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<document id="05A963363D7FB256C647253DFD6598BA" ID-DOI="10.15298/rusentj.28.3.03" ID-ISSN="0132-8069" ID-Zenodo-Dep="13164009" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722579496367" checkinUser="felipe" docAuthor="Fominykh, D. D., Zamotajlov, A. S. &amp; Titarenko, A. Yu." docDate="2019" docId="2B00EE45FFDD5415FF0BC1E2C6E78467" docLanguage="en" docName="RussEntomolJ.28.3.251-257.pdf" docOrigin="Russian Entomological Journal 28 (3)" docSource="http://dx.doi.org/10.15298/rusentj.28.3.03" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.2:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="2" docTitle="Carabus (Neoplectes) martviliensis subsp. mzekalae Fominykh, Zamotajlov &amp; Titarenko, 2019, ssp.n." docType="treatment" docVersion="2" lastPageNumber="257" masterDocId="D739963DFFDF5413FFE6C062C466877C" masterDocTitle="Description of a new subspecies of Carabus (Neoplectes) martviliensis Retezár et Djavelidze, 1992 from Central Georgia (Coleoptera: Carabidae)" masterLastPageNumber="257" masterPageNumber="251" pageNumber="253" updateTime="1722580853212" updateUser="ExternalLinkService">
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<mods:title id="14B7ED1D5DEBB4F4D9B4AC2F566608E8">Description of a new subspecies of Carabus (Neoplectes) martviliensis Retezár et Djavelidze, 1992 from Central Georgia (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="BE980BCB25CE914ACC5637C0C923C7A5">Fominykh, D. D.</mods:namePart>
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<mods:namePart id="1EFC48E9EEED6413E0F5A9F27151A574">Zamotajlov, A. S.</mods:namePart>
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<mods:namePart id="D277CF727F5E098F6C7259C45A3BBB6B">Titarenko, A. Yu.</mods:namePart>
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<mods:title id="1EB1B38F589A4FF809262F975A2FE87D">Russian Entomological Journal</mods:title>
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<mods:date id="4409EEFF2F239D3A47D06A330DDB9E49">2019</mods:date>
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<mods:number id="73FE49C190605BBD5F9A04EF577EFF54">2019-09-30</mods:number>
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<mods:identifier id="B3A47693024B74D07D930AAF69861902" type="DOI">10.15298/rusentj.28.3.03</mods:identifier>
<mods:identifier id="8A669D7F7F10DD80B9FAEA36542CCFA9" type="ISSN">0132-8069</mods:identifier>
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<treatment id="2B00EE45FFDD5415FF0BC1E2C6E78467" LSID="urn:lsid:plazi:treatment:2B00EE45FFDD5415FF0BC1E2C6E78467" httpUri="http://treatment.plazi.org/id/2B00EE45FFDD5415FF0BC1E2C6E78467" lastPageId="6" lastPageNumber="257" pageId="2" pageNumber="253">
<subSubSection id="EBB30CD8FFDD5411FF0BC1E2C6AE86C9" pageId="2" pageNumber="253" type="nomenclature">
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<heading id="F85EE83FFFDD5411FF0BC1E2C6B086E4" box="[237,726,384,408]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="253" reason="8">
<taxonomicName id="64A924D0FFDD5411FF0BC1E2C6B086E4" ID-CoL="8MGBH" authorityName="Fominykh &amp; Zamotajlov &amp; Titarenko" authorityYear="2019" box="[237,726,384,408]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="253" phylum="Arthropoda" rank="subSpecies" species="martviliensis" status="ssp. n." subGenus="Neoplectes" subSpecies="mzekalae">
<emphasis id="91DD8341FFDD5411FF0BC1E2C52F86E4" box="[237,329,384,408]" italics="true" pageId="2" pageNumber="253">Carabus</emphasis>
(
<emphasis id="91DD8341FFDD5411FEBCC1E2C5A986E4" box="[346,463,384,408]" italics="true" pageId="2" pageNumber="253">Neoplectes</emphasis>
)
<emphasis id="91DD8341FFDD5411FE06C1E2C6B086E4" box="[480,726,384,408]" italics="true" pageId="2" pageNumber="253">martviliensis mzekalae</emphasis>
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Fominykh, Zamotajlov et Titarenko,
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<taxonomicNameLabel id="8AEE3E3AFFDD5411FD6EC1FFC6AE86C9" box="[648,712,413,437]" pageId="2" pageNumber="253" rank="subSpecies">ssp.n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="EBB30CD8FFDD5411FED3C1D9C6E986AF" box="[309,655,443,467]" pageId="2" pageNumber="253" type="description">
<paragraph id="A3165F53FFDD5411FED3C1D9C6E986AF" blockId="2.[189,776,384,691]" box="[309,655,443,467]" pageId="2" pageNumber="253">
<figureCitation id="3B9243D6FFDD5411FED3C1D9C5F386AF" box="[309,405,443,467]" captionStart-0="Fig" captionStart-1="Figs 2–9" captionStartId-0="1.[226,257,1846,1865]" captionStartId-1="2.[226,263,1966,1985]" captionTargetBox-0="[272,1331,971,1827]" captionTargetBox-1="[190,1396,720,1946]" captionTargetId-0="figure-623@1.[272,1331,971,1827]" captionTargetId-1="figure-361@2.[189,1399,717,1946]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Fig. 1. Studied localities of Carabus (Neoplectes) martviliensis in Central Georgia: 1–8 — C. martviliensis martviliensis; 9–11 — C. martviliensis mzekalae ssp.n.; 1 — Ofitsari Mt. Range, 1900 m; 2 — Ofitsari Mt. Range, 1500 m; 3 — Doberazeni env., 600 m; 4 — Doberazeni env., 500 m; 5 — Mt. Muchera, 1009 m; 6 — Zeda Gordi env., 980 m; 7 — Kinchkha-Perdi env., 816 m; 8 — Satziskvilo env., 1000 m; 9 — Dzedzileti env., 520–620 m; 10 — Dzedzileti env., 770 m; 11 — Melouri env., 475 m. Рис. 1. ИЗученное распространение Carabus (Neoplectes) martviliensis в центральной части ГруЗии: 1–8 — C. martviliensis martviliensis; 9–11 — C. martviliensis mzekalae ssp.n.; 1 — хр. Офицари, 1900 м над ур. м.; 2 — хр. Офицари, 1500 м над ур. м.; 3 — окр. ДобераЗени, 600 м над ур. м.; 4 — окр. ДобераЗени, 500 м над ур. м.; 5 — г. Мучера, 1009 м над ур. м.; 6 — окр. Зеда Горди, 980 м над ур. м.; 7 — окр. Кинчха Перди, 816 м над ур. м.; 8 — окр. Сацисквило, 1000 м над ур. м.; 9 — окр. ДЗедЗилети, 520–620 м над ур. м.; 10 — окр. ДЗедЗилети, 770 м над ур. м.; 11 — окр. Мелоури, 475 м над ур. м." captionText-1="Figs 2–9. Dorsal habitus of Carabus martviliensis mzekalae ssp.n.: 2–5 — Melouri env., 475 m; 6–9 — Dzedzileti env., 770 m; 2–3, 6–7 — males; 4–5, 8–9 — females. Scale bar: 10.0 mm. Рис. 2–9. Габитус сверху Carabus martviliensis mzekalae ssp.n.: 2–5 — окр. Мелоури, 475 м над ур. м.; 6–9 — окр. ДЗедЗилети, 770 м над ур. м.; 2–3, 6–7 — самцы; 4–5, 8–9 — самки. МасШтаб: 10,0 мм." figureDoi-0="http://doi.org/10.5281/zenodo.13164013" figureDoi-1="http://doi.org/10.5281/zenodo.13164017" httpUri-0="https://zenodo.org/record/13164013/files/figure.png" httpUri-1="https://zenodo.org/record/13164017/files/figure.png" pageId="2" pageNumber="253">Figs 1–9</figureCitation>
,
<figureCitation id="3B9243D6FFDD5411FE44C1D9C65F86AF" box="[418,569,443,467]" captionStart="Figs 18–25" captionStartId="4.[226,263,1869,1888]" captionTargetBox="[201,1384,947,1827]" captionTargetId="figure-585@4.[187,1397,943,1831]" captionTargetPageId="4" captionText="Figs 18–25. Apical portion of aedeagus and fully inflated endophallus preparation of Carabus (Neoplectes) martviliensis, right lateral view on top, left lateral view below: 18, 22 — C. martviliensis mzekalae ssp.n., Melouri env., 475 m; 19, 23 — C. martviliensis mzekalae ssp.n., Dzedzileti env., 770 m; 20, 24 — C. martviliensis martviliensis, Zeda Gordi env., 980 m; 21, 25 — C. martviliensis martviliensis, Doberazeni env., 600 m. Scale bar: 10.0 mm. Рис. 18–25. ВерШина Эдеагуса и препарат полностью вывернутого внутреннего меШка Carabus (Neoplectes) martviliensis, вверху — сбоку справа, вниЗу — сбоку слева: 18, 22 — C. martviliensis mzekalae ssp.n., окр. Мелоури, 475 м над ур.м.; 19, 23 — C. martviliensis mzekalae ssp.n., окр. ДЗедЗилети, 770 м над ур.м.; 20, 24 — C. martviliensis martviliensis, окр. Зеда Горди, 980 м над ур. м.; 21, 25 — C. martviliensis martviliensis, окр. ДобераЗени, 600 м над ур.м. МасШтаб: 10,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13164021" httpUri="https://zenodo.org/record/13164021/files/figure.png" pageId="2" pageNumber="253">18–19, 22–23</figureCitation>
,
<figureCitation id="3B9243D6FFDD5411FDA3C1D9C60786AF" box="[581,609,443,467]" captionStart="Fig" captionStartId="5.[226,257,1120,1139]" captionTargetBox="[280,1321,332,1094]" captionTargetId="figure-205@5.[270,1329,314,1108]" captionTargetPageId="5" captionText="Fig. 26. Distribution of morphometric characters of Carabus (Neoplectes) martviliensis subspecies constructed using discriminant analysis based on 9 parameters (see text for measurements details). Рис. 26. Распределение морфометрических характеристик подвидов Carabus (Neoplectes) martviliensis, построенное с испольЗованием дискриминантного аналиЗа, основанного на 9 параметрах (описание промеров см. текст)." figureDoi="http://doi.org/10.5281/zenodo.13164023" httpUri="https://zenodo.org/record/13164023/files/figure.png" pageId="2" pageNumber="253">26</figureCitation>
–28.
</paragraph>
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<paragraph id="A3165F53FFDD5411FF04C182C79086EA" blockId="2.[189,776,384,691]" lastBlockId="2.[812,1399,313,695]" pageId="2" pageNumber="253">
MATERIAL.
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(
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),
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, left bank of
<collectingMunicipality id="4372C529FFDD5411FE9EC19AC6418577" box="[376,551,504,523]" pageId="2" pageNumber="253">river Tskhenistskali</collectingMunicipality>
near
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,
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,
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</materialsCitation>
<collectionCode id="C5B8C796FFDD5411FF33C26DC480855E" box="[213,230,527,546]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="253" type="Museum">V</collectionCode>
–15.VII.2018,
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leg.;
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<typeStatus id="7C12E1F1FFDD5411FE18C272C632855F" box="[510,596,528,547]" pageId="2" pageNumber="253" type="paratype">Paratypes</typeStatus>
:
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,
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(
<collectionCode id="C5B8C796FFDD5411FD26C272C69B855F" box="[704,765,528,547]" pageId="2" pageNumber="253">CFDD</collectionCode>
), same data as holotype
</materialsCitation>
;
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<specimenCount id="B5AF94DAFFDD5411FE73C24AC5B88547" box="[405,478,549,571]" count="113" pageId="2" pageNumber="253" type="male">113♂♂</specimenCount>
,
<specimenCount id="B5AF94DAFFDD5411FE0BC24AC6438547" box="[493,549,549,571]" count="139" pageId="2" pageNumber="253" type="female">139♀♀</specimenCount>
(
<collectionCode id="C5B8C796FFDD5411FDD2C24AC6148547" box="[564,626,552,571]" country="Thailand" httpUri="http://grbio.org/cool/w22j-vay4" name="Department of Microbiology, Songkla University" pageId="2" pageNumber="253">MPSU</collectionCode>
,
<collectionCode id="C5B8C796FFDD5411FD99C24AC6DC8547" box="[639,698,552,571]" pageId="2" pageNumber="253">CFDD</collectionCode>
,
<collectionCode id="C5B8C796FFDD5411FD21C24AC7628547" box="[711,772,552,571]" pageId="2" pageNumber="253">CTAY</collectionCode>
,
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),
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, left bank of
<location id="A6760988FFDD5411FE2EC222C613852F" LSID="urn:lsid:plazi:treatment:2B00EE45FFDD5415FF0BC1E2C6E78467:A6760988FFDD5411FE2EC222C613852F" box="[456,629,576,595]" name="river Tskhenistskali" pageId="2" pageNumber="253" stateProvince="Imereti">river Tskhenistskali</location>
near Dzedzileti,
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<elevation id="2884B860FFDD5411FF5BC23AC4918517" box="[189,247,600,619]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="2" pageNumber="253" unit="m" value="620.0">620 m</elevation>
</quantity>
,
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</materialsCitation>
<collectionCode id="C5B8C796FFDD5411FEFFC23AC54C8517" box="[281,298,600,619]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="253" type="Museum">V</collectionCode>
–13.VII.2019,
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,
<collectorName id="0E5C3A85FFDD5411FDCBC23AC6E58517" box="[557,643,600,619]" pageId="2" pageNumber="253">A. Zubov</collectorName>
,
<collectorName id="0E5C3A85FFDD5411FD6BC23AC6898517" box="[653,751,600,619]" pageId="2" pageNumber="253">B. Dubinin</collectorName>
&amp;
<collectorName id="0E5C3A85FFDD5411FF5BC212C54E85FF" box="[189,296,624,643]" pageId="2" pageNumber="253">D. Safronov</collectorName>
leg.;
<materialsCitation id="13C1550EFFDD5411FEB8C212C5B585CE" collectionCode="MPSU, CFDD, CTAY, CZAM" collectorName="Dzedzileti, NW" elevation="770" location="Mt. Gormagali" municipality="river Tskhenistskali" pageId="2" pageNumber="253" specimenCount="79" specimenCount-female="63" specimenCount-male="16" stateProvince="Imereti" typeStatus="paratype">
<specimenCount id="B5AF94DAFFDD5411FEB8C212C5FA85FF" box="[350,412,621,643]" count="16" pageId="2" pageNumber="253" type="male">16♂♂</specimenCount>
,
<specimenCount id="B5AF94DAFFDD5411FE4FC212C5B185FF" box="[425,471,621,643]" count="63" pageId="2" pageNumber="253" type="female">63♀♀</specimenCount>
(
<collectionCode id="C5B8C796FFDD5411FE03C212C64585FF" box="[485,547,624,643]" country="Thailand" httpUri="http://grbio.org/cool/w22j-vay4" name="Department of Microbiology, Songkla University" pageId="2" pageNumber="253">MPSU</collectionCode>
,
<collectionCode id="C5B8C796FFDD5411FDC8C212C60F85FF" box="[558,617,624,643]" pageId="2" pageNumber="253">CFDD</collectionCode>
,
<collectionCode id="C5B8C796FFDD5411FD92C212C6D785FF" box="[628,689,624,643]" pageId="2" pageNumber="253">CTAY</collectionCode>
,
<collectionCode id="C5B8C796FFDD5411FD5AC212C69B85FF" box="[700,765,624,643]" pageId="2" pageNumber="253">CZAM</collectionCode>
),
<collectingRegion id="616D91B1FFDD5411FF5BC2EAC49885E7" box="[189,254,648,667]" country="Georgia" name="Imereti" pageId="2" pageNumber="253">Imereti</collectingRegion>
, left bank of
<collectingMunicipality id="4372C529FFDD5411FE8AC2EAC67185E7" box="[364,535,648,667]" pageId="2" pageNumber="253">river Tskhenistskali</collectingMunicipality>
near
<collectorName id="0E5C3A85FFDD5411FDA3C2EAC6AA85E7" box="[581,716,648,667]" pageId="2" pageNumber="253">Dzedzileti, NW</collectorName>
slopes of
<location id="A6760988FFDD5411FF3FC2C2C50685CF" LSID="urn:lsid:plazi:treatment:2B00EE45FFDD5415FF0BC1E2C6E78467:A6760988FFDD5411FF3FC2C2C50685CF" box="[217,352,672,691]" municipality="river Tskhenistskali" name="Mt. Gormagali" pageId="2" pageNumber="253" stateProvince="Imereti">Mt. Gormagali</location>
,
<quantity id="6451F2B6FFDD5411FE89C2C2C5CB85CF" box="[367,429,672,691]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="2" pageNumber="253" unit="m" value="770.0">
<elevation id="2884B860FFDD5411FE89C2C2C5CB85CF" box="[367,429,672,691]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="2" pageNumber="253" unit="m" value="770.0">770 m</elevation>
</quantity>
,
<date id="D7177993FFDD5411FE5CC2FDC5B585CE" box="[442,467,671,690]" pageId="2" pageNumber="253" value="2019-05-15" valueMax="2019-07-13" valueMin="2019-05-15">15.</date>
</materialsCitation>
<collectionCode id="C5B8C796FFDD5411FE34C2FDC58585CE" box="[466,483,671,690]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="253" type="Museum">V</collectionCode>
–13.VII.2019,
<collectorName id="0E5C3A85FFDD5411FD8FC2C2C68285CF" box="[617,740,672,691]" pageId="2" pageNumber="253">D. Fominykh</collectorName>
,
<collectorName id="0E5C3A85FFDD5411FD14C2C2C7008632" box="[754,870,315,691]" pageId="2" pageNumber="253">A. Zubov</collectorName>
,
<collectorName id="0E5C3A85FFDD5411FC95C15EC7BE8632" box="[883,984,315,335]" pageId="2" pageNumber="253">B. Dubinin</collectorName>
&amp;
<collectorName id="0E5C3A85FFDD5411FC1DC15EC00E8632" box="[1019,1128,315,335]" pageId="2" pageNumber="253">D. Safronov</collectorName>
leg.;
<materialsCitation id="13C1550EFFDD5411FB47C159C7DA8602" collectionCode="MPSU, CFDD, CTAY, CZAM" elevation="475" location="river Semi" pageId="2" pageNumber="253" specimenCount="192" specimenCount-female="143" specimenCount-male="49" stateProvince="Imereti" typeStatus="paratype">
<specimenCount id="B5AF94DAFFDD5411FB47C159C0B98633" box="[1185,1247,313,335]" count="49" pageId="2" pageNumber="253" type="male">49♂♂</specimenCount>
,
<specimenCount id="B5AF94DAFFDD5411FB08C159C1408633" box="[1262,1318,313,335]" count="143" pageId="2" pageNumber="253" type="female">143♀♀</specimenCount>
(
<collectionCode id="C5B8C796FFDD5411FAD3C159C1158632" box="[1333,1395,315,334]" country="Thailand" httpUri="http://grbio.org/cool/w22j-vay4" name="Department of Microbiology, Songkla University" pageId="2" pageNumber="253">MPSU</collectionCode>
,
<collectionCode id="C5B8C796FFDD5411FCCAC131C701861A" box="[812,871,339,358]" pageId="2" pageNumber="253">CFDD</collectionCode>
,
<collectionCode id="C5B8C796FFDD5411FC94C131C7C9861A" box="[882,943,339,358]" pageId="2" pageNumber="253">CTAY</collectionCode>
,
<collectionCode id="C5B8C796FFDD5411FC5FC131C79C861A" box="[953,1018,339,358]" pageId="2" pageNumber="253">CZAM</collectionCode>
),
<collectingRegion id="616D91B1FFDD5411FBEDC131C02A861A" box="[1035,1100,339,358]" country="Georgia" name="Imereti" pageId="2" pageNumber="253">Imereti</collectingRegion>
, left bank of
<location id="A6760988FFDD5411FB25C131C179861A" LSID="urn:lsid:plazi:treatment:2B00EE45FFDD5415FF0BC1E2C6E78467:A6760988FFDD5411FB25C131C179861A" box="[1219,1311,339,358]" name="river Semi" pageId="2" pageNumber="253" stateProvince="Imereti">river Semi</location>
near Melouri,
<quantity id="6451F2B6FFDD5411FC86C109C7FC8603" box="[864,922,363,383]" metricMagnitude="2" metricUnit="m" metricValue="4.75" pageId="2" pageNumber="253" unit="m" value="475.0">
<elevation id="2884B860FFDD5411FC86C109C7FC8603" box="[864,922,363,383]" metricMagnitude="2" metricUnit="m" metricValue="4.75" pageId="2" pageNumber="253" unit="m" value="475.0">475 m</elevation>
</quantity>
,
<date id="D7177993FFDD5411FC45C109C7DA8602" box="[931,956,363,382]" pageId="2" pageNumber="253" value="2019-05-19" valueMax="2019-07-12" valueMin="2019-05-19">19.</date>
</materialsCitation>
<collectionCode id="C5B8C796FFDD5411FC5DC109C7AA8602" box="[955,972,363,382]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="253" type="Museum">V</collectionCode>
–12.VII.2019,
<collectorName id="0E5C3A85FFDD5411FBABC10EC0A58602" box="[1101,1219,363,383]" pageId="2" pageNumber="253">D. Fominykh</collectorName>
,
<collectorName id="0E5C3A85FFDD5411FB2AC109C1478602" box="[1228,1313,363,382]" pageId="2" pageNumber="253">A. Zubov</collectorName>
,
<collectorName id="0E5C3A85FFDD5411FACCC10EC72186EA" pageId="2" pageNumber="253">B. Dubinin</collectorName>
&amp;
<collectorName id="0E5C3A85FFDD5411FC83C1E6C7A986EA" box="[869,975,387,407]" pageId="2" pageNumber="253">D. Safronov</collectorName>
leg.
</paragraph>
<paragraph id="A3165F53FFDD5410FCB4C1F9C61C85FA" blockId="2.[812,1399,313,695]" lastBlockId="3.[189,775,315,700]" lastPageId="3" lastPageNumber="254" pageId="2" pageNumber="253">
COMPARATIVE MATERIAL.
<taxonomicName id="64A924D0FFDD5411FB9DC1F9C17586BA" authority="Retezar et Djavelidze, 1992: 18" authorityName="Retezar et Djavelidze" authorityPageNumber="18" authorityYear="1992" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="253" phylum="Arthropoda" rank="subSpecies" species="martviliensis" subGenus="Neoplectes" subSpecies="martviliensis">
<emphasis id="91DD8341FFDD5411FB9DC1F9C0A086D2" box="[1147,1222,411,430]" italics="true" pageId="2" pageNumber="253">Carabus</emphasis>
(
<emphasis id="91DD8341FFDD5411FB3FC1F9C15F86D2" box="[1241,1337,411,430]" italics="true" pageId="2" pageNumber="253">Neoplectes</emphasis>
)
<emphasis id="91DD8341FFDD5411FAAAC1FEC79786BA" italics="true" pageId="2" pageNumber="253">martviliensis martviliensis</emphasis>
Retezár et Djavelidze, 1992:
<materialsCitation id="13C1550EFFDD5411FB18C1D1C17586BA" box="[1278,1299,435,454]" collectionCode="CFDD, CTAY" elevation="1009" location="Mt. Muchera" pageId="2" pageNumber="253" specimenCount="40" specimenCount-female="22" specimenCount-male="18">
<specimenCount id="B5AF94DAFFDD5411FB18C1D1C17586BA" box="[1278,1299,435,454]" count="18" pageId="2" pageNumber="253" type="male">18</specimenCount>
</materialsCitation>
</taxonomicName>
♂♂,
<specimenCount id="B5AF94DAFFDD5411FAAFC1D1C11086BB" box="[1353,1398,433,455]" count="22" pageId="2" pageNumber="253" type="female">22♀♀</specimenCount>
(
<collectionCode id="C5B8C796FFDD5411FCD4C1A9C70986A2" box="[818,879,459,478]" pageId="2" pageNumber="253">CFDD</collectionCode>
,
<collectionCode id="C5B8C796FFDD5411FC98C1A9C7DA86A2" box="[894,956,459,478]" pageId="2" pageNumber="253">CTAY</collectionCode>
),
<taxonomicName id="64A924D0FFDD5411FC37C1A9C0ED86A2" authority=", Samegrelo" authorityName="Samegrelo" box="[977,1163,459,479]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="253" phylum="Arthropoda" rank="species" species="georgia">
<emphasis id="91DD8341FFDD5411FC37C1A9C07A86A2" bold="true" box="[977,1052,459,478]" pageId="2" pageNumber="253">Georgia</emphasis>
, Samegrelo
</taxonomicName>
,
<quantity id="6451F2B6FFDD5411FB7FC1A9C0BD86A2" box="[1177,1243,459,478]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="253" unit="km" value="3.5">3.5 km</quantity>
of Meore-Balda
<locationDeviation id="B2A44671FFDD5411FCCAC181C02F868A" box="[812,1097,483,502]" firstLocation="Abasha" pageId="2" pageNumber="253" secondLocation="Chachkheri">between Abasha and Chachkheri</locationDeviation>
rivers, SE slopes of
<location id="A6760988FFDD5411FAE6C186C115868A" LSID="urn:lsid:plazi:treatment:2B00EE45FFDD5415FF0BC1E2C6E78467:A6760988FFDD5411FAE6C186C115868A" box="[1280,1395,483,503]" name="Mt. Muchera" pageId="2" pageNumber="253">Mt. Muchera</location>
,
<quantity id="6451F2B6FFDD5411FCCAC199C7128573" box="[812,884,507,527]" metricMagnitude="3" metricUnit="m" metricValue="1.009" pageId="2" pageNumber="253" unit="m" value="1009.0">
<elevation id="2884B860FFDD5411FCCAC199C7128573" box="[812,884,507,527]" metricMagnitude="3" metricUnit="m" metricValue="1.009" pageId="2" pageNumber="253" unit="m" value="1009.0">1009 m</elevation>
</quantity>
,
<date id="D7177993FFDD5411FC66C199C0478572" box="[896,1057,507,526]" pageId="2" pageNumber="253" value="2015-04-15" valueMax="2015-06-20" valueMin="2015-04-15">
15.IV–20.
<collectionCode id="C5B8C796FFDD5411FC31C199C7968572" box="[983,1008,507,526]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="2" pageNumber="253">VI</collectionCode>
.2015
</date>
,
<collectorName id="0E5C3A85FFDD5411FBC8C19EC0C28572" box="[1070,1188,507,527]" pageId="2" pageNumber="253">D. Fominykh</collectorName>
&amp;
<collectorName id="0E5C3A85FFDD5411FB2EC199C1798572" box="[1224,1311,507,526]" pageId="2" pageNumber="253">A. Zubov</collectorName>
leg.;
<materialsCitation id="13C1550EFFDD5411FABEC199C00A8542" collectionCode="CFDD" elevation="816" location="river Okatse" pageId="2" pageNumber="253" specimenCount="1" specimenCount-male="1">
<specimenCount id="B5AF94DAFFDD5411FABEC199C1108573" box="[1368,1398,505,527]" count="1" pageId="2" pageNumber="253" type="male">1♂</specimenCount>
(
<collectionCode id="C5B8C796FFDD5411FCD4C271C716855A" box="[818,880,531,550]" pageId="2" pageNumber="253">CFDD</collectionCode>
), Samegrelo, Kinchkha-Perdi vic., right bank of
<location id="A6760988FFDD5411FAF1C271C7588543" LSID="urn:lsid:plazi:treatment:2B00EE45FFDD5415FF0BC1E2C6E78467:A6760988FFDD5411FAF1C271C7588543" name="river Okatse" pageId="2" pageNumber="253">river Okatse</location>
(=Satsikvilo),
<quantity id="6451F2B6FFDD5411FC2CC249C0618543" box="[970,1031,555,575]" metricMagnitude="2" metricUnit="m" metricValue="8.16" pageId="2" pageNumber="253" unit="m" value="816.0">
<elevation id="2884B860FFDD5411FC2CC249C0618543" box="[970,1031,555,575]" metricMagnitude="2" metricUnit="m" metricValue="8.16" pageId="2" pageNumber="253" unit="m" value="816.0">816 m</elevation>
</quantity>
,
<date id="D7177993FFDD5411FBF3C249C00A8542" box="[1045,1132,555,574]" pageId="2" pageNumber="253" value="2015-04-15" valueMax="2015-06-20" valueMin="2015-04-15">15.IV–20.</date>
</materialsCitation>
<collectionCode id="C5B8C796FFDD5411FB8DC249C0E28542" box="[1131,1156,555,574]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="2" pageNumber="253">VI</collectionCode>
.2015,
<collectorName id="0E5C3A85FFDD5411FB22C24EC15D8542" box="[1220,1339,555,575]" pageId="2" pageNumber="253">D. Fominykh</collectorName>
&amp;
<collectorName id="0E5C3A85FFDD5411FA87C249C702852A" pageId="2" pageNumber="253">A. Zubov</collectorName>
leg.;
<materialsCitation id="13C1550EFFDD5411FC7AC221C0258512" collectionCode="CFDD" elevation="1000" location="river Okatze" pageId="2" pageNumber="253" specimenCount="2" specimenCount-female="2" stateProvince="Imereti">
<specimenCount id="B5AF94DAFFDD5411FC7AC221C7D9852B" box="[924,959,577,599]" count="2" pageId="2" pageNumber="253" type="female">2♀♀</specimenCount>
(
<collectionCode id="C5B8C796FFDD5411FC2BC221C06D852A" box="[973,1035,579,598]" pageId="2" pageNumber="253">CFDD</collectionCode>
),
<collectingRegion id="616D91B1FFDD5411FBFBC221C038852A" box="[1053,1118,579,598]" country="Georgia" name="Imereti" pageId="2" pageNumber="253">Imereti</collectingRegion>
, Satziskvilo vic., right bank of
<location id="A6760988FFDD5411FCCAC239C7FD8512" LSID="urn:lsid:plazi:treatment:2B00EE45FFDD5415FF0BC1E2C6E78467:A6760988FFDD5411FCCAC239C7FD8512" box="[812,923,603,622]" name="river Okatze" pageId="2" pageNumber="253" stateProvince="Imereti">river Okatze</location>
,
<quantity id="6451F2B6FFDD5411FC78C239C7858513" box="[926,995,603,623]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="253" unit="m" value="1000.0">
<elevation id="2884B860FFDD5411FC78C239C7858513" box="[926,995,603,623]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="253" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="D7177993FFDD5411FC0AC239C0258512" box="[1004,1091,603,622]" pageId="2" pageNumber="253" value="2016-04-19" valueMax="2016-06-12" valueMin="2016-04-19">19.IV–12.</date>
</materialsCitation>
<collectionCode id="C5B8C796FFDD5411FBA7C239C03C8512" box="[1089,1114,603,622]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="2" pageNumber="253">VI</collectionCode>
.2016,
<collectorName id="0E5C3A85FFDD5411FB71C23EC16D8512" box="[1175,1291,603,623]" pageId="2" pageNumber="253">D. Fominykh</collectorName>
leg.;
<materialsCitation id="13C1550EFFDD5411FAD8C23EC04185E2" collectionCode="CFDD" elevation="1000" location="river Tskhenistskali" pageId="2" pageNumber="253" specimenCount="17" specimenCount-female="10" specimenCount-male="7" stateProvince="Imereti">
<specimenCount id="B5AF94DAFFDD5411FAD8C23EC1178513" box="[1342,1393,601,623]" count="7" pageId="2" pageNumber="253" type="male">7♂♂</specimenCount>
,
<specimenCount id="B5AF94DAFFDD5411FCCAC211C73C85FB" box="[812,858,625,647]" count="10" pageId="2" pageNumber="253" type="female">10♀♀</specimenCount>
(
<collectionCode id="C5B8C796FFDD5411FC83C211C7C585FA" box="[869,931,627,646]" pageId="2" pageNumber="253">CFDD</collectionCode>
),
<collectingRegion id="616D91B1FFDD5411FC54C211C79585FA" box="[946,1011,627,646]" country="Georgia" name="Imereti" pageId="2" pageNumber="253">Imereti</collectingRegion>
, Zeda Gordi vic, right bank of
<location id="A6760988FFDD5411FAE7C211C70D85E2" LSID="urn:lsid:plazi:treatment:2B00EE45FFDD5415FF0BC1E2C6E78467:A6760988FFDD5411FAE7C211C70D85E2" name="river Tskhenistskali" pageId="2" pageNumber="253" stateProvince="Imereti">river Tskhenistskali</location>
,
<quantity id="6451F2B6FFDD5411FC9CC2E9C7A585E3" box="[890,963,651,671]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="253" unit="m" value="1000.0">
<elevation id="2884B860FFDD5411FC9CC2E9C7A585E3" box="[890,963,651,671]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="253" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="D7177993FFDD5411FC37C2E9C04185E2" box="[977,1063,651,670]" pageId="2" pageNumber="253" value="2017-04-15" valueMax="2017-06-17" valueMin="2017-04-15">15.IV–17.</date>
</materialsCitation>
<collectionCode id="C5B8C796FFDD5411FBC3C2E9C05885E2" box="[1061,1086,651,670]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="2" pageNumber="253">VI</collectionCode>
.2017,
<collectorName id="0E5C3A85FFDD5411FB66C2EEC09185E2" box="[1152,1271,651,671]" pageId="2" pageNumber="253">D. Fominykh</collectorName>
leg.;
<materialsCitation id="13C1550EFFDD5410FAD5C2E9C68A8632" collectionCode="CFDD, CTAY, CZAM, MPSU" collectorName="Zeda-Gordi, NE" elevation="980" lastPageId="3" lastPageNumber="254" location="Mt. Gulkuna" pageId="2" pageNumber="253" specimenCount="187" specimenCount-female="106" specimenCount-male="81" stateProvince="Imereti">
<specimenCount id="B5AF94DAFFDD5411FAD5C2E9C11785E3" box="[1331,1393,649,671]" count="81" pageId="2" pageNumber="253" type="male">81♂♂</specimenCount>
,
<specimenCount id="B5AF94DAFFDD5411FCCAC2C1C70385CB" box="[812,869,673,695]" count="106" pageId="2" pageNumber="253" type="female">106♀♀</specimenCount>
(
<collectionCode id="C5B8C796FFDD5411FC96C2C1C7CB85CA" box="[880,941,675,694]" pageId="2" pageNumber="253">CFDD</collectionCode>
,
<collectionCode id="C5B8C796FFDD5411FC50C2C1C79585CA" box="[950,1011,675,694]" pageId="2" pageNumber="253">CTAY</collectionCode>
,
<collectionCode id="C5B8C796FFDD5411FC1AC2C1C05A85CA" box="[1020,1084,675,694]" pageId="2" pageNumber="253">CZAM</collectionCode>
,
<collectionCode id="C5B8C796FFDD5411FBA0C2C1C0E285CA" box="[1094,1156,675,694]" country="Thailand" httpUri="http://grbio.org/cool/w22j-vay4" name="Department of Microbiology, Songkla University" pageId="2" pageNumber="253">MPSU</collectionCode>
),
<collectingRegion id="616D91B1FFDD5411FB75C2C1C0B585CA" box="[1171,1235,675,694]" country="Georgia" name="Imereti" pageId="2" pageNumber="253">Imereti</collectingRegion>
, S slopes of Askhi plateau near
<collectorName id="0E5C3A85FFDC5410FEC8C159C5DA8633" box="[302,444,315,335]" pageId="3" pageNumber="254">Zeda-Gordi, NE</collectorName>
slope of
<location id="A6760988FFDC5410FDF6C15EC6E48632" LSID="urn:lsid:plazi:treatment:2B00EE45FFDD5415FF0BC1E2C6E78467:A6760988FFDC5410FDF6C15EC6E48632" box="[528,642,315,335]" name="Mt. Gulkuna" pageId="3" pageNumber="254" stateProvince="Imereti">Mt. Gulkuna</location>
,
<quantity id="6451F2B6FFDC5410FD6BC159C6A18633" box="[653,711,315,335]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="3" pageNumber="254" unit="m" value="980.0">
<elevation id="2884B860FFDC5410FD6BC159C6A18633" box="[653,711,315,335]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="3" pageNumber="254" unit="m" value="980.0">980 m</elevation>
</quantity>
,
<date id="D7177993FFDC5410FD37C159C68A8632" box="[721,748,315,334]" pageId="3" pageNumber="254" value="2019-05-20" valueMax="2019-07-14" valueMin="2019-05-20">20.</date>
</materialsCitation>
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– 14.VII.2019,
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&amp;
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leg.;
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,
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(
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,
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,
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,
<collectionCode id="C5B8C796FFDC5410FEA9C109C5EB8602" box="[335,397,363,382]" country="Thailand" httpUri="http://grbio.org/cool/w22j-vay4" name="Department of Microbiology, Songkla University" pageId="3" pageNumber="254">MPSU</collectionCode>
), Samegrelo, Doberazeni vic., left bank of
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,
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</quantity>
,
<date id="D7177993FFDC5410FE9AC1E1C5B586EA" box="[380,467,387,406]" pageId="3" pageNumber="254" value="2017-04-16" valueMax="2017-05-31" valueMin="2017-04-16">16.IV–31.</date>
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<collectionCode id="C5B8C796FFDC5410FE34C1E1C58586EA" box="[466,483,387,406]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="254" type="Museum">V</collectionCode>
.2017,
<collectorName id="0E5C3A85FFDC5410FDF9C1E6C6F486EA" box="[543,658,387,407]" pageId="3" pageNumber="254">D. Fominykh</collectorName>
leg.;
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<specimenCount id="B5AF94DAFFDC5410FD22C1E1C76786EB" box="[708,769,385,407]" count="58" pageId="3" pageNumber="254" type="male">58♂♂</specimenCount>
,
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(
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,
<collectionCode id="C5B8C796FFDC5410FEA2C1F9C5E486D2" box="[324,386,411,430]" pageId="3" pageNumber="254">CTAY</collectionCode>
), Samegrelo, left bank of
<collectingMunicipality id="4372C529FFDC5410FD9FC1F9C68C86D2" box="[633,746,411,430]" pageId="3" pageNumber="254">river Shakvi</collectingMunicipality>
, N slopes of
<location id="A6760988FFDC5410FEF5C1D1C5DA86BB" LSID="urn:lsid:plazi:treatment:2B00EE45FFDD5415FF0BC1E2C6E78467:A6760988FFDC5410FEF5C1D1C5DA86BB" box="[275,444,435,455]" municipality="river Shakvi" name="Ofitsari Mt. Range" pageId="3" pageNumber="254" stateProvince="Imereti">Ofitsari Mt. Range</location>
,
<quantity id="6451F2B6FFDC5410FE21C1D1C66A86BB" box="[455,524,435,455]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="254" unit="m" value="1500.0">
<elevation id="2884B860FFDC5410FE21C1D1C66A86BB" box="[455,524,435,455]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="254" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="D7177993FFDC5410FDF1C1D1C65486BA" box="[535,562,435,454]" pageId="3" pageNumber="254" value="2017-05-29" valueMax="2017-06-17" valueMin="2017-05-29">29.</date>
</materialsCitation>
<collectionCode id="C5B8C796FFDC5410FDD7C1D1C62486BA" box="[561,578,435,454]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="254" type="Museum">V</collectionCode>
–17.
<collectionCode id="C5B8C796FFDC5410FD83C1D1C61886BA" box="[613,638,435,454]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="254">VI</collectionCode>
.2017,
<collectorName id="0E5C3A85FFDC5410FD5DC1D6C48886A2" pageId="3" pageNumber="254">D. Fominykh</collectorName>
leg.;
<materialsCitation id="13C1550EFFDC5410FEF8C1A9C68C868A" collectionCode="CFDD, CTAY" elevation="1900" location="Ofitsari Mt. Range" municipality="river Shakvi" pageId="3" pageNumber="254" specimenCount="307" specimenCount-female="217" specimenCount-male="90" stateProvince="Imereti">
<specimenCount id="B5AF94DAFFDC5410FEF8C1A9C53A86A3" box="[286,348,457,479]" count="90" pageId="3" pageNumber="254" type="male">90♂♂</specimenCount>
,
<specimenCount id="B5AF94DAFFDC5410FE81C1A9C5F986A3" box="[359,415,457,479]" count="217" pageId="3" pageNumber="254" type="female">217♀♀</specimenCount>
(
<collectionCode id="C5B8C796FFDC5410FE4CC1A9C58186A2" box="[426,487,459,478]" pageId="3" pageNumber="254">CFDD</collectionCode>
,
<collectionCode id="C5B8C796FFDC5410FE16C1A9C64886A2" box="[496,558,459,478]" pageId="3" pageNumber="254">CTAY</collectionCode>
), Samegrelo, left bank of
<collectingMunicipality id="4372C529FFDC5410FF5BC181C556868A" box="[189,304,483,502]" pageId="3" pageNumber="254">river Shakvi</collectingMunicipality>
, N slopes of
<location id="A6760988FFDC5410FE5EC181C60C868B" LSID="urn:lsid:plazi:treatment:2B00EE45FFDD5415FF0BC1E2C6E78467:A6760988FFDC5410FE5EC181C60C868B" box="[440,618,483,503]" municipality="river Shakvi" name="Ofitsari Mt. Range" pageId="3" pageNumber="254" stateProvince="Imereti">Ofitsari Mt. Range</location>
,
<quantity id="6451F2B6FFDC5410FD9FC181C6A4868B" box="[633,706,483,503]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="254" unit="m" value="1900.0">
<elevation id="2884B860FFDC5410FD9FC181C6A4868B" box="[633,706,483,503]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="254" unit="m" value="1900.0">1900 m</elevation>
</quantity>
,
<date id="D7177993FFDC5410FD37C181C68C868A" box="[721,746,483,502]" pageId="3" pageNumber="254" value="2017-05-31" valueMax="2017-06-17" valueMin="2017-05-31">31.</date>
</materialsCitation>
<collectionCode id="C5B8C796FFDC5410FD0FC181C69C868A" box="[745,762,483,502]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="254" type="Museum">V</collectionCode>
– 17.
<collectionCode id="C5B8C796FFDC5410FF32C199C48B8572" box="[212,237,507,526]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="254">VI</collectionCode>
.2017,
<collectorName id="0E5C3A85FFDC5410FEC8C19EC5C28572" box="[302,420,507,527]" pageId="3" pageNumber="254">D. Fominykh</collectorName>
leg.;
<materialsCitation id="13C1550EFFDC5410FE38C199C5998542" collectionCode="CFDD, CTAY, CZAM, MPSU" elevation="1900" location="Ofitsari Mt. Range" municipality="river Shakvi" pageId="3" pageNumber="254" specimenCount="419" specimenCount-female="254" specimenCount-male="165" stateProvince="Imereti">
<specimenCount id="B5AF94DAFFDC5410FE38C199C6408573" box="[478,550,505,527]" count="165" pageId="3" pageNumber="254" type="male">165♂♂</specimenCount>
,
<specimenCount id="B5AF94DAFFDC5410FDD3C199C6088573" box="[565,622,505,527]" count="254" pageId="3" pageNumber="254" type="female">254♀♀</specimenCount>
(
<collectionCode id="C5B8C796FFDC5410FD9BC199C6DC8572" box="[637,698,507,526]" pageId="3" pageNumber="254">CFDD</collectionCode>
,
<collectionCode id="C5B8C796FFDC5410FD21C199C7628572" box="[711,772,507,526]" pageId="3" pageNumber="254">CTAY</collectionCode>
,
<collectionCode id="C5B8C796FFDC5410FF5BC271C49B855A" box="[189,253,531,550]" pageId="3" pageNumber="254">CZAM</collectionCode>
,
<collectionCode id="C5B8C796FFDC5410FEE3C271C525855A" box="[261,323,531,550]" country="Thailand" httpUri="http://grbio.org/cool/w22j-vay4" name="Department of Microbiology, Songkla University" pageId="3" pageNumber="254">MPSU</collectionCode>
), Samegrelo, upper bank of
<collectingMunicipality id="4372C529FFDC5410FDD4C271C6F8855A" box="[562,670,531,550]" pageId="3" pageNumber="254">river Shakvi</collectingMunicipality>
, N slopes of
<location id="A6760988FFDC5410FF5BC249C50D8543" LSID="urn:lsid:plazi:treatment:2B00EE45FFDD5415FF0BC1E2C6E78467:A6760988FFDC5410FF5BC249C50D8543" box="[189,363,555,575]" municipality="river Shakvi" name="Ofitsari Mt. Range" pageId="3" pageNumber="254" stateProvince="Imereti">Ofitsari Mt. Range</location>
,
<quantity id="6451F2B6FFDC5410FE9EC249C5A78543" box="[376,449,555,575]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="254" unit="m" value="1900.0">
<elevation id="2884B860FFDC5410FE9EC249C5A78543" box="[376,449,555,575]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="254" unit="m" value="1900.0">1900 m</elevation>
</quantity>
,
<date id="D7177993FFDC5410FE28C249C5998542" box="[462,511,555,574]" pageId="3" pageNumber="254" value="2018-06-02" valueMax="2018-06-21" valueMin="2018-06-02">2–21.</date>
</materialsCitation>
<collectionCode id="C5B8C796FFDC5410FE1BC249C6708542" box="[509,534,555,574]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="254">VI</collectionCode>
.2018,
<collectorName id="0E5C3A85FFDC5410FDB0C24EC6AB8542" box="[598,717,555,575]" pageId="3" pageNumber="254">D. Fominykh</collectorName>
&amp;
<collectorName id="0E5C3A85FFDC5410FD15C24EC562852A" pageId="3" pageNumber="254">B. Dubinin</collectorName>
leg.;
<materialsCitation id="13C1550EFFDC5410FED4C221C4B085FA" collectionCode="CFDD, CTAY" collectorName="Samegrelo, N" elevation="600" location="river Tekhuri" municipality="Mt. Ogedzh" pageId="3" pageNumber="254" specimenCount="73" specimenCount-female="41" specimenCount-male="32" stateProvince="Imereti">
<specimenCount id="B5AF94DAFFDC5410FED4C221C509852B" box="[306,367,577,599]" count="32" pageId="3" pageNumber="254" type="male">32♂♂</specimenCount>
,
<specimenCount id="B5AF94DAFFDC5410FE9FC221C5C1852B" box="[377,423,577,599]" count="41" pageId="3" pageNumber="254" type="female">41♀♀</specimenCount>
(
<collectionCode id="C5B8C796FFDC5410FE57C221C588852A" box="[433,494,579,598]" pageId="3" pageNumber="254">CFDD</collectionCode>
,
<collectionCode id="C5B8C796FFDC5410FE13C221C655852A" box="[501,563,579,598]" pageId="3" pageNumber="254">CTAY</collectionCode>
),
<collectorName id="0E5C3A85FFDC5410FDA7C221C6D0852B" box="[577,694,579,599]" pageId="3" pageNumber="254">Samegrelo, N</collectorName>
slopes of
<collectingMunicipality id="4372C529FFDC5410FF5BC23EC5418512" box="[189,295,603,623]" pageId="3" pageNumber="254">Mt. Ogedzh</collectingMunicipality>
near Doberazeni, left bank of
<location id="A6760988FFDC5410FDA5C239C6DA8512" LSID="urn:lsid:plazi:treatment:2B00EE45FFDD5415FF0BC1E2C6E78467:A6760988FFDC5410FDA5C239C6DA8512" box="[579,700,603,622]" municipality="Mt. Ogedzh" name="river Tekhuri" pageId="3" pageNumber="254" stateProvince="Imereti">river Tekhuri</location>
,
<quantity id="6451F2B6FFDC5410FD21C239C7658513" box="[711,771,603,623]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="254" unit="m" value="600.0">
<elevation id="2884B860FFDC5410FD21C239C7658513" box="[711,771,603,623]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="254" unit="m" value="600.0">600 m</elevation>
</quantity>
,
<date id="D7177993FFDC5410FF5BC211C4B085FA" box="[189,214,627,646]" pageId="3" pageNumber="254" value="2019-05-21" valueMax="2019-07-15" valueMin="2019-05-21">21.</date>
</materialsCitation>
<collectionCode id="C5B8C796FFDC5410FF33C211C48085FA" box="[213,230,627,646]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="254" type="Museum">V</collectionCode>
–15.VII.2019,
<collectorName id="0E5C3A85FFDC5410FE8EC216C5AF85FA" box="[360,457,627,647]" pageId="3" pageNumber="254">B. Dubinin</collectorName>
&amp;
<collectorName id="0E5C3A85FFDC5410FE01C216C63485FA" box="[487,594,627,647]" pageId="3" pageNumber="254">D. Safronov</collectorName>
leg.
</paragraph>
</subSubSection>
<caption id="F7D60FDBFFDD5411FF04C7CCC7118F71" ID-DOI="http://doi.org/10.5281/zenodo.13164017" ID-Zenodo-Dep="13164017" httpUri="https://zenodo.org/record/13164017/files/figure.png" pageId="2" pageNumber="253" startId="2.[226,263,1966,1985]" targetBox="[190,1396,720,1946]" targetPageId="2" targetType="figure">
<paragraph id="A3165F53FFDD5411FF04C7CCC6C880A5" blockId="2.[189,1399,1965,2061]" pageId="2" pageNumber="253">
Figs 2–9. Dorsal habitus of
<taxonomicName id="64A924D0FFDD5411FE3EC7CCC68980BD" authorityName="Fominykh &amp; Zamotajlov &amp; Titarenko" authorityYear="2019" box="[472,751,1966,1985]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="253" phylum="Arthropoda" rank="subSpecies" species="martviliensis" status="subsp. nov." subSpecies="mzekalae">
<emphasis id="91DD8341FFDD5411FE3EC7CCC68980BD" box="[472,751,1966,1985]" italics="true" pageId="2" pageNumber="253">Carabus martviliensis mzekalae</emphasis>
</taxonomicName>
<emphasis id="91DD8341FFDD5411FD10C7CCC74F80BD" bold="true" box="[758,809,1966,1985]" pageId="2" pageNumber="253">
<taxonomicNameLabel id="8AEE3E3AFFDD5411FD10C7CCC74F80BD" box="[758,809,1966,1985]" pageId="2" pageNumber="253" rank="subSpecies">ssp.n.</taxonomicNameLabel>
</emphasis>
: 2–5 — Melouri env., 475 m; 6–9 — Dzedzileti env., 770 m; 2–3, 6–7 — males; 4–5, 8–9 — females. Scale bar: 10.0 mm.
</paragraph>
<paragraph id="A3165F53FFDD5411FF04C780C7118F71" blockId="2.[189,1399,1965,2061]" pageId="2" pageNumber="253">
Рис. 2–9. Габитус сверху
<taxonomicName id="64A924D0FFDD5411FE36C783C6838088" authorityName="Fominykh &amp; Zamotajlov &amp; Titarenko" authorityYear="2019" box="[464,741,2017,2036]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="253" phylum="Arthropoda" rank="subSpecies" species="martviliensis" status="subsp. nov." subSpecies="mzekalae">
<emphasis id="91DD8341FFDD5411FE36C783C6838088" box="[464,741,2017,2036]" italics="true" pageId="2" pageNumber="253">Carabus martviliensis mzekalae</emphasis>
</taxonomicName>
<emphasis id="91DD8341FFDD5411FD0AC783C7798088" bold="true" box="[748,799,2017,2036]" pageId="2" pageNumber="253">
<taxonomicNameLabel id="8AEE3E3AFFDD5411FD0AC783C7798088" box="[748,799,2017,2036]" pageId="2" pageNumber="253" rank="subSpecies">ssp.n.</taxonomicNameLabel>
</emphasis>
: 2–5 — окр. Мелоури, 475 м над ур. м.; 6–9 — окр. ДЗедЗилети, 770 м над ур. м.; 2–3, 6–7 — самцы; 4–5, 8–9 — самки. МасШтаб: 10,0 мм.
</paragraph>
</caption>
<subSubSection id="EBB30CD8FFDC5417FF04C2E9C0088520" lastPageId="4" lastPageNumber="255" pageId="3" pageNumber="254" type="description">
<paragraph id="A3165F53FFDC5417FF04C2E9C49F868D" blockId="3.[189,775,315,700]" lastBlockId="4.[189,776,315,924]" lastPageId="4" lastPageNumber="255" pageId="3" pageNumber="254">
DESCRIPTION. Big sized subspecies (
<figureCitation id="3B9243D6FFDC5410FD88C2E9C6AE85DC" box="[622,712,651,673]" captionStart="Figs 2–9" captionStartId="2.[226,263,1966,1985]" captionTargetBox="[190,1396,720,1946]" captionTargetId="figure-361@2.[189,1399,717,1946]" captionTargetPageId="2" captionText="Figs 2–9. Dorsal habitus of Carabus martviliensis mzekalae ssp.n.: 2–5 — Melouri env., 475 m; 6–9 — Dzedzileti env., 770 m; 2–3, 6–7 — males; 4–5, 8–9 — females. Scale bar: 10.0 mm. Рис. 2–9. Габитус сверху Carabus martviliensis mzekalae ssp.n.: 2–5 — окр. Мелоури, 475 м над ур. м.; 6–9 — окр. ДЗедЗилети, 770 м над ур. м.; 2–3, 6–7 — самцы; 4–5, 8–9 — самки. МасШтаб: 10,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13164017" httpUri="https://zenodo.org/record/13164017/files/figure.png" pageId="3" pageNumber="254">Figs 2–9</figureCitation>
).GBL = 38.0–
<quantity id="6451F2B6FFDC5410FEECC2C4C50285C7" box="[266,356,678,700]" metricMagnitude="-2" metricUnit="m" metricValue="4.85" pageId="3" pageNumber="254" unit="mm" value="48.5">48.5 mm</quantity>
, males 38.0–
<quantity id="6451F2B6FFDC5410FE0FC2C4C62285C7" box="[489,580,678,699]" metricMagnitude="-2" metricUnit="m" metricValue="4.4399999999999995" pageId="3" pageNumber="254" unit="mm" value="44.4">44.4 mm</quantity>
, females
<quantity id="6451F2B6FFDC5410FD42C2C4C732862D" box="[676,852,316,700]" metricMagnitude="-2" metricUnit="m" metricValue="4.42" metricValueMax="4.85" metricValueMin="3.9899999999999998" pageId="3" pageNumber="254" unit="mm" value="44.2" valueMax="48.5" valueMin="39.9">39.9–48.5 mm</quantity>
, SBL = 36.8–45.0 mm in both sexes. Underside black, dorsum with metallic luster, both in males and females, violet, blue, less frequently goldish or green, in some individuals with clear iridescence, females somewhat less bright; head mostly black dorsally, sometimes with metallic luster basally; legs, antennae, palpi, and mandibles black. Head thick, somewhat inflated, surface with coarse rugosity, HL = 6.0–
<quantity id="6451F2B6FFDC5410FC8AC194C7D08577" box="[876,950,502,524]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="254" unit="mm" value="7.5">7.5 mm</quantity>
in males,
<quantity id="6451F2B6FFDC5410FBFDC194C0F78577" box="[1051,1169,502,524]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.5" metricValueMin="6.5" pageId="3" pageNumber="254" unit="mm" value="7.5" valueMax="8.5" valueMin="6.5">6.5–8.5 mm</quantity>
in females, HW =
<quantity id="6451F2B6FFDC5410FAADC194C7E5855A" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.8" metricValueMin="6.2" pageId="3" pageNumber="254" unit="mm" value="8.5" valueMax="10.8" valueMin="6.2">6.2– 10.8 mm</quantity>
in males, 7.0–
<quantity id="6451F2B6FFDC5410FBF4C272C03A855A" box="[1042,1116,528,550]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="3" pageNumber="254" unit="mm" value="8.6">8.6 mm</quantity>
in females. Pronotum nearly parallel-sided, with coarse rugosity basally, this furrows becoming less pronounced at sides; PW =
<quantity id="6451F2B6FFDC5410FB3EC224C13A8527" box="[1240,1372,582,604]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" metricValueMax="11.8" metricValueMin="6.4" pageId="3" pageNumber="254" unit="mm" value="9.1" valueMax="11.8" valueMin="6.4">6.4–11.8 mm</quantity>
in males,
<quantity id="6451F2B6FFDC5410FC94C203C792850A" box="[882,1012,609,630]" metricMagnitude="-2" metricUnit="m" metricValue="1.1800000000000002" metricValueMax="1.3800000000000001" metricValueMin="0.9800000000000001" pageId="3" pageNumber="254" unit="mm" value="11.8" valueMax="13.8" valueMin="9.8">9.8–13.8 mm</quantity>
in females, PB =
<quantity id="6451F2B6FFDC5410FB44C202C17E850A" box="[1186,1304,608,630]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" metricValueMax="8.9" metricValueMin="6.5" pageId="3" pageNumber="254" unit="mm" value="7.7" valueMax="8.9" valueMin="6.5">6.5–8.9 mm</quantity>
in males, 7.0–
<quantity id="6451F2B6FFDC5410FCBEC219C7C485ED" box="[856,930,635,657]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="3" pageNumber="254" unit="mm" value="9.9">9.9 mm</quantity>
in females, PL =
<quantity id="6451F2B6FFDC5410FBB7C219C0A185ED" box="[1105,1223,635,657]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.2" metricValueMin="5.8" pageId="3" pageNumber="254" unit="mm" value="6.5" valueMax="7.2" valueMin="5.8">5.8–7.2 mm</quantity>
in males, 6.4–8.0 mm in females, PW/HW =
<quantity id="6451F2B6FFDC5410FBA0C2F4C0CA85D0" box="[1094,1196,662,684]" metricMagnitude="-2" metricUnit="m" metricValue="2.794" metricValueMax="4.064" metricValueMin="1.524" pageId="3" pageNumber="254" unit="in" value="1.1" valueMax="1.6" valueMin="0.6">0.6–1.6 in</quantity>
both sexes, PW/PL =
<quantity id="6451F2B6FFDC5410FCDCC2D3C7C785BB" box="[826,929,689,711]" metricMagnitude="-2" metricUnit="m" metricValue="3.429" metricValueMax="4.572" metricValueMin="2.286" pageId="3" pageNumber="254" unit="in" value="1.35" valueMax="1.8" valueMin="0.9">0.9–1.8 in</quantity>
both sexes, PW/PB =
<quantity id="6451F2B6FFDC5410FB75C2D3C09C85BB" box="[1171,1274,689,711]" metricMagnitude="-2" metricUnit="m" metricValue="2.794" metricValueMax="3.5559999999999996" metricValueMin="2.032" pageId="3" pageNumber="254" unit="in" value="1.1" valueMax="1.4" valueMin="0.8">0.8–1.4 in</quantity>
both sexes; hind angles faintly protruding posteriorly. Legs moderately long. Elytra oblong-ovate, convex in males, somewhat flattened in females, with prominent sculpture in both sexes, EW/PW =
<quantity id="6451F2B6FFDB5417FEC9C1E9C5F386DD" box="[303,405,395,417]" metricMagnitude="-2" metricUnit="m" metricValue="4.571999999999999" metricValueMax="5.842" metricValueMin="3.302" pageId="4" pageNumber="255" unit="in" value="1.7999999999999998" valueMax="2.3" valueMin="1.3">1.3–2.3 in</quantity>
both sexes; EL =
<quantity id="6451F2B6FFDB5417FDBEC1E9C68C86DD" box="[600,746,395,417]" metricMagnitude="-2" metricUnit="m" metricValue="2.395" metricValueMax="2.58" metricValueMin="2.21" pageId="4" pageNumber="255" unit="mm" value="23.95" valueMax="25.8" valueMin="22.1">22.1–25.8 mm</quantity>
in males,
<quantity id="6451F2B6FFDB5417FEE3C1C4C5F086C7" box="[261,406,422,443]" metricMagnitude="-2" metricUnit="m" metricValue="2.545" metricValueMax="2.78" metricValueMin="2.31" pageId="4" pageNumber="255" unit="mm" value="25.45" valueMax="27.8" valueMin="23.1">23.1–27.8 mm</quantity>
in females, EW =
<quantity id="6451F2B6FFDB5417FDBCC1C4C68D86C7" box="[602,747,422,444]" metricMagnitude="-2" metricUnit="m" metricValue="1.485" metricValueMax="1.66" metricValueMin="1.31" pageId="4" pageNumber="255" unit="mm" value="14.85" valueMax="16.6" valueMin="13.1">13.1–16.6 mm</quantity>
in males,
<quantity id="6451F2B6FFDB5417FEE5C1A2C5F486AA" box="[259,402,448,470]" metricMagnitude="-2" metricUnit="m" metricValue="1.6149999999999998" metricValueMax="1.75" metricValueMin="1.48" pageId="4" pageNumber="255" unit="mm" value="16.15" valueMax="17.5" valueMin="14.8">14.8–17.5 mm</quantity>
in females, EL/EW =
<quantity id="6451F2B6FFDB5417FD97C1A2C6B286AB" box="[625,724,448,471]" metricMagnitude="-2" metricUnit="m" metricValue="4.064" metricValueMax="4.318" metricValueMin="3.81" pageId="4" pageNumber="255" unit="in" value="1.6" valueMax="1.7" valueMin="1.5">1.5–1.7 in</quantity>
both sexes.
</paragraph>
<caption id="F7D60FDBFFDC5410FF04C7CCC0128F71" ID-DOI="http://doi.org/10.5281/zenodo.13164019" ID-Zenodo-Dep="13164019" httpUri="https://zenodo.org/record/13164019/files/figure.png" pageId="3" pageNumber="254" startId="3.[226,263,1966,1985]" targetBox="[190,1395,736,1960]" targetPageId="3" targetType="figure">
<paragraph id="A3165F53FFDC5410FF04C7CCC71480A5" blockId="3.[189,1398,1965,2061]" pageId="3" pageNumber="254">
Figs 10–17. Dorsal habitus of
<taxonomicName id="64A924D0FFDC5410FE0FC7CCC77B80BD" authorityName="Retezar et Djavelidze" authorityYear="1992" box="[489,797,1966,1985]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="254" phylum="Arthropoda" rank="subSpecies" species="martviliensis" subSpecies="martviliensis">
<emphasis id="91DD8341FFDC5410FE0FC7CCC77B80BD" box="[489,797,1966,1985]" italics="true" pageId="3" pageNumber="254">Carabus martviliensis martviliensis</emphasis>
</taxonomicName>
: 10–13 — Zeda Gordi env., 980 m; 14–17 — Doberazeni env., 500– 600 m; 10–11, 14–15 — males; 12–13, 16–17 — females. Scale bar 10.0 mm.
</paragraph>
<paragraph id="A3165F53FFDC5410FF04C780C0128F71" blockId="3.[189,1398,1965,2061]" pageId="3" pageNumber="254">
Рис. 10–17. Габитус сверху
<taxonomicName id="64A924D0FFDC5410FE17C783C74A8088" authorityName="Retezar et Djavelidze" authorityYear="1992" box="[497,812,2017,2036]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="254" phylum="Arthropoda" rank="subSpecies" species="martviliensis" subSpecies="martviliensis">
<emphasis id="91DD8341FFDC5410FE17C783C74A8088" box="[497,812,2017,2036]" italics="true" pageId="3" pageNumber="254">Carabus martviliensis martviliensis</emphasis>
</taxonomicName>
: 10–13 — окр. Зеда Горди, 980 м над ур. м.; 14–17 — окр. ДобераЗени, 500–600 м над ур. м.; 10–11, 14–15 — самцы; 12–13, 16–17 — самки. МасШтаб: 10,0 мм.
</paragraph>
</caption>
<paragraph id="A3165F53FFDB5417FF04C194C6A78520" blockId="4.[189,776,315,924]" pageId="4" pageNumber="255">
Aedeagus as in the other
<taxonomicName id="64A924D0FFDB5417FE35C194C65B8577" box="[467,573,502,523]" pageId="4" pageNumber="257" rank="subGenus" subGenus="Neoplectes">
<emphasis id="91DD8341FFDB5417FE35C194C65B8577" box="[467,573,502,523]" italics="true" pageId="4" pageNumber="255">Neoplectes</emphasis>
</taxonomicName>
taxa, strongly inflat- ed medioapically and tapered just before apical lamella (
<figureCitation id="3B9243D6FFDB5417FD3AC273C521853C" captionStart="Figs 18–25" captionStartId="4.[226,263,1869,1888]" captionTargetBox="[201,1384,947,1827]" captionTargetId="figure-585@4.[187,1397,943,1831]" captionTargetPageId="4" captionText="Figs 18–25. Apical portion of aedeagus and fully inflated endophallus preparation of Carabus (Neoplectes) martviliensis, right lateral view on top, left lateral view below: 18, 22 — C. martviliensis mzekalae ssp.n., Melouri env., 475 m; 19, 23 — C. martviliensis mzekalae ssp.n., Dzedzileti env., 770 m; 20, 24 — C. martviliensis martviliensis, Zeda Gordi env., 980 m; 21, 25 — C. martviliensis martviliensis, Doberazeni env., 600 m. Scale bar: 10.0 mm. Рис. 18–25. ВерШина Эдеагуса и препарат полностью вывернутого внутреннего меШка Carabus (Neoplectes) martviliensis, вверху — сбоку справа, вниЗу — сбоку слева: 18, 22 — C. martviliensis mzekalae ssp.n., окр. Мелоури, 475 м над ур.м.; 19, 23 — C. martviliensis mzekalae ssp.n., окр. ДЗедЗилети, 770 м над ур.м.; 20, 24 — C. martviliensis martviliensis, окр. Зеда Горди, 980 м над ур. м.; 21, 25 — C. martviliensis martviliensis, окр. ДобераЗени, 600 м над ур.м. МасШтаб: 10,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13164021" httpUri="https://zenodo.org/record/13164021/files/figure.png" pageId="4" pageNumber="255">Figs 18–19, 22–23</figureCitation>
); endophallus of the complicated shape, with several pronounced structures (lobes or projections).
</paragraph>
<paragraph id="A3165F53FFDB5417FF04C203C76184E0" blockId="4.[189,776,315,924]" pageId="4" pageNumber="255">
DIFFERENTIAL DIAGNOSIS AND REMARKS.A new subspecies reliably differs from the nominotypical one in more ovate and less elongate body shape (
<figureCitation id="3B9243D6FFDB5417FDBBC2F4C6D585D7" box="[605,691,662,684]" captionStart="Figs 2–9" captionStartId="2.[226,263,1966,1985]" captionTargetBox="[190,1396,720,1946]" captionTargetId="figure-361@2.[189,1399,717,1946]" captionTargetPageId="2" captionText="Figs 2–9. Dorsal habitus of Carabus martviliensis mzekalae ssp.n.: 2–5 — Melouri env., 475 m; 6–9 — Dzedzileti env., 770 m; 2–3, 6–7 — males; 4–5, 8–9 — females. Scale bar: 10.0 mm. Рис. 2–9. Габитус сверху Carabus martviliensis mzekalae ssp.n.: 2–5 — окр. Мелоури, 475 м над ур. м.; 6–9 — окр. ДЗедЗилети, 770 м над ур. м.; 2–3, 6–7 — самцы; 4–5, 8–9 — самки. МасШтаб: 10,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13164017" httpUri="https://zenodo.org/record/13164017/files/figure.png" pageId="4" pageNumber="255">Figs 2–9</figureCitation>
<emphasis id="91DD8341FFDB5417FD5DC2F5C6A985D0" box="[699,719,663,684]" italics="true" pageId="4" pageNumber="255">vs</emphasis>
.
<figureCitation id="3B9243D6FFDB5417FD3BC2F4C49A85BA" captionStart="Figs 10–17" captionStartId="3.[226,263,1966,1985]" captionTargetBox="[190,1395,736,1960]" captionTargetId="figure-401@3.[189,1399,733,1963]" captionTargetPageId="3" captionText="Figs 10–17. Dorsal habitus of Carabus martviliensis martviliensis: 10–13 — Zeda Gordi env., 980 m; 14–17 — Doberazeni env., 500– 600 m; 10–11, 14–15 — males; 12–13, 16–17 — females. Scale bar 10.0 mm. Рис. 10–17. Габитус сверху Carabus martviliensis martviliensis: 10–13 — окр. Зеда Горди, 980 м над ур. м.; 14–17 — окр. ДобераЗени, 500–600 м над ур. м.; 10–11, 14–15 — самцы; 12–13, 16–17 — самки. МасШтаб: 10,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13164019" httpUri="https://zenodo.org/record/13164019/files/figure.png" pageId="4" pageNumber="255">Figs 10–17</figureCitation>
), hind angles of pronotum less protruding posteriorly. It is also easily distinguishable by dorsal range of colours in all known populations. The nominotypical subspecies possesses violet or lilac colour with characteristic olive tint, green or bronze colours are observed very rare, ratio of green or bronze to standard violet individuals is about 1 to 50–100 ones, according to our calculations, this proportion varying in different populations. A new subspecies usually possesses blue, greenish-blue, or dark violet colour, sometimes bronze
</paragraph>
<paragraph id="A3165F53FFDB5417FF2CC77FC670804B" blockId="4.[202,534,1821,1847]" box="[202,534,1821,1847]" pageId="4" pageNumber="255">
<emphasis id="91DD8341FFDB5417FF2CC77FC670804B" bold="true" box="[202,534,1821,1847]" pageId="4" pageNumber="255">22 23</emphasis>
</paragraph>
<paragraph id="A3165F53FFDB5417FCCAC15EC0438617" blockId="4.[812,1399,315,924]" pageId="4" pageNumber="255">or purely green individuals are also observed, though their ratio is about 1 to 50 too.</paragraph>
<paragraph id="A3165F53FFDB5417FCB4C113C142868D" blockId="4.[812,1399,315,924]" pageId="4" pageNumber="255">
Apical lamella of aedeagus is more narrow and pointed in comparison with
<taxonomicName id="64A924D0FFDB5417FC30C1EEC09286DC" authorityName="Retezar et Djavelidze" authorityYear="1992" box="[982,1268,395,417]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="subSpecies" species="martviliensis" subSpecies="martviliensis">
<emphasis id="91DD8341FFDB5417FC30C1EEC09286DC" box="[982,1268,395,417]" italics="true" pageId="4" pageNumber="255">C. martviliensis martviliensis</emphasis>
</taxonomicName>
(
<figureCitation id="3B9243D6FFDB5417FAE4C1E9C70F86C7" captionStart="Figs 18–25" captionStartId="4.[226,263,1869,1888]" captionTargetBox="[201,1384,947,1827]" captionTargetId="figure-585@4.[187,1397,943,1831]" captionTargetPageId="4" captionText="Figs 18–25. Apical portion of aedeagus and fully inflated endophallus preparation of Carabus (Neoplectes) martviliensis, right lateral view on top, left lateral view below: 18, 22 — C. martviliensis mzekalae ssp.n., Melouri env., 475 m; 19, 23 — C. martviliensis mzekalae ssp.n., Dzedzileti env., 770 m; 20, 24 — C. martviliensis martviliensis, Zeda Gordi env., 980 m; 21, 25 — C. martviliensis martviliensis, Doberazeni env., 600 m. Scale bar: 10.0 mm. Рис. 18–25. ВерШина Эдеагуса и препарат полностью вывернутого внутреннего меШка Carabus (Neoplectes) martviliensis, вверху — сбоку справа, вниЗу — сбоку слева: 18, 22 — C. martviliensis mzekalae ssp.n., окр. Мелоури, 475 м над ур.м.; 19, 23 — C. martviliensis mzekalae ssp.n., окр. ДЗедЗилети, 770 м над ур.м.; 20, 24 — C. martviliensis martviliensis, окр. Зеда Горди, 980 м над ур. м.; 21, 25 — C. martviliensis martviliensis, окр. ДобераЗени, 600 м над ур.м. МасШтаб: 10,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13164021" httpUri="https://zenodo.org/record/13164021/files/figure.png" pageId="4" pageNumber="255">Figs 18–19, 22–23</figureCitation>
<emphasis id="91DD8341FFDB5417FC93C1C5C7EF86C0" box="[885,905,423,444]" italics="true" pageId="4" pageNumber="255">vs</emphasis>
. 20–21, 24–25); endophallus with some minor differences in its general view too, it is less bent towards the tube of aedeagus, ventroapical lobe less developed.
</paragraph>
<paragraph id="A3165F53FFDB5417FCB4C194C0088520" blockId="4.[812,1399,315,924]" pageId="4" pageNumber="255">
Results of the discriminant analysis (
<figureCitation id="3B9243D6FFDB5417FB29C194C17C8570" box="[1231,1306,502,524]" captionStart="Fig" captionStartId="5.[226,257,1120,1139]" captionTargetBox="[280,1321,332,1094]" captionTargetId="figure-205@5.[270,1329,314,1108]" captionTargetPageId="5" captionText="Fig. 26. Distribution of morphometric characters of Carabus (Neoplectes) martviliensis subspecies constructed using discriminant analysis based on 9 parameters (see text for measurements details). Рис. 26. Распределение морфометрических характеристик подвидов Carabus (Neoplectes) martviliensis, построенное с испольЗованием дискриминантного аналиЗа, основанного на 9 параметрах (описание промеров см. текст)." figureDoi="http://doi.org/10.5281/zenodo.13164023" httpUri="https://zenodo.org/record/13164023/files/figure.png" pageId="4" pageNumber="255">Fig. 26</figureCitation>
) showed consistent differences in distribution of the studied features of
<taxonomicName id="64A924D0FFDB5417FCAEC24EC023853C" authorityName="Fominykh &amp; Zamotajlov &amp; Titarenko" authorityYear="2019" box="[840,1093,555,577]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="subSpecies" species="martviliensis" status="ssp.n." subSpecies="mzekalae">
<emphasis id="91DD8341FFDB5417FCAEC24EC023853C" box="[840,1093,555,577]" italics="true" pageId="4" pageNumber="255">C. martviliensis mzekalae</emphasis>
</taxonomicName>
<emphasis id="91DD8341FFDB5417FBABC249C0E1853C" bold="true" box="[1101,1159,555,576]" pageId="4" pageNumber="255">
<taxonomicNameLabel id="8AEE3E3AFFDB5417FBABC249C0E1853C" box="[1101,1159,555,576]" pageId="4" pageNumber="255" rank="subSpecies">ssp.n.</taxonomicNameLabel>
</emphasis>
from the nominotypical subspecies in different localities.
</paragraph>
</subSubSection>
<subSubSection id="EBB30CD8FFDB5417FCB4C203C0E6804B" pageId="4" pageNumber="255" type="distribution">
<paragraph id="A3165F53FFDB5417FCB4C203C11084E0" blockId="4.[812,1399,315,924]" pageId="4" pageNumber="255">
DISTRIBUTION AND HABITAT. A new subspecies is known from Tskhaltubo environs on the left bank of river Tskhenistskali at
<quantity id="6451F2B6FFDB5417FC3FC2F4C02E85D7" box="[985,1096,662,684]" metricMagnitude="2" metricUnit="m" metricValue="6.45" metricValueMax="7.7" metricValueMin="5.2" pageId="4" pageNumber="255" unit="m" value="645.0" valueMax="770.0" valueMin="520.0">520–770 m</quantity>
(
<figureCitation id="3B9243D6FFDB5417FBB1C2F4C0F785D7" box="[1111,1169,662,684]" captionStart="Fig" captionStartId="1.[226,257,1846,1865]" captionTargetBox="[272,1331,971,1827]" captionTargetId="figure-623@1.[272,1331,971,1827]" captionTargetPageId="1" captionText="Fig. 1. Studied localities of Carabus (Neoplectes) martviliensis in Central Georgia: 1–8 — C. martviliensis martviliensis; 9–11 — C. martviliensis mzekalae ssp.n.; 1 — Ofitsari Mt. Range, 1900 m; 2 — Ofitsari Mt. Range, 1500 m; 3 — Doberazeni env., 600 m; 4 — Doberazeni env., 500 m; 5 — Mt. Muchera, 1009 m; 6 — Zeda Gordi env., 980 m; 7 — Kinchkha-Perdi env., 816 m; 8 — Satziskvilo env., 1000 m; 9 — Dzedzileti env., 520–620 m; 10 — Dzedzileti env., 770 m; 11 — Melouri env., 475 m. Рис. 1. ИЗученное распространение Carabus (Neoplectes) martviliensis в центральной части ГруЗии: 1–8 — C. martviliensis martviliensis; 9–11 — C. martviliensis mzekalae ssp.n.; 1 — хр. Офицари, 1900 м над ур. м.; 2 — хр. Офицари, 1500 м над ур. м.; 3 — окр. ДобераЗени, 600 м над ур. м.; 4 — окр. ДобераЗени, 500 м над ур. м.; 5 — г. Мучера, 1009 м над ур. м.; 6 — окр. Зеда Горди, 980 м над ур. м.; 7 — окр. Кинчха Перди, 816 м над ур. м.; 8 — окр. Сацисквило, 1000 м над ур. м.; 9 — окр. ДЗедЗилети, 520–620 м над ур. м.; 10 — окр. ДЗедЗилети, 770 м над ур. м.; 11 — окр. Мелоури, 475 м над ур. м." figureDoi="http://doi.org/10.5281/zenodo.13164013" httpUri="https://zenodo.org/record/13164013/files/figure.png" pageId="4" pageNumber="255">Fig. 1</figureCitation>
). This river apparently divides ranges of two known
<taxonomicName id="64A924D0FFDB5417FB86C2D3C09885BA" authorityName="Retezar et Djavelidze" authorityYear="1992" box="[1120,1278,689,710]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="species" species="martviliensis">
<emphasis id="91DD8341FFDB5417FB86C2D3C09885BA" box="[1120,1278,689,710]" italics="true" pageId="4" pageNumber="255">C. martviliensis</emphasis>
</taxonomicName>
subspecies. Distribution range of
<taxonomicName id="64A924D0FFDB5417FC11C2AEC08A859C" authorityName="Fominykh &amp; Zamotajlov &amp; Titarenko" authorityYear="2019" box="[1015,1260,715,737]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="subSpecies" species="martviliensis" status="ssp.n." subSpecies="mzekalae">
<emphasis id="91DD8341FFDB5417FC11C2AEC08A859C" box="[1015,1260,715,737]" italics="true" pageId="4" pageNumber="255">C. martviliensis mzekalae</emphasis>
</taxonomicName>
<emphasis id="91DD8341FFDB5417FB16C2A9C14C859C" bold="true" box="[1264,1322,715,736]" pageId="4" pageNumber="255">
<taxonomicNameLabel id="8AEE3E3AFFDB5417FB16C2A9C14C859C" box="[1264,1322,715,736]" pageId="4" pageNumber="255" rank="subSpecies">ssp.n.</taxonomicNameLabel>
</emphasis>
spreads eastwards nearly up to Rioni valley, representative of different species-group of
<emphasis id="91DD8341FFDB5417FC1BC363C00B846A" box="[1021,1133,769,790]" italics="true" pageId="4" pageNumber="255">
<taxonomicName id="64A924D0FFDB5417FC1BC363C00F846A" box="[1021,1129,769,790]" pageId="4" pageNumber="257" rank="subGenus" subGenus="Neoplectes">Neoplectes</taxonomicName>
,
</emphasis>
<taxonomicName id="64A924D0FFDB5417FB93C363C115846A" authority="Retezar, 2011" authorityName="Retezar" authorityYear="2011" box="[1141,1395,769,790]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="species" species="szekelyi">
<emphasis id="91DD8341FFDB5417FB93C363C0B9846A" box="[1141,1247,769,790]" italics="true" pageId="4" pageNumber="255">C. szekelyi</emphasis>
Retezar, 2011
</taxonomicName>
, was found on the opposite bank. The northern limit of the range is still unclear, sympatry with different consubgener
<taxonomicName id="64A924D0FFDB5417FA86C354C7EC841A" authorityName="Zamotajlov et Fominykh" authorityYear="2014" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="species" species="titarenkoi">
<emphasis id="91DD8341FFDB5417FA86C354C7EC841A" italics="true" pageId="4" pageNumber="255">C. titarenkoi</emphasis>
</taxonomicName>
seems to be possible at Gormagali Mt. Mass, this assumption, however, requires confirmation. A new subspecies populates moist low mountain or cloud forests on karstic
</paragraph>
<paragraph id="A3165F53FFDB5417FCDCC77FC0E6804B" blockId="4.[826,1152,1821,1847]" box="[826,1152,1821,1847]" pageId="4" pageNumber="255">
<emphasis id="91DD8341FFDB5417FCDCC77FC0E6804B" bold="true" box="[826,1152,1821,1847]" pageId="4" pageNumber="255">24 25</emphasis>
</paragraph>
</subSubSection>
<caption id="F7D60FDBFFDB5417FF04C72FC0488F71" ID-DOI="http://doi.org/10.5281/zenodo.13164021" ID-Zenodo-Dep="13164021" httpUri="https://zenodo.org/record/13164021/files/figure.png" pageId="4" pageNumber="255" startId="4.[226,263,1869,1888]" targetBox="[201,1384,947,1827]" targetPageId="4" targetType="figure">
<paragraph id="A3165F53FFDB5417FF04C72FC62B80D5" blockId="4.[189,1399,1869,2061]" pageId="4" pageNumber="255">
Figs 18–25. Apical portion of aedeagus and fully inflated endophallus preparation of
<taxonomicName id="64A924D0FFDB5417FC37C72FC162801C" authorityName="Retezar et Djavelidze" authorityYear="1992" box="[977,1284,1869,1889]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="species" species="martviliensis" subGenus="Neoplectes">
<emphasis id="91DD8341FFDB5417FC37C72FC07D801C" box="[977,1051,1869,1888]" italics="true" pageId="4" pageNumber="255">Carabus</emphasis>
(
<emphasis id="91DD8341FFDB5417FBCFC72FC0E1801C" box="[1065,1159,1869,1888]" italics="true" pageId="4" pageNumber="255">Neoplectes</emphasis>
)
<emphasis id="91DD8341FFDB5417FB73C72FC162801C" box="[1173,1284,1869,1888]" italics="true" pageId="4" pageNumber="255">martviliensis</emphasis>
</taxonomicName>
, right lateral view on top, left lateral view below: 18, 22 —
<taxonomicName id="64A924D0FFDB5417FDBFC704C75F8004" authorityName="Fominykh &amp; Zamotajlov &amp; Titarenko" authorityYear="2019" box="[601,825,1893,1913]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="subSpecies" species="martviliensis" status="ssp.n." subSpecies="mzekalae">
<emphasis id="91DD8341FFDB5417FDBFC704C75F8004" box="[601,825,1893,1913]" italics="true" pageId="4" pageNumber="255">C. martviliensis mzekalae</emphasis>
</taxonomicName>
<emphasis id="91DD8341FFDB5417FCD9C704C7148005" bold="true" box="[831,882,1894,1913]" pageId="4" pageNumber="255">
<taxonomicNameLabel id="8AEE3E3AFFDB5417FCD9C704C7148005" box="[831,882,1894,1913]" pageId="4" pageNumber="255" rank="subSpecies">ssp.n.</taxonomicNameLabel>
</emphasis>
, Melouri env., 475 m; 19, 23 —
<taxonomicName id="64A924D0FFDB5417FB71C704C1108004" authorityName="Fominykh &amp; Zamotajlov &amp; Titarenko" authorityYear="2019" box="[1175,1398,1893,1913]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="subSpecies" species="martviliensis" status="ssp.n." subSpecies="mzekalae">
<emphasis id="91DD8341FFDB5417FB71C704C1108004" box="[1175,1398,1893,1913]" italics="true" pageId="4" pageNumber="255">C. martviliensis mzekalae</emphasis>
</taxonomicName>
<emphasis id="91DD8341FFDB5417FF5BC71CC49680ED" bold="true" box="[189,240,1918,1937]" pageId="4" pageNumber="255">
<taxonomicNameLabel id="8AEE3E3AFFDB5417FF5BC71CC49680ED" box="[189,240,1918,1937]" pageId="4" pageNumber="255" rank="subSpecies">ssp.n.</taxonomicNameLabel>
</emphasis>
, Dzedzileti env., 770 m; 20, 24 —
<taxonomicName id="64A924D0FFDB5417FDC9C71CC7FB80EC" authority=", Zeda Gordi" authorityName="Zeda Gordi" box="[559,925,1917,1937]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="subSpecies" species="martviliensis" subSpecies="martviliensis">
<emphasis id="91DD8341FFDB5417FDC9C71CC74880EC" box="[559,814,1917,1937]" italics="true" pageId="4" pageNumber="255">C. martviliensis martviliensis</emphasis>
, Zeda Gordi
</taxonomicName>
env., 980 m; 21, 25 —
<taxonomicName id="64A924D0FFDB5417FB97C71CC54680D4" authority=", Doberazeni" authorityName="Doberazeni" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="subSpecies" species="martviliensis" subSpecies="martviliensis">
<emphasis id="91DD8341FFDB5417FB97C71CC11680EC" box="[1137,1392,1917,1937]" italics="true" pageId="4" pageNumber="255">C. martviliensis martviliensis</emphasis>
, Doberazeni
</taxonomicName>
env., 600 m. Scale bar: 10.0 mm.
</paragraph>
<paragraph id="A3165F53FFDB5417FF04C7D0C0488F71" blockId="4.[189,1399,1869,2061]" pageId="4" pageNumber="255">
Рис. 18–25. ВерШина Эдеагуса и препарат полностью вывернутого внутреннего меШка
<taxonomicName id="64A924D0FFDB5417FC1BC7D3C14B80B8" authorityName="Retezar et Djavelidze" authorityYear="1992" box="[1021,1325,1969,1988]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="species" species="martviliensis" subGenus="Neoplectes">
<emphasis id="91DD8341FFDB5417FC1BC7D3C02180B8" box="[1021,1095,1969,1988]" italics="true" pageId="4" pageNumber="255">Carabus</emphasis>
(
<emphasis id="91DD8341FFDB5417FBB5C7D3C0D780B8" box="[1107,1201,1969,1988]" italics="true" pageId="4" pageNumber="255">Neoplectes</emphasis>
)
<emphasis id="91DD8341FFDB5417FB58C7D3C14B80B8" box="[1214,1325,1969,1988]" italics="true" pageId="4" pageNumber="255">martviliensis</emphasis>
</taxonomicName>
, вверху — сбоку справа, вниЗу — сбоку слева: 18, 22 —
<taxonomicName id="64A924D0FFDB5417FD7BC7ABC7E480A0" authorityName="Fominykh &amp; Zamotajlov &amp; Titarenko" authorityYear="2019" box="[669,898,1993,2012]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="subSpecies" species="martviliensis" status="ssp.n." subSpecies="mzekalae">
<emphasis id="91DD8341FFDB5417FD7BC7ABC7E480A0" box="[669,898,1993,2012]" italics="true" pageId="4" pageNumber="255">C. martviliensis mzekalae</emphasis>
</taxonomicName>
<emphasis id="91DD8341FFDB5417FC6DC7ABC7D880A0" bold="true" box="[907,958,1993,2012]" pageId="4" pageNumber="255">
<taxonomicNameLabel id="8AEE3E3AFFDB5417FC6DC7ABC7D880A0" box="[907,958,1993,2012]" pageId="4" pageNumber="255" rank="subSpecies">ssp.n.</taxonomicNameLabel>
</emphasis>
, окр. Мелоури, 475 м над ур.м.; 19, 23 —
<taxonomicName id="64A924D0FFDB5417FA85C7ABC5E78088" authorityName="Fominykh &amp; Zamotajlov &amp; Titarenko" authorityYear="2019" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="subSpecies" species="martviliensis" status="ssp.n." subSpecies="mzekalae">
<emphasis id="91DD8341FFDB5417FA85C7ABC5E78088" italics="true" pageId="4" pageNumber="255">C. martviliensis mzekalae</emphasis>
</taxonomicName>
<emphasis id="91DD8341FFDB5417FE61C783C5DD8088" bold="true" box="[391,443,2017,2036]" pageId="4" pageNumber="255">
<taxonomicNameLabel id="8AEE3E3AFFDB5417FE61C783C5DD8088" box="[391,443,2017,2036]" pageId="4" pageNumber="255" rank="subSpecies">ssp.n.</taxonomicNameLabel>
</emphasis>
, окр. ДЗедЗилети, 770 м над ур.м.; 20, 24 —
<taxonomicName id="64A924D0FFDB5417FCB0C783C0358088" box="[854,1107,2017,2036]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="subSpecies" species="martviliensis" subSpecies="martviliensis">
<emphasis id="91DD8341FFDB5417FCB0C783C0358088" box="[854,1107,2017,2036]" italics="true" pageId="4" pageNumber="255">C. martviliensis martviliensis</emphasis>
</taxonomicName>
, окр. Зеда Горди, 980 м над ур. м.; 21, 25 —
<taxonomicName id="64A924D0FFDB5417FED1C79BC6518F70" box="[311,567,2041,2060]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="subSpecies" species="martviliensis" subSpecies="martviliensis">
<emphasis id="91DD8341FFDB5417FED1C79BC6518F70" box="[311,567,2041,2060]" italics="true" pageId="4" pageNumber="255">C. martviliensis martviliensis</emphasis>
</taxonomicName>
, окр. ДобераЗени, 600 м над ур.м. МасШтаб: 10,0 мм.
</paragraph>
</caption>
<caption id="F7D60FDBFFDA5416FF04C402C0FB83C3" ID-DOI="http://doi.org/10.5281/zenodo.13164023" ID-Zenodo-Dep="13164023" httpUri="https://zenodo.org/record/13164023/files/figure.png" pageId="5" pageNumber="256" startId="5.[226,257,1120,1139]" targetBox="[280,1321,332,1094]" targetPageId="5" targetType="figure">
<paragraph id="A3165F53FFDA5416FF04C402C76A83F7" blockId="5.[189,1398,1120,1216]" pageId="5" pageNumber="256">
Fig. 26. Distribution of morphometric characters of
<taxonomicName id="64A924D0FFDA5416FD58C402C791830F" authorityName="Retezar et Djavelidze" authorityYear="1992" box="[702,1015,1120,1139]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="species" species="martviliensis" subGenus="Neoplectes">
<emphasis id="91DD8341FFDA5416FD58C402C76E830F" box="[702,776,1120,1139]" italics="true" pageId="5" pageNumber="256">Carabus</emphasis>
(
<emphasis id="91DD8341FFDA5416FCFFC402C711830F" box="[793,887,1120,1139]" italics="true" pageId="5" pageNumber="256">Neoplectes</emphasis>
)
<emphasis id="91DD8341FFDA5416FC6EC402C791830F" box="[904,1015,1120,1139]" italics="true" pageId="5" pageNumber="256">martviliensis</emphasis>
</taxonomicName>
subspecies constructed using discriminant analysis based on 9 parameters (see text for measurements details).
</paragraph>
<paragraph id="A3165F53FFDA5416FF04C4F7C0FB83C3" blockId="5.[189,1398,1120,1216]" pageId="5" pageNumber="256">
Рис. 26. Распределение морфометрических характеристик подвидов
<taxonomicName id="64A924D0FFDA5416FC77C4F6C0BF83DB" authorityName="Retezar et Djavelidze" authorityYear="1992" box="[913,1241,1172,1191]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="species" species="martviliensis" subGenus="Neoplectes">
<emphasis id="91DD8341FFDA5416FC77C4F6C7BB83DB" box="[913,989,1172,1191]" italics="true" pageId="5" pageNumber="256">Carabus</emphasis>
(
<emphasis id="91DD8341FFDA5416FC16C4F6C03783DB" box="[1008,1105,1172,1191]" italics="true" pageId="5" pageNumber="256">Neoplectes</emphasis>
)
<emphasis id="91DD8341FFDA5416FB83C4F6C0BF83DB" box="[1125,1241,1172,1191]" italics="true" pageId="5" pageNumber="256">martviliensis</emphasis>
</taxonomicName>
, построенное с испольЗованием дискриминантного аналиЗа, основанного на 9 параметрах (описание промеров см. текст).
</paragraph>
</caption>
<subSubSection id="EBB30CD8FFDA5415FF2AC69DC5C486FA" lastPageId="6" lastPageNumber="257" pageId="5" pageNumber="256" type="description">
<paragraph id="A3165F53FFDA5416FF2AC69DC7CE8065" blockId="5.[204,233,1791,1817]" box="[204,936,1791,1817]" lastBlockId="5.[907,936,1791,1817]" pageId="5" pageNumber="256">
<emphasis id="91DD8341FFDA5416FF2AC69DC7CE8065" bold="true" box="[204,936,1791,1817]" pageId="5" pageNumber="256">27 28</emphasis>
</paragraph>
<caption id="F7D60FDBFFDA5416FF04C752C0FA80E5" ID-DOI="http://doi.org/10.5281/zenodo.13164025" ID-Zenodo-Dep="13164025" httpUri="https://zenodo.org/record/13164025/files/figure.png" pageId="5" pageNumber="256" startId="5.[226,257,1840,1859]" targetBox="[189,1399,1255,1782]" targetPageId="5" targetType="figure">
<paragraph id="A3165F53FFDA5416FF04C752C057803F" blockId="5.[226,1231,1840,1945]" box="[226,1073,1840,1859]" pageId="5" pageNumber="256">
Fig. 27. Habitat of
<taxonomicName id="64A924D0FFDA5416FE68C752C77C803F" authorityName="Fominykh &amp; Zamotajlov &amp; Titarenko" authorityYear="2019" box="[398,794,1840,1859]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="subSpecies" species="martviliensis" status="subsp. nov." subGenus="Neoplectes" subSpecies="mzekalae">
<emphasis id="91DD8341FFDA5416FE68C752C5BE803F" box="[398,472,1840,1859]" italics="true" pageId="5" pageNumber="256">Carabus</emphasis>
(
<emphasis id="91DD8341FFDA5416FE00C752C622803F" box="[486,580,1840,1859]" italics="true" pageId="5" pageNumber="256">Neoplectes</emphasis>
)
<emphasis id="91DD8341FFDA5416FDB5C752C77C803F" box="[595,794,1840,1859]" italics="true" pageId="5" pageNumber="256">martviliensis mzekalae</emphasis>
</taxonomicName>
<emphasis id="91DD8341FFDA5416FCC7C752C732803F" bold="true" box="[801,852,1840,1859]" pageId="5" pageNumber="256">
<taxonomicNameLabel id="8AEE3E3AFFDA5416FCC7C752C732803F" box="[801,852,1840,1859]" pageId="5" pageNumber="256" rank="subSpecies">ssp.n.</taxonomicNameLabel>
</emphasis>
, Dzedzileti env., 770 m.
</paragraph>
<paragraph id="A3165F53FFDA5416FF04C72EC0A98023" blockId="5.[226,1231,1840,1945]" box="[226,1231,1867,1887]" pageId="5" pageNumber="256">
Рис. 27. Местообитание
<taxonomicName id="64A924D0FFDA5416FE2EC729C7328022" authorityName="Fominykh &amp; Zamotajlov &amp; Titarenko" authorityYear="2019" box="[456,852,1867,1887]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="subSpecies" species="martviliensis" status="subsp. nov." subGenus="Neoplectes" subSpecies="mzekalae">
<emphasis id="91DD8341FFDA5416FE2EC729C6748022" box="[456,530,1867,1886]" italics="true" pageId="5" pageNumber="256">Carabus</emphasis>
(
<emphasis id="91DD8341FFDA5416FDC7C729C6198022" box="[545,639,1867,1886]" italics="true" pageId="5" pageNumber="256">Neoplectes</emphasis>
)
<emphasis id="91DD8341FFDA5416FD6BC729C7328022" box="[653,852,1867,1886]" italics="true" pageId="5" pageNumber="256">martviliensis mzekalae</emphasis>
</taxonomicName>
<emphasis id="91DD8341FFDA5416FCBDC72EC7E98023" bold="true" box="[859,911,1868,1887]" pageId="5" pageNumber="256">
<taxonomicNameLabel id="8AEE3E3AFFDA5416FCBDC72EC7E98023" box="[859,911,1868,1887]" pageId="5" pageNumber="256" rank="subSpecies">ssp.n.</taxonomicNameLabel>
</emphasis>
, окр. ДЗедЗилети, 770 м над ур.м.
</paragraph>
<paragraph id="A3165F53FFDA5416FF04C708C0418002" blockId="5.[226,1231,1840,1945]" box="[226,1063,1898,1918]" pageId="5" pageNumber="256">
Fig. 28.
<taxonomicName id="64A924D0FFDA5416FEC8C708C6DC8001" authorityName="Fominykh &amp; Zamotajlov &amp; Titarenko" authorityYear="2019" box="[302,698,1898,1917]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="subSpecies" species="martviliensis" status="subsp. nov." subGenus="Neoplectes" subSpecies="mzekalae">
<emphasis id="91DD8341FFDA5416FEC8C708C51E8001" box="[302,376,1898,1917]" italics="true" pageId="5" pageNumber="256">Carabus</emphasis>
(
<emphasis id="91DD8341FFDA5416FE61C708C5838001" box="[391,485,1898,1917]" italics="true" pageId="5" pageNumber="256">Neoplectes</emphasis>
)
<emphasis id="91DD8341FFDA5416FE15C708C6DC8001" box="[499,698,1898,1917]" italics="true" pageId="5" pageNumber="256">martviliensis mzekalae</emphasis>
</taxonomicName>
<emphasis id="91DD8341FFDA5416FD27C708C6938001" bold="true" box="[705,757,1898,1917]" pageId="5" pageNumber="256">
<taxonomicNameLabel id="8AEE3E3AFFDA5416FD27C708C6938001" box="[705,757,1898,1917]" pageId="5" pageNumber="256" rank="subSpecies">ssp.n.</taxonomicNameLabel>
</emphasis>
in nature, Dzedzileti env., 770 m.
</paragraph>
<paragraph id="A3165F53FFDA5416FF04C7E4C0FA80E5" blockId="5.[226,1231,1840,1945]" box="[226,1180,1926,1945]" pageId="5" pageNumber="256">
Рис. 28.
<taxonomicName id="64A924D0FFDA5416FED4C7E4C6DB80E5" authorityName="Fominykh &amp; Zamotajlov &amp; Titarenko" authorityYear="2019" box="[306,701,1926,1945]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="subSpecies" species="martviliensis" status="subsp. nov." subGenus="Neoplectes" subSpecies="mzekalae">
<emphasis id="91DD8341FFDA5416FED4C7E4C51A80E5" box="[306,380,1926,1945]" italics="true" pageId="5" pageNumber="256">Carabus</emphasis>
(
<emphasis id="91DD8341FFDA5416FE6CC7E4C58E80E5" box="[394,488,1926,1945]" italics="true" pageId="5" pageNumber="256">Neoplectes</emphasis>
)
<emphasis id="91DD8341FFDA5416FE11C7E4C6DB80E5" box="[503,701,1926,1945]" italics="true" pageId="5" pageNumber="256">martviliensis mzekalae</emphasis>
</taxonomicName>
<emphasis id="91DD8341FFDA5416FD23C7E4C69E80E5" bold="true" box="[709,760,1926,1945]" pageId="5" pageNumber="256">
<taxonomicNameLabel id="8AEE3E3AFFDA5416FD23C7E4C69E80E5" box="[709,760,1926,1945]" pageId="5" pageNumber="256" rank="subSpecies">ssp.n.</taxonomicNameLabel>
</emphasis>
в природе, окр. ДЗедЗилети, 770 м над ур.м.
</paragraph>
</caption>
<paragraph id="A3165F53FFDA5415FF5BC7A6C5C486FA" blockId="5.[189,775,1988,2063]" lastBlockId="6.[189,781,315,475]" lastPageId="6" lastPageNumber="257" pageId="5" pageNumber="256">
landforms, often with the very poor ground litter and dense underbrush of rhododendron (
<emphasis id="91DD8341FFDA5416FE05C7BCC6B28088" box="[483,724,2014,2036]" italics="true" pageId="5" pageNumber="256">
<taxonomicName id="64A924D0FFDA5416FE05C7BCC614808F" authorityName="Deuve &amp; Imura" authorityYear="1991" box="[483,626,2014,2035]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="species" species="rhododendron">Rhododendron</taxonomicName>
ponticum
</emphasis>
) and blackberry (
<taxonomicName id="64A924D0FFDA5416FED3C79BC5BF8F72" box="[309,473,2041,2062]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="5" pageNumber="256" phylum="Tracheophyta" rank="species" species="fruticosus">
<emphasis id="91DD8341FFDA5416FED3C79BC5BF8F72" box="[309,473,2041,2062]" italics="true" pageId="5" pageNumber="256">Rubus fruticosus</emphasis>
</taxonomicName>
) (
<figureCitation id="3B9243D6FFDA5416FE17C79BC6588F72" box="[497,574,2041,2063]" captionStart="Fig" captionStartId="5.[226,257,1840,1859]" captionTargetBox="[189,1399,1255,1782]" captionTargetId="figure-144@5.[189,1399,1255,1782]" captionTargetPageId="5" captionText="Fig. 27. Habitat of Carabus (Neoplectes) martviliensis mzekalae ssp.n., Dzedzileti env., 770 m. Рис. 27. Местообитание Carabus (Neoplectes) martviliensis mzekalae ssp.n., окр. ДЗедЗилети, 770 м над ур.м. Fig. 28. Carabus (Neoplectes) martviliensis mzekalae ssp.n. in nature, Dzedzileti env., 770 m. Рис. 28. Carabus (Neoplectes) martviliensis mzekalae ssp.n. в природе, окр. ДЗедЗилети, 770 м над ур.м." figureDoi="http://doi.org/10.5281/zenodo.13164025" httpUri="https://zenodo.org/record/13164025/files/figure.png" pageId="5" pageNumber="256">Figs 27</figureCitation>
–28). The adults are active from the beginning of May until the mid of July. Following
<taxonomicName id="64A924D0FFDA5416FC7FC7BCC78A808F" authorityName="Linnaeus" authorityYear="1758" box="[921,1004,2014,2035]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="genus">
<emphasis id="91DD8341FFDA5416FC7FC7BCC78A808F" box="[921,1004,2014,2035]" italics="true" pageId="5" pageNumber="256">Carabus</emphasis>
</taxonomicName>
forms occur together with this subspecies:
<taxonomicName id="64A924D0FFDA5416FC87C79BC09C8F73" authority="Shaum, 1864" authorityName="Shaum" authorityYear="1864" box="[865,1274,2041,2063]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="species" species="protensus" subGenus="Archiplectes">
<emphasis id="91DD8341FFDA5416FC87C79BC71E8F72" box="[865,888,2041,2062]" italics="true" pageId="5" pageNumber="256">C.</emphasis>
(
<emphasis id="91DD8341FFDA5416FC6EC79BC0678F72" box="[904,1025,2041,2062]" italics="true" pageId="5" pageNumber="256">Archiplectes</emphasis>
)
<emphasis id="91DD8341FFDA5416FBF7C798C0098F73" box="[1041,1135,2042,2063]" italics="true" pageId="5" pageNumber="256">protensus</emphasis>
Shaum, 1864
</taxonomicName>
,
<taxonomicName id="64A924D0FFDA5415FAE1C79BC5D6862C" authority="Chaudoir, 1877" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" lastPageId="6" lastPageNumber="257" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="species" species="fossiger" subGenus="Tribax">
<emphasis id="91DD8341FFDA5416FAE1C79BC1788F72" box="[1287,1310,2041,2062]" italics="true" pageId="5" pageNumber="256">C.</emphasis>
(
<emphasis id="91DD8341FFDA5416FAC8C79BC1088F72" box="[1326,1390,2041,2062]" italics="true" pageId="5" pageNumber="256">Tribax</emphasis>
)
<emphasis id="91DD8341FFD95415FF5BC159C56F862C" box="[189,265,315,336]" italics="true" pageId="6" pageNumber="257">fossiger</emphasis>
Chaudoir, 1877
</taxonomicName>
,
<taxonomicName id="64A924D0FFD95415FE5BC15EC5FB8617" authority="Mannerheim, 1830" authorityName="Mannerheim" authorityYear="1830" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="257" phylum="Arthropoda" rank="species" species="armeniacus" subGenus="Sphodristocarabus">
<emphasis id="91DD8341FFD95415FE5BC15EC5B2862D" box="[445,468,316,337]" italics="true" pageId="6" pageNumber="257">C.</emphasis>
(
<emphasis id="91DD8341FFD95415FE00C159C6FA862C" box="[486,668,315,336]" italics="true" pageId="6" pageNumber="257">Sphodristocarabus</emphasis>
)
<emphasis id="91DD8341FFD95415FD48C15EC4BB8610" italics="true" pageId="6" pageNumber="257">armeniacus</emphasis>
Mannerheim, 1830
</taxonomicName>
, and
<taxonomicName id="64A924D0FFD95415FE34C134C5F886FA" authority="Motschulsky, 1840" authorityName="Motschulsky" authorityYear="1840" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="257" phylum="Arthropoda" rank="species" species="septemcarinatus" subGenus="Megodontus">
<emphasis id="91DD8341FFD95415FE34C134C58E8617" box="[466,488,342,363]" italics="true" pageId="6" pageNumber="257">C.</emphasis>
(
<emphasis id="91DD8341FFD95415FE11C134C6088617" box="[503,622,342,363]" italics="true" pageId="6" pageNumber="257">Megodontus</emphasis>
)
<emphasis id="91DD8341FFD95415FD9AC135C4BF86FA" italics="true" pageId="6" pageNumber="257">septemcarinatus</emphasis>
Motschulsky, 1840
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="EBB30CD8FFD95415FF04C1E9C6E78467" pageId="6" pageNumber="257" type="etymology">
<paragraph id="A3165F53FFD95415FF04C1E9C69686A7" blockId="6.[189,781,315,475]" pageId="6" pageNumber="257">ETYMOLOGY. The subspecific epithet refers to the Georgian name Mzekala (მზექალა), goddess of Sun, patroness of the wine-making, heroine of the Georgian epos.</paragraph>
<paragraph id="A3165F53FFD95415FF04C199C6E78467" blockId="6.[189,775,507,796]" pageId="6" pageNumber="257">
<emphasis id="91DD8341FFD95415FF04C199C5CF856D" bold="true" box="[226,425,507,529]" pageId="6" pageNumber="257">Acknowledgements</emphasis>
. The authors are very grateful to Mr. A. Zubov (Kishinev,
<collectingCountry id="DBBE1FC3FFD95415FE77C274C5978550" box="[401,497,534,556]" name="Moldova" pageId="6" pageNumber="257">Moldavia</collectingCountry>
), Mr. B. Dubinin (
<collectingRegion id="616D91B1FFD95415FD4AC274C7658557" box="[684,771,534,555]" country="Russia" name="Moscow" pageId="6" pageNumber="257">Moscow</collectingRegion>
,
<collectingCountry id="DBBE1FC3FFD95415FF5BC252C567853A" box="[189,257,560,582]" name="Russia" pageId="6" pageNumber="257">Russia</collectingCountry>
), Mr. D. Safronov (
<collectingRegion id="616D91B1FFD95415FE25C252C595853A" box="[451,499,560,582]" country="Russia" name="Tula" pageId="6" pageNumber="257">Tula</collectingRegion>
,
<collectingCountry id="DBBE1FC3FFD95415FE18C252C626853A" box="[510,576,560,582]" name="Russia" pageId="6" pageNumber="257">Russia</collectingCountry>
) for their assistance during the collecting trips to
<collectingCountry id="DBBE1FC3FFD95415FE3CC229C64D851D" box="[474,555,587,609]" name="Georgia" pageId="6" pageNumber="257">Georgia</collectingCountry>
. We wish to acknowledge here also the unselfish help of Prof. K. Makarov (
<collectingRegion id="616D91B1FFD95415FD34C204C48185EA" country="Russia" name="Moscow" pageId="6" pageNumber="257">Moscow</collectingRegion>
,
<collectingCountry id="DBBE1FC3FFD95415FF15C2E2C55385EA" box="[243,309,640,662]" name="Russia" pageId="6" pageNumber="257">Russia</collectingCountry>
) in taking the photographs and preparing them for publication and help of M.Sc. E. Khomitskiy (Goryachiy Klyuch,
<collectingCountry id="DBBE1FC3FFD95415FEF5C2D4C53385B0" box="[275,341,694,716]" name="Russia" pageId="6" pageNumber="257">Russia</collectingCountry>
) in various technical procedures. Some aspects of the reported study were funded by the Russian Foundation for Basic Research and Administration of
<collectingRegion id="616D91B1FFD95415FD23C289C48E8460" country="Russia" name="Krasnodar" pageId="6" pageNumber="257">Krasnodar</collectingRegion>
Territory, project number 19-44-230004.
</paragraph>
</subSubSection>
</treatment>
</document>
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