treatments-xml/data/B8/19/F3/B819F3C6AFE7AB025CD7CF8E14DE8BD0.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.143.2075" ID-GBIF-Dataset="718cdf1f-ce26-44fd-a822-381cb8f5605b" ID-PMC="PMC3208533" ID-Pensoft-Pub="1313-2970-143-47" ID-PubMed="22144866" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-143-47" ModsDocOrigin="ZooKeys 143" ModsDocTitle="The genus Anthia Weber in the Republic of South Africa, Identification, distribution, biogeography, and behavior (Coleoptera, Carabidae)" checkinTime="1451249721601" checkinUser="pensoft" docAuthor="Mawdsley, Jonathan R., Erwin, Terry L., Sithole, Hendrik, Mawdsley, James L. &amp; Mawdsley, Alice S." docDate="2011" docId="B819F3C6AFE7AB025CD7CF8E14DE8BD0" docLanguage="en" docName="ZooKeys 143: 47-81" docOrigin="ZooKeys 143" docSource="http://dx.doi.org/10.3897/zookeys.143.2075" docTitle="Anthia circumscripta Klug 1853" docType="treatment" docVersion="3" lastPageNumber="59" masterDocId="6050B464FFE1D36FAE76CE32FF60532B" masterDocTitle="The genus Anthia Weber in the Republic of South Africa, Identification, distribution, biogeography, and behavior (Coleoptera, Carabidae)" masterLastPageNumber="81" masterPageNumber="47" pageNumber="58" updateTime="1668152558385" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The genus Anthia Weber in the Republic of South Africa, Identification, distribution, biogeography, and behavior (Coleoptera, Carabidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mawdsley, Jonathan R.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Erwin, Terry L.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sithole, Hendrik</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mawdsley, James L.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mawdsley, Alice S.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>143</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>47</mods:start>
<mods:end>81</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.143.2075</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.143.2075</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-143-47</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152031829" LSID="urn:lsid:plazi:treatment:B819F3C6AFE7AB025CD7CF8E14DE8BD0" httpUri="http://treatment.plazi.org/id/B819F3C6AFE7AB025CD7CF8E14DE8BD0" lastPageId="12" lastPageNumber="59" pageId="11" pageNumber="58">
<subSubSection pageId="11" pageNumber="58" type="nomenclature">
<paragraph pageId="11" pageNumber="58">
<taxonomicName LSID="http://species-id.net/wiki/Anthia_circumscripta" authority="Klug, 1853" authorityName="Klug" authorityYear="1853" class="Insecta" family="Carabidae" genus="Anthia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthia circumscripta" order="Coleoptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="circumscripta">Anthia circumscripta Klug, 1853</taxonomicName>
Figures 23-27, 313539
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="58" type="reference_group">
<paragraph pageId="11" pageNumber="58">
<taxonomicName class="Insecta" family="Carabidae" genus="Anthia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthia circumscripta" order="Coleoptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="circumscripta">Anthia circumscripta</taxonomicName>
Klug (1853:245).
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="58" type="type locality">
<paragraph pageId="11" pageNumber="58">Type Locality.</paragraph>
<paragraph pageId="11" pageNumber="58">
<normalizedToken originalValue="“Tette”">&quot;Tette&quot;</normalizedToken>
(= Tete, Mozambique).
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="58" type="type depository">
<paragraph pageId="11" pageNumber="58">Type Depository.</paragraph>
<paragraph pageId="11" pageNumber="58">
<taxonomicName class="Insecta" family="Carabidae" genus="Anthia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthia circumscripta" order="Coleoptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="circumscripta">Anthia circumscripta</taxonomicName>
, Museum
<normalizedToken originalValue="für">fuer</normalizedToken>
Naturkunde,
<normalizedToken originalValue="Humboldt-Universität">Humboldt-Universitaet</normalizedToken>
, Berlin.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="58" type="diagnosis">
<paragraph pageId="11" pageNumber="58">Diagnosis.</paragraph>
<paragraph pageId="11" pageNumber="58">
Unrubbed specimens have scattered areas of white setae on the lateral flanges of the pronotum, which is a diagnostic feature for this species. Rubbed specimens resemble
<taxonomicName class="Insecta" family="Carabidae" genus="Anthia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthia cinctipennis" order="Coleoptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="cinctipennis">Anthia cinctipennis</taxonomicName>
but males can be separated by the structure of the male genitalia which are slender in
<taxonomicName class="Insecta" family="Carabidae" genus="Anthia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthia cinctipennis" order="Coleoptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="cinctipennis">Anthia cinctipennis</taxonomicName>
and stouter and more robust in
<taxonomicName class="Insecta" family="Carabidae" genus="Anthia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthia circumscripta" order="Coleoptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="circumscripta">Anthia circumscripta</taxonomicName>
(Figures 30, 31). This species is known from relatively few specimens from RSA; however, it is represented in museum collections by large series from Botswana and Namibia, where it is evidently more frequently collected.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="58" type="description">
<paragraph pageId="11" pageNumber="58">Description.</paragraph>
<paragraph pageId="11" pageNumber="58">Body size massive, length of male 41.3-48.0 mm (exclusive of mandibles), length of female 42.6-50.4 mm.</paragraph>
<paragraph pageId="11" pageNumber="58">
Head elongate, prognathous. Mandibles larger in males than in females. Length of right mandible in male 7.4-8.6 mm. Palpi as in
<taxonomicName class="Insecta" family="Carabidae" genus="Anthia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthia maxillosa" order="Coleoptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="maxillosa">Anthia maxillosa</taxonomicName>
. Antennae, eyes, frons, and vertex as in
<taxonomicName class="Insecta" family="Carabidae" genus="Anthia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthia thoracica" order="Coleoptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="thoracica">Anthia thoracica</taxonomicName>
.
</paragraph>
<paragraph pageId="11" pageNumber="58">
Pronotum cordiform, shape as in
<taxonomicName class="Insecta" family="Carabidae" genus="Anthia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthia cinctipennis" order="Coleoptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="cinctipennis">Anthia cinctipennis</taxonomicName>
except that the basal flanges in male are often much smaller; apical margin of flanges transverse, with distinct emargination at apex of both flanges. Lateral flanges of pronotum with a more-or-less distinct patch of scattered short white reclinate setae. Pronotal surface sculpture smooth and shining, markedly punctate with large round punctures. Scutellum as in
<taxonomicName class="Insecta" family="Carabidae" genus="Anthia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthia thoracica" order="Coleoptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="thoracica">Anthia thoracica</taxonomicName>
. Elytra elliptical-oval, convex medially. Vestiture as in
<taxonomicName class="Insecta" family="Carabidae" genus="Anthia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthia thoracica" order="Coleoptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="thoracica">Anthia thoracica</taxonomicName>
. Each elytron with 8 distinct longitudinal striate interneurs, which may be feebly to somewhat markedly impressed; each interneur with a row of small round punctures, remainder of surface with scattered small round punctures. Legs as in
<taxonomicName class="Insecta" family="Carabidae" genus="Anthia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthia maxillosa" order="Coleoptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="maxillosa">Anthia maxillosa</taxonomicName>
, mesotibiae modified in both sexes as described for
<taxonomicName class="Insecta" family="Carabidae" genus="Anthia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthia cinctipennis" order="Coleoptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="cinctipennis">Anthia cinctipennis</taxonomicName>
.
</paragraph>
<paragraph pageId="11" pageNumber="58">
Abdomen as in
<taxonomicName class="Insecta" family="Carabidae" genus="Anthia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthia thoracica" order="Coleoptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="thoracica">Anthia thoracica</taxonomicName>
. Male aedeagus stout, robust (Figure 31).
</paragraph>
<caption pageId="11" pageNumber="58">
<paragraph pageId="11" pageNumber="58">
Figures 28-31. Dorsal view of abdominal apex, with aedeagus extended 28
<taxonomicName class="Insecta" family="Carabidae" genus="Anthia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthia thoracica" order="Coleoptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="thoracica">Anthia thoracica</taxonomicName>
(Thunberg) 29
<taxonomicName class="Insecta" family="Carabidae" genus="Anthia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthia maxillosa" order="Coleoptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="maxillosa">Anthia maxillosa</taxonomicName>
(Fabricius) 30
<taxonomicName class="Insecta" family="Carabidae" genus="Anthia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthia cinctipennis" order="Coleoptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="cinctipennis">Anthia cinctipennis</taxonomicName>
Lequien 31
<taxonomicName class="Insecta" family="Carabidae" genus="Anthia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthia circumscripta" order="Coleoptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="circumscripta">Anthia circumscripta</taxonomicName>
Klug.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="11" pageNumber="58" type="variation">
<paragraph pageId="11" pageNumber="58">Variation.</paragraph>
<paragraph pageId="11" pageNumber="58">Males exhibit considerable variation in the size and length of mandibles and in the size of the basal flange on the pronotum (Figures 23-25). Females also exhibit some variability in overall body size (Figures 26-27).</paragraph>
</subSubSection>
<subSubSection lastPageId="12" lastPageNumber="59" pageId="11" pageNumber="58" type="adult activity patterns">
<paragraph pageId="11" pageNumber="58">Adult activity patterns.</paragraph>
<paragraph lastPageId="12" lastPageNumber="59" pageId="11" pageNumber="58">
Distinctly bimodal (Figure 39), with populations from the Namib and Kalahari deserts having a July activity peak while populations from eastern Botswana, Zimbabwe and the few RSA records have an activity peak in
<pageBreakToken pageId="12" pageNumber="59" start="start">November-March</pageBreakToken>
. Figure 39 shows records from throughout southern Africa, since only a very few specimens from RSA were available for study.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="59" type="material examined">
<paragraph pageId="12" pageNumber="59">Material Examined.</paragraph>
<paragraph pageId="12" pageNumber="59">2 pinned adult specimens from the following localities: Republic of South Africa: Free State Province: Golden Gate. KwaZulu-Natal Province: Maritzburg. [Additional material was examined from Botswana, Kenya, Namibia, Tanzania, Zambia, and Zimbabwe].</paragraph>
</subSubSection>
</treatment>
</document>