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<document id="255ABE363BDE449FC142114FE692C4A5" ID-DOI="10.5281/zenodo.193406" ID-GBIF-Dataset="5e8043a5-e685-4a21-b696-0f7a706ab223" ID-ISSN="1175-5326" ID-Zenodo-Dep="193406" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460138703176" checkinUser="plazi" docAuthor="Toledo, Mario, Spangler, Paul J. &amp; Balke, Michael" docDate="2010" docId="11578788FE55FFCF7DB5FBAF1220DA09" docLanguage="en" docName="zt02347p058.pdf" docOrigin="Zootaxa 2347" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Laccodytes americanus Peschet 1919" docType="treatment" docVersion="9" lastPageNumber="50" masterDocId="ED6EFFF0FE5CFFC27D22FFF3160ED852" masterDocTitle="Taxonomic revision of the Neotropical diving beetles genus Laccodytes Régimbart, 1895 (Coleoptera: Dytiscidae)" masterLastPageNumber="58" masterPageNumber="37" pageNumber="46" updateTime="1698235333712" updateUser="plazi">
<mods:mods id="00E7C37D10A6D5A49E0C6F74E605E2DC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7A3DE614EDEFC152A6D84C6BD1FCC3A5">
<mods:title id="FE03FBB1EF1F68E7AD8245B65ABCAE6D">Taxonomic revision of the Neotropical diving beetles genus Laccodytes Régimbart, 1895 (Coleoptera: Dytiscidae)</mods:title>
</mods:titleInfo>
<mods:name id="0B52EFFAC7BFF81C254E33AE0A355E47" type="personal">
<mods:role id="9302597B76F995D74F9B613717509588">
<mods:roleTerm id="8B965807616633B40F32170B95A7C1C1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="990AB1E0F12FCEFD9D2B3304F630CAD1">Toledo, Mario</mods:namePart>
</mods:name>
<mods:name id="793BBD3B2F99E7F9FF9C602414A5FE16" type="personal">
<mods:role id="9EA003E5466DEEB22F94A849FCBA6505">
<mods:roleTerm id="0493B72B7AEA6698511884B52966BC9C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D160FA4AFF238C9B3D1846EA3FBFB17F">Spangler, Paul J.</mods:namePart>
</mods:name>
<mods:name id="CBD3C1FBE9FC5DA6E2A3AF862E49FAB7" type="personal">
<mods:role id="030987FFC11FF595B1C0CD0F28B06B5D">
<mods:roleTerm id="48A8599743AF611457BDC8E22F3D19AF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="46D9D8698F8726F991D91CBDB08F484D">Balke, Michael</mods:namePart>
</mods:name>
<mods:typeOfResource id="0C9B9D364E3C000BC091C78C5B66D4AC">text</mods:typeOfResource>
<mods:relatedItem id="E4440C5F52546E4B33037C7465848BF5" type="host">
<mods:titleInfo id="2956141FF046272AD729734463F2EDD6">
<mods:title id="9BF67CF46A418E4A18D675CE5031930B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="61AC6CAFF8DED0AA3EC301C37D24A39A">
<mods:date id="7C337D0256238A678217DB1F0DB78709">2010</mods:date>
<mods:detail id="00864EA933814366722E3740F6943F0F" type="volume">
<mods:number id="C17FD273AEC1799CF389BD22B36CAB84">2347</mods:number>
</mods:detail>
<mods:extent id="729599C3ACFE61C24E41F37E9F9E4C87" unit="page">
<mods:start id="80A8605BFE1B01AA4D4859A25148F630">37</mods:start>
<mods:end id="FFE4A97F4E5FC981656554566D0A4F3A">58</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="171ABACA37245BBB37BD4C262806E27D">journal article</mods:classification>
<mods:identifier id="4234773F32BAC5728935D6CB14E7011F" type="DOI">10.5281/zenodo.193406</mods:identifier>
<mods:identifier id="9E879D69D2F30366F52A89ACE7F8B1DF" type="GBIF-Dataset">5e8043a5-e685-4a21-b696-0f7a706ab223</mods:identifier>
<mods:identifier id="3535B3A35CFB5207372B1F9D95986FDF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E0E97DD4D8647116E4C1F05A8D8F89BA" type="Zenodo-Dep">193406</mods:identifier>
</mods:mods>
<treatment id="11578788FE55FFCF7DB5FBAF1220DA09" ID-DOI="http://doi.org/10.5281/zenodo.6198368" ID-GBIF-Taxon="124955825" ID-Zenodo-Dep="6198368" LSID="urn:lsid:plazi:treatment:11578788FE55FFCF7DB5FBAF1220DA09" httpUri="http://treatment.plazi.org/id/11578788FE55FFCF7DB5FBAF1220DA09" lastPageId="13" lastPageNumber="50" pageId="9" pageNumber="46">
<subSubSection id="D1E46515FE55FFCB7DB5FBAF1461DCCB" pageId="9" pageNumber="46" type="nomenclature">
<paragraph id="9941369EFE55FFCB7DB5FBAF146EDC24" blockId="9.[151,623,1116,1177]" box="[151,608,1116,1142]" pageId="9" pageNumber="46">
<heading id="C20981F2FE55FFCB7DB5FBAF146EDC24" bold="true" box="[151,608,1116,1142]" fontSize="11" level="1" pageId="9" pageNumber="46" reason="1">
<taxonomicName id="5EFE4D1DFE55FFCB7DB5FBAF146EDC24" ID-CoL="3RLZD" authority="Peschet, 1919" authorityName="Peschet" authorityYear="1919" box="[151,608,1116,1142]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="AB8AEA8CFE55FFCB7DB5FBAF146EDC24" bold="true" box="[151,608,1116,1142]" pageId="9" pageNumber="46">
<emphasis id="AB8AEA8CFE55FFCB7DB5FBAF17BFDC24" bold="true" box="[151,433,1116,1142]" italics="true" pageId="9" pageNumber="46">Laccodytes americanus</emphasis>
<bibRefCitation id="FD6F4B6FFE55FFCB7C9AFBAF146EDC24" author="Peschet" box="[440,608,1116,1142]" pageId="9" pageNumber="58" refString="Peschet, R. (1919) Description d'un Dytiscidae nouveaux de l'Amerique du Sud (Col.). Bulletin de la Societe Entomologique de France, 1919 (8), 145 - 147." type="journal article" year="1919">Peschet, 1919</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="9941369EFE55FFCB7DB5FB8C1461DCCB" blockId="9.[151,623,1116,1177]" box="[151,623,1151,1177]" pageId="9" pageNumber="46">Figs (3, 5, 10, 13, 16–21, 30, 46, 56, 62)</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE55FFCB7DB5FB3F1507DD51" pageId="9" pageNumber="46" type="reference_group">
<paragraph id="9941369EFE55FFCB7DB5FB3F1507DD51" blockId="9.[151,896,1228,1283]" pageId="9" pageNumber="46">
<treatmentCitationGroup id="B9EE11B0FE55FFCB7DB5FB3F1507DD51" pageId="9" pageNumber="46">
<taxonomicName id="5EFE4D1DFE55FFCB7DB5FB3F147EDCB1" ID-CoL="3RLZD" authority="Peschet (1919: 145)" authorityName="Peschet" authorityPageNumber="145" authorityYear="1919" box="[151,624,1228,1251]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="AB8AEA8CFE55FFCB7DB5FB3F179FDCB1" box="[151,401,1228,1251]" italics="true" pageId="9" pageNumber="46">Laccodytes americanus</emphasis>
<treatmentCitation id="185F108FFE55FFCB7CBAFB3F147EDCB1" author="Peschet" box="[408,624,1228,1251]" page="145" pageId="9" pageNumber="46" year="1919">
<bibRefCitation id="FD6F4B6FFE55FFCB7CBAFB3F147EDCB1" author="Peschet" box="[408,624,1228,1251]" pageId="9" pageNumber="58" refString="Peschet, R. (1919) Description d'un Dytiscidae nouveaux de l'Amerique du Sud (Col.). Bulletin de la Societe Entomologique de France, 1919 (8), 145 - 147." type="journal article" year="1919">Peschet (1919: 145)</bibRefCitation>
</treatmentCitation>
</taxonomicName>
; Blackwelder (1944: 74).
<taxonomicName id="5EFE4D1DFE55FFCB7DB5FB1F1789DD51" box="[151,391,1260,1283]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="AB8AEA8CFE55FFCB7DB5FB1F1789DD51" box="[151,391,1260,1283]" italics="true" pageId="9" pageNumber="46">Laccodytes americana</emphasis>
</taxonomicName>
(sic!) Peschet:
<treatmentCitation id="185F108FFE55FFCB7F0EFB1F150BDD51" author="Nilsson" box="[556,773,1260,1283]" page="239" pageId="9" pageNumber="46" year="2001">
<bibRefCitation id="FD6F4B6FFE55FFCB7F0EFB1F150BDD51" author="Nilsson" box="[556,773,1260,1283]" pageId="9" pageNumber="58" refString="Nilsson, A. N. (2001) World catalogue of insects. Vol. 3. Dytiscidae Coleoptera. Stenstrup, Apollo Books, 395 pp." type="book" year="2001">Nilsson (2001: 239)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE55FFCB7DB5FAC115FBDF38" pageId="9" pageNumber="46" type="materials_examined">
<paragraph id="9941369EFE55FFCB7DB5FAC115A4DD1F" blockId="9.[151,1436,1330,1975]" box="[151,938,1330,1357]" pageId="9" pageNumber="46">
<emphasis id="AB8AEA8CFE55FFCB7DB5FAC11733DD1E" bold="true" box="[151,317,1330,1356]" pageId="9" pageNumber="46">
<typeStatus id="4645883CFE55FFCB7DB5FAC116DADD1E" box="[151,212,1330,1356]" pageId="9" pageNumber="46">Type</typeStatus>
locality.
</emphasis>
French
<collectingCountry id="E1E9760EFE55FFCB7CBEFAC017F2DD1F" box="[412,508,1331,1357]" name="Guyana" pageId="9" pageNumber="46">Guyana</collectingCountry>
: Charvein, Saint-Laurent de Maroni.
</paragraph>
<paragraph id="9941369EFE55FFCB7DE4FAAA1747DE67" blockId="9.[151,1436,1330,1975]" pageId="9" pageNumber="46">
<emphasis id="AB8AEA8CFE55FFCB7DE4FAAA1771DD21" bold="true" box="[198,383,1369,1395]" pageId="9" pageNumber="46">
<typeStatus id="4645883CFE55FFCB7DE4FAAA170FDD21" box="[198,257,1369,1395]" pageId="9" pageNumber="46">Type</typeStatus>
material.
</emphasis>
<typeStatus id="4645883CFE55FFCB7CAFFAAA17E3DD21" box="[397,493,1369,1395]" pageId="9" pageNumber="46" type="syntype">Syntype</typeStatus>
3 (
<collectionCode id="FFEFAE5BFE55FFCB7F06FAAA148DDD21" LSID="urn:lsid:biocol.org:col:34988" box="[548,643,1369,1395]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="46">MNHN</collectionCode>
, Peschet collection):
<collectingCountry id="E1E9760EFE55FFCB7EA8FAAA15EBDD21" box="[906,997,1369,1395]" name="French Guiana" pageId="9" pageNumber="46">Guyane</collectingCountry>
francaise, St. Laurent de Maroni //
<typeStatus id="4645883CFE55FFCB7DB5FA7316ECDDC8" box="[151,226,1408,1434]" pageId="9" pageNumber="46">
<collectionCode id="FFEFAE5BFE55FFCB7DB5FA7316ECDDC8" box="[151,226,1408,1434]" pageId="9" pageNumber="46">TYPE</collectionCode>
</typeStatus>
// Museum Paris //
<taxonomicName id="5EFE4D1DFE55FFCB7CF0FA7314E1DDC8" box="[466,751,1408,1434]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="americanus">Laccodytes americanus</taxonomicName>
. According to Peschet, there are altogether
<specimenCount id="8FF8FD17FE55FFCB7822FA73135FDDC8" box="[1280,1361,1408,1434]" pageId="9" pageNumber="46" type="male">2 male</specimenCount>
and
<specimenCount id="8FF8FD17FE55FFCB78AFFA7316E9DD93" pageId="9" pageNumber="46" type="female">5 female</specimenCount>
<typeStatus id="4645883CFE55FFCB7DCFFA541725DD93" box="[237,299,1447,1473]" pageId="9" pageNumber="46">types</typeStatus>
in his collection. One male
<typeStatus id="4645883CFE55FFCB7F51FA5414C0DD93" box="[627,718,1447,1473]" pageId="9" pageNumber="46" type="syntype">syntype</typeStatus>
was studied and illustrated by Paul Spangler in 1980, and we base our redescription of
<taxonomicName id="5EFE4D1DFE55FFCB7CE4FA3D1461DDB5" box="[454,623,1486,1511]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="AB8AEA8CFE55FFCB7CE4FA3D1461DDB5" box="[454,623,1486,1511]" italics="true" pageId="9" pageNumber="46">L. americanus</emphasis>
</taxonomicName>
on his notes and
<collectionCode id="FFEFAE5BFE55FFCB7E63FA3E1598DDB5" LSID="urn:lsid:biocol.org:col:34871" box="[833,918,1485,1511]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="46">USNM</collectionCode>
specimens which agree well with the notes on the
<typeStatus id="4645883CFE55FFCB7DC5FA071748DE5C" box="[231,326,1524,1550]" pageId="9" pageNumber="46" type="syntype">syntype</typeStatus>
. Specimens were not available from the
<collectionCode id="FFEFAE5BFE55FFCB7E00FA071572DE5C" LSID="urn:lsid:biocol.org:col:34988" box="[802,892,1524,1550]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="46">MNHN</collectionCode>
for study any more, due to shortage of staff at that institution.
</paragraph>
<paragraph id="9941369EFE55FFCB7DE4F9B21570DEFB" blockId="9.[151,1436,1330,1975]" pageId="9" pageNumber="46">
<emphasis id="AB8AEA8CFE55FFCB7DE4F9B2140ADE09" bold="true" box="[198,516,1601,1627]" pageId="9" pageNumber="46">Other material studied.</emphasis>
<collectingCountry id="E1E9760EFE55FFCB7F3AF9B21476DE09" box="[536,632,1601,1627]" name="Guyana" pageId="9" pageNumber="46">Guyana</collectingCountry>
: Mazaruni-Potaro District, Takutu Mountains,
<geoCoordinate id="FCCA5059FE55FFCB79FBF9B21325DE09" box="[1241,1323,1601,1627]" direction="north" orientation="latitude" pageId="9" pageNumber="46" precision="925" value="6.25">6°15'N</geoCoordinate>
<geoCoordinate id="FCCA5059FE55FFCB781DF9B21399DE09" box="[1343,1431,1601,1627]" direction="west" orientation="longitude" pageId="9" pageNumber="46" precision="925" value="-59.083332">59°5'W</geoCoordinate>
, Earthwatch research expedition,
<date id="ED40105EFE55FFCB7F3AF99B14C1DED0" box="[536,719,1640,1666]" pageId="9" pageNumber="46" value="1983-12-06" valueMax="1983-12-18" valueMin="1983-12-06">6.–18.XII.1983</date>
, P.J. Spangler, W.E. Steiner &amp; R.A. Faitoute (20 exs
<collectionCode id="FFEFAE5BFE55FFCB7862F99B1396DED0" LSID="urn:lsid:biocol.org:col:34871" box="[1344,1432,1640,1666]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="46">USNM</collectionCode>
,
<collectionCode id="FFEFAE5BFE55FFCB7DB5F97C16E9DEFB" box="[151,231,1679,1705]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="9" pageNumber="46">NMW</collectionCode>
); same but
<geoCoordinate id="FCCA5059FE55FFCB7C49F97C17B3DEFB" box="[363,445,1679,1705]" direction="north" orientation="latitude" pageId="9" pageNumber="46" precision="925" value="6.25">6°15'N</geoCoordinate>
<geoCoordinate id="FCCA5059FE55FFCB7CE6F97C1422DEFB" box="[452,556,1679,1705]" direction="west" orientation="longitude" pageId="9" pageNumber="46" precision="925" value="-58.916668">58°55'W</geoCoordinate>
,
<date id="ED40105EFE55FFCB7F15F97C14C9DEFB" box="[567,711,1679,1705]" pageId="9" pageNumber="46" value="1983-12-19">19.XII.1983</date>
(1 ex.
<collectionCode id="FFEFAE5BFE55FFCB7E3AF97C157CDEFB" LSID="urn:lsid:biocol.org:col:34871" box="[792,882,1679,1705]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="46">USNM</collectionCode>
).
</paragraph>
<paragraph id="9941369EFE55FFCB7DE4F94615FBDF38" blockId="9.[151,1436,1330,1975]" pageId="9" pageNumber="46">
<collectingCountry id="E1E9760EFE55FFCB7DE4F9461735DE9D" box="[198,315,1717,1743]" name="Suriname" pageId="9" pageNumber="46">Suriname</collectingCountry>
: Commewijne, Akintosoela, Celos camp,
<quantity id="5E069B7BFE55FFCB7E0DF9461574DE9D" box="[815,890,1717,1743]" metricMagnitude="4" metricUnit="m" metricValue="3.9" pageId="9" pageNumber="46" unit="km" value="39.0">39 km</quantity>
SE
<collectingCountry id="E1E9760EFE55FFCB7E8DF9461211DE9D" box="[943,1055,1717,1743]" name="Suriname" pageId="9" pageNumber="46">Suriname</collectingCountry>
river bridge, road to Redi Doti,
<quantity id="5E069B7BFE55FFCB7DB5F92F16DBDEA4" box="[151,213,1756,1782]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="46" unit="m" value="40.0">40 m</quantity>
,
<geoCoordinate id="FCCA5059FE55FFCB7DC3F92F1753DEA4" box="[225,349,1756,1782]" direction="north" orientation="latitude" pageId="9" pageNumber="46" precision="15" value="5.271389">5°16'17''N</geoCoordinate>
,
<geoCoordinate id="FCCA5059FE55FFCB7C48F92F17F6DEA4" box="[362,504,1756,1782]" direction="west" orientation="longitude" pageId="9" pageNumber="46" precision="15" value="-54.920834">54°55'15''W</geoCoordinate>
,
<date id="ED40105EFE55FFCB7F26F92F1486DEA4" box="[516,648,1756,1782]" pageId="9" pageNumber="46" value="1999-07-01">1.VII.1999</date>
, Z.H. Falin, SUR1F99 140 ex: uv light trap (2 exs.
<collectionCode id="FFEFAE5BFE55FFCB79C0F92F132EDEA4" box="[1250,1312,1756,1782]" pageId="9" pageNumber="46">KUL</collectionCode>
)
<collectingCountry id="E1E9760EFE55FFCB7DB5F8F0171BDF4F" box="[151,277,1795,1821]" name="Venezuela" pageId="9" pageNumber="46">Venezuela</collectingCountry>
: TF Amazonas, Cerro de la Neblina, basecamp,
<quantity id="5E069B7BFE55FFCB7E74F8F015AADF4F" box="[854,932,1795,1821]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="46" unit="m" value="140.0">140 m</quantity>
,
<geoCoordinate id="FCCA5059FE55FFCB7E92F8F0120DDF4F" box="[944,1027,1795,1821]" direction="north" orientation="latitude" pageId="9" pageNumber="46" precision="925" value="0.8333333">0°50'N</geoCoordinate>
<geoCoordinate id="FCCA5059FE55FFCB792EF8F0127ADF4F" box="[1036,1140,1795,1821]" direction="west" orientation="longitude" pageId="9" pageNumber="46" precision="925" value="-66.166664">66°10'W</geoCoordinate>
,
<date id="ED40105EFE55FFCB79A2F8F012FFDF4F" box="[1152,1265,1795,1821]" pageId="9" pageNumber="46" value="1985-01-27">27.I.1985</date>
&amp;
<date id="ED40105EFE55FFCB783BF8F01399DF4F" box="[1305,1431,1795,1821]" pageId="9" pageNumber="46" value="1985-02-19">19.II.1985</date>
, P.J. Spangler, P.M. Spangler, R. Faitoute, W.E. Steiner (33 exs
<collectionCode id="FFEFAE5BFE55FFCB7E59F8DA15B2DF11" box="[891,956,1833,1859]" pageId="9" pageNumber="46">CMT</collectionCode>
,
<collectionCode id="FFEFAE5BFE55FFCB7EE2F8DA1219DF11" LSID="urn:lsid:biocol.org:col:34871" box="[960,1047,1833,1859]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="46">USNM</collectionCode>
,
<collectionCode id="FFEFAE5BFE55FFCB7906F8DA126BDF11" box="[1060,1125,1833,1859]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="9" pageNumber="46">ZSM</collectionCode>
); Puerto Ayacucho,
<quantity id="5E069B7BFE55FFCB7873F8DA1392DF11" box="[1361,1436,1833,1859]" metricMagnitude="4" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="46" unit="km" value="37.0">37 km</quantity>
SE Rio Gavilan,
<date id="ED40105EFE55FFCB7C7FF8A317D4DF38" box="[349,474,1872,1898]" pageId="9" pageNumber="46" value="1986-02-20">20.II.1986</date>
, P.J. Spangler &amp; W. Sanchez (16 exs
<collectionCode id="FFEFAE5BFE55FFCB7EADF8A315E7DF38" LSID="urn:lsid:biocol.org:col:34871" box="[911,1001,1872,1898]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="46">USNM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE55FFCF7DE4F88517D5D992" lastPageId="13" lastPageNumber="50" pageId="9" pageNumber="46" type="diagnosis">
<paragraph id="9941369EFE55FFCB7DE4F88515E4DFE5" blockId="9.[151,1436,1330,1975]" pageId="9" pageNumber="46">
<emphasis id="AB8AEA8CFE55FFCB7DE4F8851748DFC2" bold="true" box="[198,326,1910,1936]" pageId="9" pageNumber="46">Diagnosis.</emphasis>
Habitus (
<figureCitation id="01C52A1BFE55FFCB7C9FF8841419DFC3" box="[445,535,1911,1937]" captionStart="FIGURES 28 – 39" captionStartId="8.[151,272,1875,1899]" captionTargetBox="[154,1429,467,1844]" captionTargetId="figure@8.[151,1436,467,1848]" captionTargetPageId="8" captionText="FIGURES 28 – 39. Subschematic body shape and colour patterns: 28) Laccodytes apalodes; 29) L. rondonia; 30) L. americanus; 31) L. phalacroides; 32) L. takutuanus; 33 a – 33 b) L. neblinae (variation of elytral patterns); 34) L. bassignanii; 35) L. obscuratus; 36) L. olibroides; 37) L. androginus; 38) Laccodytes sp. 1.; 39) Laccodytes sp. 2." httpUri="https://zenodo.org/record/193411/files/figure.png" pageId="9" pageNumber="46">Fig. 30</figureCitation>
). Body length 2.3–2.4 mm. Body slightly drop-shaped, tip of elytra almost truncate, but still with rounded angles.
<collectingCountry id="E1E9760EFE55FFCB7F42F86E1495DFE5" box="[608,667,1949,1975]" name="India" pageId="9" pageNumber="46">Hind</collectingCountry>
angle of pronotum rounded.
</paragraph>
<caption id="CD816616FE56FFC87DB5F8EC1273DFD2" httpUri="https://zenodo.org/record/193413/files/figure.png" pageId="10" pageNumber="47" targetBox="[180,1399,210,1765]" targetPageId="10">
<paragraph id="9941369EFE56FFC87DB5F8EC1273DFD2" blockId="10.[151,1436,1823,1921]" pageId="10" pageNumber="47">
<emphasis id="AB8AEA8CFE56FFC87DB5F8EC175BDF65" bold="true" box="[151,341,1823,1847]" pageId="10" pageNumber="47">FIGURES 44–47</emphasis>
. Male genitals in
<taxonomicName id="5EFE4D1DFE56FFC87F36F8EC1482DF64" box="[532,652,1823,1846]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="AB8AEA8CFE56FFC87F36F8EC1482DF64" box="[532,652,1823,1846]" italics="true" pageId="10" pageNumber="47">Laccodytes</emphasis>
</taxonomicName>
species:
<emphasis id="AB8AEA8CFE56FFC87FCEF8EC1509DF65" bold="true" box="[748,775,1823,1847]" pageId="10" pageNumber="47">44</emphasis>
)
<taxonomicName id="5EFE4D1DFE56FFC87E34F8EC159FDF64" box="[790,913,1823,1846]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="apalodes">
<emphasis id="AB8AEA8CFE56FFC87E34F8EC159FDF64" box="[790,913,1823,1846]" italics="true" pageId="10" pageNumber="47">L. apalodes</emphasis>
</taxonomicName>
:
<emphasis id="AB8AEA8CFE56FFC87EBDF8EC15A2DF65" bold="true" box="[927,940,1823,1847]" pageId="10" pageNumber="47">a</emphasis>
) median lobe of aedeagus in lateral view,
<emphasis id="AB8AEA8CFE56FFC87849F8EC139DDF65" bold="true" box="[1387,1427,1823,1847]" pageId="10" pageNumber="47">b–c</emphasis>
) right and left parameres;
<emphasis id="AB8AEA8CFE56FFC87C84F8B017CEDF09" bold="true" box="[422,448,1859,1883]" pageId="10" pageNumber="47">45</emphasis>
)
<taxonomicName id="5EFE4D1DFE56FFC87CF3F8B01441DF08" box="[465,591,1859,1882]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="rondonia">
<emphasis id="AB8AEA8CFE56FFC87CF3F8B01441DF08" box="[465,591,1859,1882]" italics="true" pageId="10" pageNumber="47">L. rondonia</emphasis>
</taxonomicName>
: idem;
<emphasis id="AB8AEA8CFE56FFC87F80F8B014B3DF09" bold="true" box="[674,701,1859,1883]" pageId="10" pageNumber="47">46</emphasis>
)
<taxonomicName id="5EFE4D1DFE56FFC87FEEF8B01569DF08" box="[716,871,1859,1882]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="AB8AEA8CFE56FFC87FEEF8B01569DF08" box="[716,871,1859,1882]" italics="true" pageId="10" pageNumber="47">L. americanus</emphasis>
</taxonomicName>
: idem;
<emphasis id="AB8AEA8CFE56FFC87E9BF8B015DADF09" bold="true" box="[953,980,1859,1883]" pageId="10" pageNumber="47">47</emphasis>
)
<taxonomicName id="5EFE4D1DFE56FFC87EC6F8B01285DF08" box="[996,1163,1859,1882]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="phalacroides">
<emphasis id="AB8AEA8CFE56FFC87EC6F8B01285DF08" box="[996,1163,1859,1882]" italics="true" pageId="10" pageNumber="47">L. phalacroides</emphasis>
</taxonomicName>
:
<emphasis id="AB8AEA8CFE56FFC879B8F8B012A9DF09" bold="true" box="[1178,1191,1859,1883]" pageId="10" pageNumber="47">a</emphasis>
) apical half of median lobe of aedeagus in dorsal view,
<emphasis id="AB8AEA8CFE56FFC87CD1F89A140CDFD3" bold="true" box="[499,514,1897,1921]" pageId="10" pageNumber="47">b</emphasis>
) median lobe in lateral view,
<emphasis id="AB8AEA8CFE56FFC87E1DF89A1569DFD3" bold="true" box="[831,871,1897,1921]" pageId="10" pageNumber="47">c–d</emphasis>
) right and left parameres;
</paragraph>
</caption>
<caption id="CD816616FE57FFC97DB5F9F41393DE99" httpUri="https://zenodo.org/record/193414/files/figure.png" pageId="11" pageNumber="48" targetBox="[189,1376,223,1499]" targetPageId="11">
<paragraph id="9941369EFE57FFC97DB5F9F41393DE99" blockId="11.[151,1437,1543,1740]" pageId="11" pageNumber="48">
<emphasis id="AB8AEA8CFE57FFC97DB5F9F41758DE4D" bold="true" box="[151,342,1543,1567]" pageId="11" pageNumber="48">FIGURES 48–53</emphasis>
. Male genitals in
<taxonomicName id="5EFE4D1DFE57FFC97F37F9FB1483DE4D" box="[533,653,1544,1567]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="AB8AEA8CFE57FFC97F37F9FB1483DE4D" box="[533,653,1544,1567]" italics="true" pageId="11" pageNumber="48">Laccodytes</emphasis>
</taxonomicName>
species:
<emphasis id="AB8AEA8CFE57FFC97FCCF9F41507DE4D" bold="true" box="[750,777,1543,1567]" pageId="11" pageNumber="48">48</emphasis>
)
<taxonomicName id="5EFE4D1DFE57FFC97E3BF9FB15A7DE4D" box="[793,937,1544,1567]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="takutuanus">
<emphasis id="AB8AEA8CFE57FFC97E3BF9FB15A7DE4D" box="[793,937,1544,1567]" italics="true" pageId="11" pageNumber="48">L. takutuanus</emphasis>
</taxonomicName>
:
<emphasis id="AB8AEA8CFE57FFC97E95F9F415CADE4D" bold="true" box="[951,964,1543,1567]" pageId="11" pageNumber="48">a</emphasis>
) median lobe of aedeagus in dorsal view,
<emphasis id="AB8AEA8CFE57FFC978A6F9F4139DDE4D" bold="true" box="[1412,1427,1543,1567]" pageId="11" pageNumber="48">b</emphasis>
) in lateral view,
<emphasis id="AB8AEA8CFE57FFC97C62F9D9176ADE10" bold="true" box="[320,356,1578,1602]" pageId="11" pageNumber="48">c-d</emphasis>
) right and left parameres;
<emphasis id="AB8AEA8CFE57FFC97FABF9D914AADE10" bold="true" box="[649,676,1578,1602]" pageId="11" pageNumber="48">49</emphasis>
)
<taxonomicName id="5EFE4D1DFE57FFC97F97F9D91521DE13" box="[693,815,1578,1601]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="neblinae">
<emphasis id="AB8AEA8CFE57FFC97F97F9D91521DE13" box="[693,815,1578,1601]" italics="true" pageId="11" pageNumber="48">L. neblinae</emphasis>
</taxonomicName>
:
<emphasis id="AB8AEA8CFE57FFC97E1DF9D91542DE10" bold="true" box="[831,844,1578,1602]" pageId="11" pageNumber="48">a</emphasis>
) median lobe of aedeagus in dorsal view,
<emphasis id="AB8AEA8CFE57FFC9783EF9D91325DE10" bold="true" box="[1308,1323,1578,1602]" pageId="11" pageNumber="48">b</emphasis>
) in lateral view,
<emphasis id="AB8AEA8CFE57FFC97DF5F9BF16F4DE36" bold="true" box="[215,250,1612,1636]" pageId="11" pageNumber="48">c-d</emphasis>
) right and left parameres;
<emphasis id="AB8AEA8CFE57FFC97F38F9BF143BDE36" bold="true" box="[538,565,1612,1636]" pageId="11" pageNumber="48">50</emphasis>
)
<taxonomicName id="5EFE4D1DFE57FFC97F66F9BE14D2DE36" box="[580,732,1613,1636]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="bassignanii">
<emphasis id="AB8AEA8CFE57FFC97F66F9BE14D2DE36" box="[580,732,1613,1636]" italics="true" pageId="11" pageNumber="48">L. bassignanii</emphasis>
</taxonomicName>
:
<emphasis id="AB8AEA8CFE57FFC97FC9F9BF14F6DE36" bold="true" box="[747,760,1612,1636]" pageId="11" pageNumber="48">a</emphasis>
) median lobe of aedeagus in dorsal view,
<emphasis id="AB8AEA8CFE57FFC9799DF9BF12ECDE36" bold="true" box="[1215,1250,1612,1636]" pageId="11" pageNumber="48">b-c</emphasis>
) in dorsal-lateral and lateral view,
<emphasis id="AB8AEA8CFE57FFC97C6EF99C1761DED5" bold="true" box="[332,367,1647,1671]" pageId="11" pageNumber="48">d-e</emphasis>
) right and left parameres;
<emphasis id="AB8AEA8CFE57FFC97FA8F99C14AADED5" bold="true" box="[650,676,1647,1671]" pageId="11" pageNumber="48">51</emphasis>
)
<taxonomicName id="5EFE4D1DFE57FFC97F96F983154BDED5" box="[692,837,1648,1671]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="obscuratus">
<emphasis id="AB8AEA8CFE57FFC97F96F983154BDED5" box="[692,837,1648,1671]" italics="true" pageId="11" pageNumber="48">L. obscuratus</emphasis>
</taxonomicName>
:
<emphasis id="AB8AEA8CFE57FFC97E71F99C156EDED5" bold="true" box="[851,864,1647,1671]" pageId="11" pageNumber="48">a</emphasis>
) median lobe of aedeagus in dorsal view,
<emphasis id="AB8AEA8CFE57FFC97802F99C1321DED5" bold="true" box="[1312,1327,1647,1671]" pageId="11" pageNumber="48">b</emphasis>
) in lateral view,
<emphasis id="AB8AEA8CFE57FFC97DF8F96116F1DEF8" bold="true" box="[218,255,1682,1706]" pageId="11" pageNumber="48">c-d</emphasis>
) right and left parameres;
<emphasis id="AB8AEA8CFE57FFC97F0EF9611446DEF8" bold="true" box="[556,584,1682,1706]" pageId="11" pageNumber="48">52</emphasis>
)
<taxonomicName id="5EFE4D1DFE57FFC97F7BF96114E9DEFB" box="[601,743,1682,1705]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="olibroides">
<emphasis id="AB8AEA8CFE57FFC97F7BF96114E9DEFB" box="[601,743,1682,1705]" italics="true" pageId="11" pageNumber="48">L. olibroides</emphasis>
</taxonomicName>
:
<emphasis id="AB8AEA8CFE57FFC97FD5F961150ADEF8" bold="true" box="[759,772,1682,1706]" pageId="11" pageNumber="48">a</emphasis>
) median lobe of aedeagus in lateral view,
<emphasis id="AB8AEA8CFE57FFC979C0F9611309DEF8" bold="true" box="[1250,1287,1682,1706]" pageId="11" pageNumber="48">b-c</emphasis>
) right an left parameres;
<emphasis id="AB8AEA8CFE57FFC97C30F9471722DE9E" bold="true" box="[274,300,1716,1740]" pageId="11" pageNumber="48">53</emphasis>
)
<taxonomicName id="5EFE4D1DFE57FFC97C19F94617DEDE9E" box="[315,464,1717,1740]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="androginus">
<emphasis id="AB8AEA8CFE57FFC97C19F94617DEDE9E" box="[315,464,1717,1740]" italics="true" pageId="11" pageNumber="48">L. androginus</emphasis>
</taxonomicName>
:
<emphasis id="AB8AEA8CFE57FFC97CFFF94717E4DE9E" bold="true" box="[477,490,1716,1740]" pageId="11" pageNumber="48">a</emphasis>
) median lobe of aedeagus in dorsal view,
<emphasis id="AB8AEA8CFE57FFC97E84F94715BBDE9E" bold="true" box="[934,949,1716,1740]" pageId="11" pageNumber="48">b</emphasis>
) in lateral view,
<emphasis id="AB8AEA8CFE57FFC97945F9471284DE9E" bold="true" box="[1127,1162,1716,1740]" pageId="11" pageNumber="48">c-d</emphasis>
) right and left parameres.
</paragraph>
</caption>
<caption id="CD816616FE50FFCE7DB5FBA0172EDCE2" httpUri="https://zenodo.org/record/193415/files/figure.png" pageId="12" pageNumber="49" targetBox="[153,1431,196,1081]" targetPageId="12">
<paragraph id="9941369EFE50FFCE7DB5FBA0172EDCE2" blockId="12.[151,1436,1107,1200]" pageId="12" pageNumber="49">
<emphasis id="AB8AEA8CFE50FFCE7DB5FBA01750DC39" bold="true" box="[151,350,1107,1131]" pageId="12" pageNumber="49">FIGURES 54–61</emphasis>
. Last abdominal segment (left male; right female) in
<taxonomicName id="5EFE4D1DFE50FFCE7E94FBA0123CDC38" box="[950,1074,1107,1130]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="AB8AEA8CFE50FFCE7E94FBA0123CDC38" box="[950,1074,1107,1130]" italics="true" pageId="12" pageNumber="49">Laccodytes</emphasis>
</taxonomicName>
species:
<emphasis id="AB8AEA8CFE50FFCE79B9FBA012B9DC39" bold="true" box="[1179,1207,1107,1131]" pageId="12" pageNumber="49">54</emphasis>
)
<taxonomicName id="5EFE4D1DFE50FFCE79E5FBA01347DC38" box="[1223,1353,1107,1130]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="apalodes">
<emphasis id="AB8AEA8CFE50FFCE79E5FBA01347DC38" box="[1223,1353,1107,1130]" italics="true" pageId="12" pageNumber="49">L. apalodes</emphasis>
</taxonomicName>
;
<emphasis id="AB8AEA8CFE50FFCE787BFBA0137BDC39" bold="true" box="[1369,1397,1107,1131]" pageId="12" pageNumber="49">55</emphasis>
)
<taxonomicName id="5EFE4D1DFE50FFCE78A4FBA016F6DCDF" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="rondonia">
<emphasis id="AB8AEA8CFE50FFCE78A4FBA016F6DCDF" italics="true" pageId="12" pageNumber="49">L. rondonia</emphasis>
</taxonomicName>
;
<emphasis id="AB8AEA8CFE50FFCE7C27FB861711DCDF" bold="true" box="[261,287,1141,1165]" pageId="12" pageNumber="49">56</emphasis>
)
<taxonomicName id="5EFE4D1DFE50FFCE7C0DFB8517C9DCDF" box="[303,455,1142,1165]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="AB8AEA8CFE50FFCE7C0DFB8517C9DCDF" box="[303,455,1142,1165]" italics="true" pageId="12" pageNumber="49">L. americanus</emphasis>
</taxonomicName>
;
<emphasis id="AB8AEA8CFE50FFCE7CF7FB8617FEDCDF" bold="true" box="[469,496,1141,1165]" pageId="12" pageNumber="49">57</emphasis>
)
<taxonomicName id="5EFE4D1DFE50FFCE7CDDFB8514ABDCDF" box="[511,677,1142,1165]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="phalacroides">
<emphasis id="AB8AEA8CFE50FFCE7CDDFB8514ABDCDF" box="[511,677,1142,1165]" italics="true" pageId="12" pageNumber="49">L. phalacroides</emphasis>
</taxonomicName>
;
<emphasis id="AB8AEA8CFE50FFCE7F91FB8614C0DCDF" bold="true" box="[691,718,1141,1165]" pageId="12" pageNumber="49">58</emphasis>
)
<taxonomicName id="5EFE4D1DFE50FFCE7FFFFB851560DCDF" box="[733,878,1142,1165]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="takutuanus">
<emphasis id="AB8AEA8CFE50FFCE7FFFFB851560DCDF" box="[733,878,1142,1165]" italics="true" pageId="12" pageNumber="49">L. takutuanus</emphasis>
</taxonomicName>
;
<emphasis id="AB8AEA8CFE50FFCE7E5EFB861599DCDF" bold="true" box="[892,919,1141,1165]" pageId="12" pageNumber="49">59</emphasis>
)
<taxonomicName id="5EFE4D1DFE50FFCE7E84FB851213DCDF" box="[934,1053,1142,1165]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="neblinae">
<emphasis id="AB8AEA8CFE50FFCE7E84FB851213DCDF" box="[934,1053,1142,1165]" italics="true" pageId="12" pageNumber="49">L. neblinae</emphasis>
</taxonomicName>
;
<emphasis id="AB8AEA8CFE50FFCE790EFB861248DCDF" bold="true" box="[1068,1094,1141,1165]" pageId="12" pageNumber="49">60</emphasis>
)
<taxonomicName id="5EFE4D1DFE50FFCE7977FB8512E2DCDF" box="[1109,1260,1142,1165]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="bassignanii">
<emphasis id="AB8AEA8CFE50FFCE7977FB8512E2DCDF" box="[1109,1260,1142,1165]" italics="true" pageId="12" pageNumber="49">L. bassignanii</emphasis>
</taxonomicName>
;
<emphasis id="AB8AEA8CFE50FFCE79D8FB86131BDCDF" bold="true" box="[1274,1301,1141,1165]" pageId="12" pageNumber="49">61</emphasis>
)
<taxonomicName id="5EFE4D1DFE50FFCE7806FB851392DCDF" box="[1316,1436,1142,1165]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="AB8AEA8CFE50FFCE7806FB851392DCDF" box="[1316,1436,1142,1165]" italics="true" pageId="12" pageNumber="49">Laccodytes</emphasis>
</taxonomicName>
sp. 1 female.
</paragraph>
</caption>
<paragraph id="9941369EFE50FFCE7DE4FB2D1248DE55" blockId="12.[151,1437,1246,2007]" pageId="12" pageNumber="49">
Color (
<figureCitation id="01C52A1BFE50FFCE7C02FB2D1771DCAA" box="[288,383,1246,1272]" captionStart="FIGURES 28 – 39" captionStartId="8.[151,272,1875,1899]" captionTargetBox="[154,1429,467,1844]" captionTargetId="figure@8.[151,1436,467,1848]" captionTargetPageId="8" captionText="FIGURES 28 – 39. Subschematic body shape and colour patterns: 28) Laccodytes apalodes; 29) L. rondonia; 30) L. americanus; 31) L. phalacroides; 32) L. takutuanus; 33 a – 33 b) L. neblinae (variation of elytral patterns); 34) L. bassignanii; 35) L. obscuratus; 36) L. olibroides; 37) L. androginus; 38) Laccodytes sp. 1.; 39) Laccodytes sp. 2." httpUri="https://zenodo.org/record/193411/files/figure.png" pageId="12" pageNumber="49">Fig. 30</figureCitation>
). Head ferruginous, sometimes slightly darkened laterally along the eyes. Pronotum yellow; base darker, middle of base dark brown to blackish, sometimes medially with dark band extending over disc of pronotum to reach the foremargin; foremargin also sometimes appearing slightly darkened; Venezuelan specimens only with middle of pronotal base blackish. Elytron blackish with contrasting yellow marks: a subbasal transverse band; usually a small to moderately large medio-discal patch; and an apical mark. The subbasal band reaches the margin and the anterior angle of elytron. The medio-discal patch is absent in some
<collectingCountry id="E1E9760EFE50FFCE7C6EFA3517A9DDB2" box="[332,423,1478,1504]" name="Guyana" pageId="12" pageNumber="49">Guyana</collectingCountry>
specimens, but usually visible; it is rather large in the Venezuelan individuals studied. Appendages and venter yellowish to ferruginous; metaventrite darker, brownish.
</paragraph>
<paragraph id="9941369EFE50FFCE7DE4F9E012E9DFDB" blockId="12.[151,1437,1246,2007]" pageId="12" pageNumber="49">
Sculpture. Head with MR consisting of small polygonal meshes; few punctures visible, the diameter of which equals that of the cells. Pronotum with very faint MR and numerous longitudinal lines, or shallow cuts. Elytron with MR as on head; no punctures visible but longitudinal lines which are, however, less distinct than on pronotum. SEM images reveal that the MR consists of networks composed of tiny serial cuticular holes and the lines are made by similar, but smaller holes, densely packed; also setae are surrounded by concentrically arranged tiny holes (
<figureCitation id="01C52A1BFE50FFCE7F15F92614B1DEBD" box="[567,703,1749,1775]" captionStart="FIGURES 17 – 25" captionStartId="4.[151,270,1905,1929]" captionTargetBox="[163,1426,187,1879]" captionTargetId="figure@4.[163,1428,187,1880]" captionTargetPageId="4" captionText="FIGURES 17 – 25. SEMs of MR in some Laccodytes species: 17) detail of elytron of L. americanus (magn. 2100 x); 18) idem, (magn. 2600 x); 19) metacoxa (MTX) and metaventrite (MTS) of L. americanus; 20) idem, detail at higher magnification; 21) idem, detail of metaventrite (MTX) at higher magnification; 22) detail of elytron of L. apalodes; 23) detail of metacoxa of L. apalodes; 24) idem, detail at higher magnification; 25) detail of metacoxa of L. neblinae." httpUri="https://zenodo.org/record/193409/files/figure.png" pageId="12" pageNumber="49">Figs 17, 18</figureCitation>
). Metaventrite rather shiny, with faint MR and few punctures; postcoxal process with MR; metacoxal plates with numerous rather diagonally orientated irregular longish cells (
<date id="ED40105EFE50FFCE7DFFF8D11700DF6E" box="[221,270,1826,1852]" pageId="12" pageNumber="49" value="1940-10">40x</date>
), giving the impression of a dense diagonal striation (
<figureCitation id="01C52A1BFE50FFCE7EA0F8D11202DF6E" box="[898,1036,1826,1852]" captionStart="FIGURES 17 – 25" captionStartId="4.[151,270,1905,1929]" captionTargetBox="[163,1426,187,1879]" captionTargetId="figure@4.[163,1428,187,1880]" captionTargetPageId="4" captionText="FIGURES 17 – 25. SEMs of MR in some Laccodytes species: 17) detail of elytron of L. americanus (magn. 2100 x); 18) idem, (magn. 2600 x); 19) metacoxa (MTX) and metaventrite (MTS) of L. americanus; 20) idem, detail at higher magnification; 21) idem, detail of metaventrite (MTX) at higher magnification; 22) detail of elytron of L. apalodes; 23) detail of metacoxa of L. apalodes; 24) idem, detail at higher magnification; 25) detail of metacoxa of L. neblinae." httpUri="https://zenodo.org/record/193409/files/figure.png" pageId="12" pageNumber="49">Figs 19, 20</figureCitation>
). The SEM (540x) images reveals indeed long lines, which actually consist of tiny holes (
<date id="ED40105EFE50FFCE7E4CF8BA15CBDF31" box="[878,965,1865,1891]" pageId="12" pageNumber="49" value="1830-10">1830x</date>
); the same is true for the meshes on metaventrite, which are also in fact networks composed of tiny serial cuticular holes (
<figureCitation id="01C52A1BFE50FFCE79A3F89C12D7DFDB" box="[1153,1241,1903,1929]" captionStart="FIGURES 17 – 25" captionStartId="4.[151,270,1905,1929]" captionTargetBox="[163,1426,187,1879]" captionTargetId="figure@4.[163,1428,187,1880]" captionTargetPageId="4" captionText="FIGURES 17 – 25. SEMs of MR in some Laccodytes species: 17) detail of elytron of L. americanus (magn. 2100 x); 18) idem, (magn. 2600 x); 19) metacoxa (MTX) and metaventrite (MTS) of L. americanus; 20) idem, detail at higher magnification; 21) idem, detail of metaventrite (MTX) at higher magnification; 22) detail of elytron of L. apalodes; 23) detail of metacoxa of L. apalodes; 24) idem, detail at higher magnification; 25) detail of metacoxa of L. neblinae." httpUri="https://zenodo.org/record/193409/files/figure.png" pageId="12" pageNumber="49">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="9941369EFE50FFCF7DE4F86515C8D8AD" blockId="12.[151,1437,1246,2007]" lastBlockId="13.[151,1437,152,603]" lastPageId="13" lastPageNumber="50" pageId="12" pageNumber="49">
Structures. Pronotum with narrow lateral bead. Prosternum and base of prosternal process with small ridge; prosternal process broadly carinate and with a short, needle-shaped, acute tip (
<figureCitation id="01C52A1BFE50FFCE79AFF84E12D8DF85" box="[1165,1238,1981,2007]" captionStart="FIGURES 3 – 16" captionStartId="3.[151,270,1860,1884]" captionTargetBox="[248,1343,197,1819]" captionTargetId="figure@3.[248,1348,197,1827]" captionTargetPageId="3" captionText="FIGURES 3 – 16. SEMs of some Laccodytes species structures: 3) segments 1 - 3 in male protarsus of L. americanus; 4) prosternal process of L. apalodes; 5) idem, L. americanus; 6) idem, L. neblinae; 7) metaventrite of L. apalodes, the arrow indicates the groove in which the tip of prosternal process fits; 8) idem, L. neblinae; 9) metacoxal process of L. apalodes; 10) idem, L. americanus; 11) idem, L. neblinae; 12) hind tarsus of L. apalodes (male); 13) idem, L. americanus (male); 14) idem, L. neblinae (female); 15) ovipositor of L. apalodes; 16) idem, L. americanus." httpUri="https://zenodo.org/record/193408/files/figure.png" pageId="12" pageNumber="49">Fig. 5</figureCitation>
). Elytra apically rounded. Epipleuron broad up to level of sternite 6 or 7.
<collectingCountry id="E1E9760EFE51FFCF7E04FF6B156FD8E0" box="[806,865,152,178]" name="India" pageId="13" pageNumber="50">Hind</collectingCountry>
margin of sternite 7 different in male and female (see below). Metatarsomeres 1–4 with apicolateral angle distinctly lobed (
<figureCitation id="01C52A1BFE51FFCF792BFF4D126FD88A" box="[1033,1121,190,216]" captionStart="FIGURES 3 – 16" captionStartId="3.[151,270,1860,1884]" captionTargetBox="[248,1343,197,1819]" captionTargetId="figure@3.[248,1348,197,1827]" captionTargetPageId="3" captionText="FIGURES 3 – 16. SEMs of some Laccodytes species structures: 3) segments 1 - 3 in male protarsus of L. americanus; 4) prosternal process of L. apalodes; 5) idem, L. americanus; 6) idem, L. neblinae; 7) metaventrite of L. apalodes, the arrow indicates the groove in which the tip of prosternal process fits; 8) idem, L. neblinae; 9) metacoxal process of L. apalodes; 10) idem, L. americanus; 11) idem, L. neblinae; 12) hind tarsus of L. apalodes (male); 13) idem, L. americanus (male); 14) idem, L. neblinae (female); 15) ovipositor of L. apalodes; 16) idem, L. americanus." httpUri="https://zenodo.org/record/193408/files/figure.png" pageId="13" pageNumber="50">Fig. 13</figureCitation>
).
<collectingCountry id="E1E9760EFE51FFCF795AFF4D12BDD88A" box="[1144,1203,190,216]" name="India" pageId="13" pageNumber="50">Hind</collectingCountry>
lobes of metacoxal process straight, truncate, with only a slight medial incision (
<figureCitation id="01C52A1BFE51FFCF7E43FF1615B7D8AD" box="[865,953,229,255]" captionStart="FIGURES 3 – 16" captionStartId="3.[151,270,1860,1884]" captionTargetBox="[248,1343,197,1819]" captionTargetId="figure@3.[248,1348,197,1827]" captionTargetPageId="3" captionText="FIGURES 3 – 16. SEMs of some Laccodytes species structures: 3) segments 1 - 3 in male protarsus of L. americanus; 4) prosternal process of L. apalodes; 5) idem, L. americanus; 6) idem, L. neblinae; 7) metaventrite of L. apalodes, the arrow indicates the groove in which the tip of prosternal process fits; 8) idem, L. neblinae; 9) metacoxal process of L. apalodes; 10) idem, L. americanus; 11) idem, L. neblinae; 12) hind tarsus of L. apalodes (male); 13) idem, L. americanus (male); 14) idem, L. neblinae (female); 15) ovipositor of L. apalodes; 16) idem, L. americanus." httpUri="https://zenodo.org/record/193408/files/figure.png" pageId="13" pageNumber="50">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="9941369EFE51FFCF7DE4FEFF1282D921" blockId="13.[151,1437,152,603]" pageId="13" pageNumber="50">
Male. Pro- and mesotarsi not dilated.
<collectingCountry id="E1E9760EFE51FFCF7FA5FEFF14CCD974" box="[647,706,268,294]" name="India" pageId="13" pageNumber="50">Hind</collectingCountry>
margin of sternite 7 slightly concave in the middle (
<figureCitation id="01C52A1BFE51FFCF7817FEFF1380D974" box="[1333,1422,268,294]" captionStart="FIGURES 54 – 61" captionStartId="12.[151,273,1107,1131]" captionTargetBox="[153,1431,196,1081]" captionTargetId="figure@12.[151,1436,194,1083]" captionTargetPageId="12" captionText="FIGURES 54 – 61. Last abdominal segment (left male; right female) in Laccodytes species: 54) L. apalodes; 55) L. rondonia; 56) L. americanus; 57) L. phalacroides; 58) L. takutuanus; 59) L. neblinae; 60) L. bassignanii; 61) Laccodytes sp. 1 female." httpUri="https://zenodo.org/record/193415/files/figure.png" pageId="13" pageNumber="50">Fig. 56</figureCitation>
). Aedeagus (
<figureCitation id="01C52A1BFE51FFCF7C03FEC1178ED91E" box="[289,384,306,332]" captionStart="FIGURES 44 – 47" captionStartId="10.[151,270,1823,1847]" captionTargetBox="[180,1399,210,1765]" captionTargetId="figure@10.[171,1415,197,1798]" captionTargetPageId="10" captionText="FIGURES 44 – 47. Male genitals in Laccodytes species: 44) L. apalodes: a) median lobe of aedeagus in lateral view, b – c) right and left parameres; 45) L. rondonia: idem; 46) L. americanus: idem; 47) L. phalacroides: a) apical half of median lobe of aedeagus in dorsal view, b) median lobe in lateral view, c – d) right and left parameres;" httpUri="https://zenodo.org/record/193413/files/figure.png" pageId="13" pageNumber="50">Fig. 46</figureCitation>
): median lobe apically with a slender, flat process, as long as about 1/3 of its length. Parameres visibly different in sizes, both club-shaped, apparently without apical setae.
</paragraph>
<paragraph id="9941369EFE51FFCF7DE4FE7317D5D992" blockId="13.[151,1437,152,603]" pageId="13" pageNumber="50">
Female. Sternite 7 (
<figureCitation id="01C52A1BFE51FFCF7C92FE731409D9C8" box="[432,519,384,410]" captionStart="FIGURES 54 – 61" captionStartId="12.[151,273,1107,1131]" captionTargetBox="[153,1431,196,1081]" captionTargetId="figure@12.[151,1436,194,1083]" captionTargetPageId="12" captionText="FIGURES 54 – 61. Last abdominal segment (left male; right female) in Laccodytes species: 54) L. apalodes; 55) L. rondonia; 56) L. americanus; 57) L. phalacroides; 58) L. takutuanus; 59) L. neblinae; 60) L. bassignanii; 61) Laccodytes sp. 1 female." httpUri="https://zenodo.org/record/193415/files/figure.png" pageId="13" pageNumber="50">Fig. 56</figureCitation>
) normally rounded on both sides, hind margin with a narrow, deep U-shaped emargination in the middle.
</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE51FFCF7DE4FE3E1209D9B5" box="[198,1031,461,487]" pageId="13" pageNumber="50" type="distribution">
<paragraph id="9941369EFE51FFCF7DE4FE3E1209D9B5" blockId="13.[151,1437,152,603]" box="[198,1031,461,487]" pageId="13" pageNumber="50">
<emphasis id="AB8AEA8CFE51FFCF7DE4FE3E1751D9B5" bold="true" box="[198,351,461,487]" pageId="13" pageNumber="50">Distribution</emphasis>
(
<figureCitation id="01C52A1BFE51FFCF7C4DFE3E17C9D9B5" box="[367,455,461,487]" captionStart="FIGURE 62" captionStartId="18.[151,255,1941,1965]" captionText="FIGURE 62. Known distribution of Laccodytes species." pageId="13" pageNumber="50">Fig. 62</figureCitation>
). French
<collectingCountry id="E1E9760EFE51FFCF7F16FE3E149CD9B5" box="[564,658,461,487]" name="Guyana" pageId="13" pageNumber="50">Guyana</collectingCountry>
,
<collectingCountry id="E1E9760EFE51FFCF7FBCFE3E14F2D9B5" box="[670,764,461,487]" name="Guyana" pageId="13" pageNumber="50">Guyana</collectingCountry>
,
<collectingCountry id="E1E9760EFE51FFCF7E2AFE3E1575D9B5" box="[776,891,461,487]" name="Suriname" pageId="13" pageNumber="50">Suriname</collectingCountry>
,
<collectingCountry id="E1E9760EFE51FFCF7EA5FE3E120DD9B5" box="[903,1027,461,487]" name="Venezuela" pageId="13" pageNumber="50">Venezuela</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE51FFCF7DE4FE001220DA09" pageId="13" pageNumber="50" type="biology_ecology">
<paragraph id="9941369EFE51FFCF7DE4FE001220DA09" blockId="13.[151,1437,152,603]" pageId="13" pageNumber="50">
<emphasis id="AB8AEA8CFE51FFCF7DE4FE001724DA5F" bold="true" box="[198,298,499,525]" pageId="13" pageNumber="50">Biology.</emphasis>
In
<collectingCountry id="E1E9760EFE51FFCF7C70FE0717BEDA5C" box="[338,432,500,526]" name="Guyana" pageId="13" pageNumber="50">Guyana</collectingCountry>
, the species was collected at blacklight in a forest clearing, close to a stream, and in a stream. Venezuelan specimens were seined from rocks in the rapids of Rio Baria, some
<specimenCount id="8FF8FD17FE51FFCF799AFDE91354DA66" box="[1208,1370,538,564]" pageId="13" pageNumber="50" type="generic">30 specimens</specimenCount>
from Cerro de la Neblina were picked by hand from leaf packs taken from a stream.
</paragraph>
</subSubSection>
</treatment>
</document>
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