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<document id="78C9519FD8885E51EE68EEB6CA16587C" ID-CLB-Dataset="5789" ID-DOI="10.11646/zootaxa.3783.1.1" ID-GBIF-Dataset="414491b2-5167-4605-95ab-f679bf476618" ID-ISSN="1175-5326" ID-Zenodo-Dep="4910562" ID-ZooBank="E6155BDC-AEAE-475D-BC83-61B3B863344C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623122599895" checkinUser="felipe" docAuthor="Karanovic, Tomislav &amp; Kim, Kichoon" docDate="2014" docId="6878D460FFE7FF8064D0FE0901D2FACB" docLanguage="en" docName="zt03783p096.pdf" docOrigin="Zootaxa 3783 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Willenstenhelia thalia Karanovic &amp; Kim 2014, sp. nov." docType="treatment" docVersion="7" lastPageNumber="84" masterDocId="9441AC18FFAEFFD36447FFF6027BFFBC" masterDocTitle="New insights into polyphyly of the harpacticoid genus Delavalia (Crustacea, Copepoda) through morphological and molecular study of an unprecedented diversity of sympatric species in a small South Korean bay" masterLastPageNumber="96" masterPageNumber="1" pageNumber="74" updateTime="1699004038958" updateUser="ExternalLinkService">
<mods:mods id="47DAAC57F9280A7EAA105D0AD1B4E8CC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5716BADAF736DC010A108106BDC60045">
<mods:title id="3173B11FB3B14DDF9256EA3BB207A77B">New insights into polyphyly of the harpacticoid genus Delavalia (Crustacea, Copepoda) through morphological and molecular study of an unprecedented diversity of sympatric species in a small South Korean bay</mods:title>
</mods:titleInfo>
<mods:name id="EB7F261C8DFDBA99FE14A8F7751B3CDF" type="personal">
<mods:role id="FC4F0B05B49C7D91FF0AB9FCF5E2E209">
<mods:roleTerm id="78F641AE38259E6D0D1DD8C93788FDA2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0CD1BF91CCCD84C4BB20BC26BC57A35E">Karanovic, Tomislav</mods:namePart>
<mods:nameIdentifier id="48349933982BEAB2F8EFD2CDCC77AFB5" type="email">aranovic@utas.edu.au</mods:nameIdentifier>
</mods:name>
<mods:name id="C63647F7C1FDC512C654B050A325A94D" type="personal">
<mods:role id="39B4FA93C01820821D567CA3A2160649">
<mods:roleTerm id="A8CAC6B07ED8027D45E5E2E427F247EF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FD58208098A6361658012E508C407B52">Kim, Kichoon</mods:namePart>
</mods:name>
<mods:typeOfResource id="984A1332C8728C336CD27E7F101811FD">text</mods:typeOfResource>
<mods:relatedItem id="C4C653A8A43D7F4EE2543E404876B1C4" type="host">
<mods:titleInfo id="BF756CD2760A5B14208352CEA40E8F32">
<mods:title id="8579F386BAC90078822FCB2733ED2252">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C5DDE5C9682A1EFD5EE831BBCB5A89AB">
<mods:date id="1F8417D79B59445EB6A404C8E0AEBBBA">2014</mods:date>
<mods:detail id="4CE2D75CBC1173D4AB0E440B5623BB50" type="pubDate">
<mods:number id="FE0BBBB570980E95C5A874A376CF5F87">2014-03-25</mods:number>
</mods:detail>
<mods:detail id="E8E3F025F879B75E0525F656014BE441" type="volume">
<mods:number id="06702354A5208314CFB107800EB96464">3783</mods:number>
</mods:detail>
<mods:detail id="0BA7AD8830AC38D632FC5F6D02279344" type="issue">
<mods:number id="EB306D67A4939C0A4E4ED8E5718A86EB">1</mods:number>
</mods:detail>
<mods:extent id="94F651425AF0BB425DEF411E26C62939" unit="page">
<mods:start id="377E8122669506FAA2AD016CEC83F1FD">1</mods:start>
<mods:end id="FE68A425574483974B4C0459D7C9E091">96</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A885C9E7E093C220991EE8FE24914797">journal article</mods:classification>
<mods:identifier id="576616F363756B9BB42951C488D9E1CC" type="CLB-Dataset">5789</mods:identifier>
<mods:identifier id="A9657192BB601310067EDDF216E4D334" type="DOI">10.11646/zootaxa.3783.1.1</mods:identifier>
<mods:identifier id="8663125CF985B28981B127A564A7ADF5" type="GBIF-Dataset">414491b2-5167-4605-95ab-f679bf476618</mods:identifier>
<mods:identifier id="06E70A6DE9A5AFA3E49EF212B39E4EAD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="844287255562847EF2E08E3C41826826" type="Zenodo-Dep">4910562</mods:identifier>
<mods:identifier id="2FDFD9C7FC26322192A15E7DBF96E29E" type="ZooBank">E6155BDC-AEAE-475D-BC83-61B3B863344C</mods:identifier>
</mods:mods>
<treatment id="6878D460FFE7FF8064D0FE0901D2FACB" ID-DOI="http://doi.org/10.5281/zenodo.5062501" ID-GBIF-Taxon="183238039" ID-Zenodo-Dep="5062501" LSID="urn:lsid:plazi:treatment:6878D460FFE7FF8064D0FE0901D2FACB" httpUri="http://treatment.plazi.org/id/6878D460FFE7FF8064D0FE0901D2FACB" lastPageId="83" lastPageNumber="84" pageId="73" pageNumber="74">
<subSubSection id="A8CB36FDFFE7FF9A64D0FE09007EFDA6" box="[151,517,511,538]" pageId="73" pageNumber="74" type="nomenclature">
<paragraph id="E06E6576FFE7FF9A64D0FE09007EFDA6" blockId="73.[151,517,511,571]" box="[151,517,511,538]" pageId="73" pageNumber="74">
<heading id="BB26D21AFFE7FF9A64D0FE09007EFDA6" bold="true" box="[151,517,511,538]" fontSize="11" level="2" pageId="73" pageNumber="74" reason="4">
<emphasis id="D2A5B964FFE7FF9A64D0FE09007EFDA6" bold="true" box="[151,517,511,538]" pageId="73" pageNumber="74">
<taxonomicName id="27D11EF5FFE7FF9A64D0FE0903DBFDA5" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[151,416,511,537]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="thalia" status="sp. nov.">
<emphasis id="D2A5B964FFE7FF9A64D0FE0903DBFDA5" bold="true" box="[151,416,511,537]" italics="true" pageId="73" pageNumber="74">Willenstenhelia thalia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C996041FFFE7FF9A65E0FDF6007EFDA6" box="[423,517,512,538]" pageId="73" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A8CB36FDFFE7FF9A64D0FDD40355FD87" box="[151,302,546,571]" pageId="73" pageNumber="74" type="description">
<paragraph id="E06E6576FFE7FF9A64D0FDD40355FD87" blockId="73.[151,517,511,571]" box="[151,302,546,571]" pageId="73" pageNumber="74">
(
<figureCitation id="78EA79F3FFE7FF9A64D8FDD4035DFD87" box="[159,294,546,571]" captionStart-0="FIGURE 48" captionStart-1="FIGURE 49" captionStart-2="FIGURE 50" captionStart-3="FIGURE 51" captionStart-4="FIGURE 52" captionStart-5="FIGURE 53" captionStart-6="FIGURE 54" captionStartId-0="75.[151,249,1810,1832]" captionStartId-1="76.[151,249,1854,1876]" captionStartId-2="77.[151,249,1918,1940]" captionStartId-3="78.[151,249,1916,1938]" captionStartId-4="79.[151,249,1858,1880]" captionStartId-5="80.[151,249,1559,1581]" captionStartId-6="81.[151,249,1560,1582]" captionTargetBox-0="[239,1370,199,1785]" captionTargetBox-1="[216,1372,198,1825]" captionTargetBox-2="[189,1401,185,1886]" captionTargetBox-3="[191,1394,189,1884]" captionTargetBox-4="[208,1370,205,1829]" captionTargetBox-5="[151,1436,193,1538]" captionTargetBox-6="[151,1436,193,1539]" captionTargetId-0="figure-129@75.[228,1370,189,1791]" captionTargetId-1="figure-124@76.[210,1374,193,1832]" captionTargetId-2="figure-82@77.[177,1405,173,1889]" captionTargetId-3="figure-90@78.[184,1401,179,1891]" captionTargetId-4="figure-120@79.[208,1374,193,1836]" captionTargetId-5="figure-204@80.[151,1436,193,1538]" captionTargetId-6="figure-216@81.[151,1436,193,1539]" captionTargetPageId-0="75" captionTargetPageId-1="76" captionTargetPageId-2="77" captionTargetPageId-3="78" captionTargetPageId-4="79" captionTargetPageId-5="80" captionTargetPageId-6="81" captionText-0="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." captionText-1="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." captionText-2="FIGURE 50. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, mandibula, posterior view; B, mandibula, antero-ventral view; C, maxilla, posterior view; D, first swimming leg, anterior view; E, endopod of second swimming leg, anterior view; F, third exopodal segment of second swimming leg, anterior view; G, third swimming leg, anterior view. Arrowheads point to most prominent specific features. Scale bars 100 µm." captionText-3="FIGURE 51. Willenstenhelia thalia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, fourth swimming leg, anterior view; B, fifth swimming leg, anterior view; C, fifth swimming legs, anterior view; D, rostrum and antennula, dorsal view. Arabic numeral indicates sensillum presumably homologous to that in Itostenhelia polyhymnia. Greek letter (α) indicates a unique pore. Arrowheads point to most prominent specific features. Scale bars 100 µm." captionText-4="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." captionText-5="FIGURE 53. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–D, female paratype 1, E–F, female paratype 2: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, genital double-somite, dorsal view; D, preanal and anal somites and caudal rami, dorsal view; E, mouth appendages, ventral view; F, genital double-somite and fifth legs, ventral view. Scale bars 20 µm (E), 30 µm (D), 40 µm (C), 50 µm (B, F), and 100 µm (A)." captionText-6="FIGURE 54. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–B, female paratype 3, C, female paratype 4, D–F, male paratype: A, cephalothorax, dorsal view; B, anal somite and caudal rami, posterior view; C, cephalothorax, lateral view; D, antennula, lateral view; E, genital somite and fifth leg, lateral view; F, posterior part of caudal rami, lateral view. Scale bars 10 µm (F), 30 µm (D, E), and 50 µm (A, B, C)." figureDoi-0="http://doi.org/10.5281/zenodo.4910666" figureDoi-1="http://doi.org/10.5281/zenodo.4910668" figureDoi-2="http://doi.org/10.5281/zenodo.4910670" figureDoi-3="http://doi.org/10.5281/zenodo.4910672" figureDoi-4="http://doi.org/10.5281/zenodo.4910674" figureDoi-5="http://doi.org/10.5281/zenodo.4910676" figureDoi-6="http://doi.org/10.5281/zenodo.4910678" httpUri-0="https://zenodo.org/record/4910666/files/figure.png" httpUri-1="https://zenodo.org/record/4910668/files/figure.png" httpUri-2="https://zenodo.org/record/4910670/files/figure.png" httpUri-3="https://zenodo.org/record/4910672/files/figure.png" httpUri-4="https://zenodo.org/record/4910674/files/figure.png" httpUri-5="https://zenodo.org/record/4910676/files/figure.png" httpUri-6="https://zenodo.org/record/4910678/files/figure.png" pageId="73" pageNumber="74">Figs. 48–54</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A8CB36FDFFE7FF9A64D0FD9C07E7FAB7" pageId="73" pageNumber="74" type="materials_examined">
<paragraph id="E06E6576FFE7FF9A64D0FD9C033BFD1B" blockId="73.[151,1436,618,2011]" pageId="73" pageNumber="74">
<materialsCitation id="50B96F2BFFE7FF9A64D0FD9C0347FD1B" ID-GBIF-Occurrence="3313373314" country="South Korea" latitude="37.53923" location="South Sea" longLatPrecision="1" longitude="126.343414" municipality="Gwangyang Bay" pageId="73" pageNumber="74" specimenCount="1" typeStatus="holotype">
<emphasis id="D2A5B964FFE7FF9A64D0FD9C0342FD3F" bold="true" box="[151,313,618,643]" pageId="73" pageNumber="74">
<typeStatus id="3F6ADBD4FFE7FF9A64D0FD9C02A9FD3F" box="[151,210,618,643]" pageId="73" pageNumber="74">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="98C625E6FFE7FF9A6504FD9C03AEFD3E" box="[323,469,618,643]" name="South Korea" pageId="73" pageNumber="74">South Korea</collectingCountry>
,
<location id="E50E33ADFFE7FF9A65A3FD9C0020FD3F" LSID="urn:lsid:plazi:treatment:6878D460FFE7FF8064D0FE0901D2FACB:E50E33ADFFE7FF9A65A3FD9C0020FD3F" box="[484,603,618,643]" country="South Korea" latitude="37.53923" longLatPrecision="1" longitude="126.343414" municipality="Gwangyang Bay" name="South Sea" pageId="73" pageNumber="74">South Sea</location>
,
<collectingMunicipality id="000AFF0CFFE7FF9A662DFD9C0152FD3F" box="[618,809,618,643]" pageId="73" pageNumber="74">Gwangyang Bay</collectingMunicipality>
, sampling station 10, muddy sediments,
<geoCoordinate id="85E503B1FFE7FF9A6142FD9C07E7FD3F" box="[1285,1436,618,643]" degrees="37.539231" direction="north" orientation="latitude" pageId="73" pageNumber="74" precision="1" value="37.53923">37.539231°N</geoCoordinate>
<geoCoordinate id="85E503B1FFE7FF9A64D0FD780347FD1B" box="[151,316,654,679]" degrees="126.343417" direction="east" orientation="longitude" pageId="73" pageNumber="74" precision="1" value="126.343414">126.343417°E</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="E06E6576FFE7FF9A6480FD440012FCE7" blockId="73.[151,1436,618,2011]" pageId="73" pageNumber="74">
<emphasis id="D2A5B964FFE7FF9A6480FD4403BDFD77" bold="true" box="[199,454,690,715]" pageId="73" pageNumber="74">Specimens examined.</emphasis>
Female
<typeStatus id="3F6ADBD4FFE7FF9A6662FD4400FCFD77" box="[549,647,690,715]" pageId="73" pageNumber="74" type="holotype">holotype</typeStatus>
dissected on one slide (collection number NIBRIV0000232706), male
<typeStatus id="3F6ADBD4FFE7FF9A64D0FD20028AFD53" box="[151,241,726,751]" pageId="73" pageNumber="74" type="allotype">allotype</typeStatus>
dissected on one slide (collection number NIBRIV0000232707),
<specimenCount id="F6D7AEFFFFE7FF9A67B2FD21061BFD53" box="[1013,1120,726,751]" pageId="73" pageNumber="74" type="male">two male</specimenCount>
<typeStatus id="3F6ADBD4FFE7FF9A602BFD2106A3FD53" box="[1132,1240,727,751]" pageId="73" pageNumber="74" type="paratype">paratypes</typeStatus>
and
<specimenCount id="F6D7AEFFFFE7FF9A6151FD2007E7FD53" box="[1302,1436,726,751]" pageId="73" pageNumber="74" type="female">four female</specimenCount>
<typeStatus id="3F6ADBD4FFE7FF9A64D0FD0D0379FCAF" box="[151,258,763,787]" pageId="73" pageNumber="74" type="paratype">paratypes</typeStatus>
together on one SEM stub (collection number NIBRIV0000232708),
<specimenCount id="F6D7AEFFFFE7FF9A6004FD0C06B2FCAF" box="[1091,1225,762,787]" pageId="73" pageNumber="74" type="male">three males</specimenCount>
and
<specimenCount id="F6D7AEFFFFE7FF9A614DFD0C07E7FCAF" box="[1290,1436,762,787]" pageId="73" pageNumber="74" type="female">four females</specimenCount>
destroyed for DNA sequence (
<specimenCount id="F6D7AEFFFFE7FF9A6647FCE800EFFC8B" box="[512,660,798,823]" pageId="73" pageNumber="74" type="female">four females</specimenCount>
amplification successful, Codes 0245, 0444, 0146 &amp; 0348), type locality,
<date id="946F43B6FFE7FF9A64B0FCB403A9FCE7" box="[247,466,834,859]" pageId="73" pageNumber="74" value="2012-11-18">18 November 2012</date>
, leg. K. Kim.
</paragraph>
<paragraph id="E06E6576FFE7FF9A6480FC9103E5FC7B" blockId="73.[151,1436,618,2011]" pageId="73" pageNumber="74">
Two male
<typeStatus id="3F6ADBD4FFE7FF9A657AFC9103D3FCC3" box="[317,424,871,895]" pageId="73" pageNumber="74" type="paratype">paratypes</typeStatus>
and
<specimenCount id="F6D7AEFFFFE7FF9A6599FC91000AFCC3" box="[478,625,870,895]" pageId="73" pageNumber="74" type="female">seven female</specimenCount>
<typeStatus id="3F6ADBD4FFE7FF9A663FFC91009FFCC3" box="[632,740,871,895]" pageId="73" pageNumber="74" type="paratype">paratypes</typeStatus>
together in ethanol (collection number NIBRIV0000232709),
<specimenCount id="F6D7AEFFFFE7FF9A64D0FC7C0356FC1F" box="[151,301,906,931]" pageId="73" pageNumber="74" type="female">three females</specimenCount>
destroyed for DNA sequence (amplification successful, Codes 0143, 0241 &amp; 0342), type locality,
<date id="946F43B6FFE7FF9A61C7FC7C037CFC7B" pageId="73" pageNumber="74" value="2012-07-30">30 July 2012</date>
, leg. K. Kim.
</paragraph>
<paragraph id="E06E6576FFE7FF9A6480FC240351FBEB" blockId="73.[151,1436,618,2011]" pageId="73" pageNumber="74">
<materialsCitation id="50B96F2BFFE7FF9A6480FC24035EFBEB" ID-GBIF-Occurrence="3313373301" collectingDate="2012-11-18" collectorName="K. Kim" country="South Korea" latitude="34.920944" location="South Sea" longLatPrecision="1" longitude="127.78553" municipality="Gwangyang Bay" pageId="73" pageNumber="74" specimenCount="31" specimenCount-female="31" typeStatus="paratype">
Thirty male
<typeStatus id="3F6ADBD4FFE7FF9A6513FC2503C4FC57" box="[340,447,979,1003]" pageId="73" pageNumber="74" type="paratype">paratypes</typeStatus>
and
<specimenCount id="F6D7AEFFFFE7FF9A65BFFC240013FC57" box="[504,616,978,1003]" pageId="73" pageNumber="74" type="female">30 female</specimenCount>
<typeStatus id="3F6ADBD4FFE7FF9A6636FC2500A7FC57" box="[625,732,979,1003]" pageId="73" pageNumber="74" type="paratype">paratypes</typeStatus>
together in ethanol (collection number NIBRIV0000232710),
<specimenCount id="F6D7AEFFFFE7FF9A64D0FC010361FBB3" box="[151,282,1014,1039]" pageId="73" pageNumber="74" type="female">one female</specimenCount>
destroyed for DNA sequence (amplification successful, Code 0547),
<collectingCountry id="98C625E6FFE7FF9A6036FC000773FBB2" box="[1137,1288,1014,1039]" name="South Korea" pageId="73" pageNumber="74">South Korea</collectingCountry>
,
<location id="E50E33ADFFE7FF9A615BFC0007E3FBB3" LSID="urn:lsid:plazi:treatment:6878D460FFE7FF8064D0FE0901D2FACB:E50E33ADFFE7FF9A615BFC0007E3FBB3" box="[1308,1432,1014,1039]" country="South Korea" latitude="34.920944" longLatPrecision="1" longitude="127.78553" municipality="Gwangyang Bay" name="South Sea" pageId="73" pageNumber="74">South Sea</location>
,
<collectingMunicipality id="000AFF0CFFE7FF9A64D0FBEC0328FB8F" box="[151,339,1050,1075]" pageId="73" pageNumber="74">Gwangyang Bay</collectingMunicipality>
, sampling station 10, muddy sediments,
<geoCoordinate id="85E503B1FFE7FF9A675BFBEC01C9FB8F" box="[796,946,1050,1075]" degrees="34.920944" direction="north" orientation="latitude" pageId="73" pageNumber="74" precision="1" value="34.920944">34.920944°N</geoCoordinate>
<geoCoordinate id="85E503B1FFE7FF9A67FDFBEC0625FB8F" box="[954,1118,1050,1075]" degrees="127.785528" direction="east" orientation="longitude" pageId="73" pageNumber="74" precision="1" value="127.78553">127.785528°E</geoCoordinate>
,
<date id="946F43B6FFE7FF9A602EFBEC073EFB8F" box="[1129,1349,1050,1075]" pageId="73" pageNumber="74" value="2012-11-18">
<collectingDate id="842BBA5EFFE7FF9A602EFBEC073EFB8F" box="[1129,1349,1050,1075]" pageId="73" pageNumber="74" value="2012-11-18">18 November 2012</collectingDate>
</date>
, leg.
<collectorName id="4D2400A0FFE7FF9A61C6FBED02B2FBEB" pageId="73" pageNumber="74">K. Kim</collectorName>
(
<figureCitation id="78EA79F3FFE7FF9A649FFBC80362FBEB" box="[216,281,1086,1111]" captionStart="FIGURE 1" captionStartId="7.[151,249,1481,1503]" captionTargetBox="[151,1436,589,1460]" captionTargetId="figure-399@7.[151,1436,589,1460]" captionTargetPageId="7" captionText="FIGURE 1. Map of 17 sampling locations in Gwangyang Bay. Inset showing location of the bay in South Korea. All maps from Google Earth." figureDoi="http://doi.org/10.5281/zenodo.4910564" httpUri="https://zenodo.org/record/4910564/files/figure.png" pageId="73" pageNumber="74">Fig. 1</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="E06E6576FFE7FF9A6480FB94077BFB7F" blockId="73.[151,1436,618,2011]" pageId="73" pageNumber="74">
<materialsCitation id="50B96F2BFFE7FF9A6480FB94077BFB7F" ID-GBIF-Occurrence="3313373309" collectingDate="2012-02-18" collectorName="K. Kim." country="South Korea" latitude="34.920944" location="South Sea" longLatPrecision="1" longitude="127.78553" municipality="Gwangyang Bay" pageId="73" pageNumber="74" specimenCount="13" specimenCount-female="13" typeStatus="paratype">
Three male
<typeStatus id="3F6ADBD4FFE7FF9A6517FB9503C0FBC7" box="[336,443,1123,1147]" pageId="73" pageNumber="74" type="paratype">paratypes</typeStatus>
and
<specimenCount id="F6D7AEFFFFE7FF9A65B2FB94001DFBC7" box="[501,614,1122,1147]" pageId="73" pageNumber="74" type="female">12 female</specimenCount>
<typeStatus id="3F6ADBD4FFE7FF9A6628FB9500A1FBC7" box="[623,730,1123,1147]" pageId="73" pageNumber="74" type="paratype">paratypes</typeStatus>
together in ethanol (collection number NIBRIV0000232711),
<specimenCount id="F6D7AEFFFFE7FF9A64D0FB710368FB23" box="[151,275,1158,1183]" pageId="73" pageNumber="74" type="female">one female</specimenCount>
destroyed for DNA sequence (amplification unsuccessful),
<collectingCountry id="98C625E6FFE7FF9A67FBFB700637FB22" box="[956,1100,1158,1183]" name="South Korea" pageId="73" pageNumber="74">South Korea</collectingCountry>
,
<location id="E50E33ADFFE7FF9A601EFB7006B6FB23" LSID="urn:lsid:plazi:treatment:6878D460FFE7FF8064D0FE0901D2FACB:E50E33ADFFE7FF9A601EFB7006B6FB23" box="[1113,1229,1158,1183]" country="South Korea" latitude="34.920944" longLatPrecision="1" longitude="127.78553" municipality="Gwangyang Bay" name="South Sea" pageId="73" pageNumber="74">South Sea</location>
,
<collectingMunicipality id="000AFF0CFFE7FF9A609DFB7007ECFB23" box="[1242,1431,1158,1183]" pageId="73" pageNumber="74">Gwangyang Bay</collectingMunicipality>
, sampling station 10, muddy sediments,
<geoCoordinate id="85E503B1FFE7FF9A6614FB5C0092FB7F" box="[595,745,1194,1219]" degrees="34.920944" direction="north" orientation="latitude" pageId="73" pageNumber="74" precision="1" value="34.920944">34.920944°N</geoCoordinate>
<geoCoordinate id="85E503B1FFE7FF9A66B7FB5C01EEFB7F" box="[752,917,1194,1219]" degrees="127.785528" direction="east" orientation="longitude" pageId="73" pageNumber="74" precision="1" value="127.78553">127.785528°E</geoCoordinate>
,
<date id="946F43B6FFE7FF9A67E7FB5C0611FB7F" box="[928,1130,1194,1219]" pageId="73" pageNumber="74" value="2012-02-18">
<collectingDate id="842BBA5EFFE7FF9A67E7FB5C0611FB7F" box="[928,1130,1194,1219]" pageId="73" pageNumber="74" value="2012-02-18">18 February 2012</collectingDate>
</date>
, leg.
<collectorName id="4D2400A0FFE7FF9A60E2FB5D077BFB7F" box="[1189,1280,1194,1219]" pageId="73" pageNumber="74">K. Kim.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="E06E6576FFE7FF9A6480FB3807E7FAB7" blockId="73.[151,1436,618,2011]" pageId="73" pageNumber="74">
<materialsCitation id="50B96F2BFFE7FF9A6480FB3807ECFAB7" ID-GBIF-Occurrence="3313373323" collectingDate="2012-11-18" collectorName="K. Kim" country="South Korea" county="Gwangyang Bay" latitude="34.797195" location="One" longLatPrecision="1" longitude="127.786446" municipality="South Sea" pageId="73" pageNumber="74" specimenCount="1">
<location id="E50E33ADFFE7FF9A6480FB38028DFB5B" LSID="urn:lsid:plazi:treatment:6878D460FFE7FF8064D0FE0901D2FACB:E50E33ADFFE7FF9A6480FB38028DFB5B" box="[199,246,1230,1255]" country="South Korea" county="Gwangyang Bay" latitude="34.797195" longLatPrecision="1" longitude="127.786446" municipality="South Sea" name="One" pageId="73" pageNumber="74">One</location>
female destroyed for DNA sequence (amplification unsuccessful),
<collectingCountry id="98C625E6FFE7FF9A67B2FB3806FDFB5A" box="[1013,1158,1230,1255]" name="South Korea" pageId="73" pageNumber="74">South Korea</collectingCountry>
,
<collectingMunicipality id="000AFF0CFFE7FF9A60D4FB38077CFB5B" box="[1171,1287,1230,1255]" pageId="73" pageNumber="74">South Sea</collectingMunicipality>
,
<collectingCounty id="090F1DFAFFE7FF9A6153FB3802BEFAB7" pageId="73" pageNumber="74">Gwangyang Bay</collectingCounty>
, sampling station 15, muddy sediments,
<geoCoordinate id="85E503B1FFE7FF9A66CFFB040164FAB7" box="[648,799,1266,1291]" degrees="34.797194" direction="north" orientation="latitude" pageId="73" pageNumber="74" precision="1" value="34.797195">34.797194°N</geoCoordinate>
<geoCoordinate id="85E503B1FFE7FF9A6762FB0401B1FAB7" box="[805,970,1266,1291]" degrees="127.786444" direction="east" orientation="longitude" pageId="73" pageNumber="74" precision="1" value="127.786446">127.786444°E</geoCoordinate>
,
<date id="946F43B6FFE7FF9A6793FB0406D5FAB7" box="[980,1198,1266,1291]" pageId="73" pageNumber="74" value="2012-11-18">
<collectingDate id="842BBA5EFFE7FF9A6793FB0406D5FAB7" box="[980,1198,1266,1291]" pageId="73" pageNumber="74" value="2012-11-18">18 November 2012</collectingDate>
</date>
, leg.
<collectorName id="4D2400A0FFE7FF9A60AFFB050740FAB7" box="[1256,1339,1266,1291]" pageId="73" pageNumber="74">K. Kim</collectorName>
(
<figureCitation id="78EA79F3FFE7FF9A610DFB0407F0FAB7" box="[1354,1419,1266,1291]" captionStart="FIGURE 1" captionStartId="7.[151,249,1481,1503]" captionTargetBox="[151,1436,589,1460]" captionTargetId="figure-399@7.[151,1436,589,1460]" captionTargetPageId="7" captionText="FIGURE 1. Map of 17 sampling locations in Gwangyang Bay. Inset showing location of the bay in South Korea. All maps from Google Earth." figureDoi="http://doi.org/10.5281/zenodo.4910564" httpUri="https://zenodo.org/record/4910564/files/figure.png" pageId="73" pageNumber="74">Fig. 1</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A8CB36FDFFE7FF9A6480FAE00728FAEF" pageId="73" pageNumber="74" type="etymology">
<paragraph id="E06E6576FFE7FF9A6480FAE00728FAEF" blockId="73.[151,1436,618,2011]" pageId="73" pageNumber="74">
<emphasis id="D2A5B964FFE7FF9A6480FAE00335FA93" bold="true" box="[199,334,1302,1327]" pageId="73" pageNumber="74">Etymology.</emphasis>
The species is named after Thalia (Ancient Greek: Θάλεια), one of nine Muses from Greek mythology, who was a patron of comedy. The species name is a noun in apposition (in the nominative case).
</paragraph>
</subSubSection>
<subSubSection id="A8CB36FDFFE7FF806480FAA801D2FACB" lastPageId="83" lastPageNumber="84" pageId="73" pageNumber="74" type="description">
<paragraph id="E06E6576FFE7FF9A6480FAA803E9F907" blockId="73.[151,1436,618,2011]" pageId="73" pageNumber="74">
<emphasis id="D2A5B964FFE7FF9A6480FAA80322FACB" bold="true" box="[199,345,1374,1399]" pageId="73" pageNumber="74">Description.</emphasis>
Female (based on
<typeStatus id="3F6ADBD4FFE7FF9A667EFAA800E0FACB" box="[569,667,1374,1399]" pageId="73" pageNumber="74" type="holotype">holotype</typeStatus>
and
<specimenCount id="F6D7AEFFFFE7FF9A669FFAA80108FACB" box="[728,883,1374,1399]" pageId="73" pageNumber="74" type="generic" typeStatus="paratypes">six paratypes</specimenCount>
). Body length from
<quantity id="2729C893FFE7FF9A6026FAA80774FACB" box="[1121,1295,1374,1399]" metricMagnitude="-1" metricUnit="m" metricValue="4.07" metricValueMax="4.41" metricValueMin="3.7299999999999995" pageId="73" pageNumber="74" unit="mm" value="407.0" valueMax="441.0" valueMin="373.0">373 to 441 µm</quantity>
(
<quantity id="2729C893FFE7FF9A6166FAA80707FACB" box="[1313,1404,1374,1399]" metricMagnitude="-1" metricUnit="m" metricValue="4.279999999999999" pageId="73" pageNumber="74" unit="mm" value="428.0">428 µm</quantity>
in
<typeStatus id="3F6ADBD4FFE7FF9A64D0FA740285FA27" box="[151,254,1410,1435]" pageId="73" pageNumber="74" type="holotype">holotype</typeStatus>
). Body segmentation, colour, nauplius eye, hyaline fringes, integument thickness as in
<taxonomicName id="27D11EF5FFE7FF9A6141FA75028AFA02" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="calliope" status="sp. nov.">
<emphasis id="D2A5B964FFE7FF9A6141FA75028AFA02" italics="true" pageId="73" pageNumber="74">Wellstenhelia calliope</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFE7FF9A64B8FA50031BFA03" bold="true" box="[255,352,1446,1471]" pageId="73" pageNumber="74">
<taxonomicNameLabel id="C996041FFFE7FF9A64B8FA50031BFA03" box="[255,352,1446,1471]" pageId="73" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Surface appearance of prosomites extremely smooth and without minute pits even when examined on highest magnifications with scanning electron microscope; surface of urosomites mostly covered with numerous irregular rows of minute pits on dorsal and lateral sides, except for hyaline fringes which generally smooth and slightly frilled. Most somite ornamentation also similar to
<taxonomicName id="27D11EF5FFE7FF9A67AEF9E5069EF997" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1001,1253,1555,1579]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE7FF9A67AEF9E5069EF997" box="[1001,1253,1555,1579]" italics="true" pageId="73" pageNumber="74">Wellstenhelia calliope</emphasis>
</taxonomicName>
, and presumed homologous pore and sensilla numbered with same Arabic numerals (see
<figureCitation id="78EA79F3FFE7FF9A679FF9C006F6F9F3" box="[984,1165,1590,1615]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="73" pageNumber="74">Figs. 48A, B, C</figureCitation>
,
<figureCitation id="78EA79F3FFE7FF9A60DEF9C0074BF9F3" box="[1177,1328,1590,1615]" captionStart="FIGURE 49" captionStartId="76.[151,249,1854,1876]" captionTargetBox="[216,1372,198,1825]" captionTargetId="figure-124@76.[210,1374,193,1832]" captionTargetPageId="76" captionText="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="73" pageNumber="74">49A, B, C, D</figureCitation>
) to allow easier comparison. Habitus (
<figureCitation id="78EA79F3FFE7FF9A659EF9AC0015F9CE" box="[473,622,1626,1651]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="73" pageNumber="74">Figs. 48A, B</figureCitation>
,
<figureCitation id="78EA79F3FFE7FF9A663CF9AC00D6F9CF" box="[635,685,1626,1651]" captionStart="FIGURE 53" captionStartId="80.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-204@80.[151,1436,193,1538]" captionTargetPageId="80" captionText="FIGURE 53. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–D, female paratype 1, E–F, female paratype 2: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, genital double-somite, dorsal view; D, preanal and anal somites and caudal rami, dorsal view; E, mouth appendages, ventral view; F, genital double-somite and fifth legs, ventral view. Scale bars 20 µm (E), 30 µm (D), 40 µm (C), 50 µm (B, F), and 100 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910676" httpUri="https://zenodo.org/record/4910676/files/figure.png" pageId="73" pageNumber="74">53A</figureCitation>
) less robust than in
<taxonomicName id="27D11EF5FFE7FF9A67D5F9AD06F1F9CF" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[914,1162,1627,1651]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE7FF9A67D5F9AD06F1F9CF" box="[914,1162,1627,1651]" italics="true" pageId="73" pageNumber="74">Wellstenhelia calliope</emphasis>
</taxonomicName>
, with prosome/urosome length ratio about 1.1, body length/width ratio from 2.6 to 2.9, and cephalothorax from 1.6 to 1.7 times as wide as genital double-somite.
</paragraph>
<paragraph id="E06E6576FFE7FF9A6480F9310614F8BF" blockId="73.[151,1436,618,2011]" pageId="73" pageNumber="74">
Rostrum (
<figureCitation id="78EA79F3FFE7FF9A6571F9300391F963" box="[310,490,1734,1759]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="73" pageNumber="74">Figs. 48A, B, C</figureCitation>
,
<figureCitation id="78EA79F3FFE7FF9A65B2F930005EF963" box="[501,549,1734,1759]" captionStart="FIGURE 53" captionStartId="80.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-204@80.[151,1436,193,1538]" captionTargetPageId="80" captionText="FIGURE 53. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–D, female paratype 1, E–F, female paratype 2: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, genital double-somite, dorsal view; D, preanal and anal somites and caudal rami, dorsal view; E, mouth appendages, ventral view; F, genital double-somite and fifth legs, ventral view. Scale bars 20 µm (E), 30 µm (D), 40 µm (C), 50 µm (B, F), and 100 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910676" httpUri="https://zenodo.org/record/4910676/files/figure.png" pageId="73" pageNumber="74">53B</figureCitation>
) general shape and position of anterior pair of sensilla no. 1 as in
<taxonomicName id="27D11EF5FFE7FF9A6141F930028AF8BF" authorityName="Karanovic &amp; Kim" authorityYear="2014" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE7FF9A6141F930028AF8BF" italics="true" pageId="73" pageNumber="74">Wellstenhelia calliope</emphasis>
</taxonomicName>
but without dorsal pore no. 2 and with two dorso-lateral pores centrally (no. α).
</paragraph>
<paragraph id="E06E6576FFE7FF996480F8F803ACFF69" blockId="73.[151,1436,618,2011]" lastBlockId="74.[151,1436,151,1977]" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74">
Cephalothorax (
<figureCitation id="78EA79F3FFE7FF9A653AF8F8006EF89A" box="[381,533,1806,1831]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="73" pageNumber="74">Figs. 48A, B</figureCitation>
,
<figureCitation id="78EA79F3FFE7FF9A6664F8F8002FF89B" box="[547,596,1806,1831]" captionStart="FIGURE 49" captionStartId="76.[151,249,1854,1876]" captionTargetBox="[216,1372,198,1825]" captionTargetId="figure-124@76.[210,1374,193,1832]" captionTargetPageId="76" captionText="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="73" pageNumber="74">49C</figureCitation>
,
<figureCitation id="78EA79F3FFE7FF9A6625F8F800E8F89B" box="[610,659,1806,1831]" captionStart="FIGURE 53" captionStartId="80.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-204@80.[151,1436,193,1538]" captionTargetPageId="80" captionText="FIGURE 53. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–D, female paratype 1, E–F, female paratype 2: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, genital double-somite, dorsal view; D, preanal and anal somites and caudal rami, dorsal view; E, mouth appendages, ventral view; F, genital double-somite and fifth legs, ventral view. Scale bars 20 µm (E), 30 µm (D), 40 µm (C), 50 µm (B, F), and 100 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910676" httpUri="https://zenodo.org/record/4910676/files/figure.png" pageId="73" pageNumber="74">53B</figureCitation>
,
<figureCitation id="78EA79F3FFE7FF9A66E6F8F8008DF89B" box="[673,758,1806,1831]" captionStart="FIGURE 54" captionStartId="81.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-216@81.[151,1436,193,1539]" captionTargetPageId="81" captionText="FIGURE 54. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–B, female paratype 3, C, female paratype 4, D–F, male paratype: A, cephalothorax, dorsal view; B, anal somite and caudal rami, posterior view; C, cephalothorax, lateral view; D, antennula, lateral view; E, genital somite and fifth leg, lateral view; F, posterior part of caudal rami, lateral view. Scale bars 10 µm (F), 30 µm (D, E), and 50 µm (A, B, C)." figureDoi="http://doi.org/10.5281/zenodo.4910678" httpUri="https://zenodo.org/record/4910678/files/figure.png" pageId="73" pageNumber="74">54A, C</figureCitation>
) about 0.9 times as long as wide; comprising 30% of total body length. Surface of cephalothoracic shield with 28 unpaired and paired sensilla and pores, most of which probably homologous to those in
<taxonomicName id="27D11EF5FFE7FF9A665DF8A00168F8D2" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[538,787,1878,1902]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE7FF9A665DF8A00168F8D2" box="[538,787,1878,1902]" italics="true" pageId="73" pageNumber="74">Wellstenhelia calliope</emphasis>
</taxonomicName>
(indicated with Arabic numerals in illustrations), but 17 pores and sensilla missing (nos. 3, 6, 10, 14, 16, 17, 18, 19, 21, 23, 26, 27, 29, 33, 34, 37, 40); absolute and relative positions of some pores and sensilla differ; one lateral pair of sensilla probably homologous to that in
<taxonomicName id="27D11EF5FFE7FF9A6141F8680290F866" authorityName="Karanovic &amp; Kim" authorityYear="2014" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="euterpe">
<emphasis id="D2A5B964FFE7FF9A6141F8680290F866" italics="true" pageId="73" pageNumber="74">Wellstenhelia euterpe</emphasis>
</taxonomicName>
(indicated with geometric shape in illustrations); one unpaired dorsal sensillum (no. II) probably homologous to that in
<taxonomicName id="27D11EF5FFE4FF9965D3FF6E00D8FF0C" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[404,675,152,176]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="polyhymnia">
<emphasis id="D2A5B964FFE4FF9965D3FF6E00D8FF0C" box="[404,675,152,176]" italics="true" pageId="74" pageNumber="75">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
; three unique pairs of pores located in anterior half, one lateral (no. β) and two dorsal (nos. γ, δ).
</paragraph>
<paragraph id="E06E6576FFE4FF996480FF29038BFEFC" blockId="74.[151,1436,151,1977]" pageId="74" pageNumber="75">
Pleuron of second pedigerous somite (
<figureCitation id="78EA79F3FFE4FF99663CFF29016BFF4B" box="[635,784,223,248]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="74" pageNumber="75">Figs. 48A, B</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF996759FF290135FF44" box="[798,846,223,248]" captionStart="FIGURE 49" captionStartId="76.[151,249,1854,1876]" captionTargetBox="[216,1372,198,1825]" captionTargetId="figure-124@76.[210,1374,193,1832]" captionTargetPageId="76" captionText="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="74" pageNumber="75">49D</figureCitation>
) relatively narrow, with only four pairs of posterior sensilla, three probably homologous to
<taxonomicName id="27D11EF5FFE4FF99661CFEF20128FEA0" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[603,851,260,284]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE4FF99661CFEF20128FEA0" box="[603,851,260,284]" italics="true" pageId="74" pageNumber="75">Wellstenhelia calliope</emphasis>
</taxonomicName>
(nos. 46, 49, 50) and one probably homologous to
<taxonomicName id="27D11EF5FFE4FF9964D0FEDE03F3FEFC" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[151,392,296,320]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="euterpe">
<emphasis id="D2A5B964FFE4FF9964D0FEDE03F3FEFC" box="[151,392,296,320]" italics="true" pageId="74" pageNumber="75">Wellstenhelia euterpe</emphasis>
</taxonomicName>
(no. IV).
</paragraph>
<paragraph id="E06E6576FFE4FF996480FEBA039AFE11" blockId="74.[151,1436,151,1977]" pageId="74" pageNumber="75">
Pleuron of third pedigerous somite (
<figureCitation id="78EA79F3FFE4FF996629FEBA0173FEDF" box="[622,776,332,357]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="74" pageNumber="75">Figs. 48A, B</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF996750FEBA0131FED9" box="[791,842,332,357]" captionStart="FIGURE 49" captionStartId="76.[151,249,1854,1876]" captionTargetBox="[216,1372,198,1825]" captionTargetId="figure-124@76.[210,1374,193,1832]" captionTargetPageId="76" captionText="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="74" pageNumber="75">49D</figureCitation>
) similar to second pedigerous somite in shape and ornamentation, with only four pairs of posterior sensilla, all probably homologous to those in
<taxonomicName id="27D11EF5FFE4FF996141FE86028AFE10" authorityName="Karanovic &amp; Kim" authorityYear="2014" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE4FF996141FE86028AFE10" italics="true" pageId="74" pageNumber="75">Wellstenhelia calliope</emphasis>
</taxonomicName>
(nos. 53, 54, 55, 56).
</paragraph>
<paragraph id="E06E6576FFE4FF996480FE4100C9FE49" blockId="74.[151,1436,151,1977]" pageId="74" pageNumber="75">
Pleuron of fourth pedigerous somite (
<figureCitation id="78EA79F3FFE4FF99662FFE410096FE6C" box="[616,749,439,464]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="74" pageNumber="75">Fig. 48A, B</figureCitation>
) with only three pairs of posterior sensilla, all homologous to those in
<taxonomicName id="27D11EF5FFE4FF9964B1FE2A0396FE48" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[246,493,476,500]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE4FF9964B1FE2A0396FE48" box="[246,493,476,500]" italics="true" pageId="74" pageNumber="75">Wellstenhelia calliope</emphasis>
</taxonomicName>
(nos. 59, 60, 61).
</paragraph>
<paragraph id="E06E6576FFE4FF996480FE0900DAFDDC" blockId="74.[151,1436,151,1977]" pageId="74" pageNumber="75">
First urosomite (
<figureCitation id="78EA79F3FFE4FF9965C5FE09006EFDAB" box="[386,533,511,536]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="74" pageNumber="75">Figs. 48A, B</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF996665FE09000BFDA4" box="[546,624,511,536]" captionStart="FIGURE 53" captionStartId="80.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-204@80.[151,1436,193,1538]" captionTargetPageId="80" captionText="FIGURE 53. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–D, female paratype 1, E–F, female paratype 2: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, genital double-somite, dorsal view; D, preanal and anal somites and caudal rami, dorsal view; E, mouth appendages, ventral view; F, genital double-somite and fifth legs, ventral view. Scale bars 20 µm (E), 30 µm (D), 40 µm (C), 50 µm (B, F), and 100 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910676" httpUri="https://zenodo.org/record/4910676/files/figure.png" pageId="74" pageNumber="75">53C, F</figureCitation>
) as in
<taxonomicName id="27D11EF5FFE4FF9966FBFDF601C8FDA4" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[700,947,512,536]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE4FF9966FBFDF601C8FDA4" box="[700,947,512,536]" italics="true" pageId="74" pageNumber="75">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except pair of anterior lateral pores no. 63 and pair of dorso-lateral sensilla no. 66 missing; dorsal surface additionally ornamented with several rows of minute spinules, while ventral surface smooth.
</paragraph>
<paragraph id="E06E6576FFE4FF996480FD9A018CFC1F" blockId="74.[151,1436,151,1977]" pageId="74" pageNumber="75">
Genital double-somite (
<figureCitation id="78EA79F3FFE4FF996595FD9A001CFD3F" box="[466,615,620,645]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="74" pageNumber="75">Figs. 48A, B</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF996633FD9A00B3FD3F" box="[628,712,620,645]" captionStart="FIGURE 49" captionStartId="76.[151,249,1854,1876]" captionTargetBox="[216,1372,198,1825]" captionTargetId="figure-124@76.[210,1374,193,1832]" captionTargetPageId="76" captionText="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="74" pageNumber="75">49A, B</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF996692FD9A015FFD39" box="[725,804,620,645]" captionStart="FIGURE 53" captionStartId="80.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-204@80.[151,1436,193,1538]" captionTargetPageId="80" captionText="FIGURE 53. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–D, female paratype 1, E–F, female paratype 2: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, genital double-somite, dorsal view; D, preanal and anal somites and caudal rami, dorsal view; E, mouth appendages, ventral view; F, genital double-somite and fifth legs, ventral view. Scale bars 20 µm (E), 30 µm (D), 40 µm (C), 50 µm (B, F), and 100 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910676" httpUri="https://zenodo.org/record/4910676/files/figure.png" pageId="74" pageNumber="75">53C, F</figureCitation>
) shape as in
<taxonomicName id="27D11EF5FFE4FF9967FEFD9A06CAFD38" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[953,1201,620,644]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE4FF9967FEFD9A06CAFD38" box="[953,1201,620,644]" italics="true" pageId="74" pageNumber="75">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except anterior part much more inflated laterally, almost as wide as first urosomite, with lateral expansions produced into posteriorly directed, large spiniform processes (arrowed in
<figureCitation id="78EA79F3FFE4FF996683FD42014CFD71" box="[708,823,692,717]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="74" pageNumber="75">Figs. 48B</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF996701FD420100FD71" box="[838,891,692,717]" captionStart="FIGURE 49" captionStartId="76.[151,249,1854,1876]" captionTargetBox="[216,1372,198,1825]" captionTargetId="figure-124@76.[210,1374,193,1832]" captionTargetPageId="76" captionText="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="74" pageNumber="75">49A</figureCitation>
); paired copulatory pores small ovoid holes in anterior part, situated ventro-laterally; paired genital apertures situated laterally, covered by reduced sixth legs; only more sclerotized part of genital apparatus visible inside; ornamentation similar to that in
<taxonomicName id="27D11EF5FFE4FF996141FD0A028AFC84" authorityName="Karanovic &amp; Kim" authorityYear="2014" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE4FF996141FD0A028AFC84" italics="true" pageId="74" pageNumber="75">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except most large spinules missing (only three at base of ventral sensillum no. 73, arrowed in
<figureCitation id="78EA79F3FFE4FF996157FCE90712FC84" box="[1296,1385,799,824]" captionStart="FIGURE 49" captionStartId="76.[151,249,1854,1876]" captionTargetBox="[216,1372,198,1825]" captionTargetId="figure-124@76.[210,1374,193,1832]" captionTargetPageId="76" captionText="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="74" pageNumber="75">Fig49A</figureCitation>
, and two close to dorsal sensillum no. 71), lateral pair of sensilla no. 72 also missing, and dorsal anterior pairs of sensilla nos. 69 &amp; 70 more widely spaced; dorsal surface additionally ornamented with many parallel rows of minute spinules, ventral surface smooth; no ventral suture between genital apertures.
</paragraph>
<paragraph id="E06E6576FFE4FF996480FC5901BDFBAC" blockId="74.[151,1436,151,1977]" pageId="74" pageNumber="75">
Third urosomite (
<figureCitation id="78EA79F3FFE4FF9965CDFC590060FC7B" box="[394,539,943,968]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="74" pageNumber="75">Figs. 48A, B</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF996660FC590022FC74" box="[551,601,943,968]" captionStart="FIGURE 49" captionStartId="76.[151,249,1854,1876]" captionTargetBox="[216,1372,198,1825]" captionTargetId="figure-124@76.[210,1374,193,1832]" captionTargetPageId="76" captionText="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="74" pageNumber="75">49A</figureCitation>
) as in
<taxonomicName id="27D11EF5FFE4FF9966E6FC4601ECFC74" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[673,919,944,968]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE4FF9966E6FC4601ECFC74" box="[673,919,944,968]" italics="true" pageId="74" pageNumber="75">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except lateral pair of sensilla no. 76 and most large spinules missing, two or three large spinules only at base of sensilla; dorsal surface additionally ornamented with four parallel rows of minute spinules, while ventral surface smooth.
</paragraph>
<paragraph id="E06E6576FFE4FF996480FBEA033DFBC1" blockId="74.[151,1436,151,1977]" pageId="74" pageNumber="75">
Preanal somite (
<figureCitation id="78EA79F3FFE4FF996539FBEA0069FB8F" box="[382,530,1052,1077]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="74" pageNumber="75">Figs. 48A, B</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF996658FBEA0028FB89" box="[543,595,1052,1077]" captionStart="FIGURE 49" captionStartId="76.[151,249,1854,1876]" captionTargetBox="[216,1372,198,1825]" captionTargetId="figure-124@76.[210,1374,193,1832]" captionTargetPageId="76" captionText="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="74" pageNumber="75">49A</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF996619FBEA00EBFB89" box="[606,656,1052,1077]" captionStart="FIGURE 53" captionStartId="80.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-204@80.[151,1436,193,1538]" captionTargetPageId="80" captionText="FIGURE 53. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–D, female paratype 1, E–F, female paratype 2: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, genital double-somite, dorsal view; D, preanal and anal somites and caudal rami, dorsal view; E, mouth appendages, ventral view; F, genital double-somite and fifth legs, ventral view. Scale bars 20 µm (E), 30 µm (D), 40 µm (C), 50 µm (B, F), and 100 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910676" httpUri="https://zenodo.org/record/4910676/files/figure.png" pageId="74" pageNumber="75">53D</figureCitation>
) without sensilla as in
<taxonomicName id="27D11EF5FFE4FF9967D0FBEA06F4FB88" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[919,1167,1052,1076]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE4FF9967D0FBEA06F4FB88" box="[919,1167,1052,1076]" italics="true" pageId="74" pageNumber="75">Wellstenhelia calliope</emphasis>
</taxonomicName>
but with pair of unique lateral pores (no. ε), two large posterior spinules dorso-laterally on each side, and three parallel rows of minute dorsal spinules.
</paragraph>
<paragraph id="E06E6576FFE4FF996480FB7106DEFB54" blockId="74.[151,1436,151,1977]" pageId="74" pageNumber="75">
Anal somite (
<figureCitation id="78EA79F3FFE4FF996522FB710385FB23" box="[357,510,1159,1184]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="74" pageNumber="75">Figs. 48A, B</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF99664AFB71003AFB1C" box="[525,577,1159,1184]" captionStart="FIGURE 49" captionStartId="76.[151,249,1854,1876]" captionTargetBox="[216,1372,198,1825]" captionTargetId="figure-124@76.[210,1374,193,1832]" captionTargetPageId="76" captionText="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="74" pageNumber="75">49A</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF996608FB7100F9FB1C" box="[591,642,1159,1184]" captionStart="FIGURE 53" captionStartId="80.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-204@80.[151,1436,193,1538]" captionTargetPageId="80" captionText="FIGURE 53. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–D, female paratype 1, E–F, female paratype 2: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, genital double-somite, dorsal view; D, preanal and anal somites and caudal rami, dorsal view; E, mouth appendages, ventral view; F, genital double-somite and fifth legs, ventral view. Scale bars 20 µm (E), 30 µm (D), 40 µm (C), 50 µm (B, F), and 100 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910676" httpUri="https://zenodo.org/record/4910676/files/figure.png" pageId="74" pageNumber="75">53D</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF9966D7FB7100BBFB1C" box="[656,704,1159,1184]" captionStart="FIGURE 54" captionStartId="81.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-216@81.[151,1436,193,1539]" captionTargetPageId="81" captionText="FIGURE 54. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–B, female paratype 3, C, female paratype 4, D–F, male paratype: A, cephalothorax, dorsal view; B, anal somite and caudal rami, posterior view; C, cephalothorax, lateral view; D, antennula, lateral view; E, genital somite and fifth leg, lateral view; F, posterior part of caudal rami, lateral view. Scale bars 10 µm (F), 30 µm (D, E), and 50 µm (A, B, C)." figureDoi="http://doi.org/10.5281/zenodo.4910678" httpUri="https://zenodo.org/record/4910678/files/figure.png" pageId="74" pageNumber="75">54B</figureCitation>
) general shape and ornamentation as in
<taxonomicName id="27D11EF5FFE4FF9960DCFB7E07EEFB1C" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1179,1429,1160,1184]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE4FF9960DCFB7E07EEFB1C" box="[1179,1429,1160,1184]" italics="true" pageId="74" pageNumber="75">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except for fewer spinules along medial cleft and distal margin, lateral pores nos. 79 &amp; 80 missing, and anal operculum well developed, broad and smooth, reaching beyond distal margin of anal somite.
</paragraph>
<paragraph id="E06E6576FFE4FF996480FB0201A4FA21" blockId="74.[151,1436,151,1977]" pageId="74" pageNumber="75">
Caudal rami (
<figureCitation id="78EA79F3FFE4FF996521FB020384FAB7" box="[358,511,1268,1293]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="74" pageNumber="75">Figs. 48A, B</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF996649FB02003BFAB1" box="[526,576,1268,1293]" captionStart="FIGURE 49" captionStartId="76.[151,249,1854,1876]" captionTargetBox="[216,1372,198,1825]" captionTargetId="figure-124@76.[210,1374,193,1832]" captionTargetPageId="76" captionText="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="74" pageNumber="75">49A</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF996608FB0200F8FAB1" box="[591,643,1268,1293]" captionStart="FIGURE 53" captionStartId="80.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-204@80.[151,1436,193,1538]" captionTargetPageId="80" captionText="FIGURE 53. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–D, female paratype 1, E–F, female paratype 2: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, genital double-somite, dorsal view; D, preanal and anal somites and caudal rami, dorsal view; E, mouth appendages, ventral view; F, genital double-somite and fifth legs, ventral view. Scale bars 20 µm (E), 30 µm (D), 40 µm (C), 50 µm (B, F), and 100 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910676" httpUri="https://zenodo.org/record/4910676/files/figure.png" pageId="74" pageNumber="75">53D</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF9966D6FB0200BBFAB1" box="[657,704,1268,1293]" captionStart="FIGURE 54" captionStartId="81.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-216@81.[151,1436,193,1539]" captionTargetPageId="81" captionText="FIGURE 54. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–B, female paratype 3, C, female paratype 4, D–F, male paratype: A, cephalothorax, dorsal view; B, anal somite and caudal rami, posterior view; C, cephalothorax, lateral view; D, antennula, lateral view; E, genital somite and fifth leg, lateral view; F, posterior part of caudal rami, lateral view. Scale bars 10 µm (F), 30 µm (D, E), and 50 µm (A, B, C)." figureDoi="http://doi.org/10.5281/zenodo.4910678" httpUri="https://zenodo.org/record/4910678/files/figure.png" pageId="74" pageNumber="75">54B</figureCitation>
) shape and armature similar to those in
<emphasis id="D2A5B964FFE4FF9960DCFB0207E7FAB0" box="[1179,1436,1268,1292]" italics="true" pageId="74" pageNumber="75">
<taxonomicName id="27D11EF5FFE4FF9960DCFB0207E3FAB0" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1179,1432,1268,1292]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="calliope">Wellstenhelia calliope</taxonomicName>
,
</emphasis>
except more elongated, middle lateral seta much shorter and more slender, inner margin without spinules, anterior ventral pore no. 82 moved more towards outer margin (arrowed in
<figureCitation id="78EA79F3FFE4FF9967E6FACA0670FAE9" box="[929,1035,1340,1365]" captionStart="FIGURE 49" captionStartId="76.[151,249,1854,1876]" captionTargetBox="[216,1372,198,1825]" captionTargetId="figure-124@76.[210,1374,193,1832]" captionTargetPageId="76" captionText="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="74" pageNumber="75">Fig. 49A</figureCitation>
), and posterior ventral pore no. 83 missing; rami cylindrical, about 1.9 times as long as anal somite, 4.6 times as long as wide (ventral view), slightly divergent or parallel, and with space between them about one ramus width.
</paragraph>
<paragraph id="E06E6576FFE4FF996480FA5100BCF9EC" blockId="74.[151,1436,151,1977]" pageId="74" pageNumber="75">
Antennula (
<figureCitation id="78EA79F3FFE4FF99650AFA5103BAFA7C" box="[333,449,1447,1472]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="74" pageNumber="75">Figs. 48D</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF996588FA510066FA03" box="[463,541,1447,1472]" captionStart="FIGURE 49" captionStartId="76.[151,249,1854,1876]" captionTargetBox="[216,1372,198,1825]" captionTargetId="figure-124@76.[210,1374,193,1832]" captionTargetPageId="76" captionText="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="74" pageNumber="75">49E, F</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF99666DFA510021FA7C" box="[554,602,1447,1472]" captionStart="FIGURE 53" captionStartId="80.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-204@80.[151,1436,193,1538]" captionTargetPageId="80" captionText="FIGURE 53. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–D, female paratype 1, E–F, female paratype 2: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, genital double-somite, dorsal view; D, preanal and anal somites and caudal rami, dorsal view; E, mouth appendages, ventral view; F, genital double-somite and fifth legs, ventral view. Scale bars 20 µm (E), 30 µm (D), 40 µm (C), 50 µm (B, F), and 100 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910676" httpUri="https://zenodo.org/record/4910676/files/figure.png" pageId="74" pageNumber="75">53B</figureCitation>
) similar to that in
<taxonomicName id="27D11EF5FFE4FF996773FA5E0656FA7C" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[820,1069,1448,1472]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE4FF996773FA5E0656FA7C" box="[820,1069,1448,1472]" italics="true" pageId="74" pageNumber="75">Wellstenhelia calliope</emphasis>
</taxonomicName>
but sixth and seventh segments almost completely fused, and eighth segment with only three lateral setae and without apical aesthetasc; only ornamentation short row of small spinules in proximal half of first segment and one central row of minute dorsal spinules, distal row of spinules and spiniform process missing; aesthetasc on fourth segment just reaching distal tip of appendage; setal formula 1.10.9.6+ae.3.(4.4).5.
</paragraph>
<paragraph id="E06E6576FFE4FF996480F9AA0369F901" blockId="74.[151,1436,151,1977]" pageId="74" pageNumber="75">
Antenna (
<figureCitation id="78EA79F3FFE4FF996570F9AA03E7F9C9" box="[311,412,1628,1653]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="74" pageNumber="75">Fig. 48E</figureCitation>
) as in
<taxonomicName id="27D11EF5FFE4FF9965ABF9AA009EF9C8" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[492,741,1628,1652]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE4FF9965ABF9AA009EF9C8" box="[492,741,1628,1652]" italics="true" pageId="74" pageNumber="75">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except allobasis and first exopodal segment proportionately shorter, endopod with single slender lateral seta, and inner apical fused seta much longer, spiniform, and geniculate.
</paragraph>
<paragraph id="E06E6576FFE4FF996480F9310678F963" blockId="74.[151,1436,151,1977]" box="[199,1027,1735,1760]" pageId="74" pageNumber="75">
Labrum (
<figureCitation id="78EA79F3FFE4FF996568F93103EAF95C" box="[303,401,1735,1760]" captionStart="FIGURE 53" captionStartId="80.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-204@80.[151,1436,193,1538]" captionTargetPageId="80" captionText="FIGURE 53. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–D, female paratype 1, E–F, female paratype 2: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, genital double-somite, dorsal view; D, preanal and anal somites and caudal rami, dorsal view; E, mouth appendages, ventral view; F, genital double-somite and fifth legs, ventral view. Scale bars 20 µm (E), 30 µm (D), 40 µm (C), 50 µm (B, F), and 100 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910676" httpUri="https://zenodo.org/record/4910676/files/figure.png" pageId="74" pageNumber="75">Fig. 53E</figureCitation>
) and paragnaths (
<figureCitation id="78EA79F3FFE4FF99661EF93100C0F95C" box="[601,699,1735,1760]" captionStart="FIGURE 53" captionStartId="80.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-204@80.[151,1436,193,1538]" captionTargetPageId="80" captionText="FIGURE 53. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–D, female paratype 1, E–F, female paratype 2: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, genital double-somite, dorsal view; D, preanal and anal somites and caudal rami, dorsal view; E, mouth appendages, ventral view; F, genital double-somite and fifth legs, ventral view. Scale bars 20 µm (E), 30 µm (D), 40 µm (C), 50 µm (B, F), and 100 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910676" httpUri="https://zenodo.org/record/4910676/files/figure.png" pageId="74" pageNumber="75">Fig. 53E</figureCitation>
) as in
<taxonomicName id="27D11EF5FFE4FF996742F93E0187F95C" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[773,1020,1736,1760]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE4FF996742F93E0187F95C" box="[773,1020,1736,1760]" italics="true" pageId="74" pageNumber="75">Wellstenhelia calliope</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="E06E6576FFE4FF996480F91A0020F8F1" blockId="74.[151,1436,151,1977]" pageId="74" pageNumber="75">
Mandibula (
<figureCitation id="78EA79F3FFE4FF996515F91A0392F8BF" box="[338,489,1772,1797]" captionStart="FIGURE 50" captionStartId="77.[151,249,1918,1940]" captionTargetBox="[189,1401,185,1886]" captionTargetId="figure-82@77.[177,1405,173,1889]" captionTargetPageId="77" captionText="FIGURE 50. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, mandibula, posterior view; B, mandibula, antero-ventral view; C, maxilla, posterior view; D, first swimming leg, anterior view; E, endopod of second swimming leg, anterior view; F, third exopodal segment of second swimming leg, anterior view; G, third swimming leg, anterior view. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910670" httpUri="https://zenodo.org/record/4910670/files/figure.png" pageId="74" pageNumber="75">Figs. 50A, B</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF9965B0F91A005EF8B9" box="[503,549,1772,1797]" captionStart="FIGURE 53" captionStartId="80.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-204@80.[151,1436,193,1538]" captionTargetPageId="80" captionText="FIGURE 53. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–D, female paratype 1, E–F, female paratype 2: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, genital double-somite, dorsal view; D, preanal and anal somites and caudal rami, dorsal view; E, mouth appendages, ventral view; F, genital double-somite and fifth legs, ventral view. Scale bars 20 µm (E), 30 µm (D), 40 µm (C), 50 µm (B, F), and 100 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910676" httpUri="https://zenodo.org/record/4910676/files/figure.png" pageId="74" pageNumber="75">53E</figureCitation>
) similar to that in
<emphasis id="D2A5B964FFE4FF9966B8F91A0184F8B8" box="[767,1023,1772,1796]" italics="true" pageId="74" pageNumber="75">
<taxonomicName id="27D11EF5FFE4FF9966B8F91A0180F8B8" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[767,1019,1772,1796]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="calliope">Wellstenhelia calliope</taxonomicName>
,
</emphasis>
except cutting edge without central seta and with shorter dorsalmost seta, basis with additional row of slender spinules, and exopod with three lateral setae inserted much closer to each other.
</paragraph>
<paragraph id="E06E6576FFE4FF996480F8A10002F805" blockId="74.[151,1436,151,1977]" pageId="74" pageNumber="75">
Maxillula (
<figureCitation id="78EA79F3FFE4FF996503F8A103CBF8CC" box="[324,432,1879,1904]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="74" pageNumber="75">Figs. 48F</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF9965FAF8A10396F8CC" box="[445,493,1879,1904]" captionStart="FIGURE 49" captionStartId="76.[151,249,1854,1876]" captionTargetBox="[216,1372,198,1825]" captionTargetId="figure-124@76.[210,1374,193,1832]" captionTargetPageId="76" captionText="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="74" pageNumber="75">49G</figureCitation>
,
<figureCitation id="78EA79F3FFE4FF9965BEF8A1005CF8CC" box="[505,551,1879,1904]" captionStart="FIGURE 53" captionStartId="80.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-204@80.[151,1436,193,1538]" captionTargetPageId="80" captionText="FIGURE 53. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–D, female paratype 1, E–F, female paratype 2: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, genital double-somite, dorsal view; D, preanal and anal somites and caudal rami, dorsal view; E, mouth appendages, ventral view; F, genital double-somite and fifth legs, ventral view. Scale bars 20 µm (E), 30 µm (D), 40 µm (C), 50 µm (B, F), and 100 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910676" httpUri="https://zenodo.org/record/4910676/files/figure.png" pageId="74" pageNumber="75">53E</figureCitation>
) as in
<taxonomicName id="27D11EF5FFE4FF996633F8AE0117F8CC" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[628,876,1880,1904]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE4FF996633F8AE0117F8CC" box="[628,876,1880,1904]" italics="true" pageId="74" pageNumber="75">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except cutting edge with stronger combs on two ventralmost spines (arrowed in
<figureCitation id="78EA79F3FFE4FF9965B9F88A0019F829" box="[510,610,1916,1941]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="74" pageNumber="75">Fig. 48F</figureCitation>
), basis almost completely fused with endopod and exopod, and endopod much wider distally (arrowed in
<figureCitation id="78EA79F3FFE4FF996642F8560016F805" box="[517,621,1952,1977]" captionStart="FIGURE 49" captionStartId="76.[151,249,1854,1876]" captionTargetBox="[216,1372,198,1825]" captionTargetId="figure-124@76.[210,1374,193,1832]" captionTargetPageId="76" captionText="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="74" pageNumber="75">Fig. 49G</figureCitation>
).
</paragraph>
<caption id="B4AE35FEFFE5FF9864D0F8E40322F85C" ID-DOI="http://doi.org/10.5281/zenodo.4910666" ID-Zenodo-Dep="4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="75" pageNumber="76" startId="75.[151,249,1810,1832]" targetBox="[239,1370,199,1785]" targetPageId="75">
<paragraph id="E06E6576FFE5FF9864D0F8E40322F85C" blockId="75.[151,1436,1810,2016]" pageId="75" pageNumber="76">
<emphasis id="D2A5B964FFE5FF9864D0F8E4035BF89B" bold="true" box="[151,288,1810,1832]" pageId="75" pageNumber="76">FIGURE 48.</emphasis>
<taxonomicName id="27D11EF5FFE5FF986561F8E4007FF894" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[294,516,1810,1832]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="thalia" status="sp. nov.">
<emphasis id="D2A5B964FFE5FF986561F8E4007FF894" box="[294,516,1810,1832]" italics="true" pageId="75" pageNumber="76">Willenstenhelia thalia</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFE5FF98664CF8E40020F89B" bold="true" box="[523,603,1810,1831]" pageId="75" pageNumber="76">
<taxonomicNameLabel id="C996041FFFE5FF98664CF8E40020F89B" box="[523,603,1810,1831]" pageId="75" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in
<taxonomicName id="27D11EF5FFE5FF986643F8980098F838" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[516,739,1902,1924]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE5FF986643F8980098F838" box="[516,739,1902,1924]" italics="true" pageId="75" pageNumber="76">Wellstenhelia calliope</emphasis>
</taxonomicName>
. Geometric shape (◊) indicates sensillum presumably homologous to that in
<taxonomicName id="27D11EF5FFE5FF9864A1F87A03BEF81E" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[230,453,1932,1954]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="euterpe">
<emphasis id="D2A5B964FFE5FF9864A1F87A03BEF81E" box="[230,453,1932,1954]" italics="true" pageId="75" pageNumber="76">Wellstenhelia euterpe</emphasis>
</taxonomicName>
. Roman numerals (II, IV) indicate sensilla presumably homologous to those in
<taxonomicName id="27D11EF5FFE5FF986163F87A0376F87D" authorityName="Karanovic &amp; Kim" authorityYear="2014" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="polyhymnia">
<emphasis id="D2A5B964FFE5FF986163F87A0376F87D" italics="true" pageId="75" pageNumber="76">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm.
</paragraph>
</caption>
<caption id="B4AE35FEFFE2FF9F64D0F8C806FCF851" ID-DOI="http://doi.org/10.5281/zenodo.4910668" ID-Zenodo-Dep="4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="76" pageNumber="77" startId="76.[151,249,1854,1876]" targetBox="[216,1372,198,1825]" targetPageId="76">
<paragraph id="E06E6576FFE2FF9F64D0F8C806FCF851" blockId="76.[151,1436,1854,2029]" pageId="76" pageNumber="77">
<emphasis id="D2A5B964FFE2FF9F64D0F8C80364F8EF" bold="true" box="[151,287,1854,1876]" pageId="76" pageNumber="77">FIGURE 49.</emphasis>
<taxonomicName id="27D11EF5FFE2FF9F6562F8C80078F8E8" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[293,515,1854,1876]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="thalia" status="sp. nov.">
<emphasis id="D2A5B964FFE2FF9F6562F8C80078F8E8" box="[293,515,1854,1876]" italics="true" pageId="76" pageNumber="77">Willenstenhelia thalia</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFE2FF9F664EF8C80022F8EF" bold="true" box="[521,601,1854,1875]" pageId="76" pageNumber="77">
<taxonomicNameLabel id="C996041FFFE2FF9F664EF8C80022F8EF" box="[521,601,1854,1875]" pageId="76" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in
<taxonomicName id="27D11EF5FFE2FF9F65C9F84F0014F873" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[398,623,1977,1999]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE2FF9F65C9F84F0014F873" box="[398,623,1977,1999]" italics="true" pageId="76" pageNumber="77">Wellstenhelia calliope</emphasis>
</taxonomicName>
. Roman numerals (II, IV) indicate sensilla presumably homologous to those in
<taxonomicName id="27D11EF5FFE2FF9F64D0F82103F1F851" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[151,394,2007,2029]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="polyhymnia">
<emphasis id="D2A5B964FFE2FF9F64D0F82103F1F851" box="[151,394,2007,2029]" italics="true" pageId="76" pageNumber="77">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
. Arrowheads point to most prominent specific features. Scale bars 100 µm.
</paragraph>
</caption>
<caption id="B4AE35FEFFE3FF9E64D0F88801FCF84C" ID-DOI="http://doi.org/10.5281/zenodo.4910670" ID-Zenodo-Dep="4910670" httpUri="https://zenodo.org/record/4910670/files/figure.png" pageId="77" pageNumber="78" startId="77.[151,249,1918,1940]" targetBox="[189,1401,185,1886]" targetPageId="77">
<paragraph id="E06E6576FFE3FF9E64D0F88801FCF84C" blockId="77.[151,1436,1918,2032]" pageId="77" pageNumber="78">
<emphasis id="D2A5B964FFE3FF9E64D0F8880359F82F" bold="true" box="[151,290,1918,1940]" pageId="77" pageNumber="78">FIGURE 50.</emphasis>
<taxonomicName id="27D11EF5FFE3FF9E656CF8880070F828" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[299,523,1918,1940]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="thalia" status="sp. nov.">
<emphasis id="D2A5B964FFE3FF9E656CF8880070F828" box="[299,523,1918,1940]" italics="true" pageId="77" pageNumber="78">Willenstenhelia thalia</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFE3FF9E6653F8890013F828" bold="true" box="[532,616,1919,1940]" pageId="77" pageNumber="78">
<taxonomicNameLabel id="C996041FFFE3FF9E6653F8890013F828" box="[532,616,1919,1940]" pageId="77" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, line drawings, female holotype: A, mandibula, posterior view; B, mandibula, antero-ventral view; C, maxilla, posterior view; D, first swimming leg, anterior view; E, endopod of second swimming leg, anterior view; F, third exopodal segment of second swimming leg, anterior view; G, third swimming leg, anterior view. Arrowheads point to most prominent specific features. Scale bars 100 µm.
</paragraph>
</caption>
<caption id="B4AE35FEFFE0FF9D64D0F88A0606F851" ID-DOI="http://doi.org/10.5281/zenodo.4910672" ID-Zenodo-Dep="4910672" httpUri="https://zenodo.org/record/4910672/files/figure.png" pageId="78" pageNumber="79" startId="78.[151,249,1916,1938]" targetBox="[191,1394,189,1884]" targetPageId="78">
<paragraph id="E06E6576FFE0FF9D64D0F88A0606F851" blockId="78.[151,1436,1916,2029]" pageId="78" pageNumber="79">
<emphasis id="D2A5B964FFE0FF9D64D0F88A035BF82D" bold="true" box="[151,288,1916,1938]" pageId="78" pageNumber="79">FIGURE 51.</emphasis>
<taxonomicName id="27D11EF5FFE0FF9D656FF88A007DF82E" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[296,518,1916,1938]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="thalia" status="sp. nov.">
<emphasis id="D2A5B964FFE0FF9D656FF88A007DF82E" box="[296,518,1916,1938]" italics="true" pageId="78" pageNumber="79">Willenstenhelia thalia</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFE0FF9D6649F88A0024F82D" bold="true" box="[526,607,1916,1937]" pageId="78" pageNumber="79">
<taxonomicNameLabel id="C996041FFFE0FF9D6649F88A0024F82D" box="[526,607,1916,1937]" pageId="78" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, line drawings, A–B, female holotype, C–D, male allotype: A, fourth swimming leg, anterior view; B, fifth swimming leg, anterior view; C, fifth swimming legs, anterior view; D, rostrum and antennula, dorsal view. Arabic numeral indicates sensillum presumably homologous to that in
<taxonomicName id="27D11EF5FFE0FF9D67B0F84F0696F873" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1015,1261,1977,1999]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="polyhymnia">
<emphasis id="D2A5B964FFE0FF9D67B0F84F0696F873" box="[1015,1261,1977,1999]" italics="true" pageId="78" pageNumber="79">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
. Greek letter (α) indicates a unique pore. Arrowheads point to most prominent specific features. Scale bars 100 µm.
</paragraph>
</caption>
<caption id="B4AE35FEFFE1FF9C64D0F8B401B1F84D" ID-DOI="http://doi.org/10.5281/zenodo.4910674" ID-Zenodo-Dep="4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="79" pageNumber="80" startId="79.[151,249,1858,1880]" targetBox="[208,1370,205,1829]" targetPageId="79">
<paragraph id="E06E6576FFE1FF9C64D0F8B401B1F84D" blockId="79.[151,1436,1858,2034]" pageId="79" pageNumber="80">
<emphasis id="D2A5B964FFE1FF9C64D0F8B4035BF8EB" bold="true" box="[151,288,1858,1880]" pageId="79" pageNumber="80">FIGURE 52.</emphasis>
<taxonomicName id="27D11EF5FFE1FF9C6560F8B4007EF8E4" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[295,517,1858,1880]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="thalia" status="sp. nov.">
<emphasis id="D2A5B964FFE1FF9C6560F8B4007EF8E4" box="[295,517,1858,1880]" italics="true" pageId="79" pageNumber="80">Willenstenhelia thalia</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFE1FF9C664BF8B50026F8E4" bold="true" box="[524,605,1859,1880]" pageId="79" pageNumber="80">
<taxonomicNameLabel id="C996041FFFE1FF9C664BF8B50026F8E4" box="[524,605,1859,1880]" pageId="79" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in
<taxonomicName id="27D11EF5FFE1FF9C60F4F87607EDF82A" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1203,1430,1920,1942]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFE1FF9C60F4F87607EDF82A" box="[1203,1430,1920,1942]" italics="true" pageId="79" pageNumber="80">Wellstenhelia calliope</emphasis>
</taxonomicName>
. Geometric shape (◊) indicates sensillum presumably homologous to that in
<taxonomicName id="27D11EF5FFE1FF9C67EAF86806F2F808" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[941,1161,1950,1972]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="euterpe">
<emphasis id="D2A5B964FFE1FF9C67EAF86806F2F808" box="[941,1161,1950,1972]" italics="true" pageId="79" pageNumber="80">Wellstenhelia euterpe</emphasis>
</taxonomicName>
. Roman numerals (II, IV) indicate sensilla presumably homologous to those in
<taxonomicName id="27D11EF5FFE1FF9C66F1F84B01D0F86F" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[694,939,1981,2003]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="polyhymnia">
<emphasis id="D2A5B964FFE1FF9C66F1F84B01D0F86F" box="[694,939,1981,2003]" italics="true" pageId="79" pageNumber="80">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm.
</paragraph>
</caption>
<caption id="B4AE35FEFFFEFF8364D0F9E1014BF935" ID-DOI="http://doi.org/10.5281/zenodo.4910676" ID-Zenodo-Dep="4910676" httpUri="https://zenodo.org/record/4910676/files/figure.png" pageId="80" pageNumber="81" startId="80.[151,249,1559,1581]" targetBox="[151,1436,193,1538]" targetPageId="80">
<paragraph id="E06E6576FFFEFF8364D0F9E1014BF935" blockId="80.[151,1436,1559,1673]" pageId="80" pageNumber="81">
<emphasis id="D2A5B964FFFEFF8364D0F9E1035BF990" bold="true" box="[151,288,1559,1581]" pageId="80" pageNumber="81">FIGURE 53.</emphasis>
<taxonomicName id="27D11EF5FFFEFF836562F9E10078F991" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[293,515,1559,1581]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="thalia" status="sp. nov.">
<emphasis id="D2A5B964FFFEFF836562F9E10078F991" box="[293,515,1559,1581]" italics="true" pageId="80" pageNumber="81">Willenstenhelia thalia</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFFEFF83664EF9E10021F990" bold="true" box="[521,602,1559,1580]" pageId="80" pageNumber="81">
<taxonomicNameLabel id="C996041FFFFEFF83664EF9E10021F990" box="[521,602,1559,1580]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, scanning electron micrographs, A–D, female paratype 1, E–F, female paratype 2: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, genital double-somite, dorsal view; D, preanal and anal somites and caudal rami, dorsal view; E, mouth appendages, ventral view; F, genital double-somite and fifth legs, ventral view. Scale bars 20 µm (E), 30 µm (D), 40 µm (C), 50 µm (B, F), and 100 µm (A).
</paragraph>
</caption>
<paragraph id="E06E6576FFFEFF836480F94606FCF950" blockId="80.[151,1436,1712,1989]" pageId="80" pageNumber="81">
Maxilla (
<figureCitation id="78EA79F3FFFEFF836577F94603D8F975" box="[304,419,1712,1737]" captionStart="FIGURE 50" captionStartId="77.[151,249,1918,1940]" captionTargetBox="[189,1401,185,1886]" captionTargetId="figure-82@77.[177,1405,173,1889]" captionTargetPageId="77" captionText="FIGURE 50. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, mandibula, posterior view; B, mandibula, antero-ventral view; C, maxilla, posterior view; D, first swimming leg, anterior view; E, endopod of second swimming leg, anterior view; F, third exopodal segment of second swimming leg, anterior view; G, third swimming leg, anterior view. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910670" httpUri="https://zenodo.org/record/4910670/files/figure.png" pageId="80" pageNumber="81">Figs. 50C</figureCitation>
,
<figureCitation id="78EA79F3FFFEFF8365F5F9460399F975" box="[434,482,1712,1737]" captionStart="FIGURE 53" captionStartId="80.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-204@80.[151,1436,193,1538]" captionTargetPageId="80" captionText="FIGURE 53. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–D, female paratype 1, E–F, female paratype 2: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, genital double-somite, dorsal view; D, preanal and anal somites and caudal rami, dorsal view; E, mouth appendages, ventral view; F, genital double-somite and fifth legs, ventral view. Scale bars 20 µm (E), 30 µm (D), 40 µm (C), 50 µm (B, F), and 100 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910676" httpUri="https://zenodo.org/record/4910676/files/figure.png" pageId="80" pageNumber="81">53E</figureCitation>
,
<figureCitation id="78EA79F3FFFEFF8365B7F9460059F975" box="[496,546,1712,1737]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="80" pageNumber="81">41A</figureCitation>
) as in
<taxonomicName id="27D11EF5FFFEFF836633F9460115F974" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[628,878,1712,1736]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFFEFF836633F9460115F974" box="[628,878,1712,1736]" italics="true" pageId="80" pageNumber="81">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except dorsal endite on syncoxa with only two setae, syncoxa with only several outer spinules, and endopod with only four slender setae.
</paragraph>
<paragraph id="E06E6576FFFEFF836480F90E068DF8E5" blockId="80.[151,1436,1712,1989]" pageId="80" pageNumber="81">
Maxilliped (
<figureCitation id="78EA79F3FFFEFF836512F90E03B2F8AD" box="[341,457,1784,1809]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="80" pageNumber="81">Figs. 48G</figureCitation>
,
<figureCitation id="78EA79F3FFFEFF83659EF90E007CF8AD" box="[473,519,1784,1809]" captionStart="FIGURE 53" captionStartId="80.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-204@80.[151,1436,193,1538]" captionTargetPageId="80" captionText="FIGURE 53. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–D, female paratype 1, E–F, female paratype 2: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, genital double-somite, dorsal view; D, preanal and anal somites and caudal rami, dorsal view; E, mouth appendages, ventral view; F, genital double-somite and fifth legs, ventral view. Scale bars 20 µm (E), 30 µm (D), 40 µm (C), 50 µm (B, F), and 100 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910676" httpUri="https://zenodo.org/record/4910676/files/figure.png" pageId="80" pageNumber="81">53E</figureCitation>
) segmentation, armature and most ornamentation as in
<emphasis id="D2A5B964FFFEFF8360DEF90E07E7F8AC" box="[1177,1436,1784,1808]" italics="true" pageId="80" pageNumber="81">
<taxonomicName id="27D11EF5FFFEFF8360DEF90E07E3F8AC" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1177,1432,1784,1808]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="calliope">Wellstenhelia calliope</taxonomicName>
,
</emphasis>
except outer distal corner of coxa produced into pronounced and blunt process and second endopodal segment fused basally to first endopodal; also setae on first endopodal segment proportionately much longer.
</paragraph>
<paragraph id="E06E6576FFFEFF836480F8920296F879" blockId="80.[151,1436,1712,1989]" pageId="80" pageNumber="81">
First swimming leg (
<figureCitation id="78EA79F3FFFEFF8365F0F892005BF8C1" box="[439,544,1892,1917]" captionStart="FIGURE 50" captionStartId="77.[151,249,1918,1940]" captionTargetBox="[189,1401,185,1886]" captionTargetId="figure-82@77.[177,1405,173,1889]" captionTargetPageId="77" captionText="FIGURE 50. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, mandibula, posterior view; B, mandibula, antero-ventral view; C, maxilla, posterior view; D, first swimming leg, anterior view; E, endopod of second swimming leg, anterior view; F, third exopodal segment of second swimming leg, anterior view; G, third swimming leg, anterior view. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910670" httpUri="https://zenodo.org/record/4910670/files/figure.png" pageId="80" pageNumber="81">Fig. 50D</figureCitation>
) as in
<taxonomicName id="27D11EF5FFFEFF836637F89301FAF8C1" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[624,897,1893,1917]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="polyhymnia">
<emphasis id="D2A5B964FFFEFF836637F89301FAF8C1" box="[624,897,1893,1917]" italics="true" pageId="80" pageNumber="81">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
, except coxa without proximal row of anterior spinules (arrowed in
<figureCitation id="78EA79F3FFFEFF8365D7F87E0381F81D" box="[400,506,1928,1953]" captionStart="FIGURE 50" captionStartId="77.[151,249,1918,1940]" captionTargetBox="[189,1401,185,1886]" captionTargetId="figure-82@77.[177,1405,173,1889]" captionTargetPageId="77" captionText="FIGURE 50. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, mandibula, posterior view; B, mandibula, antero-ventral view; C, maxilla, posterior view; D, first swimming leg, anterior view; E, endopod of second swimming leg, anterior view; F, third exopodal segment of second swimming leg, anterior view; G, third swimming leg, anterior view. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910670" httpUri="https://zenodo.org/record/4910670/files/figure.png" pageId="80" pageNumber="81">Fig. 50D</figureCitation>
) and distal inner seta on second endopodal segmented inserted closer to distal margin.
</paragraph>
<caption id="B4AE35FEFFFFFF8264D0F9EE00DFF936" ID-DOI="http://doi.org/10.5281/zenodo.4910678" ID-Zenodo-Dep="4910678" httpUri="https://zenodo.org/record/4910678/files/figure.png" pageId="81" pageNumber="82" startId="81.[151,249,1560,1582]" targetBox="[151,1436,193,1539]" targetPageId="81">
<paragraph id="E06E6576FFFFFF8264D0F9EE00DFF936" blockId="81.[151,1436,1560,1674]" pageId="81" pageNumber="82">
<emphasis id="D2A5B964FFFFFF8264D0F9EE035AF991" bold="true" box="[151,289,1560,1582]" pageId="81" pageNumber="82">FIGURE 54.</emphasis>
<taxonomicName id="27D11EF5FFFFFF82656EF9EE0073F992" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[297,520,1560,1582]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="thalia" status="sp. nov.">
<emphasis id="D2A5B964FFFFFF82656EF9EE0073F992" box="[297,520,1560,1582]" italics="true" pageId="81" pageNumber="82">Willenstenhelia thalia</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFFFFF826657F9EF0019F992" bold="true" box="[528,610,1561,1582]" pageId="81" pageNumber="82">
<taxonomicNameLabel id="C996041FFFFFFF826657F9EF0019F992" box="[528,610,1561,1582]" pageId="81" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, scanning electron micrographs, A–B, female paratype 3, C, female paratype 4, D–F, male paratype: A, cephalothorax, dorsal view; B, anal somite and caudal rami, posterior view; C, cephalothorax, lateral view; D, antennula, lateral view; E, genital somite and fifth leg, lateral view; F, posterior part of caudal rami, lateral view. Scale bars 10 µm (F), 30 µm (D, E), and 50 µm (A, B, C).
</paragraph>
</caption>
<paragraph id="E06E6576FFFFFF826480F94700FFF8AE" blockId="81.[151,1436,1713,2027]" pageId="81" pageNumber="82">
Second swimming leg (
<figureCitation id="78EA79F3FFFFFF826589F9470035F976" box="[462,590,1713,1738]" captionStart="FIGURE 50" captionStartId="77.[151,249,1918,1940]" captionTargetBox="[189,1401,185,1886]" captionTargetId="figure-82@77.[177,1405,173,1889]" captionTargetPageId="77" captionText="FIGURE 50. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, mandibula, posterior view; B, mandibula, antero-ventral view; C, maxilla, posterior view; D, first swimming leg, anterior view; E, endopod of second swimming leg, anterior view; F, third exopodal segment of second swimming leg, anterior view; G, third swimming leg, anterior view. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910670" httpUri="https://zenodo.org/record/4910670/files/figure.png" pageId="81" pageNumber="82">Fig. 50E, F</figureCitation>
) as in
<taxonomicName id="27D11EF5FFFFFF8266D1F94401DFF976" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[662,932,1714,1738]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="polyhymnia">
<emphasis id="D2A5B964FFFFFF8266D1F94401DFF976" box="[662,932,1714,1738]" italics="true" pageId="81" pageNumber="82">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
, except first exopodal segment without inner seta, third exopodal segment without distal inner seta (arrowed in
<figureCitation id="78EA79F3FFFFFF8267CBF9230195F952" box="[908,1006,1749,1774]" captionStart="FIGURE 50" captionStartId="77.[151,249,1918,1940]" captionTargetBox="[189,1401,185,1886]" captionTargetId="figure-82@77.[177,1405,173,1889]" captionTargetPageId="77" captionText="FIGURE 50. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, mandibula, posterior view; B, mandibula, antero-ventral view; C, maxilla, posterior view; D, first swimming leg, anterior view; E, endopod of second swimming leg, anterior view; F, third exopodal segment of second swimming leg, anterior view; G, third swimming leg, anterior view. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910670" httpUri="https://zenodo.org/record/4910670/files/figure.png" pageId="81" pageNumber="82">Fig. 50F</figureCitation>
) and second endopodal segment with shorter proximal seta (arrowed in
<figureCitation id="78EA79F3FFFFFF826654F90F0003F8AE" box="[531,632,1785,1810]" captionStart="FIGURE 50" captionStartId="77.[151,249,1918,1940]" captionTargetBox="[189,1401,185,1886]" captionTargetId="figure-82@77.[177,1405,173,1889]" captionTargetPageId="77" captionText="FIGURE 50. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, mandibula, posterior view; B, mandibula, antero-ventral view; C, maxilla, posterior view; D, first swimming leg, anterior view; E, endopod of second swimming leg, anterior view; F, third exopodal segment of second swimming leg, anterior view; G, third swimming leg, anterior view. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910670" httpUri="https://zenodo.org/record/4910670/files/figure.png" pageId="81" pageNumber="82">Fig. 50E</figureCitation>
).
</paragraph>
<paragraph id="E06E6576FFFFFF826480F8EB0370F87A" blockId="81.[151,1436,1713,2027]" pageId="81" pageNumber="82">
Third swimming leg (
<figureCitation id="78EA79F3FFFFFF826583F8EB0055F88A" box="[452,558,1821,1846]" captionStart="FIGURE 50" captionStartId="77.[151,249,1918,1940]" captionTargetBox="[189,1401,185,1886]" captionTargetId="figure-82@77.[177,1405,173,1889]" captionTargetPageId="77" captionText="FIGURE 50. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, mandibula, posterior view; B, mandibula, antero-ventral view; C, maxilla, posterior view; D, first swimming leg, anterior view; E, endopod of second swimming leg, anterior view; F, third exopodal segment of second swimming leg, anterior view; G, third swimming leg, anterior view. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910670" httpUri="https://zenodo.org/record/4910670/files/figure.png" pageId="81" pageNumber="82">Fig. 50G</figureCitation>
) as in
<taxonomicName id="27D11EF5FFFFFF8266C6F8E801E9F88A" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[641,914,1822,1846]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="polyhymnia">
<emphasis id="D2A5B964FFFFFF8266C6F8E801E9F88A" box="[641,914,1822,1846]" italics="true" pageId="81" pageNumber="82">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
, except intercoxal sclerite with shorter distal projections, coxa without proximal row of spinules (arrowed in
<figureCitation id="78EA79F3FFFFFF826730F8B7019AF8E6" box="[887,993,1857,1882]" captionStart="FIGURE 50" captionStartId="77.[151,249,1918,1940]" captionTargetBox="[189,1401,185,1886]" captionTargetId="figure-82@77.[177,1405,173,1889]" captionTargetPageId="77" captionText="FIGURE 50. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, mandibula, posterior view; B, mandibula, antero-ventral view; C, maxilla, posterior view; D, first swimming leg, anterior view; E, endopod of second swimming leg, anterior view; F, third exopodal segment of second swimming leg, anterior view; G, third swimming leg, anterior view. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910670" httpUri="https://zenodo.org/record/4910670/files/figure.png" pageId="81" pageNumber="82">Fig. 50G</figureCitation>
), first exopodal segment without inner seta (arrowed in
<figureCitation id="78EA79F3FFFFFF826511F89303C4F8C2" box="[342,447,1893,1918]" captionStart="FIGURE 50" captionStartId="77.[151,249,1918,1940]" captionTargetBox="[189,1401,185,1886]" captionTargetId="figure-82@77.[177,1405,173,1889]" captionTargetPageId="77" captionText="FIGURE 50. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, mandibula, posterior view; B, mandibula, antero-ventral view; C, maxilla, posterior view; D, first swimming leg, anterior view; E, endopod of second swimming leg, anterior view; F, third exopodal segment of second swimming leg, anterior view; G, third swimming leg, anterior view. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910670" httpUri="https://zenodo.org/record/4910670/files/figure.png" pageId="81" pageNumber="82">Fig. 50G</figureCitation>
), third exopodal segment without distal exopodal setae (arrowed in
<figureCitation id="78EA79F3FFFFFF826084F8930752F8C2" box="[1219,1321,1893,1918]" captionStart="FIGURE 50" captionStartId="77.[151,249,1918,1940]" captionTargetBox="[189,1401,185,1886]" captionTargetId="figure-82@77.[177,1405,173,1889]" captionTargetPageId="77" captionText="FIGURE 50. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, mandibula, posterior view; B, mandibula, antero-ventral view; C, maxilla, posterior view; D, first swimming leg, anterior view; E, endopod of second swimming leg, anterior view; F, third exopodal segment of second swimming leg, anterior view; G, third swimming leg, anterior view. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910670" httpUri="https://zenodo.org/record/4910670/files/figure.png" pageId="81" pageNumber="82">Fig. 50G</figureCitation>
) and with three outer spines, and third endopodal segment without proximal or distal inner setae (both positions arrowed in
<figureCitation id="78EA79F3FFFFFF8264D0F85B0284F87A" box="[151,255,1965,1990]" captionStart="FIGURE 50" captionStartId="77.[151,249,1918,1940]" captionTargetBox="[189,1401,185,1886]" captionTargetId="figure-82@77.[177,1405,173,1889]" captionTargetPageId="77" captionText="FIGURE 50. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, mandibula, posterior view; B, mandibula, antero-ventral view; C, maxilla, posterior view; D, first swimming leg, anterior view; E, endopod of second swimming leg, anterior view; F, third exopodal segment of second swimming leg, anterior view; G, third swimming leg, anterior view. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910670" httpUri="https://zenodo.org/record/4910670/files/figure.png" pageId="81" pageNumber="82">Fig. 50G</figureCitation>
).
</paragraph>
<paragraph id="E06E6576FFFFFF816480F8240353FEFC" blockId="81.[151,1436,1713,2027]" lastBlockId="82.[151,1436,151,2013]" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82">
Fourth swimming leg (
<figureCitation id="78EA79F3FFFFFF826590F824003AF857" box="[471,577,2002,2027]" captionStart="FIGURE 51" captionStartId="78.[151,249,1916,1938]" captionTargetBox="[191,1394,189,1884]" captionTargetId="figure-90@78.[184,1401,179,1891]" captionTargetPageId="78" captionText="FIGURE 51. Willenstenhelia thalia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, fourth swimming leg, anterior view; B, fifth swimming leg, anterior view; C, fifth swimming legs, anterior view; D, rostrum and antennula, dorsal view. Arabic numeral indicates sensillum presumably homologous to that in Itostenhelia polyhymnia. Greek letter (α) indicates a unique pore. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910672" httpUri="https://zenodo.org/record/4910672/files/figure.png" pageId="81" pageNumber="82">Fig. 51A</figureCitation>
) as in
<taxonomicName id="27D11EF5FFFFFF8266DCF82401ECF856" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[667,919,2002,2026]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFFFFF8266DCF82401ECF856" box="[667,919,2002,2026]" italics="true" pageId="81" pageNumber="82">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except intercoxal sclerite with sharp distal projections, basis without inner spinules, first exopodal segment without inner setae (arrowed in
<figureCitation id="78EA79F3FFFCFF8160AEFF61072AFF0C" box="[1257,1361,151,176]" captionStart="FIGURE 51" captionStartId="78.[151,249,1916,1938]" captionTargetBox="[191,1394,189,1884]" captionTargetId="figure-90@78.[184,1401,179,1891]" captionTargetPageId="78" captionText="FIGURE 51. Willenstenhelia thalia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, fourth swimming leg, anterior view; B, fifth swimming leg, anterior view; C, fifth swimming legs, anterior view; D, rostrum and antennula, dorsal view. Arabic numeral indicates sensillum presumably homologous to that in Itostenhelia polyhymnia. Greek letter (α) indicates a unique pore. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910672" httpUri="https://zenodo.org/record/4910672/files/figure.png" pageId="82" pageNumber="83">Fig. 51A</figureCitation>
), third exopodal segment with only two outer spines and without proximal or distal inner setae (both arrowed in
<figureCitation id="78EA79F3FFFCFF81616EFF4A07F4FF69" box="[1321,1423,188,213]" captionStart="FIGURE 51" captionStartId="78.[151,249,1916,1938]" captionTargetBox="[191,1394,189,1884]" captionTargetId="figure-90@78.[184,1401,179,1891]" captionTargetPageId="78" captionText="FIGURE 51. Willenstenhelia thalia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, fourth swimming leg, anterior view; B, fifth swimming leg, anterior view; C, fifth swimming legs, anterior view; D, rostrum and antennula, dorsal view. Arabic numeral indicates sensillum presumably homologous to that in Itostenhelia polyhymnia. Greek letter (α) indicates a unique pore. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910672" httpUri="https://zenodo.org/record/4910672/files/figure.png" pageId="82" pageNumber="83">Fig. 51A</figureCitation>
), first endopodal segment smaller and with longer and stronger inner seta (arrowed in
<figureCitation id="78EA79F3FFFCFF816012FF2906C6FF44" box="[1109,1213,223,248]" captionStart="FIGURE 51" captionStartId="78.[151,249,1916,1938]" captionTargetBox="[191,1394,189,1884]" captionTargetId="figure-90@78.[184,1401,179,1891]" captionTargetPageId="78" captionText="FIGURE 51. Willenstenhelia thalia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, fourth swimming leg, anterior view; B, fifth swimming leg, anterior view; C, fifth swimming legs, anterior view; D, rostrum and antennula, dorsal view. Arabic numeral indicates sensillum presumably homologous to that in Itostenhelia polyhymnia. Greek letter (α) indicates a unique pore. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910672" httpUri="https://zenodo.org/record/4910672/files/figure.png" pageId="82" pageNumber="83">Fig. 51A</figureCitation>
), second endopodal segment without inner seta (arrowed in
<figureCitation id="78EA79F3FFFCFF816610FEF200C5FEA1" box="[599,702,260,285]" captionStart="FIGURE 51" captionStartId="78.[151,249,1916,1938]" captionTargetBox="[191,1394,189,1884]" captionTargetId="figure-90@78.[184,1401,179,1891]" captionTargetPageId="78" captionText="FIGURE 51. Willenstenhelia thalia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, fourth swimming leg, anterior view; B, fifth swimming leg, anterior view; C, fifth swimming legs, anterior view; D, rostrum and antennula, dorsal view. Arabic numeral indicates sensillum presumably homologous to that in Itostenhelia polyhymnia. Greek letter (α) indicates a unique pore. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910672" httpUri="https://zenodo.org/record/4910672/files/figure.png" pageId="82" pageNumber="83">Fig. 51A</figureCitation>
), and third endopodal segment with only one inner seta (arrowed in
<figureCitation id="78EA79F3FFFCFF8164F3FED10360FEFC" box="[180,283,295,320]" captionStart="FIGURE 51" captionStartId="78.[151,249,1916,1938]" captionTargetBox="[191,1394,189,1884]" captionTargetId="figure-90@78.[184,1401,179,1891]" captionTargetPageId="78" captionText="FIGURE 51. Willenstenhelia thalia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, fourth swimming leg, anterior view; B, fifth swimming leg, anterior view; C, fifth swimming legs, anterior view; D, rostrum and antennula, dorsal view. Arabic numeral indicates sensillum presumably homologous to that in Itostenhelia polyhymnia. Greek letter (α) indicates a unique pore. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910672" httpUri="https://zenodo.org/record/4910672/files/figure.png" pageId="82" pageNumber="83">Fig. 51A</figureCitation>
).
</paragraph>
<paragraph id="E06E6576FFFCFF816480FEBA012CFDDC" blockId="82.[151,1436,151,2013]" pageId="82" pageNumber="83">
Fifth leg (
<figureCitation id="78EA79F3FFFCFF81657BFEBA03CAFED9" box="[316,433,332,357]" captionStart="FIGURE 49" captionStartId="76.[151,249,1854,1876]" captionTargetBox="[216,1372,198,1825]" captionTargetId="figure-124@76.[210,1374,193,1832]" captionTargetPageId="76" captionText="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="82" pageNumber="83">Figs. 49A</figureCitation>
,
<figureCitation id="78EA79F3FFFCFF816587FEBA038AFED9" box="[448,497,332,357]" captionStart="FIGURE 51" captionStartId="78.[151,249,1916,1938]" captionTargetBox="[191,1394,189,1884]" captionTargetId="figure-90@78.[184,1401,179,1891]" captionTargetPageId="78" captionText="FIGURE 51. Willenstenhelia thalia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, fourth swimming leg, anterior view; B, fifth swimming leg, anterior view; C, fifth swimming legs, anterior view; D, rostrum and antennula, dorsal view. Arabic numeral indicates sensillum presumably homologous to that in Itostenhelia polyhymnia. Greek letter (α) indicates a unique pore. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910672" httpUri="https://zenodo.org/record/4910672/files/figure.png" pageId="82" pageNumber="83">51B</figureCitation>
,
<figureCitation id="78EA79F3FFFCFF8165B8FEBA0056FED9" box="[511,557,332,357]" captionStart="FIGURE 53" captionStartId="80.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-204@80.[151,1436,193,1538]" captionTargetPageId="80" captionText="FIGURE 53. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–D, female paratype 1, E–F, female paratype 2: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, genital double-somite, dorsal view; D, preanal and anal somites and caudal rami, dorsal view; E, mouth appendages, ventral view; F, genital double-somite and fifth legs, ventral view. Scale bars 20 µm (E), 30 µm (D), 40 µm (C), 50 µm (B, F), and 100 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910676" httpUri="https://zenodo.org/record/4910676/files/figure.png" pageId="82" pageNumber="83">53F</figureCitation>
) segmentation as in
<taxonomicName id="27D11EF5FFFCFF816758FEBA0662FED8" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[799,1049,332,356]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFFCFF816758FEBA0662FED8" box="[799,1049,332,356]" italics="true" pageId="82" pageNumber="83">Wellstenhelia calliope</emphasis>
</taxonomicName>
but general shape, armature, and ornamentation very different. Endopodal lobes completely fused, each with three elements, with enormous gap (arrowed in
<figureCitation id="78EA79F3FFFCFF816564FE6203FCFE11" box="[291,391,404,429]" captionStart="FIGURE 51" captionStartId="78.[151,249,1916,1938]" captionTargetBox="[191,1394,189,1884]" captionTargetId="figure-90@78.[184,1401,179,1891]" captionTargetPageId="78" captionText="FIGURE 51. Willenstenhelia thalia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, fourth swimming leg, anterior view; B, fifth swimming leg, anterior view; C, fifth swimming legs, anterior view; D, rostrum and antennula, dorsal view. Arabic numeral indicates sensillum presumably homologous to that in Itostenhelia polyhymnia. Greek letter (α) indicates a unique pore. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910672" httpUri="https://zenodo.org/record/4910672/files/figure.png" pageId="82" pageNumber="83">Fig. 51B</figureCitation>
) between two inner spiniform ones; outer element minute, smooth, and slender (arrowed in
<figureCitation id="78EA79F3FFFCFF8164D0FE410286FE6C" box="[151,253,439,464]" captionStart="FIGURE 51" captionStartId="78.[151,249,1916,1938]" captionTargetBox="[191,1394,189,1884]" captionTargetId="figure-90@78.[184,1401,179,1891]" captionTargetPageId="78" captionText="FIGURE 51. Willenstenhelia thalia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, fourth swimming leg, anterior view; B, fifth swimming leg, anterior view; C, fifth swimming legs, anterior view; D, rostrum and antennula, dorsal view. Arabic numeral indicates sensillum presumably homologous to that in Itostenhelia polyhymnia. Greek letter (α) indicates a unique pore. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910672" httpUri="https://zenodo.org/record/4910672/files/figure.png" pageId="82" pageNumber="83">Fig. 51B</figureCitation>
); middle element strongest, unipinnate along outer margin, twice as long as inner element, and 3.5 times as long as outer slender seta. Exopod almost rectangular (arrowed in
<figureCitation id="78EA79F3FFFCFF8167F1FE2A0666FE49" box="[950,1053,476,501]" captionStart="FIGURE 51" captionStartId="78.[151,249,1916,1938]" captionTargetBox="[191,1394,189,1884]" captionTargetId="figure-90@78.[184,1401,179,1891]" captionTargetPageId="78" captionText="FIGURE 51. Willenstenhelia thalia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, fourth swimming leg, anterior view; B, fifth swimming leg, anterior view; C, fifth swimming legs, anterior view; D, rostrum and antennula, dorsal view. Arabic numeral indicates sensillum presumably homologous to that in Itostenhelia polyhymnia. Greek letter (α) indicates a unique pore. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910672" httpUri="https://zenodo.org/record/4910672/files/figure.png" pageId="82" pageNumber="83">Fig. 51B</figureCitation>
), 1.5 times as long as wide, with longitudinal row of outer spinules in distal part, transverse row of inner spinules, one anterior pore and one posterior pore, with five setae, innermost one larger than some spinules (arrowed in
<figureCitation id="78EA79F3FFFCFF816029FDD206ADFD81" box="[1134,1238,548,573]" captionStart="FIGURE 51" captionStartId="78.[151,249,1916,1938]" captionTargetBox="[191,1394,189,1884]" captionTargetId="figure-90@78.[184,1401,179,1891]" captionTargetPageId="78" captionText="FIGURE 51. Willenstenhelia thalia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, fourth swimming leg, anterior view; B, fifth swimming leg, anterior view; C, fifth swimming legs, anterior view; D, rostrum and antennula, dorsal view. Arabic numeral indicates sensillum presumably homologous to that in Itostenhelia polyhymnia. Greek letter (α) indicates a unique pore. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910672" httpUri="https://zenodo.org/record/4910672/files/figure.png" pageId="82" pageNumber="83">Fig. 51B</figureCitation>
). Length ratio of exopodal setae, starting from inner side, 1: 5.5: 1.5: 1.8: 3.1.
</paragraph>
<paragraph id="E06E6576FFFCFF816480FD9A065CFD1B" blockId="82.[151,1436,151,2013]" pageId="82" pageNumber="83">
Sixth leg (
<figureCitation id="78EA79F3FFFCFF816506FD9A03D3FD39" box="[321,424,620,645]" captionStart="FIGURE 49" captionStartId="76.[151,249,1854,1876]" captionTargetBox="[216,1372,198,1825]" captionTargetId="figure-124@76.[210,1374,193,1832]" captionTargetPageId="76" captionText="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="82" pageNumber="83">Fig. 49B</figureCitation>
) simple small cuticular plate covering genital aperture, inserted at midlength of genital double-somite laterally, with single smooth seta flanking short spiniform process.
</paragraph>
<paragraph id="E06E6576FFFCFF8164D0FD420334FC3C" blockId="82.[151,1436,151,2013]" pageId="82" pageNumber="83">
Male (based on
<typeStatus id="3F6ADBD4FFFCFF816508FD4203D2FD71" box="[335,425,692,717]" pageId="82" pageNumber="83" type="allotype">allotype</typeStatus>
and
<specimenCount id="F6D7AEFFFFFCFF8165A5FD42000DFD71" box="[482,630,692,717]" pageId="82" pageNumber="83" type="generic" typeStatus="paratypes">16 paratypes</specimenCount>
). Body length from
<quantity id="2729C893FFFCFF81671AFD42067EFD70" box="[861,1029,692,717]" metricMagnitude="-1" metricUnit="m" metricValue="3.8899999999999997" metricValueMax="4.029999999999999" metricValueMin="3.75" pageId="82" pageNumber="83" unit="mm" value="389.0" valueMax="403.0" valueMin="375.0">375 to 403 µm</quantity>
(
<quantity id="2729C893FFFCFF816052FD420616FD70" box="[1045,1133,692,717]" metricMagnitude="-1" metricUnit="m" metricValue="3.85" pageId="82" pageNumber="83" unit="mm" value="385.0">385 µm</quantity>
in
<typeStatus id="3F6ADBD4FFFCFF8160D3FD420688FD71" box="[1172,1267,692,717]" pageId="82" pageNumber="83" type="allotype">allotype</typeStatus>
). Habitus (
<figureCitation id="78EA79F3FFFCFF816128FD4202B0FD4C" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Fig. 52A</figureCitation>
), colour, rostrum (
<figureCitation id="78EA79F3FFFCFF8165E8FD21005BFD4C" box="[431,544,727,752]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Fig. 52D</figureCitation>
), shape and ornamentation of cephalothorax (
<figureCitation id="78EA79F3FFFCFF81600DFD2106BEFD4C" box="[1098,1221,727,752]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Figs. 52A</figureCitation>
,
<figureCitation id="78EA79F3FFFCFF816090FD210749FD4C" box="[1239,1330,727,752]" captionStart="FIGURE 54" captionStartId="81.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-216@81.[151,1436,193,1539]" captionTargetPageId="81" captionText="FIGURE 54. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–B, female paratype 3, C, female paratype 4, D–F, male paratype: A, cephalothorax, dorsal view; B, anal somite and caudal rami, posterior view; C, cephalothorax, lateral view; D, antennula, lateral view; E, genital somite and fifth leg, lateral view; F, posterior part of caudal rami, lateral view. Scale bars 10 µm (F), 30 µm (D, E), and 50 µm (A, B, C)." figureDoi="http://doi.org/10.5281/zenodo.4910678" httpUri="https://zenodo.org/record/4910678/files/figure.png" pageId="82" pageNumber="83">54A, C</figureCitation>
), second pedigerous somite (
<figureCitation id="78EA79F3FFFCFF8165C3FD0A007BFCA9" box="[388,512,764,789]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Figs. 52A</figureCitation>
), third pedigerous somite (
<figureCitation id="78EA79F3FFFCFF816708FD0A01BBFCA9" box="[847,960,764,789]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Fig. 52A</figureCitation>
), fourth pedigerous somite (
<figureCitation id="78EA79F3FFFCFF816159FD0A07F4FCA9" box="[1310,1423,764,789]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Fig. 52A</figureCitation>
), ornamentation of first urosomite (
<figureCitation id="78EA79F3FFFCFF816653FCE900D2FC8B" box="[532,681,799,824]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Figs. 52A, B</figureCitation>
,
<figureCitation id="78EA79F3FFFCFF8166F2FCE9009DFC84" box="[693,742,799,824]" captionStart="FIGURE 54" captionStartId="81.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-216@81.[151,1436,193,1539]" captionTargetPageId="81" captionText="FIGURE 54. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–B, female paratype 3, C, female paratype 4, D–F, male paratype: A, cephalothorax, dorsal view; B, anal somite and caudal rami, posterior view; C, cephalothorax, lateral view; D, antennula, lateral view; E, genital somite and fifth leg, lateral view; F, posterior part of caudal rami, lateral view. Scale bars 10 µm (F), 30 µm (D, E), and 50 µm (A, B, C)." figureDoi="http://doi.org/10.5281/zenodo.4910678" httpUri="https://zenodo.org/record/4910678/files/figure.png" pageId="82" pageNumber="83">54E</figureCitation>
), anal somite (
<figureCitation id="78EA79F3FFFCFF8167CBFCE9066EFC84" box="[908,1045,799,824]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Fig. 52A, B</figureCitation>
), caudal rami (
<figureCitation id="78EA79F3FFFCFF8160F8FCE90728FC8B" box="[1215,1363,799,824]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Figs. 52A, B</figureCitation>
,
<figureCitation id="78EA79F3FFFCFF816127FCE907F4FC84" box="[1376,1423,799,824]" captionStart="FIGURE 54" captionStartId="81.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-216@81.[151,1436,193,1539]" captionTargetPageId="81" captionText="FIGURE 54. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–B, female paratype 3, C, female paratype 4, D–F, male paratype: A, cephalothorax, dorsal view; B, anal somite and caudal rami, posterior view; C, cephalothorax, lateral view; D, antennula, lateral view; E, genital somite and fifth leg, lateral view; F, posterior part of caudal rami, lateral view. Scale bars 10 µm (F), 30 µm (D, E), and 50 µm (A, B, C)." figureDoi="http://doi.org/10.5281/zenodo.4910678" httpUri="https://zenodo.org/record/4910678/files/figure.png" pageId="82" pageNumber="83">54F</figureCitation>
), antenna, labrum, paragnaths, mandibula, maxillula, maxilla, maxilliped, first swimming leg, and third swimming leg as in female.
</paragraph>
<paragraph id="E06E6576FFFCFF816480FC7A0748FC74" blockId="82.[151,1436,151,2013]" pageId="82" pageNumber="83">
Genital somite (
<figureCitation id="78EA79F3FFFCFF81653DFC7A0070FC1F" box="[378,523,908,933]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Figs. 52A, B</figureCitation>
,
<figureCitation id="78EA79F3FFFCFF816650FC7A003EFC19" box="[535,581,908,933]" captionStart="FIGURE 54" captionStartId="81.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-216@81.[151,1436,193,1539]" captionTargetPageId="81" captionText="FIGURE 54. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–B, female paratype 3, C, female paratype 4, D–F, male paratype: A, cephalothorax, dorsal view; B, anal somite and caudal rami, posterior view; C, cephalothorax, lateral view; D, antennula, lateral view; E, genital somite and fifth leg, lateral view; F, posterior part of caudal rami, lateral view. Scale bars 10 µm (F), 30 µm (D, E), and 50 µm (A, B, C)." figureDoi="http://doi.org/10.5281/zenodo.4910678" httpUri="https://zenodo.org/record/4910678/files/figure.png" pageId="82" pageNumber="83">54E</figureCitation>
) shorter and wider than in
<taxonomicName id="27D11EF5FFFCFF816736FC7A0613FC18" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[881,1128,908,932]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFFCFF816736FC7A0613FC18" box="[881,1128,908,932]" italics="true" pageId="82" pageNumber="83">Wellstenhelia calliope</emphasis>
</taxonomicName>
and without large spinules; lateral pore no. 68 missing; additionally ornamented with several rows of minute spinules dorso-laterally.
</paragraph>
<paragraph id="E06E6576FFFCFF816480FC220181FB89" blockId="82.[151,1436,151,2013]" pageId="82" pageNumber="83">
Third urosomite (
<figureCitation id="78EA79F3FFFCFF8165D3FC220058FC51" box="[404,547,980,1005]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Fig. 52A, B</figureCitation>
) as in
<emphasis id="D2A5B964FFFCFF81663CFC220106FC50" box="[635,893,980,1004]" italics="true" pageId="82" pageNumber="83">
<taxonomicName id="27D11EF5FFFCFF81663CFC220102FC50" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[635,889,980,1004]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="calliope">Wellstenhelia calliope</taxonomicName>
,
</emphasis>
except lateral pair of sensilla no. 72 missing, ventral row of spinules not interrupted between sensilla pair no. 73, and these sensilla also more widely spaced; additionally ornamented with several rows of minute spinules dorso-laterally.
</paragraph>
<paragraph id="E06E6576FFFCFF816480FBC90139FBC1" blockId="82.[151,1436,151,2013]" pageId="82" pageNumber="83">
Fourth urosomite (
<figureCitation id="78EA79F3FFFCFF8165D9FBC90050FBE4" box="[414,555,1087,1112]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Fig. 52A, B</figureCitation>
) pore and sensilla pattern as in female (i.e. lateral sensilla no. 76 missing), except for continuous ventral posterior row of large spinules.
</paragraph>
<paragraph id="E06E6576FFFCFF816480FB710337FB79" blockId="82.[151,1436,151,2013]" pageId="82" pageNumber="83">
Fifth urosomite (
<figureCitation id="78EA79F3FFFCFF8165C2FB710076FB1C" box="[389,525,1159,1184]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Fig. 52A, B</figureCitation>
) with lateral pore (no. ε) as in female, but with posterior ventral row of spinules of various sizes.
</paragraph>
<paragraph id="E06E6576FFFCFF816480FB390181FAB1" blockId="82.[151,1436,151,2013]" pageId="82" pageNumber="83">
Caudal rami (
<figureCitation id="78EA79F3FFFCFF816524FB390382FB5B" box="[355,505,1231,1256]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Figs. 52A, B</figureCitation>
,
<figureCitation id="78EA79F3FFFCFF816640FB390048FB54" box="[519,563,1231,1256]" captionStart="FIGURE 54" captionStartId="81.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-216@81.[151,1436,193,1539]" captionTargetPageId="81" captionText="FIGURE 54. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–B, female paratype 3, C, female paratype 4, D–F, male paratype: A, cephalothorax, dorsal view; B, anal somite and caudal rami, posterior view; C, cephalothorax, lateral view; D, antennula, lateral view; E, genital somite and fifth leg, lateral view; F, posterior part of caudal rami, lateral view. Scale bars 10 µm (F), 30 µm (D, E), and 50 µm (A, B, C)." figureDoi="http://doi.org/10.5281/zenodo.4910678" httpUri="https://zenodo.org/record/4910678/files/figure.png" pageId="82" pageNumber="83">54F</figureCitation>
) long and slender (arrowed in
<figureCitation id="78EA79F3FFFCFF8167D1FB390184FB54" box="[918,1023,1231,1256]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Fig. 52A</figureCitation>
), about 4.5 times as long as wide in ventral view, with middle lateral seta hardly longer than spinules that flank it.
</paragraph>
<paragraph id="E06E6576FFFCFF816480FAE103C9FAC4" blockId="82.[151,1436,151,2013]" pageId="82" pageNumber="83">
Antennula (
<figureCitation id="78EA79F3FFFCFF816509FAE103B8FA8C" box="[334,451,1303,1328]" captionStart="FIGURE 51" captionStartId="78.[151,249,1916,1938]" captionTargetBox="[191,1394,189,1884]" captionTargetId="figure-90@78.[184,1401,179,1891]" captionTargetPageId="78" captionText="FIGURE 51. Willenstenhelia thalia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, fourth swimming leg, anterior view; B, fifth swimming leg, anterior view; C, fifth swimming legs, anterior view; D, rostrum and antennula, dorsal view. Arabic numeral indicates sensillum presumably homologous to that in Itostenhelia polyhymnia. Greek letter (α) indicates a unique pore. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910672" httpUri="https://zenodo.org/record/4910672/files/figure.png" pageId="82" pageNumber="83">Figs. 51D</figureCitation>
,
<figureCitation id="78EA79F3FFFCFF816595FAE1007FFA8C" box="[466,516,1303,1328]" captionStart="FIGURE 54" captionStartId="81.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-216@81.[151,1436,193,1539]" captionTargetPageId="81" captionText="FIGURE 54. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–B, female paratype 3, C, female paratype 4, D–F, male paratype: A, cephalothorax, dorsal view; B, anal somite and caudal rami, posterior view; C, cephalothorax, lateral view; D, antennula, lateral view; E, genital somite and fifth leg, lateral view; F, posterior part of caudal rami, lateral view. Scale bars 10 µm (F), 30 µm (D, E), and 50 µm (A, B, C)." figureDoi="http://doi.org/10.5281/zenodo.4910678" httpUri="https://zenodo.org/record/4910678/files/figure.png" pageId="82" pageNumber="83">54D</figureCitation>
) as in
<emphasis id="D2A5B964FFFCFF81661FFAEE0108FA8C" box="[600,883,1304,1328]" italics="true" pageId="82" pageNumber="83">
<taxonomicName id="27D11EF5FFFCFF81661FFAEE0114FA8C" box="[600,879,1304,1328]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="terpisichore">Itostenhelia terpisichore</taxonomicName>
,
</emphasis>
except aesthetascs on third and fourth segment shorter, sixth segment with more spinules, and seventh segment with one more seta; setal formula 1.11.5+ae.9+ae.1.2.3.4.5.
</paragraph>
<paragraph id="E06E6576FFFCFF816480FA720318FA5F" blockId="82.[151,1436,151,2013]" pageId="82" pageNumber="83">
Second swimming leg (
<figureCitation id="78EA79F3FFFCFF816593FA720041FA21" box="[468,570,1412,1437]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Fig. 52C</figureCitation>
) with coxa, basis, and exopod as in female. Endopod with second and third segment fused, with only three elements on ancestral segment, and with inner apical element (arrowed in
<figureCitation id="78EA79F3FFFCFF816176FA5107EFFA7C" box="[1329,1428,1447,1472]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Fig. 52C</figureCitation>
) shorter than outer.
</paragraph>
<paragraph id="E06E6576FFFCFF816480FA190760F991" blockId="82.[151,1436,151,2013]" pageId="82" pageNumber="83">
Fourth swimming leg (
<figureCitation id="78EA79F3FFFCFF816580FA190056F9B4" box="[455,557,1519,1544]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Fig. 52D</figureCitation>
) as in female, except outer spine on second exopodal segment strongly curved and sclerotized, highly characteristic and visible even on undissected specimens (arrowed in
<figureCitation id="78EA79F3FFFCFF8160E1F9E20775F991" box="[1190,1294,1556,1581]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Fig. 52D</figureCitation>
).
</paragraph>
<paragraph id="E06E6576FFFCFF816480F9C10664F8F1" blockId="82.[151,1436,151,2013]" pageId="82" pageNumber="83">
Fifth leg (
<figureCitation id="78EA79F3FFFCFF816579F9C103C8F9EC" box="[318,435,1591,1616]" captionStart="FIGURE 51" captionStartId="78.[151,249,1916,1938]" captionTargetBox="[191,1394,189,1884]" captionTargetId="figure-90@78.[184,1401,179,1891]" captionTargetPageId="78" captionText="FIGURE 51. Willenstenhelia thalia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, fourth swimming leg, anterior view; B, fifth swimming leg, anterior view; C, fifth swimming legs, anterior view; D, rostrum and antennula, dorsal view. Arabic numeral indicates sensillum presumably homologous to that in Itostenhelia polyhymnia. Greek letter (α) indicates a unique pore. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910672" httpUri="https://zenodo.org/record/4910672/files/figure.png" pageId="82" pageNumber="83">Figs. 51C</figureCitation>
,
<figureCitation id="78EA79F3FFFCFF816584F9C1038DF9EC" box="[451,502,1591,1616]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">52A</figureCitation>
,
<figureCitation id="78EA79F3FFFCFF816642F9C1004FF9EC" box="[517,564,1591,1616]" captionStart="FIGURE 54" captionStartId="81.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-216@81.[151,1436,193,1539]" captionTargetPageId="81" captionText="FIGURE 54. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–B, female paratype 3, C, female paratype 4, D–F, male paratype: A, cephalothorax, dorsal view; B, anal somite and caudal rami, posterior view; C, cephalothorax, lateral view; D, antennula, lateral view; E, genital somite and fifth leg, lateral view; F, posterior part of caudal rami, lateral view. Scale bars 10 µm (F), 30 µm (D, E), and 50 µm (A, B, C)." figureDoi="http://doi.org/10.5281/zenodo.4910678" httpUri="https://zenodo.org/record/4910678/files/figure.png" pageId="82" pageNumber="83">54E</figureCitation>
) general shape, segmentation, and most armature and ornamentation as in
<emphasis id="D2A5B964FFFCFF8164D0F9AA03D7F9C8" box="[151,428,1628,1652]" italics="true" pageId="82" pageNumber="83">
<taxonomicName id="27D11EF5FFFCFF8164D0F9AA03D2F9C8" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[151,425,1628,1652]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="polyhymnia">Itostenhelia polyhymnia</taxonomicName>
,
</emphasis>
except endopodal lobes slightly more pronounced, exopod slightly larger, and number of exopodal and endopodal armature elements often asymmetrical. Endopodal lobe mostly with two elements, inner one twice as long as outer one; third element always innermost and smallest. Exopod 1.4 times as long as wide, with outer margin smooth or with several strong spinules, inner margin always with several small spinules, with four or five armature elements; innermost armature element as in female minute and slender seta, next one longest and slender seta (arrowed in
<figureCitation id="78EA79F3FFFCFF816590F8F90045F894" box="[471,574,1807,1832]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Fig. 52A</figureCitation>
), third short and slender seta, followed by one or two strong and equally long apical spines. Length ratio of exopodal armature, from inner side, 1: 6.8: 2: 3.5.
</paragraph>
<paragraph id="E06E6576FFFCFF816480F8A106BAF8CC" blockId="82.[151,1436,151,2013]" box="[199,1217,1879,1904]" pageId="82" pageNumber="83">
Sixth legs (
<figureCitation id="78EA79F3FFFCFF816502F8A103CDF8CC" box="[325,438,1879,1904]" captionStart="FIGURE 52" captionStartId="79.[151,249,1858,1880]" captionTargetBox="[208,1370,205,1829]" captionTargetId="figure-120@79.[208,1374,193,1836]" captionTargetPageId="79" captionText="FIGURE 52. Willenstenhelia thalia sp. nov., line drawings, male allotype: A, habitus, lateral view; B, urosome, ventral view; C, endopod of second swimming leg, anterior view; D, last two exopodal segment of fourth swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (β, ε) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910674" httpUri="https://zenodo.org/record/4910674/files/figure.png" pageId="82" pageNumber="83">Figs. 52B</figureCitation>
,
<figureCitation id="78EA79F3FFFCFF816586F8A1038BF8CC" box="[449,496,1879,1904]" captionStart="FIGURE 54" captionStartId="81.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-216@81.[151,1436,193,1539]" captionTargetPageId="81" captionText="FIGURE 54. Willenstenhelia thalia sp. nov., scanning electron micrographs, A–B, female paratype 3, C, female paratype 4, D–F, male paratype: A, cephalothorax, dorsal view; B, anal somite and caudal rami, posterior view; C, cephalothorax, lateral view; D, antennula, lateral view; E, genital somite and fifth leg, lateral view; F, posterior part of caudal rami, lateral view. Scale bars 10 µm (F), 30 µm (D, E), and 50 µm (A, B, C)." figureDoi="http://doi.org/10.5281/zenodo.4910678" httpUri="https://zenodo.org/record/4910678/files/figure.png" pageId="82" pageNumber="83">54E</figureCitation>
) as in
<emphasis id="D2A5B964FFFCFF81667DF8AE0135F8CC" box="[570,846,1880,1904]" italics="true" pageId="82" pageNumber="83">
<taxonomicName id="27D11EF5FFFCFF81667DF8AE0131F8CC" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[570,842,1880,1904]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="polyhymnia">Itostenhelia polyhymnia</taxonomicName>
,
</emphasis>
except both legs distinct at base.
</paragraph>
<paragraph id="E06E6576FFFCFF806480F88A0752FF69" blockId="82.[151,1436,151,2013]" lastBlockId="83.[151,1436,151,1400]" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83">
<emphasis id="D2A5B964FFFCFF816480F88A0336F829" bold="true" box="[199,333,1916,1941]" pageId="82" pageNumber="83">Variability.</emphasis>
Fusion between sixth and seventh antennular segment in female is variable, but is never complete (
<figureCitation id="78EA79F3FFFCFF8164D8F856035EF805" box="[159,293,1952,1977]" captionStart="FIGURE 49" captionStartId="76.[151,249,1854,1876]" captionTargetBox="[216,1372,198,1825]" captionTargetId="figure-124@76.[210,1374,193,1832]" captionTargetPageId="76" captionText="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="82" pageNumber="83">Fig. 49E, F</figureCitation>
). Male fifth leg is often asymmetrical (
<figureCitation id="78EA79F3FFFCFF8166A1F8560135F805" box="[742,846,1952,1977]" captionStart="FIGURE 51" captionStartId="78.[151,249,1916,1938]" captionTargetBox="[191,1394,189,1884]" captionTargetId="figure-90@78.[184,1401,179,1891]" captionTargetPageId="78" captionText="FIGURE 51. Willenstenhelia thalia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, fourth swimming leg, anterior view; B, fifth swimming leg, anterior view; C, fifth swimming legs, anterior view; D, rostrum and antennula, dorsal view. Arabic numeral indicates sensillum presumably homologous to that in Itostenhelia polyhymnia. Greek letter (α) indicates a unique pore. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910672" httpUri="https://zenodo.org/record/4910672/files/figure.png" pageId="82" pageNumber="83">Fig. 51C</figureCitation>
), with more or less spinules on the endopodal lobe, with or without spinules on the outer margin of exopod, with two or three endopodal armature elements, and with four or five exopodal armature elements. Most morphological features are, however, extremely conservative, including the sensilla and pores pattern of somites, and length ratio of different armature on appendages.
</paragraph>
<paragraph id="E06E6576FFFDFF806480FF290172FC7B" blockId="83.[151,1436,151,1400]" pageId="83" pageNumber="84">
<emphasis id="D2A5B964FFFDFF806480FF290394FF44" bold="true" box="[199,495,223,248]" pageId="83" pageNumber="84">Morphological affinities.</emphasis>
<taxonomicName id="27D11EF5FFFDFF8065B0FF160096FF44" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[503,749,224,248]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="thalia" status="sp. nov.">
<emphasis id="D2A5B964FFFDFF8065B0FF160096FF44" box="[503,749,224,248]" italics="true" pageId="83" pageNumber="84">Willenstenhelia thalia</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFFDFF8066B1FF29012AFF44" bold="true" box="[758,849,223,248]" pageId="83" pageNumber="84">
<taxonomicNameLabel id="C996041FFFFDFF8066B1FF29012AFF44" box="[758,849,223,248]" pageId="83" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all congeners by the minute inner seta on the female fifth leg exopod (
<figureCitation id="78EA79F3FFFDFF806647FEF2001CFEA1" box="[512,615,260,285]" captionStart="FIGURE 51" captionStartId="78.[151,249,1916,1938]" captionTargetBox="[191,1394,189,1884]" captionTargetId="figure-90@78.[184,1401,179,1891]" captionTargetPageId="78" captionText="FIGURE 51. Willenstenhelia thalia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, fourth swimming leg, anterior view; B, fifth swimming leg, anterior view; C, fifth swimming legs, anterior view; D, rostrum and antennula, dorsal view. Arabic numeral indicates sensillum presumably homologous to that in Itostenhelia polyhymnia. Greek letter (α) indicates a unique pore. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910672" httpUri="https://zenodo.org/record/4910672/files/figure.png" pageId="83" pageNumber="84">Fig. 51B</figureCitation>
), minute outermost seta on the female fifth leg endopod (
<figureCitation id="78EA79F3FFFDFF8060B1FEF20725FEA1" box="[1270,1374,260,285]" captionStart="FIGURE 51" captionStartId="78.[151,249,1916,1938]" captionTargetBox="[191,1394,189,1884]" captionTargetId="figure-90@78.[184,1401,179,1891]" captionTargetPageId="78" captionText="FIGURE 51. Willenstenhelia thalia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, fourth swimming leg, anterior view; B, fifth swimming leg, anterior view; C, fifth swimming legs, anterior view; D, rostrum and antennula, dorsal view. Arabic numeral indicates sensillum presumably homologous to that in Itostenhelia polyhymnia. Greek letter (α) indicates a unique pore. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910672" httpUri="https://zenodo.org/record/4910672/files/figure.png" pageId="83" pageNumber="84">Fig. 51B</figureCitation>
), and robust and spiniform second seta from outer side on the female fifth leg endopod (
<figureCitation id="78EA79F3FFFDFF80601CFED106BEFEFC" box="[1115,1221,295,320]" captionStart="FIGURE 51" captionStartId="78.[151,249,1916,1938]" captionTargetBox="[191,1394,189,1884]" captionTargetId="figure-90@78.[184,1401,179,1891]" captionTargetPageId="78" captionText="FIGURE 51. Willenstenhelia thalia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, fourth swimming leg, anterior view; B, fifth swimming leg, anterior view; C, fifth swimming legs, anterior view; D, rostrum and antennula, dorsal view. Arabic numeral indicates sensillum presumably homologous to that in Itostenhelia polyhymnia. Greek letter (α) indicates a unique pore. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910672" httpUri="https://zenodo.org/record/4910672/files/figure.png" pageId="83" pageNumber="84">Fig. 51B</figureCitation>
), which can all be considered as autapomorphic character states. It also has a more elongated female fifth leg exopod than any other congener, but this may be a plesiomophy in a larger group of stenheliins. Posteriorly pointed, large spiniform processes on the anterior part of the genital double-somite (
<figureCitation id="78EA79F3FFFDFF806770FE6201D2FE11" box="[823,937,404,429]" captionStart="FIGURE 48" captionStartId="75.[151,249,1810,1832]" captionTargetBox="[239,1370,199,1785]" captionTargetId="figure-129@75.[228,1370,189,1791]" captionTargetPageId="75" captionText="FIGURE 48. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and partly flattened, lateral view; D, proximal part of antennula, ventral view; E, antenna, anterior view; F, praecoxa of maxillula, posterior view; G, maxillipeds, posterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicates sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Greek letters (α, β, γ, δ) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910666" httpUri="https://zenodo.org/record/4910666/files/figure.png" pageId="83" pageNumber="84">Figs. 48B</figureCitation>
,
<figureCitation id="78EA79F3FFFDFF8067F2FE62067CFE11" box="[949,1031,404,429]" captionStart="FIGURE 49" captionStartId="76.[151,249,1854,1876]" captionTargetBox="[216,1372,198,1825]" captionTargetId="figure-124@76.[210,1374,193,1832]" captionTargetPageId="76" captionText="FIGURE 49. Willenstenhelia thalia sp. nov., line drawings, female holotype: A, urosome, ventral view; B, right part of genital double-somite, ventro-lateral view; C, posterior central part of cephalothoracic shield, dissected and flattened; D, pleurons of second and third pedigerous somites, dissected and flattened; E, distal part of antennula, ventral view; F, distal part of antennula, dorso-anterior view; G, maxillular palp, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (II, IV) indicate sensilla presumably homologous to those in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910668" httpUri="https://zenodo.org/record/4910668/files/figure.png" pageId="83" pageNumber="84">49A, B</figureCitation>
) also may be an autapomorphy, but this feature was not described or illustrated in at least two congeners. All these characters would suggest a somewhat isolated position of
<taxonomicName id="27D11EF5FFFDFF8065ACFE2A00A4FE48" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[491,735,476,500]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="thalia">
<emphasis id="D2A5B964FFFDFF8065ACFE2A00A4FE48" box="[491,735,476,500]" italics="true" pageId="83" pageNumber="84">Willenstenhelia thalia</emphasis>
</taxonomicName>
in the genus
<taxonomicName id="27D11EF5FFFDFF806733FE2A065AFE48" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[884,1057,476,500]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="D2A5B964FFFDFF806733FE2A065AFE48" box="[884,1057,476,500]" italics="true" pageId="83" pageNumber="84">Willenstenhelia</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFFDFF806060FE2A06EBFE49" bold="true" box="[1063,1168,476,501]" pageId="83" pageNumber="84">
<taxonomicNameLabel id="C996041FFFFDFF806060FE2A06EBFE49" box="[1063,1168,476,501]" pageId="83" pageNumber="84" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, which is not surprising given that this is the only Pacific representative. Other congeners include
<taxonomicName id="27D11EF5FFFDFF80679BFDF607E7FDA4" authority="(Wells, 1967)" authorityName="Karanovic &amp; Kim" authorityYear="2014" baseAuthorityName="Wells" baseAuthorityYear="1967" box="[988,1436,511,536]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="unisetosa" status="comb. nov.">
<emphasis id="D2A5B964FFFDFF80679BFDF60680FDAB" box="[988,1275,512,536]" italics="true" pageId="83" pageNumber="84">Willenstenhelia unisetosa</emphasis>
(
<bibRefCitation id="84401887FFFDFF80614CFE0907EFFDA4" author="Wells, J. B. J." box="[1291,1428,511,536]" pageId="83" pageNumber="84" refId="ref50213" refString="Wells, J. B. J. (1967) The littoral Copepoda (Crustacea) of Inhaca Island, Mozambique. Transactions of the Royal Society of Edinburgh, 67,189 - 358." type="book" year="1967">Wells, 1967</bibRefCitation>
)
</taxonomicName>
<emphasis id="D2A5B964FFFDFF8064D0FDD20362FD81" bold="true" box="[151,281,548,573]" pageId="83" pageNumber="84">
<taxonomicNameLabel id="C996041FFFFDFF8064D0FDD20362FD81" box="[151,281,548,573]" pageId="83" pageNumber="84" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
from
<collectingCountry id="98C625E6FFFDFF806526FDD2038EFD81" box="[353,501,548,573]" name="Mozambique" pageId="83" pageNumber="84">Mozambique</collectingCountry>
(Indian Ocean),
<taxonomicName id="27D11EF5FFFDFF8066FFFDD201C3FD80" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[696,952,548,572]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="urania" status="sp. nov.">
<emphasis id="D2A5B964FFFDFF8066FFFDD201C3FD80" box="[696,952,548,572]" italics="true" pageId="83" pageNumber="84">Willenstenhelia urania</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFFDFF806786FDD20666FD81" bold="true" box="[961,1053,548,573]" pageId="83" pageNumber="84">
<taxonomicNameLabel id="C996041FFFFDFF806786FDD20666FD81" box="[961,1053,548,573]" pageId="83" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from the Mediterranean Coast of
<collectingCountry id="98C625E6FFFDFF8064D0FDB102A3FDDC" box="[151,216,583,608]" name="Israel" pageId="83" pageNumber="84">Israel</collectingCountry>
,
<taxonomicName id="27D11EF5FFFDFF8064A4FDBE006FFDDC" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[227,532,584,608]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="terpsichore" status="sp. nov.">
<emphasis id="D2A5B964FFFDFF8064A4FDBE006FFDDC" box="[227,532,584,608]" italics="true" pageId="83" pageNumber="84">Willenstenhelia terpsichore</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFFDFF80665BFDB1000EFDDC" bold="true" box="[540,629,583,608]" pageId="83" pageNumber="84">
<taxonomicNameLabel id="C996041FFFFDFF80665BFDB1000EFDDC" box="[540,629,583,608]" pageId="83" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from the Slovenian Coast of the Adriatic Sea (northern Mediterranean), and
<taxonomicName id="27D11EF5FFFDFF806480FD9A0009FD39" authority="T. Scott, 1902" authorityName="T. Scott" authorityYear="1902" box="[199,626,620,645]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="D2A5B964FFFDFF806480FD9A03B1FD38" box="[199,458,620,644]" italics="true" pageId="83" pageNumber="84">Willenstenhelia minuta</emphasis>
T.
<bibRefCitation id="84401887FFFDFF8065B6FD9A0009FD39" author="Scott, A." box="[497,626,620,645]" pageId="83" pageNumber="84" pagination="397 - 428" refId="ref49985" refString="Scott, A. (1902) On some Red Sea and Indian Ocean Copepoda. Proceedings and Transactions of the Liverpool Biological Society, 16, 397 - 428." type="journal article" year="1902">Scott, 1902</bibRefCitation>
</taxonomicName>
from the Indian Ocean and the Suez Canal (see below and
<bibRefCitation id="84401887FFFDFF80615DFD9A07ECFD39" author="Scott, A." box="[1306,1431,620,645]" pageId="83" pageNumber="84" pagination="397 - 428" refId="ref49985" refString="Scott, A. (1902) On some Red Sea and Indian Ocean Copepoda. Proceedings and Transactions of the Liverpool Biological Society, 16, 397 - 428." type="journal article" year="1902">Scott 1902</bibRefCitation>
;
<bibRefCitation id="84401887FFFDFF8064D0FD790354FD14" author="Gurney, R." box="[151,303,655,680]" pageId="83" pageNumber="84" pagination="451 - 577" refId="ref47728" refString="Gurney, R. (1927) Report on the Crustacea. Copepoda (littoral and semiparasitic); Zoological Results of the Cambridge Expedition to the Suez Canal, 1924, no. 33. Transactions of the Zoological Society of London, 22, 451 - 577. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1927. tb 00207. x" type="journal article" year="1927">Gurney 1927</bibRefCitation>
;
<bibRefCitation id="84401887FFFDFF80657AFD6603C0FD14" author="Lang, K." box="[317,443,655,680]" pageId="83" pageNumber="84" refId="ref49308" refString="Lang, K. (1948) Monographie der Harpacticiden, 1 - 2. Nordiska Bokhandeln, Lund, 1682 pp." type="book" year="1948">Lang 1948</bibRefCitation>
; Por 1964;
<bibRefCitation id="84401887FFFDFF806605FD7900BDFD14" author="Wells, J. B. J." box="[578,710,655,680]" pageId="83" pageNumber="84" refId="ref50213" refString="Wells, J. B. J. (1967) The littoral Copepoda (Crustacea) of Inhaca Island, Mozambique. Transactions of the Royal Society of Edinburgh, 67,189 - 358." type="book" year="1967">Wells 1967</bibRefCitation>
;
<bibRefCitation id="84401887FFFDFF806693FD79066DFD14" author="Marinov, T. M. &amp; Apostolov, A. M." box="[724,1046,655,680]" pageId="83" pageNumber="84" pagination="66 - 72" refId="ref49723" refString="Marinov, T. M. &amp; Apostolov, A. M. (1981) Contribution a l'etude des Copepodes Harpacticoides de la mer Adriatique (cote yougoslave), I. Le genre Stenhelia Boeck. Acta Zoologica Bulgarica, 17, 66 - 72." type="journal article" year="1981">Marinov &amp; Apostolov 1981</bibRefCitation>
). The males are still unknown for
<taxonomicName id="27D11EF5FFFDFF8064D0FD4203E0FD70" authorityName="T. Scott" authorityYear="1902" box="[151,411,692,716]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="D2A5B964FFFDFF8064D0FD4203E0FD70" box="[151,411,692,716]" italics="true" pageId="83" pageNumber="84">Willenstenhelia minuta</emphasis>
</taxonomicName>
and
<taxonomicName id="27D11EF5FFFDFF806592FD4200ADFD70" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[469,726,692,716]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="D2A5B964FFFDFF806592FD4200ADFD70" box="[469,726,692,716]" italics="true" pageId="83" pageNumber="84">Willenstenhelia urania</emphasis>
</taxonomicName>
, and those for
<taxonomicName id="27D11EF5FFFDFF8067C1FD4206C1FD70" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[902,1210,692,716]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="terpsichore">
<emphasis id="D2A5B964FFFDFF8067C1FD4206C1FD70" box="[902,1210,692,716]" italics="true" pageId="83" pageNumber="84">Willenstenhelia terpsichore</emphasis>
</taxonomicName>
are described quite incompletely. However, the latter species apparently has both exopodal and endopodal spines (or spiniform setae?) on the male fifth leg fused basally, forming strong spiniform processes, which is very different from the male fifth legs of
<taxonomicName id="27D11EF5FFFDFF8064AFFCD603A6FC84" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[232,477,800,824]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="thalia">
<emphasis id="D2A5B964FFFDFF8064AFFCD603A6FC84" box="[232,477,800,824]" italics="true" pageId="83" pageNumber="84">Willenstenhelia thalia</emphasis>
</taxonomicName>
and
<emphasis id="D2A5B964FFFDFF806655FCD6014DFC8B" box="[530,822,800,824]" italics="true" pageId="83" pageNumber="84">
<taxonomicName id="27D11EF5FFFDFF806655FCD60148FC8B" authorityName="Karanovic &amp; Kim" authorityYear="2014" baseAuthorityName="Wells" baseAuthorityYear="1967" box="[530,819,800,824]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="unisetosa">Willenstenhelia unisetosa</taxonomicName>
.
</emphasis>
Similarities in the male fifth leg between the latter two species, in addition to an identical armature formula of both male and female swimming legs, and a very similar anal operculum, may suggest that they are more closely related to each other than to other congeners, while the other three species seem to form a relatively compact group, at least as one can judge from a limited set of morphological characters given in their original reports.
</paragraph>
<paragraph id="E06E6576FFFDFF806480FC2201AFFB79" blockId="83.[151,1436,151,1400]" pageId="83" pageNumber="84">
<taxonomicName id="27D11EF5FFFDFF806480FC2203C5FC50" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[199,446,980,1004]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="thalia">
<emphasis id="D2A5B964FFFDFF806480FC2203C5FC50" box="[199,446,980,1004]" italics="true" pageId="83" pageNumber="84">Willenstenhelia thalia</emphasis>
</taxonomicName>
differs from
<taxonomicName id="27D11EF5FFFDFF80661FFC220103FC50" authorityName="Karanovic &amp; Kim" authorityYear="2014" baseAuthorityName="Wells" baseAuthorityYear="1967" box="[600,888,980,1004]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="unisetosa">
<emphasis id="D2A5B964FFFDFF80661FFC220103FC50" box="[600,888,980,1004]" italics="true" pageId="83" pageNumber="84">Willenstenhelia unisetosa</emphasis>
</taxonomicName>
additionally by much longer caudal rami, more slender habitus, shorter and more spiniform innermost seta on the female fifth leg endopod, larger and more deeply concave space between two inner setae on the female fifth leg endopod, much shorter outer endopodal element on the male fifth leg endopod, and much longer second seta from inner side on the male fifth leg exopod.
<bibRefCitation id="84401887FFFDFF80614BFBC907E7FBE4" author="Wells, J. B. J." box="[1292,1436,1087,1112]" pageId="83" pageNumber="84" refId="ref50213" refString="Wells, J. B. J. (1967) The littoral Copepoda (Crustacea) of Inhaca Island, Mozambique. Transactions of the Royal Society of Edinburgh, 67,189 - 358." type="book" year="1967">Wells (1967)</bibRefCitation>
also reported only three endopodal setae on the maxilliped and only one setae on the mandibular basis, but this may be because some setae may have been broken off on his specimen, as all other well-studied stenheliids have four endopodal setae on the maxilliped and three basal setae on the mandibula.
</paragraph>
<paragraph id="E06E6576FFFDFF806480FB2601D2FACB" blockId="83.[151,1436,151,1400]" pageId="83" pageNumber="84">
<taxonomicName id="27D11EF5FFFDFF806480FB2603BEFB54" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[199,453,1232,1256]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="thalia">
<emphasis id="D2A5B964FFFDFF806480FB2603BEFB54" box="[199,453,1232,1256]" italics="true" pageId="83" pageNumber="84">Willenstenhelia thalia</emphasis>
</taxonomicName>
differs from
<emphasis id="D2A5B964FFFDFF806632FB2606DDFB5B" box="[629,1190,1232,1256]" italics="true" pageId="83" pageNumber="84">
<taxonomicName id="27D11EF5FFFDFF806632FB2601F8FB5B" authorityName="T. Scott" authorityYear="1902" box="[629,899,1232,1256]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="minuta">Willenstenhelia minuta</taxonomicName>
,
<taxonomicName id="27D11EF5FFFDFF8067D0FB2606D9FB5B" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[919,1186,1232,1256]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="urania">Willenstenhelia urania</taxonomicName>
,
</emphasis>
and
<taxonomicName id="27D11EF5FFFDFF8060A8FB26036DFAB0" authorityName="Karanovic &amp; Kim" authorityYear="2014" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="terpsichore">
<emphasis id="D2A5B964FFFDFF8060A8FB26036DFAB0" italics="true" pageId="83" pageNumber="84">Willenstenhelia terpsichore</emphasis>
</taxonomicName>
additionally by the spiniform nature of the innermost element on the female fifth leg endopod, shorter second seta from outer side on the female fifth leg exopod, and presence of long spinules on the female fifth leg exopod. Its caudal rami are also significantly longer than those in
<taxonomicName id="27D11EF5FFFDFF8067EDFACA06CBFAE8" authorityName="T. Scott" authorityYear="1902" box="[938,1200,1340,1364]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="D2A5B964FFFDFF8067EDFACA06CBFAE8" box="[938,1200,1340,1364]" italics="true" pageId="83" pageNumber="84">Willenstenhelia minuta</emphasis>
</taxonomicName>
and
<taxonomicName id="27D11EF5FFFDFF8060A8FACA036EFAC4" authorityName="Karanovic &amp; Kim" authorityYear="2014" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="terpsichore">
<emphasis id="D2A5B964FFFDFF8060A8FACA036EFAC4" italics="true" pageId="83" pageNumber="84">Willenstenhelia terpsichore</emphasis>
</taxonomicName>
, but comparable in size to those in
<emphasis id="D2A5B964FFFDFF8066E4FA9601D2FACB" box="[675,937,1376,1400]" italics="true" pageId="83" pageNumber="84">
<taxonomicName id="27D11EF5FFFDFF8066E4FA9601DFFACB" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[675,932,1376,1400]" class="Maxillopoda" family="Miraciidae" genus="Willenstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="urania">Willenstenhelia urania</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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