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<document id="92A5E76B193884AE8624B984255C1130" ID-DOI="10.5281/zenodo.193406" ID-GBIF-Dataset="5e8043a5-e685-4a21-b696-0f7a706ab223" ID-ISSN="1175-5326" ID-Zenodo-Dep="193406" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460138703176" checkinUser="plazi" docAuthor="Toledo, Mario, Spangler, Paul J. &amp; Balke, Michael" docDate="2010" docId="11578788FE4FFFD17DB5FE8F1425DD5D" docLanguage="en" docName="zt02347p058.pdf" docOrigin="Zootaxa 2347" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Laccodytes" docType="treatment" docVersion="9" lastPageNumber="56" masterDocId="ED6EFFF0FE5CFFC27D22FFF3160ED852" masterDocTitle="Taxonomic revision of the Neotropical diving beetles genus Laccodytes Régimbart, 1895 (Coleoptera: Dytiscidae)" masterLastPageNumber="58" masterPageNumber="37" pageNumber="56" updateTime="1698235333712" updateUser="plazi">
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<mods:title id="F40AF2C3F267CE0F9B709A8DC7F18082">Taxonomic revision of the Neotropical diving beetles genus Laccodytes Régimbart, 1895 (Coleoptera: Dytiscidae)</mods:title>
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<mods:namePart id="46DA92F01627F826D8C823748D2ECB8F">Toledo, Mario</mods:namePart>
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<mods:namePart id="E66B82B5D2FBB19BCA38C2BC985706C1">Spangler, Paul J.</mods:namePart>
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<mods:namePart id="D4507D1A36DA4640CF1590756910AED5">Balke, Michael</mods:namePart>
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<mods:title id="8C26E48F2DD740EED0AC3B9B29E8E751">Zootaxa</mods:title>
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<mods:date id="69FABEB9589E213BA7912BC982103193">2010</mods:date>
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<treatment id="11578788FE4FFFD17DB5FE8F1425DD5D" ID-DOI="http://doi.org/10.5281/zenodo.6198394" ID-GBIF-Taxon="124955821" ID-Zenodo-Dep="6198394" LSID="urn:lsid:plazi:treatment:11578788FE4FFFD17DB5FE8F1425DD5D" httpUri="http://treatment.plazi.org/id/11578788FE4FFFD17DB5FE8F1425DD5D" lastPageNumber="56" pageId="19" pageNumber="56">
<subSubSection id="D1E46515FE4FFFD17DB5FE8F1757D9EE" pageId="19" pageNumber="56" type="nomenclature">
<paragraph id="9941369EFE4FFFD17DB5FE8F1753D9C7" blockId="19.[151,349,379,444]" box="[151,349,379,406]" pageId="19" pageNumber="56">
<heading id="C20981F2FE4FFFD17DB5FE8F1753D9C7" bold="true" box="[151,349,379,406]" fontSize="11" level="1" pageId="19" pageNumber="56" reason="1">
<emphasis id="AB8AEA8CFE4FFFD17DB5FE8F1753D9C7" bold="true" box="[151,349,379,406]" pageId="19" pageNumber="56">
<taxonomicName id="5EFE4D1DFE4FFFD17DB5FE8F1713D9C4" ID-CoL="59RJ" box="[151,285,380,406]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="AB8AEA8CFE4FFFD17DB5FE8F1713D9C4" bold="true" box="[151,285,380,406]" italics="true" pageId="19" pageNumber="56">Laccodytes</emphasis>
</taxonomicName>
sp. 1
</emphasis>
</heading>
</paragraph>
<paragraph id="9941369EFE4FFFD17DB5FE511757D9EE" blockId="19.[151,349,379,444]" box="[151,345,418,444]" pageId="19" pageNumber="56">Figs (38, 61, 62)</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE4FFFD17DB5FE1C1313DC90" pageId="19" pageNumber="56" type="materials_examined">
<paragraph id="9941369EFE4FFFD17DB5FE1C15D3DA62" blockId="19.[151,1436,495,1295]" pageId="19" pageNumber="56">
<emphasis id="AB8AEA8CFE4FFFD17DB5FE1C179DDA5B" bold="true" box="[151,403,495,521]" pageId="19" pageNumber="56">Material examined.</emphasis>
1 Ƥ (
<collectionCode id="FFEFAE5BFE4FFFD17CF8FE031424DA58" box="[474,554,496,522]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="19" pageNumber="56">NMW</collectionCode>
)
<collectingCountry id="E1E9760EFE4FFFD17F19FE0314B2DA58" box="[571,700,496,522]" name="Venezuela" pageId="19" pageNumber="56">Venezuela</collectingCountry>
: Bolivar, Quebrada, near Paraitepui Village, stream crossing tourist pathway to Mt. Roraima,
<quantity id="5E069B7BFE4FFFD17F34FDE5147CDA62" box="[534,626,534,560]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="56" unit="m" value="1000.0">1000 m</quantity>
,
<date id="ED40105EFE4FFFD17F5FFDE514FEDA62" box="[637,752,534,560]" pageId="19" pageNumber="56" value="1999-01-31">31.I.1999</date>
, Derka &amp; Reitmeier
</paragraph>
<paragraph id="9941369EFE4FFFD17DE4FDCE16F4DAA0" blockId="19.[151,1436,495,1295]" pageId="19" pageNumber="56">
<emphasis id="AB8AEA8CFE4FFFD17DE4FDCE17D7DA05" bold="true" box="[198,473,573,599]" pageId="19" pageNumber="56">Provisional diagnosis.</emphasis>
Habitus (
<figureCitation id="01C52A1BFE4FFFD17F73FDCE14A5DA05" box="[593,683,573,599]" captionStart="FIGURES 28 39" captionStartId="8.[151,272,1875,1899]" captionTargetBox="[154,1429,467,1844]" captionTargetId="figure@8.[151,1436,467,1848]" captionTargetPageId="8" captionText="FIGURES 28 39. Subschematic body shape and colour patterns: 28) Laccodytes apalodes; 29) L. rondonia; 30) L. americanus; 31) L. phalacroides; 32) L. takutuanus; 33 a 33 b) L. neblinae (variation of elytral patterns); 34) L. bassignanii; 35) L. obscuratus; 36) L. olibroides; 37) L. androginus; 38) Laccodytes sp. 1.; 39) Laccodytes sp. 2." httpUri="https://zenodo.org/record/193411/files/figure.png" pageId="19" pageNumber="56">Fig. 38</figureCitation>
). Similar to
<taxonomicName id="5EFE4D1DFE4FFFD17E66FDCE15E3DA04" box="[836,1005,573,598]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="56" phylum="Arthropoda" rank="species" species="bassignanii" status="sp. nov.">
<emphasis id="AB8AEA8CFE4FFFD17E66FDCE15E3DA04" box="[836,1005,573,598]" italics="true" pageId="19" pageNumber="56">L. bassignanii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0B957F7FE4FFFD17ED5FDCE1221DA05" box="[1015,1071,573,599]" pageId="19" pageNumber="56" rank="species">n.sp.</taxonomicNameLabel>
and
<taxonomicName id="5EFE4D1DFE4FFFD1794FFDCE131EDA04" box="[1133,1296,573,598]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="56" phylum="Arthropoda" rank="species" species="obscuratus" status="sp. nov.">
<emphasis id="AB8AEA8CFE4FFFD1794FFDCE131EDA04" box="[1133,1296,573,598]" italics="true" pageId="19" pageNumber="56">L. obscuratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0B957F7FE4FFFD17838FDCE135DDA05" box="[1306,1363,573,599]" pageId="19" pageNumber="56" rank="species">n.sp.</taxonomicNameLabel>
, from which it is distinguished by the extension of the orange patterns on the elytra and the sharp contrast of these with the black background; from
<taxonomicName id="5EFE4D1DFE4FFFD17F39FD7914B5DAF1" box="[539,699,650,675]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="56" phylum="Arthropoda" rank="species" species="obscuratus" status="sp. nov.">
<emphasis id="AB8AEA8CFE4FFFD17F39FD7914B5DAF1" box="[539,699,650,675]" italics="true" pageId="19" pageNumber="56">L. obscuratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0B957F7FE4FFFD17FE3FD7914F6DAF6" box="[705,760,650,676]" pageId="19" pageNumber="56" rank="species">n.sp.</taxonomicNameLabel>
also by the pale head. Body length 1.7 mm. Body slightly drop-shaped, tip of elytron rounded.
<collectingCountry id="E1E9760EFE4FFFD17F73FD421483DA99" box="[593,653,689,715]" name="India" pageId="19" pageNumber="56">Hind</collectingCountry>
angle of pronotum rounded. Angle between pronotum and elytra missing.
</paragraph>
<paragraph id="9941369EFE4FFFD17DE4FD0D14D3DBDE" blockId="19.[151,1436,495,1295]" pageId="19" pageNumber="56">
Color. Head orange. Pronotum orange, narrow band along base blackish. Elytron blackish with sharply contrasting orange markings: a broad subbasal band, that hardly reaches margin and does not reach suture; the apex, reaching the last third of elytra and including the subapical patches (
<figureCitation id="01C52A1BFE4FFFD17922FCBF1256DB34" box="[1024,1112,844,870]" captionStart="FIGURES 28 39" captionStartId="8.[151,272,1875,1899]" captionTargetBox="[154,1429,467,1844]" captionTargetId="figure@8.[151,1436,467,1848]" captionTargetPageId="8" captionText="FIGURES 28 39. Subschematic body shape and colour patterns: 28) Laccodytes apalodes; 29) L. rondonia; 30) L. americanus; 31) L. phalacroides; 32) L. takutuanus; 33 a 33 b) L. neblinae (variation of elytral patterns); 34) L. bassignanii; 35) L. obscuratus; 36) L. olibroides; 37) L. androginus; 38) Laccodytes sp. 1.; 39) Laccodytes sp. 2." httpUri="https://zenodo.org/record/193411/files/figure.png" pageId="19" pageNumber="56">Fig. 38</figureCitation>
). Body appendages orange. Venter orange, metaventrite and metacoxa darker.
</paragraph>
<paragraph id="9941369EFE4FFFD17DE4FC6A143DDC52" blockId="19.[151,1436,495,1295]" pageId="19" pageNumber="56">Sculpture. MR as in the preceding three species; metaventrite with fine and sparse dots, becoming closer and deeper on metasternal lateral expansion a. A series of transverse shallow grooves is visible at about the anterior fourth of metacoxal plates.</paragraph>
<paragraph id="9941369EFE4FFFD17DE4FBFE14E2DCC9" blockId="19.[151,1436,495,1295]" pageId="19" pageNumber="56">
Structures. Pronotum with narrow lateral bead; posterior angle rounded. Prosternum and prosternal process with ridge; prosternal process broadly carinate and with a long, acute tip. Epipleuron broad up to level of sternite 7. Metatarsomeres 14 with apico-lateral angle slightly lobed.
<collectingCountry id="E1E9760EFE4FFFD17933FBA91243DC26" box="[1041,1101,1114,1140]" name="India" pageId="19" pageNumber="56">Hind</collectingCountry>
lobes of metacoxal process rather straight, with a deep medial V-shaped notch.
</paragraph>
<paragraph id="9941369EFE4FFFD17DE4FB5B1313DC90" blockId="19.[151,1436,495,1295]" box="[198,1309,1192,1218]" pageId="19" pageNumber="56">
<collectingCountry id="E1E9760EFE4FFFD17DE4FB5B170FDC90" box="[198,257,1192,1218]" name="India" pageId="19" pageNumber="56">Hind</collectingCountry>
margin of sternite 7 (
<figureCitation id="01C52A1BFE4FFFD17CDCFB5B145DDC90" box="[510,595,1192,1218]" captionStart="FIGURES 54 61" captionStartId="12.[151,273,1107,1131]" captionTargetBox="[153,1431,196,1081]" captionTargetId="figure@12.[151,1436,194,1083]" captionTargetPageId="12" captionText="FIGURES 54 61. Last abdominal segment (left male; right female) in Laccodytes species: 54) L. apalodes; 55) L. rondonia; 56) L. americanus; 57) L. phalacroides; 58) L. takutuanus; 59) L. neblinae; 60) L. bassignanii; 61) Laccodytes sp. 1 female." httpUri="https://zenodo.org/record/193415/files/figure.png" pageId="19" pageNumber="56">Fig. 61</figureCitation>
) forming a V-shaped emargination, as in
<taxonomicName id="5EFE4D1DFE4FFFD1791BFB5B12D1DC93" box="[1081,1247,1192,1217]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="56" phylum="Arthropoda" rank="species" species="bassignanii" status="sp. nov.">
<emphasis id="AB8AEA8CFE4FFFD1791BFB5B12D1DC93" box="[1081,1247,1192,1217]" italics="true" pageId="19" pageNumber="56">L. bassignanii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0B957F7FE4FFFD179C4FB5B1313DC90" box="[1254,1309,1192,1218]" pageId="19" pageNumber="56" rank="species">n.sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE4FFFD17DE4FB3D128EDCBA" box="[198,1152,1230,1256]" pageId="19" pageNumber="56" type="distribution">
<paragraph id="9941369EFE4FFFD17DE4FB3D128EDCBA" blockId="19.[151,1436,495,1295]" box="[198,1152,1230,1256]" pageId="19" pageNumber="56">
<emphasis id="AB8AEA8CFE4FFFD17DE4FB3D1751DCBA" bold="true" box="[198,351,1230,1256]" pageId="19" pageNumber="56">Distribution</emphasis>
(
<figureCitation id="01C52A1BFE4FFFD17C4DFB3D17C9DCBA" box="[367,455,1230,1256]" captionStart="FIGURE 62" captionStartId="18.[151,255,1941,1965]" captionText="FIGURE 62. Known distribution of Laccodytes species." pageId="19" pageNumber="56">Fig. 62</figureCitation>
). Southeastern
<collectingCountry id="E1E9760EFE4FFFD17F58FB3D14FBDCBA" box="[634,757,1230,1256]" name="Venezuela" pageId="19" pageNumber="56">Venezuela</collectingCountry>
, close to the border with
<collectingCountry id="E1E9760EFE4FFFD1793FFB3D1272DCBA" box="[1053,1148,1230,1256]" name="Guyana" pageId="19" pageNumber="56">Guyana</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE4FFFD17DE4FB061425DD5D" box="[198,555,1269,1295]" pageId="19" pageNumber="56" type="biology_ecology">
<paragraph id="9941369EFE4FFFD17DE4FB061425DD5D" blockId="19.[151,1436,495,1295]" box="[198,555,1269,1295]" pageId="19" pageNumber="56">
<emphasis id="AB8AEA8CFE4FFFD17DE4FB061724DD5D" bold="true" box="[198,298,1269,1295]" pageId="19" pageNumber="56">Biology.</emphasis>
Taken from a stream.
</paragraph>
</subSubSection>
</treatment>
</document>