<emphasis id="B966C322FFEBFFE4FCEBA4A3550E3DE2" box="[807,863,1486,1507]" italics="true" pageId="17" pageNumber="294">lewisi</emphasis>
|
||
</taxonomicName>
|
||
(O–R): A, E, I, O = aedeagus, ventral view, B, F, J, P = aedeagus, lateral view, C, G, K, Q = male inner sternite VIII,
|
||
</paragraph>
|
||
<paragraph id="8BAD1F30FFEBFFE4FF63A76552373E1C" blockId="17.[174,1127,1457,1594]" box="[175,1126,1543,1565]" pageId="17" pageNumber="294">
|
||
D, H, L, R = gastral spicula, M = female genital tubes (
|
||
<emphasis id="B966C322FFEBFFE4FD2FA76554AB3E1C" box="[739,762,1544,1565]" italics="true" pageId="17" pageNumber="294">ag</emphasis>
|
||
– accessory gland of spermatheca,
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAD1F30FFE8FFE7FF44A1D2544D3A85" blockId="18.[135,1090,190,846]" pageId="18" pageNumber="295">
|
||
Pronotum usually weakly convex. Prothoracic hypomera with longitudinal or irregular (
|
||
<taxonomicName id="4C1264B3FFE8FFE7FF5CA1B6574638F1" authorityName="Nabozhenko" authorityYear="2001" box="[144,279,219,240]" class="Insecta" family="Tenebrionidae" genus="Caucasonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="295" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966C322FFE8FFE7FF5CA1B6574638F1" box="[144,279,219,240]" italics="true" pageId="18" pageNumber="295">Caucasonotus</emphasis>
|
||
</taxonomicName>
|
||
) wrinkles. Prosternum punctured, sometimes with smooth irregular wrinkles before procoxae. Prosternal process weakly convex, not projected apically, without teeth. Each elytron devoid of humeral callosity, with clear striae; punctures in striae normally confluent or connected (by furrow) with each other, often free from each other; interstriae with simple coeloconic sensilla, but without tubercles, eighth interstria apically more convex than other ones, connected with elytral margin; dorsal epipleural carina reaching eighth interval at apex; epipleura not reaching elytral apex and finished at level of abdominal ventrites 4 and 5. Hind wings completely developed or absent. The most of species are flightless, but some Japanese and probably Taiwanese species can fly (observations of junior author, K. A). Mesoventrite slightly depressed at middle. Metaventrite transverse, glabrous or rarely densely pubescent (subgenus
|
||
<taxonomicName id="4C1264B3FFE8FFE7FD4DA09655413A11" box="[641,784,507,528]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="295" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Nipponalassus">
|
||
<emphasis id="B966C322FFE8FFE7FD4DA09655413A11" box="[641,784,507,528]" italics="true" pageId="18" pageNumber="295">Nipponalassus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2557E59FFE8FFE7FCDAA09655D33A11" box="[790,898,507,529]" pageId="18" pageNumber="295" rank="subGenus">subgen. n.</taxonomicNameLabel>
|
||
). Male abdominal ventrites 1, 2 and sometimes 5 often with dense or sparse (
|
||
<taxonomicName id="4C1264B3FFE8FFE7FD20A37555223A2C" authorityName="Nabozhenko" authorityYear="2001" box="[748,883,536,557]" class="Insecta" family="Tenebrionidae" genus="Caucasonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="295" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966C322FFE8FFE7FD20A37555223A2C" box="[748,883,536,557]" italics="true" pageId="18" pageNumber="295">Caucasonotus</emphasis>
|
||
</taxonomicName>
|
||
) hair brushes. Legs moderately slender, tibiae simple, without teeth, not flattened nor curved. Male tarsi not widened or protarsomeres II–III slightly widened. Trochanters with single long seta, rarely (
|
||
<taxonomicName id="4C1264B3FFE8FFE7FF5CA303574E3A82" box="[144,287,622,643]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="308" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Nipponalassus">
|
||
<emphasis id="B966C322FFE8FFE7FF5CA303574E3A82" box="[144,287,622,643]" italics="true" pageId="18" pageNumber="295">Nipponalassus</emphasis>
|
||
</taxonomicName>
|
||
) with dense hair brush.
|
||
</paragraph>
|
||
<paragraph id="8BAD1F30FFE8FFE7FF09A3E655423ADA" blockId="18.[135,1090,190,846]" pageId="18" pageNumber="295">
|
||
Aedeagus ‘nalassoid’ (
|
||
<figureCitation id="132903B5FFE8FFE7FE7FA3E6540F3AA1" box="[435,606,651,673]" captionStart="Fig" captionStartId="17.[174,210,1457,1479]" captionTargetBox="[190,1108,185,1431]" captionTargetId="figure-17@17.[188,1111,182,1432]" captionTargetPageId="17" captionText="Fig. 7. The genus Nalassus, genital structure: N. (s. str.) pilushenmuus (A–D), N. (s. str.) zoltani (E–H), N. (Nipponalassus) andoi (I–M), N. (Nipponalassus) lewisi (O–R): A, E, I, O = aedeagus, ventral view, B, F, J, P = aedeagus, lateral view, C, G, K, Q = male inner sternite VIII, D, H, L, R = gastral spicula, M = female genital tubes (ag – accessory gland of spermatheca," figureDoi="http://doi.org/10.5281/zenodo.12626333" httpUri="https://zenodo.org/record/12626333/files/figure.png" pageId="18" pageNumber="295">Figs 7A–D, E–H</figureCitation>
|
||
) (subgenera
|
||
<taxonomicName id="4C1264B3FFE8FFE7FD2BA3E6556E3AA1" authorityName="Mulsant" authorityYear="1854" box="[743,831,651,672]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="295" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966C322FFE8FFE7FD2BA3E6556E3AA1" box="[743,831,651,672]" italics="true" pageId="18" pageNumber="295">Nalassus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1264B3FFE8FFE7FC87A3E655833AA1" authorityName="Nabozhenko" authorityYear="2001" box="[843,978,651,672]" class="Insecta" family="Tenebrionidae" genus="Caucasonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="295" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966C322FFE8FFE7FC87A3E655833AA1" box="[843,978,651,672]" italics="true" pageId="18" pageNumber="295">Caucasonotus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B966C322FFE8FFE7FC12A3E656ED3ABC" italics="true" pageId="18" pageNumber="295">Nipponalassus</emphasis>
|
||
in part) or ‘cylindrinotoid’ (
|
||
<figureCitation id="132903B5FFE8FFE7FE2FA3C554683ABC" box="[483,569,680,702]" captionStart="Fig" captionStartId="17.[174,210,1457,1479]" captionTargetBox="[190,1108,185,1431]" captionTargetId="figure-17@17.[188,1111,182,1432]" captionTargetPageId="17" captionText="Fig. 7. The genus Nalassus, genital structure: N. (s. str.) pilushenmuus (A–D), N. (s. str.) zoltani (E–H), N. (Nipponalassus) andoi (I–M), N. (Nipponalassus) lewisi (O–R): A, E, I, O = aedeagus, ventral view, B, F, J, P = aedeagus, lateral view, C, G, K, Q = male inner sternite VIII, D, H, L, R = gastral spicula, M = female genital tubes (ag – accessory gland of spermatheca," figureDoi="http://doi.org/10.5281/zenodo.12626333" httpUri="https://zenodo.org/record/12626333/files/figure.png" pageId="18" pageNumber="295">Fig. 7I, J</figureCitation>
|
||
) (subgenus
|
||
<taxonomicName id="4C1264B3FFE8FFE7FD74A3C555913ABC" authority="Reitter, 1922" authorityName="Reitter" authorityYear="1922" box="[696,960,680,702]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="295" phylum="Arthropoda" rank="subGenus" subGenus="Helopondrus">
|
||
<emphasis id="B966C322FFE8FFE7FD74A3C555643ABC" box="[696,821,680,701]" italics="true" pageId="18" pageNumber="295">Helopondrus</emphasis>
|
||
<bibRefCitation id="EF8362C1FFE8FFE7FCF7A3C555913ABC" author="REITTER, E." box="[827,960,680,702]" pageId="18" pageNumber="295" pagination="113 - 171" refId="ref20299" refString="REITTER, E. (1922): Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). - Wiener Entomologische Zeitung 39: 1 - 44, 113 - 171. https: // doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter, 1922</bibRefCitation>
|
||
</taxonomicName>
|
||
, one species of
|
||
<taxonomicName id="4C1264B3FFE8FFE7FF6FA3A857633ADB" box="[163,306,709,730]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="308" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Nipponalassus">
|
||
<emphasis id="B966C322FFE8FFE7FF6FA3A857633ADB" box="[163,306,709,730]" italics="true" pageId="18" pageNumber="295">Nipponalassus</emphasis>
|
||
</taxonomicName>
|
||
, one species of the nominotypical subgenus).
|
||
</paragraph>
|
||
<paragraph id="8BAD1F30FFE8FFE7FF09A38C54423B4F" blockId="18.[135,1090,190,846]" pageId="18" pageNumber="295">
|
||
Female genital tubes (
|
||
<figureCitation id="132903B5FFE8FFE7FE60A38C54513AF6" box="[428,512,737,759]" captionStart="Fig" captionStartId="17.[174,210,1457,1479]" captionTargetBox="[190,1108,185,1431]" captionTargetId="figure-17@17.[188,1111,182,1432]" captionTargetPageId="17" captionText="Fig. 7. The genus Nalassus, genital structure: N. (s. str.) pilushenmuus (A–D), N. (s. str.) zoltani (E–H), N. (Nipponalassus) andoi (I–M), N. (Nipponalassus) lewisi (O–R): A, E, I, O = aedeagus, ventral view, B, F, J, P = aedeagus, lateral view, C, G, K, Q = male inner sternite VIII, D, H, L, R = gastral spicula, M = female genital tubes (ag – accessory gland of spermatheca," figureDoi="http://doi.org/10.5281/zenodo.12626333" httpUri="https://zenodo.org/record/12626333/files/figure.png" pageId="18" pageNumber="295">Fig. 7M</figureCitation>
|
||
). Spermatheca consists of only one non-branched tube (the most of all subgenera) or the tube with several very short branches at the base. Accessory gland short, a little longer than spermatheca or subequal to spermathecal length. Basal duct of spermatheca present or short.
|
||
</paragraph>
|
||
<paragraph id="8BAD1F30FFE8FFE7FF09A21E56803C57" blockId="18.[135,1090,883,1279]" pageId="18" pageNumber="295">
|
||
Comments. The genus is divided into four subgenera: nominotypical,
|
||
<taxonomicName id="4C1264B3FFE8FFE7FF44A2F9574B3BAF" authorityName="Reitter" authorityYear="1922" box="[136,282,916,942]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="326" phylum="Arthropoda" rank="subGenus" subGenus="Helopondrus">
|
||
<emphasis id="B966C322FFE8FFE7FF44A2F9574B3BAF" box="[136,282,916,942]" italics="true" pageId="18" pageNumber="295">Helopondrus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1264B3FFE8FFE7FEE7A2F857983BAE" authorityName="Nabozhenko" authorityYear="2001" box="[299,457,917,943]" class="Insecta" family="Tenebrionidae" genus="Caucasonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="295" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966C322FFE8FFE7FEE7A2F857983BAE" box="[299,457,917,943]" italics="true" pageId="18" pageNumber="295">Caucasonotus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1264B3FFE8FFE7FDC1A2F954E23BAF" box="[525,691,916,942]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="295" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Nipponalassus">
|
||
<emphasis id="B966C322FFE8FFE7FDC1A2F954E23BAF" box="[525,691,916,942]" italics="true" pageId="18" pageNumber="295">Nipponalassus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2557E59FFE8FFE7FD71A2F9556E3BAE" box="[701,831,916,943]" pageId="18" pageNumber="295" rank="subGenus">subgen. n.</taxonomicNameLabel>
|
||
The subgenus
|
||
<taxonomicName id="4C1264B3FFE8FFE7FBCEA2F957C23BD1" authority="Reitter, 1922" authorityName="Reitter" authorityYear="1922" class="Insecta" family="Tenebrionidae" genus="Helopocerodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="295" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966C322FFE8FFE7FBCEA2F956A53BD1" italics="true" pageId="18" pageNumber="295">Helopocerodes</emphasis>
|
||
<bibRefCitation id="EF8362C1FFE8FFE7FF36A2DB57C23BD1" author="REITTER, E." box="[250,403,950,976]" pageId="18" pageNumber="295" pagination="113 - 171" refId="ref20299" refString="REITTER, E. (1922): Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). - Wiener Entomologische Zeitung 39: 1 - 44, 113 - 171. https: // doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter, 1922</bibRefCitation>
|
||
</taxonomicName>
|
||
was recently placed in the junior synonym of the nominotypical subgenus (
|
||
<bibRefCitation id="EF8362C1FFE8FFE7FE40A2B554393BF3" author="KESKIN, B. & NABOZHENKO, M. & ALPAGUT-KESKIN, N." box="[396,616,983,1010]" pageId="18" pageNumber="295" pagination="725 - 747" refId="ref18336" refString="KESKIN, B., NABOZHENKO, M. & ALPAGUT-KESKIN, N. (2017): Taxonomic review of the genera Nalassus Mulsant, 1854 and Turkonalassus gen. nov. of Turkey (Coleoptera: Tenebrionidae). - Annales zoologici 67: 725 - 747. https: // doi. org / 10.3161 / 00034541 A NZ 2017.67.4.009" type="journal article" year="2017">
|
||
KESKIN
|
||
<emphasis id="B966C322FFE8FFE7FE27A2B554723BF0" box="[491,547,983,1009]" italics="true" pageId="18" pageNumber="295">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
). Species of the subgenus
|
||
<taxonomicName id="4C1264B3FFE8FFE7FC7CA2BA52133BF0" authorityName="Reitter" authorityYear="1922" box="[944,1090,983,1009]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="295" phylum="Arthropoda" rank="subGenus" subGenus="Helopondrus">
|
||
<emphasis id="B966C322FFE8FFE7FC7CA2BA52133BF0" box="[944,1090,983,1009]" italics="true" pageId="18" pageNumber="295">Helopondrus</emphasis>
|
||
</taxonomicName>
|
||
have ‘cylindrinotoid’ male genitalia and female genital tubes but clearly belong to the genus
|
||
<taxonomicName id="4C1264B3FFE8FFE7FEA2A57657843C34" authorityName="Mulsant" authorityYear="1854" box="[366,469,1051,1077]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="295" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966C322FFE8FFE7FEA2A57657843C34" box="[366,469,1051,1077]" italics="true" pageId="18" pageNumber="295">Nalassus</emphasis>
|
||
</taxonomicName>
|
||
on larval morphology (
|
||
<bibRefCitation id="EF8362C1FFE8FFE7FD33A57656943C57" author="NABOZHENKO, M. V. & ARTOKHIN, K. S." pageId="18" pageNumber="295" pagination="72 - 79" refId="ref19712" refString="NABOZHENKO, M. V. & ARTOKHIN, K. S. (2017): Description of the larva of Nalassus (Helopondrus) sareptanus (Allard, 1876) and a position of the subgenus Helopondrus Reitter, 1922 in the system of the tribe Helopini (Coleoptera: Tenebrionidae). - Proceedings of the Russian Entomological Society 88: 72 - 79." type="journal article" year="2017">NABOZHENKO & ARTOKHIN 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAD1F30FFE8FFE7FF09A53357E93CFF" blockId="18.[135,1090,883,1279]" pageId="18" pageNumber="295">
|
||
Composition. The genus contains recent species and subspecies: ten species and one subspecies in the Pacific Palaearctic region (
|
||
<collectingCountry id="F3055FA0FFE8FFE7FCF6A51255D93C98" box="[826,904,1151,1177]" name="China" pageId="18" pageNumber="295">China</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3055FA0FFE8FFE7FC5FA5ED558D3C9B" box="[915,988,1152,1178]" name="Japan" pageId="18" pageNumber="295">Japan</collectingCountry>
|
||
, Korean Peninsula,
|
||
<collectingCountry id="F3055FA0FFE8FFE7FEDEA5CC573E3CBA" box="[274,367,1185,1211]" name="Taiwan" pageId="18" pageNumber="295">Taiwan</collectingCountry>
|
||
, and the Russian Far East), three species in the Nearctic region (NABOZHENKO
|
||
<emphasis id="B966C322FFE8FFE7FE5CA5AE57993CDC" box="[400,456,1219,1245]" italics="true" pageId="18" pageNumber="295">et al.</emphasis>
|
||
2016) and 68 species and subspecies in the Central and Western Palaearctic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |