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<document id="E22CEDB78DA7EC1787D2F876E501F820" ID-CLB-Dataset="5789" ID-DOI="10.11646/zootaxa.3783.1.1" ID-GBIF-Dataset="414491b2-5167-4605-95ab-f679bf476618" ID-ISSN="1175-5326" ID-Zenodo-Dep="4910562" ID-ZooBank="E6155BDC-AEAE-475D-BC83-61B3B863344C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623122599895" checkinUser="felipe" docAuthor="Karanovic, Tomislav &amp; Kim, Kichoon" docDate="2014" docId="6878D460FF9BFF9364D0FAE102A7FCE1" docLanguage="en" docName="zt03783p096.pdf" docOrigin="Zootaxa 3783 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Itostenhelia polyhymnia Karanovic &amp; Kim 2014, sp. nov." docType="treatment" docVersion="7" lastPageNumber="65" masterDocId="9441AC18FFAEFFD36447FFF6027BFFBC" masterDocTitle="New insights into polyphyly of the harpacticoid genus Delavalia (Crustacea, Copepoda) through morphological and molecular study of an unprecedented diversity of sympatric species in a small South Korean bay" masterLastPageNumber="96" masterPageNumber="1" pageNumber="54" updateTime="1699004038958" updateUser="ExternalLinkService">
<mods:mods id="1EAA56247229BEEF290A477AF9A5940D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="382976DB84495BCF84C35C173A5D3046">
<mods:title id="C041657EE9AA7654995DBCAF9AC1F4AB">New insights into polyphyly of the harpacticoid genus Delavalia (Crustacea, Copepoda) through morphological and molecular study of an unprecedented diversity of sympatric species in a small South Korean bay</mods:title>
</mods:titleInfo>
<mods:name id="D66748020BB227631619DD03147C5BB6" type="personal">
<mods:role id="FCE76380D55F502743627D39DDB9A260">
<mods:roleTerm id="2A3C9ADE14CA3DBAE40FD59841B207A3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9CC66B8A5E90FA73AC43FF470118C7DC">Karanovic, Tomislav</mods:namePart>
<mods:nameIdentifier id="EE2C3CD9BA5B8A4B15D0F4D05D545465" type="email">aranovic@utas.edu.au</mods:nameIdentifier>
</mods:name>
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</mods:role>
<mods:namePart id="FBF24F801DA0A1C1B2055DA665CE3065">Kim, Kichoon</mods:namePart>
</mods:name>
<mods:typeOfResource id="1C9F5FAE02975232A015605824BD2E0D">text</mods:typeOfResource>
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<mods:titleInfo id="4E9A157DDF9AE2DB158844032C89CF3A">
<mods:title id="6C3BC769ABE1BC75D4C873F6C21D024E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A65498DDD34840E1D67519B5FC775476">
<mods:date id="1011ECBFFA7F24A5ECFFE96B5363CB97">2014</mods:date>
<mods:detail id="D104B607BB7228FB9A9DA778D04F5624" type="pubDate">
<mods:number id="B81470F52C7C8E60F8559DBA62A1ABC7">2014-03-25</mods:number>
</mods:detail>
<mods:detail id="E6996A3B121A0E1B993FBF78435F9AD2" type="volume">
<mods:number id="7E6BC818EC01037BF74CEE238C90E679">3783</mods:number>
</mods:detail>
<mods:detail id="0CFA87B68F9BF5F57A51334A7E1BA5FC" type="issue">
<mods:number id="E4AAE7386B62055CD5412E59A719293C">1</mods:number>
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<mods:start id="798BC536525E595554FAAFFF0DF78082">1</mods:start>
<mods:end id="46311CA4C5E7ECB28A2645B66188B7B5">96</mods:end>
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<mods:classification id="8FCDE6B8C1B8CEB26D9DE38996E59E8C">journal article</mods:classification>
<mods:identifier id="4DFBD2204FFC4784E2BC9685EEB1801F" type="CLB-Dataset">5789</mods:identifier>
<mods:identifier id="BB560634AA1CF79B1D1EB37F3A6BFD5F" type="DOI">10.11646/zootaxa.3783.1.1</mods:identifier>
<mods:identifier id="87F1515AF094FCE3E784F65AB144C8E5" type="GBIF-Dataset">414491b2-5167-4605-95ab-f679bf476618</mods:identifier>
<mods:identifier id="097B2932F3690DAEF5277B2DBE39A4BD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="537C5556A1DE04E806B9F434EDB304DA" type="Zenodo-Dep">4910562</mods:identifier>
<mods:identifier id="2986B3A131367B63E76A39A6EE149D1E" type="ZooBank">E6155BDC-AEAE-475D-BC83-61B3B863344C</mods:identifier>
</mods:mods>
<treatment id="6878D460FF9BFF9364D0FAE102A7FCE1" ID-DOI="http://doi.org/10.5281/zenodo.5062495" ID-GBIF-Taxon="183238043" ID-Zenodo-Dep="5062495" LSID="urn:lsid:plazi:treatment:6878D460FF9BFF9364D0FAE102A7FCE1" httpUri="http://treatment.plazi.org/id/6878D460FF9BFF9364D0FAE102A7FCE1" lastPageId="64" lastPageNumber="65" pageId="53" pageNumber="54">
<subSubSection id="A8CB36FDFF9BFFE664D0FAE1005BFA8E" box="[151,544,1303,1330]" pageId="53" pageNumber="54" type="nomenclature">
<paragraph id="E06E6576FF9BFFE664D0FAE1005BFA8E" blockId="53.[151,544,1303,1363]" box="[151,544,1303,1330]" pageId="53" pageNumber="54">
<heading id="BB26D21AFF9BFFE664D0FAE1005BFA8E" bold="true" box="[151,544,1303,1330]" fontSize="11" level="2" pageId="53" pageNumber="54" reason="4">
<emphasis id="D2A5B964FF9BFFE664D0FAE1005BFA8E" bold="true" box="[151,544,1303,1330]" pageId="53" pageNumber="54">
<taxonomicName id="27D11EF5FF9BFFE664D0FAE103C0FA8D" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[151,443,1303,1329]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="polyhymnia" status="sp. nov.">
<emphasis id="D2A5B964FF9BFFE664D0FAE103C0FA8D" bold="true" box="[151,443,1303,1329]" italics="true" pageId="53" pageNumber="54">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C996041FFF9BFFE66585FAEE005BFA8E" box="[450,544,1304,1330]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A8CB36FDFF9BFFE664D0FACC0355FAEF" box="[151,302,1338,1363]" pageId="53" pageNumber="54" type="description">
<paragraph id="E06E6576FF9BFFE664D0FACC0355FAEF" blockId="53.[151,544,1303,1363]" box="[151,302,1338,1363]" pageId="53" pageNumber="54">
(
<figureCitation id="78EA79F3FF9BFFE664D8FACC035DFAEF" box="[159,294,1338,1363]" captionStart-0="FIGURE 34" captionStart-1="FIGURE 35" captionStart-2="FIGURE 36" captionStart-3="FIGURE 37" captionStart-4="FIGURE 38" captionStart-5="FIGURE 39" captionStart-6="FIGURE 40" captionStart-7="FIGURE 41" captionStart-8="FIGURE 42" captionStartId-0="54.[151,249,1885,1907]" captionStartId-1="55.[151,249,1829,1851]" captionStartId-2="56.[151,249,1889,1911]" captionStartId-3="57.[151,249,1915,1937]" captionStartId-4="58.[151,249,1884,1906]" captionStartId-5="59.[151,249,1559,1581]" captionStartId-6="61.[151,249,1596,1618]" captionStartId-7="62.[151,249,1560,1582]" captionStartId-8="65.[151,249,1858,1880]" captionTargetBox-0="[209,1381,197,1846]" captionTargetBox-1="[218,1368,192,1805]" captionTargetBox-2="[185,1400,184,1868]" captionTargetBox-3="[189,1399,185,1890]" captionTargetBox-4="[192,1391,186,1878]" captionTargetBox-5="[151,1436,193,1538]" captionTargetBox-6="[151,1436,218,1563]" captionTargetBox-7="[151,1436,193,1539]" captionTargetBox-8="[215,1371,206,1826]" captionTargetId-0="figure-102@54.[202,1381,193,1850]" captionTargetId-1="figure-138@55.[210,1371,179,1809]" captionTargetId-2="figure-101@56.[183,1402,173,1873]" captionTargetId-3="figure-91@57.[181,1405,175,1893]" captionTargetId-4="figure-97@58.[183,1404,179,1893]" captionTargetId-5="figure-211@59.[151,1436,193,1538]" captionTargetId-6="figure-185@61.[151,1436,218,1563]" captionTargetId-7="figure-244@62.[151,1436,193,1539]" captionTargetId-8="figure-105@65.[207,1371,193,1832]" captionTargetPageId-0="54" captionTargetPageId-1="55" captionTargetPageId-2="56" captionTargetPageId-3="57" captionTargetPageId-4="58" captionTargetPageId-5="59" captionTargetPageId-6="61" captionTargetPageId-7="62" captionTargetPageId-8="65" captionText-0="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." captionText-1="FIGURE 35. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, right side of cephalothoracic shiled, dissected, flattened, and partly broken; B, pleurons of third and fourth pedigerous somites, dissected and flattened; C, genital double-somite, ventral view; D, genital double-somite, lateral view; E, syncoxa of mandibula, antero-ventral view; F, syncoxa of mandibula, posterior view; G, mandibular palp, anterior view; H, maxillula, posterior view; I, mandibula, anterior view; J, maxilliped, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I, V) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." captionText-2="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." captionText-3="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." captionText-4="FIGURE 38. Itostenhelia polyhymnia sp. nov., line drawings, male allotype: A, cephalothoracic shield and pleurons of second to fourth pedigerous somites, lateral view; B, urosome, ventral view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." captionText-5="FIGURE 39. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, female paratype 1: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, pedigerous somites, dorsal view; D, first urosomite and genital double-somite, dorsal view; E, anal somite and caudal rami, dorsal view; F, antennula, dorsal view. Scale bars 30 µm (E, F), 40 µm (D), 50 µm (B, C), and 200 µm (A)." captionText-6="FIGURE 40. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–E, female paratype 2, F, female paratype 3: A, habitus, lateral view; B, cephalothorax, lateral view; C pedigerous somites, lateral view; D, first urosomite and genital double-somite, lateral view (exopod of fifth leg broken off); E, anal somite and caudal rami, lateral view; F, fifth legs and genital double-somite with one empty egg-sack, ventral view. Scale bars 50 µm (C, D, E, F), 100 µm (B), and 200 µm (A)." captionText-7="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." captionText-8="FIGURE 42. Itostenhelia polyhymnia sp. nov., L-form, line drawings, female holotype: A, habitus, lateral view; B, cephalothoracic shield, lateral view; C, labrum, dissected and flattened, anterior view; D, baseoendopod of fifth leg, anterior view. Arabic numerals indicating sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shape (◊) indicating sensillum presumably homologous to that in Wellstenhelia euterpe. Roman numerals (I, IV–VII) indicating sensilla and pores presumably homologous to those in Itostenhelia polyhymnia. Arrowheads pointing most prominent specific features. Scale bars 100 µm." figureDoi-0="http://doi.org/10.5281/zenodo.4910636" figureDoi-1="http://doi.org/10.5281/zenodo.4910638" figureDoi-2="http://doi.org/10.5281/zenodo.4910640" figureDoi-3="http://doi.org/10.5281/zenodo.4910642" figureDoi-4="http://doi.org/10.5281/zenodo.4910644" figureDoi-5="http://doi.org/10.5281/zenodo.4910646" figureDoi-6="http://doi.org/10.5281/zenodo.4910648" figureDoi-7="http://doi.org/10.5281/zenodo.4910650" figureDoi-8="http://doi.org/10.5281/zenodo.4910652" httpUri-0="https://zenodo.org/record/4910636/files/figure.png" httpUri-1="https://zenodo.org/record/4910638/files/figure.png" httpUri-2="https://zenodo.org/record/4910640/files/figure.png" httpUri-3="https://zenodo.org/record/4910642/files/figure.png" httpUri-4="https://zenodo.org/record/4910644/files/figure.png" httpUri-5="https://zenodo.org/record/4910646/files/figure.png" httpUri-6="https://zenodo.org/record/4910648/files/figure.png" httpUri-7="https://zenodo.org/record/4910650/files/figure.png" httpUri-8="https://zenodo.org/record/4910652/files/figure.png" pageId="53" pageNumber="54">Figs. 34–42</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A8CB36FDFF9BFFE664D0FA7407E6F867" pageId="53" pageNumber="54" type="materials_examined">
<paragraph id="E06E6576FF9BFFE664D0FA7403E1FA03" blockId="53.[151,1437,1410,2011]" pageId="53" pageNumber="54">
<materialsCitation id="50B96F2BFF9BFFE664D0FA7403EEFA03" ID-GBIF-Occurrence="3313373302" country="South Korea" latitude="34.897057" location="South Sea" longLatPrecision="1" longitude="127.75772" municipality="Gwangyang Bay" pageId="53" pageNumber="54" specimenCount="1" typeStatus="holotype">
<emphasis id="D2A5B964FF9BFFE664D0FA740342FA27" bold="true" box="[151,313,1410,1435]" pageId="53" pageNumber="54">
<typeStatus id="3F6ADBD4FF9BFFE664D0FA7402A9FA27" box="[151,210,1410,1435]" pageId="53" pageNumber="54">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="98C625E6FF9BFFE66504FA7403AEFA26" box="[323,469,1410,1435]" name="South Korea" pageId="53" pageNumber="54">South Korea</collectingCountry>
,
<location id="E50E33ADFF9BFFE665A3FA740020FA27" LSID="urn:lsid:plazi:treatment:6878D460FF9BFF9364D0FAE102A7FCE1:E50E33ADFF9BFFE665A3FA740020FA27" box="[484,603,1410,1435]" country="South Korea" latitude="34.897057" longLatPrecision="1" longitude="127.75772" municipality="Gwangyang Bay" name="South Sea" pageId="53" pageNumber="54">South Sea</location>
,
<collectingMunicipality id="000AFF0CFF9BFFE6662DFA740152FA27" box="[618,809,1410,1435]" pageId="53" pageNumber="54">Gwangyang Bay</collectingMunicipality>
, sampling station 10, muddy sediments,
<geoCoordinate id="85E503B1FF9BFFE66142FA7407E7FA27" box="[1285,1436,1410,1435]" degrees="34.897056" direction="north" orientation="latitude" pageId="53" pageNumber="54" precision="1" value="34.897057">34.897056°N</geoCoordinate>
<geoCoordinate id="85E503B1FF9BFFE664D0FA500343FA03" box="[151,312,1446,1471]" degrees="127.757722" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="1" value="127.75772">127.757722°E</geoCoordinate>
(
<figureCitation id="78EA79F3FF9BFFE66500FA5003F2FA03" box="[327,393,1446,1471]" captionStart="FIGURE 1" captionStartId="7.[151,249,1481,1503]" captionTargetBox="[151,1436,589,1460]" captionTargetId="figure-399@7.[151,1436,589,1460]" captionTargetPageId="7" captionText="FIGURE 1. Map of 17 sampling locations in Gwangyang Bay. Inset showing location of the bay in South Korea. All maps from Google Earth." figureDoi="http://doi.org/10.5281/zenodo.4910564" httpUri="https://zenodo.org/record/4910564/files/figure.png" pageId="53" pageNumber="54">Fig. 1</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="E06E6576FF9BFFE66480FA3C0615F9CF" blockId="53.[151,1437,1410,2011]" pageId="53" pageNumber="54">
<emphasis id="D2A5B964FF9BFFE66480FA3C03BDFA5F" bold="true" box="[199,454,1482,1507]" pageId="53" pageNumber="54">Specimens examined.</emphasis>
Female
<typeStatus id="3F6ADBD4FF9BFFE66662FA3C00FCFA5F" box="[549,647,1482,1507]" pageId="53" pageNumber="54" type="holotype">holotype</typeStatus>
dissected on one slide (collection number NIBRIV0000232691), male
<typeStatus id="3F6ADBD4FF9BFFE664D0FA18028AF9BB" box="[151,241,1518,1543]" pageId="53" pageNumber="54" type="allotype">allotype</typeStatus>
dissected on one slide (collection number NIBRIV0000232692), female
<typeStatus id="3F6ADBD4FF9BFFE6607FFA1906E3F9BB" box="[1080,1176,1519,1543]" pageId="53" pageNumber="54" type="paratype">paratype</typeStatus>
dissected on one slide (collection number NIBRIV0000232693),
<specimenCount id="F6D7AEFFFF9BFFE66632F9E500A1F997" box="[629,730,1554,1579]" pageId="53" pageNumber="54" type="male">one male</specimenCount>
<typeStatus id="3F6ADBD4FF9BFFE666A7F9E5013BF997" box="[736,832,1555,1579]" pageId="53" pageNumber="54" type="paratype">paratype</typeStatus>
and
<specimenCount id="F6D7AEFFFF9BFFE66731F9E4067AF997" box="[886,1025,1554,1579]" pageId="53" pageNumber="54" type="female">three female</specimenCount>
<typeStatus id="3F6ADBD4FF9BFFE66040F9E50608F997" box="[1031,1139,1555,1579]" pageId="53" pageNumber="54" type="paratype">paratypes</typeStatus>
together on one SEM stub (collection number NIBRIV0000232694),
<specimenCount id="F6D7AEFFFF9BFFE666C4F9C00088F9F3" box="[643,755,1590,1615]" pageId="53" pageNumber="54" type="male">four male</specimenCount>
<typeStatus id="3F6ADBD4FF9BFFE666B9F9C10111F9F3" box="[766,874,1591,1615]" pageId="53" pageNumber="54" type="paratype">paratypes</typeStatus>
and
<specimenCount id="F6D7AEFFFF9BFFE667EFF9C1063BF9F3" box="[936,1088,1590,1615]" pageId="53" pageNumber="54" type="female">seven female</specimenCount>
<typeStatus id="3F6ADBD4FF9BFFE6600CF9C106CDF9F3" box="[1099,1206,1591,1615]" pageId="53" pageNumber="54" type="paratype">paratypes</typeStatus>
together in ethanol (collection number NIBRIV0000232695), type locality,
<date id="946F43B6FF9BFFE6674AF9AC01ACF9CF" box="[781,983,1626,1651]" pageId="53" pageNumber="54" value="2012-02-18">18 February 2012</date>
, leg. K. Kim.
</paragraph>
<paragraph id="E06E6576FF9BFFE66480F98807F7F907" blockId="53.[151,1437,1410,2011]" pageId="53" pageNumber="54">
One male
<typeStatus id="3F6ADBD4FF9BFFE6657DF98903E1F92B" box="[314,410,1663,1687]" pageId="53" pageNumber="54" type="paratype">paratype</typeStatus>
and
<specimenCount id="F6D7AEFFFF9BFFE66596F988003BF92B" box="[465,576,1662,1687]" pageId="53" pageNumber="54" type="female">15 female</specimenCount>
<typeStatus id="3F6ADBD4FF9BFFE6660FF98900C8F92B" box="[584,691,1663,1687]" pageId="53" pageNumber="54" type="paratype">paratypes</typeStatus>
together in ethanol (collection number NIBRIV0000232696),
<specimenCount id="F6D7AEFFFF9BFFE66134F9890298F907" pageId="53" pageNumber="54" type="female">one female</specimenCount>
destroyed for DNA sequence (amplification unsuccessful), type locality,
<date id="946F43B6FF9BFFE6605DF954068EF907" box="[1050,1269,1698,1723]" pageId="53" pageNumber="54" value="2013-11-18">18 November 2013</date>
, leg. K. Kim.
</paragraph>
<paragraph id="E06E6576FF9BFFE66480F93002ABF89B" blockId="53.[151,1437,1410,2011]" pageId="53" pageNumber="54">
Three female
<typeStatus id="3F6ADBD4FF9BFFE6652DF93103ADF963" box="[362,470,1735,1759]" pageId="53" pageNumber="54" type="paratype">paratypes</typeStatus>
(one ovigerous) together in ethanol (collection number NIBRIV0000232697),
<specimenCount id="F6D7AEFFFF9BFFE66137F9300295F8BF" pageId="53" pageNumber="54" type="female">five females</specimenCount>
destroyed for DNA sequence (two successful, Codes 0176 &amp; 0273), type locality,
<date id="946F43B6FF9BFFE660EEF91C073AF8BF" box="[1193,1345,1770,1795]" pageId="53" pageNumber="54" value="2012-07-30">30 July 2012</date>
, leg. K. Kim.
</paragraph>
<paragraph id="E06E6576FF9BFFE66480F8C40312F82F" blockId="53.[151,1437,1410,2011]" pageId="53" pageNumber="54">
<materialsCitation id="50B96F2BFF9BFFE66480F8C40312F82F" ID-GBIF-Occurrence="3313373316" collectingDate="2012-11-18" collectorName="K. Kim." country="South Korea" county="Gwangyang Bay" latitude="34.95139" location="Three" longLatPrecision="1" longitude="127.73436" municipality="South Sea" pageId="53" pageNumber="54" specimenCount="1">
<location id="E50E33ADFF9BFFE66480F8C40373F8F7" LSID="urn:lsid:plazi:treatment:6878D460FF9BFF9364D0FAE102A7FCE1:E50E33ADFF9BFFE66480F8C40373F8F7" box="[199,264,1842,1867]" country="South Korea" county="Gwangyang Bay" latitude="34.95139" longLatPrecision="1" longitude="127.73436" municipality="South Sea" name="Three" pageId="53" pageNumber="54">Three</location>
females (two ovigerous) destroyed for DNA sequence (amplification unsuccessful),
<collectingCountry id="98C625E6FF9BFFE66087F8C40734F8F6" box="[1216,1359,1842,1867]" name="South Korea" pageId="53" pageNumber="54">South Korea</collectingCountry>
,
<collectingMunicipality id="000AFF0CFF9BFFE6611DF8C402B9F8D3" pageId="53" pageNumber="54">South Sea</collectingMunicipality>
,
<collectingCounty id="090F1DFAFF9BFFE66488F8A003F6F8D3" box="[207,397,1878,1903]" pageId="53" pageNumber="54">Gwangyang Bay</collectingCounty>
, sampling station 9, muddy sediments,
<geoCoordinate id="85E503B1FF9BFFE66715F8A00193F8D3" box="[850,1000,1878,1903]" degrees="34.951389" direction="north" orientation="latitude" pageId="53" pageNumber="54" precision="1" value="34.95139">34.951389°N</geoCoordinate>
<geoCoordinate id="85E503B1FF9BFFE667B6F8A006E9F8D3" box="[1009,1170,1878,1903]" degrees="127.734361" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="1" value="127.73436">127.734361°E</geoCoordinate>
(
<figureCitation id="78EA79F3FF9BFFE660E3F8A0069CF8D3" box="[1188,1255,1878,1903]" captionStart="FIGURE 1" captionStartId="7.[151,249,1481,1503]" captionTargetBox="[151,1436,589,1460]" captionTargetId="figure-399@7.[151,1436,589,1460]" captionTargetPageId="7" captionText="FIGURE 1. Map of 17 sampling locations in Gwangyang Bay. Inset showing location of the bay in South Korea. All maps from Google Earth." figureDoi="http://doi.org/10.5281/zenodo.4910564" httpUri="https://zenodo.org/record/4910564/files/figure.png" pageId="53" pageNumber="54">Fig. 1</figureCitation>
),
<date id="946F43B6FF9BFFE66146F8A002A9F82F" pageId="53" pageNumber="54" value="2012-11-18">
<collectingDate id="842BBA5EFF9BFFE66146F8A002A9F82F" pageId="53" pageNumber="54" value="2012-11-18">18 November 2012</collectingDate>
</date>
, leg.
<collectorName id="4D2400A0FF9BFFE6654AF88D0312F82F" box="[269,361,1914,1939]" pageId="53" pageNumber="54">K. Kim.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="E06E6576FF9BFFE66480F86807E6F867" blockId="53.[151,1437,1410,2011]" pageId="53" pageNumber="54">
<materialsCitation id="50B96F2BFF9BFFE66480F86807E6F867" ID-GBIF-Occurrence="3313373317" collectingDate="2012-11-18" collectorName="K. Kim." country="South Korea" county="Gwangyang Bay" latitude="34.76889" location="One" longLatPrecision="1" longitude="127.783806" municipality="South Sea" pageId="53" pageNumber="54" specimenCount="1">
<location id="E50E33ADFF9BFFE66480F868028DF80B" LSID="urn:lsid:plazi:treatment:6878D460FF9BFF9364D0FAE102A7FCE1:E50E33ADFF9BFFE66480F868028DF80B" box="[199,246,1950,1975]" country="South Korea" county="Gwangyang Bay" latitude="34.76889" longLatPrecision="1" longitude="127.783806" municipality="South Sea" name="One" pageId="53" pageNumber="54">One</location>
female destroyed for DNA sequence (amplification unsuccessful),
<collectingCountry id="98C625E6FF9BFFE667B2F86806FDF80A" box="[1013,1158,1950,1975]" name="South Korea" pageId="53" pageNumber="54">South Korea</collectingCountry>
,
<collectingMunicipality id="000AFF0CFF9BFFE660D4F868077CF80B" box="[1171,1287,1950,1975]" pageId="53" pageNumber="54">South Sea</collectingMunicipality>
,
<collectingCounty id="090F1DFAFF9BFFE66153F86802BEF867" pageId="53" pageNumber="54">Gwangyang Bay</collectingCounty>
, sampling station 16, muddy sediments,
<geoCoordinate id="85E503B1FF9BFFE666CFF8340164F867" box="[648,799,1986,2011]" degrees="34.768889" direction="north" orientation="latitude" pageId="53" pageNumber="54" precision="1" value="34.76889">34.768889°N</geoCoordinate>
<geoCoordinate id="85E503B1FF9BFFE66762F83401BDF867" box="[805,966,1986,2011]" degrees="127.783806" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="1" value="127.783806">127.783806°E</geoCoordinate>
(
<figureCitation id="78EA79F3FF9BFFE66792F834066DF867" box="[981,1046,1986,2011]" captionStart="FIGURE 1" captionStartId="7.[151,249,1481,1503]" captionTargetBox="[151,1436,589,1460]" captionTargetId="figure-399@7.[151,1436,589,1460]" captionTargetPageId="7" captionText="FIGURE 1. Map of 17 sampling locations in Gwangyang Bay. Inset showing location of the bay in South Korea. All maps from Google Earth." figureDoi="http://doi.org/10.5281/zenodo.4910564" httpUri="https://zenodo.org/record/4910564/files/figure.png" pageId="53" pageNumber="54">Fig. 1</figureCitation>
),
<date id="946F43B6FF9BFFE66069F8340773F867" box="[1070,1288,1986,2011]" pageId="53" pageNumber="54" value="2012-11-18">
<collectingDate id="842BBA5EFF9BFFE66069F8340773F867" box="[1070,1288,1986,2011]" pageId="53" pageNumber="54" value="2012-11-18">18 November 2012</collectingDate>
</date>
, leg.
<collectorName id="4D2400A0FF9BFFE66105F83507E6F867" box="[1346,1437,1986,2011]" pageId="53" pageNumber="54">K. Kim.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="B4AE35FEFF98FFE564D0F8AB01FCF851" ID-DOI="http://doi.org/10.5281/zenodo.4910636" ID-Zenodo-Dep="4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="54" pageNumber="55" startId="54.[151,249,1885,1907]" targetBox="[209,1381,197,1846]" targetPageId="54">
<paragraph id="E06E6576FF98FFE564D0F8AB01FCF851" blockId="54.[151,1437,1885,2030]" pageId="54" pageNumber="55">
<emphasis id="D2A5B964FF98FFE564D0F8AB0359F8CE" bold="true" box="[151,290,1885,1907]" pageId="54" pageNumber="55">FIGURE 34.</emphasis>
<taxonomicName id="27D11EF5FF98FFE5656CF8AB0059F8CF" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[299,546,1885,1907]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="polyhymnia" status="sp. nov.">
<emphasis id="D2A5B964FF98FFE5656CF8AB0059F8CF" box="[299,546,1885,1907]" italics="true" pageId="54" pageNumber="55">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FF98FFE5666CF8AB0004F8CE" bold="true" box="[555,639,1885,1906]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="C996041FFF98FFE5666CF8AB0004F8CE" box="[555,639,1885,1906]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in
<taxonomicName id="27D11EF5FF98FFE56752F86C018FF80C" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[789,1012,1946,1968]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF98FFE56752F86C018FF80C" box="[789,1012,1946,1968]" italics="true" pageId="54" pageNumber="55">Wellstenhelia calliope</emphasis>
</taxonomicName>
. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in
<taxonomicName id="27D11EF5FF98FFE56663F84F0178F873" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[548,771,1977,1999]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="euterpe">
<emphasis id="D2A5B964FF98FFE56663F84F0178F873" box="[548,771,1977,1999]" italics="true" pageId="54" pageNumber="55">Wellstenhelia euterpe</emphasis>
</taxonomicName>
. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm.
</paragraph>
</caption>
<caption id="B4AE35FEFF99FFE464D0F8D3075AF84F" ID-DOI="http://doi.org/10.5281/zenodo.4910638" ID-Zenodo-Dep="4910638" httpUri="https://zenodo.org/record/4910638/files/figure.png" pageId="55" pageNumber="56" startId="55.[151,249,1829,1851]" targetBox="[218,1368,192,1805]" targetPageId="55">
<paragraph id="E06E6576FF99FFE464D0F8D3075AF84F" blockId="55.[151,1436,1829,2035]" pageId="55" pageNumber="56">
<emphasis id="D2A5B964FF99FFE464D0F8D3035EF886" bold="true" box="[151,293,1829,1851]" pageId="55" pageNumber="56">FIGURE 35.</emphasis>
<taxonomicName id="27D11EF5FF99FFE46577F8D30052F887" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[304,553,1829,1851]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="polyhymnia" status="sp. nov.">
<emphasis id="D2A5B964FF99FFE46577F8D30052F887" box="[304,553,1829,1851]" italics="true" pageId="55" pageNumber="56">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FF99FFE46672F8D000F1F887" bold="true" box="[565,650,1830,1851]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="C996041FFF99FFE46672F8D000F1F887" box="[565,650,1830,1851]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, line drawings, female holotype: A, right side of cephalothoracic shiled, dissected, flattened, and partly broken; B, pleurons of third and fourth pedigerous somites, dissected and flattened; C, genital double-somite, ventral view; D, genital double-somite, lateral view; E, syncoxa of mandibula, antero-ventral view; F, syncoxa of mandibula, posterior view; G, mandibular palp, anterior view; H, maxillula, posterior view; I, mandibula, anterior view; J, maxilliped, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in
<taxonomicName id="27D11EF5FF99FFE46153F8560292F868" authorityName="Karanovic &amp; Kim" authorityYear="2014" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF99FFE46153F8560292F868" italics="true" pageId="55" pageNumber="56">Wellstenhelia calliope</emphasis>
</taxonomicName>
. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in
<taxonomicName id="27D11EF5FF99FFE4604EF8480698F868" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1033,1251,1982,2004]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="euterpe">
<emphasis id="D2A5B964FF99FFE4604EF8480698F868" box="[1033,1251,1982,2004]" italics="true" pageId="55" pageNumber="56">Wellstenhelia euterpe</emphasis>
</taxonomicName>
. Roman numerals (I, V) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm.
</paragraph>
</caption>
<caption id="B4AE35FEFF96FFEB64D0F89703D7F84E" ID-DOI="http://doi.org/10.5281/zenodo.4910640" ID-Zenodo-Dep="4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="56" pageNumber="57" startId="56.[151,249,1889,1911]" targetBox="[185,1400,184,1868]" targetPageId="56">
<paragraph id="E06E6576FF96FFEB64D0F89703D7F84E" blockId="56.[151,1436,1889,2034]" pageId="56" pageNumber="57">
<emphasis id="D2A5B964FF96FFEB64D0F897035FF8CA" bold="true" box="[151,292,1889,1911]" pageId="56" pageNumber="57">FIGURE 36.</emphasis>
<taxonomicName id="27D11EF5FF96FFEB6569F897005DF8CB" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[302,550,1889,1911]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="polyhymnia" status="sp. nov.">
<emphasis id="D2A5B964FF96FFEB6569F897005DF8CB" box="[302,550,1889,1911]" italics="true" pageId="56" pageNumber="57">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FF96FFEB6677F89400FEF8CB" bold="true" box="[560,645,1890,1911]" pageId="56" pageNumber="57">
<taxonomicNameLabel id="C996041FFF96FFEB6677F89400FEF8CB" box="[560,645,1890,1911]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm.
</paragraph>
</caption>
<caption id="B4AE35FEFF97FFEA64D0F88D0751F850" ID-DOI="http://doi.org/10.5281/zenodo.4910642" ID-Zenodo-Dep="4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="57" pageNumber="58" startId="57.[151,249,1915,1937]" targetBox="[189,1399,185,1890]" targetPageId="57">
<paragraph id="E06E6576FF97FFEA64D0F88D0751F850" blockId="57.[151,1436,1915,2029]" pageId="57" pageNumber="58">
<emphasis id="D2A5B964FF97FFEA64D0F88D035BF82C" bold="true" box="[151,288,1915,1937]" pageId="57" pageNumber="58">FIGURE 37.</emphasis>
<taxonomicName id="27D11EF5FF97FFEA6562F88D0062F82D" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[293,537,1915,1937]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="polyhymnia" status="sp. nov.">
<emphasis id="D2A5B964FF97FFEA6562F88D0062F82D" box="[293,537,1915,1937]" italics="true" pageId="57" pageNumber="58">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FF97FFEA6667F88D000BF82C" bold="true" box="[544,624,1915,1936]" pageId="57" pageNumber="58">
<taxonomicNameLabel id="C996041FFF97FFEA6667F88D000BF82C" box="[544,624,1915,1936]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in
<taxonomicName id="27D11EF5FF97FFEA6026F84E073FF872" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1121,1348,1976,1998]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF97FFEA6026F84E073FF872" box="[1121,1348,1976,1998]" italics="true" pageId="57" pageNumber="58">Wellstenhelia calliope</emphasis>
</taxonomicName>
. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm.
</paragraph>
</caption>
<caption id="B4AE35FEFF94FFE964D0F8AA0070F850" ID-DOI="http://doi.org/10.5281/zenodo.4910644" ID-Zenodo-Dep="4910644" httpUri="https://zenodo.org/record/4910644/files/figure.png" pageId="58" pageNumber="59" startId="58.[151,249,1884,1906]" targetBox="[192,1391,186,1878]" targetPageId="58">
<paragraph id="E06E6576FF94FFE964D0F8AA0070F850" blockId="58.[151,1436,1884,2029]" pageId="58" pageNumber="59">
<emphasis id="D2A5B964FF94FFE964D0F8AA035BF8CD" bold="true" box="[151,288,1884,1906]" pageId="58" pageNumber="59">FIGURE 38.</emphasis>
<taxonomicName id="27D11EF5FF94FFE96562F8AA0062F8CE" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[293,537,1884,1906]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="polyhymnia" status="sp. nov.">
<emphasis id="D2A5B964FF94FFE96562F8AA0062F8CE" box="[293,537,1884,1906]" italics="true" pageId="58" pageNumber="59">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FF94FFE96667F8AB000BF8CE" bold="true" box="[544,624,1885,1906]" pageId="58" pageNumber="59">
<taxonomicNameLabel id="C996041FFF94FFE96667F8AB000BF8CE" box="[544,624,1885,1906]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, line drawings, male allotype: A, cephalothoracic shield and pleurons of second to fourth pedigerous somites, lateral view; B, urosome, ventral view. Arabic numerals indicate sensilla and pores presumably homologous to those in
<taxonomicName id="27D11EF5FF94FFE965D7F86F0009F813" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[400,626,1945,1967]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF94FFE965D7F86F0009F813" box="[400,626,1945,1967]" italics="true" pageId="58" pageNumber="59">Wellstenhelia calliope</emphasis>
</taxonomicName>
. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in
<taxonomicName id="27D11EF5FF94FFE964D0F84E030DF872" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[151,374,1976,1998]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="euterpe">
<emphasis id="D2A5B964FF94FFE964D0F84E030DF872" box="[151,374,1976,1998]" italics="true" pageId="58" pageNumber="59">Wellstenhelia euterpe</emphasis>
</taxonomicName>
. Roman numerals (I–VII) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm.
</paragraph>
</caption>
<caption id="B4AE35FEFF95FFE864D0F9E10293F935" ID-DOI="http://doi.org/10.5281/zenodo.4910646" ID-Zenodo-Dep="4910646" httpUri="https://zenodo.org/record/4910646/files/figure.png" pageId="59" pageNumber="60" startId="59.[151,249,1559,1581]" targetBox="[151,1436,193,1538]" targetPageId="59">
<paragraph id="E06E6576FF95FFE864D0F9E10293F935" blockId="59.[151,1435,1559,1673]" pageId="59" pageNumber="60">
<emphasis id="D2A5B964FF95FFE864D0F9E1035BF990" bold="true" box="[151,288,1559,1581]" pageId="59" pageNumber="60">FIGURE 39.</emphasis>
<taxonomicName id="27D11EF5FF95FFE86560F9E10060F991" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[295,539,1559,1581]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="polyhymnia" status="sp. nov.">
<emphasis id="D2A5B964FF95FFE86560F9E10060F991" box="[295,539,1559,1581]" italics="true" pageId="59" pageNumber="60">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FF95FFE86665F9E10008F990" bold="true" box="[546,627,1559,1580]" pageId="59" pageNumber="60">
<taxonomicNameLabel id="C996041FFF95FFE86665F9E10008F990" box="[546,627,1559,1580]" pageId="59" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, scanning electron micrographs, female paratype 1: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, pedigerous somites, dorsal view; D, first urosomite and genital double-somite, dorsal view; E, anal somite and caudal rami, dorsal view; F, antennula, dorsal view. Scale bars 30 µm (E, F), 40 µm (D), 50 µm (B, C), and 200 µm (A).
</paragraph>
</caption>
<subSubSection id="A8CB36FDFF95FFE86480F9430326F8AA" pageId="59" pageNumber="60" type="etymology">
<paragraph id="E06E6576FF95FFE86480F9430326F8AA" blockId="59.[151,1436,1717,2030]" pageId="59" pageNumber="60">
<emphasis id="D2A5B964FF95FFE86480F9430335F972" bold="true" box="[199,334,1717,1742]" pageId="59" pageNumber="60">Etymology.</emphasis>
The species is named after Polyhymnia (Ancient Greek: Πολυύµνια), one of nine Muses from Greek mythology, who was a patron of sacred poetry and song. The species name is a noun in apposition (in the nominative case).
</paragraph>
</subSubSection>
<subSubSection id="A8CB36FDFF95FF936480F8D402A7FCE1" lastPageId="64" lastPageNumber="65" pageId="59" pageNumber="60" type="description">
<paragraph id="E06E6576FF95FFEF6480F8D4002BFEA1" blockId="59.[151,1436,1717,2030]" lastBlockId="60.[151,1436,151,2037]" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
<emphasis id="D2A5B964FF95FFE86480F8D40322F887" bold="true" box="[199,345,1826,1851]" pageId="59" pageNumber="60">Description.</emphasis>
Female (based on
<typeStatus id="3F6ADBD4FF95FFE8667DF8D400E7F887" box="[570,668,1826,1851]" pageId="59" pageNumber="60" type="holotype">holotype</typeStatus>
and
<specimenCount id="F6D7AEFFFF95FFE8669DF8D4010AF887" box="[730,881,1826,1851]" pageId="59" pageNumber="60" type="generic" typeStatus="paratypes">12 paratypes</specimenCount>
). Body length from
<quantity id="2729C893FF95FFE86027F8D40775F886" box="[1120,1294,1826,1851]" metricMagnitude="-1" metricUnit="m" metricValue="4.165" metricValueMax="4.279999999999999" metricValueMin="4.05" pageId="59" pageNumber="60" unit="mm" value="416.5" valueMax="428.0" valueMin="405.0">405 to 428 µm</quantity>
(
<quantity id="2729C893FF95FFE86167F8D40707F886" box="[1312,1404,1826,1851]" metricMagnitude="-1" metricUnit="m" metricValue="4.24" pageId="59" pageNumber="60" unit="mm" value="424.0">424 µm</quantity>
in
<typeStatus id="3F6ADBD4FF95FFE864D0F8B30285F8E2" box="[151,254,1861,1886]" pageId="59" pageNumber="60" type="holotype">holotype</typeStatus>
). Body segmentation, colour, nauplius eye, hyaline fringes, integument thickness as in
<taxonomicName id="27D11EF5FF95FFE86141F8B0028AF83E" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="calliope" status="sp. nov.">
<emphasis id="D2A5B964FF95FFE86141F8B0028AF83E" italics="true" pageId="59" pageNumber="60">Wellstenhelia calliope</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FF95FFE864B8F89C031BF83F" bold="true" box="[255,352,1898,1923]" pageId="59" pageNumber="60">
<taxonomicNameLabel id="C996041FFF95FFE864B8F89C031BF83F" box="[255,352,1898,1923]" pageId="59" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Surface appearance of prosomites extremely smooth and without minute pits even when examined on highest magnifications with scanning electron microscope; surface of urosomites mostly covered with numerous irregular rows of minute pits dorsally and laterally (see inset in
<figureCitation id="78EA79F3FF95FFE867B2F8440625F877" box="[1013,1118,1970,1995]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="59" pageNumber="60">Fig. 34A</figureCitation>
), except for hyaline fringes which generally smooth and slightly frilled. Most somite ornamentation similar to
<taxonomicName id="27D11EF5FF95FFE86022F820071AF852" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1125,1377,2006,2030]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF95FFE86022F820071AF852" box="[1125,1377,2006,2030]" italics="true" pageId="59" pageNumber="60">Wellstenhelia calliope</emphasis>
</taxonomicName>
, and presumed homologous pores and sensilla numbered with same Arabic numerals (see
<figureCitation id="78EA79F3FF92FFEF600FFF610683FF0C" box="[1096,1272,151,176]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="60" pageNumber="61">Figs. 34A, B, C</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF6144FF6107E3FF13" box="[1283,1432,151,176]" captionStart="FIGURE 35" captionStartId="55.[151,249,1829,1851]" captionTargetBox="[218,1368,192,1805]" captionTargetId="figure-138@55.[210,1371,179,1809]" captionTargetPageId="55" captionText="FIGURE 35. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, right side of cephalothoracic shiled, dissected, flattened, and partly broken; B, pleurons of third and fourth pedigerous somites, dissected and flattened; C, genital double-somite, ventral view; D, genital double-somite, lateral view; E, syncoxa of mandibula, antero-ventral view; F, syncoxa of mandibula, posterior view; G, mandibular palp, anterior view; H, maxillula, posterior view; I, mandibula, anterior view; J, maxilliped, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I, V) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910638" httpUri="https://zenodo.org/record/4910638/files/figure.png" pageId="60" pageNumber="61">35A, B, C, D</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF64D0FF4A02B3FF69" box="[151,200,188,213]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="60" pageNumber="61">37A</figureCitation>
) to allow easier comparison. Habitus (
<figureCitation id="78EA79F3FF92FFEF6631FF4A017CFF6F" box="[630,775,188,213]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="60" pageNumber="61">Figs. 34A, B</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF6754FF4A013DFF69" box="[787,838,188,213]" captionStart="FIGURE 39" captionStartId="59.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-211@59.[151,1436,193,1538]" captionTargetPageId="59" captionText="FIGURE 39. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, female paratype 1: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, pedigerous somites, dorsal view; D, first urosomite and genital double-somite, dorsal view; E, anal somite and caudal rami, dorsal view; F, antennula, dorsal view. Scale bars 30 µm (E, F), 40 µm (D), 50 µm (B, C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910646" httpUri="https://zenodo.org/record/4910646/files/figure.png" pageId="60" pageNumber="61">39A</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF6717FF4A01F8FF69" box="[848,899,188,213]" captionStart="FIGURE 40" captionStartId="61.[151,249,1596,1618]" captionTargetBox="[151,1436,218,1563]" captionTargetId="figure-185@61.[151,1436,218,1563]" captionTargetPageId="61" captionText="FIGURE 40. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–E, female paratype 2, F, female paratype 3: A, habitus, lateral view; B, cephalothorax, lateral view; C pedigerous somites, lateral view; D, first urosomite and genital double-somite, lateral view (exopod of fifth leg broken off); E, anal somite and caudal rami, lateral view; F, fifth legs and genital double-somite with one empty egg-sack, ventral view. Scale bars 50 µm (C, D, E, F), 100 µm (B), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910648" httpUri="https://zenodo.org/record/4910648/files/figure.png" pageId="60" pageNumber="61">40A</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF67C9FF4A01BBFF69" box="[910,960,188,213]" captionStart="FIGURE 16" captionStartId="28.[151,249,1573,1595]" captionTargetBox="[151,1436,193,1534]" captionTargetId="figure-209@28.[151,1436,193,1534]" captionTargetPageId="28" captionText="FIGURE 16. Wellstenhelia clio sp. nov., scanning electron micrographs, female paratype: A, habitus, dorsal view; B, rostrum, dorsal view; C, cephalothorax, dorsal view; D, pedigerous somites, dorsal view; E, first urosomite and genital double-somite, dorsal view; F, anal somite and caudal rami, dorsal view. Scale bars 40 µm (B), 50 µm (D, E, F), 100 µm (C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910600" httpUri="https://zenodo.org/record/4910600/files/figure.png" pageId="60" pageNumber="61">16A</figureCitation>
) less robust than in
<taxonomicName id="27D11EF5FF92FFEF60D9FF4A07EEFF68" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1182,1429,188,212]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF92FFEF60D9FF4A07EEFF68" box="[1182,1429,188,212]" italics="true" pageId="60" pageNumber="61">Wellstenhelia calliope</emphasis>
</taxonomicName>
, prosome/urosome length ratio about one, body length/width ratio from 3.1 to 3.6, cephalothorax from 1.5 to 1.7 times as wide as genital double-somite.
</paragraph>
<paragraph id="E06E6576FF92FFEF6480FEDE018CFE34" blockId="60.[151,1436,151,2037]" pageId="60" pageNumber="61">
Rostrum (
<figureCitation id="78EA79F3FF92FFEF657CFED103D5FEFC" box="[315,430,295,320]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="60" pageNumber="61">Figs. 34C</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF65F9FED10396FEFC" box="[446,493,295,320]" captionStart="FIGURE 39" captionStartId="59.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-211@59.[151,1436,193,1538]" captionTargetPageId="59" captionText="FIGURE 39. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, female paratype 1: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, pedigerous somites, dorsal view; D, first urosomite and genital double-somite, dorsal view; E, anal somite and caudal rami, dorsal view; F, antennula, dorsal view. Scale bars 30 µm (E, F), 40 µm (D), 50 µm (B, C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910646" httpUri="https://zenodo.org/record/4910646/files/figure.png" pageId="60" pageNumber="61">39B</figureCitation>
) general shape and position of anterior pair of sensilla no. 1 as in
<taxonomicName id="27D11EF5FF92FFEF6141FEDE028AFED8" authorityName="Karanovic &amp; Kim" authorityYear="2014" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF92FFEF6141FEDE028AFED8" italics="true" pageId="60" pageNumber="61">Wellstenhelia calliope</emphasis>
</taxonomicName>
but with additional bunch of ventral spinules just beyond tip (arrowed in
<figureCitation id="78EA79F3FF92FFEF601DFEBD06BFFED8" box="[1114,1220,331,356]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="60" pageNumber="61">Fig. 34C</figureCitation>
), with pronounced transverse dorsal suture (arrowed in
<figureCitation id="78EA79F3FF92FFEF6668FE9900EEFE34" box="[559,661,367,392]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="60" pageNumber="61">Fig. 34C</figureCitation>
), and without dorsal pore no. 2.
</paragraph>
<paragraph id="E06E6576FF92FFEF6480FE650776FD3D" blockId="60.[151,1436,151,2037]" pageId="60" pageNumber="61">
Cephalothorax (
<figureCitation id="78EA79F3FF92FFEF653BFE65006BFE16" box="[380,528,403,428]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="60" pageNumber="61">Figs. 34A, B</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF665BFE650034FE10" box="[540,591,403,428]" captionStart="FIGURE 35" captionStartId="55.[151,249,1829,1851]" captionTargetBox="[218,1368,192,1805]" captionTargetId="figure-138@55.[210,1371,179,1809]" captionTargetPageId="55" captionText="FIGURE 35. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, right side of cephalothoracic shiled, dissected, flattened, and partly broken; B, pleurons of third and fourth pedigerous somites, dissected and flattened; C, genital double-somite, ventral view; D, genital double-somite, lateral view; E, syncoxa of mandibula, antero-ventral view; F, syncoxa of mandibula, posterior view; G, mandibular palp, anterior view; H, maxillula, posterior view; I, mandibula, anterior view; J, maxilliped, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I, V) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910638" httpUri="https://zenodo.org/record/4910638/files/figure.png" pageId="60" pageNumber="61">35A</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF661CFE6500F7FE10" box="[603,652,403,428]" captionStart="FIGURE 39" captionStartId="59.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-211@59.[151,1436,193,1538]" captionTargetPageId="59" captionText="FIGURE 39. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, female paratype 1: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, pedigerous somites, dorsal view; D, first urosomite and genital double-somite, dorsal view; E, anal somite and caudal rami, dorsal view; F, antennula, dorsal view. Scale bars 30 µm (E, F), 40 µm (D), 50 µm (B, C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910646" httpUri="https://zenodo.org/record/4910646/files/figure.png" pageId="60" pageNumber="61">39B</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF66DFFE6500B3FE10" box="[664,712,403,428]" captionStart="FIGURE 40" captionStartId="61.[151,249,1596,1618]" captionTargetBox="[151,1436,218,1563]" captionTargetId="figure-185@61.[151,1436,218,1563]" captionTargetPageId="61" captionText="FIGURE 40. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–E, female paratype 2, F, female paratype 3: A, habitus, lateral view; B, cephalothorax, lateral view; C pedigerous somites, lateral view; D, first urosomite and genital double-somite, lateral view (exopod of fifth leg broken off); E, anal somite and caudal rami, lateral view; F, fifth legs and genital double-somite with one empty egg-sack, ventral view. Scale bars 50 µm (C, D, E, F), 100 µm (B), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910648" httpUri="https://zenodo.org/record/4910648/files/figure.png" pageId="60" pageNumber="61">40B</figureCitation>
) about 1.1 times as long as wide; comprising 28% of total body length. Surface of cephalothoracic shield with 30 paired or unpaired sensilla and pores, most probably homologous to those in
<taxonomicName id="27D11EF5FF92FFEF655FFE2D006BFE4F" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[280,528,475,499]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF92FFEF655FFE2D006BFE4F" box="[280,528,475,499]" italics="true" pageId="60" pageNumber="61">Wellstenhelia calliope</emphasis>
</taxonomicName>
(indicated with Arabic numerals in illustrations), 14 pores and sensilla missing (nos. 6, 10, 12, 16, 17, 18, 19, 21, 26, 29, 33, 34, 37, 40); absolute and relative positions of some pores and sensilla different; two lateral pairs of sensilla probably homologous to those in
<taxonomicName id="27D11EF5FF92FFEF67ABFDD40699FD86" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1004,1250,546,570]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="euterpe">
<emphasis id="D2A5B964FF92FFEF67ABFDD40699FD86" box="[1004,1250,546,570]" italics="true" pageId="60" pageNumber="61">Wellstenhelia euterpe</emphasis>
</taxonomicName>
(indicated with geometric shapes in illustrations); one unique pair of dorsal pores (no. I) and one unique unpaired dorsal sensillum (no. II) located in central part of cephalothorax (also indicated with Roman numerals in illustrations).
</paragraph>
<paragraph id="E06E6576FF92FFEF6480FD7B0328FD51" blockId="60.[151,1436,151,2037]" pageId="60" pageNumber="61">
Pleuron of second pedigerous somite (
<figureCitation id="78EA79F3FF92FFEF6635FD7B0178FD19" box="[626,771,653,678]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="60" pageNumber="61">Figs. 34A, B</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF6748FD7B013BFD1A" box="[783,832,653,678]" captionStart="FIGURE 39" captionStartId="59.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-211@59.[151,1436,193,1538]" captionTargetPageId="59" captionText="FIGURE 39. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, female paratype 1: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, pedigerous somites, dorsal view; D, first urosomite and genital double-somite, dorsal view; E, anal somite and caudal rami, dorsal view; F, antennula, dorsal view. Scale bars 30 µm (E, F), 40 µm (D), 50 µm (B, C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910646" httpUri="https://zenodo.org/record/4910646/files/figure.png" pageId="60" pageNumber="61">39C</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF670CFD7B0107FD1A" box="[843,892,653,678]" captionStart="FIGURE 40" captionStartId="61.[151,249,1596,1618]" captionTargetBox="[151,1436,218,1563]" captionTargetId="figure-185@61.[151,1436,218,1563]" captionTargetPageId="61" captionText="FIGURE 40. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–E, female paratype 2, F, female paratype 3: A, habitus, lateral view; B, cephalothorax, lateral view; C pedigerous somites, lateral view; D, first urosomite and genital double-somite, lateral view (exopod of fifth leg broken off); E, anal somite and caudal rami, lateral view; F, fifth legs and genital double-somite with one empty egg-sack, ventral view. Scale bars 50 µm (C, D, E, F), 100 µm (B), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910648" httpUri="https://zenodo.org/record/4910648/files/figure.png" pageId="60" pageNumber="61">40C</figureCitation>
) ornamented as in
<taxonomicName id="27D11EF5FF92FFEF6009FD7B073EFD19" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1102,1349,653,677]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF92FFEF6009FD7B073EFD19" box="[1102,1349,653,677]" italics="true" pageId="60" pageNumber="61">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except anterior lateral pair of sensilla no. 44 missing, unique pair of lateral pores (no. III), and unique pair of dorsal sensilla (no. IV).
</paragraph>
<paragraph id="E06E6576FF92FFEF6480FD0E00A5FCE4" blockId="60.[151,1436,151,2037]" pageId="60" pageNumber="61">
Pleuron of third pedigerous somite (
<figureCitation id="78EA79F3FF92FFEF662FFD0E0084FCAC" box="[616,767,760,785]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="60" pageNumber="61">Figs. 34A, B</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF674AFD0E0145FCAD" box="[781,830,760,785]" captionStart="FIGURE 35" captionStartId="55.[151,249,1829,1851]" captionTargetBox="[218,1368,192,1805]" captionTargetId="figure-138@55.[210,1371,179,1809]" captionTargetPageId="55" captionText="FIGURE 35. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, right side of cephalothoracic shiled, dissected, flattened, and partly broken; B, pleurons of third and fourth pedigerous somites, dissected and flattened; C, genital double-somite, ventral view; D, genital double-somite, lateral view; E, syncoxa of mandibula, antero-ventral view; F, syncoxa of mandibula, posterior view; G, mandibular palp, anterior view; H, maxillula, posterior view; I, mandibula, anterior view; J, maxilliped, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I, V) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910638" httpUri="https://zenodo.org/record/4910638/files/figure.png" pageId="60" pageNumber="61">35B</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF670AFD0E0105FCAD" box="[845,894,760,785]" captionStart="FIGURE 39" captionStartId="59.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-211@59.[151,1436,193,1538]" captionTargetPageId="59" captionText="FIGURE 39. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, female paratype 1: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, pedigerous somites, dorsal view; D, first urosomite and genital double-somite, dorsal view; E, anal somite and caudal rami, dorsal view; F, antennula, dorsal view. Scale bars 30 µm (E, F), 40 µm (D), 50 µm (B, C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910646" httpUri="https://zenodo.org/record/4910646/files/figure.png" pageId="60" pageNumber="61">39C</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF67CBFD0E01C7FCAD" box="[908,956,760,785]" captionStart="FIGURE 40" captionStartId="61.[151,249,1596,1618]" captionTargetBox="[151,1436,218,1563]" captionTargetId="figure-185@61.[151,1436,218,1563]" captionTargetPageId="61" captionText="FIGURE 40. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–E, female paratype 2, F, female paratype 3: A, habitus, lateral view; B, cephalothorax, lateral view; C pedigerous somites, lateral view; D, first urosomite and genital double-somite, lateral view (exopod of fifth leg broken off); E, anal somite and caudal rami, lateral view; F, fifth legs and genital double-somite with one empty egg-sack, ventral view. Scale bars 50 µm (C, D, E, F), 100 µm (B), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910648" httpUri="https://zenodo.org/record/4910648/files/figure.png" pageId="60" pageNumber="61">40C</figureCitation>
) ornamented as in
<taxonomicName id="27D11EF5FF92FFEF60DCFD0E07EEFCAC" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1179,1429,760,784]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF92FFEF60DCFD0E07EEFCAC" box="[1179,1429,760,784]" italics="true" pageId="60" pageNumber="61">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except anterior pair of dorsal pores no. 51 and anterior lateral pair of sensilla no. 52 missing (latter arrowed in
<figureCitation id="78EA79F3FF92FFEF6128FCEA02B3FCE4" captionStart="FIGURE 35" captionStartId="55.[151,249,1829,1851]" captionTargetBox="[218,1368,192,1805]" captionTargetId="figure-138@55.[210,1371,179,1809]" captionTargetPageId="55" captionText="FIGURE 35. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, right side of cephalothoracic shiled, dissected, flattened, and partly broken; B, pleurons of third and fourth pedigerous somites, dissected and flattened; C, genital double-somite, ventral view; D, genital double-somite, lateral view; E, syncoxa of mandibula, antero-ventral view; F, syncoxa of mandibula, posterior view; G, mandibular palp, anterior view; H, maxillula, posterior view; I, mandibula, anterior view; J, maxilliped, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I, V) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910638" httpUri="https://zenodo.org/record/4910638/files/figure.png" pageId="60" pageNumber="61">Fig. 35B</figureCitation>
), and one unique dorsal pair of sensilla (no. V).
</paragraph>
<paragraph id="E06E6576FF92FFEF6480FC95073BFC1C" blockId="60.[151,1436,151,2037]" pageId="60" pageNumber="61">
Pleuron of fourth pedigerous somite (
<figureCitation id="78EA79F3FF92FFEF662CFC950086FCC7" box="[619,765,867,892]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="60" pageNumber="61">Figs. 34A, B</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF674EFC950141FCC0" box="[777,826,867,892]" captionStart="FIGURE 35" captionStartId="55.[151,249,1829,1851]" captionTargetBox="[218,1368,192,1805]" captionTargetId="figure-138@55.[210,1371,179,1809]" captionTargetPageId="55" captionText="FIGURE 35. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, right side of cephalothoracic shiled, dissected, flattened, and partly broken; B, pleurons of third and fourth pedigerous somites, dissected and flattened; C, genital double-somite, ventral view; D, genital double-somite, lateral view; E, syncoxa of mandibula, antero-ventral view; F, syncoxa of mandibula, posterior view; G, mandibular palp, anterior view; H, maxillula, posterior view; I, mandibula, anterior view; J, maxilliped, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I, V) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910638" httpUri="https://zenodo.org/record/4910638/files/figure.png" pageId="60" pageNumber="61">35B</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF6701FC95010CFCC0" box="[838,887,867,892]" captionStart="FIGURE 39" captionStartId="59.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-211@59.[151,1436,193,1538]" captionTargetPageId="59" captionText="FIGURE 39. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, female paratype 1: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, pedigerous somites, dorsal view; D, first urosomite and genital double-somite, dorsal view; E, anal somite and caudal rami, dorsal view; F, antennula, dorsal view. Scale bars 30 µm (E, F), 40 µm (D), 50 µm (B, C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910646" httpUri="https://zenodo.org/record/4910646/files/figure.png" pageId="60" pageNumber="61">39C</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF67C4FC9501C8FCC0" box="[899,947,867,892]" captionStart="FIGURE 40" captionStartId="61.[151,249,1596,1618]" captionTargetBox="[151,1436,218,1563]" captionTargetId="figure-185@61.[151,1436,218,1563]" captionTargetPageId="61" captionText="FIGURE 40. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–E, female paratype 2, F, female paratype 3: A, habitus, lateral view; B, cephalothorax, lateral view; C pedigerous somites, lateral view; D, first urosomite and genital double-somite, lateral view (exopod of fifth leg broken off); E, anal somite and caudal rami, lateral view; F, fifth legs and genital double-somite with one empty egg-sack, ventral view. Scale bars 50 µm (C, D, E, F), 100 µm (B), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910648" httpUri="https://zenodo.org/record/4910648/files/figure.png" pageId="60" pageNumber="61">40C</figureCitation>
) as in
<taxonomicName id="27D11EF5FF92FFEF67BAFC92068FFCC0" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1021,1268,868,892]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF92FFEF67BAFC92068FFCC0" box="[1021,1268,868,892]" italics="true" pageId="60" pageNumber="61">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except pair of anterior lateral pores no. 57 and pair of anterior lateral sensilla no. 62 missing (latter arrowed in
<figureCitation id="78EA79F3FF92FFEF608AFC710748FC1C" box="[1229,1331,903,928]" captionStart="FIGURE 35" captionStartId="55.[151,249,1829,1851]" captionTargetBox="[218,1368,192,1805]" captionTargetId="figure-138@55.[210,1371,179,1809]" captionTargetPageId="55" captionText="FIGURE 35. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, right side of cephalothoracic shiled, dissected, flattened, and partly broken; B, pleurons of third and fourth pedigerous somites, dissected and flattened; C, genital double-somite, ventral view; D, genital double-somite, lateral view; E, syncoxa of mandibula, antero-ventral view; F, syncoxa of mandibula, posterior view; G, mandibular palp, anterior view; H, maxillula, posterior view; I, mandibula, anterior view; J, maxilliped, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I, V) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910638" httpUri="https://zenodo.org/record/4910638/files/figure.png" pageId="60" pageNumber="61">Fig. 35B</figureCitation>
).
</paragraph>
<paragraph id="E06E6576FF92FFEF64D0FC5D028EFC5B" blockId="60.[151,1436,151,2037]" pageId="60" pageNumber="61">
First urosomite (
<figureCitation id="78EA79F3FF92FFEF6514FC5D0392FC7E" box="[339,489,939,964]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="60" pageNumber="61">Figs. 34A, B</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF65B2FC5D0053FC78" box="[501,552,939,964]" captionStart="FIGURE 39" captionStartId="59.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-211@59.[151,1436,193,1538]" captionTargetPageId="59" captionText="FIGURE 39. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, female paratype 1: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, pedigerous somites, dorsal view; D, first urosomite and genital double-somite, dorsal view; E, anal somite and caudal rami, dorsal view; F, antennula, dorsal view. Scale bars 30 µm (E, F), 40 µm (D), 50 µm (B, C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910646" httpUri="https://zenodo.org/record/4910646/files/figure.png" pageId="60" pageNumber="61">39D</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF6672FC5D001CFC78" box="[565,615,939,964]" captionStart="FIGURE 40" captionStartId="61.[151,249,1596,1618]" captionTargetBox="[151,1436,218,1563]" captionTargetId="figure-185@61.[151,1436,218,1563]" captionTargetPageId="61" captionText="FIGURE 40. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–E, female paratype 2, F, female paratype 3: A, habitus, lateral view; B, cephalothorax, lateral view; C pedigerous somites, lateral view; D, first urosomite and genital double-somite, lateral view (exopod of fifth leg broken off); E, anal somite and caudal rami, lateral view; F, fifth legs and genital double-somite with one empty egg-sack, ventral view. Scale bars 50 µm (C, D, E, F), 100 µm (B), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910648" httpUri="https://zenodo.org/record/4910648/files/figure.png" pageId="60" pageNumber="61">40D</figureCitation>
) as in
<taxonomicName id="27D11EF5FF92FFEF66F2FC5D01D6FC7F" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[693,941,939,963]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF92FFEF66F2FC5D01D6FC7F" box="[693,941,939,963]" italics="true" pageId="60" pageNumber="61">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except pair of anterior lateral pores no. 63 missing.
</paragraph>
<paragraph id="E06E6576FF92FFEF6480FC040757FBCA" blockId="60.[151,1436,151,2037]" pageId="60" pageNumber="61">
Genital double-somite (
<figureCitation id="78EA79F3FF92FFEF6588FC04001BFBB6" box="[463,608,1010,1035]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="60" pageNumber="61">Figs. 34A, B</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF662BFC0400C4FBB6" box="[620,703,1010,1035]" captionStart="FIGURE 35" captionStartId="55.[151,249,1829,1851]" captionTargetBox="[218,1368,192,1805]" captionTargetId="figure-138@55.[210,1371,179,1809]" captionTargetPageId="55" captionText="FIGURE 35. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, right side of cephalothoracic shiled, dissected, flattened, and partly broken; B, pleurons of third and fourth pedigerous somites, dissected and flattened; C, genital double-somite, ventral view; D, genital double-somite, lateral view; E, syncoxa of mandibula, antero-ventral view; F, syncoxa of mandibula, posterior view; G, mandibular palp, anterior view; H, maxillula, posterior view; I, mandibula, anterior view; J, maxilliped, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I, V) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910638" httpUri="https://zenodo.org/record/4910638/files/figure.png" pageId="60" pageNumber="61">35C, D</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF668EFC040087FBB7" box="[713,764,1010,1035]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="60" pageNumber="61">37A</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF6740FC040141FBB7" box="[775,826,1010,1035]" captionStart="FIGURE 39" captionStartId="59.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-211@59.[151,1436,193,1538]" captionTargetPageId="59" captionText="FIGURE 39. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, female paratype 1: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, pedigerous somites, dorsal view; D, first urosomite and genital double-somite, dorsal view; E, anal somite and caudal rami, dorsal view; F, antennula, dorsal view. Scale bars 30 µm (E, F), 40 µm (D), 50 µm (B, C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910646" httpUri="https://zenodo.org/record/4910646/files/figure.png" pageId="60" pageNumber="61">39D</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF6702FC04010DFBB7" box="[837,886,1010,1035]" captionStart="FIGURE 40" captionStartId="61.[151,249,1596,1618]" captionTargetBox="[151,1436,218,1563]" captionTargetId="figure-185@61.[151,1436,218,1563]" captionTargetPageId="61" captionText="FIGURE 40. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–E, female paratype 2, F, female paratype 3: A, habitus, lateral view; B, cephalothorax, lateral view; C pedigerous somites, lateral view; D, first urosomite and genital double-somite, lateral view (exopod of fifth leg broken off); E, anal somite and caudal rami, lateral view; F, fifth legs and genital double-somite with one empty egg-sack, ventral view. Scale bars 50 µm (C, D, E, F), 100 µm (B), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910648" httpUri="https://zenodo.org/record/4910648/files/figure.png" pageId="60" pageNumber="61">40D</figureCitation>
) as in
<taxonomicName id="27D11EF5FF92FFEF67F8FC0506CDFBB7" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[959,1206,1011,1035]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF92FFEF67F8FC0506CDFBB7" box="[959,1206,1011,1035]" italics="true" pageId="60" pageNumber="61">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except anterior part less inflated laterally, genital apertures situated more ventrally, all large spinules missing (arrowed in
<figureCitation id="78EA79F3FF92FFEF6161FBE007ECFB93" box="[1318,1431,1046,1071]" captionStart="FIGURE 35" captionStartId="55.[151,249,1829,1851]" captionTargetBox="[218,1368,192,1805]" captionTargetId="figure-138@55.[210,1371,179,1809]" captionTargetPageId="55" captionText="FIGURE 35. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, right side of cephalothoracic shiled, dissected, flattened, and partly broken; B, pleurons of third and fourth pedigerous somites, dissected and flattened; C, genital double-somite, ventral view; D, genital double-somite, lateral view; E, syncoxa of mandibula, antero-ventral view; F, syncoxa of mandibula, posterior view; G, mandibular palp, anterior view; H, maxillula, posterior view; I, mandibula, anterior view; J, maxilliped, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I, V) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910638" httpUri="https://zenodo.org/record/4910638/files/figure.png" pageId="60" pageNumber="61">Figs. 35C</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF64D0FBCC02B1FBEF" box="[151,202,1082,1107]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="60" pageNumber="61">37A</figureCitation>
), anterior pair of pores no. 68 missing, and dorsal anterior pairs of sensilla nos. 69 &amp; 70 more widely spaced; only more sclerotized part of genital apparatus visible inside and copulatory pores not visible on surface.
</paragraph>
<paragraph id="E06E6576FF92FFEF6480FB7701C2FB02" blockId="60.[151,1436,151,2037]" pageId="60" pageNumber="61">
Third urosomite (
<figureCitation id="78EA79F3FF92FFEF65C9FB77005FFB25" box="[398,548,1153,1178]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="60" pageNumber="61">Figs. 34A, B</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF6676FB77001FFB26" box="[561,612,1153,1178]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="60" pageNumber="61">37A</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF6636FB7700D9FB26" box="[625,674,1153,1178]" captionStart="FIGURE 40" captionStartId="61.[151,249,1596,1618]" captionTargetBox="[151,1436,218,1563]" captionTargetId="figure-185@61.[151,1436,218,1563]" captionTargetPageId="61" captionText="FIGURE 40. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–E, female paratype 2, F, female paratype 3: A, habitus, lateral view; B, cephalothorax, lateral view; C pedigerous somites, lateral view; D, first urosomite and genital double-somite, lateral view (exopod of fifth leg broken off); E, anal somite and caudal rami, lateral view; F, fifth legs and genital double-somite with one empty egg-sack, ventral view. Scale bars 50 µm (C, D, E, F), 100 µm (B), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910648" httpUri="https://zenodo.org/record/4910648/files/figure.png" pageId="60" pageNumber="61">40D</figureCitation>
) as in
<taxonomicName id="27D11EF5FF92FFEF66B6FB770191FB25" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[753,1002,1153,1177]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF92FFEF66B6FB770191FB25" box="[753,1002,1153,1177]" italics="true" pageId="60" pageNumber="61">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except all large spinules missing and one unique pair of ventral pores (no. VI) present (arrowed in
<figureCitation id="78EA79F3FF92FFEF6703FB5301D7FB02" box="[836,940,1189,1214]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="60" pageNumber="61">Fig. 37A</figureCitation>
).
</paragraph>
<paragraph id="E06E6576FF92FFEF6480FB3F00FEFAB9" blockId="60.[151,1436,151,2037]" pageId="60" pageNumber="61">
Preanal somite (
<figureCitation id="78EA79F3FF92FFEF65C5FB3F0060FB5D" box="[386,539,1225,1250]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="60" pageNumber="61">Figs. 34A, B</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF666DFB3F0020FB5E" box="[554,603,1225,1250]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="60" pageNumber="61">37A</figureCitation>
) without pores and sensilla, with numerous rows of minute spinules on dorsal and lateral sides, as in all urosomites.
</paragraph>
<paragraph id="E06E6576FF92FFEF6480FAE606DEFACC" blockId="60.[151,1436,151,2037]" pageId="60" pageNumber="61">
Anal somite (
<figureCitation id="78EA79F3FF92FFEF6522FAE6007BFA94" box="[357,512,1296,1321]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="60" pageNumber="61">Figs. 34A, B</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF6648FAE60039FA95" box="[527,578,1296,1321]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="60" pageNumber="61">37A</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF6616FAE600FAFA95" box="[593,641,1296,1321]" captionStart="FIGURE 39" captionStartId="59.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-211@59.[151,1436,193,1538]" captionTargetPageId="59" captionText="FIGURE 39. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, female paratype 1: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, pedigerous somites, dorsal view; D, first urosomite and genital double-somite, dorsal view; E, anal somite and caudal rami, dorsal view; F, antennula, dorsal view. Scale bars 30 µm (E, F), 40 µm (D), 50 µm (B, C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910646" httpUri="https://zenodo.org/record/4910646/files/figure.png" pageId="60" pageNumber="61">39E</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF66C8FAE600C5FA95" box="[655,702,1296,1321]" captionStart="FIGURE 40" captionStartId="61.[151,249,1596,1618]" captionTargetBox="[151,1436,218,1563]" captionTargetId="figure-185@61.[151,1436,218,1563]" captionTargetPageId="61" captionText="FIGURE 40. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–E, female paratype 2, F, female paratype 3: A, habitus, lateral view; B, cephalothorax, lateral view; C pedigerous somites, lateral view; D, first urosomite and genital double-somite, lateral view (exopod of fifth leg broken off); E, anal somite and caudal rami, lateral view; F, fifth legs and genital double-somite with one empty egg-sack, ventral view. Scale bars 50 µm (C, D, E, F), 100 µm (B), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910648" httpUri="https://zenodo.org/record/4910648/files/figure.png" pageId="60" pageNumber="61">40E</figureCitation>
) general shape and ornamentation as in
<taxonomicName id="27D11EF5FF92FFEF60DDFAE607EEFA94" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1178,1429,1296,1320]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF92FFEF60DDFAE607EEFA94" box="[1178,1429,1296,1320]" italics="true" pageId="60" pageNumber="61">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except ventral spinules along medial cleft (arrowed in
<figureCitation id="78EA79F3FF92FFEF6746FAC2011CFAF1" box="[769,871,1332,1357]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="60" pageNumber="61">Fig. 37A</figureCitation>
) and lateral pores nos. 79 &amp; 80 missing, and anal operculum well developed, broad and smooth, reaching beyond distal margin of anal somite.
</paragraph>
<paragraph id="E06E6576FF92FFEF6480FA8D0644F9FB" blockId="60.[151,1436,151,2037]" pageId="60" pageNumber="61">
Caudal rami (
<figureCitation id="78EA79F3FF92FFEF652FFA8D0078FA2F" box="[360,515,1403,1428]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="60" pageNumber="61">Figs. 34A, B</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF6654FA8D0011FA2F" box="[531,618,1403,1428]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="60" pageNumber="61">37A, B</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF663DFA8D00D1FA28" box="[634,682,1403,1428]" captionStart="FIGURE 39" captionStartId="59.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-211@59.[151,1436,193,1538]" captionTargetPageId="59" captionText="FIGURE 39. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, female paratype 1: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, pedigerous somites, dorsal view; D, first urosomite and genital double-somite, dorsal view; E, anal somite and caudal rami, dorsal view; F, antennula, dorsal view. Scale bars 30 µm (E, F), 40 µm (D), 50 µm (B, C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910646" httpUri="https://zenodo.org/record/4910646/files/figure.png" pageId="60" pageNumber="61">39E</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF66FEFA8D0093FA28" box="[697,744,1403,1428]" captionStart="FIGURE 40" captionStartId="61.[151,249,1596,1618]" captionTargetBox="[151,1436,218,1563]" captionTargetId="figure-185@61.[151,1436,218,1563]" captionTargetPageId="61" captionText="FIGURE 40. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–E, female paratype 2, F, female paratype 3: A, habitus, lateral view; B, cephalothorax, lateral view; C pedigerous somites, lateral view; D, first urosomite and genital double-somite, lateral view (exopod of fifth leg broken off); E, anal somite and caudal rami, lateral view; F, fifth legs and genital double-somite with one empty egg-sack, ventral view. Scale bars 50 µm (C, D, E, F), 100 µm (B), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910648" httpUri="https://zenodo.org/record/4910648/files/figure.png" pageId="60" pageNumber="61">40E</figureCitation>
) shape and armature very similar to those in
<emphasis id="D2A5B964FF92FFEF6141FA8A0283FA0B" italics="true" pageId="60" pageNumber="61">
<taxonomicName id="27D11EF5FF92FFEF6141FA8A028FFA0B" authorityName="Karanovic &amp; Kim" authorityYear="2014" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="calliope">Wellstenhelia calliope</taxonomicName>
,
</emphasis>
except dorsalmost lateral setae much stronger and conspicuously directed outwards (arrowed in
<figureCitation id="78EA79F3FF92FFEF6137FA6902B0FA60" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="60" pageNumber="61">Fig. 37A</figureCitation>
); rami cylindrical, about 1.7 times as long as anal somite, 3.5 times as long as wide (ventral view), slightly divergent or parallel, with space between them about one ramus width; ornamentation consisting of several large inner spinules in anterior part, single unique anterior pore in distal part (no. VII; arrowed in
<figureCitation id="78EA79F3FF92FFEF60E4F9FC0773F99F" box="[1187,1288,1546,1571]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="60" pageNumber="61">Fig. 37B</figureCitation>
), and several small spinules at base of dorsal and lateral setae and along dorsal and inner surface.
</paragraph>
<paragraph id="E06E6576FF92FFEF6480F9A403F3F946" blockId="60.[151,1436,151,2037]" pageId="60" pageNumber="61">
Antennula (
<figureCitation id="78EA79F3FF92FFEF6509F9A403B9F9D7" box="[334,450,1618,1643]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="60" pageNumber="61">Figs. 34D</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF6596F9A40384F9D7" box="[465,511,1618,1643]" captionStart="FIGURE 39" captionStartId="59.[151,249,1559,1581]" captionTargetBox="[151,1436,193,1538]" captionTargetId="figure-211@59.[151,1436,193,1538]" captionTargetPageId="59" captionText="FIGURE 39. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, female paratype 1: A, habitus, dorsal view; B, cephalothorax, dorsal view; C, pedigerous somites, dorsal view; D, first urosomite and genital double-somite, dorsal view; E, anal somite and caudal rami, dorsal view; F, antennula, dorsal view. Scale bars 30 µm (E, F), 40 µm (D), 50 µm (B, C), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910646" httpUri="https://zenodo.org/record/4910646/files/figure.png" pageId="60" pageNumber="61">39F</figureCitation>
) similar to that in
<taxonomicName id="27D11EF5FF92FFEF669CF9A401AEF9D6" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[731,981,1618,1642]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF92FFEF669CF9A401AEF9D6" box="[731,981,1618,1642]" italics="true" pageId="60" pageNumber="61">Wellstenhelia calliope</emphasis>
</taxonomicName>
but sixth and seventh segments almost completely fused, and eighth segment with only three lateral setae and without apical aesthetasc (all three features arrowed in
<figureCitation id="78EA79F3FF92FFEF655AF96F03F3F90E" box="[285,392,1689,1714]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="60" pageNumber="61">Fig. 34D</figureCitation>
); only ornamentation single short row of small spinules in proximal half of first segment, distal row of spinules missing; aesthetasc on fourth segment reaching distal tip of appendage; setal formula 1.10.8.6+ae.3.(2.5).5.
</paragraph>
<paragraph id="E06E6576FF92FFEF6480F8F2033DF8FD" blockId="60.[151,1436,151,2037]" pageId="60" pageNumber="61">
Antenna (
<figureCitation id="78EA79F3FF92FFEF6570F8F203E7F8A1" box="[311,412,1796,1821]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="60" pageNumber="61">Fig. 34E</figureCitation>
) as in
<taxonomicName id="27D11EF5FF92FFEF65ABF8F3009EF8A1" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[492,741,1797,1821]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF92FFEF65ABF8F3009EF8A1" box="[492,741,1797,1821]" italics="true" pageId="60" pageNumber="61">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except allobasis and first exopodal segment proportionately slightly shorter.
</paragraph>
<paragraph id="E06E6576FF92FFEF6480F8BA016BF835" blockId="60.[151,1436,151,2037]" pageId="60" pageNumber="61">
Labrum (
<figureCitation id="78EA79F3FF92FFEF6574F8BA03E6F8D9" box="[307,413,1868,1893]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="60" pageNumber="61">Fig. 41A</figureCitation>
) and paragnaths (
<figureCitation id="78EA79F3FF92FFEF6636F8BA00A0F8D9" box="[625,731,1868,1893]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="60" pageNumber="61">Fig. 41A</figureCitation>
) as in
<taxonomicName id="27D11EF5FF92FFEF6775F8BA0655F8D8" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[818,1070,1868,1892]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF92FFEF6775F8BA0655F8D8" box="[818,1070,1868,1892]" italics="true" pageId="60" pageNumber="61">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except labrum with additional spiniform process on anterior surface near distal margin.
</paragraph>
<paragraph id="E06E6576FF92FFEF6480F86200F7F848" blockId="60.[151,1436,151,2037]" pageId="60" pageNumber="61">
Mandibula (
<figureCitation id="78EA79F3FF92FFEF6517F8620381F811" box="[336,506,1940,1965]" captionStart="FIGURE 35" captionStartId="55.[151,249,1829,1851]" captionTargetBox="[218,1368,192,1805]" captionTargetId="figure-138@55.[210,1371,179,1809]" captionTargetPageId="55" captionText="FIGURE 35. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, right side of cephalothoracic shiled, dissected, flattened, and partly broken; B, pleurons of third and fourth pedigerous somites, dissected and flattened; C, genital double-somite, ventral view; D, genital double-somite, lateral view; E, syncoxa of mandibula, antero-ventral view; F, syncoxa of mandibula, posterior view; G, mandibular palp, anterior view; H, maxillula, posterior view; I, mandibula, anterior view; J, maxilliped, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I, V) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910638" httpUri="https://zenodo.org/record/4910638/files/figure.png" pageId="60" pageNumber="61">Figs. 35E, F, G</figureCitation>
,
<figureCitation id="78EA79F3FF92FFEF6642F862004CF811" box="[517,567,1940,1965]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="60" pageNumber="61">41A</figureCitation>
) similar to that in
<taxonomicName id="27D11EF5FF92FFEF6741F8620186F810" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[774,1021,1940,1964]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF92FFEF6741F8620186F810" box="[774,1021,1940,1964]" italics="true" pageId="60" pageNumber="61">Wellstenhelia calliope</emphasis>
</taxonomicName>
but cutting edge without central seta (arrowed in
<figureCitation id="78EA79F3FF92FFEF6563F84E03F3F86D" box="[292,392,1976,2001]" captionStart="FIGURE 35" captionStartId="55.[151,249,1829,1851]" captionTargetBox="[218,1368,192,1805]" captionTargetId="figure-138@55.[210,1371,179,1809]" captionTargetPageId="55" captionText="FIGURE 35. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, right side of cephalothoracic shiled, dissected, flattened, and partly broken; B, pleurons of third and fourth pedigerous somites, dissected and flattened; C, genital double-somite, ventral view; D, genital double-somite, lateral view; E, syncoxa of mandibula, antero-ventral view; F, syncoxa of mandibula, posterior view; G, mandibular palp, anterior view; H, maxillula, posterior view; I, mandibula, anterior view; J, maxilliped, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I, V) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910638" httpUri="https://zenodo.org/record/4910638/files/figure.png" pageId="60" pageNumber="61">Fig. 35E</figureCitation>
) and exopod with only one proximal seta (arrowed in
<figureCitation id="78EA79F3FF92FFEF67B9F84E0612F86D" box="[1022,1129,1976,2001]" captionStart="FIGURE 35" captionStartId="55.[151,249,1829,1851]" captionTargetBox="[218,1368,192,1805]" captionTargetId="figure-138@55.[210,1371,179,1809]" captionTargetPageId="55" captionText="FIGURE 35. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, right side of cephalothoracic shiled, dissected, flattened, and partly broken; B, pleurons of third and fourth pedigerous somites, dissected and flattened; C, genital double-somite, ventral view; D, genital double-somite, lateral view; E, syncoxa of mandibula, antero-ventral view; F, syncoxa of mandibula, posterior view; G, mandibular palp, anterior view; H, maxillula, posterior view; I, mandibula, anterior view; J, maxilliped, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I, V) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910638" httpUri="https://zenodo.org/record/4910638/files/figure.png" pageId="60" pageNumber="61">Fig. 35G</figureCitation>
); both coxa and basis with more spinules than in
<taxonomicName id="27D11EF5FF92FFEF65C9F82A00FEF848" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[398,645,2012,2036]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF92FFEF65C9F82A00FEF848" box="[398,645,2012,2036]" italics="true" pageId="60" pageNumber="61">Wellstenhelia calliope</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="B4AE35FEFF93FFEE64D0F9CA070BF912" ID-DOI="http://doi.org/10.5281/zenodo.4910648" ID-Zenodo-Dep="4910648" httpUri="https://zenodo.org/record/4910648/files/figure.png" pageId="61" pageNumber="62" startId="61.[151,249,1596,1618]" targetBox="[151,1436,218,1563]" targetPageId="61">
<paragraph id="E06E6576FF93FFEE64D0F9CA070BF912" blockId="61.[151,1436,1596,1710]" pageId="61" pageNumber="62">
<emphasis id="D2A5B964FF93FFEE64D0F9CA035BF9ED" bold="true" box="[151,288,1596,1618]" pageId="61" pageNumber="62">FIGURE 40.</emphasis>
<taxonomicName id="27D11EF5FF93FFEE6560F9CA0060F9EE" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[295,539,1596,1618]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="polyhymnia" status="sp. nov.">
<emphasis id="D2A5B964FF93FFEE6560F9CA0060F9EE" box="[295,539,1596,1618]" italics="true" pageId="61" pageNumber="62">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FF93FFEE6665F9CB0008F9EE" bold="true" box="[546,627,1597,1618]" pageId="61" pageNumber="62">
<taxonomicNameLabel id="C996041FFF93FFEE6665F9CB0008F9EE" box="[546,627,1597,1618]" pageId="61" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, scanning electron micrographs, A–E, female paratype 2, F, female paratype 3: A, habitus, lateral view; B, cephalothorax, lateral view; C pedigerous somites, lateral view; D, first urosomite and genital double-somite, lateral view (exopod of fifth leg broken off); E, anal somite and caudal rami, lateral view; F, fifth legs and genital double-somite with one empty egg-sack, ventral view. Scale bars 50 µm (C, D, E, F), 100 µm (B), and 200 µm (A).
</paragraph>
</caption>
<paragraph id="E06E6576FF93FFEE6480F92300E4F8AF" blockId="61.[151,1436,1749,1990]" pageId="61" pageNumber="62">
Maxillula (
<figureCitation id="78EA79F3FF93FFEE6504F92303CFF952" box="[323,436,1749,1774]" captionStart="FIGURE 35" captionStartId="55.[151,249,1829,1851]" captionTargetBox="[218,1368,192,1805]" captionTargetId="figure-138@55.[210,1371,179,1809]" captionTargetPageId="55" captionText="FIGURE 35. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, right side of cephalothoracic shiled, dissected, flattened, and partly broken; B, pleurons of third and fourth pedigerous somites, dissected and flattened; C, genital double-somite, ventral view; D, genital double-somite, lateral view; E, syncoxa of mandibula, antero-ventral view; F, syncoxa of mandibula, posterior view; G, mandibular palp, anterior view; H, maxillula, posterior view; I, mandibula, anterior view; J, maxilliped, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I, V) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910638" httpUri="https://zenodo.org/record/4910638/files/figure.png" pageId="61" pageNumber="62">Figs. 35H</figureCitation>
,
<figureCitation id="78EA79F3FF93FFEE65F8F923038BF952" box="[447,496,1749,1774]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="61" pageNumber="62">41A</figureCitation>
) as in
<taxonomicName id="27D11EF5FF93FFEE667EF9200154F952" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[569,815,1750,1774]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF93FFEE667EF9200154F952" box="[569,815,1750,1774]" italics="true" pageId="61" pageNumber="62">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except exopod and endopod not fused (arrowed in
<figureCitation id="78EA79F3FF93FFEE6137F92302B3F8AF" captionStart="FIGURE 35" captionStartId="55.[151,249,1829,1851]" captionTargetBox="[218,1368,192,1805]" captionTargetId="figure-138@55.[210,1371,179,1809]" captionTargetPageId="55" captionText="FIGURE 35. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, right side of cephalothoracic shiled, dissected, flattened, and partly broken; B, pleurons of third and fourth pedigerous somites, dissected and flattened; C, genital double-somite, ventral view; D, genital double-somite, lateral view; E, syncoxa of mandibula, antero-ventral view; F, syncoxa of mandibula, posterior view; G, mandibular palp, anterior view; H, maxillula, posterior view; I, mandibula, anterior view; J, maxilliped, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I, V) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910638" httpUri="https://zenodo.org/record/4910638/files/figure.png" pageId="61" pageNumber="62">Fig. 35H</figureCitation>
) and coxa without large anterior spinules.
</paragraph>
<paragraph id="E06E6576FF93FFEE6480F8EB02ADF8C1" blockId="61.[151,1436,1749,1990]" pageId="61" pageNumber="62">
Maxilla (
<figureCitation id="78EA79F3FF93FFEE6577F8EB03E0F88A" box="[304,411,1821,1846]" captionStart="FIGURE 35" captionStartId="55.[151,249,1829,1851]" captionTargetBox="[218,1368,192,1805]" captionTargetId="figure-138@55.[210,1371,179,1809]" captionTargetPageId="55" captionText="FIGURE 35. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, right side of cephalothoracic shiled, dissected, flattened, and partly broken; B, pleurons of third and fourth pedigerous somites, dissected and flattened; C, genital double-somite, ventral view; D, genital double-somite, lateral view; E, syncoxa of mandibula, antero-ventral view; F, syncoxa of mandibula, posterior view; G, mandibular palp, anterior view; H, maxillula, posterior view; I, mandibula, anterior view; J, maxilliped, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I, V) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910638" httpUri="https://zenodo.org/record/4910638/files/figure.png" pageId="61" pageNumber="62">Figs. 35I</figureCitation>
,
<figureCitation id="78EA79F3FF93FFEE65EDF8EB03A0F88A" box="[426,475,1821,1846]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="61" pageNumber="62">41A</figureCitation>
) as in
<taxonomicName id="27D11EF5FF93FFEE6668F8E80152F88A" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[559,809,1822,1846]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF93FFEE6668F8E80152F88A" box="[559,809,1822,1846]" italics="true" pageId="61" pageNumber="62">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except dorsal endite on syncoxa with only two setae (arrowed in
<figureCitation id="78EA79F3FF93FFEE6563F8B403F8F8E7" box="[292,387,1858,1883]" captionStart="FIGURE 35" captionStartId="55.[151,249,1829,1851]" captionTargetBox="[218,1368,192,1805]" captionTargetId="figure-138@55.[210,1371,179,1809]" captionTargetPageId="55" captionText="FIGURE 35. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, right side of cephalothoracic shiled, dissected, flattened, and partly broken; B, pleurons of third and fourth pedigerous somites, dissected and flattened; C, genital double-somite, ventral view; D, genital double-somite, lateral view; E, syncoxa of mandibula, antero-ventral view; F, syncoxa of mandibula, posterior view; G, mandibular palp, anterior view; H, maxillula, posterior view; I, mandibula, anterior view; J, maxilliped, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I, V) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910638" httpUri="https://zenodo.org/record/4910638/files/figure.png" pageId="61" pageNumber="62">Fig. 35I</figureCitation>
), syncoxa with only one arched row of outer spinules, and endopod with only four slender setae.
</paragraph>
<paragraph id="E06E6576FF93FFEE6480F87C01C7F879" blockId="61.[151,1436,1749,1990]" pageId="61" pageNumber="62">
Maxilliped (
<figureCitation id="78EA79F3FF93FFEE6515F87C03C7F81F" box="[338,444,1930,1955]" captionStart="FIGURE 35" captionStartId="55.[151,249,1829,1851]" captionTargetBox="[218,1368,192,1805]" captionTargetId="figure-138@55.[210,1371,179,1809]" captionTargetPageId="55" captionText="FIGURE 35. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, right side of cephalothoracic shiled, dissected, flattened, and partly broken; B, pleurons of third and fourth pedigerous somites, dissected and flattened; C, genital double-somite, ventral view; D, genital double-somite, lateral view; E, syncoxa of mandibula, antero-ventral view; F, syncoxa of mandibula, posterior view; G, mandibular palp, anterior view; H, maxillula, posterior view; I, mandibula, anterior view; J, maxilliped, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I, V) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910638" httpUri="https://zenodo.org/record/4910638/files/figure.png" pageId="61" pageNumber="62">Figs. 35J</figureCitation>
,
<figureCitation id="78EA79F3FF93FFEE658EF87C0381F81F" box="[457,506,1930,1955]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="61" pageNumber="62">41A</figureCitation>
) segmentation, armature and most ornamentation as in
<emphasis id="D2A5B964FF93FFEE6036F87C070BF81E" box="[1137,1392,1930,1954]" italics="true" pageId="61" pageNumber="62">
<taxonomicName id="27D11EF5FF93FFEE6036F87C0717F81E" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1137,1388,1930,1954]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="calliope">Wellstenhelia calliope</taxonomicName>
,
</emphasis>
but outer distal corner of coxa produced into pronounced and blunt process.
</paragraph>
<caption id="B4AE35FEFF90FFED64D0F9EE013DF936" ID-DOI="http://doi.org/10.5281/zenodo.4910650" ID-Zenodo-Dep="4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="62" pageNumber="63" startId="62.[151,249,1560,1582]" targetBox="[151,1436,193,1539]" targetPageId="62">
<paragraph id="E06E6576FF90FFED64D0F9EE013DF936" blockId="62.[151,1437,1560,1674]" pageId="62" pageNumber="63">
<emphasis id="D2A5B964FF90FFED64D0F9EE035BF991" bold="true" box="[151,288,1560,1582]" pageId="62" pageNumber="63">FIGURE 41.</emphasis>
<taxonomicName id="27D11EF5FF90FFED656FF9EE0066F992" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[296,541,1560,1582]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="polyhymnia" status="sp. nov.">
<emphasis id="D2A5B964FF90FFED656FF9EE0066F992" box="[296,541,1560,1582]" italics="true" pageId="62" pageNumber="63">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FF90FFED6663F9EF000DF992" bold="true" box="[548,630,1561,1582]" pageId="62" pageNumber="63">
<taxonomicNameLabel id="C996041FFF90FFED6663F9EF000DF992" box="[548,630,1561,1582]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C).
</paragraph>
</caption>
<paragraph id="E06E6576FF90FFED6480F9470751F8AE" blockId="62.[151,1436,1713,2027]" pageId="62" pageNumber="63">
First swimming leg (
<figureCitation id="78EA79F3FF90FFED65E9F947005BF976" box="[430,544,1713,1738]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="62" pageNumber="63">Figs. 36A</figureCitation>
,
<figureCitation id="78EA79F3FF90FFED666DF9470021F976" box="[554,602,1713,1738]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="62" pageNumber="63">41B</figureCitation>
) as in
<taxonomicName id="27D11EF5FF90FFED66E5F94401E3F976" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[674,920,1714,1738]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF90FFED66E5F94401E3F976" box="[674,920,1714,1738]" italics="true" pageId="62" pageNumber="63">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except coxa and basis without inner spinules, first endopodal segment much shorter and its inner seta longer and stronger (both arrowed in
<figureCitation id="78EA79F3FF90FFED60F4F9230760F952" box="[1203,1307,1749,1774]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="62" pageNumber="63">Fig. 36A</figureCitation>
), and distal inner seta on second endopodal segment inserted further away from distal margin (arrowed in
<figureCitation id="78EA79F3FF90FFED60F1F90F0766F8AE" box="[1206,1309,1785,1810]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="62" pageNumber="63">Fig. 36A</figureCitation>
).
</paragraph>
<paragraph id="E06E6576FF90FFED6480F8EB00E4F81E" blockId="62.[151,1436,1713,2027]" pageId="62" pageNumber="63">
Second swimming leg (
<figureCitation id="78EA79F3FF90FFED6589F8EB00FAF889" box="[462,641,1821,1846]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="62" pageNumber="63">Figs. 36B, C, D</figureCitation>
,
<figureCitation id="78EA79F3FF90FFED66CBF8EB00C0F88A" box="[652,699,1821,1846]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="62" pageNumber="63">41B</figureCitation>
) as in
<taxonomicName id="27D11EF5FF90FFED6743F8E80180F88A" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[772,1019,1822,1846]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF90FFED6743F8E80180F88A" box="[772,1019,1822,1846]" italics="true" pageId="62" pageNumber="63">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except intercoxal sclerite with sharp distal projections (arrowed in
<figureCitation id="78EA79F3FF90FFED65B7F8B7002DF8E6" box="[496,598,1857,1882]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="62" pageNumber="63">Fig. 36B</figureCitation>
) and smooth, first endopodal segment with slender seta (arrowed in
<figureCitation id="78EA79F3FF90FFED6137F8B702B2F8C2" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="62" pageNumber="63">Fig. 36C</figureCitation>
), second endopodal segment with two inner setae (arrowed in
<figureCitation id="78EA79F3FF90FFED67E5F8930671F8C2" box="[930,1034,1893,1918]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="62" pageNumber="63">Fig. 36C</figureCitation>
), and third exopodal segment with shorter distal inner seta (arrowed in
<figureCitation id="78EA79F3FF90FFED666CF87F00E8F81E" box="[555,659,1929,1954]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="62" pageNumber="63">Fig. 36D</figureCitation>
).
</paragraph>
<paragraph id="E06E6576FF90FFEC6480F85B008FFF44" blockId="62.[151,1436,1713,2027]" lastBlockId="63.[151,1437,151,2013]" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
Third swimming leg (
<figureCitation id="78EA79F3FF90FFED65F9F85B0056F87A" box="[446,557,1965,1990]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="62" pageNumber="63">Figs. 36E</figureCitation>
,
<figureCitation id="78EA79F3FF90FFED667DF85B0011F87A" box="[570,618,1965,1990]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="62" pageNumber="63">41B</figureCitation>
) as in
<taxonomicName id="27D11EF5FF90FFED66F0F85801D4F87A" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[695,943,1966,1990]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF90FFED66F0F85801D4F87A" box="[695,943,1966,1990]" italics="true" pageId="62" pageNumber="63">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except intercoxal sclerite with sharp distal projections (arrowed in
<figureCitation id="78EA79F3FF90FFED65F5F8240067F857" box="[434,540,2002,2027]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="62" pageNumber="63">Fig. 36E</figureCitation>
), inner distal row of spinules on coxa missing (arrowed in
<figureCitation id="78EA79F3FF90FFED60A2F8240735F857" box="[1253,1358,2002,2027]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="62" pageNumber="63">Fig. 36E</figureCitation>
), third exopodal segment with only two outer spines and distal inner seta short and slender (both arrowed in
<figureCitation id="78EA79F3FF91FFEC616EFF6107F4FF0C" box="[1321,1423,151,176]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="63" pageNumber="64">Fig. 36E</figureCitation>
), first endopodal segment with slender inner seta (arrowed in
<figureCitation id="78EA79F3FF91FFEC6715FF4A01C2FF69" box="[850,953,188,213]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="63" pageNumber="64">Fig. 36E</figureCitation>
), and distal inner seta on third endopodal segment very short and slender (arrowed in
<figureCitation id="78EA79F3FF91FFEC66C4FF290093FF44" box="[643,744,223,248]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="63" pageNumber="64">Fig. 36E</figureCitation>
).
</paragraph>
<paragraph id="E06E6576FF91FFEC6480FEF203F4FE34" blockId="63.[151,1437,151,2013]" pageId="63" pageNumber="64">
Fourth swimming leg (
<figureCitation id="78EA79F3FF91FFEC659FFEF20044FEA1" box="[472,575,260,285]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="63" pageNumber="64">Fig. 36F</figureCitation>
) as in
<taxonomicName id="27D11EF5FF91FFEC66DEFEF201EEFEA0" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[665,917,260,284]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF91FFEC66DEFEF201EEFEA0" box="[665,917,260,284]" italics="true" pageId="63" pageNumber="64">Wellstenhelia calliope</emphasis>
</taxonomicName>
, except intercoxal sclerite with sharp distal projections (arrowed in
<figureCitation id="78EA79F3FF91FFEC65E5FED10078FEFC" box="[418,515,295,320]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="63" pageNumber="64">Fig. 36F</figureCitation>
), third exopodal segment with only two outer spines and distal inner seta short and slender (both arrowed in
<figureCitation id="78EA79F3FF91FFEC6582FEBA0057FED9" box="[453,556,332,357]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="63" pageNumber="64">Fig. 36F</figureCitation>
), and distal inner seta on third endopodal segment very short and slender (arrowed in
<figureCitation id="78EA79F3FF91FFEC6567FE9903F9FE34" box="[288,386,367,392]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="63" pageNumber="64">Fig. 36F</figureCitation>
).
</paragraph>
<paragraph id="E06E6576FF91FFEC6480FE62012CFD39" blockId="63.[151,1437,151,2013]" pageId="63" pageNumber="64">
Fifth leg (
<figureCitation id="78EA79F3FF91FFEC6570FE6203B0FE17" box="[311,459,404,429]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="63" pageNumber="64">Figs. 34A, B</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC6590FE620051FE17" box="[471,554,404,429]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="63" pageNumber="64">36G, H</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC6671FE620012FE11" box="[566,617,404,429]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="63" pageNumber="64">37A</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC6632FE6200BEFE11" box="[629,709,404,429]" captionStart="FIGURE 40" captionStartId="61.[151,249,1596,1618]" captionTargetBox="[151,1436,218,1563]" captionTargetId="figure-185@61.[151,1436,218,1563]" captionTargetPageId="61" captionText="FIGURE 40. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–E, female paratype 2, F, female paratype 3: A, habitus, lateral view; B, cephalothorax, lateral view; C pedigerous somites, lateral view; D, first urosomite and genital double-somite, lateral view (exopod of fifth leg broken off); E, anal somite and caudal rami, lateral view; F, fifth legs and genital double-somite with one empty egg-sack, ventral view. Scale bars 50 µm (C, D, E, F), 100 µm (B), and 200 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.4910648" httpUri="https://zenodo.org/record/4910648/files/figure.png" pageId="63" pageNumber="64">40D, F</figureCitation>
) segmentation, general shape, and number of armature elements on endopodal lobe as in
<emphasis id="D2A5B964FF91FFEC65EFFE4E00DDFE6C" box="[424,678,440,464]" italics="true" pageId="63" pageNumber="64">
<taxonomicName id="27D11EF5FF91FFEC65EFFE4E00D9FE6C" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[424,674,440,464]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="calliope">Wellstenhelia calliope</taxonomicName>
,
</emphasis>
except baseoendopod without spiniform process at base of exopod, with row of strong spinules instead (arrowed in
<figureCitation id="78EA79F3FF91FFEC66F9FE2A015CFE49" box="[702,807,476,501]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="63" pageNumber="64">Fig. 36H</figureCitation>
), additional anterior pore at base of basal seta. Exopod ovoid, about 2.1 times as long as greatest width, with long outer spinules (arrowed in
<figureCitation id="78EA79F3FF91FFEC6021FE090682FDAB" box="[1126,1273,511,536]" captionStart="FIGURE 34" captionStartId="54.[151,249,1885,1907]" captionTargetBox="[209,1381,197,1846]" captionTargetId="figure-102@54.[202,1381,193,1850]" captionTargetPageId="54" captionText="FIGURE 34. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, habitus, dorsal view; B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, antennula, dorsal view; E, antenna, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910636" httpUri="https://zenodo.org/record/4910636/files/figure.png" pageId="63" pageNumber="64">Figs. 34A, B</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC6141FE090741FDA4" box="[1286,1338,511,536]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="63" pageNumber="64">36G</figureCitation>
), shorter inner spinules, and anterior pore, with five armature elements, second from inner side much longer than third (arrowed in
<figureCitation id="78EA79F3FF91FFEC6564FDB103F7FDDC" box="[291,396,583,608]" captionStart="FIGURE 36" captionStartId="56.[151,249,1889,1911]" captionTargetBox="[185,1400,184,1868]" captionTargetId="figure-101@56.[183,1402,173,1873]" captionTargetPageId="56" captionText="FIGURE 36. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, first swimming leg, anterior view; B, intercoxal sclerite of second swimming leg, anterior view; C, endopod of second swimming leg, anterior view; D, third exopodal segment of second swimming leg, anterior view; E, third swimming leg, anterior view; F, fourth swimming leg, anterior view; G, exopod of fifth leg, anterior view; H, fifth leg, anterior view;. Arrowheads point to most prominent specific features. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910640" httpUri="https://zenodo.org/record/4910640/files/figure.png" pageId="63" pageNumber="64">Fig. 36G</figureCitation>
). Length ratio of endopodal setae, starting from inner side, 1: 1: 2.4: 1.5. Length ratio of exopodal setae, starting from inner side, 1: 0.5: 0.2: 0.7: 0.3.
</paragraph>
<paragraph id="E06E6576FF91FFEC6480FD79036DFD4C" blockId="63.[151,1437,151,2013]" pageId="63" pageNumber="64">
Sixth legs (
<figureCitation id="78EA79F3FF91FFEC650CFD7903ACFD14" box="[331,471,655,680]" captionStart="FIGURE 35" captionStartId="55.[151,249,1829,1851]" captionTargetBox="[218,1368,192,1805]" captionTargetId="figure-138@55.[210,1371,179,1809]" captionTargetPageId="55" captionText="FIGURE 35. Itostenhelia polyhymnia sp. nov., line drawings, female holotype: A, right side of cephalothoracic shiled, dissected, flattened, and partly broken; B, pleurons of third and fourth pedigerous somites, dissected and flattened; C, genital double-somite, ventral view; D, genital double-somite, lateral view; E, syncoxa of mandibula, antero-ventral view; F, syncoxa of mandibula, posterior view; G, mandibular palp, anterior view; H, maxillula, posterior view; I, mandibula, anterior view; J, maxilliped, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I, V) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910638" httpUri="https://zenodo.org/record/4910638/files/figure.png" pageId="63" pageNumber="64">Fig. 35C, D</figureCitation>
) simple cuticular plates covering paired genital apertures, inserted close to anterior margin of genital double-somite ventro-laterally, with single bipinnate seta, joined together into short but wide operculum.
</paragraph>
<paragraph id="E06E6576FF91FFEC6480FD0A0681FC19" blockId="63.[151,1437,151,2013]" pageId="63" pageNumber="64">
Male (based on
<typeStatus id="3F6ADBD4FF91FFEC6539FD0A03A3FCA9" box="[382,472,764,789]" pageId="63" pageNumber="64" type="allotype">allotype</typeStatus>
and
<specimenCount id="F6D7AEFFFF91FFEC6656FD0A00C3FCA9" box="[529,696,764,789]" pageId="63" pageNumber="64" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
). Body length from
<quantity id="2729C893FF91FFEC67DAFD0A063FFCA8" box="[925,1092,764,789]" metricMagnitude="-1" metricUnit="m" metricValue="3.845" metricValueMax="4.12" metricValueMin="3.57" pageId="63" pageNumber="64" unit="mm" value="384.5" valueMax="412.0" valueMin="357.0">357 to 412 µm</quantity>
(
<quantity id="2729C893FF91FFEC6013FD0A06D7FCA8" box="[1108,1196,764,789]" metricMagnitude="-1" metricUnit="m" metricValue="3.82" pageId="63" pageNumber="64" unit="mm" value="382.0">382 µm</quantity>
in
<typeStatus id="3F6ADBD4FF91FFEC6095FD0A074AFCA9" box="[1234,1329,764,789]" pageId="63" pageNumber="64" type="allotype">allotype</typeStatus>
). Habitus (
<figureCitation id="78EA79F3FF91FFEC64D8FCE9037FFC84" box="[159,260,799,824]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="63" pageNumber="64">Fig. 41C</figureCitation>
), colour, rostrum, shape and ornamentation of cephalothorax (
<figureCitation id="78EA79F3FF91FFEC67FEFCE90651FC84" box="[953,1066,799,824]" captionStart="FIGURE 38" captionStartId="58.[151,249,1884,1906]" captionTargetBox="[192,1391,186,1878]" captionTargetId="figure-97@58.[183,1404,179,1893]" captionTargetPageId="58" captionText="FIGURE 38. Itostenhelia polyhymnia sp. nov., line drawings, male allotype: A, cephalothoracic shield and pleurons of second to fourth pedigerous somites, lateral view; B, urosome, ventral view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910644" httpUri="https://zenodo.org/record/4910644/files/figure.png" pageId="63" pageNumber="64">Figs. 38A</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC6072FCE90613FC84" box="[1077,1128,799,824]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="63" pageNumber="64">41D</figureCitation>
), second pedigerous somite (
<figureCitation id="78EA79F3FF91FFEC64D8FCB2036AFCE1" box="[159,273,836,861]" captionStart="FIGURE 38" captionStartId="58.[151,249,1884,1906]" captionTargetBox="[192,1391,186,1878]" captionTargetId="figure-97@58.[183,1404,179,1893]" captionTargetPageId="58" captionText="FIGURE 38. Itostenhelia polyhymnia sp. nov., line drawings, male allotype: A, cephalothoracic shield and pleurons of second to fourth pedigerous somites, lateral view; B, urosome, ventral view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910644" httpUri="https://zenodo.org/record/4910644/files/figure.png" pageId="63" pageNumber="64">Figs. 38A</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC655AFCB2032BFCE1" box="[285,336,836,861]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="63" pageNumber="64">41D</figureCitation>
), third pedigerous somite (
<figureCitation id="78EA79F3FF91FFEC663AFCB2009FFCE1" box="[637,740,836,861]" captionStart="FIGURE 38" captionStartId="58.[151,249,1884,1906]" captionTargetBox="[192,1391,186,1878]" captionTargetId="figure-97@58.[183,1404,179,1893]" captionTargetPageId="58" captionText="FIGURE 38. Itostenhelia polyhymnia sp. nov., line drawings, male allotype: A, cephalothoracic shield and pleurons of second to fourth pedigerous somites, lateral view; B, urosome, ventral view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910644" httpUri="https://zenodo.org/record/4910644/files/figure.png" pageId="63" pageNumber="64">Fig. 38A</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC66B7FCB20158FCE1" box="[752,803,836,861]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="63" pageNumber="64">41D</figureCitation>
), fourth pedigerous somite (
<figureCitation id="78EA79F3FF91FFEC6018FCB206BCFCE1" box="[1119,1223,836,861]" captionStart="FIGURE 38" captionStartId="58.[151,249,1884,1906]" captionTargetBox="[192,1391,186,1878]" captionTargetId="figure-97@58.[183,1404,179,1893]" captionTargetPageId="58" captionText="FIGURE 38. Itostenhelia polyhymnia sp. nov., line drawings, male allotype: A, cephalothoracic shield and pleurons of second to fourth pedigerous somites, lateral view; B, urosome, ventral view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910644" httpUri="https://zenodo.org/record/4910644/files/figure.png" pageId="63" pageNumber="64">Fig. 38A</figureCitation>
), ornamentation of first urosomite (
<figureCitation id="78EA79F3FF91FFEC650BFC9103C8FC3C" box="[332,435,871,896]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="63" pageNumber="64">Fig. 37C</figureCitation>
), caudal rami (
<figureCitation id="78EA79F3FF91FFEC661BFC9100B6FC3C" box="[604,717,871,896]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="63" pageNumber="64">Figs. 37C</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC669EFC910171FC3C" box="[729,778,871,896]" captionStart="FIGURE 38" captionStartId="58.[151,249,1884,1906]" captionTargetBox="[192,1391,186,1878]" captionTargetId="figure-97@58.[183,1404,179,1893]" captionTargetPageId="58" captionText="FIGURE 38. Itostenhelia polyhymnia sp. nov., line drawings, male allotype: A, cephalothoracic shield and pleurons of second to fourth pedigerous somites, lateral view; B, urosome, ventral view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910644" httpUri="https://zenodo.org/record/4910644/files/figure.png" pageId="63" pageNumber="64">38B</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC6750FC91013DFC3C" box="[791,838,871,896]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="63" pageNumber="64">41F</figureCitation>
), antenna, labrum, paragnaths, mandibula, maxillula, maxilla, maxilliped, first swimming leg, third swimming leg, and fourth swimming leg as in female.
</paragraph>
<paragraph id="E06E6576FF91FFEC6480FC590115FBE4" blockId="63.[151,1437,151,2013]" pageId="63" pageNumber="64">
Genital somite (
<figureCitation id="78EA79F3FF91FFEC65C0FC590386FC74" box="[391,509,943,968]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="63" pageNumber="64">Figs. 37C</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC6648FC59003BFC74" box="[527,576,943,968]" captionStart="FIGURE 38" captionStartId="58.[151,249,1884,1906]" captionTargetBox="[192,1391,186,1878]" captionTargetId="figure-97@58.[183,1404,179,1893]" captionTargetPageId="58" captionText="FIGURE 38. Itostenhelia polyhymnia sp. nov., line drawings, male allotype: A, cephalothoracic shield and pleurons of second to fourth pedigerous somites, lateral view; B, urosome, ventral view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910644" httpUri="https://zenodo.org/record/4910644/files/figure.png" pageId="63" pageNumber="64">38B</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC6615FC5900FBFC74" box="[594,640,943,968]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="63" pageNumber="64">41E</figureCitation>
) somewhat wider than in
<taxonomicName id="27D11EF5FF91FFEC6785FC4606C4FC74" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[962,1215,944,968]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF91FFEC6785FC4606C4FC74" box="[962,1215,944,968]" italics="true" pageId="63" pageNumber="64">Wellstenhelia calliope</emphasis>
</taxonomicName>
and without large spinules; lateral pore no. 68 missing; additionally ornamented with several rows of minute spinules dorso-laterally. Third urosomite (
<figureCitation id="78EA79F3FF91FFEC651AFC0103B5FBAC" box="[349,462,1015,1040]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="63" pageNumber="64">Figs. 37C</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC659DFC010070FBAC" box="[474,523,1015,1040]" captionStart="FIGURE 38" captionStartId="58.[151,249,1884,1906]" captionTargetBox="[192,1391,186,1878]" captionTargetId="figure-97@58.[183,1404,179,1893]" captionTargetPageId="58" captionText="FIGURE 38. Itostenhelia polyhymnia sp. nov., line drawings, male allotype: A, cephalothoracic shield and pleurons of second to fourth pedigerous somites, lateral view; B, urosome, ventral view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910644" httpUri="https://zenodo.org/record/4910644/files/figure.png" pageId="63" pageNumber="64">38B</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC665FFC01003DFBAC" box="[536,582,1015,1040]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="63" pageNumber="64">41E</figureCitation>
) as in
<emphasis id="D2A5B964FF91FFEC66D4FC0E01EAFBAC" box="[659,913,1016,1040]" italics="true" pageId="63" pageNumber="64">
<taxonomicName id="27D11EF5FF91FFEC66D4FC0E01F6FBAC" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[659,909,1016,1040]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="calliope">Wellstenhelia calliope</taxonomicName>
,
</emphasis>
except ventral row of spinules not interrupted between sensilla pair no. 73 (arrowed in
<figureCitation id="78EA79F3FF91FFEC66C2FBEA0095FB89" box="[645,750,1052,1077]" captionStart="FIGURE 38" captionStartId="58.[151,249,1884,1906]" captionTargetBox="[192,1391,186,1878]" captionTargetId="figure-97@58.[183,1404,179,1893]" captionTargetPageId="58" captionText="FIGURE 38. Itostenhelia polyhymnia sp. nov., line drawings, male allotype: A, cephalothoracic shield and pleurons of second to fourth pedigerous somites, lateral view; B, urosome, ventral view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910644" httpUri="https://zenodo.org/record/4910644/files/figure.png" pageId="63" pageNumber="64">Fig. 38B</figureCitation>
) and these sensilla also more widely spaced; additionally ornamented with several rows of minute spinules dorso-laterally.
</paragraph>
<paragraph id="E06E6576FF91FFEC6480FB9206ECFB1C" blockId="63.[151,1437,151,2013]" pageId="63" pageNumber="64">
Fourth urosomite (
<figureCitation id="78EA79F3FF91FFEC65DFFB920072FBC1" box="[408,521,1124,1149]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="63" pageNumber="64">Figs. 37C</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC6653FB92003FFBC1" box="[532,580,1124,1149]" captionStart="FIGURE 38" captionStartId="58.[151,249,1884,1906]" captionTargetBox="[192,1391,186,1878]" captionTargetId="figure-97@58.[183,1404,179,1893]" captionTargetPageId="58" captionText="FIGURE 38. Itostenhelia polyhymnia sp. nov., line drawings, male allotype: A, cephalothoracic shield and pleurons of second to fourth pedigerous somites, lateral view; B, urosome, ventral view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910644" httpUri="https://zenodo.org/record/4910644/files/figure.png" pageId="63" pageNumber="64">38B</figureCitation>
) pore and sensilla pattern as in female (i.e. unique pore pair no. VI present), but additionally ornamented with ventral row of spinules of various sizes, some quite large.
</paragraph>
<paragraph id="E06E6576FF91FFEC6480FB5A000DFB54" blockId="63.[151,1437,151,2013]" pageId="63" pageNumber="64">
Fifth urosomite (
<figureCitation id="78EA79F3FF91FFEC65C3FB5A038DFB79" box="[388,502,1196,1221]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="63" pageNumber="64">Figs. 37C</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC6646FB5A0049FB79" box="[513,562,1196,1221]" captionStart="FIGURE 38" captionStartId="58.[151,249,1884,1906]" captionTargetBox="[192,1391,186,1878]" captionTargetId="figure-97@58.[183,1404,179,1893]" captionTargetPageId="58" captionText="FIGURE 38. Itostenhelia polyhymnia sp. nov., line drawings, male allotype: A, cephalothoracic shield and pleurons of second to fourth pedigerous somites, lateral view; B, urosome, ventral view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910644" httpUri="https://zenodo.org/record/4910644/files/figure.png" pageId="63" pageNumber="64">38B</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC6679FB5A0010FB79" box="[574,619,1196,1221]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="63" pageNumber="64">41F</figureCitation>
) without pores or sensilla as in female, but with posterior ventral row of spinules of various sizes, some quite large.
</paragraph>
<paragraph id="E06E6576FF91FFEC6480FB02034BFA8C" blockId="63.[151,1437,151,2013]" pageId="63" pageNumber="64">
Anal somite (
<figureCitation id="78EA79F3FF91FFEC6518FB0203ABFAB1" box="[351,464,1268,1293]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="63" pageNumber="64">Figs. 37C</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC659CFB020077FAB1" box="[475,524,1268,1293]" captionStart="FIGURE 38" captionStartId="58.[151,249,1884,1906]" captionTargetBox="[192,1391,186,1878]" captionTargetId="figure-97@58.[183,1404,179,1893]" captionTargetPageId="58" captionText="FIGURE 38. Itostenhelia polyhymnia sp. nov., line drawings, male allotype: A, cephalothoracic shield and pleurons of second to fourth pedigerous somites, lateral view; B, urosome, ventral view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910644" httpUri="https://zenodo.org/record/4910644/files/figure.png" pageId="63" pageNumber="64">38B</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC665FFB02003EFAB1" box="[536,581,1268,1293]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="63" pageNumber="64">41F</figureCitation>
) as in female, except lateral pores nos. 79 &amp; 80 present in
<typeStatus id="3F6ADBD4FF91FFEC609DFB02074FFAB1" box="[1242,1332,1268,1293]" pageId="63" pageNumber="64" type="allotype">allotype</typeStatus>
(missing in
<typeStatus id="3F6ADBD4FF91FFEC64F3FAE1035EFA8C" box="[180,293,1303,1328]" pageId="63" pageNumber="64" type="paratype">paratypes</typeStatus>
).
</paragraph>
<paragraph id="E06E6576FF91FFEC6480FACA004AFAC4" blockId="63.[151,1437,151,2013]" pageId="63" pageNumber="64">
Caudal rami (
<figureCitation id="78EA79F3FF91FFEC6519FACA03B5FAE9" box="[350,462,1340,1365]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="63" pageNumber="64">Figs. 37C</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC659EFACA0071FAE9" box="[473,522,1340,1365]" captionStart="FIGURE 38" captionStartId="58.[151,249,1884,1906]" captionTargetBox="[192,1391,186,1878]" captionTargetId="figure-97@58.[183,1404,179,1893]" captionTargetPageId="58" captionText="FIGURE 38. Itostenhelia polyhymnia sp. nov., line drawings, male allotype: A, cephalothoracic shield and pleurons of second to fourth pedigerous somites, lateral view; B, urosome, ventral view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910644" httpUri="https://zenodo.org/record/4910644/files/figure.png" pageId="63" pageNumber="64">38B</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC6652FACA0039FAE9" box="[533,578,1340,1365]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="63" pageNumber="64">41F</figureCitation>
) about four times as long as wide in ventral view, with two rows of relatively short inner spinules in anterior third.
</paragraph>
<paragraph id="E06E6576FF91FFEC6480FA72060EFA21" blockId="63.[151,1437,151,2013]" box="[199,1141,1412,1437]" pageId="63" pageNumber="64">
Antennula (
<figureCitation id="78EA79F3FF91FFEC650CFA7203A7FA21" box="[331,476,1412,1437]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="63" pageNumber="64">Figs. 41C, D</figureCitation>
) as in
<emphasis id="D2A5B964FF91FFEC6661FA720145FA20" box="[550,830,1412,1436]" italics="true" pageId="63" pageNumber="64">
<taxonomicName id="27D11EF5FF91FFEC6661FA720141FA20" box="[550,826,1412,1436]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="terpisichore">Itostenhelia terpisichore</taxonomicName>
,
</emphasis>
except completely clasped.
</paragraph>
<paragraph id="E06E6576FF91FFEC6480FA510190F924" blockId="63.[151,1437,151,2013]" pageId="63" pageNumber="64">
Second swimming leg (
<figureCitation id="78EA79F3FF91FFEC65ACFA510021FA7C" box="[491,602,1447,1472]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="63" pageNumber="64">Fig. 37D</figureCitation>
) with coxa, basis, and exopod as in female. Endopod significantly transformed, with second and third segment fused (arrowed in
<figureCitation id="78EA79F3FF91FFEC671BFA3A01BFFA59" box="[860,964,1484,1509]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="63" pageNumber="64">Fig. 37D</figureCitation>
), third segment smaller and without outer spinules (arrowed in
<figureCitation id="78EA79F3FF91FFEC65C1FA190395F9B4" box="[390,494,1519,1544]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="63" pageNumber="64">Fig. 37D</figureCitation>
), outer apical spine enlarged, with small inner hump at proximal fifth of its length, mostly smooth, with only two anterior spinules at midlength and short row of slender inner spinules near distal tip; small notch on either side as remnant of ancestral segmentation; proximal seta on ancestral second segment short and slender, about as long as segment’s width; inner spiniform element on ancestral third segment also strong but bipinnate and not transformed, about 1.7 times as long as outer apical spine.
</paragraph>
<paragraph id="E06E6576FF91FFEC6480F95201EBF829" blockId="63.[151,1437,151,2013]" pageId="63" pageNumber="64">
Fifth leg (
<figureCitation id="78EA79F3FF91FFEC6570F95203D2F901" box="[311,425,1700,1725]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="63" pageNumber="64">Figs. 37C</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC65F2F952039DF901" box="[437,486,1700,1725]" captionStart="FIGURE 38" captionStartId="58.[151,249,1884,1906]" captionTargetBox="[192,1391,186,1878]" captionTargetId="figure-97@58.[183,1404,179,1893]" captionTargetPageId="58" captionText="FIGURE 38. Itostenhelia polyhymnia sp. nov., line drawings, male allotype: A, cephalothoracic shield and pleurons of second to fourth pedigerous somites, lateral view; B, urosome, ventral view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910644" httpUri="https://zenodo.org/record/4910644/files/figure.png" pageId="63" pageNumber="64">38B</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC65B4F952005BF901" box="[499,544,1700,1725]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="63" pageNumber="64">41E</figureCitation>
) general shape and segmentation as in
<taxonomicName id="27D11EF5FF91FFEC679AF95206AEF900" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[989,1237,1700,1724]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="calliope">
<emphasis id="D2A5B964FF91FFEC679AF95206AEF900" box="[989,1237,1700,1724]" italics="true" pageId="63" pageNumber="64">Wellstenhelia calliope</emphasis>
</taxonomicName>
but armature and ornamentation very different; fused endopodal lobes forming slightly convex plate, with four spines (two belonging to each leg; arrowed in
<figureCitation id="78EA79F3FF91FFEC6658F91A00FFF8B9" box="[543,644,1772,1797]" captionStart="FIGURE 38" captionStartId="58.[151,249,1884,1906]" captionTargetBox="[192,1391,186,1878]" captionTargetId="figure-97@58.[183,1404,179,1893]" captionTargetPageId="58" captionText="FIGURE 38. Itostenhelia polyhymnia sp. nov., line drawings, male allotype: A, cephalothoracic shield and pleurons of second to fourth pedigerous somites, lateral view; B, urosome, ventral view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910644" httpUri="https://zenodo.org/record/4910644/files/figure.png" pageId="63" pageNumber="64">Fig. 38B</figureCitation>
) and long posterior row of spinules between endopodal armature and exopod (arrowed in
<figureCitation id="78EA79F3FF91FFEC65C7F8F90393F894" box="[384,488,1807,1832]" captionStart="FIGURE 38" captionStartId="58.[151,249,1884,1906]" captionTargetBox="[192,1391,186,1878]" captionTargetId="figure-97@58.[183,1404,179,1893]" captionTargetPageId="58" captionText="FIGURE 38. Itostenhelia polyhymnia sp. nov., line drawings, male allotype: A, cephalothoracic shield and pleurons of second to fourth pedigerous somites, lateral view; B, urosome, ventral view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910644" httpUri="https://zenodo.org/record/4910644/files/figure.png" pageId="63" pageNumber="64">Fig. 38B</figureCitation>
); inner endopodal spine almost six times as long as outer one; exopod ovoid, with several outer spinules, with three inner setae (all short, slender, and smooth) and two apical spines, inner one fused basally to segment; outer apical exopodal spine only slightly shorter than inner endopodal spine and length ratio of exopodal armature, starting from inner side, 1: 0.45: 0.6: 0.3: 0.3.
</paragraph>
<paragraph id="E06E6576FF91FFEC6480F8560284F861" blockId="63.[151,1437,151,2013]" pageId="63" pageNumber="64">
Sixth legs (
<figureCitation id="78EA79F3FF91FFEC6516F85603C7F805" box="[337,444,1952,1977]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="63" pageNumber="64">Fig. 37C</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC658AF8560385F805" box="[461,510,1952,1977]" captionStart="FIGURE 38" captionStartId="58.[151,249,1884,1906]" captionTargetBox="[192,1391,186,1878]" captionTargetId="figure-97@58.[183,1404,179,1893]" captionTargetPageId="58" captionText="FIGURE 38. Itostenhelia polyhymnia sp. nov., line drawings, male allotype: A, cephalothoracic shield and pleurons of second to fourth pedigerous somites, lateral view; B, urosome, ventral view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910644" httpUri="https://zenodo.org/record/4910644/files/figure.png" pageId="63" pageNumber="64">38B</figureCitation>
,
<figureCitation id="78EA79F3FF91FFEC6648F8560045F805" box="[527,574,1952,1977]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="63" pageNumber="64">41E</figureCitation>
) as in
<emphasis id="D2A5B964FF91FFEC66DEF85601E7F804" box="[665,924,1952,1976]" italics="true" pageId="63" pageNumber="64">
<taxonomicName id="27D11EF5FF91FFEC66DEF85601E3F804" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[665,920,1952,1976]" class="Maxillopoda" family="Miraciidae" genus="Wellstenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="calliope">Wellstenhelia calliope</taxonomicName>
,
</emphasis>
except right leg distinct at base and more movable.
</paragraph>
<paragraph id="E06E6576FFEEFF936480FF6106B6FF44" blockId="64.[151,1436,151,861]" pageId="64" pageNumber="65">
<emphasis id="D2A5B964FFEEFF936480FF610336FF0C" bold="true" box="[199,333,151,176]" pageId="64" pageNumber="65">Variability.</emphasis>
Most morphological features are extremely conservative, including the sensilla and pores pattern of somites, and length ratio of different armature on appendages. Except for body length, the only other variable feature was the presence/absence of lateral pores on the male anal somite (
<figureCitation id="78EA79F3FFEEFF93679EFF290632FF44" box="[985,1097,223,248]" captionStart="FIGURE 37" captionStartId="57.[151,249,1915,1937]" captionTargetBox="[189,1399,185,1890]" captionTargetId="figure-91@57.[181,1405,175,1893]" captionTargetPageId="57" captionText="FIGURE 37. Itostenhelia polyhymnia sp. nov., line drawings, A–B, female holotype, C–D, male allotype: A, urosome, ventral view; B, right caudal ramus, lateral view; C, urosome, lateral view; D, endopod of second swimming leg, anterior view. Arabic numerals and hash mark (#) indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate unique pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910642" httpUri="https://zenodo.org/record/4910642/files/figure.png" pageId="64" pageNumber="65">Figs. 37C</figureCitation>
,
<figureCitation id="78EA79F3FFEEFF936013FF2906FEFF44" box="[1108,1157,223,248]" captionStart="FIGURE 38" captionStartId="58.[151,249,1884,1906]" captionTargetBox="[192,1391,186,1878]" captionTargetId="figure-97@58.[183,1404,179,1893]" captionTargetPageId="58" captionText="FIGURE 38. Itostenhelia polyhymnia sp. nov., line drawings, male allotype: A, cephalothoracic shield and pleurons of second to fourth pedigerous somites, lateral view; B, urosome, ventral view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–VII) indicate unique sensilla and pores Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910644" httpUri="https://zenodo.org/record/4910644/files/figure.png" pageId="64" pageNumber="65">38B</figureCitation>
,
<figureCitation id="78EA79F3FFEEFF9360D6FF2906BAFF44" box="[1169,1217,223,248]" captionStart="FIGURE 41" captionStartId="62.[151,249,1560,1582]" captionTargetBox="[151,1436,193,1539]" captionTargetId="figure-244@62.[151,1436,193,1539]" captionTargetPageId="62" captionText="FIGURE 41. Itostenhelia polyhymnia sp. nov., scanning electron micrographs, A–B, female paratype 3, C–F, male paratype: A, mouth appendages, ventral view; B, tips of swimming legs, ventral view; C, habitus, lateral view; D, prosome, lateral view; E, second (genital) and third urosomites with fifth and sixth legs, lateral view; F, anal somite and caudal rami, lateral view. Scale bars 20 µm (A, B, E), 30 µm (F), 50 µm (D), and 100 µm (C)." figureDoi="http://doi.org/10.5281/zenodo.4910650" httpUri="https://zenodo.org/record/4910650/files/figure.png" pageId="64" pageNumber="65">41F</figureCitation>
).
</paragraph>
<paragraph id="E06E6576FFEEFF936480FEF202A7FCE1" blockId="64.[151,1436,151,861]" pageId="64" pageNumber="65">
<emphasis id="D2A5B964FFEEFF936480FEF20395FEA1" bold="true" box="[199,494,260,285]" pageId="64" pageNumber="65">Morphological affinities.</emphasis>
The Korean
<taxonomicName id="27D11EF5FFEEFF9366C5FEF201EAFEA0" authority="Karanovic &amp; Kim, 2014" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[642,913,260,284]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="polyhymnia" status="sp. nov.">
<emphasis id="D2A5B964FFEEFF9366C5FEF201EAFEA0" box="[642,913,260,284]" italics="true" pageId="64" pageNumber="65">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
<emphasis id="D2A5B964FFEEFF9367DFFEF20188FEA1" bold="true" box="[920,1011,260,285]" pageId="64" pageNumber="65">
<taxonomicNameLabel id="C996041FFFEEFF9367DFFEF20188FEA1" box="[920,1011,260,285]" pageId="64" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is morphologically very similar to its only congener, the Russian
<taxonomicName id="27D11EF5FFEEFF936592FEDE01E8FEFC" authority="(Chislenko, 1978)" authorityName="Karanovic &amp; Kim" authorityYear="2014" baseAuthorityName="Chislenko" baseAuthorityYear="1978" box="[469,915,295,320]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="golikovi" status="comb. nov.">
<emphasis id="D2A5B964FFEEFF936592FEDE00C6FEFC" box="[469,701,296,320]" italics="true" pageId="64" pageNumber="65">Itostenhelia golikovi</emphasis>
(
<bibRefCitation id="84401887FFEEFF936689FED101F0FEFC" author="Chislenko, L. L." box="[718,907,295,320]" pageId="64" pageNumber="65" pagination="161 - 192" refId="ref47383" refString="Chislenko, L. L. (1978) New species of harpacticoid copepods (Copepoda, Harpacticoida) from Posyet Bay, Sea of Japan. Trudy Zoologicheskogo Instituta, Akademii Nauk SSSR, Leningrad, 61, 161 - 192. [in Russian]" type="journal article" year="1978">Chislenko, 1978</bibRefCitation>
)
</taxonomicName>
<emphasis id="D2A5B964FFEEFF9367DBFED10664FEFC" bold="true" box="[924,1055,295,320]" pageId="64" pageNumber="65">
<taxonomicNameLabel id="C996041FFFEEFF9367DBFED10664FEFC" box="[924,1055,295,320]" pageId="64" pageNumber="65" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
They show no differences in the antennula, antenna, mouth appendages or swimming legs, and even their sensilla and pores pattern on somites is exactly the same. That, and the fact that the original description of the Russian species by
<bibRefCitation id="84401887FFEEFF9360E4FE990713FE34" author="Chislenko, L. L." box="[1187,1384,367,392]" pageId="64" pageNumber="65" pagination="161 - 192" refId="ref47383" refString="Chislenko, L. L. (1978) New species of harpacticoid copepods (Copepoda, Harpacticoida) from Posyet Bay, Sea of Japan. Trudy Zoologicheskogo Instituta, Akademii Nauk SSSR, Leningrad, 61, 161 - 192. [in Russian]" type="journal article" year="1978">Chislenko (1978)</bibRefCitation>
was lacking in detail and contained several inaccuracies, was the reason we redescribe
<taxonomicName id="27D11EF5FFEEFF936075FE62076CFE10" authorityName="Karanovic &amp; Kim" authorityYear="2014" baseAuthorityName="Chislenko" baseAuthorityYear="1978" box="[1074,1303,404,428]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="golikovi">
<emphasis id="D2A5B964FFEEFF936075FE62076CFE10" box="[1074,1303,404,428]" italics="true" pageId="64" pageNumber="65">Itostenhelia golikovi</emphasis>
</taxonomicName>
below, also providing the first description of the male. Most major differences between these two species are all male characters, and they include proportions of the caudal rami (slightly longer in
<taxonomicName id="27D11EF5FFEEFF936044FE2A076AFE48" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1027,1297,476,500]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="polyhymnia">
<emphasis id="D2A5B964FFEEFF936044FE2A076AFE48" box="[1027,1297,476,500]" italics="true" pageId="64" pageNumber="65">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
), size of the patch of spinules on the inner margin of the caudal rami (smaller in
<taxonomicName id="27D11EF5FFEEFF9367E5FDF606CAFDA4" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[930,1201,512,536]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="polyhymnia">
<emphasis id="D2A5B964FFEEFF9367E5FDF606CAFDA4" box="[930,1201,512,536]" italics="true" pageId="64" pageNumber="65">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
), relative size of the proximal inner seta on the ancestral second endopodal segment of the second leg (smaller in
<taxonomicName id="27D11EF5FFEEFF93615FFDD20361FDDC" authorityName="Karanovic &amp; Kim" authorityYear="2014" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="polyhymnia">
<emphasis id="D2A5B964FFEEFF93615FFDD20361FDDC" italics="true" pageId="64" pageNumber="65">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
), relative length of the inner spiniform seta on the ancestral third endopodal segment of the second leg (shorter in
<taxonomicName id="27D11EF5FFEEFF936553FD9A005FFD38" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[276,548,620,644]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="polyhymnia">
<emphasis id="D2A5B964FFEEFF936553FD9A005FFD38" box="[276,548,620,644]" italics="true" pageId="64" pageNumber="65">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
), shape of the inner margin of the fused ancestral second and third endopodal segments of the second leg (with a noch in
<taxonomicName id="27D11EF5FFEEFF936638FD6601F5FD14" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[639,910,656,680]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="polyhymnia">
<emphasis id="D2A5B964FFEEFF936638FD6601F5FD14" box="[639,910,656,680]" italics="true" pageId="64" pageNumber="65">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
), and length of the outer endopodal seta of the fifth leg (shorter in
<taxonomicName id="27D11EF5FFEEFF936528FD420006FD70" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[367,637,692,716]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="polyhymnia">
<emphasis id="D2A5B964FFEEFF936528FD420006FD70" box="[367,637,692,716]" italics="true" pageId="64" pageNumber="65">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
). The females are extremely similar and can be distinguished mostly by the habitus size (smaller in
<taxonomicName id="27D11EF5FFEEFF9365A5FD2E008EFD4C" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[482,757,728,752]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="polyhymnia">
<emphasis id="D2A5B964FFEEFF9365A5FD2E008EFD4C" box="[482,757,728,752]" italics="true" pageId="64" pageNumber="65">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
) and shape (more slender in
<taxonomicName id="27D11EF5FFEEFF936010FD2E0711FD4C" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1111,1386,728,752]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="polyhymnia">
<emphasis id="D2A5B964FFEEFF936010FD2E0711FD4C" box="[1111,1386,728,752]" italics="true" pageId="64" pageNumber="65">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
), in addition to the shape of the sclerotized parts of the genital receptacles (wider posterior parts in
<taxonomicName id="27D11EF5FFEEFF93615FFD0A0361FC84" authorityName="Karanovic &amp; Kim" authorityYear="2014" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="polyhymnia">
<emphasis id="D2A5B964FFEEFF93615FFD0A0361FC84" italics="true" pageId="64" pageNumber="65">Itostenhelia polyhymnia</emphasis>
</taxonomicName>
). All these differences are arrowed in the drawings of
<taxonomicName id="27D11EF5FFEEFF9367C2FCD60610FC84" authorityName="Karanovic &amp; Kim" authorityYear="2014" baseAuthorityName="Chislenko" baseAuthorityYear="1978" box="[901,1131,800,824]" class="Maxillopoda" family="Miraciidae" genus="Itostenhelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="golikovi">
<emphasis id="D2A5B964FFEEFF9367C2FCD60610FC84" box="[901,1131,800,824]" italics="true" pageId="64" pageNumber="65">Itostenhelia golikovi</emphasis>
</taxonomicName>
(see
<figureCitation id="78EA79F3FFEEFF9360EFFCE90761FC84" box="[1192,1306,799,824]" captionStart="FIGURE 43" captionStartId="66.[151,249,1825,1847]" captionTargetBox="[226,1353,201,1797]" captionTargetId="figure-128@66.[220,1354,193,1804]" captionTargetPageId="66" captionText="FIGURE 43. Itostenhelia golikovi (Chislenko, 1978) comb. nov., line drawings, female holotype: A, habitus, dorsal view (arrow points to separately drawn distal part of inner principal caudal seta); B, habitus, lateral view; C, rostrum, dissected and flattened, dorsal view; D, distal part of antennula, dorsal view; E, labrum, dissected and flattened, anterior view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Geometric shapes (○, ◊) indicate sensilla presumably homologous to those in Wellstenhelia euterpe. Roman numerals (I–V) indicate sensilla and pores presumably homologous to those in Itostenhelia polyhymnia. Arrowhead point to prominent specific feature. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910654" httpUri="https://zenodo.org/record/4910654/files/figure.png" pageId="64" pageNumber="65">Figs. 43A</figureCitation>
,
<figureCitation id="78EA79F3FFEEFF936161FCE90723FC84" box="[1318,1368,799,824]" captionStart="FIGURE 44" captionStartId="67.[151,249,1853,1875]" captionTargetBox="[211,1372,189,1831]" captionTargetId="figure-118@67.[204,1372,183,1835]" captionTargetPageId="67" captionText="FIGURE 44. Itostenhelia golikovi (Chislenko, 1978) comb. nov., line drawings, A–E, female holotype, F, male allotype: A, abdomen, ventral view; B, coxa of third swimmingleg, anterior view; C, third endopodal segment of fourth swimming leg, anterior view; D, fifth leg, anterior view; E, sixth leg undissected on genital double-somite, ventro-lateral view; F, antennula, slightly uncoiled and flattened, dorsal view. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numerals (VI, VII) indicate sensilla and pores presumably homologous to those in Itostenhelia polyhymnia. Arrowhead points to prominent specific feature. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910658" httpUri="https://zenodo.org/record/4910658/files/figure.png" pageId="64" pageNumber="65">44A</figureCitation>
,
<figureCitation id="78EA79F3FFEEFF936122FCE902B4FCE1" captionStart="FIGURE 45" captionStartId="68.[151,249,1892,1914]" captionTargetBox="[204,1390,190,1859]" captionTargetId="figure-100@68.[196,1393,183,1868]" captionTargetPageId="68" captionText="FIGURE 45. Itostenhelia golikovi (Chislenko, 1978) comb. nov., line drawings, male allotype: A, urosome with spermatophore visible inside, ventral view; B, urosome with spermatophore visible inside, lateral view; C, maxilliped, posterior view; D, endopod of second swimming leg. Arabic numerals indicate sensilla and pores presumably homologous to those in Wellstenhelia calliope. Roman numeral (VII) indicates a pore presumably homologous to that in Itostenhelia polyhymnia. Arrowheads point to most prominent specific features. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4910660" httpUri="https://zenodo.org/record/4910660/files/figure.png" pageId="64" pageNumber="65">45A, B, D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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