360 lines
63 KiB
XML
360 lines
63 KiB
XML
<document id="76D05CBACF37A525ADA616CAFBE0FE01" ID-CLB-Dataset="53164" ID-DOI="10.11646/zootaxa.5124.1.1" ID-GBIF-Dataset="9666d599-2c7d-4a52-bce2-a8794f03265b" ID-ISSN="1175-5326" ID-Zenodo-Dep="6404674" ID-ZooBank="C015F8CB-799B-4A92-90AE-02B4C576089E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648795871367" checkinUser="plazi" docAuthor="O’Hara, Timothy D. & Thuy, Ben" docDate="2022" docId="C72D87A1534AFF87FEA385DCFAEDFC08" docLanguage="en" docName="zootaxa.5124.1.1.pdf" docOrigin="Zootaxa 5124 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ophiolebes felli O’Hara & Thuy 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="24" masterDocId="3B14FFD9535EFF90FE348503FFA8FFAE" masterDocTitle="Biogeography and taxonomy of Ophiuroidea (Echinodermata) from the Îles Saint- Paul and Amsterdam in the southern Indian Ocean" masterLastPageNumber="49" masterPageNumber="1" pageNumber="21" updateTime="1699310728982" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="7947C440E65F71D48535C723EE37DAF1">Biogeography and taxonomy of Ophiuroidea (Echinodermata) from the Îles Saint- Paul and Amsterdam in the southern Indian Ocean</mods:title>
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<mods:name id="9FBBD97512ECE80D959C737A2CD09690" type="personal">
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<mods:namePart id="CC0A78301B94C9A6499A69C84E357944">O’Hara, Timothy D.</mods:namePart>
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<mods:affiliation id="2362649191088BC804412E2DDF59365C">Museums Victoria, GPO Box 666 E, Melbourne, 3001, AUSTRALIA,</mods:affiliation>
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<mods:namePart id="4663D91995F259E60C005EACF7BE44CF">Thuy, Ben</mods:namePart>
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<mods:affiliation id="8DA2E18D2CE57BCE80547C1EC6602570">Natural History Museum of Luxembourg, 24 Rue Münster, 2160 Luxembourg</mods:affiliation>
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<mods:title id="B2F75F3926385D6F054DF7645EAB1595">Zootaxa</mods:title>
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<mods:date id="4EE7B02479DE74D83688AF0D31AC4E23">2022</mods:date>
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<treatment id="C72D87A1534AFF87FEA385DCFAEDFC08" ID-DOI="http://doi.org/10.5281/zenodo.6404718" ID-GBIF-Taxon="194442338" ID-Zenodo-Dep="6404718" LSID="urn:lsid:plazi:treatment:C72D87A1534AFF87FEA385DCFAEDFC08" httpUri="http://treatment.plazi.org/id/C72D87A1534AFF87FEA385DCFAEDFC08" lastPageId="23" lastPageNumber="24" pageId="20" pageNumber="21">
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<subSubSection id="079E653C534AFF84FEA385DCFE12FF57" box="[151,442,223,250]" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="4F3B36B7534AFF84FEA385DCFE12FF57" blockId="20.[151,1193,223,321]" box="[151,442,223,250]" pageId="20" pageNumber="21">
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<heading id="147381DB534AFF84FEA385DCFE12FF57" bold="true" box="[151,442,223,250]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
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<emphasis id="7DF0EAA5534AFF84FEA385DCFE12FF57" bold="true" box="[151,442,223,250]" pageId="20" pageNumber="21">
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<taxonomicName id="88844D34534AFF84FEA385DCFEFDFF54" ID-CoL="8ZDNK" authority="O’Hara & Thuy, 2022" authorityName="O’Hara & Thuy" authorityYear="2022" box="[151,341,223,250]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="20" pageNumber="21" phylum="Echinodermata" rank="species" species="felli" status="sp. nov.">
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<emphasis id="7DF0EAA5534AFF84FEA385DCFEFDFF54" bold="true" box="[151,341,223,250]" italics="true" pageId="20" pageNumber="21">Ophiolebes felli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="66C357DE534AFF84FF6885DCFE12FF57" box="[348,442,223,249]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="079E653C534AFF84FEA38400FB01FEEF" pageId="20" pageNumber="21" type="description">
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<paragraph id="4F3B36B7534AFF84FEA38400FEF8FEB3" blockId="20.[151,1193,223,321]" box="[151,336,259,285]" pageId="20" pageNumber="21">
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<figureCitation id="D7BF2A32534AFF84FEA38400FEADFEB3" box="[151,261,259,285]" captionStart="FIGURE 7" captionStartId="21.[151,250,1920,1944]" captionTargetBox="[151,1435,181,1896]" captionTargetId="figure-17@21.[151,1435,181,1896]" captionTargetPageId="21" captionText="FIGURE 7. (A–B) Ophiosemnotes conferta, MNHN IE.2009.1574, A: dorsal and B: ventral views. (C–G) Ophiolebes felli sp. nov., holotype, MNHN IE.2019.4746, C: dorsal outline on black coral, D: dorsal, E: ventral view, F: proximal arm, G: arm tip. (H–I) paratype, MNHN IE.2009.1599, H: dorsal and I: ventral views." figureDoi="http://doi.org/10.5281/zenodo.6404694" httpUri="https://zenodo.org/record/6404694/files/figure.png" pageId="20" pageNumber="21">Fig. 7C–I</figureCitation>
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,
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<figureCitation id="D7BF2A32534AFF84FF258400FEF8FEB3" box="[273,336,259,285]" captionStart="FIGURE 8" captionStartId="22.[151,250,1824,1848]" captionTargetBox="[177,1411,181,1800]" captionTargetId="figure-26@22.[177,1411,181,1800]" captionTargetPageId="22" captionText="FIGURE 8. (A–S) Ophiolebes felli sp. nov. paratype, IE.2009.1569. A–B: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D–E: arm spines. F: tentacle scale. G–K: proximal vertebra in distal (G), proximal (H), lateral (I), dorsal (J) and ventral (K) views. L: disc granule. M: ventral arm plate. N: dorsal arm plate. O: oral plate split in proximal and distal halves. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404698" httpUri="https://zenodo.org/record/6404698/files/figure.png" pageId="20" pageNumber="21">8A–S</figureCitation>
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</paragraph>
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<paragraph id="4F3B36B7534AFF84FEA38424FB01FEEF" blockId="20.[151,1193,223,321]" box="[151,1193,295,321]" pageId="20" pageNumber="21">
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http://zoobank.org/
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<subSubSection id="079E653C534AFF84FEA3846CFE52F817" pageId="20" pageNumber="21" type="materials_examined">
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<paragraph id="4F3B36B7534AFF84FEA3846CFBB6FE00" blockId="20.[151,1437,367,1978]" pageId="20" pageNumber="21">
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<emphasis id="7DF0EAA5534AFF84FEA3846CFEEAFE27" bold="true" box="[151,322,367,393]" pageId="20" pageNumber="21">Type material</emphasis>
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.
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<materialsCitation id="FFEC3CEA534AFF84FF61846CFF55FE03" ID-GBIF-Occurrence="3746276313" collectingDate="1986-07-09" collectionCode="MD" elevation="940" latitude="-37.786667" longLatPrecision="92" longitude="77.649666" pageId="20" pageNumber="21" specimenCode="MD50" specimenCount="1">
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<specimenCode id="1F229ECC534AFF84FF61846CFE37FE24" box="[341,415,367,394]" collectionCode="MD" country="Germany" name="Museum Donaueschingen" pageId="20" pageNumber="21">MD50</specimenCode>
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CP7, north-east of Île Amsterdam,
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,
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<geoCoordinate id="2AB05070534AFF84FDE3846CFBCEFE27" box="[983,1126,367,393]" degrees="77" direction="east" minutes="38.98" orientation="longitude" pageId="20" pageNumber="21" precision="9" value="77.649666">77° 38.98´E</geoCoordinate>
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, 1680–
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<quantity id="887C9B52534AFF84FA89846CFB57FE27" box="[1213,1279,367,393]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="20" pageNumber="21" unit="m" value="940.0">
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<elevation id="C4A9D184534AFF84FA89846CFB57FE27" box="[1213,1279,367,393]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="20" pageNumber="21" unit="m" value="940.0">940m</elevation>
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</quantity>
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, Beam trawl,
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<date id="3B3A1077534AFF84FEA38490FF55FE03" box="[151,253,403,429]" pageId="20" pageNumber="21" value="1986-07-09">
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<collectingDate id="2B7EE99F534AFF84FEA38490FF55FE03" box="[151,253,403,429]" pageId="20" pageNumber="21" value="1986-07-09">9/7/1986</collectingDate>
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,
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<materialsCitation id="FFEC3CEA534AFF84FF3D8497FDD4FE00" ID-GBIF-Occurrence="3746276306" box="[265,636,403,430]" collectionCode="MNHN, IE" pageId="20" pageNumber="21" specimenCount="1" typeStatus="holotype">
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<collectionCode id="2995AE72534AFF84FF3D8497FEF7FE00" box="[265,351,404,430]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="21">MNHN</collectionCode>
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<collectionCode id="2995AE72534AFF84FF518490FE28FE03" box="[357,384,403,429]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="20" pageNumber="21">IE</collectionCode>
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.2019.4746 (
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<typeStatus id="903F8815534AFF84FC398490FDDDFE00" box="[525,629,403,430]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
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)
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</materialsCitation>
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,
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<materialsCitation id="FFEC3CEA534AFF84FCB38497FBB2FE00" ID-GBIF-Occurrence="3746276311" box="[647,1050,403,430]" collectionCode="MNHN, IE" pageId="20" pageNumber="21" specimenCount="9" typeStatus="paratype">
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<collectionCode id="2995AE72534AFF84FCB38497FD75FE00" box="[647,733,404,430]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="21">MNHN</collectionCode>
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<collectionCode id="2995AE72534AFF84FCD08490FD57FE00" box="[740,767,403,430]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="20" pageNumber="21">IE</collectionCode>
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.2009.1569 (
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<specimenCount id="5982FD3E534AFF84FDB88490FBBBFE00" box="[908,1043,403,430]" pageId="20" pageNumber="21" type="generic" typeStatus="paratypes">9 paratypes</specimenCount>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="4F3B36B7534AFF84FEF384B4FF76FD5F" blockId="20.[151,1437,367,1978]" pageId="20" pageNumber="21">
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<emphasis id="7DF0EAA5534AFF84FEF384B4FE68FE7F" bold="true" box="[199,448,439,465]" pageId="20" pageNumber="21">
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<typeStatus id="903F8815534AFF84FEF384B4FE9CFE7F" box="[199,308,439,465]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
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description
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</emphasis>
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. Disc is
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||
<quantity id="887C9B52534AFF84FC2B84B4FDDCFE7C" box="[543,628,439,466]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="20" pageNumber="21" unit="mm" value="8.3">8.3 mm</quantity>
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dd, arms are coiled distally but are at least
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<quantity id="887C9B52534AFF84FA6484B4FB36FE7C" box="[1104,1182,439,466]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="21" unit="mm" value="25.0">25 mm</quantity>
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long. Both sides of the disc and the arms are covered in skin that obscures underlying plates. Disc is five lobed, constricted interradially, covered in skin with embedded low conical granules (
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<figureCitation id="D7BF2A32534AFF84FCC384FCFCFAFDB4" box="[759,850,511,538]" captionStart="FIGURE 7" captionStartId="21.[151,250,1920,1944]" captionTargetBox="[151,1435,181,1896]" captionTargetId="figure-17@21.[151,1435,181,1896]" captionTargetPageId="21" captionText="FIGURE 7. (A–B) Ophiosemnotes conferta, MNHN IE.2009.1574, A: dorsal and B: ventral views. (C–G) Ophiolebes felli sp. nov., holotype, MNHN IE.2019.4746, C: dorsal outline on black coral, D: dorsal, E: ventral view, F: proximal arm, G: arm tip. (H–I) paratype, MNHN IE.2009.1599, H: dorsal and I: ventral views." figureDoi="http://doi.org/10.5281/zenodo.6404694" httpUri="https://zenodo.org/record/6404694/files/figure.png" pageId="20" pageNumber="21">Fig. 7D</figureCitation>
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). The granules are largest on and between the radial shields and in the centre of the disc (
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<quantity id="887C9B52534AFF84FC008720FD63FD90" box="[564,715,547,574]" metricMagnitude="-4" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" pageId="20" pageNumber="21" unit="mm" value="0.285" valueMax="0.3" valueMin="0.27">0.27–0.3 mm</quantity>
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dia), on the dorsal and ventral interradii they are smaller (
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<quantity id="887C9B52534AFF84FB6F8720FEAAFDCC" metricMagnitude="-4" metricUnit="m" metricValue="2.15" metricValueMax="2.5" metricValueMin="1.7999999999999998" pageId="20" pageNumber="21" unit="mm" value="0.215" valueMax="0.25" valueMin="0.18">0.18– 0.25 mm</quantity>
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), sparser and separated by skin and small scales. The wide genital slits are contiguous with the first 2 arm segments (
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<figureCitation id="D7BF2A32534AFF84FF768768FE31FD28" box="[322,409,619,646]" captionStart="FIGURE 7" captionStartId="21.[151,250,1920,1944]" captionTargetBox="[151,1435,181,1896]" captionTargetId="figure-17@21.[151,1435,181,1896]" captionTargetPageId="21" captionText="FIGURE 7. (A–B) Ophiosemnotes conferta, MNHN IE.2009.1574, A: dorsal and B: ventral views. (C–G) Ophiolebes felli sp. nov., holotype, MNHN IE.2019.4746, C: dorsal outline on black coral, D: dorsal, E: ventral view, F: proximal arm, G: arm tip. (H–I) paratype, MNHN IE.2009.1599, H: dorsal and I: ventral views." figureDoi="http://doi.org/10.5281/zenodo.6404694" httpUri="https://zenodo.org/record/6404694/files/figure.png" pageId="20" pageNumber="21">Fig. 7E</figureCitation>
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). Oral shields are rhomboid, 1.5 times wider than long; adorals are almost of the same size, trapezoid, widest proximally, 2 times as wide as long, meeting broadly interradially, slightly tumid; teeth robust, pentagonal, pointed at tip, 3 conical papillae on the ventral margin of the oral plate, the adoral shield spine is slightly larger.
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</paragraph>
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||
<paragraph id="4F3B36B7534AFF84FEF387F8FE7DFBBC" blockId="20.[151,1437,367,1978]" pageId="20" pageNumber="21">
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||
DAPs are fan-shaped, as wide as long, with a rounded distal margin and acute angle proximally, separate. Some disc granules are present along the base of the arm and between DAPs of the first few segments. VAPs are wider than long, pentagonal with an obtuse proximal angle, rounded distal margin and straight lateral margins, separate. Arm spines are rounded in cross section and tapering to a blunt rounded tip, rugose but not thorny. First segments have 3 short spines, 4 on next 2 segments; beyond the disc there are 5–6 arm spines, uppermost to as long as a segment, lower spines 2/3 to half as long (
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<figureCitation id="D7BF2A32534AFF84FC5186ACFD11FC64" box="[613,697,943,970]" captionStart="FIGURE 7" captionStartId="21.[151,250,1920,1944]" captionTargetBox="[151,1435,181,1896]" captionTargetId="figure-17@21.[151,1435,181,1896]" captionTargetPageId="21" captionText="FIGURE 7. (A–B) Ophiosemnotes conferta, MNHN IE.2009.1574, A: dorsal and B: ventral views. (C–G) Ophiolebes felli sp. nov., holotype, MNHN IE.2019.4746, C: dorsal outline on black coral, D: dorsal, E: ventral view, F: proximal arm, G: arm tip. (H–I) paratype, MNHN IE.2009.1599, H: dorsal and I: ventral views." figureDoi="http://doi.org/10.5281/zenodo.6404694" httpUri="https://zenodo.org/record/6404694/files/figure.png" pageId="20" pageNumber="21">Fig. 7F</figureCitation>
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). Near the arm tip, the 3 spines become slender, subequal, less than a segment long and have thorns on the ventral side (
|
||
<figureCitation id="D7BF2A32534AFF84FCD786D0FC95FC43" box="[739,829,979,1005]" captionStart="FIGURE 7" captionStartId="21.[151,250,1920,1944]" captionTargetBox="[151,1435,181,1896]" captionTargetId="figure-17@21.[151,1435,181,1896]" captionTargetPageId="21" captionText="FIGURE 7. (A–B) Ophiosemnotes conferta, MNHN IE.2009.1574, A: dorsal and B: ventral views. (C–G) Ophiolebes felli sp. nov., holotype, MNHN IE.2019.4746, C: dorsal outline on black coral, D: dorsal, E: ventral view, F: proximal arm, G: arm tip. (H–I) paratype, MNHN IE.2009.1599, H: dorsal and I: ventral views." figureDoi="http://doi.org/10.5281/zenodo.6404694" httpUri="https://zenodo.org/record/6404694/files/figure.png" pageId="20" pageNumber="21">Fig. 7G</figureCitation>
|
||
). Tentacle pores are small with a tiny pointed tentacle scale on proximal segments.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B7534AFF84FEF38118FCFFFBD3" blockId="20.[151,1437,367,1978]" pageId="20" pageNumber="21">
|
||
<emphasis id="7DF0EAA5534AFF84FEF38118FE12FB9B" bold="true" box="[199,442,1051,1077]" pageId="20" pageNumber="21">
|
||
<typeStatus id="903F8815534AFF84FEF38118FE9CFB9B" box="[199,308,1051,1077]" pageId="20" pageNumber="21" type="paratype">Paratype</typeStatus>
|
||
variations.
|
||
</emphasis>
|
||
<quantity id="887C9B52534AFF84FFF58118FDE1FB98" box="[449,585,1051,1078]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="20" pageNumber="21" unit="mm" value="5.0" valueMax="5.5" valueMin="4.5">4.5–5.5 mm</quantity>
|
||
dd (
|
||
<figureCitation id="D7BF2A32534AFF84FC4F8118FD46FB98" box="[635,750,1051,1078]" captionStart="FIGURE 7" captionStartId="21.[151,250,1920,1944]" captionTargetBox="[151,1435,181,1896]" captionTargetId="figure-17@21.[151,1435,181,1896]" captionTargetPageId="21" captionText="FIGURE 7. (A–B) Ophiosemnotes conferta, MNHN IE.2009.1574, A: dorsal and B: ventral views. (C–G) Ophiolebes felli sp. nov., holotype, MNHN IE.2019.4746, C: dorsal outline on black coral, D: dorsal, E: ventral view, F: proximal arm, G: arm tip. (H–I) paratype, MNHN IE.2009.1599, H: dorsal and I: ventral views." figureDoi="http://doi.org/10.5281/zenodo.6404694" httpUri="https://zenodo.org/record/6404694/files/figure.png" pageId="20" pageNumber="21">Fig. 7H–I</figureCitation>
|
||
). On one broken specimen, a vertical row of 5 flattened teeth are visible with a rounded distal edge. On some specimens there are only 2 lateral oral papillae on some jaws. The tentacle scales are difficult to see and rare on many specimens.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B7534AFF84FEF38184FC34F9A4" blockId="20.[151,1437,367,1978]" pageId="20" pageNumber="21">
|
||
<emphasis id="7DF0EAA5534AFF84FEF38184FD8BFB0F" bold="true" box="[199,547,1159,1185]" pageId="20" pageNumber="21">
|
||
<typeStatus id="903F8815534AFF84FEF38184FE9CFB0F" box="[199,308,1159,1185]" pageId="20" pageNumber="21" type="paratype">Paratype</typeStatus>
|
||
ossicle description.
|
||
</emphasis>
|
||
Proximal LAPs (
|
||
<figureCitation id="D7BF2A32534AFF84FCC68184FCC6FB0F" box="[754,878,1159,1185]" captionStart="FIGURE 8" captionStartId="22.[151,250,1824,1848]" captionTargetBox="[177,1411,181,1800]" captionTargetId="figure-26@22.[177,1411,181,1800]" captionTargetPageId="22" captionText="FIGURE 8. (A–S) Ophiolebes felli sp. nov. paratype, IE.2009.1569. A–B: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D–E: arm spines. F: tentacle scale. G–K: proximal vertebra in distal (G), proximal (H), lateral (I), dorsal (J) and ventral (K) views. L: disc granule. M: ventral arm plate. N: dorsal arm plate. O: oral plate split in proximal and distal halves. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404698" httpUri="https://zenodo.org/record/6404698/files/figure.png" pageId="20" pageNumber="21">Fig. 8A–B</figureCitation>
|
||
) are approximately as high as long, of rounded outline, tumid, thick, with strongly convex distal edge; ventral edge convex, not protruding; dorsal edge oblique, very weakly concave; outer surface with coarsely meshed stereom, with trabecular intersections transformed into small, inconspicuous tubercles, no vertical striation; outer proximal edge with band of more finely meshed stereom, with a single, poorly defined spur in the dorsal half of the proximal edge. Spine articulations (
|
||
<figureCitation id="D7BF2A32534AFF84FA6C8014FB04FA9F" box="[1112,1196,1303,1329]" captionStart="FIGURE 8" captionStartId="22.[151,250,1824,1848]" captionTargetBox="[177,1411,181,1800]" captionTargetId="figure-26@22.[177,1411,181,1800]" captionTargetPageId="22" captionText="FIGURE 8. (A–S) Ophiolebes felli sp. nov. paratype, IE.2009.1569. A–B: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D–E: arm spines. F: tentacle scale. G–K: proximal vertebra in distal (G), proximal (H), lateral (I), dorsal (J) and ventral (K) views. L: disc granule. M: ventral arm plate. N: dorsal arm plate. O: oral plate split in proximal and distal halves. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404698" httpUri="https://zenodo.org/record/6404698/files/figure.png" pageId="20" pageNumber="21">Fig. 8C</figureCitation>
|
||
) are freestanding in a continuous vertical row on a raised distal portion of the LAP, ear-shaped, relatively small, composed of proximally merged dorsal and ventral lobes enclosing a large muscle opening and a smaller nerve opening, with a weakly developed sigmoidal fold. Inner side of LAPs with a large, prominent, moderately well-defined, digit-1-shaped vertebral articular ridge; poorly-defined spur in the dorsal half of the inner distal edge corresponding to the spur on the outer distal edge of the LAP; tentacle notch small, oblique, shallow. Very small, inconspicuous perforations occur close to the ventro-distal tip of the vertebral articular structure.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B7534AFF84FEF38310FD8CF937" blockId="20.[151,1437,367,1978]" pageId="20" pageNumber="21">
|
||
Vertebrae (
|
||
<figureCitation id="D7BF2A32534AFF84FF758310FE12F983" box="[321,442,1555,1581]" captionStart="FIGURE 8" captionStartId="22.[151,250,1824,1848]" captionTargetBox="[177,1411,181,1800]" captionTargetId="figure-26@22.[177,1411,181,1800]" captionTargetPageId="22" captionText="FIGURE 8. (A–S) Ophiolebes felli sp. nov. paratype, IE.2009.1569. A–B: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D–E: arm spines. F: tentacle scale. G–K: proximal vertebra in distal (G), proximal (H), lateral (I), dorsal (J) and ventral (K) views. L: disc granule. M: ventral arm plate. N: dorsal arm plate. O: oral plate split in proximal and distal halves. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404698" httpUri="https://zenodo.org/record/6404698/files/figure.png" pageId="20" pageNumber="21">Fig. 8G–K</figureCitation>
|
||
) compact, tumid, with small, narrow distal and proximal muscle fossae; large, hourglassshaped (streptospondylous) distal and proximal articulations; oblique P-shaped lateral articulation structure corresponding with articular structure on the inner side of the LAP; large, deep dorsal and ventral furrows, with small, inconspicuous podial basins.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B7534AFF84FEF383A0FE52F817" blockId="20.[151,1437,367,1978]" pageId="20" pageNumber="21">
|
||
Oral plates (
|
||
<figureCitation id="D7BF2A32534AFF84FF6483A0FE00F913" box="[336,424,1699,1725]" captionStart="FIGURE 8" captionStartId="22.[151,250,1824,1848]" captionTargetBox="[177,1411,181,1800]" captionTargetId="figure-26@22.[177,1411,181,1800]" captionTargetPageId="22" captionText="FIGURE 8. (A–S) Ophiolebes felli sp. nov. paratype, IE.2009.1569. A–B: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D–E: arm spines. F: tentacle scale. G–K: proximal vertebra in distal (G), proximal (H), lateral (I), dorsal (J) and ventral (K) views. L: disc granule. M: ventral arm plate. N: dorsal arm plate. O: oral plate split in proximal and distal halves. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404698" httpUri="https://zenodo.org/record/6404698/files/figure.png" pageId="20" pageNumber="21">Fig. 8O</figureCitation>
|
||
) split into distal and proximal halves, approximately as high as long, with adradial muscle fossa lining the ventro-distal edge of articulation area. Dental plates (
|
||
<figureCitation id="D7BF2A32534AFF84FDA883C4FC47F94F" box="[924,1007,1735,1761]" captionStart="FIGURE 8" captionStartId="22.[151,250,1824,1848]" captionTargetBox="[177,1411,181,1800]" captionTargetId="figure-26@22.[177,1411,181,1800]" captionTargetPageId="22" captionText="FIGURE 8. (A–S) Ophiolebes felli sp. nov. paratype, IE.2009.1569. A–B: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D–E: arm spines. F: tentacle scale. G–K: proximal vertebra in distal (G), proximal (H), lateral (I), dorsal (J) and ventral (K) views. L: disc granule. M: ventral arm plate. N: dorsal arm plate. O: oral plate split in proximal and distal halves. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404698" httpUri="https://zenodo.org/record/6404698/files/figure.png" pageId="20" pageNumber="21">Fig. 8P</figureCitation>
|
||
) are undivided, narrow, rounded, with single row of small, shallow tooth sockets surrounded by weakly-defined protruding ring. Abradial genital plates (
|
||
<figureCitation id="D7BF2A32534AFF84FE94820CFF50F887" box="[160,248,1807,1833]" captionStart="FIGURE 8" captionStartId="22.[151,250,1824,1848]" captionTargetBox="[177,1411,181,1800]" captionTargetId="figure-26@22.[177,1411,181,1800]" captionTargetPageId="22" captionText="FIGURE 8. (A–S) Ophiolebes felli sp. nov. paratype, IE.2009.1569. A–B: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D–E: arm spines. F: tentacle scale. G–K: proximal vertebra in distal (G), proximal (H), lateral (I), dorsal (J) and ventral (K) views. L: disc granule. M: ventral arm plate. N: dorsal arm plate. O: oral plate split in proximal and distal halves. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404698" httpUri="https://zenodo.org/record/6404698/files/figure.png" pageId="20" pageNumber="21">Fig. 8Q</figureCitation>
|
||
) are almost as long as the adradial ones (
|
||
<figureCitation id="D7BF2A32534AFF84FCF8820CFC8DF887" box="[716,805,1807,1833]" captionStart="FIGURE 8" captionStartId="22.[151,250,1824,1848]" captionTargetBox="[177,1411,181,1800]" captionTargetId="figure-26@22.[177,1411,181,1800]" captionTargetPageId="22" captionText="FIGURE 8. (A–S) Ophiolebes felli sp. nov. paratype, IE.2009.1569. A–B: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D–E: arm spines. F: tentacle scale. G–K: proximal vertebra in distal (G), proximal (H), lateral (I), dorsal (J) and ventral (K) views. L: disc granule. M: ventral arm plate. N: dorsal arm plate. O: oral plate split in proximal and distal halves. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404698" httpUri="https://zenodo.org/record/6404698/files/figure.png" pageId="20" pageNumber="21">Fig. 8R</figureCitation>
|
||
), relatively thick, bar-like, without ridges or groves and with a straight adradio-distal tip. Radial shields (
|
||
<figureCitation id="D7BF2A32534AFF84FC8C8230FCA3F8E3" box="[696,779,1843,1869]" captionStart="FIGURE 8" captionStartId="22.[151,250,1824,1848]" captionTargetBox="[177,1411,181,1800]" captionTargetId="figure-26@22.[177,1411,181,1800]" captionTargetPageId="22" captionText="FIGURE 8. (A–S) Ophiolebes felli sp. nov. paratype, IE.2009.1569. A–B: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D–E: arm spines. F: tentacle scale. G–K: proximal vertebra in distal (G), proximal (H), lateral (I), dorsal (J) and ventral (K) views. L: disc granule. M: ventral arm plate. N: dorsal arm plate. O: oral plate split in proximal and distal halves. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404698" httpUri="https://zenodo.org/record/6404698/files/figure.png" pageId="20" pageNumber="21">Fig. 8S</figureCitation>
|
||
) are strongly elongated, isosceles-triangular outline with a slightly oblique proximal tip; no ornamentation on the outer surface. Proximal dorsal arm spines (
|
||
<figureCitation id="D7BF2A32534AFF84FAD48254FA91F8DF" box="[1248,1337,1879,1905]" captionStart="FIGURE 8" captionStartId="22.[151,250,1824,1848]" captionTargetBox="[177,1411,181,1800]" captionTargetId="figure-26@22.[177,1411,181,1800]" captionTargetPageId="22" captionText="FIGURE 8. (A–S) Ophiolebes felli sp. nov. paratype, IE.2009.1569. A–B: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D–E: arm spines. F: tentacle scale. G–K: proximal vertebra in distal (G), proximal (H), lateral (I), dorsal (J) and ventral (K) views. L: disc granule. M: ventral arm plate. N: dorsal arm plate. O: oral plate split in proximal and distal halves. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404698" httpUri="https://zenodo.org/record/6404698/files/figure.png" pageId="20" pageNumber="21">Fig. 8D</figureCitation>
|
||
), ventral arm spines (
|
||
<figureCitation id="D7BF2A32534AFF84FF148278FEDFF83B" box="[288,375,1915,1941]" captionStart="FIGURE 8" captionStartId="22.[151,250,1824,1848]" captionTargetBox="[177,1411,181,1800]" captionTargetId="figure-26@22.[177,1411,181,1800]" captionTargetPageId="22" captionText="FIGURE 8. (A–S) Ophiolebes felli sp. nov. paratype, IE.2009.1569. A–B: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D–E: arm spines. F: tentacle scale. G–K: proximal vertebra in distal (G), proximal (H), lateral (I), dorsal (J) and ventral (K) views. L: disc granule. M: ventral arm plate. N: dorsal arm plate. O: oral plate split in proximal and distal halves. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404698" httpUri="https://zenodo.org/record/6404698/files/figure.png" pageId="20" pageNumber="21">Fig. 8E</figureCitation>
|
||
), tentacle scales (
|
||
<figureCitation id="D7BF2A32534AFF84FC0A8278FD3CF83B" box="[574,660,1915,1941]" captionStart="FIGURE 8" captionStartId="22.[151,250,1824,1848]" captionTargetBox="[177,1411,181,1800]" captionTargetId="figure-26@22.[177,1411,181,1800]" captionTargetPageId="22" captionText="FIGURE 8. (A–S) Ophiolebes felli sp. nov. paratype, IE.2009.1569. A–B: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D–E: arm spines. F: tentacle scale. G–K: proximal vertebra in distal (G), proximal (H), lateral (I), dorsal (J) and ventral (K) views. L: disc granule. M: ventral arm plate. N: dorsal arm plate. O: oral plate split in proximal and distal halves. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404698" httpUri="https://zenodo.org/record/6404698/files/figure.png" pageId="20" pageNumber="21">Fig. 8F</figureCitation>
|
||
), disc granules (
|
||
<figureCitation id="D7BF2A32534AFF84FD788278FC0BF83B" box="[844,931,1915,1941]" captionStart="FIGURE 8" captionStartId="22.[151,250,1824,1848]" captionTargetBox="[177,1411,181,1800]" captionTargetId="figure-26@22.[177,1411,181,1800]" captionTargetPageId="22" captionText="FIGURE 8. (A–S) Ophiolebes felli sp. nov. paratype, IE.2009.1569. A–B: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D–E: arm spines. F: tentacle scale. G–K: proximal vertebra in distal (G), proximal (H), lateral (I), dorsal (J) and ventral (K) views. L: disc granule. M: ventral arm plate. N: dorsal arm plate. O: oral plate split in proximal and distal halves. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404698" httpUri="https://zenodo.org/record/6404698/files/figure.png" pageId="20" pageNumber="21">Fig. 8L</figureCitation>
|
||
), ventral arm plate (
|
||
<figureCitation id="D7BF2A32534AFF84FABC8278FB4CF83B" box="[1160,1252,1915,1941]" captionStart="FIGURE 8" captionStartId="22.[151,250,1824,1848]" captionTargetBox="[177,1411,181,1800]" captionTargetId="figure-26@22.[177,1411,181,1800]" captionTargetPageId="22" captionText="FIGURE 8. (A–S) Ophiolebes felli sp. nov. paratype, IE.2009.1569. A–B: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D–E: arm spines. F: tentacle scale. G–K: proximal vertebra in distal (G), proximal (H), lateral (I), dorsal (J) and ventral (K) views. L: disc granule. M: ventral arm plate. N: dorsal arm plate. O: oral plate split in proximal and distal halves. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404698" httpUri="https://zenodo.org/record/6404698/files/figure.png" pageId="20" pageNumber="21">Fig. 8M</figureCitation>
|
||
) and dorsal arm plate (
|
||
<figureCitation id="D7BF2A32534AFF84FEE9829CFE9DF817" box="[221,309,1951,1977]" captionStart="FIGURE 8" captionStartId="22.[151,250,1824,1848]" captionTargetBox="[177,1411,181,1800]" captionTargetId="figure-26@22.[177,1411,181,1800]" captionTargetPageId="22" captionText="FIGURE 8. (A–S) Ophiolebes felli sp. nov. paratype, IE.2009.1569. A–B: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D–E: arm spines. F: tentacle scale. G–K: proximal vertebra in distal (G), proximal (H), lateral (I), dorsal (J) and ventral (K) views. L: disc granule. M: ventral arm plate. N: dorsal arm plate. O: oral plate split in proximal and distal halves. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles." figureDoi="http://doi.org/10.5281/zenodo.6404698" httpUri="https://zenodo.org/record/6404698/files/figure.png" pageId="20" pageNumber="21">Fig. 8N</figureCitation>
|
||
) are also figured.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="1BFB663F534BFF85FEA38283FCF0F84E" ID-DOI="http://doi.org/10.5281/zenodo.6404694" ID-Zenodo-Dep="6404694" httpUri="https://zenodo.org/record/6404694/files/figure.png" pageId="21" pageNumber="22" startId="21.[151,250,1920,1944]" targetBox="[151,1435,181,1896]" targetPageId="21">
|
||
<paragraph id="4F3B36B7534BFF85FEA38283FCF0F84E" blockId="21.[151,1437,1920,2017]" pageId="21" pageNumber="22">
|
||
<emphasis id="7DF0EAA5534BFF85FEA38283FEBBF836" bold="true" box="[151,275,1920,1944]" pageId="21" pageNumber="22">FIGURE 7.</emphasis>
|
||
(A–B)
|
||
<taxonomicName id="88844D34534BFF85FF558283FD7DF836" authority=", MNHN IE." authorityName="MNHN IE." box="[353,725,1920,1944]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiosemnotes" kingdom="Animalia" order="Ophiacanthida" pageId="21" pageNumber="22" phylum="Echinodermata" rank="species" species="conferta">
|
||
<emphasis id="7DF0EAA5534BFF85FF558283FDFFF836" box="[353,599,1920,1944]" italics="true" pageId="21" pageNumber="22">Ophiosemnotes conferta</emphasis>
|
||
, MNHN IE.
|
||
</taxonomicName>
|
||
2009.1574, A: dorsal and B: ventral views. (C–G)
|
||
<taxonomicName id="88844D34534BFF85FAE28283FADFF836" authority="O’Hara & Thuy, 2022" authorityName="O’Hara & Thuy" authorityYear="2022" box="[1238,1399,1920,1944]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="21" pageNumber="22" phylum="Echinodermata" rank="species" species="felli" status="sp. nov.">
|
||
<emphasis id="7DF0EAA5534BFF85FAE28283FADFF836" box="[1238,1399,1920,1944]" italics="true" pageId="21" pageNumber="22">Ophiolebes felli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7DF0EAA5534BFF85FB4A8283FF6BF812" bold="true" pageId="21" pageNumber="22">
|
||
<taxonomicNameLabel id="66C357DE534BFF85FB4A8283FF6BF812" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, MNHN IE.2019.4746, C: dorsal outline on black coral, D: dorsal, E: ventral view, F: proximal arm, G: arm tip. (H–I) paratype, MNHN IE.2009.1599, H: dorsal and I: ventral views.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="1BFB663F5348FF86FEA38223FB2CF842" ID-DOI="http://doi.org/10.5281/zenodo.6404698" ID-Zenodo-Dep="6404698" httpUri="https://zenodo.org/record/6404698/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,250,1824,1848]" targetBox="[177,1411,181,1800]" targetPageId="22">
|
||
<paragraph id="4F3B36B75348FF86FEA38223FB2CF842" blockId="22.[151,1437,1824,2029]" pageId="22" pageNumber="23">
|
||
<emphasis id="7DF0EAA55348FF86FEA38223FEBEF896" bold="true" box="[151,278,1824,1848]" pageId="22" pageNumber="23">FIGURE 8.</emphasis>
|
||
(A–S)
|
||
<taxonomicName id="88844D345348FF86FF5C8223FDA5F896" authority="O’Hara & Thuy, 2022" authorityName="O’Hara & Thuy" authorityYear="2022" box="[360,525,1824,1848]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="22" pageNumber="23" phylum="Echinodermata" rank="species" species="felli" status="sp. nov.">
|
||
<emphasis id="7DF0EAA55348FF86FF5C8223FDA5F896" box="[360,525,1824,1848]" italics="true" pageId="22" pageNumber="23">Ophiolebes felli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7DF0EAA55348FF86FC238222FDC3F897" bold="true" box="[535,619,1825,1849]" pageId="22" pageNumber="23">
|
||
<taxonomicNameLabel id="66C357DE5348FF86FC238222FDC3F897" box="[535,619,1825,1849]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
paratype, IE.2009.1569. A–B: proximal lateral arm plate in external (A) and internal (B) views. C: detail of spine articulations. D–E: arm spines. F: tentacle scale. G–K: proximal vertebra in distal (G), proximal (H), lateral (I), dorsal (J) and ventral (K) views. L: disc granule. M: ventral arm plate. N: dorsal arm plate. O: oral plate split in proximal and distal halves. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="079E653C5349FF87FEF38594FC62FCF3" pageId="23" pageNumber="24" type="discussion">
|
||
<paragraph id="4F3B36B75349FF87FEF38594FB31FE03" blockId="23.[151,1437,151,934]" pageId="23" pageNumber="24">
|
||
<emphasis id="7DF0EAA55349FF87FEF38594FE9CFF1F" bold="true" box="[199,308,151,177]" pageId="23" pageNumber="24">Remarks</emphasis>
|
||
. This species is similar to the Atlantic species
|
||
<taxonomicName id="88844D345349FF87FD648594FBD2FF1F" authority="Koehler, 1921" authorityName="Koehler" authorityYear="1921" box="[848,1146,151,177]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="species" species="baccata">
|
||
<emphasis id="7DF0EAA55349FF87FD648594FC65FF1F" box="[848,973,151,177]" italics="true" pageId="23" pageNumber="24">O. baccata</emphasis>
|
||
<bibRefCitation id="2B154B465349FF87FDE28594FBD2FF1F" author="Koehler, R." box="[982,1146,151,177]" pageId="23" pageNumber="24" pagination="1 - 8" refId="ref32488" refString="Koehler, R. (1921) Echinodermes (asteries, ophiures, echinides, et crinoides) des dernieres campagnes de la Princesse-Alice et de l'Hirondelle II. Bulletin de l'Institut Oceanographique, Monaco, 396, 1 - 8." type="journal article" year="1921">Koehler, 1921</bibRefCitation>
|
||
</taxonomicName>
|
||
. However, that species is generally smaller, to
|
||
<quantity id="887C9B525349FF87FFB385B9FE61FF7B" box="[391,457,186,213]" metricMagnitude="0" metricUnit="m" metricValue="4.5" pageId="23" pageNumber="24" unit="m" value="4.5">4.5 m</quantity>
|
||
dd, with smaller (
|
||
<quantity id="887C9B525349FF87FCAF85B8FD5CFF7B" box="[667,756,187,213]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="24" unit="mm" value="0.2">0.2 mm</quantity>
|
||
dia) disc granules, and distally lobed oral shields (
|
||
<bibRefCitation id="2B154B465349FF87FB0E85B8FF7BFF57" author="Paterson, G. L. J." pageId="23" pageNumber="24" pagination="1 - 162" refId="ref35334" refString="Paterson, G. L. J. (1985) The deep-sea Ophiuroidea of the North Atlantic Ocean. Bulletin of the British Museum (Natural History) Zoology, 49, 1 - 162." type="journal article" year="1985">Paterson 1985</bibRefCitation>
|
||
). Other similar species include
|
||
<taxonomicName id="88844D345349FF87FC1C85DCFC9BFF57" authority="Lyman, 1878" authorityName="Lyman" authorityYear="1878" box="[552,819,223,249]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="species" species="vestitus">
|
||
<emphasis id="7DF0EAA55349FF87FC1C85DCFD34FF56" box="[552,668,223,249]" italics="true" pageId="23" pageNumber="24">O. vestitus</emphasis>
|
||
Lyman, 1878
|
||
</taxonomicName>
|
||
from off southern
|
||
<collectingCountry id="379376275349FF87FDC985DCFB92FF57" box="[1021,1082,223,249]" name="Chile" pageId="23" pageNumber="24">Chile</collectingCountry>
|
||
(
|
||
<quantity id="887C9B525349FF87FA7285DCFB6EFF57" box="[1094,1222,223,249]" metricMagnitude="2" metricUnit="m" metricValue="6.0649999999999995" metricValueMax="9.4" metricValueMin="2.73" pageId="23" pageNumber="24" unit="m" value="606.5" valueMax="940.0" valueMin="273.0">273–940 m</quantity>
|
||
) and
|
||
<taxonomicName id="88844D345349FF87FB3485DCFEF3FEB3" authority="A. H. Clark, 1916" authorityName="A. H. Clark" authorityYear="1916" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="species" species="mortenseni">
|
||
<emphasis id="7DF0EAA55349FF87FB3485DCFA34FF56" box="[1280,1436,223,249]" italics="true" pageId="23" pageNumber="24">O. mortenseni</emphasis>
|
||
A.H.
|
||
<bibRefCitation id="2B154B465349FF87FEE28400FEF3FEB3" author="Clark, A. H." box="[214,347,259,285]" pageId="23" pageNumber="24" pagination="115 - 122" refId="ref29774" refString="Clark, A. H. (1916) On a new starfish and five new brittle-stars from the Galapagos Islands. Annals and Magazine of Natural History, 18, 115 - 122. https: // doi. org / 10.1080 / 00222931608693831" type="journal article" year="1916">Clark, 1916</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Galapagos (
|
||
<quantity id="887C9B525349FF87FC678400FD09FEB3" box="[595,673,259,285]" metricMagnitude="2" metricUnit="m" metricValue="7.29" pageId="23" pageNumber="24" unit="m" value="729.0">729 m</quantity>
|
||
). These differ in having a more even covering of disc granules that have the form of short cylindrical stumps with a crown of thorns. The species named as “
|
||
<taxonomicName id="88844D345349FF87FA5F8424FA8BFEEF" box="[1131,1315,295,321]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="species" species="undetermined">Ophiacantha sp.</taxonomicName>
|
||
” in
|
||
<bibRefCitation id="2B154B465349FF87FB648424FF73FECB" author="Manso, C. L. C." pageId="23" pageNumber="24" pagination="1 - 15" refId="ref33583" refString="Manso, C. L. C. (2010) Deep-water Ophiuroidea (Echinodermata) from off Chile in the Eastern South Pacific. Biota Neotropica, 10, 1 - 15. https: // doi. org / 10.1590 / S 1676 - 06032010000200023" type="journal article" year="2010">Manso (2010</bibRefCitation>
|
||
, fig. 6) is also likely to be either
|
||
<taxonomicName id="88844D345349FF87FC7A8448FD6CFECA" authorityName="Lyman" authorityYear="1878" box="[590,708,331,357]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="species" species="vestitus">
|
||
<emphasis id="7DF0EAA55349FF87FC7A8448FD6CFECA" box="[590,708,331,357]" italics="true" pageId="23" pageNumber="24">O. vestitus</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="88844D345349FF87FCDE8448FC21FECA" authorityName="A. H. Clark" authorityYear="1916" box="[746,905,331,357]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="species" species="mortenseni">
|
||
<emphasis id="7DF0EAA55349FF87FCDE8448FC21FECA" box="[746,905,331,357]" italics="true" pageId="23" pageNumber="24">O. mortenseni</emphasis>
|
||
</taxonomicName>
|
||
. The
|
||
<typeStatus id="903F88155349FF87FDFE8448FC52FECB" box="[970,1018,331,357]" pageId="23" pageNumber="24">type</typeStatus>
|
||
of
|
||
<taxonomicName id="88844D345349FF87FA148448FB08FECB" authorityName="Lyman" authorityYear="1878" box="[1056,1184,331,357]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="7DF0EAA55349FF87FA148448FB08FECB" box="[1056,1184,331,357]" italics="true" pageId="23" pageNumber="24">Ophiolebes</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<taxonomicName id="88844D345349FF87FAF48448FF75FE27" authority="Lyman, 1878 b" authorityName="Lyman" authorityYear="1878" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="species" species="scorteus">
|
||
<emphasis id="7DF0EAA55349FF87FAF48448FAE8FECA" box="[1216,1344,331,357]" italics="true" pageId="23" pageNumber="24">O. scorteus</emphasis>
|
||
<bibRefCitation id="2B154B465349FF87FB738448FF75FE27" author="Lyman, T." pageId="23" pageNumber="24" pagination="65 - 168" refId="ref33150" refString="Lyman, T. (1878 b) Ophiuridae and Astrophytidae of the exploring voyage of H. M. S. Challenger, under Prof. Sir Wyville Thomson, F. R. S. Part 1. Bulletin of the Museum of Comparative Zoology, Harvard University, 5, 65 - 168, 10 pls." type="journal article" year="1878">Lyman, 1878b</bibRefCitation>
|
||
</taxonomicName>
|
||
from off subantarctic Marion Island (
|
||
<quantity id="887C9B525349FF87FCB8846DFCA7FE27" box="[652,783,366,393]" metricMagnitude="2" metricUnit="m" metricValue="4.275" metricValueMax="5.76" metricValueMin="2.79" pageId="23" pageNumber="24" unit="m" value="427.5" valueMax="576.0" valueMin="279.0">279–576 m</quantity>
|
||
) which has not been reported since. The granules on that species are smaller and restricted to the radial shields with only a few occurring marginally.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B75349FF87FEF384B4FBE4FD2B" blockId="23.[151,1437,151,934]" pageId="23" pageNumber="24">
|
||
The other Atlantic species,
|
||
<taxonomicName id="88844D345349FF87FFCF84B4FCDBFE7F" authority="H. L. Clark, 1915" authorityName="H. L. Clark" authorityYear="1915" box="[507,883,438,465]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="species" species="pachyphylax">
|
||
<emphasis id="7DF0EAA55349FF87FFCF84B4FD04FE7F" box="[507,684,439,465]" italics="true" pageId="23" pageNumber="24">O. pachyphylax</emphasis>
|
||
H.L.
|
||
<bibRefCitation id="2B154B465349FF87FCDB84B4FCDBFE7F" author="Clark, H. L." box="[751,883,438,465]" pageId="23" pageNumber="24" pagination="165 - 376" refId="ref29982" refString="Clark, H. L. (1915) Catalogue of Recent ophiurans: based on the collection of the Museum of Comparative Zoology. Memoirs of the Museum of Comparative Zoology, Harvard University, 25, 165 - 376, pls. 1 - 20. https: // doi. org / 10.5962 / bhl. title. 48598" type="journal article" year="1915">Clark, 1915</bibRefCitation>
|
||
</taxonomicName>
|
||
has much longer arm spines (to 2 segments) and obvious disc plates bearing the granules, and
|
||
<taxonomicName id="88844D345349FF87FCB884D8FC35FE5B" authority="Koehler, 1895" authorityName="Koehler" authorityYear="1895" box="[652,925,474,501]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="species" species="retecta">
|
||
<emphasis id="7DF0EAA55349FF87FCB884D8FD52FE5A" box="[652,762,475,501]" italics="true" pageId="23" pageNumber="24">O. retecta</emphasis>
|
||
<bibRefCitation id="2B154B465349FF87FCCB84D8FC35FE5B" author="Koehler, R." box="[767,925,474,501]" pageId="23" pageNumber="24" pagination="439 - 496" refId="ref31999" refString="Koehler, R. (1895) Dragages profonds executes a bord du Caudan dans le Golfe de Gascogne Revue biologique du Nord de la France, 7, 439 - 496." type="journal article" year="1895">Koehler, 1895</bibRefCitation>
|
||
</taxonomicName>
|
||
has DAPs that are split into two sections (also present in
|
||
<taxonomicName id="88844D345349FF87FF3F84FCFDE0FDB7" authority="Koehler, 1901" authorityName="Koehler" authorityYear="1901" box="[267,584,511,537]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="species" species="antarctica">
|
||
<emphasis id="7DF0EAA55349FF87FF3F84FCFE37FDB6" box="[267,415,511,537]" italics="true" pageId="23" pageNumber="24">O. antarctica</emphasis>
|
||
<bibRefCitation id="2B154B465349FF87FF9184FCFDE0FDB7" author="Koehler, R." box="[421,584,511,537]" pageId="23" pageNumber="24" pagination="222 - 231" refId="ref32117" refString="Koehler, R. (1901) Note preliminaire sur quelque Ophiures nouvelles provenant des compagnes de la Princesse Alice. Bulletin de la Societe Zoologique de France, 26, 222 - 231." type="journal article" year="1901">Koehler, 1901</bibRefCitation>
|
||
</taxonomicName>
|
||
). There are a number of North Pacific species also referred to
|
||
<taxonomicName id="88844D345349FF87FACE84FCFAD2FDB7" authorityName="Lyman" authorityYear="1878" box="[1274,1402,511,537]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="7DF0EAA55349FF87FACE84FCFAD2FDB7" box="[1274,1402,511,537]" italics="true" pageId="23" pageNumber="24">Ophiolebes</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="2B154B465349FF87FEA38720FE8FFD93" author="Clark, H. L." box="[151,295,547,573]" pageId="23" pageNumber="24" pagination="1 - 302" refId="ref29926" refString="Clark, H. L. (1911) North Pacific ophiurans in the collection of the United States National Museum. Bulletin of the United States National Museum, 75, 1 - 302, figs. 1 - 144. https: // doi. org / 10.5962 / bhl. title. 32556" type="journal article" year="1911">Clark (1911)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="2B154B465349FF87FF078720FDADFD93" author="Matsumoto, H." box="[307,517,547,573]" pageId="23" pageNumber="24" pagination="1 - 408" refId="ref33961" refString="Matsumoto, H. (1917) A monograph of Japanese Ophiuroidea, arranged according to a new classification. Journal of the College of Science, Imperial University Tokyo, 38, 1 - 408, pls. 1 - 7." type="journal article" year="1917">Matsumoto (1917)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="2B154B465349FF87FC258720FD78FD93" author="Djakonov, A. M." box="[529,720,547,573]" pageId="23" pageNumber="24" pagination="4 - 127" refId="ref30635" refString="Djakonov, A. M. (1949) Key to the Echinoderms of the Far Eastern Seas. Izvestiya Tikhookeanskogo Instituta Rybnogo Khozyaistva i Okeanografii, 30, 4 - 127." type="journal article" year="1949">Djakonov (1949)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="2B154B465349FF87FD328720FC02FD93" author="Irimura, S." box="[774,938,547,573]" pageId="23" pageNumber="24" refId="ref31779" refString="Irimura, S. (1982) The Brittle-stars of Sagami Bay. Biological Laboratory, Imperial Household, Tokyo, 95 + 53 pp, 15 leaves of pls." type="book" year="1982">Irimura (1982)</bibRefCitation>
|
||
but none of these have the low disc granules embedded in thick skin of the present species. The
|
||
<collectingCountry id="379376275349FF87FCF98744FCCAFDCF" box="[717,866,583,609]" name="New Zealand" pageId="23" pageNumber="24">New Zealand</collectingCountry>
|
||
species,
|
||
<taxonomicName id="88844D345349FF87FDF38744FA80FDCF" authority="McKnight, 2003" authorityName="McKnight" authorityYear="2003" box="[967,1320,583,609]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="species" species="comatulina">
|
||
<emphasis id="7DF0EAA55349FF87FDF38744FBCEFDCF" box="[967,1126,583,609]" italics="true" pageId="23" pageNumber="24">O. comatulina</emphasis>
|
||
<bibRefCitation id="2B154B465349FF87FA598744FA80FDCF" author="McKnight, D. G." box="[1133,1320,583,609]" pageId="23" pageNumber="24" pagination="1 - 36" refId="ref34183" refString="McKnight, D. G. (2003) New brittle-stars (Echinodermata: Ophiuroidea) from New Zealand waters. Zootaxa, 352, 1 - 36. https: // doi. org / 10.11646 / zootaxa. 352.1.1" type="journal article" year="2003">McKnight, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
, has much longer disc spines. All these species require reassessment of their generic placement.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B75349FF87FEF3878CFD95FC97" blockId="23.[151,1437,151,934]" pageId="23" pageNumber="24">
|
||
Conversely,
|
||
<taxonomicName id="88844D345349FF87FF69878CFCAAFD07" authority="Fell, 1958" authorityName="Fell" authorityYear="1958" box="[349,770,654,681]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiacanthida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="species" species="yaldwyni">
|
||
<emphasis id="7DF0EAA55349FF87FF69878CFDCDFD07" box="[349,613,655,681]" italics="true" pageId="23" pageNumber="24">Ophiacantha yaldwyni</emphasis>
|
||
<bibRefCitation id="2B154B465349FF87FC42878CFCAAFD07" author="Fell, H. B." box="[630,770,654,681]" pageId="23" pageNumber="24" pagination="1 - 40" refId="ref31031" refString="Fell, H. B. (1958) Deep-sea echinoderms of New Zealand. Zoological Publications of the Victoria University Wellington, 24, 1 - 40." type="journal article" year="1958">Fell, 1958</bibRefCitation>
|
||
</taxonomicName>
|
||
from southern
|
||
<collectingCountry id="379376275349FF87FDFE878CFB99FD07" box="[970,1073,655,681]" name="Australia" pageId="23" pageNumber="24">Australia</collectingCountry>
|
||
and
|
||
<collectingCountry id="379376275349FF87FA4E878CFAB2FD07" box="[1146,1306,655,681]" name="New Zealand" pageId="23" pageNumber="24">New Zealand</collectingCountry>
|
||
is similar morphologically to the current species and is herein transferred to
|
||
<taxonomicName id="88844D345349FF87FD4A87B0FC56FD63" authorityName="Lyman" authorityYear="1878" box="[894,1022,691,717]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiolebes" kingdom="Animalia" order="Ophiacanthida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="7DF0EAA55349FF87FD4A87B0FC56FD63" box="[894,1022,691,717]" italics="true" pageId="23" pageNumber="24">Ophiolebes</emphasis>
|
||
</taxonomicName>
|
||
, sharing the tiny tentacle scale, short arm spines, and conical granules embedded in disc skin (
|
||
<bibRefCitation id="2B154B465349FF87FD0487D4FC02FD5F" author="Fell, H. B." box="[816,938,726,753]" pageId="23" pageNumber="24" pagination="1 - 40" refId="ref31031" refString="Fell, H. B. (1958) Deep-sea echinoderms of New Zealand. Zoological Publications of the Victoria University Wellington, 24, 1 - 40." type="journal article" year="1958">Fell, 1958</bibRefCitation>
|
||
). It differs from the new species in having divergent thorns arising from the centre of the disc granules, thorns at the tip of the arm spines and numerous lateral oral papillae and adoral shield spines.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B75349FF87FEF38640FC62FCF3" blockId="23.[151,1437,151,934]" box="[199,970,834,861]" pageId="23" pageNumber="24">
|
||
<materialsCitation id="FFEC3CEA5349FF87FEF38640FC6EFCF3" ID-GBIF-Occurrence="3746276319" box="[199,966,834,861]" collectionCode="MD" pageId="23" pageNumber="24" specimenCount="50">
|
||
The
|
||
<collectionCode id="2995AE725349FF87FECE8641FE8FFCF3" box="[250,295,834,861]" country="Germany" name="Museum Donaueschingen" pageId="23" pageNumber="24">MD</collectionCode>
|
||
<specimenCount id="5982FD3E5349FF87FF138641FE17FCF3" box="[295,447,834,861]" pageId="23" pageNumber="24" type="generic">50 specimens</specimenCount>
|
||
were found epizoic on a black coral (
|
||
<figureCitation id="D7BF2A325349FF87FD508640FC15FCF3" box="[868,957,835,861]" captionStart="FIGURE 7" captionStartId="21.[151,250,1920,1944]" captionTargetBox="[151,1435,181,1896]" captionTargetId="figure-17@21.[151,1435,181,1896]" captionTargetPageId="21" captionText="FIGURE 7. (A–B) Ophiosemnotes conferta, MNHN IE.2009.1574, A: dorsal and B: ventral views. (C–G) Ophiolebes felli sp. nov., holotype, MNHN IE.2019.4746, C: dorsal outline on black coral, D: dorsal, E: ventral view, F: proximal arm, G: arm tip. (H–I) paratype, MNHN IE.2009.1599, H: dorsal and I: ventral views." figureDoi="http://doi.org/10.5281/zenodo.6404694" httpUri="https://zenodo.org/record/6404694/files/figure.png" pageId="23" pageNumber="24">Fig. 7C</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="079E653C5349FF87FEF38664FAEDFC08" pageId="23" pageNumber="24" type="etymology">
|
||
<paragraph id="4F3B36B75349FF87FEF38664FAEDFC08" blockId="23.[151,1437,151,934]" pageId="23" pageNumber="24">
|
||
<emphasis id="7DF0EAA55349FF87FEF38664FEE0FC2F" bold="true" box="[199,328,871,897]" pageId="23" pageNumber="24">Etymology</emphasis>
|
||
. Named after H.B. Fell, a pioneer of echinoderm taxonomy and biogeography in
|
||
<collectingCountry id="379376275349FF87FAFE8664FACAFC2F" box="[1226,1378,871,897]" name="New Zealand" pageId="23" pageNumber="24">New Zealand</collectingCountry>
|
||
, who promoted the idea of long distance dispersal around southern oceans on the “West Wind Drift” (
|
||
<bibRefCitation id="2B154B465349FF87FAFC8688FA90FC08" author="Fell, H. B." box="[1224,1336,907,934]" pageId="23" pageNumber="24" pagination="759 - 761" refId="ref31060" refString="Fell, H. B. (1962) West-wind-drift dispersal of echinoderms in the southern hemisphere. Nature, 193, 759 - 761. https: // doi. org / 10.1038 / 193759 a 0" type="journal article" year="1962">Fell 1962</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |