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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.340.5470" ID-GBIF-Dataset="35e1f7dd-6c8a-457f-a648-4cb716ffe363" ID-PMC="PMC3800796" ID-Pensoft-Pub="1313-2970-340-1" ID-PubMed="24146589" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-340-1" ModsDocOrigin="ZooKeys 340" ModsDocTitle="Taxonomic and nomenclatural notes on the genera Themus Motschulsky and Lycocerus Gorham (Coleoptera, Cantharidae)" checkinTime="1451246833308" checkinUser="pensoft" docAuthor="Yang, Yuxia, Kopetz, Andreas &amp; Yang, Xingke" docDate="2013" docId="FC672358EE221534F8BD3EC3CB404713" docLanguage="en" docName="ZooKeys 340: 1-19" docOrigin="ZooKeys 340" docSource="http://dx.doi.org/10.3897/zookeys.340.5470" docTitle="Themus (Telephorops) coelestis Gorham 1889" docType="treatment" docVersion="4" lastPageNumber="3" masterDocId="FFE3C7763B12FFC4A9220B69FFF7FFC8" masterDocTitle="Taxonomic and nomenclatural notes on the genera Themus Motschulsky and Lycocerus Gorham (Coleoptera, Cantharidae)" masterLastPageNumber="19" masterPageNumber="1" pageNumber="1" updateTime="1668156592509" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic and nomenclatural notes on the genera Themus Motschulsky and Lycocerus Gorham (Coleoptera, Cantharidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yang, Yuxia</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kopetz, Andreas</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yang, Xingke</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>340</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>19</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.340.5470</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.340.5470</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-340-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152048628" LSID="urn:lsid:plazi:treatment:FC672358EE221534F8BD3EC3CB404713" httpUri="http://treatment.plazi.org/id/FC672358EE221534F8BD3EC3CB404713" lastPageId="2" lastPageNumber="3" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://species-id.net/wiki/Themus_coelestis" authority="Gorham, 1889" authorityName="Gorham" authorityYear="1889" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus (Telephorops) coelestis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="coelestis" subGenus="Telephorops">Themus (Telephorops) coelestis (Gorham, 1889)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Cantharidae" genus="Telephorus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Telephorus coelestis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="coelestis">Telephorus coelestis</taxonomicName>
<bibRefCitation author="Gorham, HS" journalOrPublisher="Proceedings of the Zoological Society" pageId="11" pageNumber="12" pagination="96 - 111" title="Descriptions of new species and a new genus of Coleoptera of the family Telephoridae." volume="1889" year="1889">Gorham 1889</bibRefCitation>
: 104, t. 10, fig. 7.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus coelestis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="coelestis">Themus coelestis</taxonomicName>
:
<bibRefCitation author="Jacobson, GG" journalOrPublisher="A. F. Devrjen, St-Petersburg" pageId="12" pageNumber="13" pagination="641 - 720" title="Zhuki Rossii i Zapadnoy Evropy. Rukovodstvo k opredeleniyu zhukov." year="1911">Jacobson 1911</bibRefCitation>
: 675.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus rugosus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="rugosus">Themus rugosus</taxonomicName>
<bibRefCitation author="Pic, M" journalOrPublisher="L'Echange" pageId="12" pageNumber="13" pagination="7 - 8" title="Coleopteres exotiques en partie nouveaux (Suite)." volume="45" year="1929">Pic 1929</bibRefCitation>
: 8. [Synonymized by
<bibRefCitation author="Wittmer, W" journalOrPublisher="Entomologischen Arbeiten aus dem Museum G. Frey" pageId="13" pageNumber="14" pagination="189 - 239" title="Beitrag zur einer Revision der Gattung Themus Motsch. Coleoptera: Cantharidae." volume="31 - 32" year="1983 a">Wittmer 1983a</bibRefCitation>
: 197.]
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus (Telephorops) coelestis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="coelestis" subGenus="Telephorops">Themus (Telephorops) coelestis</taxonomicName>
:
<bibRefCitation author="Wittmer, W" journalOrPublisher="Entomologischen Arbeiten aus dem Museum G. Frey" pageId="13" pageNumber="14" pagination="189 - 239" title="Beitrag zur einer Revision der Gattung Themus Motsch. Coleoptera: Cantharidae." volume="31 - 32" year="1983 a">Wittmer 1983a</bibRefCitation>
: 197, figs. 1, 59.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus violetipennis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="violetipennis">Themus violetipennis</taxonomicName>
<bibRefCitation pageId="0" pageNumber="1">Wang and Yang 1992</bibRefCitation>
: 265, fig 2.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Themus</taxonomicName>
(s.str.) violetipennis:
<bibRefCitation author="Svihla, V" journalOrPublisher="Veroeffentlichungen des Naturkundemuseums Erfurt" pageId="12" pageNumber="13" pagination="183 - 190" title="Redescription of the subgenera of the genus Themus Motschulsky, 1858, with description of five new species (Coleoptera: Cantharidae)." volume="27" year="2008">
<normalizedToken originalValue="Švihla">Svihla</normalizedToken>
2008
</bibRefCitation>
: 186. syn. n.
</paragraph>
</subSubSection>
<subSubSection lastPageId="1" lastPageNumber="2" pageId="0" pageNumber="1" type="type material examined">
<paragraph pageId="0" pageNumber="1">Type material examined.</paragraph>
<paragraph pageId="1" pageNumber="2">
<taxonomicName class="Insecta" family="Cantharidae" genus="Telephorus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Telephorus coelestis" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="coelestis">
<pageBreakToken pageId="1" pageNumber="2" start="start">Telephorus</pageBreakToken>
coelestis
</taxonomicName>
: Lectotype ♂ (NHMB): without locality information, [h]&quot;coelestis ♂&quot;, [h]
<normalizedToken originalValue="“♂”">&quot;♂&quot;</normalizedToken>
, [h]&quot;Themus \ (Telephorops) \ coelestis \ (Gorh.) \ det. W. Wittmer&quot;, [h]
<normalizedToken originalValue="“Type”">&quot;Type&quot;</normalizedToken>
, [p]
<normalizedToken originalValue="“LECTOTYPUS”">&quot;LECTOTYPUS&quot;</normalizedToken>
, [p]&quot;Naturhist. \ Museum Basel \ coll. W. Wittmer&quot;, [p]&quot;CANTHARIDAE \ CANTH00001277&quot;. Paralectotype: 1♀ (MNHN): [p]&quot;Kiukiang \ June, 1887 \ A. E. Pratt&quot;, [h]&quot;coelestis \ Gorh.&quot;, [h]&quot;Themus \ (Telephorops) \ coelestis \ (Gorh.) \ det. W. Wittmer&quot;, [h]
<normalizedToken originalValue="“TYPE”">&quot;TYPE&quot;</normalizedToken>
, [p]
<normalizedToken originalValue="“PARALECTOTYPUS”">&quot;PARALECTOTYPUS&quot;</normalizedToken>
.
</paragraph>
<paragraph pageId="1" pageNumber="2">
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus rugosus" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="rugosus">Themus rugosus</taxonomicName>
: Holotype ♀ (MNHN): [h]
<normalizedToken originalValue="“Fokien”">&quot;Fokien&quot;</normalizedToken>
, [h]&quot;Themus \ rugosus \ n. sp.&quot;, [h]&quot;Themus \ (Telephorops) \ coelestis \ (Gorh.) \ det. W. Wittmer&quot;, [h]
<normalizedToken originalValue="“type”">&quot;type&quot;</normalizedToken>
, [p]
<normalizedToken originalValue="“TYPE”">&quot;TYPE&quot;</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="2" type="neotype designation">
<paragraph pageId="1" pageNumber="2">Neotype designation.</paragraph>
<paragraph pageId="1" pageNumber="2">
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus violetipennis" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="violetipennis">Themus violetipennis</taxonomicName>
: Neotype ♀ (here designated, IZAS): &quot;湖南永顺杉木河林场 \ 600m \
<normalizedToken originalValue="中国科学院”">中国科学院&quot;</normalizedToken>
[Hunan, Yongshun, Shanmuhe forestry station \ 600m], &quot;1988.VIII.4 \
<normalizedToken originalValue="采集者：王书永”">采集者：王书永&quot;</normalizedToken>
[4.VIII.1988 \ leg. Shu-Yong Wang].
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="2" type="additional material examined">
<paragraph pageId="1" pageNumber="2">Additional material examined.</paragraph>
<paragraph pageId="1" pageNumber="2">
CHINA: Shaanxi: 2♀♀ (IZAS): Foping, 16.VIII.2007, leg. Yu-Xia Yang. Gansu: 1♂ (IZAS): Kangxian, Baiyunshan, 1250-1750m, 12.VII.1998, leg. Shu-Yong Wang; 1♀ (IZAS): Kangxian, Heimaguan, 1450-1550m, 13.VII.1998, leg. De-Cheng Yuan. Henan: 1♂ (IZAS): Jigongshan, 700m, 14.VII.2001, leg. Si-Qin Ge; 1♀ (IZAS): Tongbaishan, 500m, 16.VII.2001, leg. Si-Qin Ge; 1♀ (IZAS): Neixiang, Baoyunman, 21.VII.2001, leg. Fu-Qiang Chen. Anhui: 2♂♂ (NHMB): &quot;Dabieshan, 65km SW Huoshan, 1400m, 21.-24.VI.1995, leg. Bolm&quot;. Zhejiang: 1♀ (IZAS): Xitianmushan, 23.VI.1998, leg. Ming-Shui Zhao. Hubei: 1♂, 1♀ (NHMB): &quot;Hupeh, Lichuan Distr., Suisapa, 1000m, 22.VII.1948, Gressitt&amp;Djou Collra&quot;; 1♂ (NHMB): same data, 24.VII.1948; 1♂ (NHMB): same data, 29.VII.1948; 2♂♂, 1♀ (NHMB): same data, 31.VII.1948; 1♂, 1♀ (NHMB): same data, 4.VIII.1948; 1♂ (IZAS): Hefeng, Fenshuiling, 1400m, 1.VIII.1981, leg. Long-Long Yang; 1♀ (IZAS): Hefeng, Shayuan, 30.VII.1989, leg. Shu-Yong Wang. Jiangxi: 1♂ (IZAS): Jiulianshan, Huangniushi, 19.VI.1975, leg. You-Wei Zhang; 1♀ (IZAS): Longnan, Jiulianshan, 17.VI.1975, leg. You-Wei Zhang. Hunan: 1♂, 1♀ (NHMB): &quot;Yon-ping, 12.VI.1917&quot;; 1♂ (NHMB): same data, &quot;Yon-ping, 14.VI.1917&quot;; 1♂ (IZAS): Sangzhi, Tianpingshan, 700-1450m, 14.VIII.1988, leg. Shu-Yong Wang. Fujian: 1♂ (IZAS):
<normalizedToken originalValue="Chong’an">Chong'an</normalizedToken>
, Xingcun, Xianfengling, 1170m, 14.VII.1960, leg. Cheng-Lin Ma; 1♀ (IZAS): Dehua, Chengguan, 510-550m, 1.VI.1960, leg. Fu-Ji Pu; 11 spec. (NHMB): &quot;Fukien, Kuatun, 15.VIII.1946, Tschung-Sen leg.&quot;; 5 spec. (NHMB): &quot;Kuatun, 26.VII.1946&quot;; 6 spec. (NHMB): same data, 11.VII.1946; 5 spec. (NHMB): same data, 16.VIII.1946; 4 spec. (NHMB): same data, 18.IX.1946. Hainan: 1♀ (IZAS): Wanning, 10m, 9.VI.1960, leg. Chang-Qing Li. Guangxi: 1♂ (IZAS): Longsheng, Tianpingshan, 740m, 17.VI.1963, leg. Shu-Yong Wang; 1♀ (IZAS): Maoershan, Tongmujiang, 800m, 15.VII.1985, leg. Su-Bai Liao. Sichuan: 1♂, 1♀ (IZAS): Youyang, 780m, 15.VII.1989, leg. Shu-Yong Wang. Guizhou: 1♂ (NHMB):
<normalizedToken originalValue="“Kouy-Tchéou”">&quot;Kouy-Tcheou&quot;</normalizedToken>
; 1♂ (IZAS): Fanjingshan, Huguosi, 1350m, 3.VIII.2001, leg. Qiong-Zhang Song; 1♀ (IZAS): Fanjingshan, Huixiangping, 1600m, 2.VIII.2001, leg. Kang-Zhen Dong.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="2" type="distribution">
<paragraph pageId="1" pageNumber="2">Distribution.</paragraph>
<paragraph pageId="1" pageNumber="2">China (Shaanxi, Gansu, Henan, Anhui, Zhejiang, Hubei, Jiangxi, Hunan, Fujian, Hainan, Guangxi, Sichuan, Guizhou).</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="remarks">
<paragraph pageId="2" pageNumber="3">
<pageBreakToken pageId="2" pageNumber="3" start="start">Remarks</pageBreakToken>
.
</paragraph>
<paragraph pageId="2" pageNumber="3">
According to the original publication, the types of
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus violetipennis" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="violetipennis">Themus violetipennis</taxonomicName>
Wang &amp; Yang, 1992 were deposited in the IZAS and China Agriculture University, Beijing, China (CAUB), but our search of the types in the two Chinese museums have been long, repeated and with no results. The original description of
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus violetipennis" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="violetipennis">Themus violetipennis</taxonomicName>
was in accord with the standard of that time but insufficient considering the present level, and neotype allows us to satisfy a better comparision. Fortunately, a female specimen, which was collected at the same locality and date as that of one paratype designated by
<bibRefCitation pageId="2" pageNumber="3">Wang and Yang (1992)</bibRefCitation>
, was found in IZAS during our study. Its morphological characters are consistent with the original description, so it is designated as the neotype here, in order to clarify the taxonomic status of this species (ICZN, 4th, article 75.3). Furthermore, a careful examination of the types shows that
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus violetipennis" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="violetipennis">Themus violetipennis</taxonomicName>
Wang &amp; Yang, 1992 is a junior synonym of
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus (Telephorops) coelestis" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="coelestis" subGenus="Telephorops">Themus (Telephorops) coelestis</taxonomicName>
(Gorham, 1889), which is widely distributed in China based on the data from a large series of examined specimens.
</paragraph>
</subSubSection>
</treatment>
</document>
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