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<document id="64F98FC6AAB645277D1FBD7C537AED53" ID-DOI="10.1093/zoolinnean/zlad093" ID-ISSN="0024-4082" ID-Zenodo-Dep="11240465" ID-ZooBank="3C4B3B10-EED7-405E-9A2B-88B2C1A63B07C" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1716362213817" checkinUser="plazi" docAuthor="Hájek, Jiří, Alarie, Yves, Benetti, Cesar J., Hamada, Neusa, Springer, Monika, Hendrich, Lars, Villastrigo, Adrián, Torres, Rodulfo Ospina, Basantes, Michael S. &amp; Balke, Michael" docDate="2024" docId="3D188782FFCCEC39F6433D735CBE41EE" docLanguage="en" docName="zlad093.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 200 (3)" docSource="https://www.mendeley.com/catalogue/3de85c86-0e4b-316a-bea1-fa325bfbddf3/" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.5:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="5" docTitle="Copelatus florae Hajek, Alarie, Benetti, Basantes, Hendrich &amp; Balke 2024, sp. nov." docType="treatment" docVersion="1" lastPageNumber="729" masterDocId="C121FFFAFFCAEC30F63A39655F2C4073" masterDocTitle="Underestimated diversity and range size of diving beetles in tank bromeliads-Coleoptera of ‘ hygrofloric’ lifestyle (Dytiscidae)" masterLastPageNumber="735" masterPageNumber="720" pageNumber="726" updateTime="1716577774993" updateUser="jonas">
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<mods:title id="BF1A60669BA34D77D79B8C815C7379F8">Underestimated diversity and range size of diving beetles in tank bromeliads-Coleoptera of ‘ hygrofloric’ lifestyle (Dytiscidae)</mods:title>
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<mods:namePart id="869D4469282B845A1C22508F5E43348D">Hájek, Jiří</mods:namePart>
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<mods:namePart id="3FF30CC54C71AE8407F4F82A0BD3C969">Alarie, Yves</mods:namePart>
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<mods:namePart id="A7887321105F13B30F11BDFD57C984B3">Basantes, Michael S.</mods:namePart>
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Hájek, Alarie, Benetti, Basantes, Hendrich &amp; Balke
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(
<figureCitation id="2D8A2A11FFCCEC36F7743D005E91440C" box="[334,445,1125,1151]" captionStart="Figure 4" captionStartId="7.[129,194,1772,1796]" captionTargetBox="[252,1354,147,1740]" captionTargetId="figure-68@7.[248,1354,144,1744]" captionTargetPageId="7" captionText="Figure 4. Habitus of bromeliadicolous Copelatus. A, C. florae (holotype). B, instar III larva of C. florae (Alto de Piedra).C, C. florae (Manaus). D,C. panguana (holotype). Scale bar = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11240473" httpUri="https://zenodo.org/record/11240473/files/figure.png" pageId="6" pageNumber="726">Figs 4A–C</figureCitation>
, 
<figureCitation id="2D8A2A11FFCCEC36F7FD3D005D2A440C" box="[455,518,1125,1151]" captionStart="Figure 2" captionStartId="4.[113,178,1836,1860]" captionTargetBox="[163,1408,153,1801]" captionTargetId="figure-7@4.[157,1415,144,1808]" captionTargetPageId="4" captionText="Figure 2. Male genitalia of Copelatus. A, C. bimaculatus (Santo Amaro da Imperatriz). B, C. bimaculatus (Nova Friburgo).C, C.bromeliarum (El Tucuche).D, C. bromeliarum (Henri Pittier). E, C. espinhasso (holotype).F, C. florae (Alto de Piedra). G, C. florae (Manaus). H, C.panguana (holotype). a, median lobe in lateral view. b, median lobe in ventral view.c, paramere.Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11240469" httpUri="https://zenodo.org/record/11240469/files/figure.png" pageId="6" pageNumber="726">2F–G</figureCitation>
, 
<figureCitation id="2D8A2A11FFCCEC36F42A3D005D32440C" box="[528,542,1125,1151]" captionStart="Figure 5" captionStartId="8.[113,178,1677,1701]" captionTargetBox="[147,1426,145,1648]" captionTargetId="figure-7@8.[146,1426,144,1649]" captionTargetPageId="8" captionText="Figure 5. Details of instar III larva of C. florae.A, head capsule, dorsal aspect (scale-like microsculptures not represented). B, maxilla, dorsal aspect. C, labium, dorsal aspect. D, metathoracic leg, anterior surface. E, same, posterior surface. F, last abdominal segment, dorsal aspect. G, same, ventral aspect. Numbers and lowercase letters refer to primary setae and pores, respectively; asterisks refer to secondary seta (including additional setae) or pores. AB, abdominal segment VIII (= LAS). CO, coxa.FE, femur.GA, galea. LA, labium. MX, maxilla.PT, pretarsus.TA, tarsus. TI, tibia. TR, trochanter. UR, urogomphus.sp, spinulae.Pores TAc, TAd, Tae, and TAf not represented. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11240475" httpUri="https://zenodo.org/record/11240475/files/figure.png" pageId="6" pageNumber="726">5</figureCitation>
)
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<emphasis id="87C5EA86FFCCEC36F64B3DC05E7444CE" box="[113,344,1189,1213]" italics="true" pageId="6" pageNumber="726">Zoobank registration:</emphasis>
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urn:lsid:zoobank.org:act: 
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(
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), labelled: ‘ 
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: 
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<collectingMunicipality id="556AACEEFFCCEC36F7993C785D2A4546" box="[419,518,1309,1333]" pageId="6" pageNumber="726">Santa Fé</collectingMunicipality>
env., below 
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/ 08°31.451ʹN, 81°7.650ʹW, 
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/ pasture with solitary trees; in bromeliads; Fikáček, Hájek, Seidel &amp; Sekerka lgt. [p]// HOLOTYPE/ 
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/ 
<collectorName id="18445342FFCCEC36F4D73C195F8245C0" pageId="6" pageNumber="726">J. Hájek</collectorName>
<emphasis id="87C5EA86FFCCEC36F6813CFE5FDE45C0" box="[187,242,1435,1459]" italics="true" pageId="6" pageNumber="726">et al.</emphasis>
det. 2022 [red label, p]’.
</materialsCitation>
<typeStatus id="6A0A8836FFCCEC36F42D3CFE5D5345C0" box="[535,639,1435,1459]" pageId="6" pageNumber="726" type="paratype">Paratypes</typeStatus>
: 
<collectingCountry id="CDA67604FFCCEC36F4B43CFE5DD645C0" box="[654,762,1435,1459]" name="Panama" pageId="6" pageNumber="726">PANAMA</collectingCountry>
<specimenCount id="A3B7FD1DFFCCEC36F64B3CDF5F9F45A1" box="[113,179,1463,1490]" count="7" pageId="6" pageNumber="726" type="male">7♂♂</specimenCount>
, 
<specimenCount id="A3B7FD1DFFCCEC36F6FB3CDF5E2F45A1" box="[193,259,1463,1490]" count="6" pageId="6" pageNumber="726" type="female">6♀♀</specimenCount>
, same label data as 
<typeStatus id="6A0A8836FFCCEC36F7ED3CDF5D1F45A1" box="[471,563,1466,1490]" pageId="6" pageNumber="726" type="holotype">holotype</typeStatus>
(NMPC, ZSMG); 
<specimenCount id="A3B7FD1DFFCCEC36F64B3CBC5F9F4582" box="[113,179,1494,1521]" count="3" pageId="6" pageNumber="726" type="female">3♀♀</specimenCount>
, labelled: ‘ 
<collectingCountry id="CDA67604FFCCEC36F7113CBC5EB24582" box="[299,414,1497,1521]" name="Panama" pageId="6" pageNumber="726">PANAMA</collectingCountry>
: 
<collectingRegion id="7775F876FFCCEC36F7963CBC5D2A4582" box="[428,518,1497,1521]" country="Panama" name="Panama" pageId="6" pageNumber="726">PANAMá</collectingRegion>
<date id="C10F1054FFCCEC36F4283CBC5DAE4582" box="[530,642,1497,1521]" pageId="6" pageNumber="726" value="2017-09-13">13.ix.2017</date>
/ PN Altos de Campana: Cerro Campana/ 08°41.1ʹN, 79°56.0ʹW, 
<quantity id="72499B71FFCCEC36F4FC3C9C5F944643" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="6" pageNumber="726" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800– 900 m</quantity>
/ lower montane forest; in bromeliads/ Fikáček, Hájek, Seidel &amp; Sekerka lgt. [p]’ (NMPC, ZSMG). 
<collectingCountry id="CDA67604FFCCEC36F4783F525DF5463C" box="[578,729,1591,1615]" name="Costa Rica" pageId="6" pageNumber="726">COSTA RICA</collectingCountry>
17 exs, labelled: ‘Costa Rica: 
<collectingRegion id="7775F876FFCCEC36F7BD3F325ECC461C" box="[391,480,1623,1647]" country="Costa Rica" name="Alajuela" pageId="6" pageNumber="726">Alajuela</collectingRegion>
,/ NNW San Ramon,/ 
<quantity id="72499B71FFCCEC36F4E93F325FAB46FE" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="6" pageNumber="726" unit="m" value="810.0">810 m</quantity>
, 
<date id="C10F1054FFCCEC36F6AF3F135E2846FD" box="[149,260,1654,1678]" pageId="6" pageNumber="726" value="2022-03-13">13.iii.2022</date>
, [p]// 10.2253° -84.5553°,/ Springer &amp; Balke/ (CR_MB_2022_06) [p]’, 
<specimenCount id="A3B7FD1DFFCCEC36F7AB3FF35D2D46DE" box="[401,513,1685,1709]" count="2" pageId="6" pageNumber="726" type="male">two males</specimenCount>
with additional labels ‘MB 8902 [p, green typing]’ and ‘MB8903 [p, green typing]’, respectively (UCR, ZSMG). 
<collectingCountry id="CDA67604FFCCEC36F7953FB15D25469F" box="[431,521,1748,1772]" name="Brazil" pageId="6" pageNumber="726">BRAZIL</collectingCountry>
<specimenCount id="A3B7FD1DFFCCEC36F42C3FB15D74469F" box="[534,600,1745,1772]" count="2" pageId="6" pageNumber="726" type="male">2♂♂</specimenCount>
, 
<specimenCount id="A3B7FD1DFFCCEC36F4533FB15DBC469F" box="[617,656,1745,1772]" count="1" pageId="6" pageNumber="726" type="female">1♀</specimenCount>
, labelled: ‘SNSB-Zoologische, Staatssammlung München, Digital imaging, ZSM-COL-00182’/ ‘Brazil: Amazonas, Manaus; Reserva Ducke, proximo ao Igarapé Acará, 
<geoCoordinate id="D0855053FFCCEC36F7D13E575D4E4739" box="[491,610,1842,1866]" degrees="02" direction="south" minutes="56" orientation="latitude" pageId="6" pageNumber="726" precision="15" seconds="06" value="-2.9350002">02º56’06”S</geoCoordinate>
/ 
<geoCoordinate id="D0855053FFCCEC36F44C3E575DDA4739" box="[630,758,1842,1866]" degrees="59" direction="west" minutes="57" orientation="longitude" pageId="6" pageNumber="726" precision="15" seconds="12" value="-59.953335">59º57’12”W</geoCoordinate>
, 
<date id="C10F1054FFCCEC36F64B3E345FF1471A" box="[113,221,1873,1897]" pageId="6" pageNumber="726" value="2017-04-17">17.iv.2017</date>
, 
<quantity id="72499B71FFCCEC36F6D13E345E0A471A" box="[235,294,1873,1897]" metricMagnitude="1" metricUnit="m" metricValue="7.2" pageId="6" pageNumber="726" unit="m" value="72.0">72 m</quantity>
, G. Jorge’, 
<specimenCount id="A3B7FD1DFFCCEC36F7A43E375D27471A" box="[414,523,1873,1897]" count="2" pageId="6" pageNumber="726" type="male">two males</specimenCount>
with additional labels ‘MB 8946 [p, green typing]’ and ‘MB8948 [p, green typing]’ (INPA, ZSMG); 2 exs, labelled; ‘Brazil: Amazonas, Manaus, Reserva da Campina, BR 174, 
<geoCoordinate id="D0855053FFCCEC36F25539F55BC840DB" box="[1135,1252,144,168]" degrees="02" direction="south" minutes="35" orientation="latitude" pageId="6" pageNumber="726" precision="15" seconds="52" value="-2.5977776">02º35’52”S</geoCoordinate>
, 
<geoCoordinate id="D0855053FFCCEC36F2CB39F55A5D40DB" box="[1265,1393,144,168]" degrees="60" direction="west" minutes="02" orientation="longitude" pageId="6" pageNumber="726" precision="15" seconds="05" value="-60.03472">60º02’05”W</geoCoordinate>
, Jorge G., 
<date id="C10F1054FFCCEC36F56E39CA5C9A40B4" box="[852,950,175,199]" pageId="6" pageNumber="726" value="2022-01-06">06.i.2022</date>
, in bromelia’ (INPA, ZSMG). 
<collectingCountry id="CDA67604FFCCEC36F33539CA5AB540B4" box="[1295,1433,175,199]" name="Colombia" pageId="6" pageNumber="726">COLOMBIA</collectingCountry>
4 exs: ‘Colombia: Boya., Sta Maria, 
<quantity id="72499B71FFCCEC36F2AB39AA5BC84095" box="[1169,1252,207,231]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="726" unit="m" value="1200.0">1200 m</quantity>
, 
<date id="C10F1054FFCCEC36F2CA39AA5A7B4094" box="[1264,1367,207,231]" pageId="6" pageNumber="726" value="2022-10-29">29.x.2022</date>
, 4,8807° -73,2533°, Balke 
<emphasis id="87C5EA86FFCCEC36F5E9398A5B2D4175" box="[979,1025,238,262]" italics="true" pageId="6" pageNumber="726">et al.</emphasis>
(COL_MB_2022_10)’; 2 exs: ‘Colombia: Boya., Sta Maria, 
<quantity id="72499B71FFCCEC36F5D238685B024156" box="[1000,1070,269,293]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="6" pageNumber="726" unit="m" value="880.0">880 m</quantity>
, 
<date id="C10F1054FFCCEC36F20638685B8F4156" box="[1084,1187,269,293]" pageId="6" pageNumber="726" value="2022-10-29">29.x.2022</date>
, 4,8818° -73,2434°, Balke 
<emphasis id="87C5EA86FFCCEC36F51038485C734137" box="[810,863,300,324]" italics="true" pageId="6" pageNumber="726">et al.</emphasis>
(COL_MB_2022_11)’; 19 exs: ‘Colombia: Boya., Sta Maria, 
<quantity id="72499B71FFCCEC36F54838295C994110" box="[882,949,332,356]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="6" pageNumber="726" unit="m" value="740.0">740 m</quantity>
, 
<date id="C10F1054FFCCEC36F58538295B0A4117" box="[959,1062,332,356]" pageId="6" pageNumber="726" value="2022-10-30">30.x.2022</date>
, 4,8271° -73,2582°, Balke 
<emphasis id="87C5EA86FFCCEC36F30A38295A734117" box="[1328,1375,332,356]" italics="true" pageId="6" pageNumber="726">et al.</emphasis>
(COL_ MB_2022_12)’ (UNAL, ZSMG). 
<collectingCountry id="CDA67604FFCCEC36F2B4380E5A2741F0" box="[1166,1291,363,387]" name="Ecuador" pageId="6" pageNumber="726">ECUADOR</collectingCountry>
2 exs: ‘Ecuador: 
<collectingRegion id="7775F876FFCCEC36F51038EE5C4841D1" box="[810,868,395,418]" country="Ecuador" name="Napo" pageId="6" pageNumber="726">Napo</collectingRegion>
, Road side to Archidona, Bromelia, 
<quantity id="72499B71FFCCEC36F2DD38EE5A0141D1" box="[1255,1325,395,419]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="6" pageNumber="726" unit="m" value="770.0">770 m</quantity>
, 
<date id="C10F1054FFCCEC36F30038EF5A8341D1" box="[1338,1455,394,418]" pageId="6" pageNumber="726" value="2022-12-13">13.xii.2022</date>
, -0,840° -77,7792°, Basantes (ECU_MB_2022_13)’ (INABIO, ZSMG); 2 exs: ‘Ecuador: 
<collectingRegion id="7775F876FFCCEC36F27E38AF5B524192" box="[1092,1150,458,481]" country="Ecuador" name="Napo" pageId="6" pageNumber="726">Napo</collectingRegion>
, Tena, Amazon Park La Isla, Bromelia, 502, 
<date id="C10F1054FFCCEC36F5FD388D5B104273" box="[967,1084,488,512]" pageId="6" pageNumber="726" value="2022-12-15">15.xii.2022</date>
, -0,994° -77,8174°, Basantes (ECU_ MB_2022_15)’ (INABIO, ZSMG). Each 
<typeStatus id="6A0A8836FFCCEC36F2C83B6D5A674253" box="[1266,1355,520,544]" pageId="6" pageNumber="726" type="paratype">paratype</typeStatus>
provided with the respective red printed label.
</paragraph>
</subSubSection>
<subSubSection id="FDAB651FFFCCEC39F5103B045EE545E8" lastPageId="9" lastPageNumber="729" pageId="6" pageNumber="726" type="description">
<paragraph id="B50E3694FFCCEC36F5103B045B4142C4" blockId="6.[810,1459,608,695]" pageId="6" pageNumber="726">
<emphasis id="87C5EA86FFCCEC36F5103B045B79420B" box="[810,1109,608,632]" italics="true" pageId="6" pageNumber="726">
Description of male 
<typeStatus id="6A0A8836FFCCEC36F5C13B055B7C420B" box="[1019,1104,608,632]" pageId="6" pageNumber="726" type="holotype">holotype</typeStatus>
:
</emphasis>
Habitus (
<figureCitation id="2D8A2A11FFCCEC36F2FD3B055A3A420B" box="[1223,1302,608,632]" captionStart="Figure 4" captionStartId="7.[129,194,1772,1796]" captionTargetBox="[252,1354,147,1740]" captionTargetId="figure-68@7.[248,1354,144,1744]" captionTargetPageId="7" captionText="Figure 4. Habitus of bromeliadicolous Copelatus. A, C. florae (holotype). B, instar III larva of C. florae (Alto de Piedra).C, C. florae (Manaus). D,C. panguana (holotype). Scale bar = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11240473" httpUri="https://zenodo.org/record/11240473/files/figure.png" pageId="6" pageNumber="726">Fig. 4A</figureCitation>
) drop-shaped, strongly attenuated posteriorly, broadest at base of elytra, slightly convex. Dorsal surface submatt.
</paragraph>
<paragraph id="B50E3694FFCCEC36F5103BBD5B58433D" blockId="6.[810,1459,728,846]" pageId="6" pageNumber="726">
<emphasis id="87C5EA86FFCCEC36F5103BBD5CB64283" box="[810,922,728,752]" italics="true" pageId="6" pageNumber="726">Coloration:</emphasis>
Body black; sides of pronotum and abdomen somewhat reddish translucent; ventral part of head, prosternum, and appendages testaceous; elytra with oblong transverse orange spot on base between striae 1–5.
</paragraph>
<paragraph id="B50E3694FFCCEC36F5103A155A4444D1" blockId="6.[810,1459,879,1186]" pageId="6" pageNumber="726">
<emphasis id="87C5EA86FFCCEC36F5103A155C4B43FB" box="[810,871,880,904]" italics="true" pageId="6" pageNumber="726">Head:</emphasis>
Small, 
<emphasis id="87C5EA86FFCCEC36F58B3A155C9643F4" box="[945,954,880,903]" italics="true" pageId="6" pageNumber="726">c</emphasis>
. 0.54 × width of pronotum, trapezoidal. Anterior margin of clypeus slightly concave. Antennae with antennomeres rather broad (antennomeres II–X 
<emphasis id="87C5EA86FFCCEC36F2A93ACA5BB043B5" box="[1171,1180,943,966]" italics="true" pageId="6" pageNumber="726">c</emphasis>
. 1.5 × longer than width), club-shaped. Reticulation consisting of fine, well impressed isodiametric polygonal meshes. Numerous short, shallow, and isolated strioles present between eyes. Punctation double; several large setigerous punctures present in fronto-clypeal depressions, frontal depressions at level of anterior margin of eyes, and in depressions along inner margin of eyes; very fine and sparsely distributed punctures placed among meshes of reticulation.
</paragraph>
<paragraph id="B50E3694FFCCEC36F5103DA15B884667" blockId="6.[810,1460,1219,1557]" pageId="6" pageNumber="726">
<emphasis id="87C5EA86FFCCEC36F5103DA15CBF44A8" box="[810,915,1220,1243]" italics="true" pageId="6" pageNumber="726">Pronotum:</emphasis>
Transverse, broadest at posterior angles. Anterior angles acute, posterior angles rectangular. Sides strongly and evenly curved, lateral beading distinct except for anterior corners. Anterior margin straight, posterior margin nearly straight with only indistinct sinuation medially. Reticulation similar to that of head, meshes longitudinally elongate. Surface of pronotum with numerous long shallow strioles; centre of disc with shallow longitudinal scratch. Punctation double; row of coarse setigerous punctures present along anterior margin, laterally close to sides, and in basolateral depressions along basal margin; fine punctures placed among meshes of reticulation.
</paragraph>
<paragraph id="B50E3694FFCCEC36F5103F505BBC47D5" blockId="6.[810,1459,1589,1959]" pageId="6" pageNumber="726">
<emphasis id="87C5EA86FFCCEC36F5103F505C41463E" box="[810,877,1589,1613]" italics="true" pageId="6" pageNumber="726">Elytra:</emphasis>
Elytral striation consisting of six superficially impressed discal striae and a submarginal stria: all discal striae complete, except for stria 5 which is absent at base; submarginal stria long, starting at 1/3 of elytral length. Whole surface of elytra covered with fine, shallow long longitudinal striolae; striolae somewhat sparser and shorter and transverse in apical part of elytra. Surface reticulation consisting of very fine, superficially impressed polygonal meshes; meshes longitudinally elongate in basal half, but isodiametric or transverse in apical half of elytra. Punctation double; large setigerous punctures present along lateral margin of elytra; badly perceptible, very fine punctures sparsely distributed among meshes of reticulation.
</paragraph>
<caption id="E1CE661CFFCDEC37F6BB3F895D69476C" ID-DOI="http://doi.org/10.5281/zenodo.11240473" ID-Zenodo-Dep="11240473" httpUri="https://zenodo.org/record/11240473/files/figure.png" pageId="7" pageNumber="727" startId="7.[129,194,1772,1796]" targetBox="[252,1354,147,1740]" targetPageId="7" targetType="figure">
<paragraph id="B50E3694FFCDEC37F6BB3F895D69476C" blockId="7.[129,1455,1772,1824]" pageId="7" pageNumber="727">
<emphasis id="87C5EA86FFCDEC37F6BB3F895FF44777" bold="true" box="[129,216,1772,1796]" pageId="7" pageNumber="727">Figure 4.</emphasis>
Habitus of bromeliadicolous 
<taxonomicName id="72B14D17FFCDEC37F7D53F895D654777" authorityName="Erichson" authorityYear="1832" box="[495,585,1772,1796]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="727" phylum="Arthropoda" rank="genus">
<emphasis id="87C5EA86FFCDEC37F7D53F895D654777" box="[495,585,1772,1796]" italics="true" pageId="7" pageNumber="727">Copelatus</emphasis>
</taxonomicName>
. A, 
<taxonomicName id="72B14D17FFCDEC37F4573F895D944777" authorityName="Hájek &amp; Alarie &amp; Benetti &amp; Hamada &amp; Springer &amp; Hendrich &amp; Villastrigo &amp; Torres &amp; Basantes &amp; Balke" authorityYear="2024" box="[621,696,1772,1796]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="727" phylum="Arthropoda" rank="species" species="florae">
<emphasis id="87C5EA86FFCDEC37F4573F895D944777" box="[621,696,1772,1796]" italics="true" pageId="7" pageNumber="727">C. florae</emphasis>
</taxonomicName>
(holotype). B, instar III larva of 
<taxonomicName id="72B14D17FFCDEC37F5D13F895BF44777" authority="(Alto de Piedra)" baseAuthorityName="Alto de Piedra" box="[1003,1240,1772,1796]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="727" phylum="Arthropoda" rank="species" species="florae">
<emphasis id="87C5EA86FFCDEC37F5D13F895B1A4777" box="[1003,1078,1772,1796]" italics="true" pageId="7" pageNumber="727">C. florae</emphasis>
(Alto de Piedra)
</taxonomicName>
. C, 
<taxonomicName id="72B14D17FFCDEC37F2C03F895A874777" authority="(Manaus)" baseAuthorityName="Manaus" box="[1274,1451,1772,1796]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="727" phylum="Arthropoda" rank="species" species="florae">
<emphasis id="87C5EA86FFCDEC37F2C03F895A6A4777" box="[1274,1350,1772,1796]" italics="true" pageId="7" pageNumber="727">C. florae</emphasis>
(Manaus)
</taxonomicName>
. D, 
<taxonomicName id="72B14D17FFCDEC37F6A13E6D5E234753" authorityName="Hájek &amp; Alarie &amp; Benetti &amp; Hamada &amp; Springer &amp; Hendrich &amp; Villastrigo &amp; Torres &amp; Basantes &amp; Balke" authorityYear="2024" box="[155,271,1800,1824]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="727" phylum="Arthropoda" rank="species" species="panguana">
<emphasis id="87C5EA86FFCDEC37F6A13E6D5E234753" box="[155,271,1800,1824]" italics="true" pageId="7" pageNumber="727">C. panguana</emphasis>
</taxonomicName>
(holotype). Scale bar = 2.0 mm.
</paragraph>
</caption>
<paragraph id="B50E3694FFCDEC37F6BB3E5B5DBD47C1" blockId="7.[129,778,1853,1971]" pageId="7" pageNumber="727">
<emphasis id="87C5EA86FFCDEC37F6BB3E5B5F9F4726" box="[129,179,1854,1877]" italics="true" pageId="7" pageNumber="727">Legs:</emphasis>
Protibia modified, slightly angled near base, distinctly broadened distally, club shaped. Pro- and mesotarsomeres 1–3 distinctly broadened, ventrally with adhesive setae. Natatorial setae present only on dorsal side of metatarsomeres.
</paragraph>
<paragraph id="B50E3694FFCDEC39F5033E585EB64110" blockId="7.[825,1474,1853,1971]" lastBlockId="9.[129,778,144,356]" lastPageId="9" lastPageNumber="729" pageId="7" pageNumber="727">
<emphasis id="87C5EA86FFCDEC37F5033E585C964726" box="[825,954,1853,1877]" italics="true" pageId="7" pageNumber="727">Ventral side:</emphasis>
Prosternum sinuate anteriorly, obtusely keeled medially. Prosternal process shortly lanceolate, in cross-section convex, apex obtuse; process distinctly bordered; reticulation consisting of well impressed polygonal meshes; punctation consisting of setigerous punctures along margin. Metaventrite with microsculpture consisting of polygonal meshes, meshes distinctly transverse laterally; lateral parts of metaventrite (ʻmetasternal wingsʼ) tongue-shaped, slender. Metacoxal lines well impressed, abbreviated (absent in basal fourth). Metacoxal plates covered with long longitudinal strioles; reticulation consisting of extremely elongated, longitudinal polygonal meshes. Metacoxal processes rounded and incised at posterior margin. Abdominal ventrites I–II with longitudinal strioles; ventrites III–IV with oblique strioles laterally. Abdominal reticulation consisting of elongate polygonal meshes, longitudinal on ventrites I–II, oblique on ventrite III and transverse on ventrites IV–VI. Punctation double; coarse setigerous punctures present medially and medio-laterally on ventrites III–VI; fine punctures sparsely distributed on surface of ventrites, clusters of somewhat coarser punctures present laterally on ventrite V and basolaterally on ventrite VI.
</paragraph>
<caption id="E1CE661CFFC2EC38F64B3FE85A644767" ID-DOI="http://doi.org/10.5281/zenodo.11240475" ID-Zenodo-Dep="11240475" httpUri="https://zenodo.org/record/11240475/files/figure.png" pageId="8" pageNumber="728" startId="8.[113,178,1677,1701]" targetBox="[147,1426,145,1648]" targetPageId="8" targetType="figure">
<paragraph id="B50E3694FFC2EC38F64B3FE85A644767" blockId="8.[113,1449,1677,1813]" pageId="8" pageNumber="728">
<emphasis id="87C5EA86FFC2EC38F64B3FE85FE546D6" bold="true" box="[113,201,1677,1701]" pageId="8" pageNumber="728">Figure 5.</emphasis>
Details of instar III larva of 
<emphasis id="87C5EA86FFC2EC38F7F53FE85D0D46D6" box="[463,545,1677,1701]" italics="true" pageId="8" pageNumber="728">
<taxonomicName id="72B14D17FFC2EC38F7F53FE85D3146D6" box="[463,541,1677,1701]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="728" phylum="Arthropoda" rank="species" species="florae">C. florae</taxonomicName>
.
</emphasis>
A, head capsule, dorsal aspect (scale-like microsculptures not represented). B, maxilla, dorsal aspect. C, labium, dorsal aspect. D, metathoracic leg, anterior surface. E, same, posterior surface. F, last abdominal segment, dorsal aspect. G, same, ventral aspect. Numbers and lowercase letters refer to primary setae and pores, respectively; asterisks refer to secondary seta (including additional setae) or pores. AB, abdominal segment VIII (= LAS). CO, coxa. FE, femur. GA, galea. LA, labium. MX, maxilla. PT, pretarsus. TA, tarsus. TI, tibia. TR, trochanter. UR, urogomphus. sp, spinulae. Pores TAc, TAd, Tae, and TAf not represented. Scale bars = 0.2 mm.
</paragraph>
</caption>
<paragraph id="B50E3694FFC3EC39F6BB38E05FD44188" blockId="9.[129,778,388,507]" pageId="9" pageNumber="729">
<emphasis id="87C5EA86FFC3EC39F6BB38E05E3A41EE" box="[129,278,389,413]" italics="true" pageId="9" pageNumber="729">Male genitalia:</emphasis>
Median lobe in lateral aspect ʻCʼ- shaped, apex pointed; in ventral aspect median lobe attenuated in apical fourth, apex twisted left (as in 
<figureCitation id="2D8A2A11FFC3EC39F7C938A15D1141AF" box="[499,573,452,476]" captionStart="Figure 2" captionStartId="4.[113,178,1836,1860]" captionTargetBox="[163,1408,153,1801]" captionTargetId="figure-7@4.[157,1415,144,1808]" captionTargetPageId="4" captionText="Figure 2. Male genitalia of Copelatus. A, C. bimaculatus (Santo Amaro da Imperatriz). B, C. bimaculatus (Nova Friburgo).C, C.bromeliarum (El Tucuche).D, C. bromeliarum (Henri Pittier). E, C. espinhasso (holotype).F, C. florae (Alto de Piedra). G, C. florae (Manaus). H, C.panguana (holotype). a, median lobe in lateral view. b, median lobe in ventral view.c, paramere.Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11240469" httpUri="https://zenodo.org/record/11240469/files/figure.png" pageId="9" pageNumber="729">Fig. 2F</figureCitation>
). Parameres broad, ʻDʼ- shaped.
</paragraph>
<paragraph id="B50E3694FFC3EC39F6BB3B785EAD42C1" blockId="9.[129,777,541,690]" pageId="9" pageNumber="729">
<emphasis id="87C5EA86FFC3EC39F6BB3B785FE14246" box="[129,205,541,565]" italics="true" pageId="9" pageNumber="729">Female:</emphasis>
Females do not differ in external morphology from males except for nearly straight, apically less broadened protibia, slender pro- and mesotarsi without adhesive setae, and sparser and shorter striolation of pronotum and elytra. Females are generally smaller than males.
</paragraph>
<paragraph id="B50E3694FFC3EC39F6BB3BB65E7F43B4" blockId="9.[129,777,723,967]" pageId="9" pageNumber="729">
<emphasis id="87C5EA86FFC3EC39F6BB3BB65FDF4298" box="[129,243,723,747]" italics="true" pageId="9" pageNumber="729">Variability:</emphasis>
This species displays allometric variability—larger specimens are more attenuated posteriorly (more drop-shaped), whereas the smaller specimens are more regularly oval (cf. 
<figureCitation id="2D8A2A11FFC3EC39F6ED3A545E13433A" box="[215,319,817,841]" captionStart="Figure 4" captionStartId="7.[129,194,1772,1796]" captionTargetBox="[252,1354,147,1740]" captionTargetId="figure-68@7.[248,1354,144,1744]" captionTargetPageId="7" captionText="Figure 4. Habitus of bromeliadicolous Copelatus. A, C. florae (holotype). B, instar III larva of C. florae (Alto de Piedra).C, C. florae (Manaus). D,C. panguana (holotype). Scale bar = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11240473" httpUri="https://zenodo.org/record/11240473/files/figure.png" pageId="9" pageNumber="729">Fig. 4A, C</figureCitation>
). In addition, the specimens vary in size and shape of orange spot at the base of elytra, and in the density of strioles on pronotum and elytra. The median lobe of aedeagus could appear more of less stout even within the same population (cf. 
<figureCitation id="2D8A2A11FFC3EC39F6E33ACB5E6E43B4" box="[217,322,942,967]" captionStart="Figure 2" captionStartId="4.[113,178,1836,1860]" captionTargetBox="[163,1408,153,1801]" captionTargetId="figure-7@4.[157,1415,144,1808]" captionTargetPageId="4" captionText="Figure 2. Male genitalia of Copelatus. A, C. bimaculatus (Santo Amaro da Imperatriz). B, C. bimaculatus (Nova Friburgo).C, C.bromeliarum (El Tucuche).D, C. bromeliarum (Henri Pittier). E, C. espinhasso (holotype).F, C. florae (Alto de Piedra). G, C. florae (Manaus). H, C.panguana (holotype). a, median lobe in lateral view. b, median lobe in ventral view.c, paramere.Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11240469" httpUri="https://zenodo.org/record/11240469/files/figure.png" pageId="9" pageNumber="729">Fig. 2F–G</figureCitation>
).
</paragraph>
<paragraph id="B50E3694FFC3EC39F6BB3A8C5D84442E" blockId="9.[129,778,997,1118]" pageId="9" pageNumber="729">
<emphasis id="87C5EA86FFC3EC39F6BB3A8C5E7B4473" box="[129,343,1000,1024]" italics="true" pageId="9" pageNumber="729">
Measurements (N = 
<specimenCount id="A3B7FD1DFFC3EC39F7703A8D5E7B4473" box="[330,343,1000,1024]" count="8" pageId="9" pageNumber="729" type="male">8</specimenCount>
</emphasis>
♂♂ 
<specimenCount id="A3B7FD1DFFC3EC39F7AA3A8D5EFE4473" box="[400,466,997,1024]" count="9" pageId="9" pageNumber="729" type="female">
<emphasis id="87C5EA86FFC3EC39F7AA3A8D5EB14473" box="[400,413,1000,1024]" italics="true" pageId="9" pageNumber="729">9</emphasis>
♀♀
</specimenCount>
<emphasis id="87C5EA86FFC3EC39F7E83A8D5EC8438C" box="[466,484,1000,1023]" italics="true" pageId="9" pageNumber="729">):</emphasis>
TL 
<specimenCount id="A3B7FD1DFFC3EC39F4333A805D124473" box="[521,574,997,1024]" count="1" pageId="9" pageNumber="729" type="male">♂♂</specimenCount>
: 
<quantity id="72499B71FFC3EC39F4733A8D5DEF438C" box="[585,707,1000,1024]" metricMagnitude="-3" metricUnit="m" metricValue="6.35" metricValueMax="6.5" metricValueMin="6.2" pageId="9" pageNumber="729" unit="mm" value="6.35" valueMax="6.5" valueMin="6.2">6.2–6.5 mm</quantity>
(
<typeStatus id="6A0A8836FFC3EC39F4F53A8D5F9F4453" pageId="9" pageNumber="729" type="holotype">holotype</typeStatus>
: 
<quantity id="72499B71FFC3EC39F6853D625E39446C" box="[191,277,1031,1055]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="729" unit="mm" value="6.5">6.5 mm</quantity>
); TL-h 
<specimenCount id="A3B7FD1DFFC3EC39F7583D615EBB446C" box="[354,407,1028,1055]" count="1" pageId="9" pageNumber="729" type="male">♂♂</specimenCount>
: 
<quantity id="72499B71FFC3EC39F79D3D625D09446C" box="[423,549,1031,1055]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" metricValueMax="5.9" metricValueMin="5.6" pageId="9" pageNumber="729" unit="mm" value="5.75" valueMax="5.9" valueMin="5.6">5.6–5.9 mm</quantity>
(
<typeStatus id="6A0A8836FFC3EC39F40C3D625DB6446C" box="[566,666,1031,1055]" pageId="9" pageNumber="729" type="holotype">holotype</typeStatus>
: 
<quantity id="72499B71FFC3EC39F49D3D625DD1446C" box="[679,765,1031,1055]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="9" pageNumber="729" unit="mm" value="5.9">5.9 mm</quantity>
); MW 
<specimenCount id="A3B7FD1DFFC3EC39F68A3D465FC9444D" box="[176,229,1059,1086]" count="1" pageId="9" pageNumber="729" type="male">♂♂</specimenCount>
: 
<quantity id="72499B71FFC3EC39F6CE3D425E5D444D" box="[244,369,1063,1087]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.4" metricValueMin="3.1" pageId="9" pageNumber="729" unit="mm" value="3.25" valueMax="3.4" valueMin="3.1">3.1–3.4 mm</quantity>
(
<typeStatus id="6A0A8836FFC3EC39F7BB3D435EC9444D" box="[385,485,1062,1086]" pageId="9" pageNumber="729" type="holotype">holotype</typeStatus>
: 
<quantity id="72499B71FFC3EC39F7CB3D425D69444C" box="[497,581,1063,1087]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="9" pageNumber="729" unit="mm" value="3.4">3.4 mm</quantity>
). TL 
<specimenCount id="A3B7FD1DFFC3EC39F4433D465D82444D" box="[633,686,1059,1086]" count="1" pageId="9" pageNumber="729" type="female">♀♀</specimenCount>
: 
<quantity id="72499B71FFC3EC39F4863D435F81442E" metricMagnitude="-3" metricUnit="m" metricValue="5.85" metricValueMax="6.1" metricValueMin="5.6" pageId="9" pageNumber="729" unit="mm" value="5.85" valueMax="6.1" valueMin="5.6">5.6–6.1 mm</quantity>
; TL-h 
<specimenCount id="A3B7FD1DFFC3EC39F6D73D265E0E442D" box="[237,290,1091,1118]" count="1" pageId="9" pageNumber="729" type="female">♀♀</specimenCount>
: 5.0– 
<quantity id="72499B71FFC3EC39F7663D235E81442E" box="[348,429,1094,1118]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="729" unit="mm" value="5.5">5.5 mm</quantity>
; MW 
<specimenCount id="A3B7FD1DFFC3EC39F7DC3D265D37442D" box="[486,539,1091,1118]" count="1" pageId="9" pageNumber="729" type="female">♀♀</specimenCount>
: 
<quantity id="72499B71FFC3EC39F41D3D235D89442E" box="[551,677,1094,1118]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" pageId="9" pageNumber="729" unit="mm" value="2.95" valueMax="3.1" valueMin="2.8">2.8–3.1 mm</quantity>
.
</paragraph>
<paragraph id="B50E3694FFC3EC39F6BB3DED5EE545E8" blockId="9.[129,778,1160,1435]" pageId="9" pageNumber="729">
<emphasis id="87C5EA86FFC3EC39F6BB3DED5E2444D3" box="[129,264,1160,1184]" italics="true" pageId="9" pageNumber="729">Identification:</emphasis>
Compared with the other bromeliadicolous 
<taxonomicName id="72B14D17FFC3EC39F6BB3DC25FCE44CC" authorityName="Erichson" authorityYear="1832" box="[129,226,1191,1215]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="729" phylum="Arthropoda" rank="genus">
<emphasis id="87C5EA86FFC3EC39F6BB3DC25FCE44CC" box="[129,226,1191,1215]" italics="true" pageId="9" pageNumber="729">Copelatus</emphasis>
</taxonomicName>
, this new species can be easily recognized based on strongly drop-shaped habitus, elytra with transverse basal orange spot, well impressed six discal and a submarginal stria on each elytron, pronotum and elytra densely covered with fine longitudinal striolae (
<figureCitation id="2D8A2A11FFC3EC39F77B3C415E82454E" box="[321,430,1316,1341]" captionStart="Figure 4" captionStartId="7.[129,194,1772,1796]" captionTargetBox="[252,1354,147,1740]" captionTargetId="figure-68@7.[248,1354,144,1744]" captionTargetPageId="7" captionText="Figure 4. Habitus of bromeliadicolous Copelatus. A, C. florae (holotype). B, instar III larva of C. florae (Alto de Piedra).C, C. florae (Manaus). D,C. panguana (holotype). Scale bar = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11240473" httpUri="https://zenodo.org/record/11240473/files/figure.png" pageId="9" pageNumber="729">Fig. 4A, C</figureCitation>
), and by the characteristic shape of the median lobe of the aedeagus—in lateral view, the median lobe is in apical half slightly attenuating to pointed, dorsally distinctly curved apex (
<figureCitation id="2D8A2A11FFC3EC39F7683CE75E9545E8" box="[338,441,1410,1435]" captionStart="Figure 2" captionStartId="4.[113,178,1836,1860]" captionTargetBox="[163,1408,153,1801]" captionTargetId="figure-7@4.[157,1415,144,1808]" captionTargetPageId="4" captionText="Figure 2. Male genitalia of Copelatus. A, C. bimaculatus (Santo Amaro da Imperatriz). B, C. bimaculatus (Nova Friburgo).C, C.bromeliarum (El Tucuche).D, C. bromeliarum (Henri Pittier). E, C. espinhasso (holotype).F, C. florae (Alto de Piedra). G, C. florae (Manaus). H, C.panguana (holotype). a, median lobe in lateral view. b, median lobe in ventral view.c, paramere.Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11240469" httpUri="https://zenodo.org/record/11240469/files/figure.png" pageId="9" pageNumber="729">Fig. 2F–G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="FDAB651FFFC3EC39F6BB3CA15FD84649" pageId="9" pageNumber="729" type="etymology">
<paragraph id="B50E3694FFC3EC39F6BB3CA15FD84649" blockId="9.[129,778,1476,1594]" pageId="9" pageNumber="729">
<emphasis id="87C5EA86FFC3EC39F6BB3CA15FDC45AF" box="[129,240,1476,1500]" italics="true" pageId="9" pageNumber="729">Etymology:</emphasis>
The new species is dedicated to the partner of the senior author ‘Květa’ which is the Slavic form of the Latin name ‘Flora’; the name is a noun in the genitive case, standing in apposition.
</paragraph>
</subSubSection>
<subSubSection id="FDAB651FFFC3EC39F6BB3F395C8340B4" pageId="9" pageNumber="729" type="biology_ecology">
<paragraph id="B50E3694FFC3EC39F6BB3F395C8340B4" blockId="9.[129,778,1628,1996]" lastBlockId="9.[825,1474,144,199]" pageId="9" pageNumber="729">
<emphasis id="87C5EA86FFC3EC39F6BB3F395E574600" box="[129,379,1628,1652]" italics="true" pageId="9" pageNumber="729">Collecting circumstances:</emphasis>
The specimens from 
<collectingCountry id="CDA67604FFC3EC39F4433F395DE64600" box="[633,714,1628,1651]" name="Panama" pageId="9" pageNumber="729">Panama</collectingCountry>
were collected at two medium altitude localities (
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a.s.l.) in the thin water layer between leaf axils of a bromeliad species tentatively identified as 
<taxonomicName id="72B14D17FFC3EC39F7973FDF5FC74682" authority="(H. Wendl.) J. R. Grant" authorityName="J. R. Grant" baseAuthorityName="H. Wendl." class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="9" pageNumber="729" phylum="Tracheophyta" rank="species" species="gladioliflora">
<emphasis id="87C5EA86FFC3EC39F7973FDF5DA546A1" box="[429,649,1722,1746]" italics="true" pageId="9" pageNumber="729">Werauhia gladioliflora</emphasis>
(H.Wendl.) J.R. Grant
</taxonomicName>
, growing on solitary trees on pasture, close to forest (Alto de Piedra, 
<figureCitation id="2D8A2A11FFC3EC39F7113F9D5E484763" box="[299,356,1784,1808]" captionStart="Figure 6" captionStartId="10.[113,178,1836,1860]" captionTargetBox="[167,1406,147,1805]" captionTargetId="figure-63@10.[164,1409,144,1808]" captionTargetPageId="10" captionText="Figure 6. Habitat of C. florae in Alto de Piedra, Panama.A, general view of the pasture with solitary trees.B, large cf.Werauhia bromeliad. C, collecting of beetles from leaf axils. D, specimen of C. florae crawling on the leaf." figureDoi="http://doi.org/10.5281/zenodo.11240477" httpUri="https://zenodo.org/record/11240477/files/figure.png" pageId="9" pageNumber="729">Fig. 6</figureCitation>
), and on trees along a forest road (Cerro Campana). The specimens from 
<collectingCountry id="CDA67604FFC3EC39F7E83E7D5D214743" box="[466,525,1816,1840]" name="Brazil" pageId="9" pageNumber="729">Brazil</collectingCountry>
were collected at 
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a.s.l. altitude, in the water layer between leaf axils of bromeliads, growing on trees on the banks of stream, inside the Ducke Forest Reserve. In 
<collectingCountry id="CDA67604FFC3EC39F6C73E135E4A47FD" box="[253,358,1910,1934]" name="Colombia" pageId="9" pageNumber="729">Colombia</collectingCountry>
, the beetles were collected from bromeliads which were tentatively identified as a 
<taxonomicName id="72B14D17FFC3EC39F4013EF05FE647BF" pageId="9" pageNumber="729">
<emphasis id="87C5EA86FFC3EC39F4013EF05DB647DE" box="[571,666,1941,1965]" italics="true" pageId="9" pageNumber="729">Werauhia</emphasis>
J.R. Grant species
</taxonomicName>
as well as others but without inflorescences that would aid identification. The latter is also true for the samples from 
<collectingCountry id="CDA67604FFC3EC39F50339CA5C8740B4" box="[825,939,175,199]" name="Costa Rica" pageId="9" pageNumber="729">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="FDAB651FFFC3EC39F503398D5CBE41EE" pageId="9" pageNumber="729" type="distribution">
<paragraph id="B50E3694FFC3EC39F503398D5CBE41EE" blockId="9.[825,1474,232,413]" pageId="9" pageNumber="729">
<emphasis id="87C5EA86FFC3EC39F503398D5C974173" box="[825,955,232,256]" italics="true" pageId="9" pageNumber="729">Distribution:</emphasis>
The species has apparently a very wide distribution, covering the area from central 
<collectingCountry id="CDA67604FFC3EC39F2D1386D5A4F4153" box="[1259,1379,264,288]" name="Costa Rica" pageId="9" pageNumber="729">Costa Rica</collectingCountry>
through 
<collectingCountry id="CDA67604FFC3EC39F50338425CA2414D" box="[825,910,295,318]" name="Panama" pageId="9" pageNumber="729">Panama</collectingCountry>
, eastern 
<collectingCountry id="CDA67604FFC3EC39F5DC38425B62414C" box="[998,1102,295,319]" name="Colombia" pageId="9" pageNumber="729">Colombia</collectingCountry>
and 
<collectingCountry id="CDA67604FFC3EC39F24538425BFB414C" box="[1151,1239,295,319]" name="Ecuador" pageId="9" pageNumber="729">Ecuador</collectingCountry>
to Manaus in the heart of Brazilian Amazonia, more than 
<quantity id="72499B71FFC3EC39F29138235A25412D" box="[1195,1289,326,350]" metricMagnitude="6" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="729" unit="km" value="2600.0">2600 km</quantity>
apart (
<figureCitation id="2D8A2A11FFC3EC39F36138235AB4412D" box="[1371,1432,326,350]" captionStart="Figure 3" captionStartId="5.[129,194,1003,1027]" captionTargetBox="[144,762,147,972]" captionTargetId="figure-671@5.[141,765,144,975]" captionTargetPageId="5" captionText="Figure 3. Distributional map of bromeliadicolous Copelatus species." figureDoi="http://doi.org/10.5281/zenodo.11240471" httpUri="https://zenodo.org/record/11240471/files/figure.png" pageId="9" pageNumber="729">Fig. 3</figureCitation>
). It inhabits places from lowlands to medium altitude (
<emphasis id="87C5EA86FFC3EC39F36A38035A75410E" box="[1360,1369,358,381]" italics="true" pageId="9" pageNumber="729">c</emphasis>
. 
<quantity id="72499B71FFC3EC39F35F38035C6241EF" metricMagnitude="2" metricUnit="m" metricValue="6.35" metricValueMax="12.0" metricValueMin="0.7" pageId="9" pageNumber="729" unit="m" value="635.0" valueMax="1200.0" valueMin="70.0">70–1200 m</quantity>
a.s.l.).
</paragraph>
</subSubSection>
</treatment>
</document>