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<document id="ABF3AB3CF74583BDE70559DF0C5126C2" ID-CLB-Dataset="31674" ID-DOI="10.11646/zootaxa.4341.3.11" ID-GBIF-Dataset="c3bf98e8-c0c1-4276-9f9f-e06a8249a7bc" ID-ISSN="1175-5326" ID-Zenodo-Dep="1040027" ID-ZooBank="7315B301-2918-4F78-BB60-433C832E9D03" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509519833341" checkinUser="plazi" docAuthor="Ramírez, José &amp; Asenjo, Angélico" docDate="2017" docId="365915086115FFCFFF05FEC2151CF8B2" docLanguage="en" docName="zootaxa.4341.3.11.pdf" docOrigin="Zootaxa 4341 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cylindroxystus janualis Ramírez &amp; Asenjo 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="438" masterDocId="CA606D706114FFCEFF92FF871441FFC6" masterDocTitle="First record and a new species of Cylindroxystus Bierig (Coleoptera, Staphylinidae, Paederinae) in Colombia" masterLastPageNumber="440" masterPageNumber="437" pageNumber="438" updateTime="1698470604384" updateUser="ExternalLinkService">
<mods:mods id="AF8E2D303D0A01B2DA3BA27B587FE3A4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1E33EA1C32054FD19FBFACD9BCBFE3A1">
<mods:title id="E96B5D8342556EEDD2E8EB998355950B">First record and a new species of Cylindroxystus Bierig (Coleoptera, Staphylinidae, Paederinae) in Colombia</mods:title>
</mods:titleInfo>
<mods:name id="B3476DFFB1574B213485CA99285F7AED" type="personal">
<mods:role id="4D260091ABEC7201E9B149FBEF9FE193">
<mods:roleTerm id="0FF3F625F8F8EB9428BC8B0AB1F4C47C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4F32417BF9AA9692368157FDEA3AB1A5">Ramírez, José</mods:namePart>
</mods:name>
<mods:name id="5C28831F9944596FCEC322A29B603B6A" type="personal">
<mods:role id="0AF40016426C53121253917D43E2B779">
<mods:roleTerm id="6F72D765EE42F74EDE1875EAB4FF9E8E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="941D11B8E7C6AC81FE55F13E4E5BDDF2">Asenjo, Angélico</mods:namePart>
</mods:name>
<mods:typeOfResource id="4175859DEC4610E709AEA201D0DFD0FB">text</mods:typeOfResource>
<mods:relatedItem id="887E9240E7DE8781CFC8EDC0B6016607" type="host">
<mods:titleInfo id="A7656195AB460DAC2F99DF08BADF279E">
<mods:title id="0B69BA489E71A3BCD9A99A8A18B77FCF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A78668F9A1C7C43BFAF42188B6B690E8">
<mods:date id="9392EF9005F07E9F0D19F2EC7B3A6C15">2017</mods:date>
<mods:detail id="EBB378B2C077CB9D110D1FCDC4D64395" type="pubDate">
<mods:number id="3476049FC9CD9A66498B932EFDDF7385">2017-11-01</mods:number>
</mods:detail>
<mods:detail id="1BE06403DF64E76922D9A0352F38E17C" type="volume">
<mods:number id="4C41DC8FBAE7E68455BB653C71AC406A">4341</mods:number>
</mods:detail>
<mods:detail id="29A942502DDBEBA77BA1356B1F0D3B7D" type="issue">
<mods:number id="BF3FE81C771846563A5F05BDAEDC6803">3</mods:number>
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<mods:start id="29CB30D754EA7330F7A6E6A7CDC1306B">437</mods:start>
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<mods:classification id="B1C8E798E46FB6BACE74743EA310E015">journal article</mods:classification>
<mods:identifier id="5879089FC76CCBED585511726393E0A7" type="CLB-Dataset">31674</mods:identifier>
<mods:identifier id="6095AB1E963C3AA29666C18D208B23DB" type="DOI">10.11646/zootaxa.4341.3.11</mods:identifier>
<mods:identifier id="0ED56DEB7148E73EEA899947321FC9BB" type="GBIF-Dataset">c3bf98e8-c0c1-4276-9f9f-e06a8249a7bc</mods:identifier>
<mods:identifier id="2E91FBE9598BEAA7D99681527B1558A2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D4D7276CF61F3968D8B38FB0CB86BF27" type="Zenodo-Dep">1040027</mods:identifier>
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<treatment id="365915086115FFCFFF05FEC2151CF8B2" ID-DOI="http://doi.org/10.5281/zenodo.6004279" ID-GBIF-Taxon="135557928" ID-Zenodo-Dep="6004279" LSID="urn:lsid:plazi:treatment:365915086115FFCFFF05FEC2151CF8B2" httpUri="http://treatment.plazi.org/id/365915086115FFCFFF05FEC2151CF8B2" lastPageNumber="438" pageId="1" pageNumber="438">
<subSubSection id="F6EAF7956115FFCFFF05FEC2154DFEB9" pageId="1" pageNumber="438" type="nomenclature">
<paragraph id="BE4FA41E6115FFCFFF05FEC21751FE99" blockId="1.[151,784,290,383]" box="[151,784,325,351]" pageId="1" pageNumber="438">
<heading id="E50713726115FFCFFF05FEC21751FE99" bold="true" box="[151,784,325,351]" fontSize="11" level="1" pageId="1" pageNumber="438" reason="1">
<emphasis id="8C84780C6115FFCFFF05FEC21751FE99" bold="true" box="[151,784,325,351]" pageId="1" pageNumber="438">
<taxonomicName id="79F0DF9D6115FFCFFF05FEC216E7FE99" ID-CoL="334J3" authority="Ramirez &amp; Asenjo" authorityName="Ramírez &amp; Asenjo" authorityYear="2017" box="[151,678,325,351]" class="Insecta" family="Staphylinidae" genus="Cylindroxystus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="438" phylum="Arthropoda" rank="species" species="janualis" status="sp. nov.">
<emphasis id="8C84780C6115FFCFFF05FEC215F4FE99" bold="true" box="[151,437,325,351]" italics="true" pageId="1" pageNumber="438">Cylindroxystus janualis</emphasis>
Ramírez &amp; Asenjo
</taxonomicName>
,
<taxonomicNameLabel id="97B7C5776115FFCFFD21FEC21751FE99" box="[691,784,325,351]" pageId="1" pageNumber="438" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="BE4FA41E6115FFCFFF05FEEF154DFEB9" blockId="1.[151,784,290,383]" box="[151,268,360,383]" pageId="1" pageNumber="438">
(
<figureCitation id="26CBB89B6115FFCFFF0DFEEF1542FEB9" box="[159,259,360,383]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,258,1522,1544]" captionTargetBox="[180,1419,208,1492]" captionTargetId="figure@2.[151,1436,194,1498]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Cylindroxystus janualis sp. nov., holotype male. 1, habitus, dorsal view; 2, Right antenna, ss spinelike setae, ts thrid segment cupulate; 3, Right maxilla, la lacinia, g galea, fs fourth segment nipple-shaped; 4, Tergum III, pt paratergites, bc three internal canals at base; 5, Sternite II and III, mlc medial longitudinal carina, ldl lateral longitudinal depression; 6, Tergum VIII; 7, Sternum VIII; 8, aedeagus in ventral view; 9, aedeagus in dorsal view, op operculum. Scale bars: 1, 1 mm; 6 – 7, 0.5 mm; 8, 0.05 mm." httpUri="https://zenodo.org/record/1040029/files/figure.png" pageId="1" pageNumber="438">Figs. 1–9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F6EAF7956115FFCFFF05FE2A1152FD52" pageId="1" pageNumber="438" type="materials_examined">
<paragraph id="BE4FA41E6115FFCFFF05FE2A15D4FDEA" blockId="1.[151,1436,429,1908]" pageId="1" pageNumber="438">
<emphasis id="8C84780C6115FFCFFF05FE2A1515FE02" bold="true" box="[151,340,429,452]" pageId="1" pageNumber="438">Type material (6</emphasis>
♂
<emphasis id="8C84780C6115FFCFFEFAFE2A16CEFE02" bold="true" box="[360,655,429,452]" pageId="1" pageNumber="438">). Holotype: COLOMBIA</emphasis>
: ♂, labeled: “Colombia, / Chocó, / Acandí, / Capurganá, / Camino a los rios, / Bosque, /
<quantity id="790809FB6115FFCFFED7FE5715C3FE21" box="[325,386,464,487]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="438" unit="m" value="250.0">250m</quantity>
[meters of altitude above sea level], / Pitfall trap /
<date id="CA4E82DE6115FFCFFC0CFE571795FE21" box="[926,980,464,487]" pageId="1" pageNumber="438">14. x</date>
[October]. 2008 / C. Becerra, M. Betancur, A. López, J. Ruíz collectors”; “HOLOTYPE [red label]/
<taxonomicName id="79F0DF9D6115FFCFFD60FE7617D0FDCF" authorityName="Bierig" authorityYear="1943" box="[754,913,497,521]" class="Insecta" family="Staphylinidae" genus="Cylindroxystus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="438" phylum="Arthropoda" rank="genus">
<emphasis id="8C84780C6115FFCFFD60FE7617D0FDCF" box="[754,913,497,521]" italics="true" pageId="1" pageNumber="438">Cylindroxystus</emphasis>
</taxonomicName>
/
<taxonomicName id="79F0DF9D6115FFCFFC36FE761089FDCF" authority="Ramirez &amp; Asenjo" authorityName="Ramirez &amp; Asenjo" box="[932,1224,497,521]" class="Insecta" family="Staphylinidae" genus="Cylindroxystus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="438" phylum="Arthropoda" rank="species" species="janualis">
<emphasis id="8C84780C6115FFCFFC36FE7617BBFDCF" box="[932,1018,497,521]" italics="true" pageId="1" pageNumber="438">janualis</emphasis>
Ramírez &amp; Asenjo
</taxonomicName>
/ Desig. Ramírez &amp; Asenjo, 2017” (MPUJ).
</paragraph>
<paragraph id="BE4FA41E6115FFCFFF55FDBF1152FD52" blockId="1.[151,1436,429,1908]" pageId="1" pageNumber="438">
<emphasis id="8C84780C6115FFCFFF55FDBF1503FD89" bold="true" box="[199,322,568,591]" pageId="1" pageNumber="438">Paratypes:</emphasis>
COLOMBIA: 5♂. Labeled: “Colombia, / Chocó, / Acandí, / Capurganá, / Los Girasoles, /
<quantity id="790809FB6115FFCFFA9BFDBF1104FD89" box="[1289,1349,568,591]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="438" unit="m" value="300.0">300m</quantity>
[meters of altitude above sea level], / Bosque, / Pitfall trap /
<date id="CA4E82DE6115FFCFFD51FDDD16B9FDB7" box="[707,760,602,625]" pageId="1" pageNumber="438">11. x</date>
[October]. 2008 / A. Quintero collector”; “PARATYPE [yellow label]/
<taxonomicName id="79F0DF9D6115FFCFFF70FDFB15C0FD52" authorityName="Bierig" authorityYear="1943" box="[226,385,636,660]" class="Insecta" family="Staphylinidae" genus="Cylindroxystus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="438" phylum="Arthropoda" rank="genus">
<emphasis id="8C84780C6115FFCFFF70FDFB15C0FD52" box="[226,385,636,660]" italics="true" pageId="1" pageNumber="438">Cylindroxystus</emphasis>
</taxonomicName>
/
<taxonomicName id="79F0DF9D6115FFCFFE07FDFB16FBFD52" authority="Ramirez &amp; Asenjo" authorityName="Ramirez &amp; Asenjo" box="[405,698,636,660]" class="Insecta" family="Staphylinidae" genus="Cylindroxystus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="438" phylum="Arthropoda" rank="species" species="janualis">
<emphasis id="8C84780C6115FFCFFE07FDFB15AAFD52" box="[405,491,636,660]" italics="true" pageId="1" pageNumber="438">janualis</emphasis>
Ramírez &amp; Asenjo
</taxonomicName>
/ Desig. Ramírez &amp; Asenjo, 2017” (4 MPUJ, 1 CEMT).
</paragraph>
</subSubSection>
<subSubSection id="F6EAF7956115FFCFFF55FD271656FD3A" pageId="1" pageNumber="438" type="diagnosis">
<paragraph id="BE4FA41E6115FFCFFF55FD271656FD3A" blockId="1.[151,1436,429,1908]" pageId="1" pageNumber="438">
<emphasis id="8C84780C6115FFCFFF55FD27157DFD71" bold="true" box="[199,316,672,695]" pageId="1" pageNumber="438">Diagnosis.</emphasis>
<taxonomicName id="79F0DF9D6115FFCFFED6FD181603FD71" authority="Ramírez &amp; Asenjo, 2017" authorityName="Ramírez &amp; Asenjo" authorityYear="2017" box="[324,578,671,695]" class="Insecta" family="Staphylinidae" genus="Cylindroxystus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="438" phylum="Arthropoda" rank="species" species="janualis" status="sp. nov.">
<emphasis id="8C84780C6115FFCFFED6FD181603FD71" box="[324,578,671,695]" italics="true" pageId="1" pageNumber="438">Cylindroxystus janualis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="97B7C5776115FFCFFDDEFD2716E5FD71" box="[588,676,672,695]" pageId="1" pageNumber="438" rank="species">
<emphasis id="8C84780C6115FFCFFDDEFD2716E5FD71" bold="true" box="[588,676,672,695]" pageId="1" pageNumber="438">sp. nov.</emphasis>
</taxonomicNameLabel>
resembles
<taxonomicName id="79F0DF9D6115FFCFFCB3FD2717ACFD71" authority="Herman" authorityName="Herman" box="[801,1005,672,695]" class="Insecta" family="Staphylinidae" genus="Cylindroxystus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="438" phylum="Arthropoda" rank="species" species="messus">
<emphasis id="8C84780C6115FFCFFCB3FD2717CEFD71" box="[801,911,672,695]" italics="true" pageId="1" pageNumber="438">C. messus</emphasis>
Herman
</taxonomicName>
but differs as follows: elytral length is greater than the pronotum length; foramen is larger occupying the basal third, two transverse carinae at the lateral part of the aedeagal operculum (
<figureCitation id="26CBB89B6115FFCFFE30FD62164BFD3A" box="[418,522,741,764]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,258,1522,1544]" captionTargetBox="[180,1419,208,1492]" captionTargetId="figure@2.[151,1436,194,1498]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Cylindroxystus janualis sp. nov., holotype male. 1, habitus, dorsal view; 2, Right antenna, ss spinelike setae, ts thrid segment cupulate; 3, Right maxilla, la lacinia, g galea, fs fourth segment nipple-shaped; 4, Tergum III, pt paratergites, bc three internal canals at base; 5, Sternite II and III, mlc medial longitudinal carina, ldl lateral longitudinal depression; 6, Tergum VIII; 7, Sternum VIII; 8, aedeagus in ventral view; 9, aedeagus in dorsal view, op operculum. Scale bars: 1, 1 mm; 6 – 7, 0.5 mm; 8, 0.05 mm." httpUri="https://zenodo.org/record/1040029/files/figure.png" pageId="1" pageNumber="438">Figs. 8–9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F6EAF7956115FFCFFF55FC8F1723F999" pageId="1" pageNumber="438" type="description">
<paragraph id="BE4FA41E6115FFCFFF55FC8F1628FCD9" blockId="1.[151,1436,429,1908]" box="[199,617,776,799]" pageId="1" pageNumber="438">
<emphasis id="8C84780C6115FFCFFF55FC8F1513FCD9" bold="true" box="[199,338,776,799]" pageId="1" pageNumber="438">Description.</emphasis>
<typeStatus id="614B1ABC6115FFCFFECAFC8F15FAFCD9" box="[344,443,776,799]" pageId="1" pageNumber="438" type="holotype">Holotype</typeStatus>
male, BL: 3.76.
</paragraph>
<paragraph id="BE4FA41E6115FFCFFF55FCAD1712FC87" blockId="1.[151,1436,429,1908]" box="[199,851,810,833]" pageId="1" pageNumber="438">
Body reddish-brown (
<figureCitation id="26CBB89B6115FFCFFE23FCAD15B1FC87" box="[433,496,810,833]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,258,1522,1544]" captionTargetBox="[180,1419,208,1492]" captionTargetId="figure@2.[151,1436,194,1498]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Cylindroxystus janualis sp. nov., holotype male. 1, habitus, dorsal view; 2, Right antenna, ss spinelike setae, ts thrid segment cupulate; 3, Right maxilla, la lacinia, g galea, fs fourth segment nipple-shaped; 4, Tergum III, pt paratergites, bc three internal canals at base; 5, Sternite II and III, mlc medial longitudinal carina, ldl lateral longitudinal depression; 6, Tergum VIII; 7, Sternum VIII; 8, aedeagus in ventral view; 9, aedeagus in dorsal view, op operculum. Scale bars: 1, 1 mm; 6 – 7, 0.5 mm; 8, 0.05 mm." httpUri="https://zenodo.org/record/1040029/files/figure.png" pageId="1" pageNumber="438">Fig. 1</figureCitation>
). Antennae, legs and palps paler.
</paragraph>
<paragraph id="BE4FA41E6115FFCFFF55FCCA1779FA12" blockId="1.[151,1436,429,1908]" pageId="1" pageNumber="438">
Head: Vertex with setigerous punctation confined to edges of the frontal region. Postocular margin of head converging gradually rounded to basal margin of head. Eye length longer than postocular length of head. Gena with setigerous punctation surrounding the eye. Gular sutures widely separated. Labrum edentate and deeply and broadly emarginate at anterior margin. Mandible long, slender, falcate and edentate. Galea and lacinia of maxilla moderately long, forming a bulb with a set of bristles in apical third. Maxillary palpus moderately long, second segment shorter than third and gradually expanded apically, third segment strongly expanded from base to near middle then slightly narrowed towards apex (
<figureCitation id="26CBB89B6115FFCFFEA0FB9A1533FBF2" box="[306,370,1053,1076]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,258,1522,1544]" captionTargetBox="[180,1419,208,1492]" captionTargetId="figure@2.[151,1436,194,1498]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Cylindroxystus janualis sp. nov., holotype male. 1, habitus, dorsal view; 2, Right antenna, ss spinelike setae, ts thrid segment cupulate; 3, Right maxilla, la lacinia, g galea, fs fourth segment nipple-shaped; 4, Tergum III, pt paratergites, bc three internal canals at base; 5, Sternite II and III, mlc medial longitudinal carina, ldl lateral longitudinal depression; 6, Tergum VIII; 7, Sternum VIII; 8, aedeagus in ventral view; 9, aedeagus in dorsal view, op operculum. Scale bars: 1, 1 mm; 6 – 7, 0.5 mm; 8, 0.05 mm." httpUri="https://zenodo.org/record/1040029/files/figure.png" pageId="1" pageNumber="438">Fig. 3</figureCitation>
); Labial palpus with first segment shorter than second and second longer than third; first and second segments of similar thickness and third more slender. Antenna with segments 4–10 each with short, stout, spine-like setae around apex (
<figureCitation id="26CBB89B6115FFCFFEF5FBE515E8FBBF" box="[359,425,1122,1145]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,258,1522,1544]" captionTargetBox="[180,1419,208,1492]" captionTargetId="figure@2.[151,1436,194,1498]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Cylindroxystus janualis sp. nov., holotype male. 1, habitus, dorsal view; 2, Right antenna, ss spinelike setae, ts thrid segment cupulate; 3, Right maxilla, la lacinia, g galea, fs fourth segment nipple-shaped; 4, Tergum III, pt paratergites, bc three internal canals at base; 5, Sternite II and III, mlc medial longitudinal carina, ldl lateral longitudinal depression; 6, Tergum VIII; 7, Sternum VIII; 8, aedeagus in ventral view; 9, aedeagus in dorsal view, op operculum. Scale bars: 1, 1 mm; 6 – 7, 0.5 mm; 8, 0.05 mm." httpUri="https://zenodo.org/record/1040029/files/figure.png" pageId="1" pageNumber="438">Fig. 2</figureCitation>
), segment 10 approximately as long as 9. Thorax: Pronotum longer than wide (PL: 0.43; PW: 0.32) with parallel lateral margins. Disk shinning, with a pair of paramedial longitudinal rows of setigerous punctation and with a few scattered punctures either side of the row. Prosternum with median carina present but short and moderately developed. Hypomeron polished and without punctures or sculpturing. Elytra longer than pronotum (EL: 0.64; EW: 0.5), surface with two discal longitudinal rows of setigerous punctures. Epipleural elytral present. Abdomen: Abdominal segments VII and VIII with tergum and sternum separated (
<figureCitation id="26CBB89B6115FFCFFC07FA97104CFAE1" box="[917,1037,1296,1319]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,258,1522,1544]" captionTargetBox="[180,1419,208,1492]" captionTargetId="figure@2.[151,1436,194,1498]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Cylindroxystus janualis sp. nov., holotype male. 1, habitus, dorsal view; 2, Right antenna, ss spinelike setae, ts thrid segment cupulate; 3, Right maxilla, la lacinia, g galea, fs fourth segment nipple-shaped; 4, Tergum III, pt paratergites, bc three internal canals at base; 5, Sternite II and III, mlc medial longitudinal carina, ldl lateral longitudinal depression; 6, Tergum VIII; 7, Sternum VIII; 8, aedeagus in ventral view; 9, aedeagus in dorsal view, op operculum. Scale bars: 1, 1 mm; 6 – 7, 0.5 mm; 8, 0.05 mm." httpUri="https://zenodo.org/record/1040029/files/figure.png" pageId="1" pageNumber="438">Fig. 1, 6, 7</figureCitation>
). Abdominal segments III to VI with longitudinal depression at base near lateral margin (
<figureCitation id="26CBB89B6115FFCFFD5DFAB51753FA8F" box="[719,786,1330,1353]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,258,1522,1544]" captionTargetBox="[180,1419,208,1492]" captionTargetId="figure@2.[151,1436,194,1498]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Cylindroxystus janualis sp. nov., holotype male. 1, habitus, dorsal view; 2, Right antenna, ss spinelike setae, ts thrid segment cupulate; 3, Right maxilla, la lacinia, g galea, fs fourth segment nipple-shaped; 4, Tergum III, pt paratergites, bc three internal canals at base; 5, Sternite II and III, mlc medial longitudinal carina, ldl lateral longitudinal depression; 6, Tergum VIII; 7, Sternum VIII; 8, aedeagus in ventral view; 9, aedeagus in dorsal view, op operculum. Scale bars: 1, 1 mm; 6 – 7, 0.5 mm; 8, 0.05 mm." httpUri="https://zenodo.org/record/1040029/files/figure.png" pageId="1" pageNumber="438">Fig. 5</figureCitation>
). Sterna II and III with medial longitudinal carina (
<figureCitation id="26CBB89B6115FFCFFAD8FAB511CCFA8F" box="[1354,1421,1330,1353]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,258,1522,1544]" captionTargetBox="[180,1419,208,1492]" captionTargetId="figure@2.[151,1436,194,1498]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Cylindroxystus janualis sp. nov., holotype male. 1, habitus, dorsal view; 2, Right antenna, ss spinelike setae, ts thrid segment cupulate; 3, Right maxilla, la lacinia, g galea, fs fourth segment nipple-shaped; 4, Tergum III, pt paratergites, bc three internal canals at base; 5, Sternite II and III, mlc medial longitudinal carina, ldl lateral longitudinal depression; 6, Tergum VIII; 7, Sternum VIII; 8, aedeagus in ventral view; 9, aedeagus in dorsal view, op operculum. Scale bars: 1, 1 mm; 6 – 7, 0.5 mm; 8, 0.05 mm." httpUri="https://zenodo.org/record/1040029/files/figure.png" pageId="1" pageNumber="438">Fig. 5</figureCitation>
). Tergum III with three internal canals at base (
<figureCitation id="26CBB89B6115FFCFFDEEFAD216FCFAAA" box="[636,701,1365,1388]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,258,1522,1544]" captionTargetBox="[180,1419,208,1492]" captionTargetId="figure@2.[151,1436,194,1498]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Cylindroxystus janualis sp. nov., holotype male. 1, habitus, dorsal view; 2, Right antenna, ss spinelike setae, ts thrid segment cupulate; 3, Right maxilla, la lacinia, g galea, fs fourth segment nipple-shaped; 4, Tergum III, pt paratergites, bc three internal canals at base; 5, Sternite II and III, mlc medial longitudinal carina, ldl lateral longitudinal depression; 6, Tergum VIII; 7, Sternum VIII; 8, aedeagus in ventral view; 9, aedeagus in dorsal view, op operculum. Scale bars: 1, 1 mm; 6 – 7, 0.5 mm; 8, 0.05 mm." httpUri="https://zenodo.org/record/1040029/files/figure.png" pageId="1" pageNumber="438">Fig. 4</figureCitation>
). Sternum VII with broad, shallow, median emargination at posterior margin. Tergum VIII with two lateral and two median internal canals at base and with broad, apically acute, triangular lobe on apical margin (
<figureCitation id="26CBB89B6115FFCFFE19FA1D158CFA77" box="[395,461,1434,1457]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,258,1522,1544]" captionTargetBox="[180,1419,208,1492]" captionTargetId="figure@2.[151,1436,194,1498]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Cylindroxystus janualis sp. nov., holotype male. 1, habitus, dorsal view; 2, Right antenna, ss spinelike setae, ts thrid segment cupulate; 3, Right maxilla, la lacinia, g galea, fs fourth segment nipple-shaped; 4, Tergum III, pt paratergites, bc three internal canals at base; 5, Sternite II and III, mlc medial longitudinal carina, ldl lateral longitudinal depression; 6, Tergum VIII; 7, Sternum VIII; 8, aedeagus in ventral view; 9, aedeagus in dorsal view, op operculum. Scale bars: 1, 1 mm; 6 – 7, 0.5 mm; 8, 0.05 mm." httpUri="https://zenodo.org/record/1040029/files/figure.png" pageId="1" pageNumber="438">Fig. 6</figureCitation>
). Sternum VIII with two lateral and two median internal canals at base and with moderately deep and broad median emargination on apical margin (
<figureCitation id="26CBB89B6115FFCFFD7BFA3A176AFA12" box="[745,811,1469,1492]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,258,1522,1544]" captionTargetBox="[180,1419,208,1492]" captionTargetId="figure@2.[151,1436,194,1498]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Cylindroxystus janualis sp. nov., holotype male. 1, habitus, dorsal view; 2, Right antenna, ss spinelike setae, ts thrid segment cupulate; 3, Right maxilla, la lacinia, g galea, fs fourth segment nipple-shaped; 4, Tergum III, pt paratergites, bc three internal canals at base; 5, Sternite II and III, mlc medial longitudinal carina, ldl lateral longitudinal depression; 6, Tergum VIII; 7, Sternum VIII; 8, aedeagus in ventral view; 9, aedeagus in dorsal view, op operculum. Scale bars: 1, 1 mm; 6 – 7, 0.5 mm; 8, 0.05 mm." httpUri="https://zenodo.org/record/1040029/files/figure.png" pageId="1" pageNumber="438">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="BE4FA41E6115FFCFFF55FA6716D7F9DF" blockId="1.[151,1436,429,1908]" pageId="1" pageNumber="438">
Aedeagus: with median orifice covered by operculum (
<figureCitation id="26CBB89B6115FFCFFC81FA671715FA31" box="[787,852,1504,1527]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,258,1522,1544]" captionTargetBox="[180,1419,208,1492]" captionTargetId="figure@2.[151,1436,194,1498]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Cylindroxystus janualis sp. nov., holotype male. 1, habitus, dorsal view; 2, Right antenna, ss spinelike setae, ts thrid segment cupulate; 3, Right maxilla, la lacinia, g galea, fs fourth segment nipple-shaped; 4, Tergum III, pt paratergites, bc three internal canals at base; 5, Sternite II and III, mlc medial longitudinal carina, ldl lateral longitudinal depression; 6, Tergum VIII; 7, Sternum VIII; 8, aedeagus in ventral view; 9, aedeagus in dorsal view, op operculum. Scale bars: 1, 1 mm; 6 – 7, 0.5 mm; 8, 0.05 mm." httpUri="https://zenodo.org/record/1040029/files/figure.png" pageId="1" pageNumber="438">Fig. 8</figureCitation>
). Operculum with median lobe in basal margin, with a shallow depression in both lateral sides (
<figureCitation id="26CBB89B6115FFCFFDD5F98516C8F9DF" box="[583,649,1538,1561]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,258,1522,1544]" captionTargetBox="[180,1419,208,1492]" captionTargetId="figure@2.[151,1436,194,1498]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Cylindroxystus janualis sp. nov., holotype male. 1, habitus, dorsal view; 2, Right antenna, ss spinelike setae, ts thrid segment cupulate; 3, Right maxilla, la lacinia, g galea, fs fourth segment nipple-shaped; 4, Tergum III, pt paratergites, bc three internal canals at base; 5, Sternite II and III, mlc medial longitudinal carina, ldl lateral longitudinal depression; 6, Tergum VIII; 7, Sternum VIII; 8, aedeagus in ventral view; 9, aedeagus in dorsal view, op operculum. Scale bars: 1, 1 mm; 6 – 7, 0.5 mm; 8, 0.05 mm." httpUri="https://zenodo.org/record/1040029/files/figure.png" pageId="1" pageNumber="438">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="BE4FA41E6115FFCFFF55F9A315D4F9FA" blockId="1.[151,1436,429,1908]" box="[199,405,1572,1596]" pageId="1" pageNumber="438">
<emphasis id="8C84780C6115FFCFFF55F9A3155FF9FA" box="[199,286,1572,1596]" italics="true" pageId="1" pageNumber="438">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="BE4FA41E6115FFCFFF55F9CF1723F999" blockId="1.[151,1436,429,1908]" box="[199,866,1608,1631]" pageId="1" pageNumber="438">
<emphasis id="8C84780C6115FFCFFF55F9CF1564F999" bold="true" box="[199,293,1608,1631]" pageId="1" pageNumber="438">Habitat.</emphasis>
The specimens were collected in rainforest leaf litter.
</paragraph>
</subSubSection>
<subSubSection id="F6EAF7956115FFCFFF55F9ED1502F962" pageId="1" pageNumber="438" type="etymology">
<paragraph id="BE4FA41E6115FFCFFF55F9ED1502F962" blockId="1.[151,1436,429,1908]" pageId="1" pageNumber="438">
<emphasis id="8C84780C6115FFCFFF55F9ED1506F947" bold="true" box="[199,327,1642,1665]" pageId="1" pageNumber="438">Etymology.</emphasis>
The specific epithet name is an eponym based on the name of the brother of the first author, Frank Janual Ramírez.
</paragraph>
</subSubSection>
<subSubSection id="F6EAF7956115FFCFFF55F93716B5F901" box="[199,756,1712,1735]" pageId="1" pageNumber="438" type="distribution">
<paragraph id="BE4FA41E6115FFCFFF55F93716B5F901" blockId="1.[151,1436,429,1908]" box="[199,756,1712,1735]" pageId="1" pageNumber="438">
<emphasis id="8C84780C6115FFCFFF55F9371518F901" bold="true" box="[199,345,1712,1735]" pageId="1" pageNumber="438">Distribution.</emphasis>
Northwest
<collectingCountry id="C6E7E48E6115FFCFFE47F9371603F901" box="[469,578,1712,1735]" name="Colombia" pageId="1" pageNumber="438">Colombia</collectingCountry>
:
<collectingRegion id="7C346AFC6115FFCFFDDEF93716D4F901" box="[588,661,1712,1735]" country="Colombia" name="Choco" pageId="1" pageNumber="438">Chocó</collectingRegion>
; Acandí.
</paragraph>
</subSubSection>
<subSubSection id="F6EAF7956115FFCFFF55F955151CF8B2" pageId="1" pageNumber="438" type="discussion">
<paragraph id="BE4FA41E6115FFCFFF55F955151CF8B2" blockId="1.[151,1436,429,1908]" pageId="1" pageNumber="438">
<emphasis id="8C84780C6115FFCFFF55F9551508F92F" bold="true" box="[199,329,1746,1769]" pageId="1" pageNumber="438">Comments.</emphasis>
The new species belongs to the genus
<taxonomicName id="79F0DF9D6115FFCFFD7CF95617CCF92F" authorityName="Bierig" authorityYear="1943" box="[750,909,1745,1769]" class="Insecta" family="Staphylinidae" genus="Cylindroxystus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="438" phylum="Arthropoda" rank="genus">
<emphasis id="8C84780C6115FFCFFD7CF95617CCF92F" box="[750,909,1745,1769]" italics="true" pageId="1" pageNumber="438">Cylindroxystus</emphasis>
</taxonomicName>
based on the characters used by
<bibRefCitation id="DA61D9EF6115FFCFFB62F95511D6F92F" author="Herman" box="[1264,1431,1746,1769]" pageId="1" pageNumber="438" refString="Herman, L. H. (1991) Revision of the subtribe Cylindroxystina (Coleoptera: Staphylinidae: Paederinae). Bulletin of the American Museum of Natural History, 203, 1 - 83." type="journal article" year="1991">Herman (1991)</bibRefCitation>
, and that are in the new species: presence of one pair of paratergites on segment III (fig. 4) and their absence from segments IV to VII (
<figureCitation id="26CBB89B6115FFCFFEE6F89F15F3F8E9" box="[372,434,1816,1839]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,258,1522,1544]" captionTargetBox="[180,1419,208,1492]" captionTargetId="figure@2.[151,1436,194,1498]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Cylindroxystus janualis sp. nov., holotype male. 1, habitus, dorsal view; 2, Right antenna, ss spinelike setae, ts thrid segment cupulate; 3, Right maxilla, la lacinia, g galea, fs fourth segment nipple-shaped; 4, Tergum III, pt paratergites, bc three internal canals at base; 5, Sternite II and III, mlc medial longitudinal carina, ldl lateral longitudinal depression; 6, Tergum VIII; 7, Sternum VIII; 8, aedeagus in ventral view; 9, aedeagus in dorsal view, op operculum. Scale bars: 1, 1 mm; 6 – 7, 0.5 mm; 8, 0.05 mm." httpUri="https://zenodo.org/record/1040029/files/figure.png" pageId="1" pageNumber="438">Fig. 1</figureCitation>
), and by the cupulate third antennal segment (fig. 2). Tergum and sternum IV to VI are fused to form a cylinder (
<figureCitation id="26CBB89B6115FFCFFEFFF8BD15ECF897" box="[365,429,1850,1873]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,258,1522,1544]" captionTargetBox="[180,1419,208,1492]" captionTargetId="figure@2.[151,1436,194,1498]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Cylindroxystus janualis sp. nov., holotype male. 1, habitus, dorsal view; 2, Right antenna, ss spinelike setae, ts thrid segment cupulate; 3, Right maxilla, la lacinia, g galea, fs fourth segment nipple-shaped; 4, Tergum III, pt paratergites, bc three internal canals at base; 5, Sternite II and III, mlc medial longitudinal carina, ldl lateral longitudinal depression; 6, Tergum VIII; 7, Sternum VIII; 8, aedeagus in ventral view; 9, aedeagus in dorsal view, op operculum. Scale bars: 1, 1 mm; 6 – 7, 0.5 mm; 8, 0.05 mm." httpUri="https://zenodo.org/record/1040029/files/figure.png" pageId="1" pageNumber="438">Fig. 1</figureCitation>
), the fourth segment of the maxillary palpus is nipple-shaped (fig. 3), and the mesothoracic peritreme is small.
</paragraph>
</subSubSection>
</treatment>
</document>
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