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29 KiB
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304 lines
29 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e73829" ID-Pensoft-Pub="1864-8312-80-309" ID-Pensoft-UUID="11907D1B6F665D96B5FD2F9751431366" ID-ZooBank="D5C0468B99A14EF39237D9BC51A8BDA3" ModsDocID="1864-8312-80-309" checkinTime="1660711556436" checkinUser="pensoft" docAuthor="Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo" docDate="2022" docId="968A31B5C99D54DDB93BEA656D95B60E" docLanguage="en" docName="ArthropodSystPhyl 80: 309-388" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-08-16" docSource="http://dx.doi.org/10.3897/asp.80.e73829" docTitle="Ayacucho weyrauchi Benedetti & Pinto-da-Rocha 2022, comb. nov." docType="treatment" docVersion="1" id="11907D1B6F665D96B5FD2F9751431366" lastPageNumber="309" masterDocId="11907D1B6F665D96B5FD2F9751431366" masterDocTitle="Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species" masterLastPageNumber="388" masterPageNumber="309" pageNumber="309" updateTime="1660711556436" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Benedetti, Alipio Rezende</mods:namePart>
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<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil & Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">alipiobenedetti@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pinto-da-Rocha, Ricardo</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3959-2205</mods:nameIdentifier>
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<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-16</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>80</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>309</mods:start>
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<mods:end>388</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/asp.80.e73829</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e73829</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1864-8312-80-309</mods:identifier>
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<mods:identifier type="ZooBank">D5C0468B99A14EF39237D9BC51A8BDA3</mods:identifier>
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<mods:identifier type="Pensoft-UUID">11907D1B6F665D96B5FD2F9751431366</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:968A31B5C99D54DDB93BEA656D95B60E" httpUri="http://treatment.plazi.org/id/968A31B5C99D54DDB93BEA656D95B60E" lastPageNumber="309" pageId="0" pageNumber="309">
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<subSubSection pageId="0" pageNumber="309" type="nomenclature">
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<paragraph pageId="0" pageNumber="309">
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3.27.
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<taxonomicName LSID="968A31B5-C99D-54DD-B93B-EA656D95B60E" authority="(Roewer, 1952)" authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" baseAuthorityName="Roewer" baseAuthorityYear="1952" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho weyrauchi" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="weyrauchi" status="comb. nov.">Ayacucho weyrauchi (Roewer, 1952)</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="309">comb. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="description">
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<paragraph pageId="0" pageNumber="309">
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">Figs 3F</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male trochanter-tibia IV of leg IV of Ayacucho, dorsal and ventral view, respectively. A - B A. uniseriatus (Roewer, 1959) comb. nov.; C - D A. vargasllosai sp. nov.; E - F A. weyrauchi (Roewer, 1952) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure8" httpUri="https://binary.pensoft.net/fig/729882" pageId="0" pageNumber="309">, 8E, F</figureCitation>
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B A. vargasllosai sp. nov.; C - D A. weyrauchi (Roewer, 1952) comb. nov.; E - F Incasarcus argenteus Kury & Maury, 1998; G - H Incasarcus dianae Kury & Maury, 1998; I - J: Incasarcus pictus Kury & Maury, 1998; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure13" httpUri="https://binary.pensoft.net/fig/729887" pageId="0" pageNumber="309">, 13C, D</figureCitation>
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of Ayacucho spp. in Peru" figureDoi="10.3897/asp.80.e73829.figure28" httpUri="https://binary.pensoft.net/fig/729902" pageId="0" pageNumber="309">, 28</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="reference_group">
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<paragraph pageId="0" pageNumber="309">
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<taxonomicName authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" baseAuthorityName="Benedetti & Pinto-da-Rocha" baseAuthorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Cajamarca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cajamarca weyrauchi" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="weyrauchi" status="comb. nov.">Cajamarca weyrauchi</taxonomicName>
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Roewer, 1952: 41 (desc.);
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<bibRefCitation author="Roewer, CF" journalOrPublisher="Senckenbergiana Biologica" pageId="0" pageNumber="309" pagination="67 - 94" refId="B108" refString="Roewer, CF, 1957. Arachnida Arthrogastra aus Peru, III. Senckenbergiana Biologica 38 (1-2): 67 - 94" title="Arachnida Arthrogastra aus Peru, III." volume="38" year="1957">Roewer 1957</bibRefCitation>
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: 75 (cit.), fig. 30 (male femur IV);
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<bibRefCitation author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" refId="B51" refString="Kury, AB, 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol.7: 1-337." title="Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. 7: 1 - 337." year="2003">Kury 2003</bibRefCitation>
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: 144 (cat.).
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</paragraph>
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<paragraph pageId="0" pageNumber="309">
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<taxonomicName authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" baseAuthorityName="Benedetti & Pinto-da-Rocha" baseAuthorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Cajamarca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cajamarca affinis" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="affinis" status="comb. nov.">Cajamarca affinis</taxonomicName>
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Roewer, 1957: 75 (desc.), fig. 31 (male femur IV);
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<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1998.tb01297.x" author="Kury, AB" journalOrPublisher="Zoo-logical Journal of Linnean Society" pageId="0" pageNumber="309" pagination="143 - 162" refId="B58" refString="Kury, AB, Maury, EA, 1998. A new genus and five new species of Metasarcinae from Peru (Arachnida, Opiliones, Gonyleptidae). Zoo-logical Journal of Linnean Society 123: 143 - 162, DOI: https://doi.org/10.1111/j.1096-3642.1998.tb01297.x" title="A new genus and five new species of Metasarcinae from Peru (Arachnida, Opiliones, Gonyleptidae)." url="https://doi.org/10.1111/j.1096-3642.1998.tb01297.x" volume="123" year="1998">Kury and Maury 1998</bibRefCitation>
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: 145 (cit.);
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<bibRefCitation author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" refId="B51" refString="Kury, AB, 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol.7: 1-337." title="Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. 7: 1 - 337." year="2003">Kury 2003</bibRefCitation>
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: 144 (cat.),
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<emphasis bold="true" pageId="0" pageNumber="309">syn. n.</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="redescription">
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<paragraph pageId="0" pageNumber="309">Redescription.</paragraph>
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<paragraph pageId="0" pageNumber="309">
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<emphasis bold="true" pageId="0" pageNumber="309">
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MALE:
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<emphasis italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
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</emphasis>
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(
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<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
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=2) DSW: 4.5; DSL: 4.6-5.0 (5.0); CL: 1.5-1.7 (1.5). FIVL: 4.2-4.5 (4.5). ChL: 1.5-3.0 (3.0).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Coloration</emphasis>
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(in ethanol): Predominantly yellowish. Areas I-IV, posterior margin of DS and free tergites I-IV brown.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Dorsum</emphasis>
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: (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3F</figureCitation>
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) Alpha-type DSS, with wide coda. Anterior margin of DS with median elevation with few sparsely granules. Ocularium with a pair of short spines; with granules between the spines. Carapace with few granules in the lateral regions of the ocularium, predominantly smooth in the posterior region of the ocularium. Areas I-IV few granular. Lateral margins of DS with few sparsely granules. Posterior margin of DS with 10-11 tiny tubercles. Free tergites I-III each one with a row of tiny tubercles with a wider base than their height.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Chelicerae</emphasis>
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: (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3F</figureCitation>
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) Swollen in the largest specimen (the holotype of
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<taxonomicName authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" baseAuthorityName="Benedetti & Pinto-da-Rocha" baseAuthorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Cajamarca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cajamarca weyrauchi" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="weyrauchi">
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<emphasis italics="true" pageId="0" pageNumber="309">Cajamarca weyrauchi</emphasis>
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</taxonomicName>
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); similar to the females in the other male. Segment I granular. Segment II predominantly smooth; finger with one tooth. Segment III with five teeth.
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<taxonomicName genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Pedipalpus</emphasis>
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</taxonomicName>
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: Small granules distributed on the dorsal surface of the femur and patella. Trochanter with a large ventrodistal setiferous tubercle. Femur with a ventrobasal setiferous tubercle; a ventral row of four or five setiferous tubercles, larger in larger males. Tibia: prolateral iiIi, retrolateral iIi. Tarsus: prolateral IIi, retrolateral Iii.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Venter</emphasis>
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: Coxa I with a median row of six small tubercles. Coxae II-IV smooth. Three tiny apical tubercles between the coxae II-III and III-IV. Genital area smooth. Free sternites with a row of granules. Anal operculum with granules sparsely distributed over its length.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Legs</emphasis>
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: (Figs
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3F</figureCitation>
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,
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male trochanter-tibia IV of leg IV of Ayacucho, dorsal and ventral view, respectively. A - B A. uniseriatus (Roewer, 1959) comb. nov.; C - D A. vargasllosai sp. nov.; E - F A. weyrauchi (Roewer, 1952) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure8" httpUri="https://binary.pensoft.net/fig/729882" pageId="0" pageNumber="309">8E, F</figureCitation>
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) Coxae I-III each one with a prolateral and a retrolateral apophysis (largest on coxa II); retrolateral apophysis on coxa II and prolateral apophysis on coxa III fused. Coxa IV with few granules, concentrated more on the distal part, with a prodistal spiniform apophysis. Trochanters I-III smooth. Trochanter IV with 3-4 median dorsal granules and a retroapical acuminate tubercle. Femora I-III with sparse granules throughout their length. Femur IV granular; with a median dorsal row of 5-8 large acuminate tubercles, growing apically in size; a retrolateral row with 3-4 large acuminate tubercles at distal half of this article; a retroventral row of 9-13 small tubercles, some of them similar to granules; a proventral row of 6-7 large acuminate tubercles along the apical portion. Patella I-IV with sparse granulation. Tibiae I-IV granular. Tarsal formula: (
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<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
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=2) 7, 11-12 (12), 6-7 (6), 8 (?).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Penis</emphasis>
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: (Fig.
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B A. vargasllosai sp. nov.; C - D A. weyrauchi (Roewer, 1952) comb. nov.; E - F Incasarcus argenteus Kury & Maury, 1998; G - H Incasarcus dianae Kury & Maury, 1998; I - J: Incasarcus pictus Kury & Maury, 1998; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure13" httpUri="https://binary.pensoft.net/fig/729887" pageId="0" pageNumber="309">13C, D</figureCitation>
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) VP with greater width in the median region; distal margin concave; straight in lateral view (except the apical portion). MS C1-C3 subapical long and curved or straight; MS A1-A2 median long and straight. Lateral sacs long and apically acuminated, with long T3-like microsetae. Stylus sinuous, with inflated apex, a long ventral projection and several small apical projections. Dorsal process half the length of the stylus. Promontory convex. -
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<emphasis bold="true" pageId="0" pageNumber="309">
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FEMALE:
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<emphasis italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
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</emphasis>
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(
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<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
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=7) DSW: 3.8-4.1; DSL: 4.2-4.5; CL: 1.1-1.7. FIVL: 3.4-3.6. ChL: 1.3-1.6. Chelicerae similar to that of small males.
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<taxonomicName genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">Pedipalpus</taxonomicName>
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femur with a proapical spine. Femur IV with a dorsal row of 3-4 acuminate tubercles, spaced apart from each other; a retrolateral row of granules, with two acuminate tubercles in the median portion; a proventral row of 15-17 small tubercles, larger in the apical portion. All tubercles present in females are smaller than those in males. Tarsal formula: (
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<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
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=7) 7, 11-13, 5-7, 8.
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</paragraph>
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<caption doi="10.3897/asp.80.e73829.figure20" httpUri="https://binary.pensoft.net/fig/729894" pageId="0" pageNumber="309" start="Figure 20" startId="F20">
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<paragraph pageId="0" pageNumber="309">
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<emphasis bold="true" pageId="0" pageNumber="309">Figure 20.</emphasis>
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Penis of
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<taxonomicName authorityName="Roewer" authorityYear="1913" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="309">Metasarcus</emphasis>
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</taxonomicName>
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, dorsal, lateral and ventral views, respectively.
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<emphasis bold="true" pageId="0" pageNumber="309">A</emphasis>
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-
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<emphasis bold="true" pageId="0" pageNumber="309">C</emphasis>
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<taxonomicName class="Arachnida" genus="M." kingdom="Animalia" lsidName="M. trispinosus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="trispinosus">
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<emphasis italics="true" pageId="0" pageNumber="309">M. trispinosus</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
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;
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<emphasis bold="true" pageId="0" pageNumber="309">D</emphasis>
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-
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<emphasis bold="true" pageId="0" pageNumber="309">F</emphasis>
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<taxonomicName class="Arachnida" genus="M." kingdom="Animalia" lsidName="M. vacafloresae" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="vacafloresae">
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<emphasis italics="true" pageId="0" pageNumber="309">M. vacafloresae</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
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; Legend bars = 0.1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="diagnosis">
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<paragraph pageId="0" pageNumber="309">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="309">
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Similar to
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<taxonomicName authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" baseAuthorityName="Roewer" baseAuthorityYear="1957" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho bambamarca" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="bambamarca">
|
||
<emphasis italics="true" pageId="0" pageNumber="309">Ayacucho bambamarca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
|
||
and
|
||
<taxonomicName genus="A." lsidName="A. triarmatus" pageId="0" pageNumber="309" rank="species" species="triarmatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="309">A. triarmatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="309">nom. nov.</emphasis>
|
||
for presenting three rows of acuminate tubercles in the male femur IV (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male trochanter-tibia IV of leg IV of Ayacucho, dorsal and ventral view, respectively. A - B A. uniseriatus (Roewer, 1959) comb. nov.; C - D A. vargasllosai sp. nov.; E - F A. weyrauchi (Roewer, 1952) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure8" httpUri="https://binary.pensoft.net/fig/729882" pageId="0" pageNumber="309">8E, F</figureCitation>
|
||
). It differs from
|
||
<taxonomicName genus="A." lsidName="A. bambamarca" pageId="0" pageNumber="309" rank="species" species="bambamarca">
|
||
<emphasis italics="true" pageId="0" pageNumber="309">A. bambamarca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
|
||
and
|
||
<taxonomicName genus="A." lsidName="A. triarmatus" pageId="0" pageNumber="309" rank="species" species="triarmatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="309">A. triarmatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="309">nom. nov.</emphasis>
|
||
in having a maximum of four acuminate tubercles in the retrolateral row (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male trochanter-tibia IV of leg IV of Ayacucho, dorsal and ventral view, respectively. A - B A. uniseriatus (Roewer, 1959) comb. nov.; C - D A. vargasllosai sp. nov.; E - F A. weyrauchi (Roewer, 1952) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure8" httpUri="https://binary.pensoft.net/fig/729882" pageId="0" pageNumber="309">8E, F</figureCitation>
|
||
); smaller tubercles in ocularium; free tergites with small tubercles (with wide base); areas I-IV with few granulation (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3F</figureCitation>
|
||
). It differs from
|
||
<taxonomicName genus="A." lsidName="A. bambamarca" pageId="0" pageNumber="309" rank="species" species="bambamarca">
|
||
<emphasis italics="true" pageId="0" pageNumber="309">A. bambamarca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
|
||
in having scutal areas unarmed (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="309" type="distribution">
|
||
<paragraph pageId="0" pageNumber="309">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="309">
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of Ayacucho spp. in Peru" figureDoi="10.3897/asp.80.e73829.figure28" httpUri="https://binary.pensoft.net/fig/729902" pageId="0" pageNumber="309">28</figureCitation>
|
||
) PERU.
|
||
<taxonomicName authorityName="Roewer" authorityYear="1952" class="Arachnida" family="Metasarcidae" genus="Cajamarca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cajamarca" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Cajamarca</taxonomicName>
|
||
.
|
||
<taxonomicName authorityName="Roewer" authorityYear="1952" class="Arachnida" family="Metasarcidae" genus="Cajamarca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cajamarca" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Cajamarca</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="309" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="309">Material examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="309">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Type material</emphasis>
|
||
:
|
||
<materialsCitation accessionNumber="RII 10128, RII 11646" collectingDate="1956-08-04" collectorName="Weyrauch" country="PERU" county="Celendin" elevation="2750" location="Of" municipality="Of" specimenCount="7" specimenCount-female="7" typeStatus="Holotype">
|
||
<location LSID="urn:lsid:plazi:treatment:968A31B5C99D54DDB93BEA656D95B60E:285180A97BB16EF78CACA360CD1B370A" country="PERU" county="Celendin" municipality="Of" name="Of">Of</location>
|
||
<taxonomicName genus="C." lsidName="C. weyrauchi" pageId="0" pageNumber="309" rank="species" species="weyrauchi">
|
||
<emphasis italics="true" pageId="0" pageNumber="309">C. weyrauchi</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis bold="true" pageId="0" pageNumber="309">
|
||
<typeStatus>Holotype</typeStatus>
|
||
</emphasis>
|
||
<specimenCount>♂</specimenCount>
|
||
, '
|
||
<collectingCountry name="Peru">PERU</collectingCountry>
|
||
,
|
||
<taxonomicName authorityName="Roewer" authorityYear="1952" class="Arachnida" family="Metasarcidae" genus="Cajamarca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cajamarca" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Cajamarca</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Roewer" authorityYear="1952" class="Arachnida" family="Metasarcidae" genus="Cajamarca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cajamarca" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Cajamarca</taxonomicName>
|
||
,|
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.75" unit="m" value="2750.0">
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.75" unit="m" value="2750.0">2,750 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
, without date,
|
||
<collectorName>Weyrauch</collectorName>
|
||
leg. (SMF
|
||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/RII10128">RII 10128</accessionNumber>
|
||
/22) -
|
||
<emphasis bold="true" pageId="0" pageNumber="309">
|
||
<typeStatus>Paratypes</typeStatus>
|
||
</emphasis>
|
||
<specimenCount type="female">2 ♀</specimenCount>
|
||
,
|
||
<normalizedToken originalValue="‘ditto’">'ditto'</normalizedToken>
|
||
(SMF
|
||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/RII10128">RII 10128</accessionNumber>
|
||
/22).
|
||
<collectingMunicipality>Of</collectingMunicipality>
|
||
<taxonomicName genus="C." lsidName="C. affinis" pageId="0" pageNumber="309" rank="species" species="affinis">
|
||
<emphasis italics="true" pageId="0" pageNumber="309">C. affinis</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis bold="true" pageId="0" pageNumber="309">
|
||
<typeStatus>Holotype</typeStatus>
|
||
</emphasis>
|
||
<specimenCount>♂</specimenCount>
|
||
, '
|
||
<collectingCountry name="Peru">PERU</collectingCountry>
|
||
,
|
||
<taxonomicName authorityName="Roewer" authorityYear="1952" class="Arachnida" family="Metasarcidae" genus="Cajamarca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cajamarca" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Cajamarca</taxonomicName>
|
||
, near
|
||
<taxonomicName authorityName="Roewer" authorityYear="1952" class="Arachnida" family="Metasarcidae" genus="Cajamarca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cajamarca" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Cajamarca</taxonomicName>
|
||
,
|
||
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.4" unit="km" value="24.0">24 km</quantity>
|
||
from
|
||
<taxonomicName authorityName="Roewer" authorityYear="1952" class="Arachnida" family="Metasarcidae" genus="Cajamarca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cajamarca" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Cajamarca</taxonomicName>
|
||
, road to
|
||
<collectingCounty>Celendin</collectingCounty>
|
||
,
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="3.15" unit="m" value="3150.0">
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.15" unit="m" value="3150.0">3,150 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<collectingDate value="1956-08-04">04/VIII/1956</collectingDate>
|
||
,
|
||
<collectorName>Weyrauch</collectorName>
|
||
leg. (SMF
|
||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/RII11646">RII 11646</accessionNumber>
|
||
/29) -
|
||
<emphasis bold="true" pageId="0" pageNumber="309">
|
||
<typeStatus>Paratypes</typeStatus>
|
||
</emphasis>
|
||
<specimenCount type="female">5 ♀</specimenCount>
|
||
,
|
||
<normalizedToken originalValue="‘ditto’">'ditto'</normalizedToken>
|
||
(SMF
|
||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/RII11646">RII 11646</accessionNumber>
|
||
/29)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |