treatments-xml/data/68/78/D4/6878D460FFB9FFCD64D0FE700115FF69.xml

<document id="894FD3C2C2207D74199DA1DE14525CBF" ID-CLB-Dataset="5789" ID-DOI="10.11646/zootaxa.3783.1.1" ID-GBIF-Dataset="414491b2-5167-4605-95ab-f679bf476618" ID-ISSN="1175-5326" ID-Zenodo-Dep="4910562" ID-ZooBank="E6155BDC-AEAE-475D-BC83-61B3B863344C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623122599895" checkinUser="felipe" docAuthor="Karanovic, Tomislav &amp; Kim, Kichoon" docDate="2014" docId="6878D460FFB9FFCD64D0FE700115FF69" docLanguage="en" docName="zt03783p096.pdf" docOrigin="Zootaxa 3783 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Wellstenhelia clio Karanovic &amp; Kim 2014, sp. nov." docType="treatment" docVersion="7" lastPageNumber="30" masterDocId="9441AC18FFAEFFD36447FFF6027BFFBC" masterDocTitle="New insights into polyphyly of the harpacticoid genus Delavalia (Crustacea, Copepoda) through morphological and molecular study of an unprecedented diversity of sympatric species in a small South Korean bay" masterLastPageNumber="96" masterPageNumber="1" pageNumber="24" updateTime="1699004038958" updateUser="ExternalLinkService">
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<mods:title id="57F283DEEC20C9DE15A7DFB0CD2C3F5C">New insights into polyphyly of the harpacticoid genus Delavalia (Crustacea, Copepoda) through morphological and molecular study of an unprecedented diversity of sympatric species in a small South Korean bay</mods:title>
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<mods:namePart id="22D882F16B2F65DC707CB5EF59B3D074">Karanovic, Tomislav</mods:namePart>
<mods:nameIdentifier id="57FE98463342497A97045D883587DA08" type="email">aranovic@utas.edu.au</mods:nameIdentifier>
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<mods:namePart id="B1BF93F258B312B355AC4DEEB22799E2">Kim, Kichoon</mods:namePart>
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<mods:date id="F8CD44AA13BC7D55596FCB78CC9A32A3">2014</mods:date>
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<mods:number id="B35BF476A8DA1E8485153E4C9C8401BF">3783</mods:number>
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<treatment id="6878D460FFB9FFCD64D0FE700115FF69" ID-DOI="http://doi.org/10.5281/zenodo.5062483" ID-GBIF-Taxon="183238045" ID-Zenodo-Dep="5062483" LSID="urn:lsid:plazi:treatment:6878D460FFB9FFCD64D0FE700115FF69" httpUri="http://treatment.plazi.org/id/6878D460FFB9FFCD64D0FE700115FF69" lastPageId="30" lastPageNumber="30" pageId="23" pageNumber="24">
<subSubSection id="A8CB36FDFFB9FFC464D0FE7003AAFE1C" box="[151,465,390,416]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="E06E6576FFB9FFC464D0FE7003AAFE1C" blockId="23.[151,465,390,450]" box="[151,465,390,416]" pageId="23" pageNumber="24">
<heading id="BB26D21AFFB9FFC464D0FE7003AAFE1C" bold="true" box="[151,465,390,416]" fontSize="11" level="2" pageId="23" pageNumber="24" reason="4">
<emphasis id="D2A5B964FFB9FFC464D0FE7003AAFE1C" bold="true" box="[151,465,390,416]" pageId="23" pageNumber="24">
<taxonomicName id="27D11EF5FFB9FFC464D0FE700317FE1C" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[151,364,390,416]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="clio" status="sp. nov.">
<emphasis id="D2A5B964FFB9FFC464D0FE700317FE1C" bold="true" box="[151,364,390,416]" italics="true" pageId="23" pageNumber="24">Wellstenhelia clio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C996041FFFB9FFC46533FE7003AAFE1C" box="[372,465,390,416]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="A8CB36FDFFB9FFC464D0FE5F0356FE7E" box="[151,301,425,450]" pageId="23" pageNumber="24" type="description">
<paragraph id="E06E6576FFB9FFC464D0FE5F0356FE7E" blockId="23.[151,465,390,450]" box="[151,301,425,450]" pageId="23" pageNumber="24">
(
<figureCitation id="78EA79F3FFB9FFC464D8FE5F035EFE7E" box="[159,293,425,450]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStart-2="FIGURE 14" captionStart-3="FIGURE 15" captionStart-4="FIGURE 16" captionStart-5="FIGURE 17" captionStartId-0="24.[151,249,1916,1938]" captionStartId-1="25.[151,249,1906,1928]" captionStartId-2="26.[151,249,1890,1912]" captionStartId-3="27.[151,249,1921,1943]" captionStartId-4="28.[151,249,1573,1595]" captionStartId-5="30.[151,249,1630,1652]" captionTargetBox-0="[190,1408,189,1892]" captionTargetBox-1="[198,1393,208,1882]" captionTargetBox-2="[199,1389,186,1864]" captionTargetBox-3="[195,1396,205,1892]" captionTargetBox-4="[151,1436,193,1534]" captionTargetBox-5="[151,1436,267,1608]" captionTargetId-0="figure-89@24.[185,1408,171,1899]" captionTargetId-1="figure-79@25.[190,1397,193,1885]" captionTargetId-2="figure-116@26.[186,1400,175,1868]" captionTargetId-3="figure-74@27.[186,1401,193,1900]" captionTargetId-4="figure-209@28.[151,1436,193,1534]" captionTargetId-5="figure-154@30.[151,1436,267,1608]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionTargetPageId-2="26" captionTargetPageId-3="27" captionTargetPageId-4="28" captionTargetPageId-5="30" captionText-0="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." captionText-1="FIGURE 13. Wellstenhelia clio sp. nov., line drawings, female holotype: A, abdomen, ventral view; B, left side of cephalothoracic shield, dissected and flattened; C pleuron of third pedigerous somite, dissected and flattened; D, mandibular palp, anterior view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads pointing most prominent specific features. Scale bars 100 µm." captionText-2="FIGURE 14. Wellstenhelia clio sp. nov., line drawings, A–D, female holotype, E–F, male allotype: A, pleuron of fourth pedigerous somite, dissected and flattened; B, fourth swimming leg without rami, anterior view; C, fifth leg, dissected and flattened, anterior view; D, sixth leg on genital somite, ventro-lateral view; E, cephalothoracic shield, lateral view; F, antennula, dorsal view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, €) indicating unique sensilla and pores. Arrowheads pointing most prominent specific features. Scale bars 100 µm." captionText-3="FIGURE 15. Wellstenhelia clio sp. nov., line drawings, male allotype: A, urosome, ventral view; B, urosome, lateral view; C, pleuron of second pedigerous somite, dissected and flattened. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads point to most prominent specific features. Scale bars 100 µm." captionText-4="FIGURE 16. Wellstenhelia clio sp. nov., scanning electron micrographs, female paratype: A, habitus, dorsal view; B, rostrum, dorsal view; C, cephalothorax, dorsal view; D, pedigerous somites, dorsal view; E, first urosomite and genital double-somite, dorsal view; F, anal somite and caudal rami, dorsal view. Scale bars 40 µm (B), 50 µm (D, E, F), 100 µm (C), and 200 µm (A)." captionText-5="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi-0="http://doi.org/10.5281/zenodo.4910592" figureDoi-1="http://doi.org/10.5281/zenodo.4910594" figureDoi-2="http://doi.org/10.5281/zenodo.4910596" figureDoi-3="http://doi.org/10.5281/zenodo.4910598" figureDoi-4="http://doi.org/10.5281/zenodo.4910600" figureDoi-5="http://doi.org/10.5281/zenodo.4910602" httpUri-0="https://zenodo.org/record/4910592/files/figure.png" httpUri-1="https://zenodo.org/record/4910594/files/figure.png" httpUri-2="https://zenodo.org/record/4910596/files/figure.png" httpUri-3="https://zenodo.org/record/4910598/files/figure.png" httpUri-4="https://zenodo.org/record/4910600/files/figure.png" httpUri-5="https://zenodo.org/record/4910602/files/figure.png" pageId="23" pageNumber="24">Figs. 12–17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A8CB36FDFFB9FFC464D0FE070358FBBE" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="E06E6576FFB9FFC464D0FE0703E1FD92" blockId="23.[151,1437,497,2034]" pageId="23" pageNumber="24">
<materialsCitation id="50B96F2BFFB9FFC464D0FE0703EEFD92" ID-GBIF-Occurrence="3313373320" country="South Korea" latitude="34.920944" location="South Sea" longLatPrecision="1" longitude="127.78553" municipality="Gwangyang Bay" pageId="23" pageNumber="24" specimenCount="1" typeStatus="holotype">
<emphasis id="D2A5B964FFB9FFC464D0FE070342FDB6" bold="true" box="[151,313,497,522]" pageId="23" pageNumber="24">
<typeStatus id="3F6ADBD4FFB9FFC464D0FE0702A9FDB6" box="[151,210,497,522]" pageId="23" pageNumber="24">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="98C625E6FFB9FFC46504FE0703AEFDB5" box="[323,469,497,522]" name="South Korea" pageId="23" pageNumber="24">South Korea</collectingCountry>
, 
<location id="E50E33ADFFB9FFC465A3FE070020FDB6" LSID="urn:lsid:plazi:treatment:6878D460FFB9FFCD64D0FE700115FF69:E50E33ADFFB9FFC465A3FE070020FDB6" box="[484,603,497,522]" country="South Korea" latitude="34.920944" longLatPrecision="1" longitude="127.78553" municipality="Gwangyang Bay" name="South Sea" pageId="23" pageNumber="24">South Sea</location>
, 
<collectingMunicipality id="000AFF0CFFB9FFC4662DFE070152FDB6" box="[618,809,497,522]" pageId="23" pageNumber="24">Gwangyang Bay</collectingMunicipality>
, sampling station 10, muddy sediments, 
<geoCoordinate id="85E503B1FFB9FFC46142FE0707E7FDB6" box="[1285,1436,497,522]" degrees="34.920944" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="34.920944">34.920944°N</geoCoordinate>
<geoCoordinate id="85E503B1FFB9FFC464D0FDE30343FD92" box="[151,312,533,558]" degrees="127.785528" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="127.78553">127.785528°E</geoCoordinate>
(
<figureCitation id="78EA79F3FFB9FFC46500FDE303F2FD92" box="[327,393,533,558]" captionStart="FIGURE 1" captionStartId="7.[151,249,1481,1503]" captionTargetBox="[151,1436,589,1460]" captionTargetId="figure-399@7.[151,1436,589,1460]" captionTargetPageId="7" captionText="FIGURE 1. Map of 17 sampling locations in Gwangyang Bay. Inset showing location of the bay in South Korea. All maps from Google Earth." figureDoi="http://doi.org/10.5281/zenodo.4910564" httpUri="https://zenodo.org/record/4910564/files/figure.png" pageId="23" pageNumber="24">Fig. 1</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="E06E6576FFB9FFC46480FDCF0752FD5E" blockId="23.[151,1437,497,2034]" pageId="23" pageNumber="24">
<emphasis id="D2A5B964FFB9FFC46480FDCF03BDFDEE" bold="true" box="[199,454,569,594]" pageId="23" pageNumber="24">Specimens examined.</emphasis>
Female 
<typeStatus id="3F6ADBD4FFB9FFC46662FDCF00FCFDEE" box="[549,647,569,594]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
dissected on one slide (collection number NIBRIV0000232676), male 
<typeStatus id="3F6ADBD4FFB9FFC464D0FDAB028AFDCA" box="[151,241,605,630]" pageId="23" pageNumber="24" type="allotype">allotype</typeStatus>
dissected on one slide (collection number NIBRIV0000232677), 
<specimenCount id="F6D7AEFFFFB9FFC467A8FDA80621FDCA" box="[1007,1114,605,630]" pageId="23" pageNumber="24" type="male">two male</specimenCount>
<typeStatus id="3F6ADBD4FFB9FFC46022FDA806ABFDCA" box="[1125,1232,606,630]" pageId="23" pageNumber="24" type="paratype">paratypes</typeStatus>
and 
<specimenCount id="F6D7AEFFFFB9FFC4614AFDAB07E7FDCA" box="[1293,1436,605,630]" pageId="23" pageNumber="24" type="female">three female</specimenCount>
<typeStatus id="3F6ADBD4FFB9FFC464D0FD740379FD26" box="[151,258,642,666]" pageId="23" pageNumber="24" type="paratype">paratypes</typeStatus>
together on one SEM stub (collection number NIBRIV0000232678), 
<specimenCount id="F6D7AEFFFFB9FFC4604AFD770610FD26" box="[1037,1131,641,666]" pageId="23" pageNumber="24" type="male">six male</specimenCount>
<typeStatus id="3F6ADBD4FFB9FFC46036FD7406A6FD26" box="[1137,1245,642,666]" pageId="23" pageNumber="24" type="paratype">paratypes</typeStatus>
and 
<specimenCount id="F6D7AEFFFFB9FFC46156FD7707E7FD26" box="[1297,1436,641,666]" pageId="23" pageNumber="24" type="female">eight female</specimenCount>
<typeStatus id="3F6ADBD4FFB9FFC464D0FD500379FD02" box="[151,258,678,702]" pageId="23" pageNumber="24" type="paratype">paratypes</typeStatus>
and six copepodid 
<typeStatus id="3F6ADBD4FFB9FFC465A7FD500030FD02" box="[480,587,678,702]" pageId="23" pageNumber="24" type="paratype">paratypes</typeStatus>
together in ethanol (collection number NIBRIV0000232679), 
<specimenCount id="F6D7AEFFFFB9FFC46156FD5307E7FD02" box="[1297,1436,677,702]" pageId="23" pageNumber="24" type="female">five females</specimenCount>
destroyed for DNA sequence (amplification unsuccessful); type locality, 
<date id="946F43B6FFB9FFC4678FFD3F06E9FD5E" box="[968,1170,713,738]" pageId="23" pageNumber="24" value="2012-02-18">18 February 2012</date>
, leg. K. Kim.
</paragraph>
<paragraph id="E06E6576FFB9FFC46480FD180022FCF2" blockId="23.[151,1437,497,2034]" pageId="23" pageNumber="24">
Two male 
<typeStatus id="3F6ADBD4FFB9FFC46505FD1803D6FCBA" box="[322,429,750,774]" pageId="23" pageNumber="24" type="paratype">paratypes</typeStatus>
and 
<specimenCount id="F6D7AEFFFFB9FFC465AEFD1B000CFCBA" box="[489,631,749,774]" pageId="23" pageNumber="24" type="female">three female</specimenCount>
<typeStatus id="3F6ADBD4FFB9FFC466C6FD180097FCBA" box="[641,748,750,774]" pageId="23" pageNumber="24" type="paratype">paratypes</typeStatus>
and one copepodid 
<typeStatus id="3F6ADBD4FFB9FFC4679EFD180641FCBA" box="[985,1082,750,774]" pageId="23" pageNumber="24" type="paratype">paratype</typeStatus>
together in ethanol (collection number NIBRIV0000232680), 
<specimenCount id="F6D7AEFFFFB9FFC465B9FCE40007FC96" box="[510,636,785,810]" pageId="23" pageNumber="24" type="female">one female</specimenCount>
destroyed for DNA sequence (amplification successful, Code 0187), type locality, 
<date id="946F43B6FFB9FFC46569FCC303B9FCF2" box="[302,450,821,846]" pageId="23" pageNumber="24" value="2012-07-30">30 July 2012</date>
, leg. K. Kim.
</paragraph>
<paragraph id="E06E6576FFB9FFC46480FCAF0358FBBE" blockId="23.[151,1437,497,2034]" pageId="23" pageNumber="24">
<materialsCitation id="50B96F2BFFB9FFC46480FCAF02A3FC06" ID-GBIF-Occurrence="3313373321" collectingDate="2012-11-18" collectorName="K. Kim. Three" country="South Korea" county="Gwangyang Bay" latitude="34.89014" location="One" longLatPrecision="1" longitude="127.79511" municipality="South Sea" pageId="23" pageNumber="24" specimenCount="1">
<location id="E50E33ADFFB9FFC46480FCAF028DFCCE" LSID="urn:lsid:plazi:treatment:6878D460FFB9FFCD64D0FE700115FF69:E50E33ADFFB9FFC46480FCAF028DFCCE" box="[199,246,857,882]" country="South Korea" county="Gwangyang Bay" latitude="34.89014" longLatPrecision="1" longitude="127.79511" municipality="South Sea" name="One" pageId="23" pageNumber="24">One</location>
female destroyed for DNA sequence (amplification unsuccessful), 
<collectingCountry id="98C625E6FFB9FFC467B2FCAF06FDFCCD" box="[1013,1158,857,882]" name="South Korea" pageId="23" pageNumber="24">South Korea</collectingCountry>
, 
<collectingMunicipality id="000AFF0CFFB9FFC460D4FCAF077CFCCE" box="[1171,1287,857,882]" pageId="23" pageNumber="24">South Sea</collectingMunicipality>
, 
<collectingCounty id="090F1DFAFFB9FFC46153FCAF02BEFC2A" pageId="23" pageNumber="24">Gwangyang Bay</collectingCounty>
, sampling station 15, muddy sediments, 
<geoCoordinate id="85E503B1FFB9FFC466CEFC8B015BFC2A" box="[649,800,893,918]" degrees="34.890139" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="34.89014">34.890139°N</geoCoordinate>
<geoCoordinate id="85E503B1FFB9FFC46761FC8B01BEFC2A" box="[806,965,893,918]" degrees="127.795111" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="127.79511">127.795111°E</geoCoordinate>
(
<figureCitation id="78EA79F3FFB9FFC46793FC8B066EFC2A" box="[980,1045,893,918]" captionStart="FIGURE 1" captionStartId="7.[151,249,1481,1503]" captionTargetBox="[151,1436,589,1460]" captionTargetId="figure-399@7.[151,1436,589,1460]" captionTargetPageId="7" captionText="FIGURE 1. Map of 17 sampling locations in Gwangyang Bay. Inset showing location of the bay in South Korea. All maps from Google Earth." figureDoi="http://doi.org/10.5281/zenodo.4910564" httpUri="https://zenodo.org/record/4910564/files/figure.png" pageId="23" pageNumber="24">Fig. 1</figureCitation>
), 
<date id="946F43B6FFB9FFC4606AFC8B077CFC2A" box="[1069,1287,893,918]" pageId="23" pageNumber="24" value="2012-11-18">
<collectingDate id="842BBA5EFFB9FFC4606AFC8B077CFC2A" box="[1069,1287,893,918]" pageId="23" pageNumber="24" value="2012-11-18">18 November 2012</collectingDate>
</date>
, leg. 
<collectorName id="4D2400A0FFB9FFC46106FC8802A3FC06" pageId="23" pageNumber="24">K. Kim. Three</collectorName>
</materialsCitation>
<materialsCitation id="50B96F2BFFB9FFC46498FC570358FBBE" ID-GBIF-Occurrence="3313373305" collectingDate="2012-11-18" collectorName="K. Kim." country="South Korea" latitude="34.95139" location="South Sea" longLatPrecision="1" longitude="127.73436" municipality="Gwangyang Bay" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1">
males and 
<specimenCount id="F6D7AEFFFFB9FFC46511FC570397FC06" box="[342,492,929,954]" pageId="23" pageNumber="24" type="female">three females</specimenCount>
destroyed for DNA sequence (amplification unsuccessful), 
<collectingCountry id="98C625E6FFB9FFC460CDFC570762FC05" box="[1162,1305,929,954]" name="South Korea" pageId="23" pageNumber="24">South Korea</collectingCountry>
, 
<location id="E50E33ADFFB9FFC46163FC5707ECFC06" LSID="urn:lsid:plazi:treatment:6878D460FFB9FFCD64D0FE700115FF69:E50E33ADFFB9FFC46163FC5707ECFC06" box="[1316,1431,929,954]" country="South Korea" latitude="34.95139" longLatPrecision="1" longitude="127.73436" municipality="Gwangyang Bay" name="South Sea" pageId="23" pageNumber="24">South Sea</location>
, 
<collectingMunicipality id="000AFF0CFFB9FFC464D0FC330328FC62" box="[151,339,965,990]" pageId="23" pageNumber="24">Gwangyang Bay</collectingMunicipality>
, sampling station 12, muddy sediments, 
<geoCoordinate id="85E503B1FFB9FFC4675FFC3301D5FC62" box="[792,942,965,990]" degrees="34.951389" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="34.95139">34.951389°N</geoCoordinate>
<geoCoordinate id="85E503B1FFB9FFC467F2FC33062EFC62" box="[949,1109,965,990]" degrees="127.734361" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="127.73436">127.734361°E</geoCoordinate>
(
<figureCitation id="78EA79F3FFB9FFC46022FC3306DDFC62" box="[1125,1190,965,990]" captionStart="FIGURE 1" captionStartId="7.[151,249,1481,1503]" captionTargetBox="[151,1436,589,1460]" captionTargetId="figure-399@7.[151,1436,589,1460]" captionTargetPageId="7" captionText="FIGURE 1. Map of 17 sampling locations in Gwangyang Bay. Inset showing location of the bay in South Korea. All maps from Google Earth." figureDoi="http://doi.org/10.5281/zenodo.4910564" httpUri="https://zenodo.org/record/4910564/files/figure.png" pageId="23" pageNumber="24">Fig. 1</figureCitation>
), 
<date id="946F43B6FFB9FFC460F9FC3307E3FC62" box="[1214,1432,965,990]" pageId="23" pageNumber="24" value="2012-11-18">
<collectingDate id="842BBA5EFFB9FFC460F9FC3307E3FC62" box="[1214,1432,965,990]" pageId="23" pageNumber="24" value="2012-11-18">18 November 2012</collectingDate>
</date>
, leg. 
<collectorName id="4D2400A0FFB9FFC46480FC1C0358FBBE" box="[199,291,1001,1026]" pageId="23" pageNumber="24">K. Kim.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A8CB36FDFFB9FFC46480FBFB0733FBF6" pageId="23" pageNumber="24" type="etymology">
<paragraph id="E06E6576FFB9FFC46480FBFB0733FBF6" blockId="23.[151,1437,497,2034]" pageId="23" pageNumber="24">
<emphasis id="D2A5B964FFB9FFC46480FBFB0335FB9A" bold="true" box="[199,334,1037,1062]" pageId="23" pageNumber="24">Etymology.</emphasis>
The species is named after Clio (Ancient Greek: Κλειώ), one of nine Muses from Greek mythology, who was a patron of history. The species name is a noun in apposition (in the nominative case).
</paragraph>
</subSubSection>
<subSubSection id="A8CB36FDFFB9FFCD6480FBA30115FF69" lastPageId="30" lastPageNumber="31" pageId="23" pageNumber="24" type="description">
<paragraph id="E06E6576FFB9FFC46480FBA306CCFAFA" blockId="23.[151,1437,497,2034]" pageId="23" pageNumber="24">
<emphasis id="D2A5B964FFB9FFC46480FBA30322FBD2" bold="true" box="[199,345,1109,1134]" pageId="23" pageNumber="24">Description.</emphasis>
Female (based on 
<typeStatus id="3F6ADBD4FFB9FFC46678FBA300DAFBD2" box="[575,673,1109,1134]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
and 
<specimenCount id="F6D7AEFFFFB9FFC466A6FBA001FDFBD2" box="[737,902,1109,1134]" pageId="23" pageNumber="24" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
). Body length from 
<quantity id="2729C893FFB9FFC4603DFBA30757FBD2" box="[1146,1324,1109,1134]" metricMagnitude="-1" metricUnit="m" metricValue="5.785" metricValueMax="6.17" metricValueMin="5.4" pageId="23" pageNumber="24" unit="mm" value="578.5" valueMax="617.0" valueMin="540.0">540 to 617 µm</quantity>
(latter in 
<typeStatus id="3F6ADBD4FFB9FFC464D0FB8F0285FB2E" box="[151,254,1145,1170]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
). Body segmentation, colour, nauplius eye, hyaline fringes, integument thickness and surface appearence as in 
<taxonomicName id="27D11EF5FFB9FFC4649AFB6B03A2FB09" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[221,473,1181,1205]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="calliope" status="sp. nov.">
<emphasis id="D2A5B964FFB9FFC4649AFB6B03A2FB09" box="[221,473,1181,1205]" italics="true" pageId="23" pageNumber="24">Wellstenhelia calliope</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFB9FFC465A1FB6B003EFB0A" bold="true" box="[486,581,1181,1206]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="C996041FFFB9FFC465A1FB6B003EFB0A" box="[486,581,1181,1206]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, including minute sparse pits visible only on highest magnifications on scanning electron microscope. Most somite ornamentation also similar to 
<taxonomicName id="27D11EF5FFB9FFC467B7FB370690FB65" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1008,1259,1217,1241]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFB9FFC467B7FB370690FB65" box="[1008,1259,1217,1241]" italics="true" pageId="23" pageNumber="24">Wellstenhelia calliope</emphasis>
</taxonomicName>
, and presumed homologous pores and sensilla numbered with same Arabic numerals (see 
<figureCitation id="78EA79F3FFB9FFC467B3FB1306D5FB42" box="[1012,1198,1253,1278]" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="23" pageNumber="24">Figs. 12A, B, C</figureCitation>
, 
<figureCitation id="78EA79F3FFB9FFC460FBFB13074EFB42" box="[1212,1333,1253,1278]" captionStart="FIGURE 13" captionStartId="25.[151,249,1906,1928]" captionTargetBox="[198,1393,208,1882]" captionTargetId="figure-79@25.[190,1397,193,1885]" captionTargetPageId="25" captionText="FIGURE 13. Wellstenhelia clio sp. nov., line drawings, female holotype: A, abdomen, ventral view; B, left side of cephalothoracic shield, dissected and flattened; C pleuron of third pedigerous somite, dissected and flattened; D, mandibular palp, anterior view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910594" httpUri="https://zenodo.org/record/4910594/files/figure.png" pageId="23" pageNumber="24">13A, B, C</figureCitation>
, 
<figureCitation id="78EA79F3FFB9FFC46104FB13070EFB42" box="[1347,1397,1253,1278]" captionStart="FIGURE 14" captionStartId="26.[151,249,1890,1912]" captionTargetBox="[199,1389,186,1864]" captionTargetId="figure-116@26.[186,1400,175,1868]" captionTargetPageId="26" captionText="FIGURE 14. Wellstenhelia clio sp. nov., line drawings, A–D, female holotype, E–F, male allotype: A, pleuron of fourth pedigerous somite, dissected and flattened; B, fourth swimming leg without rami, anterior view; C, fifth leg, dissected and flattened, anterior view; D, sixth leg on genital somite, ventro-lateral view; E, cephalothoracic shield, lateral view; F, antennula, dorsal view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, €) indicating unique sensilla and pores. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910596" httpUri="https://zenodo.org/record/4910596/files/figure.png" pageId="23" pageNumber="24">14A</figureCitation>
) to allow easier comparison. Habitus (
<figureCitation id="78EA79F3FFB9FFC46666FAFF00CDFA9D" box="[545,694,1289,1314]" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="23" pageNumber="24">Figs. 12A, B</figureCitation>
, 
<figureCitation id="78EA79F3FFB9FFC46684FAFF008FFA9E" box="[707,756,1289,1314]" captionStart="FIGURE 16" captionStartId="28.[151,249,1573,1595]" captionTargetBox="[151,1436,193,1534]" captionTargetId="figure-209@28.[151,1436,193,1534]" captionTargetPageId="28" captionText="FIGURE 16. Wellstenhelia clio sp. nov., scanning electron micrographs, female paratype: A, habitus, dorsal view; B, rostrum, dorsal view; C, cephalothorax, dorsal view; D, pedigerous somites, dorsal view; E, first urosomite and genital double-somite, dorsal view; F, anal somite and caudal rami, dorsal view. Scale bars 40 µm (B), 50 µm (D, E, F), 100 µm (C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910600" httpUri="https://zenodo.org/record/4910600/files/figure.png" pageId="23" pageNumber="24">16A</figureCitation>
) more robust, with prosome/urosome length ratio 1.2, body length/width ratio about 2.9, and cephalothorax nearly twice as wide as genital double-somite.
</paragraph>
<paragraph id="E06E6576FFB9FFC46480FAA4010CFA32" blockId="23.[151,1437,497,2034]" pageId="23" pageNumber="24">
Rostrum (
<figureCitation id="78EA79F3FFB9FFC4657DFAA703D5FAD6" box="[314,430,1361,1386]" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="23" pageNumber="24">Figs. 12C</figureCitation>
, 
<figureCitation id="78EA79F3FFB9FFC465F9FAA70396FAD6" box="[446,493,1361,1386]" captionStart="FIGURE 16" captionStartId="28.[151,249,1573,1595]" captionTargetBox="[151,1436,193,1534]" captionTargetId="figure-209@28.[151,1436,193,1534]" captionTargetPageId="28" captionText="FIGURE 16. Wellstenhelia clio sp. nov., scanning electron micrographs, female paratype: A, habitus, dorsal view; B, rostrum, dorsal view; C, cephalothorax, dorsal view; D, pedigerous somites, dorsal view; E, first urosomite and genital double-somite, dorsal view; F, anal somite and caudal rami, dorsal view. Scale bars 40 µm (B), 50 µm (D, E, F), 100 µm (C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910600" httpUri="https://zenodo.org/record/4910600/files/figure.png" pageId="23" pageNumber="24">16B</figureCitation>
) slightly narrower in dorsal view than in 
<taxonomicName id="27D11EF5FFB9FFC4679DFAA706AFFAD5" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[986,1236,1361,1385]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFB9FFC4679DFAA706AFFAD5" box="[986,1236,1361,1385]" italics="true" pageId="23" pageNumber="24">Wellstenhelia calliope</emphasis>
</taxonomicName>
(arrowed in 
<figureCitation id="78EA79F3FFB9FFC46137FAA702B3FA32" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="23" pageNumber="24">Fig. 12C</figureCitation>
), but without any other difference in shape or ornamentation.
</paragraph>
<paragraph id="E06E6576FFB9FFC46480FA6F06F1F9A2" blockId="23.[151,1437,497,2034]" pageId="23" pageNumber="24">
Cephalothorax (
<figureCitation id="78EA79F3FFB9FFC46539FA6F0041FA0E" box="[382,570,1433,1458]" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="23" pageNumber="24">Figs. 12A, B, C</figureCitation>
, 
<figureCitation id="78EA79F3FFB9FFC4660EFA6F0001FA0E" box="[585,634,1433,1458]" captionStart="FIGURE 13" captionStartId="25.[151,249,1906,1928]" captionTargetBox="[198,1393,208,1882]" captionTargetId="figure-79@25.[190,1397,193,1885]" captionTargetPageId="25" captionText="FIGURE 13. Wellstenhelia clio sp. nov., line drawings, female holotype: A, abdomen, ventral view; B, left side of cephalothoracic shield, dissected and flattened; C pleuron of third pedigerous somite, dissected and flattened; D, mandibular palp, anterior view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910594" httpUri="https://zenodo.org/record/4910594/files/figure.png" pageId="23" pageNumber="24">13B</figureCitation>
, 
<figureCitation id="78EA79F3FFB9FFC466CEFA6F00C2FA0E" box="[649,697,1433,1458]" captionStart="FIGURE 16" captionStartId="28.[151,249,1573,1595]" captionTargetBox="[151,1436,193,1534]" captionTargetId="figure-209@28.[151,1436,193,1534]" captionTargetPageId="28" captionText="FIGURE 16. Wellstenhelia clio sp. nov., scanning electron micrographs, female paratype: A, habitus, dorsal view; B, rostrum, dorsal view; C, cephalothorax, dorsal view; D, pedigerous somites, dorsal view; E, first urosomite and genital double-somite, dorsal view; F, anal somite and caudal rami, dorsal view. Scale bars 40 µm (B), 50 µm (D, E, F), 100 µm (C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910600" httpUri="https://zenodo.org/record/4910600/files/figure.png" pageId="23" pageNumber="24">16C</figureCitation>
) about 0.8 times as long as wide; represents 28% of total body length. Surface of cephalothoracic shield ornamented as in 
<taxonomicName id="27D11EF5FFB9FFC46778FA4B0642FA69" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[831,1081,1469,1493]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFB9FFC46778FA4B0642FA69" box="[831,1081,1469,1493]" italics="true" pageId="23" pageNumber="24">Wellstenhelia calliope</emphasis>
</taxonomicName>
; except two additional sensilla (nos. $, £) and one additional pore (no. ¥) present, two lateral pores (nos. 19, 33) missing, sensilla no. 40 paired, and somewhat different relative position of pores nos. 9, 10, 28 (arrowed in 
<figureCitation id="78EA79F3FFB9FFC467ADF9F30606F9A2" box="[1002,1149,1541,1566]" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="23" pageNumber="24">Figs. 12A, C</figureCitation>
).
</paragraph>
<paragraph id="E06E6576FFB9FFC46480F9DF0143F9DA" blockId="23.[151,1437,497,2034]" pageId="23" pageNumber="24">
Pleuron of second pedigerous somite (
<figureCitation id="78EA79F3FFB9FFC466C0F9DF015AF9FD" box="[647,801,1577,1602]" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="23" pageNumber="24">Figs. 12A, B</figureCitation>
, 
<figureCitation id="78EA79F3FFB9FFC46777F9DF011AF9FE" box="[816,865,1577,1602]" captionStart="FIGURE 16" captionStartId="28.[151,249,1573,1595]" captionTargetBox="[151,1436,193,1534]" captionTargetId="figure-209@28.[151,1436,193,1534]" captionTargetPageId="28" captionText="FIGURE 16. Wellstenhelia clio sp. nov., scanning electron micrographs, female paratype: A, habitus, dorsal view; B, rostrum, dorsal view; C, cephalothorax, dorsal view; D, pedigerous somites, dorsal view; E, first urosomite and genital double-somite, dorsal view; F, anal somite and caudal rami, dorsal view. Scale bars 40 µm (B), 50 µm (D, E, F), 100 µm (C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910600" httpUri="https://zenodo.org/record/4910600/files/figure.png" pageId="23" pageNumber="24">16D</figureCitation>
) ornamented as in 
<taxonomicName id="27D11EF5FFB9FFC46001F9DF073AF9FD" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1094,1345,1577,1601]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFB9FFC46001F9DF073AF9FD" box="[1094,1345,1577,1601]" italics="true" pageId="23" pageNumber="24">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except lateral pair of sensilla no. 48 missing (arrowed in 
<figureCitation id="78EA79F3FFB9FFC46683F9BB0150F9DA" box="[708,811,1613,1638]" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="23" pageNumber="24">Fig. 12A</figureCitation>
).
</paragraph>
<paragraph id="E06E6576FFB9FFC46480F98701B0F911" blockId="23.[151,1437,497,2034]" pageId="23" pageNumber="24">
Pleurons of third pedigerous somite (
<figureCitation id="78EA79F3FFB9FFC4662AF9870179F935" box="[621,770,1649,1674]" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="23" pageNumber="24">Figs. 12A, B</figureCitation>
, 
<figureCitation id="78EA79F3FFB9FFC46748F987013AF936" box="[783,833,1649,1674]" captionStart="FIGURE 13" captionStartId="25.[151,249,1906,1928]" captionTargetBox="[198,1393,208,1882]" captionTargetId="figure-79@25.[190,1397,193,1885]" captionTargetPageId="25" captionText="FIGURE 13. Wellstenhelia clio sp. nov., line drawings, female holotype: A, abdomen, ventral view; B, left side of cephalothoracic shield, dissected and flattened; C pleuron of third pedigerous somite, dissected and flattened; D, mandibular palp, anterior view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910594" httpUri="https://zenodo.org/record/4910594/files/figure.png" pageId="23" pageNumber="24">13C</figureCitation>
, 
<figureCitation id="78EA79F3FFB9FFC46709F98701FBF936" box="[846,896,1649,1674]" captionStart="FIGURE 16" captionStartId="28.[151,249,1573,1595]" captionTargetBox="[151,1436,193,1534]" captionTargetId="figure-209@28.[151,1436,193,1534]" captionTargetPageId="28" captionText="FIGURE 16. Wellstenhelia clio sp. nov., scanning electron micrographs, female paratype: A, habitus, dorsal view; B, rostrum, dorsal view; C, cephalothorax, dorsal view; D, pedigerous somites, dorsal view; E, first urosomite and genital double-somite, dorsal view; F, anal somite and caudal rami, dorsal view. Scale bars 40 µm (B), 50 µm (D, E, F), 100 µm (C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910600" httpUri="https://zenodo.org/record/4910600/files/figure.png" pageId="23" pageNumber="24">16D</figureCitation>
), fourth pedigerous somite (
<figureCitation id="78EA79F3FFB9FFC46085F9870723F935" box="[1218,1368,1649,1674]" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="23" pageNumber="24">Figs. 12A, B</figureCitation>
, 
<figureCitation id="78EA79F3FFB9FFC46122F98707ECF936" box="[1381,1431,1649,1674]" captionStart="FIGURE 14" captionStartId="26.[151,249,1890,1912]" captionTargetBox="[199,1389,186,1864]" captionTargetId="figure-116@26.[186,1400,175,1868]" captionTargetPageId="26" captionText="FIGURE 14. Wellstenhelia clio sp. nov., line drawings, A–D, female holotype, E–F, male allotype: A, pleuron of fourth pedigerous somite, dissected and flattened; B, fourth swimming leg without rami, anterior view; C, fifth leg, dissected and flattened, anterior view; D, sixth leg on genital somite, ventro-lateral view; E, cephalothoracic shield, lateral view; F, antennula, dorsal view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, €) indicating unique sensilla and pores. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910596" httpUri="https://zenodo.org/record/4910596/files/figure.png" pageId="23" pageNumber="24">14A</figureCitation>
, 
<figureCitation id="78EA79F3FFB9FFC464D0F96302B1F912" box="[151,202,1685,1710]" captionStart="FIGURE 16" captionStartId="28.[151,249,1573,1595]" captionTargetBox="[151,1436,193,1534]" captionTargetId="figure-209@28.[151,1436,193,1534]" captionTargetPageId="28" captionText="FIGURE 16. Wellstenhelia clio sp. nov., scanning electron micrographs, female paratype: A, habitus, dorsal view; B, rostrum, dorsal view; C, cephalothorax, dorsal view; D, pedigerous somites, dorsal view; E, first urosomite and genital double-somite, dorsal view; F, anal somite and caudal rami, dorsal view. Scale bars 40 µm (B), 50 µm (D, E, F), 100 µm (C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910600" httpUri="https://zenodo.org/record/4910600/files/figure.png" pageId="23" pageNumber="24">16D</figureCitation>
), and first urosomite (
<figureCitation id="78EA79F3FFB9FFC46586F9630032F911" box="[449,585,1685,1710]" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="23" pageNumber="24">Fig. 12A, B</figureCitation>
, 
<figureCitation id="78EA79F3FFB9FFC46613F96300F8F912" box="[596,643,1685,1710]" captionStart="FIGURE 16" captionStartId="28.[151,249,1573,1595]" captionTargetBox="[151,1436,193,1534]" captionTargetId="figure-209@28.[151,1436,193,1534]" captionTargetPageId="28" captionText="FIGURE 16. Wellstenhelia clio sp. nov., scanning electron micrographs, female paratype: A, habitus, dorsal view; B, rostrum, dorsal view; C, cephalothorax, dorsal view; D, pedigerous somites, dorsal view; E, first urosomite and genital double-somite, dorsal view; F, anal somite and caudal rami, dorsal view. Scale bars 40 µm (B), 50 µm (D, E, F), 100 µm (C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910600" httpUri="https://zenodo.org/record/4910600/files/figure.png" pageId="23" pageNumber="24">16E</figureCitation>
) as in 
<taxonomicName id="27D11EF5FFB9FFC4668AF96301BFF911" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[717,964,1685,1709]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFB9FFC4668AF96301BFF911" box="[717,964,1685,1709]" italics="true" pageId="23" pageNumber="24">Wellstenhelia calliope</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="E06E6576FFB9FFC46480F94F0180F8A6" blockId="23.[151,1437,497,2034]" pageId="23" pageNumber="24">
Genital double-somite (
<figureCitation id="78EA79F3FFB9FFC46592F94F0017F96D" box="[469,620,1721,1746]" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="23" pageNumber="24">Figs. 12A, B</figureCitation>
, 
<figureCitation id="78EA79F3FFB9FFC4663CF94F00D7F96E" box="[635,684,1721,1746]" captionStart="FIGURE 13" captionStartId="25.[151,249,1906,1928]" captionTargetBox="[198,1393,208,1882]" captionTargetId="figure-79@25.[190,1397,193,1885]" captionTargetPageId="25" captionText="FIGURE 13. Wellstenhelia clio sp. nov., line drawings, female holotype: A, abdomen, ventral view; B, left side of cephalothoracic shield, dissected and flattened; C pleuron of third pedigerous somite, dissected and flattened; D, mandibular palp, anterior view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910594" httpUri="https://zenodo.org/record/4910594/files/figure.png" pageId="23" pageNumber="24">13A</figureCitation>
, 
<figureCitation id="78EA79F3FFB9FFC466F6F94F009BF96E" box="[689,736,1721,1746]" captionStart="FIGURE 16" captionStartId="28.[151,249,1573,1595]" captionTargetBox="[151,1436,193,1534]" captionTargetId="figure-209@28.[151,1436,193,1534]" captionTargetPageId="28" captionText="FIGURE 16. Wellstenhelia clio sp. nov., scanning electron micrographs, female paratype: A, habitus, dorsal view; B, rostrum, dorsal view; C, cephalothorax, dorsal view; D, pedigerous somites, dorsal view; E, first urosomite and genital double-somite, dorsal view; F, anal somite and caudal rami, dorsal view. Scale bars 40 µm (B), 50 µm (D, E, F), 100 µm (C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910600" httpUri="https://zenodo.org/record/4910600/files/figure.png" pageId="23" pageNumber="24">16E</figureCitation>
) as in 
<taxonomicName id="27D11EF5FFB9FFC46775F94F0650F96D" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[818,1067,1721,1745]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFB9FFC46775F94F0650F96D" box="[818,1067,1721,1745]" italics="true" pageId="23" pageNumber="24">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except anterior part even more inflated laterally, forming blunt chitinous processes, ventral pair of sensilla (no. 73) much more widely spaced, and two additional rows of minute spinules in anterior half (arrowed in 
<figureCitation id="78EA79F3FFB9FFC467CFF8F70195F8A6" box="[904,1006,1793,1818]" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="23" pageNumber="24">Fig. 12B</figureCitation>
).
</paragraph>
<paragraph id="E06E6576FFB9FFC46480F8D007F3F8DE" blockId="23.[151,1437,497,2034]" pageId="23" pageNumber="24">
Last threeurosomites (
<figureCitation id="78EA79F3FFB9FFC46582F8D30026F881" box="[453,605,1829,1854]" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="23" pageNumber="24">Figs. 12A, B</figureCitation>
, 
<figureCitation id="78EA79F3FFB9FFC4662BF8D300E5F882" box="[620,670,1829,1854]" captionStart="FIGURE 13" captionStartId="25.[151,249,1906,1928]" captionTargetBox="[198,1393,208,1882]" captionTargetId="figure-79@25.[190,1397,193,1885]" captionTargetPageId="25" captionText="FIGURE 13. Wellstenhelia clio sp. nov., line drawings, female holotype: A, abdomen, ventral view; B, left side of cephalothoracic shield, dissected and flattened; C pleuron of third pedigerous somite, dissected and flattened; D, mandibular palp, anterior view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910594" httpUri="https://zenodo.org/record/4910594/files/figure.png" pageId="23" pageNumber="24">13A</figureCitation>
, 
<figureCitation id="78EA79F3FFB9FFC466EAF8D300A2F882" box="[685,729,1829,1854]" captionStart="FIGURE 16" captionStartId="28.[151,249,1573,1595]" captionTargetBox="[151,1436,193,1534]" captionTargetId="figure-209@28.[151,1436,193,1534]" captionTargetPageId="28" captionText="FIGURE 16. Wellstenhelia clio sp. nov., scanning electron micrographs, female paratype: A, habitus, dorsal view; B, rostrum, dorsal view; C, cephalothorax, dorsal view; D, pedigerous somites, dorsal view; E, first urosomite and genital double-somite, dorsal view; F, anal somite and caudal rami, dorsal view. Scale bars 40 µm (B), 50 µm (D, E, F), 100 µm (C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910600" httpUri="https://zenodo.org/record/4910600/files/figure.png" pageId="23" pageNumber="24">16F</figureCitation>
) as in 
<taxonomicName id="27D11EF5FFB9FFC4676BF8D3065EF881" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[812,1061,1829,1853]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFB9FFC4676BF8D3065EF881" box="[812,1061,1829,1853]" italics="true" pageId="23" pageNumber="24">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except for short lateral rows of spinules on preanal somite, more widely spaced sensilla nos. 74 &amp; 78, and more ventrally located pores 79 &amp; 80.
</paragraph>
<paragraph id="E06E6576FFB9FFC46480F89B074DF84E" blockId="23.[151,1437,497,2034]" pageId="23" pageNumber="24">
Caudal rami (
<figureCitation id="78EA79F3FFB9FFC4651DF89B039CF839" box="[346,487,1901,1926]" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="23" pageNumber="24">Figs. 12A, B</figureCitation>
, 
<figureCitation id="78EA79F3FFB9FFC465B4F89B005FF83A" box="[499,548,1901,1926]" captionStart="FIGURE 13" captionStartId="25.[151,249,1906,1928]" captionTargetBox="[198,1393,208,1882]" captionTargetId="figure-79@25.[190,1397,193,1885]" captionTargetPageId="25" captionText="FIGURE 13. Wellstenhelia clio sp. nov., line drawings, female holotype: A, abdomen, ventral view; B, left side of cephalothoracic shield, dissected and flattened; C pleuron of third pedigerous somite, dissected and flattened; D, mandibular palp, anterior view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910594" httpUri="https://zenodo.org/record/4910594/files/figure.png" pageId="23" pageNumber="24">13A</figureCitation>
, 
<figureCitation id="78EA79F3FFB9FFC46669F89B0021F83A" box="[558,602,1901,1926]" captionStart="FIGURE 16" captionStartId="28.[151,249,1573,1595]" captionTargetBox="[151,1436,193,1534]" captionTargetId="figure-209@28.[151,1436,193,1534]" captionTargetPageId="28" captionText="FIGURE 16. Wellstenhelia clio sp. nov., scanning electron micrographs, female paratype: A, habitus, dorsal view; B, rostrum, dorsal view; C, cephalothorax, dorsal view; D, pedigerous somites, dorsal view; E, first urosomite and genital double-somite, dorsal view; F, anal somite and caudal rami, dorsal view. Scale bars 40 µm (B), 50 µm (D, E, F), 100 µm (C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910600" httpUri="https://zenodo.org/record/4910600/files/figure.png" pageId="23" pageNumber="24">16F</figureCitation>
) short and stout, much shorter than in 
<taxonomicName id="27D11EF5FFB9FFC467BEF89B069CF839" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1017,1255,1901,1925]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFB9FFC467BEF89B069CF839" box="[1017,1255,1901,1925]" italics="true" pageId="23" pageNumber="24">Wellstenhelia calliope</emphasis>
</taxonomicName>
(arrowed in 
<figureCitation id="78EA79F3FFB9FFC46136F89B02B2F816" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="23" pageNumber="24">Fig. 12A</figureCitation>
), about 1.3 times as long as anal somite, cylindrical, 2.1 times as long as wide (ventral view), slightly divergent, and with space between them about one ramus width; ornamentation and armature as in 
<taxonomicName id="27D11EF5FFB9FFC4601EF843073CF871" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1113,1351,1973,1997]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFB9FFC4601EF843073CF871" box="[1113,1351,1973,1997]" italics="true" pageId="23" pageNumber="24">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except central part of inner margin without spinules and middle lateral seta much shorter (both arrowed in 
<figureCitation id="78EA79F3FFB9FFC46081F82F0752F84E" box="[1222,1321,2009,2034]" captionStart="FIGURE 13" captionStartId="25.[151,249,1906,1928]" captionTargetBox="[198,1393,208,1882]" captionTargetId="figure-79@25.[190,1397,193,1885]" captionTargetPageId="25" captionText="FIGURE 13. Wellstenhelia clio sp. nov., line drawings, female holotype: A, abdomen, ventral view; B, left side of cephalothoracic shield, dissected and flattened; C pleuron of third pedigerous somite, dissected and flattened; D, mandibular palp, anterior view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910594" httpUri="https://zenodo.org/record/4910594/files/figure.png" pageId="23" pageNumber="24">Fig. 13A</figureCitation>
).
</paragraph>
<caption id="B4AE35FEFFB6FFCB64D0F88A01FCF852" ID-DOI="http://doi.org/10.5281/zenodo.4910592" ID-Zenodo-Dep="4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,249,1916,1938]" targetBox="[190,1408,189,1892]" targetPageId="24">
<paragraph id="E06E6576FFB6FFCB64D0F88A01FCF852" blockId="24.[151,1436,1916,2030]" pageId="24" pageNumber="25">
<emphasis id="D2A5B964FFB6FFCB64D0F88A035BF82D" bold="true" box="[151,288,1916,1938]" pageId="24" pageNumber="25">FIGURE 12.</emphasis>
<taxonomicName id="27D11EF5FFB6FFCB6560F88A03A7F82E" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[295,476,1916,1938]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="clio" status="sp. nov.">
<emphasis id="D2A5B964FFB6FFCB6560F88A03A7F82E" box="[295,476,1916,1938]" italics="true" pageId="24" pageNumber="25">Wellstenhelia clio</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFB6FFCB65A3F88A004EF82D" bold="true" box="[484,565,1916,1937]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="C996041FFFB6FFCB65A3F88A004EF82D" box="[484,565,1916,1937]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in 
<taxonomicName id="27D11EF5FFB6FFCB665FF84F0081F873" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[536,762,1977,1999]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFB6FFCB665FF84F0081F873" box="[536,762,1977,1999]" italics="true" pageId="24" pageNumber="25">Wellstenhelia calliope</emphasis>
</taxonomicName>
. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm.
</paragraph>
</caption>
<caption id="B4AE35FEFFB7FFCA64D0F88401F6F858" ID-DOI="http://doi.org/10.5281/zenodo.4910594" ID-Zenodo-Dep="4910594" httpUri="https://zenodo.org/record/4910594/files/figure.png" pageId="25" pageNumber="26" startId="25.[151,249,1906,1928]" targetBox="[198,1393,208,1882]" targetPageId="25">
<paragraph id="E06E6576FFB7FFCA64D0F88401F6F858" blockId="25.[151,1435,1906,2020]" pageId="25" pageNumber="26">
<emphasis id="D2A5B964FFB7FFCA64D0F884035DF83B" bold="true" box="[151,294,1906,1928]" pageId="25" pageNumber="26">FIGURE 13.</emphasis>
<taxonomicName id="27D11EF5FFB7FFCA6573F8840394F834" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[308,495,1906,1928]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="clio" status="sp. nov.">
<emphasis id="D2A5B964FFB7FFCA6573F8840394F834" box="[308,495,1906,1928]" italics="true" pageId="25" pageNumber="26">Wellstenhelia clio</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFB7FFCA65BBF884002FF83B" bold="true" box="[508,596,1906,1927]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="C996041FFFB7FFCA65BBF884002FF83B" box="[508,596,1906,1927]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, line drawings, female holotype: A, abdomen, ventral view; B, left side of cephalothoracic shield, dissected and flattened; C pleuron of third pedigerous somite, dissected and flattened; D, mandibular palp, anterior view. Arabic numerals indicating sensilla and pores presumably homologous to those in 
<taxonomicName id="27D11EF5FFB7FFCA60F2F85907EEF879" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1205,1429,1967,1989]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFB7FFCA60F2F85907EEF879" box="[1205,1429,1967,1989]" italics="true" pageId="25" pageNumber="26">Wellstenhelia calliope</emphasis>
</taxonomicName>
. Arrowheads pointing most prominent specific features. Scale bars 100 µm.
</paragraph>
</caption>
<caption id="B4AE35FEFFB4FFC964D0F89407E7F84E" ID-DOI="http://doi.org/10.5281/zenodo.4910596" ID-Zenodo-Dep="4910596" httpUri="https://zenodo.org/record/4910596/files/figure.png" pageId="26" pageNumber="27" startId="26.[151,249,1890,1912]" targetBox="[199,1389,186,1864]" targetPageId="26">
<paragraph id="E06E6576FFB4FFC964D0F89407E7F84E" blockId="26.[151,1436,1890,2034]" pageId="26" pageNumber="27">
<emphasis id="D2A5B964FFB4FFC964D0F8940358F8CB" bold="true" box="[151,291,1890,1912]" pageId="26" pageNumber="27">FIGURE 14.</emphasis>
<taxonomicName id="27D11EF5FFB4FFC96569F894039FF8C4" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[302,484,1890,1912]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="clio" status="sp. nov.">
<emphasis id="D2A5B964FFB4FFC96569F894039FF8C4" box="[302,484,1890,1912]" italics="true" pageId="26" pageNumber="27">Wellstenhelia clio</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFB4FFC965A8F8940038F8CB" bold="true" box="[495,579,1890,1911]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="C996041FFFB4FFC965A8F8940038F8CB" box="[495,579,1890,1911]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, line drawings, A–D, female holotype, E–F, male allotype: A, pleuron of fourth pedigerous somite, dissected and flattened; B, fourth swimming leg without rami, anterior view; C, fifth leg, dissected and flattened, anterior view; D, sixth leg on genital somite, ventro-lateral view; E, cephalothoracic shield, lateral view; F, antennula, dorsal view. Arabic numerals indicating sensilla and pores presumably homologous to those in 
<taxonomicName id="27D11EF5FFB4FFC96012F8480749F868" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1109,1330,1982,2004]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFB4FFC96012F8480749F868" box="[1109,1330,1982,2004]" italics="true" pageId="26" pageNumber="27">Wellstenhelia calliope</emphasis>
</taxonomicName>
. Currency symbols ($, £, €) indicating unique sensilla and pores. Arrowheads pointing most prominent specific features. Scale bars 100 µm.
</paragraph>
</caption>
<caption id="B4AE35FEFFB5FFC864D0F8770297F84F" ID-DOI="http://doi.org/10.5281/zenodo.4910598" ID-Zenodo-Dep="4910598" httpUri="https://zenodo.org/record/4910598/files/figure.png" pageId="27" pageNumber="28" startId="27.[151,249,1921,1943]" targetBox="[195,1396,205,1892]" targetPageId="27">
<paragraph id="E06E6576FFB5FFC864D0F8770297F84F" blockId="27.[151,1436,1921,2035]" pageId="27" pageNumber="28">
<emphasis id="D2A5B964FFB5FFC864D0F877035BF82A" bold="true" box="[151,288,1921,1943]" pageId="27" pageNumber="28">FIGURE 15.</emphasis>
<taxonomicName id="27D11EF5FFB5FFC86561F87703A1F82B" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[294,474,1921,1943]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="clio" status="sp. nov.">
<emphasis id="D2A5B964FFB5FFC86561F87703A1F82B" box="[294,474,1921,1943]" italics="true" pageId="27" pageNumber="28">Wellstenhelia clio</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFB5FFC865A6F8740049F82B" bold="true" box="[481,562,1922,1943]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="C996041FFFB5FFC865A6F8740049F82B" box="[481,562,1922,1943]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, line drawings, male allotype: A, urosome, ventral view; B, urosome, lateral view; C, pleuron of second pedigerous somite, dissected and flattened. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in 
<taxonomicName id="27D11EF5FFB5FFC8664AF8480095F868" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[525,750,1982,2004]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFB5FFC8664AF8480095F868" box="[525,750,1982,2004]" italics="true" pageId="27" pageNumber="28">Wellstenhelia calliope</emphasis>
</taxonomicName>
. Arrowheads point to most prominent specific features. Scale bars 100 µm.
</paragraph>
</caption>
<caption id="B4AE35FEFFB2FFCF64D0F9D307ECF9C5" ID-DOI="http://doi.org/10.5281/zenodo.4910600" ID-Zenodo-Dep="4910600" httpUri="https://zenodo.org/record/4910600/files/figure.png" pageId="28" pageNumber="29" startId="28.[151,249,1573,1595]" targetBox="[151,1436,193,1534]" targetPageId="28">
<paragraph id="E06E6576FFB2FFCF64D0F9D307ECF9C5" blockId="28.[151,1436,1573,1657]" pageId="28" pageNumber="29">
<emphasis id="D2A5B964FFB2FFCF64D0F9D30364F986" bold="true" box="[151,287,1573,1595]" pageId="28" pageNumber="29">FIGURE 16.</emphasis>
<taxonomicName id="27D11EF5FFB2FFCF6561F9D303A2F987" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[294,473,1573,1595]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="clio" status="sp. nov.">
<emphasis id="D2A5B964FFB2FFCF6561F9D303A2F987" box="[294,473,1573,1595]" italics="true" pageId="28" pageNumber="29">Wellstenhelia clio</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFB2FFCF65A7F9D0004AF987" bold="true" box="[480,561,1574,1595]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="C996041FFFB2FFCF65A7F9D0004AF987" box="[480,561,1574,1595]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, scanning electron micrographs, female paratype: A, habitus, dorsal view; B, rostrum, dorsal view; C, cephalothorax, dorsal view; D, pedigerous somites, dorsal view; E, first urosomite and genital double-somite, dorsal view; F, anal somite and caudal rami, dorsal view. Scale bars 40 µm (B), 50 µm (D, E, F), 100 µm (C), and 200 µm (A).
</paragraph>
</caption>
<paragraph id="E06E6576FFB2FFCF6480F95303AAF95D" blockId="28.[151,1436,1701,2014]" pageId="28" pageNumber="29">
Antennula (
<figureCitation id="78EA79F3FFB2FFCF650BF95303CFF902" box="[332,436,1701,1726]" captionStart="FIGURE 16" captionStartId="28.[151,249,1573,1595]" captionTargetBox="[151,1436,193,1534]" captionTargetId="figure-209@28.[151,1436,193,1534]" captionTargetPageId="28" captionText="FIGURE 16. Wellstenhelia clio sp. nov., scanning electron micrographs, female paratype: A, habitus, dorsal view; B, rostrum, dorsal view; C, cephalothorax, dorsal view; D, pedigerous somites, dorsal view; E, first urosomite and genital double-somite, dorsal view; F, anal somite and caudal rami, dorsal view. Scale bars 40 µm (B), 50 µm (D, E, F), 100 µm (C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910600" httpUri="https://zenodo.org/record/4910600/files/figure.png" pageId="28" pageNumber="29">Fig. 16A</figureCitation>
), antenna, labrum, paragnaths, mandibula (
<figureCitation id="78EA79F3FFB2FFCF67DCF9530678F902" box="[923,1027,1701,1726]" captionStart="FIGURE 13" captionStartId="25.[151,249,1906,1928]" captionTargetBox="[198,1393,208,1882]" captionTargetId="figure-79@25.[190,1397,193,1885]" captionTargetPageId="25" captionText="FIGURE 13. Wellstenhelia clio sp. nov., line drawings, female holotype: A, abdomen, ventral view; B, left side of cephalothoracic shield, dissected and flattened; C pleuron of third pedigerous somite, dissected and flattened; D, mandibular palp, anterior view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910594" httpUri="https://zenodo.org/record/4910594/files/figure.png" pageId="28" pageNumber="29">Fig. 13D</figureCitation>
), maxillula, maxilla, and maxilliped as in 
<emphasis id="D2A5B964FFB2FFCF6495F93F03AAF95D" box="[210,465,1737,1761]" italics="true" pageId="28" pageNumber="29">
<taxonomicName id="27D11EF5FFB2FFCF6495F93F03B6F95D" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[210,461,1737,1761]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="calliope">Wellstenhelia calliope</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="E06E6576FFB2FFCF6480F91B061EF8F2" blockId="28.[151,1436,1701,2014]" pageId="28" pageNumber="29">
Swimming legs (
<figureCitation id="78EA79F3FFB2FFCF65CAF91B0078F8BA" box="[397,515,1773,1798]" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="28" pageNumber="29">Figs. 12D</figureCitation>
, 
<figureCitation id="78EA79F3FFB2FFCF6654F91B0038F8BA" box="[531,579,1773,1798]" captionStart="FIGURE 14" captionStartId="26.[151,249,1890,1912]" captionTargetBox="[199,1389,186,1864]" captionTargetId="figure-116@26.[186,1400,175,1868]" captionTargetPageId="26" captionText="FIGURE 14. Wellstenhelia clio sp. nov., line drawings, A–D, female holotype, E–F, male allotype: A, pleuron of fourth pedigerous somite, dissected and flattened; B, fourth swimming leg without rami, anterior view; C, fifth leg, dissected and flattened, anterior view; D, sixth leg on genital somite, ventro-lateral view; E, cephalothoracic shield, lateral view; F, antennula, dorsal view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, €) indicating unique sensilla and pores. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910596" httpUri="https://zenodo.org/record/4910596/files/figure.png" pageId="28" pageNumber="29">14B</figureCitation>
) segmentation, ornamentation, armature, and even proportions of various armature elements as in 
<emphasis id="D2A5B964FFB2FFCF65E1F8E700D8F895" box="[422,675,1809,1833]" italics="true" pageId="28" pageNumber="29">
<taxonomicName id="27D11EF5FFB2FFCF65E1F8E700E4F895" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[422,671,1809,1833]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="calliope">Wellstenhelia calliope</taxonomicName>
,
</emphasis>
except proximal rows of spinules on coxae slightly longer (arrowed in 
<figureCitation id="78EA79F3FFB2FFCF64F3F8C3036CF8F2" box="[180,279,1845,1870]" captionStart="FIGURE 14" captionStartId="26.[151,249,1890,1912]" captionTargetBox="[199,1389,186,1864]" captionTargetId="figure-116@26.[186,1400,175,1868]" captionTargetPageId="26" captionText="FIGURE 14. Wellstenhelia clio sp. nov., line drawings, A–D, female holotype, E–F, male allotype: A, pleuron of fourth pedigerous somite, dissected and flattened; B, fourth swimming leg without rami, anterior view; C, fifth leg, dissected and flattened, anterior view; D, sixth leg on genital somite, ventro-lateral view; E, cephalothoracic shield, lateral view; F, antennula, dorsal view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, €) indicating unique sensilla and pores. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910596" httpUri="https://zenodo.org/record/4910596/files/figure.png" pageId="28" pageNumber="29">Fig. 14B</figureCitation>
) and first endopodal segments without anterior pore (arrowed in 
<figureCitation id="78EA79F3FFB2FFCF67B7F8C30623F8F2" box="[1008,1112,1845,1870]" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="28" pageNumber="29">Fig. 12D</figureCitation>
).
</paragraph>
<paragraph id="E06E6576FFB2FFCE6480F8AF0339FF69" blockId="28.[151,1436,1701,2014]" lastBlockId="29.[151,1436,151,2013]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
Fifth leg (
<figureCitation id="78EA79F3FFB2FFCF6572F8AF03DCF8CE" box="[309,423,1881,1906]" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="28" pageNumber="29">Figs. 12A</figureCitation>
, 
<figureCitation id="78EA79F3FFB2FFCF65F6F8AF039AF8CE" box="[433,481,1881,1906]" captionStart="FIGURE 14" captionStartId="26.[151,249,1890,1912]" captionTargetBox="[199,1389,186,1864]" captionTargetId="figure-116@26.[186,1400,175,1868]" captionTargetPageId="26" captionText="FIGURE 14. Wellstenhelia clio sp. nov., line drawings, A–D, female holotype, E–F, male allotype: A, pleuron of fourth pedigerous somite, dissected and flattened; B, fourth swimming leg without rami, anterior view; C, fifth leg, dissected and flattened, anterior view; D, sixth leg on genital somite, ventro-lateral view; E, cephalothoracic shield, lateral view; F, antennula, dorsal view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, €) indicating unique sensilla and pores. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910596" httpUri="https://zenodo.org/record/4910596/files/figure.png" pageId="28" pageNumber="29">14C</figureCitation>
) segmentation, general shape, number of armature elements, and most ornamentation as in 
<emphasis id="D2A5B964FFB2FFCF6494F88B03A9F829" box="[211,466,1917,1941]" italics="true" pageId="28" pageNumber="29">
<taxonomicName id="27D11EF5FFB2FFCF6494F88B03B5F829" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[211,462,1917,1941]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="calliope">Wellstenhelia calliope</taxonomicName>
,
</emphasis>
except innermost endopodal seta proportionately longer (arrowed in 
<figureCitation id="78EA79F3FFB2FFCF6098F88B072BF82A" box="[1247,1360,1917,1942]" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="28" pageNumber="29">Figs. 12A</figureCitation>
, 
<figureCitation id="78EA79F3FFB2FFCF611AF88B07F4F82A" box="[1373,1423,1917,1942]" captionStart="FIGURE 14" captionStartId="26.[151,249,1890,1912]" captionTargetBox="[199,1389,186,1864]" captionTargetId="figure-116@26.[186,1400,175,1868]" captionTargetPageId="26" captionText="FIGURE 14. Wellstenhelia clio sp. nov., line drawings, A–D, female holotype, E–F, male allotype: A, pleuron of fourth pedigerous somite, dissected and flattened; B, fourth swimming leg without rami, anterior view; C, fifth leg, dissected and flattened, anterior view; D, sixth leg on genital somite, ventro-lateral view; E, cephalothoracic shield, lateral view; F, antennula, dorsal view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, €) indicating unique sensilla and pores. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910596" httpUri="https://zenodo.org/record/4910596/files/figure.png" pageId="28" pageNumber="29">14C</figureCitation>
), space between two central endopodal setae wider (arrowed in 
<figureCitation id="78EA79F3FFB2FFCF6722F85701B6F806" box="[869,973,1953,1978]" captionStart="FIGURE 14" captionStartId="26.[151,249,1890,1912]" captionTargetBox="[199,1389,186,1864]" captionTargetId="figure-116@26.[186,1400,175,1868]" captionTargetPageId="26" captionText="FIGURE 14. Wellstenhelia clio sp. nov., line drawings, A–D, female holotype, E–F, male allotype: A, pleuron of fourth pedigerous somite, dissected and flattened; B, fourth swimming leg without rami, anterior view; C, fifth leg, dissected and flattened, anterior view; D, sixth leg on genital somite, ventro-lateral view; E, cephalothoracic shield, lateral view; F, antennula, dorsal view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, €) indicating unique sensilla and pores. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910596" httpUri="https://zenodo.org/record/4910596/files/figure.png" pageId="28" pageNumber="29">Fig. 14C</figureCitation>
), exopod wider at base (arrowed in 
<figureCitation id="78EA79F3FFB2FFCF6137F85702B2F862" captionStart="FIGURE 14" captionStartId="26.[151,249,1890,1912]" captionTargetBox="[199,1389,186,1864]" captionTargetId="figure-116@26.[186,1400,175,1868]" captionTargetPageId="26" captionText="FIGURE 14. Wellstenhelia clio sp. nov., line drawings, A–D, female holotype, E–F, male allotype: A, pleuron of fourth pedigerous somite, dissected and flattened; B, fourth swimming leg without rami, anterior view; C, fifth leg, dissected and flattened, anterior view; D, sixth leg on genital somite, ventro-lateral view; E, cephalothoracic shield, lateral view; F, antennula, dorsal view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, €) indicating unique sensilla and pores. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910596" httpUri="https://zenodo.org/record/4910596/files/figure.png" pageId="28" pageNumber="29">Fig. 14C</figureCitation>
), and additional pore present on anterior surface of exopod (arrowed in 
<figureCitation id="78EA79F3FFB2FFCF67BEF833061AF862" box="[1017,1121,1989,2014]" captionStart="FIGURE 14" captionStartId="26.[151,249,1890,1912]" captionTargetBox="[199,1389,186,1864]" captionTargetId="figure-116@26.[186,1400,175,1868]" captionTargetPageId="26" captionText="FIGURE 14. Wellstenhelia clio sp. nov., line drawings, A–D, female holotype, E–F, male allotype: A, pleuron of fourth pedigerous somite, dissected and flattened; B, fourth swimming leg without rami, anterior view; C, fifth leg, dissected and flattened, anterior view; D, sixth leg on genital somite, ventro-lateral view; E, cephalothoracic shield, lateral view; F, antennula, dorsal view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, €) indicating unique sensilla and pores. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910596" httpUri="https://zenodo.org/record/4910596/files/figure.png" pageId="28" pageNumber="29">Fig. 14C</figureCitation>
). Length ratio of endopodal setae, starting from inner side, 1: 1.2: 1.7: 1. Length ratio of exopodal setae, starting from inner side, 1: 0.3: 0.3: 0.8: 0.6: 0.6.
</paragraph>
<paragraph id="E06E6576FFB3FFCE6480FF290092FF44" blockId="29.[151,1436,151,2013]" box="[199,745,223,248]" pageId="29" pageNumber="30">
Sixth leg (
<figureCitation id="78EA79F3FFB3FFCE657CFF2903DBFF44" box="[315,416,223,248]" captionStart="FIGURE 14" captionStartId="26.[151,249,1890,1912]" captionTargetBox="[199,1389,186,1864]" captionTargetId="figure-116@26.[186,1400,175,1868]" captionTargetPageId="26" captionText="FIGURE 14. Wellstenhelia clio sp. nov., line drawings, A–D, female holotype, E–F, male allotype: A, pleuron of fourth pedigerous somite, dissected and flattened; B, fourth swimming leg without rami, anterior view; C, fifth leg, dissected and flattened, anterior view; D, sixth leg on genital somite, ventro-lateral view; E, cephalothoracic shield, lateral view; F, antennula, dorsal view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, €) indicating unique sensilla and pores. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910596" httpUri="https://zenodo.org/record/4910596/files/figure.png" pageId="29" pageNumber="30">Fig. 14D</figureCitation>
) as in 
<emphasis id="D2A5B964FFB3FFCE65ADFF160092FF44" box="[490,745,224,248]" italics="true" pageId="29" pageNumber="30">
<taxonomicName id="27D11EF5FFB3FFCE65ADFF16009EFF44" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[490,741,224,248]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="calliope">Wellstenhelia calliope</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="E06E6576FFB3FFCE6480FEF20306FDDC" blockId="29.[151,1436,151,2013]" pageId="29" pageNumber="30">
Male (based on 
<typeStatus id="3F6ADBD4FFB3FFCE6539FEF203A3FEA1" box="[382,472,260,285]" pageId="29" pageNumber="30" type="allotype">allotype</typeStatus>
and 
<specimenCount id="F6D7AEFFFFB3FFCE6655FEF200CEFEA1" box="[530,693,260,285]" pageId="29" pageNumber="30" type="generic" typeStatus="paratypes">five paratypes</specimenCount>
). Body length from 
<quantity id="2729C893FFB3FFCE67DCFEF20638FEA0" box="[923,1091,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="5.415" metricValueMax="5.64" metricValueMin="5.1899999999999995" pageId="29" pageNumber="30" unit="mm" value="541.5" valueMax="564.0" valueMin="519.0">519 to 564 µm</quantity>
(
<quantity id="2729C893FFB3FFCE6014FEF206D7FEA0" box="[1107,1196,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="5.55" pageId="29" pageNumber="30" unit="mm" value="555.0">555 µm</quantity>
in 
<typeStatus id="3F6ADBD4FFB3FFCE6095FEF2074AFEA1" box="[1234,1329,260,285]" pageId="29" pageNumber="30" type="allotype">allotype</typeStatus>
). Habitus (
<figureCitation id="78EA79F3FFB3FFCE64D8FED10373FEFC" box="[159,264,295,320]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">Fig. 17A</figureCitation>
), colour, rostrum (
<figureCitation id="78EA79F3FFB3FFCE659DFED1003AFEFC" box="[474,577,295,320]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">Fig. 17C</figureCitation>
), shape and almost all ornamentation of cephalothorax (
<figureCitation id="78EA79F3FFB3FFCE60FAFED10756FEFC" box="[1213,1325,295,320]" captionStart="FIGURE 14" captionStartId="26.[151,249,1890,1912]" captionTargetBox="[199,1389,186,1864]" captionTargetId="figure-116@26.[186,1400,175,1868]" captionTargetPageId="26" captionText="FIGURE 14. Wellstenhelia clio sp. nov., line drawings, A–D, female holotype, E–F, male allotype: A, pleuron of fourth pedigerous somite, dissected and flattened; B, fourth swimming leg without rami, anterior view; C, fifth leg, dissected and flattened, anterior view; D, sixth leg on genital somite, ventro-lateral view; E, cephalothoracic shield, lateral view; F, antennula, dorsal view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, €) indicating unique sensilla and pores. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910596" httpUri="https://zenodo.org/record/4910596/files/figure.png" pageId="29" pageNumber="30">Figs. 14E</figureCitation>
, 
<figureCitation id="78EA79F3FFB3FFCE617DFED107F5FEFC" box="[1338,1422,295,320]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">17A, C</figureCitation>
), shape and ornamentation of second pedigerous somite (
<figureCitation id="78EA79F3FFB3FFCE6750FEBA01F1FED9" box="[791,906,332,357]" captionStart="FIGURE 15" captionStartId="27.[151,249,1921,1943]" captionTargetBox="[195,1396,205,1892]" captionTargetId="figure-74@27.[186,1401,193,1900]" captionTargetPageId="27" captionText="FIGURE 15. Wellstenhelia clio sp. nov., line drawings, male allotype: A, urosome, ventral view; B, urosome, lateral view; C, pleuron of second pedigerous somite, dissected and flattened. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910598" httpUri="https://zenodo.org/record/4910598/files/figure.png" pageId="29" pageNumber="30">Figs. 15C</figureCitation>
, 
<figureCitation id="78EA79F3FFB3FFCE67DFFEBA01B2FED9" box="[920,969,332,357]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">17A</figureCitation>
) (including missing sensilla pair no. 48; arrowed in 
<figureCitation id="78EA79F3FFB3FFCE6550FE990307FE34" box="[279,380,367,392]" captionStart="FIGURE 15" captionStartId="27.[151,249,1921,1943]" captionTargetBox="[195,1396,205,1892]" captionTargetId="figure-74@27.[186,1401,193,1900]" captionTargetPageId="27" captionText="FIGURE 15. Wellstenhelia clio sp. nov., line drawings, male allotype: A, urosome, ventral view; B, urosome, lateral view; C, pleuron of second pedigerous somite, dissected and flattened. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910598" httpUri="https://zenodo.org/record/4910598/files/figure.png" pageId="29" pageNumber="30">Fig. 15C</figureCitation>
), third pedigerous somite (
<figureCitation id="78EA79F3FFB3FFCE66EEFE99016AFE34" box="[681,785,367,392]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">Fig. 17A</figureCitation>
), and fourth pedigerous somite (
<figureCitation id="78EA79F3FFB3FFCE6039FE99069EFE34" box="[1150,1253,367,392]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">Fig. 17A</figureCitation>
), ornamentation of first urosomite (
<figureCitation id="78EA79F3FFB3FFCE6535FE62039EFE11" box="[370,485,404,429]" captionStart="FIGURE 15" captionStartId="27.[151,249,1921,1943]" captionTargetBox="[195,1396,205,1892]" captionTargetId="figure-74@27.[186,1401,193,1900]" captionTargetPageId="27" captionText="FIGURE 15. Wellstenhelia clio sp. nov., line drawings, male allotype: A, urosome, ventral view; B, urosome, lateral view; C, pleuron of second pedigerous somite, dissected and flattened. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910598" httpUri="https://zenodo.org/record/4910598/files/figure.png" pageId="29" pageNumber="30">Figs. 15B</figureCitation>
, 
<figureCitation id="78EA79F3FFB3FFCE65B3FE62005EFE11" box="[500,549,404,429]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">17B</figureCitation>
), ornamentation of last threeurosomites (
<figureCitation id="78EA79F3FFB3FFCE6047FE6206E2FE17" box="[1024,1177,404,429]" captionStart="FIGURE 15" captionStartId="27.[151,249,1921,1943]" captionTargetBox="[195,1396,205,1892]" captionTargetId="figure-74@27.[186,1401,193,1900]" captionTargetPageId="27" captionText="FIGURE 15. Wellstenhelia clio sp. nov., line drawings, male allotype: A, urosome, ventral view; B, urosome, lateral view; C, pleuron of second pedigerous somite, dissected and flattened. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910598" httpUri="https://zenodo.org/record/4910598/files/figure.png" pageId="29" pageNumber="30">Figs. 15A, B</figureCitation>
, 
<figureCitation id="78EA79F3FFB3FFCE60E0FE620686FE11" box="[1191,1277,404,429]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">17A, B</figureCitation>
), caudal rami (
<figureCitation id="78EA79F3FFB3FFCE64D8FE410352FE73" box="[159,297,439,464]" captionStart="FIGURE 15" captionStartId="27.[151,249,1921,1943]" captionTargetBox="[195,1396,205,1892]" captionTargetId="figure-74@27.[186,1401,193,1900]" captionTargetPageId="27" captionText="FIGURE 15. Wellstenhelia clio sp. nov., line drawings, male allotype: A, urosome, ventral view; B, urosome, lateral view; C, pleuron of second pedigerous somite, dissected and flattened. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910598" httpUri="https://zenodo.org/record/4910598/files/figure.png" pageId="29" pageNumber="30">Fig. 15A, B</figureCitation>
, 
<figureCitation id="78EA79F3FFB3FFCE6572FE410313FE6C" box="[309,360,439,464]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">17A</figureCitation>
), antenna (
<figureCitation id="78EA79F3FFB3FFCE65A3FE410036FE6C" box="[484,589,439,464]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">Fig. 17D</figureCitation>
), labrum (
<figureCitation id="78EA79F3FFB3FFCE6685FE410151FE6C" box="[706,810,439,464]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">Fig. 17D</figureCitation>
), paragnaths, mandibula, maxillula, maxilla (
<figureCitation id="78EA79F3FFB3FFCE616EFE4107F4FE6C" box="[1321,1423,439,464]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">Fig. 17E</figureCitation>
), maxilliped (
<figureCitation id="78EA79F3FFB3FFCE6567FE2A03FDFE49" box="[288,390,476,501]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">Fig. 17E</figureCitation>
), first swimming leg (
<figureCitation id="78EA79F3FFB3FFCE66C3FE2A0093FE49" box="[644,744,476,501]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">Fig. 17F</figureCitation>
), and coxae, bases, and exopods of second, third, and fourth swimming legs as in female. Prosome/urosome ratio 1.1, greatest width at posterior end of cephalothorax, body length/width ratio about 3.5; cephalothorax twice as wide as genital somite in dorsal view. Genital somite and third urosomite not fused.
</paragraph>
<paragraph id="E06E6576FFB3FFCE6480FD9A00FFFD14" blockId="29.[151,1436,151,2013]" pageId="29" pageNumber="30">
Cephalothorax (
<figureCitation id="78EA79F3FFB3FFCE653BFD9A0397FD39" box="[380,492,620,645]" captionStart="FIGURE 14" captionStartId="26.[151,249,1890,1912]" captionTargetBox="[199,1389,186,1864]" captionTargetId="figure-116@26.[186,1400,175,1868]" captionTargetPageId="26" captionText="FIGURE 14. Wellstenhelia clio sp. nov., line drawings, A–D, female holotype, E–F, male allotype: A, pleuron of fourth pedigerous somite, dissected and flattened; B, fourth swimming leg without rami, anterior view; C, fifth leg, dissected and flattened, anterior view; D, sixth leg on genital somite, ventro-lateral view; E, cephalothoracic shield, lateral view; F, antennula, dorsal view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, €) indicating unique sensilla and pores. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910596" httpUri="https://zenodo.org/record/4910596/files/figure.png" pageId="29" pageNumber="30">Figs. 14E</figureCitation>
, 
<figureCitation id="78EA79F3FFB3FFCE65BFFD9A0030FD39" box="[504,587,620,645]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">17A, C</figureCitation>
) in addition to all sensilla and pores present in female, with one additional pair of lateral pores in posterior half (no. €).
</paragraph>
<paragraph id="E06E6576FFB3FFCE6480FD42013EFD4C" blockId="29.[151,1436,151,2013]" pageId="29" pageNumber="30">
First urosomite (
<figureCitation id="78EA79F3FFB3FFCE65CEFD420386FD71" box="[393,509,692,717]" captionStart="FIGURE 15" captionStartId="27.[151,249,1921,1943]" captionTargetBox="[195,1396,205,1892]" captionTargetId="figure-74@27.[186,1401,193,1900]" captionTargetPageId="27" captionText="FIGURE 15. Wellstenhelia clio sp. nov., line drawings, male allotype: A, urosome, ventral view; B, urosome, lateral view; C, pleuron of second pedigerous somite, dissected and flattened. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910598" httpUri="https://zenodo.org/record/4910598/files/figure.png" pageId="29" pageNumber="30">Figs. 15B</figureCitation>
, 
<figureCitation id="78EA79F3FFB3FFCE6649FD420046FD71" box="[526,573,692,717]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">17B</figureCitation>
) slightly narrower and longer than in female but also with three pairs of sensilla (nos. 64, 65, 66) and two pairs of pores (nos. 63, 67).
</paragraph>
<paragraph id="E06E6576FFB3FFCE6480FD0A0643FCE1" blockId="29.[151,1436,151,2013]" pageId="29" pageNumber="30">
Genital somite (
<figureCitation id="78EA79F3FFB3FFCE653EFD0A0071FCAF" box="[377,522,764,789]" captionStart="FIGURE 15" captionStartId="27.[151,249,1921,1943]" captionTargetBox="[195,1396,205,1892]" captionTargetId="figure-74@27.[186,1401,193,1900]" captionTargetPageId="27" captionText="FIGURE 15. Wellstenhelia clio sp. nov., line drawings, male allotype: A, urosome, ventral view; B, urosome, lateral view; C, pleuron of second pedigerous somite, dissected and flattened. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910598" httpUri="https://zenodo.org/record/4910598/files/figure.png" pageId="29" pageNumber="30">Figs. 15A, B</figureCitation>
, 
<figureCitation id="78EA79F3FFB3FFCE6652FD0A003EFCA9" box="[533,581,764,789]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">17B</figureCitation>
) somewhat wider and with fewer spinules than in 
<emphasis id="D2A5B964FFB3FFCE6033FD0A070AFCA8" box="[1140,1393,764,788]" italics="true" pageId="29" pageNumber="30">
<taxonomicName id="27D11EF5FFB3FFCE6033FD0A0716FCA8" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1140,1389,764,788]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="calliope">Wellstenhelia calliope</taxonomicName>
,
</emphasis>
but with all ornamentation same, except pore no. 68 situated more ventrally (arrowed in 
<figureCitation id="78EA79F3FFB3FFCE6015FCE906CCFC84" box="[1106,1207,799,824]" captionStart="FIGURE 15" captionStartId="27.[151,249,1921,1943]" captionTargetBox="[195,1396,205,1892]" captionTargetId="figure-74@27.[186,1401,193,1900]" captionTargetPageId="27" captionText="FIGURE 15. Wellstenhelia clio sp. nov., line drawings, male allotype: A, urosome, ventral view; B, urosome, lateral view; C, pleuron of second pedigerous somite, dissected and flattened. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910598" httpUri="https://zenodo.org/record/4910598/files/figure.png" pageId="29" pageNumber="30">Fig. 15B</figureCitation>
), i.e. much closer to pore no. #, and in 
<specimenCount id="F6D7AEFFFFB3FFCE6525FCB2038AFCE0" box="[354,497,836,860]" pageId="29" pageNumber="30" type="generic" typeStatus="paratype">one paratype</specimenCount>
two pores extremely close to each other (
<figureCitation id="78EA79F3FFB3FFCE6783FCB20650FCE1" box="[964,1067,836,861]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">Fig. 17B</figureCitation>
).
</paragraph>
<paragraph id="E06E6576FFB3FFCE6480FC91076AFC19" blockId="29.[151,1436,151,2013]" pageId="29" pageNumber="30">
Third urosomite (
<figureCitation id="78EA79F3FFB3FFCE65CDFC910060FCC3" box="[394,539,871,896]" captionStart="FIGURE 15" captionStartId="27.[151,249,1921,1943]" captionTargetBox="[195,1396,205,1892]" captionTargetId="figure-74@27.[186,1401,193,1900]" captionTargetPageId="27" captionText="FIGURE 15. Wellstenhelia clio sp. nov., line drawings, male allotype: A, urosome, ventral view; B, urosome, lateral view; C, pleuron of second pedigerous somite, dissected and flattened. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910598" httpUri="https://zenodo.org/record/4910598/files/figure.png" pageId="29" pageNumber="30">Figs. 15A, B</figureCitation>
, 
<figureCitation id="78EA79F3FFB3FFCE6661FC91002EFC3C" box="[550,597,871,896]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">17B</figureCitation>
) as in 
<emphasis id="D2A5B964FFB3FFCE66D9FC9E01E0FC3C" box="[670,923,872,896]" italics="true" pageId="29" pageNumber="30">
<taxonomicName id="27D11EF5FFB3FFCE66D9FC9E01ECFC3C" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[670,919,872,896]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="calliope">Wellstenhelia calliope</taxonomicName>
,
</emphasis>
except ventral row of spinules not interrupted between sensilla pair no. 73 and not broken between sensilla nos. 72 &amp; 73 (both arrowed in 
<figureCitation id="78EA79F3FFB3FFCE60DAFC7A077EFC19" box="[1181,1285,908,933]" captionStart="FIGURE 15" captionStartId="27.[151,249,1921,1943]" captionTargetBox="[195,1396,205,1892]" captionTargetId="figure-74@27.[186,1401,193,1900]" captionTargetPageId="27" captionText="FIGURE 15. Wellstenhelia clio sp. nov., line drawings, male allotype: A, urosome, ventral view; B, urosome, lateral view; C, pleuron of second pedigerous somite, dissected and flattened. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910598" httpUri="https://zenodo.org/record/4910598/files/figure.png" pageId="29" pageNumber="30">Fig. 15A</figureCitation>
).
</paragraph>
<paragraph id="E06E6576FFB3FFCE6480FC59012CFBAC" blockId="29.[151,1436,151,2013]" pageId="29" pageNumber="30">
Antennula (
<figureCitation id="78EA79F3FFB3FFCE6515FC5903BFFC74" box="[338,452,943,968]" captionStart="FIGURE 14" captionStartId="26.[151,249,1890,1912]" captionTargetBox="[199,1389,186,1864]" captionTargetId="figure-116@26.[186,1400,175,1868]" captionTargetPageId="26" captionText="FIGURE 14. Wellstenhelia clio sp. nov., line drawings, A–D, female holotype, E–F, male allotype: A, pleuron of fourth pedigerous somite, dissected and flattened; B, fourth swimming leg without rami, anterior view; C, fifth leg, dissected and flattened, anterior view; D, sixth leg on genital somite, ventro-lateral view; E, cephalothoracic shield, lateral view; F, antennula, dorsal view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, €) indicating unique sensilla and pores. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910596" httpUri="https://zenodo.org/record/4910596/files/figure.png" pageId="29" pageNumber="30">Figs. 14F</figureCitation>
, 
<figureCitation id="78EA79F3FFB3FFCE6591FC59007DFC74" box="[470,518,943,968]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">17C</figureCitation>
) shape, segmentation, armature, and most ornamentation as in 
<emphasis id="D2A5B964FFB3FFCE6141FC460283FC50" italics="true" pageId="29" pageNumber="30">
<taxonomicName id="27D11EF5FFB3FFCE6141FC46028FFC50" authorityName="Karanovic &amp; Kim" authorityYear="2014" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="calliope">Wellstenhelia calliope</taxonomicName>
,
</emphasis>
except spiniform process on first segment smaller (arrowed in 
<figureCitation id="78EA79F3FFB3FFCE67F8FC22065AFC51" box="[959,1057,980,1005]" captionStart="FIGURE 15" captionStartId="27.[151,249,1921,1943]" captionTargetBox="[195,1396,205,1892]" captionTargetId="figure-74@27.[186,1401,193,1900]" captionTargetPageId="27" captionText="FIGURE 15. Wellstenhelia clio sp. nov., line drawings, male allotype: A, urosome, ventral view; B, urosome, lateral view; C, pleuron of second pedigerous somite, dissected and flattened. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910598" httpUri="https://zenodo.org/record/4910598/files/figure.png" pageId="29" pageNumber="30">Fig. 15F</figureCitation>
), dorsal pore missing (arrowed in 
<figureCitation id="78EA79F3FFB3FFCE64D0FC010283FBAC" box="[151,248,1015,1040]" captionStart="FIGURE 15" captionStartId="27.[151,249,1921,1943]" captionTargetBox="[195,1396,205,1892]" captionTargetId="figure-74@27.[186,1401,193,1900]" captionTargetPageId="27" captionText="FIGURE 15. Wellstenhelia clio sp. nov., line drawings, male allotype: A, urosome, ventral view; B, urosome, lateral view; C, pleuron of second pedigerous somite, dissected and flattened. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910598" httpUri="https://zenodo.org/record/4910598/files/figure.png" pageId="29" pageNumber="30">Fig. 15F</figureCitation>
), and aesthetascs on third and fourth segments longer.
</paragraph>
<paragraph id="E06E6576FFB3FFCE6480FBEA00D3FBE4" blockId="29.[151,1436,151,2013]" pageId="29" pageNumber="30">
Fifth leg (
<figureCitation id="78EA79F3FFB3FFCE6579FBEA03A1FB8F" box="[318,474,1052,1077]" captionStart="FIGURE 15" captionStartId="27.[151,249,1921,1943]" captionTargetBox="[195,1396,205,1892]" captionTargetId="figure-74@27.[186,1401,193,1900]" captionTargetPageId="27" captionText="FIGURE 15. Wellstenhelia clio sp. nov., line drawings, male allotype: A, urosome, ventral view; B, urosome, lateral view; C, pleuron of second pedigerous somite, dissected and flattened. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910598" httpUri="https://zenodo.org/record/4910598/files/figure.png" pageId="29" pageNumber="30">Figs. 15A, B</figureCitation>
, 
<figureCitation id="78EA79F3FFB3FFCE65ADFBEA0062FB89" box="[490,537,1052,1077]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">17B</figureCitation>
) shape, armature, and ornamentation as in 
<emphasis id="D2A5B964FFB3FFCE605EFBEA0760FB88" box="[1049,1307,1052,1076]" italics="true" pageId="29" pageNumber="30">
<taxonomicName id="27D11EF5FFB3FFCE605EFBEA076CFB88" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1049,1303,1052,1076]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="calliope">Wellstenhelia calliope</taxonomicName>
,
</emphasis>
except for shallow ventral notch on fused baseoendopods.
</paragraph>
<paragraph id="E06E6576FFB3FFCE6480FB92035DFB1C" blockId="29.[151,1436,151,2013]" pageId="29" pageNumber="30">
Sixth legs (
<figureCitation id="78EA79F3FFB3FFCE6502FB9203B6FBC7" box="[325,461,1124,1149]" captionStart="FIGURE 15" captionStartId="27.[151,249,1921,1943]" captionTargetBox="[195,1396,205,1892]" captionTargetId="figure-74@27.[186,1401,193,1900]" captionTargetPageId="27" captionText="FIGURE 15. Wellstenhelia clio sp. nov., line drawings, male allotype: A, urosome, ventral view; B, urosome, lateral view; C, pleuron of second pedigerous somite, dissected and flattened. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910598" httpUri="https://zenodo.org/record/4910598/files/figure.png" pageId="29" pageNumber="30">Fig. 15A, B</figureCitation>
, 
<figureCitation id="78EA79F3FFB3FFCE659FFB92007CFBC1" box="[472,519,1124,1149]" captionStart="FIGURE 17" captionStartId="30.[151,249,1630,1652]" captionTargetBox="[151,1436,267,1608]" captionTargetId="figure-154@30.[151,1436,267,1608]" captionTargetPageId="30" captionText="FIGURE 17. Wellstenhelia clio sp. nov., scanning electron micrographs, A–B, male paratype 1, C, male paratype 2, D–F, male paratype 3: A, habitus, lateral view; B, fifth pedigerous and anal somites, lateral view; C, antennula and rostrum, lateral view; cephalothorax, dorsal view; D, labrum and antennae, ventro-lateral view; E, maxilla and mailliped, ventral view; F, first and second swimming leg, ventro-anterior view. Scale bars 20 µm (E), 30 µm (D), 50 µm (B, C, F), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910602" httpUri="https://zenodo.org/record/4910602/files/figure.png" pageId="29" pageNumber="30">17B</figureCitation>
) as in 
<emphasis id="D2A5B964FFB3FFCE6615FB92012BFBC0" box="[594,848,1124,1148]" italics="true" pageId="29" pageNumber="30">
<taxonomicName id="27D11EF5FFB3FFCE6615FB920137FBC0" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[594,844,1124,1148]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="calliope">Wellstenhelia calliope</taxonomicName>
,
</emphasis>
except middle seta proportionately shorter (arrowed in 
<figureCitation id="78EA79F3FFB3FFCE64F3FB710361FB1C" box="[180,282,1159,1184]" captionStart="FIGURE 15" captionStartId="27.[151,249,1921,1943]" captionTargetBox="[195,1396,205,1892]" captionTargetId="figure-74@27.[186,1401,193,1900]" captionTargetPageId="27" captionText="FIGURE 15. Wellstenhelia clio sp. nov., line drawings, male allotype: A, urosome, ventral view; B, urosome, lateral view; C, pleuron of second pedigerous somite, dissected and flattened. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910598" httpUri="https://zenodo.org/record/4910598/files/figure.png" pageId="29" pageNumber="30">Fig. 15B</figureCitation>
).
</paragraph>
<paragraph id="E06E6576FFB3FFCE6480FB5A06B8FAB1" blockId="29.[151,1436,151,2013]" pageId="29" pageNumber="30">
<emphasis id="D2A5B964FFB3FFCE6480FB5A0336FB79" bold="true" box="[199,333,1196,1221]" pageId="29" pageNumber="30">Variability.</emphasis>
Most morphological features are extremely conservative, including the sensilla and pores pattern of somites, and length ratio of different armature on appendages. Except for body length, the only other variable feature was the position of lateral pore no. 68 on the male genital somite (arrowed in 
<figureCitation id="78EA79F3FFB3FFCE6016FB0206CDFAB1" box="[1105,1206,1268,1293]" captionStart="FIGURE 15" captionStartId="27.[151,249,1921,1943]" captionTargetBox="[195,1396,205,1892]" captionTargetId="figure-74@27.[186,1401,193,1900]" captionTargetPageId="27" captionText="FIGURE 15. Wellstenhelia clio sp. nov., line drawings, male allotype: A, urosome, ventral view; B, urosome, lateral view; C, pleuron of second pedigerous somite, dissected and flattened. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910598" httpUri="https://zenodo.org/record/4910598/files/figure.png" pageId="29" pageNumber="30">Fig. 15B</figureCitation>
).
</paragraph>
<paragraph id="E06E6576FFB3FFCE6480FAE10359F8B9" blockId="29.[151,1436,151,2013]" pageId="29" pageNumber="30">
<emphasis id="D2A5B964FFB3FFCE6480FAE10396FA8C" bold="true" box="[199,493,1303,1328]" pageId="29" pageNumber="30">Morphological affinities.</emphasis>
<taxonomicName id="27D11EF5FFB3FFCE65B4FAEE00C2FA8C" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[499,697,1304,1328]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="clio" status="sp. nov.">
<emphasis id="D2A5B964FFB3FFCE65B4FAEE00C2FA8C" box="[499,697,1304,1328]" italics="true" pageId="29" pageNumber="30">Wellstenhelia clio</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFB3FFCE66F8FAE10163FA8C" bold="true" box="[703,792,1303,1328]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="C996041FFFB3FFCE66F8FAE10163FA8C" box="[703,792,1303,1328]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has no obvious autapomorphy that would distinguish it at once from all other congeners. As mentioned above, its female fifth leg (
<figureCitation id="78EA79F3FFB3FFCE6787FACA065FFAE9" box="[960,1060,1340,1365]" captionStart="FIGURE 14" captionStartId="26.[151,249,1890,1912]" captionTargetBox="[199,1389,186,1864]" captionTargetId="figure-116@26.[186,1400,175,1868]" captionTargetPageId="26" captionText="FIGURE 14. Wellstenhelia clio sp. nov., line drawings, A–D, female holotype, E–F, male allotype: A, pleuron of fourth pedigerous somite, dissected and flattened; B, fourth swimming leg without rami, anterior view; C, fifth leg, dissected and flattened, anterior view; D, sixth leg on genital somite, ventro-lateral view; E, cephalothoracic shield, lateral view; F, antennula, dorsal view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, €) indicating unique sensilla and pores. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910596" httpUri="https://zenodo.org/record/4910596/files/figure.png" pageId="29" pageNumber="30">Fig. 14C</figureCitation>
) is relatively similar to that in the Swedish 
<taxonomicName id="27D11EF5FFB3FFCE6547FA9600C9FAC4" authority="(Lang, 1948)" authorityName="Karanovic &amp; Kim" authorityYear="2014" baseAuthorityName="Lang" baseAuthorityYear="1948" box="[256,690,1375,1400]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="hanstromi" status="comb. nov.">
<emphasis id="D2A5B964FFB3FFCE6547FA96006AFAC4" box="[256,529,1376,1400]" italics="true" pageId="29" pageNumber="30">Wellstenhelia hanstromi</emphasis>
(
<bibRefCitation id="84401887FFB3FFCE6663FAA900D1FAC4" author="Lang, K." box="[548,682,1375,1400]" pageId="29" pageNumber="30" refId="ref49308" refString="Lang, K. (1948) Monographie der Harpacticiden, 1 - 2. Nordiska Bokhandeln, Lund, 1682 pp." type="book" year="1948">Lang, 1948</bibRefCitation>
)
</taxonomicName>
<emphasis id="D2A5B964FFB3FFCE66FAFAA9013AFAC4" bold="true" box="[701,833,1375,1400]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="C996041FFFB3FFCE66FAFAA9013AFAC4" box="[701,833,1375,1400]" pageId="29" pageNumber="30" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, with two inner endopodal setae of about the same length and strength and as long as the outermost endopodal seta (see 
<bibRefCitation id="84401887FFB3FFCE67E2FA720659FA21" author="Lang, K." box="[933,1058,1412,1437]" pageId="29" pageNumber="30" refId="ref49308" refString="Lang, K. (1948) Monographie der Harpacticiden, 1 - 2. Nordiska Bokhandeln, Lund, 1682 pp." type="book" year="1948">Lang 1948</bibRefCitation>
), although not as long as those in the Artcic 
<taxonomicName id="27D11EF5FFB3FFCE6559FA5E0038FA7C" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[286,579,1448,1472]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="melpomene" status="sp. nov.">
<emphasis id="D2A5B964FFB3FFCE6559FA5E0038FA7C" box="[286,579,1448,1472]" italics="true" pageId="29" pageNumber="30">Wellstenhelia melpomene</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFB3FFCE6614FA5100CEFA7C" bold="true" box="[595,693,1447,1472]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="C996041FFFB3FFCE6614FA5100CEFA7C" box="[595,693,1447,1472]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(see 
<bibRefCitation id="84401887FFB3FFCE6746FA5E062DFA7C" author="Kornev, P. N. &amp; Chertoprud, E. C." box="[769,1110,1447,1472]" pageId="29" pageNumber="30" refId="ref49254" refString="Kornev, P. N. &amp; Chertoprud, E. C. (2008) Copepod Crustaceans of the Order Harpacticoida of the White Sea: Morphology, Systematics, Ecology. Biology Faculty, Moscow State University, Tovarishchestvo Nauchnikh Izdanii KMK, Moscow, 379 pp. [in Russian]" type="book" year="2008">Kornev &amp; Chertoprud 2008</bibRefCitation>
). However, this may be a plesiomorphic character, as a similar fifth leg endopod can be found in several lineages of the genus 
<taxonomicName id="27D11EF5FFB3FFCE616AFA3A035DF9B4" authority="Brady, 1869" authorityName="Brady" authorityYear="1869" class="Maxillopoda" family="Miraciidae" genus="Delavalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="D2A5B964FFB3FFCE616AFA3A07E0FA58" box="[1325,1435,1484,1508]" italics="true" pageId="29" pageNumber="30">Delavalia</emphasis>
Brady, 1869
</taxonomicName>
. Unfortunately, both 
<taxonomicName id="27D11EF5FFB3FFCE6659FA06014BF9B4" authorityName="Karanovic &amp; Kim" authorityYear="2014" baseAuthorityName="Lang" baseAuthorityYear="1948" box="[542,816,1520,1544]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="hanstromi">
<emphasis id="D2A5B964FFB3FFCE6659FA06014BF9B4" box="[542,816,1520,1544]" italics="true" pageId="29" pageNumber="30">Wellstenhelia hanstromi</emphasis>
</taxonomicName>
and 
<taxonomicName id="27D11EF5FFB3FFCE6729FA0606F6F9B4" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[878,1165,1520,1544]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="melpomene">
<emphasis id="D2A5B964FFB3FFCE6729FA0606F6F9B4" box="[878,1165,1520,1544]" italics="true" pageId="29" pageNumber="30">Wellstenhelia melpomene</emphasis>
</taxonomicName>
are known so far only from a very limited set of female morphological characters (
<bibRefCitation id="84401887FFB3FFCE6715F9E201A8F991" author="Lang, K." box="[850,979,1556,1581]" pageId="29" pageNumber="30" refId="ref49308" refString="Lang, K. (1948) Monographie der Harpacticiden, 1 - 2. Nordiska Bokhandeln, Lund, 1682 pp." type="book" year="1948">Lang 1948</bibRefCitation>
; 
<bibRefCitation id="84401887FFB3FFCE67A6F9E20759F991" author="Kornev, P. N. &amp; Chertoprud, E. C." box="[993,1314,1556,1581]" pageId="29" pageNumber="30" refId="ref49254" refString="Kornev, P. N. &amp; Chertoprud, E. C. (2008) Copepod Crustaceans of the Order Harpacticoida of the White Sea: Morphology, Systematics, Ecology. Biology Faculty, Moscow State University, Tovarishchestvo Nauchnikh Izdanii KMK, Moscow, 379 pp. [in Russian]" type="book" year="2008">Kornev &amp; Chertoprud 2008</bibRefCitation>
) and most features cannot be compared. Even so, there is no doubt that all three represent separate species, as 
<taxonomicName id="27D11EF5FFB3FFCE6141F9CE0373F9C8" authorityName="Karanovic &amp; Kim" authorityYear="2014" baseAuthorityName="Lang" baseAuthorityYear="1948" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="hanstromi">
<emphasis id="D2A5B964FFB3FFCE6141F9CE0373F9C8" italics="true" pageId="29" pageNumber="30">Wellstenhelia hanstromi</emphasis>
</taxonomicName>
has much longer caudal rami and a shorter seta on the first endopodal segment of the fourth leg than the other two, and 
<taxonomicName id="27D11EF5FFB3FFCE6578F9760020F924" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[319,603,1664,1688]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="melpomene">
<emphasis id="D2A5B964FFB3FFCE6578F9760020F924" box="[319,603,1664,1688]" italics="true" pageId="29" pageNumber="30">Wellstenhelia melpomene</emphasis>
</taxonomicName>
has a longer exopod and all setae on the fifth leg, while 
<taxonomicName id="27D11EF5FFB3FFCE6092F97607E7F924" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1237,1436,1664,1688]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="clio">
<emphasis id="D2A5B964FFB3FFCE6092F97607E7F924" box="[1237,1436,1664,1688]" italics="true" pageId="29" pageNumber="30">Wellstenhelia clio</emphasis>
</taxonomicName>
has a shorter endopod of the first leg. In the absence of other evidence we have to assume that 
<taxonomicName id="27D11EF5FFB3FFCE60FCF95207F9F900" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1211,1410,1700,1724]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="clio">
<emphasis id="D2A5B964FFB3FFCE60FCF95207F9F900" box="[1211,1410,1700,1724]" italics="true" pageId="29" pageNumber="30">Wellstenhelia clio</emphasis>
</taxonomicName>
is probably morphologically most similar to 
<taxonomicName id="27D11EF5FFB3FFCE663AF93E01E0F95C" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[637,923,1736,1760]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="melpomene">
<emphasis id="D2A5B964FFB3FFCE663AF93E01E0F95C" box="[637,923,1736,1760]" italics="true" pageId="29" pageNumber="30">Wellstenhelia melpomene</emphasis>
</taxonomicName>
, as these two species also have very similar caudal rami.
</paragraph>
<paragraph id="E06E6576FFB3FFCD6480F8F90115FF69" blockId="29.[151,1436,151,2013]" lastBlockId="30.[151,1436,151,213]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
Other congeners can be easily distinguished from 
<taxonomicName id="27D11EF5FFB3FFCE66B4F8E601C2F894" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[755,953,1808,1832]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="clio">
<emphasis id="D2A5B964FFB3FFCE66B4F8E601C2F894" box="[755,953,1808,1832]" italics="true" pageId="29" pageNumber="30">Wellstenhelia clio</emphasis>
</taxonomicName>
by many characters. 
<taxonomicName id="27D11EF5FFB3FFCE60E2F8E607E7F894" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1189,1436,1808,1832]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="calliope" status="sp. nov.">
<emphasis id="D2A5B964FFB3FFCE60E2F8E607E7F894" box="[1189,1436,1808,1832]" italics="true" pageId="29" pageNumber="30">Wellstenhelia calliope</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFB3FFCE64D0F8C2028AF8F1" bold="true" box="[151,241,1844,1869]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="C996041FFFB3FFCE64D0F8C2028AF8F1" box="[151,241,1844,1869]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="27D11EF5FFB3FFCE656EF8C20166F8F1" authority="(Ma &amp; Li, 2011)" authorityName="Karanovic &amp; Kim" authorityYear="2014" baseAuthorityName="Ma &amp; Li" baseAuthorityYear="2011" box="[297,797,1844,1869]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="qingdaoensis" status="comb. nov.">
<emphasis id="D2A5B964FFB3FFCE656EF8C20021F8F0" box="[297,602,1844,1868]" italics="true" pageId="29" pageNumber="30">Wellstenhelia qingdaoensis</emphasis>
(
<bibRefCitation id="84401887FFB3FFCE662EF8C2016EF8F1" author="Ma, L. &amp; Li, X. - Z." box="[617,789,1844,1869]" pageId="29" pageNumber="30" pagination="1085 - 1097" refId="ref49609" refString="Ma, L. &amp; Li, X. - Z. (2011) Delavalia qingdaoensis sp. nov. (Harpacticoida, Miraciidae), a new copepod species from Jiaozhou Bay, Yellow Sea. Crustaceana, 84, 1085 - 1097. http: // dx. doi. org / 10.1163 / 001121611 x 584334" type="journal article" year="2011">Ma &amp; Li, 2011</bibRefCitation>
)
</taxonomicName>
<emphasis id="D2A5B964FFB3FFCE6763F8C201DDF8F1" bold="true" box="[804,934,1844,1869]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="C996041FFFB3FFCE6763F8C201DDF8F1" box="[804,934,1844,1869]" pageId="29" pageNumber="30" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
have much longer caudal rami and a shorter innermost seta on the female fifth leg endopod; 
<taxonomicName id="27D11EF5FFB3FFCE6687F8AE01C8F8CC" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[704,947,1880,1904]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="euterpe" status="sp. nov.">
<emphasis id="D2A5B964FFB3FFCE6687F8AE01C8F8CC" box="[704,947,1880,1904]" italics="true" pageId="29" pageNumber="30">Wellstenhelia euterpe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C996041FFFB3FFCE67FBF8AE0674F8D3" box="[956,1039,1880,1903]" pageId="29" pageNumber="30" rank="species">sp. nov</taxonomicNameLabel>
has much shorter caudal rami and only three setae on the female fifth leg endopod; while 
<taxonomicName id="27D11EF5FFB3FFCE6759F88A0182F828" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[798,1017,1916,1940]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="erato" status="sp. nov.">
<emphasis id="D2A5B964FFB3FFCE6759F88A0182F828" box="[798,1017,1916,1940]" italics="true" pageId="29" pageNumber="30">Wellstenhelia erato</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFB3FFCE6044F88A061BF829" bold="true" box="[1027,1120,1916,1941]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="C996041FFFB3FFCE6044F88A061BF829" box="[1027,1120,1916,1941]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="27D11EF5FFB3FFCE60DBF88A034AF805" authority="(Soyer, 1971)" authorityName="Karanovic &amp; Kim" authorityYear="2014" baseAuthorityName="Soyer" baseAuthorityYear="1971" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="bocqueti" status="comb. nov.">
<emphasis id="D2A5B964FFB3FFCE60DBF88A07E7F828" box="[1180,1436,1916,1940]" italics="true" pageId="29" pageNumber="30">Wellstenhelia bocqueti</emphasis>
(
<bibRefCitation id="84401887FFB3FFCE64D8F8560352F805" author="Soyer, J." box="[159,297,1952,1977]" pageId="29" pageNumber="30" pagination="263 - 280" refId="ref50016" refString="Soyer, J. (1971) Contribution a l'etude des Copepodes Harpacticoides de Mediterranee occidentale, 5. Stenhelia (Delavalia) coineauae n. sp. Stenhelia (D.) bocqueti n. sp. et Typhlamphiascus bouligandi n. sp. (Diosaccidae, Sars). Vie et Milieu, 22, 263 - 280." type="journal article" year="1971">Soyer, 1971</bibRefCitation>
)
</taxonomicName>
<emphasis id="D2A5B964FFB3FFCE657DF85603C7F805" bold="true" box="[314,444,1952,1977]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="C996041FFFB3FFCE657DF85603C7F805" box="[314,444,1952,1977]" pageId="29" pageNumber="30" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
have a much shorter innermost seta on the female fifth leg endopod. Each species can additionally be distinguished from 
<taxonomicName id="27D11EF5FFB3FFCE6667F832009CF860" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[544,743,1988,2012]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="clio">
<emphasis id="D2A5B964FFB3FFCE6667F832009CF860" box="[544,743,1988,2012]" italics="true" pageId="29" pageNumber="30">Wellstenhelia clio</emphasis>
</taxonomicName>
by some other feature in the proportion of armature elements or ornamentation of somites. Numerous differences between this species and 
<taxonomicName id="27D11EF5FFB0FFCD604EFF6E0779FF0C" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1033,1282,152,176]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FFB0FFCD604EFF6E0779FF0C" box="[1033,1282,152,176]" italics="true" pageId="30" pageNumber="31">Wellstenhelia calliope</emphasis>
</taxonomicName>
are indicated by arrowheads in 
<figureCitation id="78EA79F3FFB0FFCD6518FF4A004FFF6F" box="[351,564,188,213]" captionStart="FIGURE 12" captionStartId="24.[151,249,1916,1938]" captionTargetBox="[190,1408,189,1892]" captionTargetId="figure-89@24.[185,1408,171,1899]" captionTargetPageId="24" captionText="FIGURE 12. Wellstenhelia clio sp. nov., line drawings, female holotype: A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, dissected and flattened, dorsal view; D, endopod of third swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, ¥) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910592" httpUri="https://zenodo.org/record/4910592/files/figure.png" pageId="30" pageNumber="31">Figs. 12A, B, C, D</figureCitation>
, 
<figureCitation id="78EA79F3FFB0FFCD6678FF4A000AFF69" box="[575,625,188,213]" captionStart="FIGURE 13" captionStartId="25.[151,249,1906,1928]" captionTargetBox="[198,1393,208,1882]" captionTargetId="figure-79@25.[190,1397,193,1885]" captionTargetPageId="25" captionText="FIGURE 13. Wellstenhelia clio sp. nov., line drawings, female holotype: A, abdomen, ventral view; B, left side of cephalothoracic shield, dissected and flattened; C pleuron of third pedigerous somite, dissected and flattened; D, mandibular palp, anterior view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910594" httpUri="https://zenodo.org/record/4910594/files/figure.png" pageId="30" pageNumber="31">13A</figureCitation>
, 
<figureCitation id="78EA79F3FFB0FFCD663AFF4A0091FF6F" box="[637,746,188,213]" captionStart="FIGURE 14" captionStartId="26.[151,249,1890,1912]" captionTargetBox="[199,1389,186,1864]" captionTargetId="figure-116@26.[186,1400,175,1868]" captionTargetPageId="26" captionText="FIGURE 14. Wellstenhelia clio sp. nov., line drawings, A–D, female holotype, E–F, male allotype: A, pleuron of fourth pedigerous somite, dissected and flattened; B, fourth swimming leg without rami, anterior view; C, fifth leg, dissected and flattened, anterior view; D, sixth leg on genital somite, ventro-lateral view; E, cephalothoracic shield, lateral view; F, antennula, dorsal view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Currency symbols ($, £, €) indicating unique sensilla and pores. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910596" httpUri="https://zenodo.org/record/4910596/files/figure.png" pageId="30" pageNumber="31">14B, C, F</figureCitation>
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<figureCitation id="78EA79F3FFB0FFCD66B1FF4A0110FF69" box="[758,875,188,213]" captionStart="FIGURE 15" captionStartId="27.[151,249,1921,1943]" captionTargetBox="[195,1396,205,1892]" captionTargetId="figure-74@27.[186,1401,193,1900]" captionTargetPageId="27" captionText="FIGURE 15. Wellstenhelia clio sp. nov., line drawings, male allotype: A, urosome, ventral view; B, urosome, lateral view; C, pleuron of second pedigerous somite, dissected and flattened. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910598" httpUri="https://zenodo.org/record/4910598/files/figure.png" pageId="30" pageNumber="31">15A, B, C</figureCitation>
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</paragraph>
</subSubSection>
</treatment>
</document>