634 lines
87 KiB
XML
634 lines
87 KiB
XML
<document id="571E4EB93ED884780152F1E2BE6103C2" ID-DOI="10.5281/zenodo.188069" ID-GBIF-Dataset="5552705a-eca2-454c-be4b-614bae40a4a9" ID-ISSN="1175-5326" ID-Zenodo-Dep="188069" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461083768863" checkinUser="plazi" docAuthor="Botero-Trujillo, Ricardo, Erazo-Moreno, Martha C. & Pérez, Gustavo A." docDate="2009" docId="E9284E34FF83FF90FEE6FC47C8CBFEC4" docLanguage="en" docName="zt02120p038.pdf" docOrigin="Zootaxa 2120" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Microtityus bivicentorum Botero-Trujillo, Erazo-Moreno & Pérez, 2009, sp. nov." docType="treatment" docVersion="9" lastPageNumber="36" masterDocId="1511364CFF81FF99FE71FFEDC90FFFB2" masterDocTitle="A new species of Microtityus Kjellesvig-Waering (Scorpiones: Buthidae) from northern Colombia" masterLastPageNumber="38" masterPageNumber="27" pageNumber="29" updateTime="1698587631261" updateUser="plazi">
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<mods:mods id="4F58C7430964C4CB7FD676F5C1D25CD0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="021CA1469139512AE3B2C3A0CD1CD227">
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<mods:title id="90859E5BA782E7E85A945D86C2A82A72">A new species of Microtityus Kjellesvig-Waering (Scorpiones: Buthidae) from northern Colombia</mods:title>
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</mods:titleInfo>
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<mods:name id="6F71C6E86E121CDFE0702BF0DAA0FAF0" type="personal">
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<mods:roleTerm id="3EFE5B384D2B2D17947EAF0B6C2F27C9">Author</mods:roleTerm>
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<mods:namePart id="DD67CECFE20A5054590950BA96986E38">Botero-Trujillo, Ricardo</mods:namePart>
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</mods:name>
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<mods:name id="1C27521BB962F9DB545B712A4A150024" type="personal">
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<mods:roleTerm id="3E87D12806850C5A447DACC827502A26">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4027FBF44C57D2B079692428228AFEB9">Erazo-Moreno, Martha C.</mods:namePart>
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</mods:name>
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<mods:name id="B9905EE971440008BC067904C088924A" type="personal">
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<mods:role id="BAAB835E94004B7E43202B2728460011">
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<mods:roleTerm id="402BA8FD562F4F48E69C3127EF6132D7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EA1F34B48D546C486441A28DF91E5CD8">Pérez, Gustavo A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="413E622D6CCBD491DA7D0E576F1BD40E">text</mods:typeOfResource>
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<mods:title id="A4FF943EBA2060928B499EAC38B0FF2F">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="9C6C2A2AFB124A23A5ED32AF536695ED">
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<mods:date id="C4F4FADCE7436FFD2772A17A9B8594B8">2009</mods:date>
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<mods:number id="06FB33EA5F4A42E53628E14CAE2904DF">2120</mods:number>
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<mods:classification id="03F3B619AF0DC128BC3F403E6CDEB960">journal article</mods:classification>
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<mods:identifier id="FA0B0225706BFB459155127C1F641478" type="DOI">10.5281/zenodo.188069</mods:identifier>
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<mods:identifier id="84E41A5A40EEFC9B4C0CE4B1C1CE648C" type="GBIF-Dataset">5552705a-eca2-454c-be4b-614bae40a4a9</mods:identifier>
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<mods:identifier id="A63EE13E0E871D7B8BE0E63BD71F8EFB" type="ISSN">1175-5326</mods:identifier>
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<treatment id="E9284E34FF83FF90FEE6FC47C8CBFEC4" ID-DOI="http://doi.org/10.5281/zenodo.6214362" ID-GBIF-Taxon="119618501" ID-Zenodo-Dep="6214362" LSID="urn:lsid:plazi:treatment:E9284E34FF83FF90FEE6FC47C8CBFEC4" httpUri="http://treatment.plazi.org/id/E9284E34FF83FF90FEE6FC47C8CBFEC4" lastPageId="9" lastPageNumber="36" pageId="2" pageNumber="29">
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<subSubSection id="299BACA9FF83FF9BFEE6FC47C898FC55" pageId="2" pageNumber="29" type="nomenclature">
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<paragraph id="613EFF22FF83FF9BFEE6FC47CB22FC76" blockId="2.[151,557,938,999]" box="[151,557,938,964]" pageId="2" pageNumber="29">
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<heading id="3A76484EFF83FF9BFEE6FC47CB22FC76" bold="true" box="[151,557,938,964]" fontSize="11" level="1" pageId="2" pageNumber="29" reason="1">
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<emphasis id="53F52330FF83FF9BFEE6FC47CB22FC76" bold="true" box="[151,557,938,964]" pageId="2" pageNumber="29">
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<taxonomicName id="A68184A1FF83FF9BFEE6FC47C8C7FC76" ID-CoL="438SQ" box="[151,456,938,964]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="bivicentorum" status="sp. nov.">
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<emphasis id="53F52330FF83FF9BFEE6FC47C8C7FC76" bold="true" box="[151,456,938,964]" italics="true" pageId="2" pageNumber="29">Microtityus bivicentorum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="48C69E4BFF83FF9BFFBEFC47CB22FC76" box="[463,557,938,964]" pageId="2" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="613EFF22FF83FF9BFEE6FC20C898FC55" blockId="2.[151,557,938,999]" box="[151,407,973,999]" pageId="2" pageNumber="29">
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<figureCitation id="F9BAE3A7FF83FF9BFEE6FC20C83CFC55" box="[151,307,973,999]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 – 5" captionStart-2="FIGURES 6 – 7" captionStart-3="FIGURES 8 – 11" captionStart-4="FIGURES 12 – 17" captionStart-5="FIGURES 18 – 23" captionStartId-0="3.[151,259,1588,1612]" captionStartId-1="5.[151,269,1631,1655]" captionStartId-2="6.[151,269,1163,1187]" captionStartId-3="7.[151,269,1299,1323]" captionStartId-4="8.[151,269,1574,1598]" captionStartId-5="9.[151,273,1902,1926]" captionTargetBox-0="[156,1430,432,1560]" captionTargetBox-1="[156,1433,201,1602]" captionTargetBox-2="[157,1431,201,1136]" captionTargetBox-3="[159,1425,370,1244]" captionTargetBox-4="[157,1429,367,1512]" captionTargetBox-5="[159,1430,456,1842]" captionTargetId-0="figure@3.[151,1436,426,1565]" captionTargetId-1="figure@5.[151,1436,194,1607]" captionTargetId-2="figure@6.[151,1436,194,1139]" captionTargetId-3="figure@7.[151,1436,355,1276]" captionTargetId-4="figure@8.[151,1436,349,1550]" captionTargetId-5="figure@9.[151,1436,432,1894]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionText-0="FIGURE 1. Known distribution of Microtityus bivicentorum sp. nov. (), Microtityus desuzeae (․), Microtityus franckei (˔) and Microtityus joseantonioi (●)." captionText-1="FIGURES 2 – 5. Microtityus bivicentorum sp. nov. 2 – 3. Male holotype, dorsal and ventral views. 4 – 5. Female paratype, dorsal and ventral views. Scale bars = 5 mm." captionText-2="FIGURES 6 – 7. Microtityus bivicentorum sp. nov., male holotype under UV light. 6. Dorsal view. 7. Ventral view. MC = Median carina; MLC = Median lateral carina; LC = Lateral carina; SMC = Submedian carina. Scale bars = 5 mm." captionText-3="FIGURES 8 – 11. Microtityus bivicentorum sp. nov. 8. Sternum, genital operculum, pectinal basal piece, and pectines of male holotype. 9. Sternum, genital operculum, pectinal basal piece, and pectines of female paratype. 10. Sternite V of male holotype. 11. Telson of male holotype, lateral view. Scale bars = 0.5 mm." captionText-4="FIGURES 12 – 17. Microtityus bivicentorum sp. nov., distribution of the trichobothria in male holotype (illustrated from the right pedipalp). 12. Femur, dorsal view. 13. Femur, internal view. 14. Patella, dorsal view. 15. Patella, external view. 16. Chela, external view. 17. Chela, ventral view. Scale bars = 0.5 mm." captionText-5="FIGURES 18 – 23. Right pedipalp chela. 18 – 21. Microtityus bivicentorum sp. nov., male holotype. 18. Detail of the movable finger dentition. 19. Detail of the fixed finger dentition. 20. Chela, external view. 21. Chela, ventral view. 22 – 23. Microtityus franckei, male paratype (MPUJ-SCO- 367). 22. Chela, external view. 23. Chela, ventral view. Scale bars = 0.5 mm." httpUri-0="https://zenodo.org/record/188070/files/figure.png" httpUri-1="https://zenodo.org/record/188071/files/figure.png" httpUri-2="https://zenodo.org/record/188072/files/figure.png" httpUri-3="https://zenodo.org/record/188073/files/figure.png" httpUri-4="https://zenodo.org/record/188074/files/figure.png" httpUri-5="https://zenodo.org/record/188075/files/figure.png" pageId="2" pageNumber="29">Figures 1–21</figureCitation>
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; Table 1
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</paragraph>
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</subSubSection>
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<subSubSection id="299BACA9FF83FF9BFEE6FBF7CAD0FB44" pageId="2" pageNumber="29" type="materials_examined">
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<paragraph id="613EFF22FF83FF9BFEE6FBF7CDF4FB30" blockId="2.[151,1437,1050,2010]" pageId="2" pageNumber="29">
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<emphasis id="53F52330FF83FF9BFEE6FBF7C821FB86" bold="true" box="[151,302,1050,1076]" pageId="2" pageNumber="29">
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<typeStatus id="BE3A4180FF83FF9BFEE6FBF7C9DBFB86" box="[151,212,1050,1076]" pageId="2" pageNumber="29">Type</typeStatus>
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series.
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</emphasis>
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—
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<emphasis id="53F52330FF83FF9BFF3CFBF7C8C8FB81" box="[333,455,1050,1075]" italics="true" pageId="2" pageNumber="29">
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<typeStatus id="BE3A4180FF83FF9BFF3CFBF7C8CFFB81" box="[333,448,1050,1075]" pageId="2" pageNumber="29" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<collectingCountry id="1996BFB2FF83FF9BFFA1FBF7CB7AFB86" box="[464,629,1050,1076]" name="Colombia" pageId="2" pageNumber="29">COLOMBIA</collectingCountry>
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:
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<emphasis id="53F52330FF83FF9BFCF0FBF7CA61FB81" box="[641,878,1050,1075]" italics="true" pageId="2" pageNumber="29">Cesar Department:</emphasis>
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adult ♂ from Valledupar, Cerro Los Besotes, Santuario de Vida Silvestre Los Besotes, Camino Los Carretos,
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<geoCoordinate id="04B599E5FF83FF9BFDB6FBACCD74FBE9" box="[967,1147,1089,1115]" direction="north" orientation="latitude" pageId="2" pageNumber="29" precision="1" value="10.573611">10º34’25.0’’N</geoCoordinate>
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<geoCoordinate id="04B599E5FF83FF9BFAF5FBACCC30FBE9" box="[1156,1343,1089,1115]" direction="west" orientation="longitude" pageId="2" pageNumber="29" precision="1" value="-73.26342">73º15’48.3’’W</geoCoordinate>
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,
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<quantity id="A67952C7FF83FF9BFB3CFBACCC92FBE9" box="[1357,1437,1089,1115]" metricMagnitude="2" metricUnit="m" metricValue="3.66" pageId="2" pageNumber="29" unit="m" value="366.0">366 m</quantity>
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elevation, Pitfall trap,
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<date id="153FD9E2FF83FF9BFFEDFB85CB4CFB30" box="[412,579,1128,1154]" pageId="2" pageNumber="29" value="2008-04-20">20 April 2008</date>
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, M.C. Erazo-Moreno & G.A. Pérez coll. (MPUJ-SC0-380).
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</paragraph>
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<paragraph id="613EFF22FF83FF9BFEB4FB62CAD0FB44" blockId="2.[151,1437,1050,2010]" pageId="2" pageNumber="29">
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<emphasis id="53F52330FF83FF9BFEB4FB62C833FB1A" box="[197,316,1167,1192]" italics="true" pageId="2" pageNumber="29">
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<typeStatus id="BE3A4180FF83FF9BFEB4FB62C839FB1A" box="[197,310,1167,1192]" pageId="2" pageNumber="29" type="paratype">Paratype</typeStatus>
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:
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</emphasis>
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<collectingCountry id="1996BFB2FF83FF9BFF35FB63C8E8FB1A" box="[324,487,1166,1192]" name="Colombia" pageId="2" pageNumber="29">COLOMBIA</collectingCountry>
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:
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<emphasis id="53F52330FF83FF9BFF82FB62CBD4FB1A" box="[499,731,1167,1192]" italics="true" pageId="2" pageNumber="29">Cesar Department:</emphasis>
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adult ♀ from Valledupar, Cerro Los Besotes, Santuario de Vida Silvestre Los Besotes, Manantial Los Zainos,
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<geoCoordinate id="04B599E5FF83FF9BFC8FFB58CAA5FB7D" box="[766,938,1205,1231]" direction="north" orientation="latitude" pageId="2" pageNumber="29" precision="1" value="10.573527">10°34’24.7’’N</geoCoordinate>
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<geoCoordinate id="04B599E5FF83FF9BFDC2FB58CD6AFB7D" box="[947,1125,1205,1231]" direction="west" orientation="longitude" pageId="2" pageNumber="29" precision="1" value="-73.273895">73°16’26.0’’W</geoCoordinate>
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,
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<quantity id="A67952C7FF83FF9BFA02FB58CDB0FB7D" box="[1139,1215,1205,1231]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="29" unit="m" value="200.0">200 m</quantity>
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elevation,
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<emphasis id="53F52330FF83FF9BFB35FB58CC9AFB7C" box="[1348,1429,1205,1230]" italics="true" pageId="2" pageNumber="29">ad hoc</emphasis>
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, under stone, at night,
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<date id="153FD9E2FF83FF9BFFE4FB31CB34FB44" box="[405,571,1244,1270]" pageId="2" pageNumber="29" value="2008-04-23">23 April 2008</date>
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, G.A. Pérez coll. (MPUJ-SC0-381).
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</paragraph>
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</subSubSection>
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<subSubSection id="299BACA9FF83FF9BFEB4FAEFCA11FAAE" box="[197,798,1282,1308]" pageId="2" pageNumber="29" type="distribution">
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<paragraph id="613EFF22FF83FF9BFEB4FAEFCA11FAAE" blockId="2.[151,1437,1050,2010]" box="[197,798,1282,1308]" pageId="2" pageNumber="29">
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<emphasis id="53F52330FF83FF9BFEB4FAEFC86AFAAE" bold="true" box="[197,357,1282,1308]" pageId="2" pageNumber="29">Distribution.</emphasis>
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—Known only from the
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<typeStatus id="BE3A4180FF83FF9BFCF6FAEFCBB5FAAE" box="[647,698,1282,1308]" pageId="2" pageNumber="29">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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<subSubSection id="299BACA9FF83FF9BFEB4FAC5CBDDFA05" pageId="2" pageNumber="29" type="etymology">
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<paragraph id="613EFF22FF83FF9BFEB4FAC5CBDDFA05" blockId="2.[151,1437,1050,2010]" pageId="2" pageNumber="29">
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<emphasis id="53F52330FF83FF9BFEB4FAC5C856FAF0" bold="true" box="[197,345,1320,1346]" pageId="2" pageNumber="29">Etymology.</emphasis>
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—The species name is dedicated to Jose Vicente Rodríguez-Mahecha and Jose Vicente Rueda-Almonacid, formerly at "Conservación Internacional" in Bogotá. It is derived from the junction of the prefix "Bi" (meaning two or double) and the personal name "Vicente", a combination that takes advantage to celebrate the academic labor of both individuals.
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</paragraph>
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</subSubSection>
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<subSubSection id="299BACA9FF83FF9BFEB4FA2ECB1EF97E" pageId="2" pageNumber="29" type="diagnosis">
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<paragraph id="613EFF22FF83FF9BFEB4FA2ECB1EF97E" blockId="2.[151,1437,1050,2010]" pageId="2" pageNumber="29">
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<emphasis id="53F52330FF83FF9BFEB4FA2EC84BFA6F" bold="true" box="[197,324,1475,1501]" pageId="2" pageNumber="29">Diagnosis.</emphasis>
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—
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<taxonomicName id="A68184A1FF83FF9BFF13FA29CB85FA6F" box="[354,650,1476,1501]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="bivicentorum" status="sp. nov.">
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<emphasis id="53F52330FF83FF9BFF13FA29CB85FA6F" box="[354,650,1476,1501]" italics="true" pageId="2" pageNumber="29">Microtityus bivicentorum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="48C69E4BFF83FF9BFCE0FA29CBE4FA6C" box="[657,747,1476,1502]" pageId="2" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
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differs from all other species described in the genus by the following unique combination of features: pedipalps with reductive neobothriotaxy, with trichobothria
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<emphasis id="53F52330FF83FF9BFBF4FA06CC93F9BA" box="[1413,1436,1515,1544]" italics="true" pageId="2" pageNumber="29">d2</emphasis>
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absent on femur and patella, and
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<emphasis id="53F52330FF83FF9BFC56F9F9CB5FF980" box="[551,592,1556,1586]" italics="true" pageId="2" pageNumber="29">Eb3</emphasis>
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,
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<emphasis id="53F52330FF83FF9BFC11F9F9CB82F99F" box="[608,653,1556,1581]" italics="true" pageId="2" pageNumber="29">Esb</emphasis>
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and
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<emphasis id="53F52330FF83FF9BFCB9F9F9CBFFF99F" box="[712,752,1556,1581]" italics="true" pageId="2" pageNumber="29">esb</emphasis>
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absent on chela (
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<figureCitation id="F9BAE3A7FF83FF9BFDB2F9F9CD59F99C" box="[963,1110,1556,1582]" captionStart="FIGURES 12 – 17" captionStartId="8.[151,269,1574,1598]" captionTargetBox="[157,1429,367,1512]" captionTargetId="figure@8.[151,1436,349,1550]" captionTargetPageId="8" captionText="FIGURES 12 – 17. Microtityus bivicentorum sp. nov., distribution of the trichobothria in male holotype (illustrated from the right pedipalp). 12. Femur, dorsal view. 13. Femur, internal view. 14. Patella, dorsal view. 15. Patella, external view. 16. Chela, external view. 17. Chela, ventral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188074/files/figure.png" pageId="2" pageNumber="29">Figs. 12–17</figureCitation>
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); trichobothrium
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<emphasis id="53F52330FF83FF9BFB59F9F9CC32F99F" box="[1320,1341,1556,1581]" italics="true" pageId="2" pageNumber="29">et</emphasis>
|
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located between
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<emphasis id="53F52330FF83FF9BFF73F9D3C810F9E5" box="[258,287,1598,1623]" italics="true" pageId="2" pageNumber="29">db</emphasis>
|
||
and
|
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<emphasis id="53F52330FF83FF9BFF2BF9D3C87EF9E5" box="[346,369,1598,1623]" italics="true" pageId="2" pageNumber="29">dt</emphasis>
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on pedipalp fixed finger (
|
||
<figureCitation id="F9BAE3A7FF83FF9BFCDAF9D3CA0BF9EA" box="[683,772,1598,1624]" captionStart="FIGURES 12 – 17" captionStartId="8.[151,269,1574,1598]" captionTargetBox="[157,1429,367,1512]" captionTargetId="figure@8.[151,1436,349,1550]" captionTargetPageId="8" captionText="FIGURES 12 – 17. Microtityus bivicentorum sp. nov., distribution of the trichobothria in male holotype (illustrated from the right pedipalp). 12. Femur, dorsal view. 13. Femur, internal view. 14. Patella, dorsal view. 15. Patella, external view. 16. Chela, external view. 17. Chela, ventral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188074/files/figure.png" pageId="2" pageNumber="29">Fig. 16</figureCitation>
|
||
); movable finger with 11–12 rows of granules; pectinal basal piece quadrangular in shape in both sexes, lacking posterior projection (
|
||
<figureCitation id="F9BAE3A7FF83FF9BFA40F989CDAAF9CC" box="[1073,1189,1636,1662]" captionStart="FIGURES 8 – 11" captionStartId="7.[151,269,1299,1323]" captionTargetBox="[159,1425,370,1244]" captionTargetId="figure@7.[151,1436,355,1276]" captionTargetPageId="7" captionText="FIGURES 8 – 11. Microtityus bivicentorum sp. nov. 8. Sternum, genital operculum, pectinal basal piece, and pectines of male holotype. 9. Sternum, genital operculum, pectinal basal piece, and pectines of female paratype. 10. Sternite V of male holotype. 11. Telson of male holotype, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188073/files/figure.png" pageId="2" pageNumber="29">Figs. 8–9</figureCitation>
|
||
); sternite V of males with three smooth, hyaline areas, of which the median is shaped like an equilateral triangle, and the laterals are large and flattened (
|
||
<figureCitation id="F9BAE3A7FF83FF9BFFDAF95FCB0CF97E" box="[427,515,1714,1740]" captionStart="FIGURES 8 – 11" captionStartId="7.[151,269,1299,1323]" captionTargetBox="[159,1425,370,1244]" captionTargetId="figure@7.[151,1436,355,1276]" captionTargetPageId="7" captionText="FIGURES 8 – 11. Microtityus bivicentorum sp. nov. 8. Sternum, genital operculum, pectinal basal piece, and pectines of male holotype. 9. Sternum, genital operculum, pectinal basal piece, and pectines of female paratype. 10. Sternite V of male holotype. 11. Telson of male holotype, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188073/files/figure.png" pageId="2" pageNumber="29">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="299BACA9FF83FF9EFEB4F935C87DF8A6" lastPageId="7" lastPageNumber="34" pageId="2" pageNumber="29" type="description">
|
||
<paragraph id="613EFF22FF83FF9BFEB4F935CB6EF940" blockId="2.[151,1437,1050,2010]" box="[197,609,1752,1778]" pageId="2" pageNumber="29">
|
||
<emphasis id="53F52330FF83FF9BFEB4F935CB6EF940" bold="true" box="[197,609,1752,1778]" pageId="2" pageNumber="29">
|
||
Description of the male
|
||
<typeStatus id="BE3A4180FF83FF9BFF9EF935CB53F940" box="[495,604,1752,1778]" pageId="2" pageNumber="29" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="613EFF22FF83FF9AFEB4F912CD59FEC1" blockId="2.[151,1437,1050,2010]" lastBlockId="3.[151,1436,152,371]" lastPageId="3" lastPageNumber="30" pageId="2" pageNumber="29">
|
||
<emphasis id="53F52330FF83FF9BFEB4F912C85DF8AB" bold="true" box="[197,338,1791,1817]" italics="true" pageId="2" pageNumber="29">Coloration.</emphasis>
|
||
—Carapace predominantly yellow with abundant dark-brown mottling throughout. Median and lateral ocular tubercles blackish. Chelicerae yellow with some brownish areas located basally on both fixed and movable fingers; teeth reddish. Tergites predominantly yellow; I–VI with brown regions restricted to the posterior margin and separated from each other by the longitudinal carinae which are yellow, and with an anteromedian brownish region which is darker in tergites IV–VI; VII with brown-colored areas laterally and dorsoanteriorly. Coxosternal region, genital operculum, pectines, pectinal basal piece and sternites completely yellow and spotless. Metasoma predominantly yellow with some brown areas. Telson with vesicle, subaculear tubercle and basal half of aculeus immaculately yellow and spotless; distal half of aculeus dark-red. Pedipalps predominantly yellow, with brownish areas over dorsal, internal and external surfaces of femur and patella, and dorsal and external surfaces of chela; ventral surfaces of all segments and internal surface of chela completely yellow; fixed and movable fingers reddish brown, darker basally. Legs with variegated pigmentation in all segments, except for the telotarsi which are yellow.
|
||
</paragraph>
|
||
<caption id="35FEAFAAFF82FF9AFEE6F9D9CB86F9DC" httpUri="https://zenodo.org/record/188070/files/figure.png" pageId="3" pageNumber="30" targetBox="[156,1430,432,1560]" targetPageId="3">
|
||
<paragraph id="613EFF22FF82FF9AFEE6F9D9CB86F9DC" blockId="3.[151,1436,1588,1647]" pageId="3" pageNumber="30">
|
||
<emphasis id="53F52330FF82FF9AFEE6F9D9C82FF9FE" bold="true" box="[151,288,1588,1612]" pageId="3" pageNumber="30">FIGURE 1.</emphasis>
|
||
Known distribution of
|
||
<taxonomicName id="A68184A1FF82FF9AFC5DF9D8CA44F9FE" box="[556,843,1589,1612]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="bivicentorum" status="sp. nov.">
|
||
<emphasis id="53F52330FF82FF9AFC5DF9D8CA44F9FE" box="[556,843,1589,1612]" italics="true" pageId="3" pageNumber="30">Microtityus bivicentorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="53F52330FF82FF9AFD22F9D9CAA1F9FE" bold="true" box="[851,942,1588,1612]" pageId="3" pageNumber="30">
|
||
<taxonomicNameLabel id="48C69E4BFF82FF9AFD22F9D9CAA1F9FE" box="[851,942,1588,1612]" pageId="3" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(),
|
||
<taxonomicName id="A68184A1FF82FF9AFD80F9D8CDEFF9FE" box="[1009,1248,1589,1612]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="desuzeae">
|
||
<emphasis id="53F52330FF82FF9AFD80F9D8CDEFF9FE" box="[1009,1248,1589,1612]" italics="true" pageId="3" pageNumber="30">Microtityus desuzeae</emphasis>
|
||
</taxonomicName>
|
||
(․),
|
||
<taxonomicName id="A68184A1FF82FF9AFB6AF9D8C9E1F9DD" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="franckei">
|
||
<emphasis id="53F52330FF82FF9AFB6AF9D8C9E1F9DD" italics="true" pageId="3" pageNumber="30">Microtityus franckei</emphasis>
|
||
</taxonomicName>
|
||
(˔) and
|
||
<taxonomicName id="A68184A1FF82FF9AFF24F9B5CB57F9DD" box="[341,600,1624,1647]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="joseantonioi">
|
||
<emphasis id="53F52330FF82FF9AFF24F9B5CB57F9DD" box="[341,600,1624,1647]" italics="true" pageId="3" pageNumber="30">Microtityus joseantonioi</emphasis>
|
||
</taxonomicName>
|
||
(●).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="613EFF22FF82FF9DFEB7F970C85DFE92" blockId="3.[151,1437,1693,2018]" lastBlockId="4.[151,1437,151,338]" lastPageId="4" lastPageNumber="31" pageId="3" pageNumber="30">
|
||
<emphasis id="53F52330FF82FF9AFEB7F970C8F3F904" box="[198,508,1693,1719]" italics="true" pageId="3" pageNumber="30">
|
||
<emphasis id="53F52330FF82FF9AFEB7F970C86FF905" bold="true" box="[198,352,1693,1719]" italics="true" pageId="3" pageNumber="30">Morphology.</emphasis>
|
||
—Carapace:
|
||
</emphasis>
|
||
Subtriangular, densely granulose throughout; anterior margin deeply emarginated; median ocular tubercle slightly anterior to the centre of the carapace; lateral ocular tubercles each with three ocelli; anterior median, supercialiary, central median and posterior median carinae pronounced and elevated; lateral ocular carinae vestigial; anterior marginal, anterior median and median ocular furrows deep; posterior marginal, posterior median, lateral ocular, central lateral and central median furrows shallow; other carinae and furrows absent.
|
||
<emphasis id="53F52330FF82FF9AFC86F8B5CA55F8C3" box="[759,858,1880,1905]" italics="true" pageId="3" pageNumber="30">Tergites:</emphasis>
|
||
With similar granulation to that of the carapace; five longitudinal carinae (median; paired median lateral and lateral carinae) are present on the posterior half of tergites I–IV, the lateral carinae vestigial (
|
||
<figureCitation id="F9BAE3A7FF82FF9AFCEEF84FCBE4F80E" box="[671,747,1954,1980]" captionStart="FIGURES 6 – 7" captionStartId="6.[151,269,1163,1187]" captionTargetBox="[157,1431,201,1136]" captionTargetId="figure@6.[151,1436,194,1139]" captionTargetPageId="6" captionText="FIGURES 6 – 7. Microtityus bivicentorum sp. nov., male holotype under UV light. 6. Dorsal view. 7. Ventral view. MC = Median carina; MLC = Median lateral carina; LC = Lateral carina; SMC = Submedian carina. Scale bars = 5 mm." httpUri="https://zenodo.org/record/188072/files/figure.png" pageId="3" pageNumber="30">Fig. 6</figureCitation>
|
||
); tergites V–VI with evident median and median lateral carinae of similar development than in the preceding tergites, but lateral carinae apparently absent (
|
||
<figureCitation id="F9BAE3A7FF82FF9AFB52F825CC64F850" box="[1315,1387,1992,2018]" captionStart="FIGURES 6 – 7" captionStartId="6.[151,269,1163,1187]" captionTargetBox="[157,1431,201,1136]" captionTargetId="figure@6.[151,1436,194,1139]" captionTargetPageId="6" captionText="FIGURES 6 – 7. Microtityus bivicentorum sp. nov., male holotype under UV light. 6. Dorsal view. 7. Ventral view. MC = Median carina; MLC = Median lateral carina; LC = Lateral carina; SMC = Submedian carina. Scale bars = 5 mm." httpUri="https://zenodo.org/record/188072/files/figure.png" pageId="3" pageNumber="30">Fig. 6</figureCitation>
|
||
);
|
||
<emphasis id="53F52330FF85FF9DFEE6FF76C81EFF01" bold="true" box="[151,273,155,179]" pageId="4" pageNumber="31">TABLE 1.</emphasis>
|
||
Meristic data for
|
||
<taxonomicName id="A68184A1FF85FF9DFF92FF76CA0AFF00" box="[483,773,155,178]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="bivicentorum" status="sp. nov.">
|
||
<emphasis id="53F52330FF85FF9DFF92FF76CA0AFF00" box="[483,773,155,178]" italics="true" pageId="4" pageNumber="31">Microtityus bivicentorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="48C69E4BFF85FF9DFD7CFF76CA66FF01" box="[781,873,155,179]" pageId="4" pageNumber="31" rank="species">
|
||
<emphasis id="53F52330FF85FF9DFD7CFF76CA66FF01" bold="true" box="[781,873,155,179]" pageId="4" pageNumber="31">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Measurements in millimeters. 1Sum of prosoma, mesosoma and metasoma. 2Sum of tergites
|
||
<date id="153FD9E2FF85FF9DFC1CFF2CCBB2FF79" box="[621,701,190,216]" pageId="4" pageNumber="31">I–VII. 3</date>
|
||
Sum of metasomal segments I–V and telson. 4Sum of femur, patella and chela. 5Measured from the commissure of the junction with the movable finger to the finger tip. 6Measured along the marginal lamella.
|
||
</paragraph>
|
||
<paragraph id="613EFF22FF85FF9DFDFCFED6CDB1F850" blockId="4.[151,1437,151,338]" lastBlockId="4.[159,1227,1865,2018]" pageId="4" pageNumber="31">
|
||
Male
|
||
<typeStatus id="BE3A4180FF85FF9DFDBAFED6CD27FEE0" box="[971,1064,315,338]" pageId="4" pageNumber="31" type="holotype">holotype</typeStatus>
|
||
Female
|
||
<typeStatus id="BE3A4180FF85FF9DFA80FED6CC42FEE0" box="[1265,1357,315,338]" pageId="4" pageNumber="31" type="paratype">paratype</typeStatus>
|
||
movable finger length: 1.44 2.00 palm length: 1.23 1.36 Pectines: length:6 0.86 0.90 teeth count (left/right): 9/9 8/8
|
||
</paragraph>
|
||
<paragraph id="613EFF22FF85FF9DFEEEFE84CDC4F880" pageId="4" pageNumber="31">
|
||
<table id="13810D82FF850066FEE6FE8BCC92F887" box="[151,1437,358,1845]" gridcols="3" gridrows="23" pageId="4" pageNumber="31">
|
||
<tr id="DFB1FD60FF850066FEE6FE8BCC92FE18" box="[151,1437,358,426]" gridrow="0" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6FE8BCBD0FE18" box="[151,735,358,426]" gridcol="0" gridrow="0" pageId="4" pageNumber="31">Total body length:1 Carapace: length:</th>
|
||
<th id="9C60941CFF850066FDFCFE8BCAC6FE18" box="[909,969,358,426]" gridcol="1" gridrow="0" pageId="4" pageNumber="31">12.94 1.80</th>
|
||
<th id="9C60941CFF850066FAEDFE8BCC92FE18" box="[1180,1437,358,426]" gridcol="2" gridrow="0" pageId="4" pageNumber="31">15.78 2.12</th>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6FE53CC92FE67" box="[151,1437,446,469]" gridrow="1" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6FE53CBD0FE67" box="[151,735,446,469]" gridcol="0" gridrow="1" pageId="4" pageNumber="31">anterior width:</th>
|
||
<td id="9C60941CFF850066FDFCFE53CAC6FE67" box="[909,969,446,469]" gridcol="1" gridrow="1" pageId="4" pageNumber="31">0.74</td>
|
||
<td id="9C60941CFF850066FAEDFE53CC92FE67" box="[1180,1437,446,469]" gridcol="2" gridrow="1" pageId="4" pageNumber="31">1.00</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6FE04CC92FD9F" box="[151,1437,489,557]" gridrow="2" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6FE04CBD0FD9F" box="[151,735,489,557]" gridcol="0" gridrow="2" pageId="4" pageNumber="31">posterior width: Mesosoma: total length:2</th>
|
||
<td id="9C60941CFF850066FDFCFE04CAC6FD9F" box="[909,969,489,557]" gridcol="1" gridrow="2" pageId="4" pageNumber="31">1.90 3.23</td>
|
||
<td id="9C60941CFF850066FAEDFE04CC92FD9F" box="[1180,1437,489,557]" gridcol="2" gridrow="2" pageId="4" pageNumber="31">2.50 4.40</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6FDACCC92FDE8" box="[151,1437,577,602]" gridrow="3" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6FDACCBD0FDE8" box="[151,735,577,602]" gridcol="0" gridrow="3" pageId="4" pageNumber="31">Metasoma: total length:3</th>
|
||
<td id="9C60941CFF850066FDFCFDACCAC6FDE8" box="[909,969,577,602]" gridcol="1" gridrow="3" pageId="4" pageNumber="31">7.91</td>
|
||
<td id="9C60941CFF850066FAEDFDACCC92FDE8" box="[1180,1437,577,602]" gridcol="2" gridrow="3" pageId="4" pageNumber="31">9.26</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6FD83CC92FD37" box="[151,1437,622,645]" gridrow="4" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6FD83CBD0FD37" box="[151,735,622,645]" gridcol="0" gridrow="4" pageId="4" pageNumber="31">Metasomal segment I: length:</th>
|
||
<td id="9C60941CFF850066FDFCFD83CAC6FD37" box="[909,969,622,645]" gridcol="1" gridrow="4" pageId="4" pageNumber="31">0.91</td>
|
||
<td id="9C60941CFF850066FAEDFD83CC92FD37" box="[1180,1437,622,645]" gridcol="2" gridrow="4" pageId="4" pageNumber="31">1.10</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6FD74CC92FD68" box="[151,1437,665,730]" gridrow="5" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6FD74CBD0FD68" box="[151,735,665,730]" gridcol="0" gridrow="5" pageId="4" pageNumber="31">width: depth:</th>
|
||
<td id="9C60941CFF850066FDFCFD74CAC6FD68" box="[909,969,665,730]" gridcol="1" gridrow="5" pageId="4" pageNumber="31">0.93 0.80</td>
|
||
<td id="9C60941CFF850066FAEDFD74CC92FD68" box="[1180,1437,665,730]" gridcol="2" gridrow="5" pageId="4" pageNumber="31">1.10 0.94</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6FD03CC92FCB7" box="[151,1437,750,773]" gridrow="6" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6FD03CBD0FCB7" box="[151,735,750,773]" gridcol="0" gridrow="6" pageId="4" pageNumber="31">Metasomal segment II: length:</th>
|
||
<td id="9C60941CFF850066FDFCFD03CAC6FCB7" box="[909,969,750,773]" gridcol="1" gridrow="6" pageId="4" pageNumber="31">1.12</td>
|
||
<td id="9C60941CFF850066FAEDFD03CC92FCB7" box="[1180,1437,750,773]" gridcol="2" gridrow="6" pageId="4" pageNumber="31">1.36</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6FCF4CC92FCE8" box="[151,1437,793,858]" gridrow="7" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6FCF4CBD0FCE8" box="[151,735,793,858]" gridcol="0" gridrow="7" pageId="4" pageNumber="31">width: depth:</th>
|
||
<td id="9C60941CFF850066FDFCFCF4CAC6FCE8" box="[909,969,793,858]" gridcol="1" gridrow="7" pageId="4" pageNumber="31">0.83 0.80</td>
|
||
<td id="9C60941CFF850066FAEDFCF4CC92FCE8" box="[1180,1437,793,858]" gridcol="2" gridrow="7" pageId="4" pageNumber="31">1.00 0.90</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6FC83CC92FC37" box="[151,1437,878,901]" gridrow="8" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6FC83CBD0FC37" box="[151,735,878,901]" gridcol="0" gridrow="8" pageId="4" pageNumber="31">Metasomal segment III: length:</th>
|
||
<td id="9C60941CFF850066FDFCFC83CAC6FC37" box="[909,969,878,901]" gridcol="1" gridrow="8" pageId="4" pageNumber="31">1.22</td>
|
||
<td id="9C60941CFF850066FAEDFC83CC92FC37" box="[1180,1437,878,901]" gridcol="2" gridrow="8" pageId="4" pageNumber="31">1.44</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6FC74CC92FC68" box="[151,1437,921,986]" gridrow="9" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6FC74CBD0FC68" box="[151,735,921,986]" gridcol="0" gridrow="9" pageId="4" pageNumber="31">width: depth:</th>
|
||
<td id="9C60941CFF850066FDFCFC74CAC6FC68" box="[909,969,921,986]" gridcol="1" gridrow="9" pageId="4" pageNumber="31">0.80 0.75</td>
|
||
<td id="9C60941CFF850066FAEDFC74CC92FC68" box="[1180,1437,921,986]" gridcol="2" gridrow="9" pageId="4" pageNumber="31">0.94 0.96</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6FC03CC92FBB7" box="[151,1437,1006,1029]" gridrow="10" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6FC03CBD0FBB7" box="[151,735,1006,1029]" gridcol="0" gridrow="10" pageId="4" pageNumber="31">Metasomal segment IV: length:</th>
|
||
<td id="9C60941CFF850066FDFCFC03CAC6FBB7" box="[909,969,1006,1029]" gridcol="1" gridrow="10" pageId="4" pageNumber="31">1.36</td>
|
||
<td id="9C60941CFF850066FAEDFC03CC92FBB7" box="[1180,1437,1006,1029]" gridcol="2" gridrow="10" pageId="4" pageNumber="31">1.56</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6FBF4CC92FBE8" box="[151,1437,1049,1114]" gridrow="11" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6FBF4CBD0FBE8" box="[151,735,1049,1114]" gridcol="0" gridrow="11" pageId="4" pageNumber="31">width: depth:</th>
|
||
<td id="9C60941CFF850066FDFCFBF4CAC6FBE8" box="[909,969,1049,1114]" gridcol="1" gridrow="11" pageId="4" pageNumber="31">0.77 0.75</td>
|
||
<td id="9C60941CFF850066FAEDFBF4CC92FBE8" box="[1180,1437,1049,1114]" gridcol="2" gridrow="11" pageId="4" pageNumber="31">0.90 0.90</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6FB83CC92FB37" box="[151,1437,1134,1157]" gridrow="12" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6FB83CBD0FB37" box="[151,735,1134,1157]" gridcol="0" gridrow="12" pageId="4" pageNumber="31">Metasomal segment V: length:</th>
|
||
<td id="9C60941CFF850066FDFCFB83CAC6FB37" box="[909,969,1134,1157]" gridcol="1" gridrow="12" pageId="4" pageNumber="31">1.80</td>
|
||
<td id="9C60941CFF850066FAEDFB83CC92FB37" box="[1180,1437,1134,1157]" gridcol="2" gridrow="12" pageId="4" pageNumber="31">2.00</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6FB74CC92FB68" box="[151,1437,1177,1242]" gridrow="13" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6FB74CBD0FB68" box="[151,735,1177,1242]" gridcol="0" gridrow="13" pageId="4" pageNumber="31">width: depth:</th>
|
||
<td id="9C60941CFF850066FDFCFB74CAC6FB68" box="[909,969,1177,1242]" gridcol="1" gridrow="13" pageId="4" pageNumber="31">0.72 0.72</td>
|
||
<td id="9C60941CFF850066FAEDFB74CC92FB68" box="[1180,1437,1177,1242]" gridcol="2" gridrow="13" pageId="4" pageNumber="31">0.84 0.84</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6FB03CC92FAB7" box="[151,1437,1262,1285]" gridrow="14" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6FB03CBD0FAB7" box="[151,735,1262,1285]" gridcol="0" gridrow="14" pageId="4" pageNumber="31">Telson: length:</th>
|
||
<td id="9C60941CFF850066FDFCFB03CAC6FAB7" box="[909,969,1262,1285]" gridcol="1" gridrow="14" pageId="4" pageNumber="31">1.50</td>
|
||
<td id="9C60941CFF850066FAEDFB03CC92FAB7" box="[1180,1437,1262,1285]" gridcol="2" gridrow="14" pageId="4" pageNumber="31">1.80</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6FAF4CC92FAE8" box="[151,1437,1305,1370]" gridrow="15" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6FAF4CBD0FAE8" box="[151,735,1305,1370]" gridcol="0" gridrow="15" pageId="4" pageNumber="31">vesicle width: vesicle depth:</th>
|
||
<td id="9C60941CFF850066FDFCFAF4CAC6FAE8" box="[909,969,1305,1370]" gridcol="1" gridrow="15" pageId="4" pageNumber="31">0.64 0.61</td>
|
||
<td id="9C60941CFF850066FAEDFAF4CC92FAE8" box="[1180,1437,1305,1370]" gridcol="2" gridrow="15" pageId="4" pageNumber="31">0.76 0.74</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6FA83CC92FA00" box="[151,1437,1390,1458]" gridrow="16" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6FA83CBD0FA00" box="[151,735,1390,1458]" gridcol="0" gridrow="16" pageId="4" pageNumber="31">Pedipalp: total length:4 Pedipalp femur: length:</th>
|
||
<td id="9C60941CFF850066FDFCFA83CAC6FA00" box="[909,969,1390,1458]" gridcol="1" gridrow="16" pageId="4" pageNumber="31">5.31 1.33</td>
|
||
<td id="9C60941CFF850066FAEDFA83CC92FA00" box="[1180,1437,1390,1458]" gridcol="2" gridrow="16" pageId="4" pageNumber="31">6.98 1.80</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6FA2BCC92FA6F" box="[151,1437,1478,1501]" gridrow="17" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6FA2BCBD0FA6F" box="[151,735,1478,1501]" gridcol="0" gridrow="17" pageId="4" pageNumber="31">width:</th>
|
||
<td id="9C60941CFF850066FDFCFA2BCAC6FA6F" box="[909,969,1478,1501]" gridcol="1" gridrow="17" pageId="4" pageNumber="31">0.48</td>
|
||
<td id="9C60941CFF850066FAEDFA2BCC92FA6F" box="[1180,1437,1478,1501]" gridcol="2" gridrow="17" pageId="4" pageNumber="31">0.61</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6FA1CCC92F980" box="[151,1437,1521,1586]" gridrow="18" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6FA1CCBD0F980" box="[151,735,1521,1586]" gridcol="0" gridrow="18" pageId="4" pageNumber="31">depth: Pedipalp patella: length:</th>
|
||
<td id="9C60941CFF850066FDFCFA1CCAC6F980" box="[909,969,1521,1586]" gridcol="1" gridrow="18" pageId="4" pageNumber="31">0.29 1.58</td>
|
||
<td id="9C60941CFF850066FAEDFA1CCC92F980" box="[1180,1437,1521,1586]" gridcol="2" gridrow="18" pageId="4" pageNumber="31">0.40 2.18</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6F9ABCC92F9EF" box="[151,1437,1606,1629]" gridrow="19" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6F9ABCBD0F9EF" box="[151,735,1606,1629]" gridcol="0" gridrow="19" pageId="4" pageNumber="31">width:</th>
|
||
<td id="9C60941CFF850066FDFCF9ABCAC6F9EF" box="[909,969,1606,1629]" gridcol="1" gridrow="19" pageId="4" pageNumber="31">0.64</td>
|
||
<td id="9C60941CFF850066FAEDF9ABCC92F9EF" box="[1180,1437,1606,1629]" gridcol="2" gridrow="19" pageId="4" pageNumber="31">0.83</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6F99CCC92F900" box="[151,1437,1649,1714]" gridrow="20" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6F99CCBD0F900" box="[151,735,1649,1714]" gridcol="0" gridrow="20" pageId="4" pageNumber="31">depth: Pedipalp chela: length:</th>
|
||
<td id="9C60941CFF850066FDFCF99CCAC6F900" box="[909,969,1649,1714]" gridcol="1" gridrow="20" pageId="4" pageNumber="31">0.48 2.40</td>
|
||
<td id="9C60941CFF850066FAEDF99CCC92F900" box="[1180,1437,1649,1714]" gridcol="2" gridrow="20" pageId="4" pageNumber="31">0.67 3.00</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6F92BCC92F96F" box="[151,1437,1734,1757]" gridrow="21" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6F92BCBD0F96F" box="[151,735,1734,1757]" gridcol="0" gridrow="21" pageId="4" pageNumber="31">width:</th>
|
||
<td id="9C60941CFF850066FDFCF92BCAC6F96F" box="[909,969,1734,1757]" gridcol="1" gridrow="21" pageId="4" pageNumber="31">0.75</td>
|
||
<td id="9C60941CFF850066FAEDF92BCC92F96F" box="[1180,1437,1734,1757]" gridcol="2" gridrow="21" pageId="4" pageNumber="31">0.80</td>
|
||
</tr>
|
||
<tr id="DFB1FD60FF850066FEE6F91CCC92F887" box="[151,1437,1777,1845]" gridrow="22" pageId="4" pageNumber="31">
|
||
<th id="9C60941CFF850066FEE6F91CCBD0F887" box="[151,735,1777,1845]" gridcol="0" gridrow="22" pageId="4" pageNumber="31">depth: fixed finger length:5</th>
|
||
<td id="9C60941CFF850066FDFCF91CCAC6F887" box="[909,969,1777,1845]" gridcol="1" gridrow="22" pageId="4" pageNumber="31">0.77 1.20</td>
|
||
<td id="9C60941CFF850066FAEDF91CCC92F887" box="[1180,1437,1777,1845]" gridcol="2" gridrow="22" pageId="4" pageNumber="31">0.83 1.54</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="35FEAFAAFF84FF9CFEE6F9B2CB79F92B" httpUri="https://zenodo.org/record/188071/files/figure.png" pageId="5" pageNumber="32" targetBox="[156,1433,201,1602]" targetPageId="5">
|
||
<paragraph id="613EFF22FF84FF9CFEE6F9B2CB79F92B" blockId="5.[151,1436,1631,1689]" pageId="5" pageNumber="32">
|
||
<emphasis id="53F52330FF84FF9CFEE6F9B2C84CF9C5" bold="true" box="[151,323,1631,1655]" pageId="5" pageNumber="32">FIGURES 2–5.</emphasis>
|
||
<taxonomicName id="A68184A1FF84FF9CFF3BF98DCB57F9C5" box="[330,600,1632,1655]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="bivicentorum" status="sp. nov.">
|
||
<emphasis id="53F52330FF84FF9CFF3BF98DCB57F9C5" box="[330,600,1632,1655]" italics="true" pageId="5" pageNumber="32">Microtityus bivicentorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="53F52330FF84FF9CFC2EF9B2CBE4F9C5" bold="true" box="[607,747,1631,1655]" pageId="5" pageNumber="32">
|
||
<taxonomicNameLabel id="48C69E4BFF84FF9CFC2EF9B2CBBAF9C5" box="[607,693,1631,1655]" pageId="5" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
2–3.
|
||
</emphasis>
|
||
Male holotype, dorsal and ventral views.
|
||
<emphasis id="53F52330FF84FF9CFADFF9B2CDD2F9C5" bold="true" box="[1198,1245,1631,1655]" pageId="5" pageNumber="32">4–5.</emphasis>
|
||
Female paratype, dorsal and ventral views. Scale bars = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="613EFF22FF84FF9EFEE6F925C8F0FE9E" blockId="5.[151,1437,1736,2023]" lastBlockId="7.[151,1436,151,300]" lastPageId="7" lastPageNumber="34" pageId="5" pageNumber="32">
|
||
tergite VII pentacarinate, with the median carina represented by an anteromedian elevation (
|
||
<figureCitation id="F9BAE3A7FF84FF9CFB4DF925CC82F950" box="[1340,1421,1736,1762]" captionStart="FIGURES 6 – 7" captionStartId="6.[151,269,1163,1187]" captionTargetBox="[157,1431,201,1136]" captionTargetId="figure@6.[151,1436,194,1139]" captionTargetPageId="6" captionText="FIGURES 6 – 7. Microtityus bivicentorum sp. nov., male holotype under UV light. 6. Dorsal view. 7. Ventral view. MC = Median carina; MLC = Median lateral carina; LC = Lateral carina; SMC = Submedian carina. Scale bars = 5 mm." httpUri="https://zenodo.org/record/188072/files/figure.png" pageId="5" pageNumber="32">Fig. 6</figureCitation>
|
||
).
|
||
<emphasis id="53F52330FF84FF9CFEE6F900C883F8B4" box="[151,396,1773,1798]" italics="true" pageId="5" pageNumber="32">Coxosternal region:</emphasis>
|
||
Sternum pentagonal (
|
||
<figureCitation id="F9BAE3A7FF84FF9CFCEAF900CBE9F8B5" box="[667,742,1773,1799]" captionStart="FIGURES 8 – 11" captionStartId="7.[151,269,1299,1323]" captionTargetBox="[159,1425,370,1244]" captionTargetId="figure@7.[151,1436,355,1276]" captionTargetPageId="7" captionText="FIGURES 8 – 11. Microtityus bivicentorum sp. nov. 8. Sternum, genital operculum, pectinal basal piece, and pectines of male holotype. 9. Sternum, genital operculum, pectinal basal piece, and pectines of female paratype. 10. Sternite V of male holotype. 11. Telson of male holotype, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188073/files/figure.png" pageId="5" pageNumber="32">Fig. 8</figureCitation>
|
||
), depressed anteriorly; all the components of this region granulose, except for coxapophyses I–II which are smooth.
|
||
<emphasis id="53F52330FF84FF9CFDE5F8FFCC39F899" box="[916,1334,1810,1835]" italics="true" pageId="5" pageNumber="32">Genital operculum and pectines:</emphasis>
|
||
Genital operculum divided longitudinally, formed by subtriangular plates (
|
||
<figureCitation id="F9BAE3A7FF84FF9CFDC9F8D5CD0EF8E0" box="[952,1025,1848,1874]" captionStart="FIGURES 8 – 11" captionStartId="7.[151,269,1299,1323]" captionTargetBox="[159,1425,370,1244]" captionTargetId="figure@7.[151,1436,355,1276]" captionTargetPageId="7" captionText="FIGURES 8 – 11. Microtityus bivicentorum sp. nov. 8. Sternum, genital operculum, pectinal basal piece, and pectines of male holotype. 9. Sternum, genital operculum, pectinal basal piece, and pectines of female paratype. 10. Sternite V of male holotype. 11. Telson of male holotype, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188073/files/figure.png" pageId="5" pageNumber="32">Fig. 8</figureCitation>
|
||
); pectines less than two thirds the length of coxae IV; pectinal marginal lamellae with 3/3 pieces, middle lamellae with 5/5, fulcra with 8/8; teeth count 9/9; pectinal basal piece almost quadrangular in shape, with anteromedian notch (
|
||
<figureCitation id="F9BAE3A7FF84FF9CFAB7F86FCC1EF82E" box="[1222,1297,1922,1948]" captionStart="FIGURES 8 – 11" captionStartId="7.[151,269,1299,1323]" captionTargetBox="[159,1425,370,1244]" captionTargetId="figure@7.[151,1436,355,1276]" captionTargetPageId="7" captionText="FIGURES 8 – 11. Microtityus bivicentorum sp. nov. 8. Sternum, genital operculum, pectinal basal piece, and pectines of male holotype. 9. Sternum, genital operculum, pectinal basal piece, and pectines of female paratype. 10. Sternite V of male holotype. 11. Telson of male holotype, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188073/files/figure.png" pageId="5" pageNumber="32">Fig. 8</figureCitation>
|
||
).
|
||
<emphasis id="53F52330FF84FF9CFB58F86FCC93F829" box="[1321,1436,1922,1947]" italics="true" pageId="5" pageNumber="32">Sternites:</emphasis>
|
||
Densely granulose throughout; sternites VI–VII with paired submedian and lateral carinae, incomplete and granulose (
|
||
<figureCitation id="F9BAE3A7FF84FF9CFF68F820C86FF855" box="[281,352,1997,2023]" captionStart="FIGURES 6 – 7" captionStartId="6.[151,269,1163,1187]" captionTargetBox="[157,1431,201,1136]" captionTargetId="figure@6.[151,1436,194,1139]" captionTargetPageId="6" captionText="FIGURES 6 – 7. Microtityus bivicentorum sp. nov., male holotype under UV light. 6. Dorsal view. 7. Ventral view. MC = Median carina; MLC = Median lateral carina; LC = Lateral carina; SMC = Submedian carina. Scale bars = 5 mm." httpUri="https://zenodo.org/record/188072/files/figure.png" pageId="5" pageNumber="32">Fig. 7</figureCitation>
|
||
); sternite V with three posterior smooth, hyaline areas, of which the median one is shaped like an equilateral triangle, and the lateral ones are large and flattened (
|
||
<figureCitation id="F9BAE3A7FF87FF9FFDA6FB1ECD21FABF" box="[983,1070,1267,1293]" captionStart="FIGURES 8 – 11" captionStartId="7.[151,269,1299,1323]" captionTargetBox="[159,1425,370,1244]" captionTargetId="figure@7.[151,1436,355,1276]" captionTargetPageId="7" captionText="FIGURES 8 – 11. Microtityus bivicentorum sp. nov. 8. Sternum, genital operculum, pectinal basal piece, and pectines of male holotype. 9. Sternum, genital operculum, pectinal basal piece, and pectines of female paratype. 10. Sternite V of male holotype. 11. Telson of male holotype, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188073/files/figure.png" pageId="6" pageNumber="33">Fig. 10</figureCitation>
|
||
); book lung spiracles short and oval.
|
||
<emphasis id="53F52330FF87FF9FFEADFAF7C869FA81" box="[220,358,1306,1331]" italics="true" pageId="6" pageNumber="33">Metasoma:</emphasis>
|
||
Segments I–II with ten carinae (paired ventrosubmedian, ventrolateral, median lateral, dorsolateral, and dorsosubmedian carinae), the median lateral less developed in segment II; segments III–IV with eight carinae (the median lateral absent); segment V with five carinae (ventromedian, paired ventrolateral and dorsolateral carinae); all carinae granulose; intercarinal spaces with abundant granulation. Telson completely smooth; subaculear tubercle strong, with two small dorsal granules; aculeus strongly curved (
|
||
<figureCitation id="F9BAE3A7FF87FF9FFE8CFA36C856FA47" box="[253,345,1499,1525]" captionStart="FIGURES 8 – 11" captionStartId="7.[151,269,1299,1323]" captionTargetBox="[159,1425,370,1244]" captionTargetId="figure@7.[151,1436,355,1276]" captionTargetPageId="7" captionText="FIGURES 8 – 11. Microtityus bivicentorum sp. nov. 8. Sternum, genital operculum, pectinal basal piece, and pectines of male holotype. 9. Sternum, genital operculum, pectinal basal piece, and pectines of female paratype. 10. Sternite V of male holotype. 11. Telson of male holotype, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188073/files/figure.png" pageId="6" pageNumber="33">Fig. 11</figureCitation>
|
||
).
|
||
<emphasis id="53F52330FF87FF9FFF00FA36CB0CFA46" box="[369,515,1499,1524]" italics="true" pageId="6" pageNumber="33">Chelicerae:</emphasis>
|
||
Cheliceral dentition characteristic of the family
|
||
<taxonomicName id="A68184A1FF87FF9FFA14FA36CC97FA47" authority="Vachon 1963" authorityName="Vachon" authorityYear="1963" box="[1125,1432,1499,1525]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="33" phylum="Arthropoda" rank="family">
|
||
Buthidae (
|
||
<bibRefCitation id="051082D3FF87FF9FFA94FA36CC81FA47" author="Vachon" box="[1253,1422,1499,1525]" pageId="6" pageNumber="38" refString="Vachon, M. (1963) De l'utilite´, en systematique, d'une nomenclature des dents des cheliceres chez les scorpions. Bulletin du Museum national de Histoire naturelle Paris (2 e ser.), 35 (2), 161 - 166." type="journal article" year="1963">Vachon 1963</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Movable finger externally with two small basal teeth, one median pronounced tooth, one subdistal tooth slightly shorter than the median, and one distal tooth. Fixed finger externally with one basal and one median tooth mounted onto a bicuspid, one subdistal, and one distal tooth. Internal surfaces of both movable and fixed fingers are hidden by the coxapophyses and pedipalp trochanters, since the chelicerae are retracted deep under the carapace and could not be pulled out successfully; therefore, dentition on the internal surfaces is not observable.
|
||
<emphasis id="53F52330FF87FF9FFF55F92EC8ABF96E" box="[292,420,1731,1756]" italics="true" pageId="6" pageNumber="33">Pedipalps:</emphasis>
|
||
Densely granulose throughout; femur with proximo-marginal carina, five longitudinal carinae (dorso-interior, dorso-exterior, ventro-exterior, ventro-interior carinae, plus an additional one on internal median position), and pronounced non-spinoid tubercle located basally on the internal surface; patella with eight longitudinal carinae [all of those identified by
|
||
<bibRefCitation id="051082D3FF87FF9FFD26F8DACD18F8E3" author="Stahnke" box="[855,1047,1847,1873]" pageId="6" pageNumber="38" refString="Stahnke, H. L. (1970) Scorpion nomenclature and mensuration. Entomological News, 81 (12), 297 - 316." type="journal article" year="1970">Stahnke (1970)</bibRefCitation>
|
||
], of which the ventro-median is vestigial) and two enlarged and spine-like granules on the dorso-interior carina and one on the ventro-interior; chela with nine longitudinal carinae; all carinae of all segments granulose. Movable and fixed fingers each with eleven slightly imbricate rows of granules in both pedipalps (including the short apical row) (
|
||
<figureCitation id="F9BAE3A7FF87FF9FFB2EF846C9E8F85E" captionStart="FIGURES 18 – 23" captionStartId="9.[151,273,1902,1926]" captionTargetBox="[159,1430,456,1842]" captionTargetId="figure@9.[151,1436,432,1894]" captionTargetPageId="9" captionText="FIGURES 18 – 23. Right pedipalp chela. 18 – 21. Microtityus bivicentorum sp. nov., male holotype. 18. Detail of the movable finger dentition. 19. Detail of the fixed finger dentition. 20. Chela, external view. 21. Chela, ventral view. 22 – 23. Microtityus franckei, male paratype (MPUJ-SCO- 367). 22. Chela, external view. 23. Chela, ventral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188075/files/figure.png" pageId="6" pageNumber="33">Figs. 18–19</figureCitation>
|
||
).
|
||
<emphasis id="53F52330FF87FF9FFE8FF83FC8BCF859" box="[254,435,2002,2027]" italics="true" pageId="6" pageNumber="33">Trichobothria:</emphasis>
|
||
Trichobothriotaxy
|
||
<typeStatus id="BE3A4180FF87FF9FFCD3F83FCBD1F85E" box="[674,734,2002,2028]" pageId="6" pageNumber="33">Type</typeStatus>
|
||
A (
|
||
<bibRefCitation id="051082D3FF87FF9FFD7FF83FCABBF85E" author="Vachon" box="[782,948,2002,2028]" pageId="6" pageNumber="38" refString="Vachon, M. (1974) Etude des caracteres utilises pour classer les familles et les genres de scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les scorpions. Bulletin du Museum national de Histoire naturelle, Paris (3 e ser.), 104, 857 - 958." type="journal article" year="1974">Vachon 1974</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="051082D3FF87FF9FFDB3F83FCD0EF85E" author="Vachon" box="[962,1025,2002,2028]" pageId="6" pageNumber="38" refString="Vachon, M. (1975) Sur l'utilisation de la trichobothriotaxie du bras des pedipalpes des scorpions (Arachnides) dans le classement des genres de la famille des Buthidae Simon. Comptes Rendus des seances de l'Academie des Sciences, Paris (ser. D) 281, 1597 - 1599." type="journal article" year="1975">1975</bibRefCitation>
|
||
) with 34 trichobothria (reductive neobothriotaxy) (
|
||
<figureCitation id="F9BAE3A7FF86FF9EFF1AFF75CB0CFF00" box="[363,515,152,178]" captionStart="FIGURES 12 – 17" captionStartId="8.[151,269,1574,1598]" captionTargetBox="[157,1429,367,1512]" captionTargetId="figure@8.[151,1436,349,1550]" captionTargetPageId="8" captionText="FIGURES 12 – 17. Microtityus bivicentorum sp. nov., distribution of the trichobothria in male holotype (illustrated from the right pedipalp). 12. Femur, dorsal view. 13. Femur, internal view. 14. Patella, dorsal view. 15. Patella, external view. 16. Chela, external view. 17. Chela, ventral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188074/files/figure.png" pageId="7" pageNumber="34">Figs. 12–17</figureCitation>
|
||
): femur with ten trichobothria in α configuration (
|
||
<emphasis id="53F52330FF86FF9EFA1FFF75CD8AFF04" box="[1134,1157,152,182]" italics="true" pageId="7" pageNumber="34">d2</emphasis>
|
||
absent) (
|
||
<figureCitation id="F9BAE3A7FF86FF9EFA89FF75CC81FF00" box="[1272,1422,152,178]" captionStart="FIGURES 12 – 17" captionStartId="8.[151,269,1574,1598]" captionTargetBox="[157,1429,367,1512]" captionTargetId="figure@8.[151,1436,349,1550]" captionTargetPageId="8" captionText="FIGURES 12 – 17. Microtityus bivicentorum sp. nov., distribution of the trichobothria in male holotype (illustrated from the right pedipalp). 12. Femur, dorsal view. 13. Femur, internal view. 14. Patella, dorsal view. 15. Patella, external view. 16. Chela, external view. 17. Chela, ventral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188074/files/figure.png" pageId="7" pageNumber="34">Figs. 12–13</figureCitation>
|
||
), patella with twelve (
|
||
<emphasis id="53F52330FF86FF9EFFFBFF2CC8AEFF6D" box="[394,417,193,223]" italics="true" pageId="7" pageNumber="34">d2</emphasis>
|
||
absent) (
|
||
<figureCitation id="F9BAE3A7FF86FF9EFC7FFF2CCB91FF69" box="[526,670,193,219]" captionStart="FIGURES 12 – 17" captionStartId="8.[151,269,1574,1598]" captionTargetBox="[157,1429,367,1512]" captionTargetId="figure@8.[151,1436,349,1550]" captionTargetPageId="8" captionText="FIGURES 12 – 17. Microtityus bivicentorum sp. nov., distribution of the trichobothria in male holotype (illustrated from the right pedipalp). 12. Femur, dorsal view. 13. Femur, internal view. 14. Patella, dorsal view. 15. Patella, external view. 16. Chela, external view. 17. Chela, ventral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188074/files/figure.png" pageId="7" pageNumber="34">Figs. 14–15</figureCitation>
|
||
), and chela with twelve (
|
||
<emphasis id="53F52330FF86FF9EFDBBFF2CCAFCFF6D" box="[970,1011,193,223]" italics="true" pageId="7" pageNumber="34">Eb3</emphasis>
|
||
,
|
||
<emphasis id="53F52330FF86FF9EFA73FF2CCD21FF68" box="[1026,1070,193,218]" italics="true" pageId="7" pageNumber="34">Esb</emphasis>
|
||
and
|
||
<emphasis id="53F52330FF86FF9EFA18FF2CCD9FFF68" box="[1129,1168,193,218]" italics="true" pageId="7" pageNumber="34">esb</emphasis>
|
||
absent) (
|
||
<figureCitation id="F9BAE3A7FF86FF9EFA8FFF2CCC80FF69" box="[1278,1423,193,219]" captionStart="FIGURES 12 – 17" captionStartId="8.[151,269,1574,1598]" captionTargetBox="[157,1429,367,1512]" captionTargetId="figure@8.[151,1436,349,1550]" captionTargetPageId="8" captionText="FIGURES 12 – 17. Microtityus bivicentorum sp. nov., distribution of the trichobothria in male holotype (illustrated from the right pedipalp). 12. Femur, dorsal view. 13. Femur, internal view. 14. Patella, dorsal view. 15. Patella, external view. 16. Chela, external view. 17. Chela, ventral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188074/files/figure.png" pageId="7" pageNumber="34">Figs. 16–17</figureCitation>
|
||
).
|
||
<emphasis id="53F52330FF86FF9EFEE6FF06C9D5FEB6" box="[151,218,235,260]" italics="true" pageId="7" pageNumber="34">Legs:</emphasis>
|
||
Tibia, basitarsus and telotarsus with abundant setation; tibial spur absent; prolateral and retrolateral pedal spurs present on all legs.
|
||
</paragraph>
|
||
<caption id="35FEAFAAFF87FF9FFEE6FB66CC5BFB76" httpUri="https://zenodo.org/record/188072/files/figure.png" pageId="6" pageNumber="33" targetBox="[157,1431,201,1136]" targetPageId="6">
|
||
<paragraph id="613EFF22FF87FF9FFEE6FB66CC5BFB76" blockId="6.[151,1436,1163,1220]" pageId="6" pageNumber="33">
|
||
<emphasis id="53F52330FF87FF9FFEE6FB66C84DFB11" bold="true" box="[151,322,1163,1187]" pageId="6" pageNumber="33">FIGURES 6–7.</emphasis>
|
||
<taxonomicName id="A68184A1FF87FF9FFF38FB66CB5AFB10" box="[329,597,1163,1186]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="bivicentorum" status="sp. nov.">
|
||
<emphasis id="53F52330FF87FF9FFF38FB66CB5AFB10" box="[329,597,1163,1186]" italics="true" pageId="6" pageNumber="33">Microtityus bivicentorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="48C69E4BFF87FF9FFC2DFB66CBBEFB11" box="[604,689,1163,1187]" pageId="6" pageNumber="33" rank="species">
|
||
<emphasis id="53F52330FF87FF9FFC2DFB66CBBEFB11" bold="true" box="[604,689,1163,1187]" pageId="6" pageNumber="33">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male holotype under UV light.
|
||
<emphasis id="53F52330FF87FF9FFA78FB66CD11FB11" bold="true" box="[1033,1054,1163,1187]" pageId="6" pageNumber="33">6.</emphasis>
|
||
Dorsal view.
|
||
<emphasis id="53F52330FF87FF9FFAC1FB66CDCBFB11" bold="true" box="[1200,1220,1163,1187]" pageId="6" pageNumber="33">7.</emphasis>
|
||
Ventral view. MC = Median carina; MLC = Median lateral carina; LC = Lateral carina; SMC = Submedian carina. Scale bars = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="35FEAFAAFF86FF9EFEE6FAFECAD3FAC2" httpUri="https://zenodo.org/record/188073/files/figure.png" pageId="7" pageNumber="34" targetBox="[159,1425,370,1244]" targetPageId="7">
|
||
<paragraph id="613EFF22FF86FF9EFEE6FAFECAD3FAC2" blockId="7.[151,1436,1299,1393]" pageId="7" pageNumber="34">
|
||
<emphasis id="53F52330FF86FF9EFEE6FAFEC840FA99" bold="true" box="[151,335,1299,1323]" pageId="7" pageNumber="34">FIGURES 8–11.</emphasis>
|
||
<taxonomicName id="A68184A1FF86FF9EFF27FAF9CB6BFA99" box="[342,612,1300,1323]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="bivicentorum" status="sp. nov.">
|
||
<emphasis id="53F52330FF86FF9EFF27FAF9CB6BFA99" box="[342,612,1300,1323]" italics="true" pageId="7" pageNumber="34">Microtityus bivicentorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="53F52330FF86FF9EFC1AFAFECBD3FA99" bold="true" box="[619,732,1299,1323]" pageId="7" pageNumber="34">
|
||
<taxonomicNameLabel id="48C69E4BFF86FF9EFC1AFAFECBCEFA99" box="[619,705,1299,1323]" pageId="7" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
8.
|
||
</emphasis>
|
||
Sternum, genital operculum, pectinal basal piece, and pectines of male holotype.
|
||
<emphasis id="53F52330FF86FF9EFF31FADBC85BFAFC" bold="true" box="[320,340,1334,1358]" pageId="7" pageNumber="34">9.</emphasis>
|
||
Sternum, genital operculum, pectinal basal piece, and pectines of female paratype.
|
||
<emphasis id="53F52330FF86FF9EFA97FADBCC08FAFC" bold="true" box="[1254,1287,1334,1358]" pageId="7" pageNumber="34">10.</emphasis>
|
||
Sternite V of male holotype.
|
||
<emphasis id="53F52330FF86FF9EFF4DFAB4C855FAC3" bold="true" box="[316,346,1369,1393]" pageId="7" pageNumber="34">11.</emphasis>
|
||
Telson of male holotype, lateral view. Scale bars = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="613EFF22FF86FF9EFEB4FA73CBC4FA0A" blockId="7.[151,1437,1438,2009]" box="[197,715,1438,1464]" pageId="7" pageNumber="34">
|
||
<emphasis id="53F52330FF86FF9EFEB4FA73CBC4FA0A" bold="true" box="[197,715,1438,1464]" pageId="7" pageNumber="34">
|
||
Female
|
||
<typeStatus id="BE3A4180FF86FF9EFF56FA73C898FA0A" box="[295,407,1438,1464]" pageId="7" pageNumber="34" type="paratype">paratype</typeStatus>
|
||
and sexual dimorphism.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="613EFF22FF86FF9EFEB4FA28CB4DF95C" blockId="7.[151,1437,1438,2009]" pageId="7" pageNumber="34">
|
||
Similar to the male
|
||
<typeStatus id="BE3A4180FF86FF9EFFC4FA28CB2EFA6D" box="[437,545,1477,1503]" pageId="7" pageNumber="34" type="holotype">holotype</typeStatus>
|
||
, but the following differences were identified and are attributed to sexual dimorphism: lateral carinae on tergites I–VI markedly vestigial; 8/8 pectinal teeth; pectinal basal piece without anteromedian notch and longer than in the male (
|
||
<figureCitation id="F9BAE3A7FF86FF9EFD46F9FFCA70F99E" box="[823,895,1554,1580]" captionStart="FIGURES 8 – 11" captionStartId="7.[151,269,1299,1323]" captionTargetBox="[159,1425,370,1244]" captionTargetId="figure@7.[151,1436,355,1276]" captionTargetPageId="7" captionText="FIGURES 8 – 11. Microtityus bivicentorum sp. nov. 8. Sternum, genital operculum, pectinal basal piece, and pectines of male holotype. 9. Sternum, genital operculum, pectinal basal piece, and pectines of female paratype. 10. Sternite V of male holotype. 11. Telson of male holotype, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188073/files/figure.png" pageId="7" pageNumber="34">Fig. 9</figureCitation>
|
||
); sternite V without posterior smooth, hyaline areas. Other differences were also detected, but cannot be reliably attributed to dimorphism: 7/7 pectinal fulcra; telson with few granules located dorsally on the posterolateral surfaces; movable finger of both pedipalps each with twelve rows of granules and fixed fingers with eleven rows. Cheliceral movable finger internally with two small teeth (one basal and one median) and one distal tooth; cheliceral fixed finger internally with only one small tooth.
|
||
</paragraph>
|
||
<paragraph id="613EFF22FF86FF9EFEB4F917C87DF8A6" blockId="7.[151,1437,1438,2009]" box="[197,370,1786,1812]" pageId="7" pageNumber="34">
|
||
<emphasis id="53F52330FF86FF9EFEB4F917C87DF8A6" bold="true" box="[197,370,1786,1812]" pageId="7" pageNumber="34">Comparisons.</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="299BACA9FF86FF90FEB4F8CCC8CBFEC4" lastPageId="9" lastPageNumber="36" pageId="7" pageNumber="34" type="reference_group">
|
||
<paragraph id="613EFF22FF86FF91FEB4F8CCCC18FE94" blockId="7.[151,1437,1438,2009]" lastBlockId="8.[151,1436,152,294]" lastPageId="8" lastPageNumber="35" pageId="7" pageNumber="34">
|
||
<taxonomicName id="A68184A1FF86FF9EFEB4F8CCC8E2F888" ID-CoL="438SQ" box="[197,493,1825,1850]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="bivicentorum" status="sp. nov.">
|
||
<emphasis id="53F52330FF86FF9EFEB4F8CCC8E2F888" box="[197,493,1825,1850]" italics="true" pageId="7" pageNumber="34">Microtityus bivicentorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="48C69E4BFF86FF9EFF84F8CCCB5EF889" box="[501,593,1825,1851]" pageId="7" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is most similar to
|
||
<taxonomicName id="A68184A1FF86FF9EFD5CF8CCCABAF888" ID-CoL="438T2" box="[813,949,1825,1850]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="franckei">
|
||
<emphasis id="53F52330FF86FF9EFD5CF8CCCABAF888" box="[813,949,1825,1850]" italics="true" pageId="7" pageNumber="34">M. franckei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A68184A1FF86FF9EFDB5F8CCC9DCF8D0" ID-CoL="73JT9" authority="Gonzalez-Sponga, 2001" authorityName="Gonzalez-Sponga" authorityYear="2001" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="desuzeae">
|
||
<emphasis id="53F52330FF86FF9EFDB5F8CCCDB3F888" box="[964,1212,1825,1850]" italics="true" pageId="7" pageNumber="34">Microtityus desuzeae</emphasis>
|
||
<bibRefCitation id="051082D3FF86FF9EFAB5F8CCC9DCF8D0" author="Gonzalez-Sponga" pageId="7" pageNumber="38" refString="Gonzalez-Sponga, M. A. (2001) Aracnidos de Venezuela. Seis nuevas especies del genero Microtityus (Scorpionida: Buthidae) del sistema montanoso de la costa. Boletin de la Academia de Ciencias Fisicas, Matematicas y Naturales, 61 (1 - 2), 45 - 66." type="journal article" year="2001">González-Sponga, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
[originally
|
||
<taxonomicName id="A68184A1FF86FF9EFF12F8A5C8EAF8D3" box="[355,485,1864,1889]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="desuzei">
|
||
<emphasis id="53F52330FF86FF9EFF12F8A5C8EAF8D3" box="[355,485,1864,1889]" italics="true" pageId="7" pageNumber="34">M. desuzei</emphasis>
|
||
</taxonomicName>
|
||
, corrected by Rojas-Runjaic & de
|
||
<bibRefCitation id="051082D3FF86FF9EFDFFF8A5CD3EF8D0" author="Sousa" box="[910,1073,1864,1890]" pageId="7" pageNumber="38" refString="Rojas-Runjaic, F. J. M. & de Sousa, L. (2007) Catalogo de los escorpiones de Venezuela (Arachnida: Scorpiones). Boletin de la Sociedad Entomologica Aragonesa, 40, 281 - 307." type="journal article" year="2007">Sousa (2007)</bibRefCitation>
|
||
] and
|
||
<taxonomicName id="A68184A1FF86FF9EFA05F8A5C8B1F83B" ID-CoL="438T7" authority="Gonzalez-Sponga, 1981" authorityName="Gonzalez-Sponga" authorityYear="1981" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="joseantonioi">
|
||
<emphasis id="53F52330FF86FF9EFA05F8A5CC93F8D3" box="[1140,1436,1864,1889]" italics="true" pageId="7" pageNumber="34">Microtityus joseantonioi</emphasis>
|
||
<bibRefCitation id="051082D3FF86FF9EFEE6F882C8B1F83B" author="Gonzalez-Sponga" box="[151,446,1903,1929]" pageId="7" pageNumber="38" refString="Gonzalez-Sponga, M. A. (1981) Un nuevo genero y dos nuevas especies de la familia Buthidae de Venezuela (Arachnida: Scorpiones). Monografias Cientificas '' Augusto Pi Suner' ' (Instituto Universitario Pedagogico de Caracas), 13, 1 - 26." type="journal article" year="1981">González-Sponga, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
, with which it shares the absence of trichobothrium
|
||
<emphasis id="53F52330FF86FF9EFA32F882CD55F83F" box="[1091,1114,1903,1933]" italics="true" pageId="7" pageNumber="34">d2</emphasis>
|
||
on femur,
|
||
<emphasis id="53F52330FF86FF9EFA90F882CC04F83F" box="[1249,1291,1903,1933]" italics="true" pageId="7" pageNumber="34">Eb3</emphasis>
|
||
and
|
||
<emphasis id="53F52330FF86FF9EFB39F882CC79F83A" box="[1352,1398,1903,1928]" italics="true" pageId="7" pageNumber="34">Esb</emphasis>
|
||
on pedipalp chela, and
|
||
<emphasis id="53F52330FF86FF9EFFF5F874C8A4F800" box="[388,427,1945,1970]" italics="true" pageId="7" pageNumber="34">esb</emphasis>
|
||
on fixed finger, a combination unique to these species. The new species differs form
|
||
<taxonomicName id="A68184A1FF86FF9EFEE6F852C82CF86A" ID-CoL="438T2" box="[151,291,1983,2008]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="franckei">
|
||
<emphasis id="53F52330FF86FF9EFEE6F852C82CF86A" box="[151,291,1983,2008]" italics="true" pageId="7" pageNumber="34">M. franckei</emphasis>
|
||
</taxonomicName>
|
||
because in the former trichobothrium
|
||
<emphasis id="53F52330FF86FF9EFC86F852CA02F86A" box="[759,781,1983,2008]" italics="true" pageId="7" pageNumber="34">et</emphasis>
|
||
is located between
|
||
<emphasis id="53F52330FF86FF9EFD8CF852CD14F86A" box="[1021,1051,1983,2008]" italics="true" pageId="7" pageNumber="34">db</emphasis>
|
||
and
|
||
<emphasis id="53F52330FF86FF9EFA26F852CD60F86A" box="[1111,1135,1983,2008]" italics="true" pageId="7" pageNumber="34">dt</emphasis>
|
||
on pedipalp fixed finger (
|
||
<figureCitation id="F9BAE3A7FF89FF91FEEEFF75C9F9FF00" box="[159,246,152,178]" captionStart="FIGURES 12 – 17" captionStartId="8.[151,269,1574,1598]" captionTargetBox="[157,1429,367,1512]" captionTargetId="figure@8.[151,1436,349,1550]" captionTargetPageId="8" captionText="FIGURES 12 – 17. Microtityus bivicentorum sp. nov., distribution of the trichobothria in male holotype (illustrated from the right pedipalp). 12. Femur, dorsal view. 13. Femur, internal view. 14. Patella, dorsal view. 15. Patella, external view. 16. Chela, external view. 17. Chela, ventral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188074/files/figure.png" pageId="8" pageNumber="35">Fig. 16</figureCitation>
|
||
), and sternite V of males exhibits three smooth, hyaline areas (
|
||
<figureCitation id="F9BAE3A7FF89FF91FDAEFF75CD39FF00" box="[991,1078,152,178]" captionStart="FIGURES 8 – 11" captionStartId="7.[151,269,1299,1323]" captionTargetBox="[159,1425,370,1244]" captionTargetId="figure@7.[151,1436,355,1276]" captionTargetPageId="7" captionText="FIGURES 8 – 11. Microtityus bivicentorum sp. nov. 8. Sternum, genital operculum, pectinal basal piece, and pectines of male holotype. 9. Sternum, genital operculum, pectinal basal piece, and pectines of female paratype. 10. Sternite V of male holotype. 11. Telson of male holotype, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188073/files/figure.png" pageId="8" pageNumber="35">Fig. 10</figureCitation>
|
||
); in contrast, in
|
||
<emphasis id="53F52330FF89FF91FA86FF75CC93FF03" box="[1271,1436,152,177]" italics="true" pageId="8" pageNumber="35">
|
||
<taxonomicName id="A68184A1FF89FF91FA86FF75CC70FF03" ID-CoL="438T2" box="[1271,1407,152,177]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="franckei">M. franckei</taxonomicName>
|
||
et
|
||
</emphasis>
|
||
is basal to
|
||
<emphasis id="53F52330FF89FF91FF62FF52C83FFF6A" box="[275,304,191,216]" italics="true" pageId="8" pageNumber="35">db</emphasis>
|
||
(
|
||
<figureCitation id="F9BAE3A7FF89FF91FF31FF53C898FF6A" box="[320,407,190,216]" captionStart="FIGURES 18 – 23" captionStartId="9.[151,273,1902,1926]" captionTargetBox="[159,1430,456,1842]" captionTargetId="figure@9.[151,1436,432,1894]" captionTargetPageId="9" captionText="FIGURES 18 – 23. Right pedipalp chela. 18 – 21. Microtityus bivicentorum sp. nov., male holotype. 18. Detail of the movable finger dentition. 19. Detail of the fixed finger dentition. 20. Chela, external view. 21. Chela, ventral view. 22 – 23. Microtityus franckei, male paratype (MPUJ-SCO- 367). 22. Chela, external view. 23. Chela, ventral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188075/files/figure.png" pageId="8" pageNumber="35">Fig. 22</figureCitation>
|
||
;
|
||
<bibRefCitation id="051082D3FF89FF91FFD2FF53CA11FF6A" author="Botero-Trujillo" box="[419,798,190,216]" pageId="8" pageNumber="37" refString="Botero-Trujillo, R. & Noriega, J. A. (2008) First record of the scorpion genus Microtityus from Colombia, with the description of a new species (Scorpiones, Buthidae). Journal of Arachnology, 36, 259 - 266." type="journal article" year="2008">Botero-Trujillo & Noriega 2008</bibRefCitation>
|
||
: fig. 13), and sternite V of males has only one smooth, hyaline area (
|
||
<bibRefCitation id="051082D3FF89FF91FF49FF08CA04FF4D" author="Botero-Trujillo" box="[312,779,229,255]" pageId="8" pageNumber="37" refString="Botero-Trujillo, R. & Noriega, J. A. (2008) First record of the scorpion genus Microtityus from Colombia, with the description of a new species (Scorpiones, Buthidae). Journal of Arachnology, 36, 259 - 266." type="journal article" year="2008">Botero-Trujillo & Noriega 2008: fig. 8</bibRefCitation>
|
||
). Also, the new species differs from
|
||
<taxonomicName id="A68184A1FF89FF91FAB7FF08CC5FFF4C" ID-CoL="438T2" box="[1222,1360,229,254]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="franckei">
|
||
<emphasis id="53F52330FF89FF91FAB7FF08CC5FFF4C" box="[1222,1360,229,254]" italics="true" pageId="8" pageNumber="35">M. franckei</emphasis>
|
||
</taxonomicName>
|
||
in the "palm to movable finger” ratio, approximately 4:
|
||
<quantity id="A67952C7FF89FF91FCA5FEE1CA0DFE94" box="[724,770,268,294]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="8" pageNumber="35" unit="in" value="5.0">5 in</quantity>
|
||
the former but 3:
|
||
<quantity id="A67952C7FF89FF91FDBCFEE1CAF3FE94" box="[973,1020,268,294]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="8" pageNumber="35" unit="in" value="5.0">5 in</quantity>
|
||
the latter (
|
||
<figureCitation id="F9BAE3A7FF89FF91FA0BFEE1CC05FE94" box="[1146,1290,268,294]" captionStart="FIGURES 18 – 23" captionStartId="9.[151,273,1902,1926]" captionTargetBox="[159,1430,456,1842]" captionTargetId="figure@9.[151,1436,432,1894]" captionTargetPageId="9" captionText="FIGURES 18 – 23. Right pedipalp chela. 18 – 21. Microtityus bivicentorum sp. nov., male holotype. 18. Detail of the movable finger dentition. 19. Detail of the fixed finger dentition. 20. Chela, external view. 21. Chela, ventral view. 22 – 23. Microtityus franckei, male paratype (MPUJ-SCO- 367). 22. Chela, external view. 23. Chela, ventral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188075/files/figure.png" pageId="8" pageNumber="35">Figs. 20–23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="35FEAFAAFF89FF91FEE6F9CBCA83F931" httpUri="https://zenodo.org/record/188074/files/figure.png" pageId="8" pageNumber="35" targetBox="[157,1429,367,1512]" targetPageId="8">
|
||
<paragraph id="613EFF22FF89FF91FEE6F9CBCA83F931" blockId="8.[151,1437,1574,1667]" pageId="8" pageNumber="35">
|
||
<emphasis id="53F52330FF89FF91FEE6F9CBC851F98C" bold="true" box="[151,350,1574,1598]" pageId="8" pageNumber="35">FIGURES 12–17.</emphasis>
|
||
<taxonomicName id="A68184A1FF89FF91FF15F9CBCB7DF98F" ID-CoL="438SQ" box="[356,626,1574,1597]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="bivicentorum" status="sp. nov.">
|
||
<emphasis id="53F52330FF89FF91FF15F9CBCB7DF98F" box="[356,626,1574,1597]" italics="true" pageId="8" pageNumber="35">Microtityus bivicentorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="48C69E4BFF89FF91FC08F9CBCBC1F98C" box="[633,718,1574,1598]" pageId="8" pageNumber="35" rank="species">
|
||
<emphasis id="53F52330FF89FF91FC08F9CBCBC1F98C" bold="true" box="[633,718,1574,1598]" pageId="8" pageNumber="35">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, distribution of the trichobothria in male holotype (illustrated from the right pedipalp).
|
||
<emphasis id="53F52330FF89FF91FF1BF9A4C884F9D3" bold="true" box="[362,395,1609,1633]" pageId="8" pageNumber="35">12.</emphasis>
|
||
Femur, dorsal view.
|
||
<emphasis id="53F52330FF89FF91FC1CF9A4CB81F9D3" bold="true" box="[621,654,1609,1633]" pageId="8" pageNumber="35">13.</emphasis>
|
||
Femur, internal view.
|
||
<emphasis id="53F52330FF89FF91FDF1F9A4CAADF9D3" bold="true" box="[896,930,1609,1633]" pageId="8" pageNumber="35">14.</emphasis>
|
||
Patella, dorsal view.
|
||
<emphasis id="53F52330FF89FF91FAF6F9A4CDA7F9D3" bold="true" box="[1159,1192,1609,1633]" pageId="8" pageNumber="35">15.</emphasis>
|
||
Patella, external view.
|
||
<emphasis id="53F52330FF89FF91FEE6F986C9B7F931" bold="true" box="[151,184,1643,1667]" pageId="8" pageNumber="35">16.</emphasis>
|
||
Chela, external view.
|
||
<emphasis id="53F52330FF89FF91FFD7F986C8C8F931" bold="true" box="[422,455,1643,1667]" pageId="8" pageNumber="35">17.</emphasis>
|
||
Chela, ventral view. Scale bars = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="613EFF22FF89FF90FEB4F95CCA47FF6A" blockId="8.[151,1437,1713,2015]" lastBlockId="9.[151,1437,152,374]" lastPageId="9" lastPageNumber="36" pageId="8" pageNumber="35">
|
||
<taxonomicName id="A68184A1FF89FF91FEB4F95CC8F6F978" ID-CoL="438SQ" box="[197,505,1713,1738]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="bivicentorum" status="sp. nov.">
|
||
<emphasis id="53F52330FF89FF91FEB4F95CC8F6F978" box="[197,505,1713,1738]" italics="true" pageId="8" pageNumber="35">Microtityus bivicentorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="48C69E4BFF89FF91FC73F95CCB6EF979" box="[514,609,1713,1739]" pageId="8" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="A68184A1FF89FF91FD74F95CCACAF978" ID-CoL="438T7" box="[773,965,1713,1738]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="joseantonioi">
|
||
<emphasis id="53F52330FF89FF91FD74F95CCACAF978" box="[773,965,1713,1738]" italics="true" pageId="8" pageNumber="35">M. joseantonioi</emphasis>
|
||
</taxonomicName>
|
||
because in the former species patellar trichobothrium
|
||
<emphasis id="53F52330FF89FF91FF23F935C866F944" box="[338,361,1752,1782]" italics="true" pageId="8" pageNumber="35">d2</emphasis>
|
||
is absent (
|
||
<figureCitation id="F9BAE3A7FF89FF91FF9AF935CB4BF940" box="[491,580,1752,1778]" captionStart="FIGURES 12 – 17" captionStartId="8.[151,269,1574,1598]" captionTargetBox="[157,1429,367,1512]" captionTargetId="figure@8.[151,1436,349,1550]" captionTargetPageId="8" captionText="FIGURES 12 – 17. Microtityus bivicentorum sp. nov., distribution of the trichobothria in male holotype (illustrated from the right pedipalp). 12. Femur, dorsal view. 13. Femur, internal view. 14. Patella, dorsal view. 15. Patella, external view. 16. Chela, external view. 17. Chela, ventral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188074/files/figure.png" pageId="8" pageNumber="35">Fig. 14</figureCitation>
|
||
), the movable finger of pedipalp chela has 11–12 rows of granules, the pectinal basal piece is quadrangular in shape in both sexes (despite sexual dimorphism) (
|
||
<figureCitation id="F9BAE3A7FF89FF91FA96F8ECCC50F8A9" box="[1255,1375,1793,1819]" captionStart="FIGURES 8 – 11" captionStartId="7.[151,269,1299,1323]" captionTargetBox="[159,1425,370,1244]" captionTargetId="figure@7.[151,1436,355,1276]" captionTargetPageId="7" captionText="FIGURES 8 – 11. Microtityus bivicentorum sp. nov. 8. Sternum, genital operculum, pectinal basal piece, and pectines of male holotype. 9. Sternum, genital operculum, pectinal basal piece, and pectines of female paratype. 10. Sternite V of male holotype. 11. Telson of male holotype, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188073/files/figure.png" pageId="8" pageNumber="35">Figs. 8–9</figureCitation>
|
||
), the median smooth, hyaline area on sternite V of males is shaped like an equilateral triangle, and the lateral areas are large and flattened (
|
||
<figureCitation id="F9BAE3A7FF89FF91FFB9F8A2CB26F8DB" box="[456,553,1871,1897]" captionStart="FIGURES 8 – 11" captionStartId="7.[151,269,1299,1323]" captionTargetBox="[159,1425,370,1244]" captionTargetId="figure@7.[151,1436,355,1276]" captionTargetPageId="7" captionText="FIGURES 8 – 11. Microtityus bivicentorum sp. nov. 8. Sternum, genital operculum, pectinal basal piece, and pectines of male holotype. 9. Sternum, genital operculum, pectinal basal piece, and pectines of female paratype. 10. Sternite V of male holotype. 11. Telson of male holotype, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188073/files/figure.png" pageId="8" pageNumber="35">Fig. 10</figureCitation>
|
||
); in contrast, in
|
||
<taxonomicName id="A68184A1FF89FF91FC8CF8A2CAC9F8DA" ID-CoL="438T7" box="[765,966,1871,1896]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="joseantonioi">
|
||
<emphasis id="53F52330FF89FF91FC8CF8A2CAC9F8DA" box="[765,966,1871,1896]" italics="true" pageId="8" pageNumber="35">M. joseantonioi</emphasis>
|
||
</taxonomicName>
|
||
patellar trichobothrium
|
||
<emphasis id="53F52330FF89FF91FB71F8A2CC17F8DF" box="[1280,1304,1871,1901]" italics="true" pageId="8" pageNumber="35">d2</emphasis>
|
||
is present (
|
||
<bibRefCitation id="051082D3FF89FF91FEEEF895CB2DF820" author="Gonzalez-Sponga" box="[159,546,1912,1938]" pageId="8" pageNumber="38" refString="Gonzalez-Sponga, M. A. (1981) Un nuevo genero y dos nuevas especies de la familia Buthidae de Venezuela (Arachnida: Scorpiones). Monografias Cientificas '' Augusto Pi Suner' ' (Instituto Universitario Pedagogico de Caracas), 13, 1 - 26." type="journal article" year="1981">González-Sponga 1981: fig. 11</bibRefCitation>
|
||
), the movable finger of pedipalp chela has nine rows (
|
||
<bibRefCitation id="051082D3FF89FF91FAB3F895C9DAF80B" author="Gonzalez-Sponga" pageId="8" pageNumber="38" refString="Gonzalez-Sponga, M. A. (1981) Un nuevo genero y dos nuevas especies de la familia Buthidae de Venezuela (Arachnida: Scorpiones). Monografias Cientificas '' Augusto Pi Suner' ' (Instituto Universitario Pedagogico de Caracas), 13, 1 - 26." type="journal article" year="1981">González-Sponga 1981</bibRefCitation>
|
||
), the pectinal basal piece is not quadrangular in either males or females [it is curved posteriorly in males and has a posterior projection in females (
|
||
<bibRefCitation id="051082D3FF89FF91FCF6F828CD38F86D" author="Gonzalez-Sponga" box="[647,1079,1989,2015]" pageId="8" pageNumber="38" refString="Gonzalez-Sponga, M. A. (1981) Un nuevo genero y dos nuevas especies de la familia Buthidae de Venezuela (Arachnida: Scorpiones). Monografias Cientificas '' Augusto Pi Suner' ' (Instituto Universitario Pedagogico de Caracas), 13, 1 - 26." type="journal article" year="1981">González-Sponga 1981: figs. 15–16</bibRefCitation>
|
||
)], the median smooth, hyaline area on sternite V of males is higher than wide (shaped like an isosceles triangle), and the lateral areas are much reduced and ovoid (
|
||
<bibRefCitation id="051082D3FF88FF90FFB6FF53CA34FF6A" author="Gonzalez-Sponga" box="[455,827,190,216]" pageId="9" pageNumber="38" refString="Gonzalez-Sponga, M. A. (1981) Un nuevo genero y dos nuevas especies de la familia Buthidae de Venezuela (Arachnida: Scorpiones). Monografias Cientificas '' Augusto Pi Suner' ' (Instituto Universitario Pedagogico de Caracas), 13, 1 - 26." type="journal article" year="1981">González-Sponga 1981: fig. 14</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="613EFF22FF88FF90FEB4FF08C8CBFEC4" blockId="9.[151,1437,152,374]" pageId="9" pageNumber="36">
|
||
Finally, the new species differs from
|
||
<taxonomicName id="A68184A1FF88FF90FC0AFF08CA02FF4C" ID-CoL="73JT9" box="[635,781,229,254]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="36" phylum="Arthropoda" rank="species" species="desuzeae">
|
||
<emphasis id="53F52330FF88FF90FC0AFF08CA02FF4C" box="[635,781,229,254]" italics="true" pageId="9" pageNumber="36">M. desuzeae</emphasis>
|
||
</taxonomicName>
|
||
on the basis of the patellar trichobothrium
|
||
<emphasis id="53F52330FF88FF90FB7DFF08CC2CFEB1" box="[1292,1315,229,259]" italics="true" pageId="9" pageNumber="36">d2</emphasis>
|
||
, absent in
|
||
<taxonomicName id="A68184A1FF88FF90FEE6FEE2C851FE9A" ID-CoL="438SQ" box="[151,350,271,296]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="36" phylum="Arthropoda" rank="species" species="bivicentorum" status="sp. nov.">
|
||
<emphasis id="53F52330FF88FF90FEE6FEE2C851FE9A" box="[151,350,271,296]" italics="true" pageId="9" pageNumber="36">M. bivicentorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="48C69E4BFF88FF90FF16FEE2C8CAFE9B" box="[359,453,271,297]" pageId="9" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="F9BAE3A7FF88FF90FFA7FEE2CB21FE9B" box="[470,558,271,297]" captionStart="FIGURES 12 – 17" captionStartId="8.[151,269,1574,1598]" captionTargetBox="[157,1429,367,1512]" captionTargetId="figure@8.[151,1436,349,1550]" captionTargetPageId="8" captionText="FIGURES 12 – 17. Microtityus bivicentorum sp. nov., distribution of the trichobothria in male holotype (illustrated from the right pedipalp). 12. Femur, dorsal view. 13. Femur, internal view. 14. Patella, dorsal view. 15. Patella, external view. 16. Chela, external view. 17. Chela, ventral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/188074/files/figure.png" pageId="9" pageNumber="36">Fig. 14</figureCitation>
|
||
) but present in
|
||
<taxonomicName id="A68184A1FF88FF90FC9CFEE2CA8CFE9A" ID-CoL="73JT9" box="[749,899,271,296]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="36" phylum="Arthropoda" rank="species" species="desuzeae">
|
||
<emphasis id="53F52330FF88FF90FC9CFEE2CA8CFE9A" box="[749,899,271,296]" italics="true" pageId="9" pageNumber="36">M. desuzeae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="051082D3FF88FF90FDE2FEE2CC23FE9B" author="Gonzalez-Sponga" box="[915,1324,271,297]" pageId="9" pageNumber="38" refString="Gonzalez-Sponga, M. A. (2001) Aracnidos de Venezuela. Seis nuevas especies del genero Microtityus (Scorpionida: Buthidae) del sistema montanoso de la costa. Boletin de la Academia de Ciencias Fisicas, Matematicas y Naturales, 61 (1 - 2), 45 - 66." type="journal article" year="2001">González-Sponga 2001: figs. 7–8</bibRefCitation>
|
||
), and the number of granular rows on the pedipalp movable finger,
|
||
<quantity id="A67952C7FF88FF90FD20FED8CAB5FEFD" box="[849,954,309,335]" metricMagnitude="-1" metricUnit="m" metricValue="2.921" metricValueMax="3.048" metricValueMin="2.794" pageId="9" pageNumber="36" unit="in" value="11.5" valueMax="12.0" valueMin="11.0">11–12 in</quantity>
|
||
the former species but nine in the latter (
|
||
<bibRefCitation id="051082D3FF88FF90FEEEFEB1C8B8FEC4" author="Gonzalez-Sponga" box="[159,439,348,374]" pageId="9" pageNumber="38" refString="Gonzalez-Sponga, M. A. (2001) Aracnidos de Venezuela. Seis nuevas especies del genero Microtityus (Scorpionida: Buthidae) del sistema montanoso de la costa. Boletin de la Academia de Ciencias Fisicas, Matematicas y Naturales, 61 (1 - 2), 45 - 66." type="journal article" year="2001">González-Sponga 2001</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |