465 lines
68 KiB
XML
465 lines
68 KiB
XML
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<mods:title id="DE545162FC637773A0541E7A7789D780">Revision of the genus Halys (Hemiptera: Pentatomidae) with description of a new species from India</mods:title>
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<treatment id="0C3D87C22239E37DFF0F23CEB151FEF0" ID-GBIF-Taxon="156788545" LSID="urn:lsid:plazi:treatment:0C3D87C22239E37DFF0F23CEB151FEF0" httpUri="http://treatment.plazi.org/id/0C3D87C22239E37DFF0F23CEB151FEF0" lastPageId="19" lastPageNumber="369" pageId="16" pageNumber="367">
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<subSubSection id="CC8E655F2239E37EFF0F23CEB02DFE6E" box="[151,539,481,507]" pageId="16" pageNumber="367" type="nomenclature">
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<paragraph id="842B36D42239E37EFF0F23CEB02DFE6E" blockId="16.[151,539,481,541]" box="[151,539,481,507]" pageId="16" pageNumber="367">
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<heading id="DF6381B82239E37EFF0F23CEB02DFE6E" bold="true" box="[151,539,481,507]" fontSize="11" level="1" pageId="16" pageNumber="367" reason="1">
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<taxonomicName id="43944D572239E37EFF0F23CEB02DFE6E" authority="(Thunberg, 1783)" baseAuthorityName="Thunberg" baseAuthorityYear="1783" box="[151,539,481,507]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="sulcata">
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<emphasis id="B6E0EAC62239E37EFF0F23CEB02DFE6E" bold="true" box="[151,539,481,507]" pageId="16" pageNumber="367">
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<emphasis id="B6E0EAC62239E37EFF0F23CEB30CFE6E" bold="true" box="[151,314,481,507]" italics="true" pageId="16" pageNumber="367">Halys sulcata</emphasis>
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(
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<bibRefCitation id="E0054B252239E37EFED123CEB025FE6E" author="Thunberg, C. P." box="[329,531,481,507]" pageId="16" pageNumber="367" pagination="29 - 52" refId="ref14448" refString="Thunberg, C. P. (1783) Dissertatio entomologica novas insectorum species, sistens, cujus partem secundam, cons. exper. facult. med. upsal. publice ventilandam exhibent Johan. Edman, Upsala, 22 pp. [pp. 29 - 52]" type="book chapter" year="1783">Thunberg, 1783</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="CC8E655F2239E37EFF0F202BB33DFD88" box="[151,267,516,541]" pageId="16" pageNumber="367" type="description">
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<paragraph id="842B36D42239E37EFF0F202BB33DFD88" blockId="16.[151,539,481,541]" box="[151,267,516,541]" pageId="16" pageNumber="367">
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(
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<figureCitation id="1CAF2A512239E37EFF07202BB334FD88" box="[159,258,516,541]" captionStart="FIGURES 1–9" captionStartId="4.[151,264,1939,1962]" captionTargetBox="[246,1377,251,1915]" captionTargetId="figure@4.[183,1409,193,1927]" captionTargetPageId="4" captionText="FIGURES 1–9. Halys Fabricius, 1803 (dorsal and ventral habitus). 1–2, Halys mudigerensis sp. nov. (male); 3, H. mudigerensis sp. nov. (female without abdomen); 4–5, H. serrigera Westwood, 1837; 6–7, H. shaista Ghauri, 1988; 8–9, H. sulcata (Thunberg, 1783)" httpUri="https://zenodo.org/record/2644556/files/figure.png" pageId="16" pageNumber="367">Figs 8, 9</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="CC8E655F2239E37EFF0F2062B17BFB5B" pageId="16" pageNumber="367" type="reference_group">
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<paragraph id="842B36D42239E37EFF0F2062B151FC01" blockId="16.[151,1055,588,981]" pageId="16" pageNumber="367">
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<treatmentCitationGroup id="A48411FA2239E37EFF0F2062B151FC01" pageId="16" pageNumber="367">
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<taxonomicName id="43944D572239E37EFF0F2062B034FDF7" authority="Thunberg, 1783: 43" authorityName="Thunberg" authorityPageNumber="43" authorityYear="1783" box="[151,514,588,610]" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="sulcatus">
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<emphasis id="B6E0EAC62239E37EFF0F2062B307FDF7" box="[151,305,588,610]" italics="true" pageId="16" pageNumber="367">Cimex sulcatus</emphasis>
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<treatmentCitation id="053510C52239E37EFEAF2063B034FDF7" author="Thunberg, C. P." box="[311,514,588,610]" page="43" pageId="16" pageNumber="367" year="1783">
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<bibRefCitation id="E0054B252239E37EFEAF2063B3EAFDF7" author="Thunberg, C. P." box="[311,476,588,610]" pageId="16" pageNumber="367" pagination="29 - 52" refId="ref14448" refString="Thunberg, C. P. (1783) Dissertatio entomologica novas insectorum species, sistens, cujus partem secundam, cons. exper. facult. med. upsal. publice ventilandam exhibent Johan. Edman, Upsala, 22 pp. [pp. 29 - 52]" type="book chapter" year="1783">Thunberg, 1783</bibRefCitation>
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: 43
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</treatmentCitation>
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</taxonomicName>
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.
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<taxonomicName id="43944D572239E37EFF0F2044B3CBFD14" authority=": Kirkaldy, 1909: 196" authorityName="Kirkaldy" authorityPageNumber="196" authorityYear="1909" box="[151,509,619,641]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="sulcata">
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<emphasis id="B6E0EAC62239E37EFF0F2044B317FD14" box="[151,289,619,641]" italics="true" pageId="16" pageNumber="367">Halys sulcata</emphasis>
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:
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<treatmentCitation id="053510C52239E37EFEB72044B3CBFD14" author="Kirkaldy, G. W." box="[303,509,619,641]" page="196" pageId="16" pageNumber="367" year="1909">
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<bibRefCitation id="E0054B252239E37EFEB72044B3FCFD14" author="Kirkaldy, G. W." box="[303,458,619,641]" pageId="16" pageNumber="367" refId="ref13331" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of food plants and parasites etc. Fol. I. Cimicidae. Berlin, XI + 392 pp." type="book" year="1909">Kirkaldy, 1909</bibRefCitation>
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: 196
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</treatmentCitation>
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</taxonomicName>
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.
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<taxonomicName id="43944D572239E37EFF0F20A6B3DAFD0A" authority="Chopra, 1974: 477" authorityName="Chopra" authorityPageNumber="477" authorityYear="1974" box="[151,492,649,671]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="magnus">
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<emphasis id="B6E0EAC62239E37EFF0F20A6B311FD0A" box="[151,295,649,671]" italics="true" pageId="16" pageNumber="367">Halys magnus</emphasis>
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<treatmentCitation id="053510C52239E37EFEB520A6B3DAFD0A" author="Chopra, N. P." box="[301,492,649,671]" page="477" pageId="16" pageNumber="367" year="1974">
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<bibRefCitation id="E0054B252239E37EFEB520A6B38CFD0A" author="Chopra, N. P." box="[301,442,649,671]" pageId="16" pageNumber="367" pagination="473 - 479" refId="ref12757" refString="Chopra, N. P. (1974) Studies on the genus Halys (Hemiptera: Pentatomidae), Oriental Insects, 8 (4), 473 - 479. https: // doi. org / 10.1080 / 00305316.1974.10434879" type="journal article" year="1974">Chopra, 1974</bibRefCitation>
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: 477
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</treatmentCitation>
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</taxonomicName>
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, 479 (syn.
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<treatmentCitation id="053510C52239E37EFDC320A6B13CFD0A" author="Ghauri, M. S. K." box="[603,778,649,671]" page="81" pageId="16" pageNumber="367" year="1988">
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<bibRefCitation id="E0054B252239E37EFDC320A6B0D2FD0A" author="Ghauri, M. S. K." box="[603,740,649,671]" pageId="16" pageNumber="367" pagination="77 - 92" refId="ref13220" refString="Ghauri, M. S. K. (1988) A revision of Asian species of the genus Halys Fabricius based on the type material (Insecta, Heteroptera, Pentatomidae, Pentatomidae). Entomologische Abhandlungen Staatliches Museum fur Tierkunde Dresden, 51 (6), 77 - 92." type="journal article" year="1988">Ghauri, 1988</bibRefCitation>
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||
: 81
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||
</treatmentCitation>
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||
).
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<taxonomicName id="43944D572239E37EFF0F2087B002FD2B" authority="Abbasi & Ahmad, 1976: 26" authorityName="Abbasi & Ahmad" authorityPageNumber="26" authorityYear="1976" box="[151,564,680,702]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="qadrii">
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<emphasis id="B6E0EAC62239E37EFF0F2087B320FD2B" box="[151,278,680,702]" italics="true" pageId="16" pageNumber="367">Halys qadrii</emphasis>
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||
<treatmentCitation id="053510C52239E37EFE842087B002FD2B" author="Abbasi, Q. A. & Ahmad, I." box="[284,564,680,702]" page="26" pageId="16" pageNumber="367" year="1976">
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<bibRefCitation id="E0054B252239E37EFE842087B027FD2B" author="Abbasi, Q. A. & Ahmad, I." box="[284,529,680,702]" pageId="16" pageNumber="367" pagination="26 - 31" refId="ref12358" refString="Abbasi, Q. A. & Ahmad, I. (1976) A new species of the genus Halys Fabr. (Pentatomidea: Halyini) from East Bengal with notes on its genitalia and their bearing on classification. Karachi University Journal of Science, 4 (1 - 2), 26 - 31." type="journal article" year="1976">Abbasi & Ahmad, 1976</bibRefCitation>
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: 26
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||
</treatmentCitation>
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</taxonomicName>
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||
–30 (syn.
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<treatmentCitation id="053510C52239E37EFD012087B17EFD2B" author="Ghauri, M. S. K." box="[665,840,680,702]" page="81" pageId="16" pageNumber="367" year="1988">
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<bibRefCitation id="E0054B252239E37EFD012087B114FD2B" author="Ghauri, M. S. K." box="[665,802,680,702]" pageId="16" pageNumber="367" pagination="77 - 92" refId="ref13220" refString="Ghauri, M. S. K. (1988) A revision of Asian species of the genus Halys Fabricius based on the type material (Insecta, Heteroptera, Pentatomidae, Pentatomidae). Entomologische Abhandlungen Staatliches Museum fur Tierkunde Dresden, 51 (6), 77 - 92." type="journal article" year="1988">Ghauri, 1988</bibRefCitation>
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||
: 81
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||
</treatmentCitation>
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).
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||
<taxonomicName id="43944D572239E37EFF0F20E9B3DEFD49" authority=": Ghauri, 1988: 81" authorityName="Ghauri" authorityPageNumber="81" authorityYear="1988" box="[151,488,710,732]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="sulcatus">
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<emphasis id="B6E0EAC62239E37EFF0F20E9B31DFD49" box="[151,299,710,732]" italics="true" pageId="16" pageNumber="367">Halys sulcatus</emphasis>
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:
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<treatmentCitation id="053510C52239E37EFEA120E9B3DEFD49" author="Ghauri, M. S. K." box="[313,488,710,732]" page="81" pageId="16" pageNumber="367" year="1988">
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<bibRefCitation id="E0054B252239E37EFEA120E9B3F7FD49" author="Ghauri, M. S. K." box="[313,449,710,732]" pageId="16" pageNumber="367" pagination="77 - 92" refId="ref13220" refString="Ghauri, M. S. K. (1988) A revision of Asian species of the genus Halys Fabricius based on the type material (Insecta, Heteroptera, Pentatomidae, Pentatomidae). Entomologische Abhandlungen Staatliches Museum fur Tierkunde Dresden, 51 (6), 77 - 92." type="journal article" year="1988">Ghauri, 1988</bibRefCitation>
|
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: 81
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</treatmentCitation>
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</taxonomicName>
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.
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||
<taxonomicName id="43944D572239E37EFF0F20CAB3D3FD6E" authority=": Rider, 2006: 308" authorityName="Rider" authorityPageNumber="308" authorityYear="2006" box="[151,485,741,763]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="sulcatus">
|
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<emphasis id="B6E0EAC62239E37EFF0F20CAB31DFD6E" box="[151,299,741,763]" italics="true" pageId="16" pageNumber="367">Halys sulcatus</emphasis>
|
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:
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<treatmentCitation id="053510C52239E37EFEA120CAB3D3FD6E" author="Rider, D. A." box="[313,485,741,763]" page="308" pageId="16" pageNumber="367" year="2006">
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<bibRefCitation id="E0054B252239E37EFEA120CAB384FD6E" author="Rider, D. A." box="[313,434,741,763]" pageId="16" pageNumber="367" pagination="233 - 402" refId="ref14017" refString="Rider, D. A. (2006) Family Pentatomidae. In: Aukema, B. & Rieger, C. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Fol. 5. The Netherlands Entomological Society, Amsterdam, pp. 233 - 402." type="book chapter" year="2006">Rider, 2006</bibRefCitation>
|
||
: 308
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||
</treatmentCitation>
|
||
</taxonomicName>
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||
.
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||
<taxonomicName id="43944D572239E37EFF0F212BB05CFC8F" authority="Memon, Meier & Manan, 2006" authorityName="Memon, Meier & Manan" authorityYear="2006" box="[151,618,772,794]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="sindilla">
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||
<emphasis id="B6E0EAC62239E37EFF0F212BB312FC8F" box="[151,292,772,794]" italics="true" pageId="16" pageNumber="367">Halys sindilla</emphasis>
|
||
<bibRefCitation id="E0054B252239E37EFEB3212AB05CFC8F" author="Memon, N. & Meier, R. & Manan, A. & Su, K. F. - Y." box="[299,618,772,794]" pageId="16" pageNumber="367" pagination="703 - 710" refId="ref13758" refString="Memon, N., Meier, R., Manan, A. & Su, K. F. - Y. (2006) On the use of DNA sequences for determining the species limits of a polymorphic new species in the stink bug genus Halys (Heteroptera: Pentatomidae) from Pakistan. Systematic Entomology, 31 (4), 703 - 710. https: // doi. org / 10.1111 / j. 1365 - 3113.2006.00350. x" type="journal article" year="2006">Memon, Meier & Manan, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
,
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||
<taxonomicNameLabel id="ADD357BD2239E37EFDED212CB0E0FC8C" box="[629,726,771,794]" pageId="16" pageNumber="367">
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||
<emphasis id="B6E0EAC62239E37EFDED212CB0E0FC8C" bold="true" box="[629,726,771,794]" pageId="16" pageNumber="367">Syn. nov.</emphasis>
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||
</taxonomicNameLabel>
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||
<taxonomicName id="43944D572239E37EFF0F210CB057FCAC" authority="Shaikh, Memon & Shah, 2011" authorityName="Shaikh, Memon & Shah" authorityYear="2011" box="[151,609,803,825]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="spinosa">
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||
<emphasis id="B6E0EAC62239E37EFF0F210CB313FCAC" box="[151,293,803,825]" italics="true" pageId="16" pageNumber="367">Halys spinosa</emphasis>
|
||
<bibRefCitation id="E0054B252239E37EFEB4210CB057FCAC" author="Shaikh, A. M. & Memon, N. & Shah, M. A." box="[300,609,803,825]" pageId="16" pageNumber="367" pagination="157 - 160" refId="ref14285" refString="Shaikh, A. M., Memon, N. & Shah, M. A. (2011) A new halyine species of Halys Fabricius (Hemiptera, Pentatomidae, Pentatominae) from Miani forest, Sindh, Pakistan. Sindh University Research Journal, Science Series, 43 (2), 157 - 160." type="journal article" year="2011">Shaikh, Memon & Shah, 2011</bibRefCitation>
|
||
</taxonomicName>
|
||
,
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||
<taxonomicNameLabel id="ADD357BD2239E37EFDF4210DB0FBFCAD" box="[620,717,802,825]" pageId="16" pageNumber="367">
|
||
<emphasis id="B6E0EAC62239E37EFDF4210DB0FBFCAD" bold="true" box="[620,717,802,825]" pageId="16" pageNumber="367">Syn. nov.</emphasis>
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||
</taxonomicNameLabel>
|
||
<taxonomicName id="43944D572239E37EFF0F216EB120FCC2" authority="Memon, Parveen, Ahmad & Shaikh, 2016" authorityName="Memon, Parveen, Ahmad & Shaikh" authorityYear="2016" box="[151,790,833,855]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="mulberriensis">
|
||
<emphasis id="B6E0EAC62239E37EFF0F216EB355FCC2" box="[151,355,833,855]" italics="true" pageId="16" pageNumber="367">Halys mulberriensis</emphasis>
|
||
<bibRefCitation id="E0054B252239E37EFEF2216DB120FCC2" author="Memon, N. & Parveen, R. & Ahmad, I. & Shaikh, A. M." box="[362,790,833,855]" pageId="16" pageNumber="367" pagination="977 - 980" refId="ref13843" refString="Memon, N., Parveen, R., Ahmad, I. & Shaikh, A. M. (2016) Description of a new species of Oriental genus Halys (Pentatomoidae, Pentatominae, Halyini) from Lahore, Punjab, Pakistan. Pakistan Journal of Zoology, 48 (4), 977 - 980." type="journal article" year="2016">Memon, Parveen, Ahmad & Shaikh, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="ADD357BD2239E37EFCB8216EB1B4FCC2" box="[800,898,833,856]" pageId="16" pageNumber="367">
|
||
<emphasis id="B6E0EAC62239E37EFCB8216EB1B4FCC2" bold="true" box="[800,898,833,856]" pageId="16" pageNumber="367">Syn. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
<taxonomicName id="43944D572239E37EFF0F214FB17BFCE3" authority="Memon, Parveen, Ahmad & Shaikh, 2016: 977" authorityName="Memon, Parveen, Ahmad & Shaikh" authorityPageNumber="977" authorityYear="2016" box="[151,845,864,886]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="mulberriences">
|
||
<emphasis id="B6E0EAC62239E37EFF0F214FB35FFCE3" box="[151,361,864,886]" italics="true" pageId="16" pageNumber="367">Halys mulberriences</emphasis>
|
||
<treatmentCitation id="053510C52239E37EFEF7214EB17BFCE3" author="Memon, N. & Parveen, R. & Ahmad, I. & Shaikh, A. M." box="[367,845,864,886]" page="977" pageId="16" pageNumber="367" year="2016">
|
||
<bibRefCitation id="E0054B252239E37EFEF7214EB128FCE3" author="Memon, N. & Parveen, R. & Ahmad, I. & Shaikh, A. M." box="[367,798,864,886]" pageId="16" pageNumber="367" pagination="977 - 980" refId="ref13843" refString="Memon, N., Parveen, R., Ahmad, I. & Shaikh, A. M. (2016) Description of a new species of Oriental genus Halys (Pentatomoidae, Pentatominae, Halyini) from Lahore, Punjab, Pakistan. Pakistan Journal of Zoology, 48 (4), 977 - 980." type="journal article" year="2016">Memon, Parveen, Ahmad & Shaikh, 2016</bibRefCitation>
|
||
: 977
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
–979 (lapsus calami)
|
||
<taxonomicName id="43944D572239E37EFF0F2151B0CDFC01" authority="Memon, Parveen, Ahmad & Shah, 2017" authorityName="Memon, Parveen, Ahmad & Shah" authorityYear="2017" box="[151,763,894,916]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="noakoatensis">
|
||
<emphasis id="B6E0EAC62239E37EFF0F2151B36AFC01" box="[151,348,894,916]" italics="true" pageId="16" pageNumber="367">Halys noakoatensis</emphasis>
|
||
<bibRefCitation id="E0054B252239E37EFEFB2150B0CDFC01" author="Memon, N. & Parveen, R. & Ahmad, I. & Shah, N. A." box="[355,763,894,916]" pageId="16" pageNumber="367" pagination="1301 - 1306" refId="ref13904" refString="Memon, N., Parveen, R., Ahmad, I. & Shah, N. A. (2017) Description of two new species of Oriental genus Halys (Pentatomoidae, Pentatominae, Halyini) from Sindh, Pakistan with a key of its nine Asian species. Pakistan Journal of Zoology, 49 (4), 1301 - 1306. https: // doi. org / 10.17582 / journal. pjz / 2017.49.4.1301.1306" type="journal article" year="2017">Memon, Parveen, Ahmad & Shah, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="ADD357BD2239E37EFC9D2151B151FC01" box="[773,871,894,917]" pageId="16" pageNumber="367">
|
||
<emphasis id="B6E0EAC62239E37EFC9D2151B151FC01" bold="true" box="[773,871,894,917]" pageId="16" pageNumber="367">Syn. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="842B36D42239E37EFF0F218CB625FC40" blockId="16.[151,1055,588,981]" pageId="16" pageNumber="367">
|
||
<treatmentCitationGroup id="A48411FA2239E37EFF0F218CB625FC40" pageId="16" pageNumber="367">
|
||
<taxonomicName id="43944D572239E37EFF0F218CB11AFC2C" authority="Memon, Parveen, Ahmad & Shah, 2017: 1304" authorityName="Memon, Parveen, Ahmad & Shah" authorityPageNumber="1304" authorityYear="2017" box="[151,812,931,953]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="naokotensis">
|
||
<emphasis id="B6E0EAC62239E37EFF0F218CB379FC2C" box="[151,335,931,953]" italics="true" pageId="16" pageNumber="367">Halys naokotensis</emphasis>
|
||
<treatmentCitation id="053510C52239E37EFECE218BB11AFC2C" author="Memon, N. & Parveen, R. & Ahmad, I. & Shah, N. A." box="[342,812,931,953]" page="1304" pageId="16" pageNumber="367" year="2017">
|
||
<bibRefCitation id="E0054B252239E37EFECE218BB0C6FC2C" author="Memon, N. & Parveen, R. & Ahmad, I. & Shah, N. A." box="[342,752,931,953]" pageId="16" pageNumber="367" pagination="1301 - 1306" refId="ref13904" refString="Memon, N., Parveen, R., Ahmad, I. & Shah, N. A. (2017) Description of two new species of Oriental genus Halys (Pentatomoidae, Pentatominae, Halyini) from Sindh, Pakistan with a key of its nine Asian species. Pakistan Journal of Zoology, 49 (4), 1301 - 1306. https: // doi. org / 10.17582 / journal. pjz / 2017.49.4.1301.1306" type="journal article" year="2017">Memon, Parveen, Ahmad & Shah, 2017</bibRefCitation>
|
||
: 1304
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
(lapsus calami)
|
||
<taxonomicName id="43944D572239E37EFF0F2190B145FC40" authority="Memon, Parveen, Ahmad & Shah, 2017: 1304" authorityName="Memon, Parveen, Ahmad & Shah" authorityPageNumber="1304" authorityYear="2017" box="[151,883,959,981]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="naokotiensis">
|
||
<emphasis id="B6E0EAC62239E37EFF0F2190B360FC40" box="[151,342,959,981]" italics="true" pageId="16" pageNumber="367">Halys naokotiensis</emphasis>
|
||
<treatmentCitation id="053510C52239E37EFEC521EFB103FC40" author="Memon, N. & Parveen, R. & Ahmad, I. & Shah, N. A." box="[349,821,959,981]" page="1304" pageId="16" pageNumber="367" year="2017">
|
||
<bibRefCitation id="E0054B252239E37EFEC521EFB0C1FC40" author="Memon, N. & Parveen, R. & Ahmad, I. & Shah, N. A." box="[349,759,959,981]" pageId="16" pageNumber="367" pagination="1301 - 1306" refId="ref13904" refString="Memon, N., Parveen, R., Ahmad, I. & Shah, N. A. (2017) Description of two new species of Oriental genus Halys (Pentatomoidae, Pentatominae, Halyini) from Sindh, Pakistan with a key of its nine Asian species. Pakistan Journal of Zoology, 49 (4), 1301 - 1306. https: // doi. org / 10.17582 / journal. pjz / 2017.49.4.1301.1306" type="journal article" year="2017">Memon, Parveen, Ahmad & Shah, 2017</bibRefCitation>
|
||
: 1304
|
||
</treatmentCitation>
|
||
, 1305
|
||
</taxonomicName>
|
||
(lapsus calami)
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="842B36D42239E37EFF0F262EB17BFB5B" blockId="16.[151,1437,1025,2022]" pageId="16" pageNumber="367">
|
||
<emphasis id="B6E0EAC62239E37EFF0F262EB315FB8F" bold="true" box="[151,291,1025,1050]" italics="true" pageId="16" pageNumber="367">Colouration</emphasis>
|
||
. Dorsum including head ochraceous with spots formed by clusters of black punctures; antennae colour variable, usually ochraceous with segments IV and V black, except basal region; scutellar pits black; connexivum concolourous with dorsum, except for black anterior and posterior third of lateral margin of segments; hemelytral membrane light brown with black veins; ventral surface including labium and legs nearly concolourous with or paler than dorsum except for black apical segment of labium, apices of tarsal segments and claws, anterior and posterior angles of abdominal ventrites and spiracular outline.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CC8E655F2239E37FFF5F26F5B1C2FA58" lastPageId="17" lastPageNumber="368" pageId="16" pageNumber="367" type="description">
|
||
<paragraph id="842B36D42239E37EFF5F26F5B375FAAF" blockId="16.[151,1437,1025,2022]" pageId="16" pageNumber="367">
|
||
<emphasis id="B6E0EAC62239E37EFF5F26F5B3C9FB67" bold="true" box="[199,511,1241,1266]" italics="true" pageId="16" pageNumber="367">Integument and vestiture.</emphasis>
|
||
Dorsum including head covered with dense, irregular dark brown to black punctures, sometimes in clusters. Ventral surface including legs with dark brown to black, coarse, dense punctures. Body glabrous.
|
||
</paragraph>
|
||
<paragraph id="842B36D42239E37EFF5F276AB01EFA33" blockId="16.[151,1437,1025,2022]" pageId="16" pageNumber="367">
|
||
<emphasis id="B6E0EAC62239E37EFF5F276AB30FFACB" bold="true" box="[199,313,1349,1374]" italics="true" pageId="16" pageNumber="367">Structure.</emphasis>
|
||
Labium surpassing anterior margin of ventrite VI. Lateral and posterior margins of pronotum nearly straight. Humeri angulate, but not elevated. Disc of pronotum as well as scutellum more or less flat. Other characters as in generic description.
|
||
</paragraph>
|
||
<paragraph id="842B36D42239E37FFF5F279EB665FF25" blockId="16.[151,1437,1025,2022]" lastBlockId="17.[151,1107,151,176]" lastPageId="17" lastPageNumber="368" pageId="16" pageNumber="367">
|
||
<emphasis id="B6E0EAC62239E37EFF5F279EB0A3FA5F" bold="true" box="[199,661,1457,1482]" pageId="16" pageNumber="367">
|
||
External male genitalia (
|
||
<figureCitation id="1CAF2A512239E37EFE66279EB0B1FA5F" box="[510,647,1457,1482]" captionStart-0="FIGURES 77–83" captionStart-1="FIGURES 84–90" captionStartId-0="17.[151,264,1157,1180]" captionStartId-1="18.[151,264,1408,1431]" captionTargetBox-0="[168,1425,263,1128]" captionTargetBox-1="[175,1422,499,1319]" captionTargetId-0="figure@17.[151,1435,229,1137]" captionTargetId-1="figure@18.[151,1435,460,1367]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="FIGURES 77–83. H. sulcata (Thunberg, 1783)—male genitalia. 77, genital capsule (dorsal); 78, genital capsule (ventral); 79, genital capsule (caudal); 80, paramere (dorsal); 81, paramere (ventrolateral); 82, paramere (ventral); 83, articulatory apparatus. Lettering: cp—capitate processes; mp—median process; t—tooth." captionText-1="FIGURES 84–90. H. sulcata (Thunberg, 1783)—male and female genitalia. 84, phallus (dorsal); 85, phallus (ventral); 86, phallus (lateral); 87, spermatheca; 88, female terminalia (before digestion); 89, female terminalia (after digestion); 90,valvulae VIII." httpUri-0="https://zenodo.org/record/2644578/files/figure.png" httpUri-1="https://zenodo.org/record/2644580/files/figure.png" pageId="16" pageNumber="367">Figs 77–90</figureCitation>
|
||
).
|
||
</emphasis>
|
||
<emphasis id="B6E0EAC62239E37EFD3A279EB157FA5F" box="[674,865,1457,1482]" italics="true" pageId="16" pageNumber="367">Genital capsule.</emphasis>
|
||
Quadrangular. Dorsal rim outlined with mat of minute, fine pubescence, widely and shallowly excavated concave, with median, small U-shaped notch, pigmented black. Dorsal rim slightly indented adjacent to caudal lobes; infoldings sclerotized black forming 1+1 kidney shaped, moderately deep impressions on sides of median notch, infoldings with sclerotized 1+1 short, black tooth submedially on sides of median notch (
|
||
<figureCitation id="1CAF2A512239E37EFDCA246EB090F9CF" box="[594,678,1601,1626]" captionStart="FIGURES 94–99" captionStartId="22.[151,264,1273,1296]" captionTargetBox="[170,1415,467,1204]" captionTargetId="figure@22.[151,1435,409,1253]" captionTargetPageId="22" captionText="FIGURES 94–99. FIGURES 94–95. Genital capsule (caudal). 94, H. shaista Ghauri, 1988; 95, H. sulcata (Thunberg, 1783). Lettering: t—tooth; ts—tooth like serrations. FIGURES 96–99. Capitate processes. 96, H. mudigerensis sp. nov.; 97, H. serrigera Westwood, 1837; 98, H. shaista Ghauri, 1988; 99, H. sulcata (Thunberg, 1783)" httpUri="https://zenodo.org/record/2644584/files/figure.png" pageId="16" pageNumber="367">Fig. 95</figureCitation>
|
||
); ventral rim together with infoldings widely and deeply excavated concave medially; infoldings with median region convex, dorsally possessing small, short, spout-like median projection (
|
||
<figureCitation id="1CAF2A512239E37EFE8224A6B346F937" box="[282,368,1673,1698]" captionStart="FIGURES 77–83" captionStartId="17.[151,264,1157,1180]" captionTargetBox="[168,1425,263,1128]" captionTargetId="figure@17.[151,1435,229,1137]" captionTargetPageId="17" captionText="FIGURES 77–83. H. sulcata (Thunberg, 1783)—male genitalia. 77, genital capsule (dorsal); 78, genital capsule (ventral); 79, genital capsule (caudal); 80, paramere (dorsal); 81, paramere (ventrolateral); 82, paramere (ventral); 83, articulatory apparatus. Lettering: cp—capitate processes; mp—median process; t—tooth." httpUri="https://zenodo.org/record/2644578/files/figure.png" pageId="16" pageNumber="367">Fig. 78</figureCitation>
|
||
); 1+1 sclerotized, obliquely straight narrow ridge on sides of median convex region on inner side, reaching caudolateral margin of infoldings; infoldings well developed, lamellate, elevated, laterally reaching adjacent to caudal lobes as small rounded projection (
|
||
<figureCitation id="1CAF2A512239E37EFC9124FEB157F97F" box="[777,865,1745,1770]" captionStart="FIGURES 77–83" captionStartId="17.[151,264,1157,1180]" captionTargetBox="[168,1425,263,1128]" captionTargetId="figure@17.[151,1435,229,1137]" captionTargetPageId="17" captionText="FIGURES 77–83. H. sulcata (Thunberg, 1783)—male genitalia. 77, genital capsule (dorsal); 78, genital capsule (ventral); 79, genital capsule (caudal); 80, paramere (dorsal); 81, paramere (ventrolateral); 82, paramere (ventral); 83, articulatory apparatus. Lettering: cp—capitate processes; mp—median process; t—tooth." httpUri="https://zenodo.org/record/2644578/files/figure.png" pageId="16" pageNumber="367">Fig. 78</figureCitation>
|
||
).
|
||
<emphasis id="B6E0EAC62239E37EFCEF24FCB1D0F97F" box="[887,998,1747,1770]" italics="true" pageId="16" pageNumber="367">Paramere</emphasis>
|
||
roughly C-shaped (
|
||
<figureCitation id="1CAF2A512239E37EFB5624FEB762F97F" box="[1230,1364,1745,1770]" captionStart="FIGURES 77–83" captionStartId="17.[151,264,1157,1180]" captionTargetBox="[168,1425,263,1128]" captionTargetId="figure@17.[151,1435,229,1137]" captionTargetPageId="17" captionText="FIGURES 77–83. H. sulcata (Thunberg, 1783)—male genitalia. 77, genital capsule (dorsal); 78, genital capsule (ventral); 79, genital capsule (caudal); 80, paramere (dorsal); 81, paramere (ventrolateral); 82, paramere (ventral); 83, articulatory apparatus. Lettering: cp—capitate processes; mp—median process; t—tooth." httpUri="https://zenodo.org/record/2644578/files/figure.png" pageId="16" pageNumber="367">Figs 80–82</figureCitation>
|
||
), with expanded lanceolate crown, short stem and expanded plate-like apodeme. Crown with one short, blunt, black, slightly curved, hook-like tooth at caudal apex and another continued as indistinct ridge or narrow black patch medially.
|
||
<emphasis id="B6E0EAC62239E37EFE942512B35FF8C3" box="[268,361,1853,1878]" italics="true" pageId="16" pageNumber="367">Phallus.</emphasis>
|
||
Articulatory apparatus with basal plates and support bridge complex, dorsal connectives widened into 1+1 roughly quadrangular capitate processes (
|
||
<figureCitation id="1CAF2A512239E37EFCC7254EB18EF8EF" box="[863,952,1889,1914]" captionStart="FIGURES 77–83" captionStartId="17.[151,264,1157,1180]" captionTargetBox="[168,1425,263,1128]" captionTargetId="figure@17.[151,1435,229,1137]" captionTargetPageId="17" captionText="FIGURES 77–83. H. sulcata (Thunberg, 1783)—male genitalia. 77, genital capsule (dorsal); 78, genital capsule (ventral); 79, genital capsule (caudal); 80, paramere (dorsal); 81, paramere (ventrolateral); 82, paramere (ventral); 83, articulatory apparatus. Lettering: cp—capitate processes; mp—median process; t—tooth." httpUri="https://zenodo.org/record/2644578/files/figure.png" pageId="16" pageNumber="367">Fig. 83</figureCitation>
|
||
,
|
||
<figureCitation id="1CAF2A512239E37EFC52254DB1DCF8EF" box="[970,1002,1890,1914]" captionStart="FIGURES 94–99" captionStartId="22.[151,264,1273,1296]" captionTargetBox="[170,1415,467,1204]" captionTargetId="figure@22.[151,1435,409,1253]" captionTargetPageId="22" captionText="FIGURES 94–99. FIGURES 94–95. Genital capsule (caudal). 94, H. shaista Ghauri, 1988; 95, H. sulcata (Thunberg, 1783). Lettering: t—tooth; ts—tooth like serrations. FIGURES 96–99. Capitate processes. 96, H. mudigerensis sp. nov.; 97, H. serrigera Westwood, 1837; 98, H. shaista Ghauri, 1988; 99, H. sulcata (Thunberg, 1783)" httpUri="https://zenodo.org/record/2644584/files/figure.png" pageId="16" pageNumber="367">99</figureCitation>
|
||
). Phallosoma sclerotized, constricted towards both ends, broadest medially and concave dorsally; two pairs of membranous conjunctival processes, dorsal pair elongate and longer than processes of aedeagus, with apex rounded, ventral pair subequal to dorsal pair (
|
||
<figureCitation id="1CAF2A512239E37EFF0725E2B312F873" box="[159,292,1997,2022]" captionStart="FIGURES 84–90" captionStartId="18.[151,264,1408,1431]" captionTargetBox="[175,1422,499,1319]" captionTargetId="figure@18.[151,1435,460,1367]" captionTargetPageId="18" captionText="FIGURES 84–90. H. sulcata (Thunberg, 1783)—male and female genitalia. 84, phallus (dorsal); 85, phallus (ventral); 86, phallus (lateral); 87, spermatheca; 88, female terminalia (before digestion); 89, female terminalia (after digestion); 90,valvulae VIII." httpUri="https://zenodo.org/record/2644580/files/figure.png" pageId="16" pageNumber="367">Figs 84–86</figureCitation>
|
||
). Processes of aedeagus hard, sclerotized, highly sclerotized and free (not fused), apically narrowly excavated concave (
|
||
<figureCitation id="1CAF2A512238E37FFEE122B8B3FAFF25" box="[377,460,151,176]" captionStart="FIGURES 84–90" captionStartId="18.[151,264,1408,1431]" captionTargetBox="[175,1422,499,1319]" captionTargetId="figure@18.[151,1435,460,1367]" captionTargetPageId="18" captionText="FIGURES 84–90. H. sulcata (Thunberg, 1783)—male and female genitalia. 84, phallus (dorsal); 85, phallus (ventral); 86, phallus (lateral); 87, spermatheca; 88, female terminalia (before digestion); 89, female terminalia (after digestion); 90,valvulae VIII." httpUri="https://zenodo.org/record/2644580/files/figure.png" pageId="17" pageNumber="368">Fig. 86</figureCitation>
|
||
); aedeagus moderately long and broadly curved upwards.
|
||
</paragraph>
|
||
<caption id="D0EB665C2238E37FFF0F26AAB103FB4C" httpUri="https://zenodo.org/record/2644578/files/figure.png" pageId="17" pageNumber="368" startId="17.[151,264,1157,1180]" targetBox="[168,1425,263,1128]" targetPageId="17">
|
||
<paragraph id="842B36D42238E37FFF0F26AAB103FB4C" blockId="17.[151,1437,1157,1241]" pageId="17" pageNumber="368">
|
||
<emphasis id="B6E0EAC62238E37FFF0F26AAB362FB0E" bold="true" box="[151,340,1157,1180]" pageId="17" pageNumber="368">FIGURES 77–83.</emphasis>
|
||
<taxonomicName id="43944D572238E37FFEC226A8B04FFB09" authority="(Thunberg, 1783)" authorityName="Thunberg" authorityYear="1783" baseAuthorityName="Thunberg" baseAuthorityYear="1783" box="[346,633,1158,1180]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="17" pageNumber="368" phylum="Chordata" rank="species" species="sulcata">
|
||
<emphasis id="B6E0EAC62238E37FFEC226A8B3F5FB09" box="[346,451,1158,1180]" italics="true" pageId="17" pageNumber="368">H. sulcata</emphasis>
|
||
(Thunberg, 1783)
|
||
</taxonomicName>
|
||
—male genitalia. 77, genital capsule (dorsal); 78, genital capsule (ventral); 79, genital capsule (caudal); 80, paramere (dorsal); 81, paramere (ventrolateral); 82, paramere (ventral); 83, articulatory apparatus. Lettering: cp—capitate processes; mp—median process; t—tooth.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="842B36D42238E37FFF5F272FB1C2FA58" blockId="17.[151,1437,1280,2025]" pageId="17" pageNumber="368">
|
||
<emphasis id="B6E0EAC62238E37FFF5F272FB0A3FA8C" bold="true" box="[199,661,1280,1305]" pageId="17" pageNumber="368">
|
||
External female genitalia (
|
||
<figureCitation id="1CAF2A512238E37FFD9C272FB0BEFA8C" box="[516,648,1280,1305]" captionStart="FIGURES 84–90" captionStartId="18.[151,264,1408,1431]" captionTargetBox="[175,1422,499,1319]" captionTargetId="figure@18.[151,1435,460,1367]" captionTargetPageId="18" captionText="FIGURES 84–90. H. sulcata (Thunberg, 1783)—male and female genitalia. 84, phallus (dorsal); 85, phallus (ventral); 86, phallus (lateral); 87, spermatheca; 88, female terminalia (before digestion); 89, female terminalia (after digestion); 90,valvulae VIII." httpUri="https://zenodo.org/record/2644580/files/figure.png" pageId="17" pageNumber="368">Figs 87–90</figureCitation>
|
||
).
|
||
</emphasis>
|
||
<emphasis id="B6E0EAC62238E37FFD05272FB128FA8C" box="[669,798,1280,1305]" italics="true" pageId="17" pageNumber="368">Terminalia.</emphasis>
|
||
Valvifers VIII roughly quadrangular, with mesal margin slightly concave, inner posterior angles rounded; posterior margin slightly convex to nearly straight. Valvulae VIII with apical (= caudal) margin uniformly rounded (
|
||
<figureCitation id="1CAF2A512238E37FFD792767B10FFAF4" box="[737,825,1352,1377]" captionStart="FIGURES 84–90" captionStartId="18.[151,264,1408,1431]" captionTargetBox="[175,1422,499,1319]" captionTargetId="figure@18.[151,1435,460,1367]" captionTargetPageId="18" captionText="FIGURES 84–90. H. sulcata (Thunberg, 1783)—male and female genitalia. 84, phallus (dorsal); 85, phallus (ventral); 86, phallus (lateral); 87, spermatheca; 88, female terminalia (before digestion); 89, female terminalia (after digestion); 90,valvulae VIII." httpUri="https://zenodo.org/record/2644580/files/figure.png" pageId="17" pageNumber="368">Fig. 90</figureCitation>
|
||
). Valvifers IX single transverse, narrow sclerite with indistinct median fusion line. Laterotergite IX obliquely placed and distinctly shorter than apex of abdomen. Laterotergite VIII broad, elongate and triangular with nearly smooth caudal margin.
|
||
<emphasis id="B6E0EAC62238E37FFBCC27BFB6D0FA3C" box="[1108,1254,1424,1449]" italics="true" pageId="17" pageNumber="368">Spermatheca</emphasis>
|
||
(
|
||
<figureCitation id="1CAF2A512238E37FFB6E27BFB77FFA3C" box="[1270,1353,1424,1449]" captionStart="FIGURES 84–90" captionStartId="18.[151,264,1408,1431]" captionTargetBox="[175,1422,499,1319]" captionTargetId="figure@18.[151,1435,460,1367]" captionTargetPageId="18" captionText="FIGURES 84–90. H. sulcata (Thunberg, 1783)—male and female genitalia. 84, phallus (dorsal); 85, phallus (ventral); 86, phallus (lateral); 87, spermatheca; 88, female terminalia (before digestion); 89, female terminalia (after digestion); 90,valvulae VIII." httpUri="https://zenodo.org/record/2644580/files/figure.png" pageId="17" pageNumber="368">Fig. 87</figureCitation>
|
||
) apical receptacle orbicular with two finger-like processes, directed towards flanges.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CC8E655F2238E37FFF5F27F7B0BFF95C" pageId="17" pageNumber="368" type="diagnosis">
|
||
<paragraph id="842B36D42238E37FFF5F27F7B0BFF95C" blockId="17.[151,1437,1280,2025]" pageId="17" pageNumber="368">
|
||
<emphasis id="B6E0EAC62238E37FFF5F27F7B3FBFA64" bold="true" box="[199,461,1496,1521]" pageId="17" pageNumber="368">Differential diagnosis</emphasis>
|
||
. This species can be recognized by the short, spout-like median process on the infoldings of ventral rim of genital capsule, expanded parameral crown with two tooth, one situated apically and another medially towards ventrolateral margin of crown, quadrangular valvifers VIII with posterior margin nearly straight (
|
||
<figureCitation id="1CAF2A512238E37FFF66246BB3B3F9C8" box="[254,389,1604,1629]" captionStart="FIGURES 84–90" captionStartId="18.[151,264,1408,1431]" captionTargetBox="[175,1422,499,1319]" captionTargetId="figure@18.[151,1435,460,1367]" captionTargetPageId="18" captionText="FIGURES 84–90. H. sulcata (Thunberg, 1783)—male and female genitalia. 84, phallus (dorsal); 85, phallus (ventral); 86, phallus (lateral); 87, spermatheca; 88, female terminalia (before digestion); 89, female terminalia (after digestion); 90,valvulae VIII." httpUri="https://zenodo.org/record/2644580/files/figure.png" pageId="17" pageNumber="368">Figs 88, 89</figureCitation>
|
||
) and valvulae VIII with caudal margin uniformly rounded (
|
||
<tableCitation id="C916036F2238E37FFBDD246BB6AAF9C8" box="[1093,1180,1604,1629]" captionStart="TABLE 1" captionStartId="20.[151,239,185,208]" captionTargetPageId="20" captionText="TABLE 1. Diagnostic characters of species of Halys from India" httpUri="http://table.plazi.org/id/D0EB665C223DE37AFF0F2296B177FF45" pageId="17" pageNumber="368" tableUuid="D0EB665C223DE37AFF0F2296B177FF45">Table 1</tableCitation>
|
||
). The females of this species can be differentiated from
|
||
<taxonomicName id="43944D572238E37FFD8E2445B0B1F914" authorityName="Ghauri" authorityYear="1988" box="[534,647,1640,1665]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="17" pageNumber="368" phylum="Chordata" rank="species" species="shaista">
|
||
<emphasis id="B6E0EAC62238E37FFD8E2445B0B1F914" box="[534,647,1640,1665]" italics="true" pageId="17" pageNumber="368">H. shaista</emphasis>
|
||
</taxonomicName>
|
||
by the shape of the valvifers IX, which appear to be divided due to the presence of a indistinct median fusion line (
|
||
<figureCitation id="1CAF2A512238E37FFD1224A3B0E9F930" box="[650,735,1676,1701]" captionStart="FIGURES 84–90" captionStartId="18.[151,264,1408,1431]" captionTargetBox="[175,1422,499,1319]" captionTargetId="figure@18.[151,1435,460,1367]" captionTargetPageId="18" captionText="FIGURES 84–90. H. sulcata (Thunberg, 1783)—male and female genitalia. 84, phallus (dorsal); 85, phallus (ventral); 86, phallus (lateral); 87, spermatheca; 88, female terminalia (before digestion); 89, female terminalia (after digestion); 90,valvulae VIII." httpUri="https://zenodo.org/record/2644580/files/figure.png" pageId="17" pageNumber="368">Fig. 88</figureCitation>
|
||
), whereas in
|
||
<taxonomicName id="43944D572238E37FFCEF24A1B1DFF930" authorityName="Ghauri" authorityYear="1988" box="[887,1001,1676,1701]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="17" pageNumber="368" phylum="Chordata" rank="species" species="shaista">
|
||
<emphasis id="B6E0EAC62238E37FFCEF24A1B1DFF930" box="[887,1001,1676,1701]" italics="true" pageId="17" pageNumber="368">H. shaista</emphasis>
|
||
</taxonomicName>
|
||
, the valvifers IX is single, continuous sclerite lacking median fusion line (
|
||
<figureCitation id="1CAF2A512238E37FFDB0249FB04AF95C" box="[552,636,1712,1737]" captionStart="FIGURES 69–76" captionStartId="15.[151,264,1121,1144]" captionTargetBox="[181,1411,227,1050]" captionTargetId="figure@15.[151,1435,193,1101]" captionTargetPageId="15" captionText="FIGURES 69–76. H. shaista Ghauri, 1988—male and female genitalia. 69, phallus (dorsal); 70, phallus (ventral); 71–72, phallus (lateral); 73, spermatheca; 74, female terminalia (before digestion); 75, female terminalia (after digestion); 76, valvulae VIII." httpUri="https://zenodo.org/record/2644576/files/figure.png" pageId="17" pageNumber="368">Fig. 74</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CC8E655F2238E37CFF5F24FBB65EFF40" lastPageId="18" lastPageNumber="369" pageId="17" pageNumber="368" type="description">
|
||
<paragraph id="842B36D42238E37CFF5F24FBB65EFF40" blockId="17.[151,1437,1280,2025]" lastBlockId="18.[151,1436,151,392]" lastPageId="18" lastPageNumber="369" pageId="17" pageNumber="368">
|
||
<emphasis id="B6E0EAC62238E37FFF5F24FBB3DBF978" bold="true" box="[199,493,1748,1773]" pageId="17" pageNumber="368">Measurements (in mm).</emphasis>
|
||
Median (minimum–maximum).
|
||
<emphasis id="B6E0EAC62238E37FFCF424FBB185F978" bold="true" box="[876,947,1748,1773]" pageId="17" pageNumber="368">Males</emphasis>
|
||
(n=4). Body length 20.36 (19.91–20.63); head length 4.80 (4.56–5.13), head width 3.29 (3.22–3.43), interocular width 1.99 (1.92–2.09), lengths of antennal segments (I) 0.95 (0.87–1.00): (II) 2.17 (2.05–2.27): (III) 2.44 (2.36–2.56): (IV) 2.10 (1.99–2.22): (V) 1.72; lengths of labial segments (I) 2.26 (2.15–2.49): (II) 4.43 (4.18–4.64): (III) 4.19 (3.90–4.35): (IV) 2.53 (1.94–2.89); pronotum length 4.20 (4.15–4.25); pronotum width 9.09 (8.81–9.29); scutellum length 7.55 (7.39–7.69); scutellum width 5.39 (5.19–5.55).
|
||
<emphasis id="B6E0EAC62238E37FFE2B25A7B023F834" bold="true" box="[435,533,1928,1953]" pageId="17" pageNumber="368">Females</emphasis>
|
||
(n=2). Body length 20.83 (20.73–20.93); head length 5.02 (4.98–5.06), head width 3.41 (3.38–3.43), interocular width 2.07 (2.02–2.11), lengths of antennal segments (I) 0.88 (0.79–0.96): (II) 2.21 (2.17–2.25): (III) 2.38 (2.31–2.44): (IV) 1.92: (V) 1.61; lengths of labial segments (I) 2.16 (2.15–2.16): (II) 3.98 (3.80–4.17): (III) 4.25 (4.18–4.32): (IV) 2.7 (2.61–2.79); pronotum length 4.22 (4.16–4.29); pronotum width 9.21 (9.08–9.33); scutellum length 7.54 (7.51–7.57); scutellum width 5.57 (5.49–5.64).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CC8E655F223BE37CFF5F22F0B3B7FE89" pageId="18" pageNumber="369" type="materials_examined">
|
||
<paragraph id="842B36D4223BE37CFF5F22F0B3B7FE89" blockId="18.[151,1436,151,392]" pageId="18" pageNumber="369">
|
||
<emphasis id="B6E0EAC6223BE37CFF5F22F0B38EFF6D" bold="true" box="[199,440,223,248]" pageId="18" pageNumber="369">
|
||
<collectingRegion id="4650F836223BE37CFF5F22F0B319FF6D" box="[199,303,223,248]" pageId="18" pageNumber="369">Material</collectingRegion>
|
||
examined.
|
||
</emphasis>
|
||
<materialsCitation id="34FC3C89223BE37CFE5F22CFB739FF6D" ID-GBIF-Occurrence="2236308468" box="[455,1295,223,248]" collectingDate="2008-05-13" collectorName="Shama, P." country="India" location="Delhi" pageId="18" pageNumber="369" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Delhi">
|
||
<collectingCountry id="FC837644223BE37CFE5F22CFB02DFF6D" box="[455,539,224,248]" name="India" pageId="18" pageNumber="369">INDIA</collectingCountry>
|
||
:
|
||
<emphasis id="B6E0EAC6223BE37CFDB522F0B087FF6D" bold="true" box="[557,689,223,248]" pageId="18" pageNumber="369">
|
||
New
|
||
<collectingRegion id="4650F836223BE37CFDE922F0B087FF6D" box="[625,689,223,248]" country="India" name="Delhi" pageId="18" pageNumber="369">Delhi</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9292FD5D223BE37CFD5F22CFB0DAFF6D" box="[711,748,224,248]" pageId="18" pageNumber="369" type="male">3♂</specimenCount>
|
||
,
|
||
<specimenCount id="9292FD5D223BE37CFC9822CFB112FF6D" box="[768,804,224,248]" pageId="18" pageNumber="369" type="female">2♀</specimenCount>
|
||
, IARI,
|
||
<collectingDate id="E06EE9FC223BE37CFC1022CFB1CFFF6D" box="[904,1017,224,248]" pageId="18" pageNumber="369" value="2008-05-13">
|
||
<date id="F02A1014223BE37CFC1022CFB1CFFF6D" box="[904,1017,224,248]" pageId="18" pageNumber="369" value="2008-05-13">13.v.2008</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="29615302223BE37CFB9522F0B6B3FF62" box="[1037,1157,223,248]" pageId="18" pageNumber="369">Shama, P.</collectorName>
|
||
,
|
||
<emphasis id="B6E0EAC6223BE37CFB0022CEB687FF6D" box="[1176,1201,225,248]" italics="true" pageId="18" pageNumber="369">ex</emphasis>
|
||
mango
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount id="9292FD5D223BE37CFABA22CFB771FF6D" box="[1314,1351,224,248]" pageId="18" pageNumber="369" type="male">1♂</specimenCount>
|
||
, IARI,
|
||
<date id="F02A1014223BE37CFF0F232BB33EFE89" box="[151,264,260,284]" pageId="18" pageNumber="369" value="2008-05-02">02.v.2008</date>
|
||
, Shama, P.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CC8E655F223BE37DFF5F2308B151FEF0" lastPageId="19" lastPageNumber="370" pageId="18" pageNumber="369" type="discussion">
|
||
<paragraph id="842B36D4223BE37CFF5F2308B350FE1D" blockId="18.[151,1436,151,392]" pageId="18" pageNumber="369">
|
||
<emphasis id="B6E0EAC6223BE37CFF5F2308B30DFED5" bold="true" box="[199,315,295,320]" pageId="18" pageNumber="369">Remarks.</emphasis>
|
||
This species is restricted to the northern region of
|
||
<collectingCountry id="FC837644223BE37CFCE32308B18EFED5" box="[891,952,295,320]" name="India" pageId="18" pageNumber="369">India</collectingCountry>
|
||
; the only record from southern
|
||
<collectingCountry id="FC837644223BE37CFA852308B761FED5" box="[1309,1367,295,320]" name="India" pageId="18" pageNumber="369">India</collectingCountry>
|
||
is the one by
|
||
<bibRefCitation id="E0054B25223BE37CFF742363B3A4FEF1" author="Chopra, N. P." box="[236,402,332,357]" pageId="18" pageNumber="369" pagination="473 - 479" refId="ref12757" refString="Chopra, N. P. (1974) Studies on the genus Halys (Hemiptera: Pentatomidae), Oriental Insects, 8 (4), 473 - 479. https: // doi. org / 10.1080 / 00305316.1974.10434879" type="journal article" year="1974">Chopra (1974)</bibRefCitation>
|
||
from Coimbatore (
|
||
<collectingRegion id="4650F836223BE37CFDF62363B0CEFEF0" box="[622,760,332,357]" country="India" name="Tamil Nadu" pageId="18" pageNumber="369">Tamil Nadu</collectingRegion>
|
||
) (as
|
||
<taxonomicName id="43944D57223BE37CFCAA2362B19BFEF1" authorityName="Chopra" authorityYear="1974" box="[818,941,333,356]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="18" pageNumber="369" phylum="Chordata" rank="species" species="magnus">
|
||
<emphasis id="B6E0EAC6223BE37CFCAA2362B19BFEF1" box="[818,941,333,356]" italics="true" pageId="18" pageNumber="369">H. magnus</emphasis>
|
||
</taxonomicName>
|
||
), but the determination is questionable and needs verification.
|
||
</paragraph>
|
||
<caption id="D0EB665C223BE37CFF0F27AFB2FCFA41" httpUri="https://zenodo.org/record/2644580/files/figure.png" pageId="18" pageNumber="369" startId="18.[151,264,1408,1431]" targetBox="[175,1422,499,1319]" targetPageId="18">
|
||
<paragraph id="842B36D4223BE37CFF0F27AFB2FCFA41" blockId="18.[151,1436,1408,1492]" pageId="18" pageNumber="369">
|
||
<emphasis id="B6E0EAC6223BE37CFF0F27AFB360FA03" bold="true" box="[151,342,1408,1431]" pageId="18" pageNumber="369">FIGURES 84–90.</emphasis>
|
||
<taxonomicName id="43944D57223BE37CFEC727ADB0B3FA02" authority="(Thunberg, 1783)" authorityName="Thunberg" authorityYear="1783" baseAuthorityName="Thunberg" baseAuthorityYear="1783" box="[351,645,1409,1431]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="18" pageNumber="369" phylum="Chordata" rank="species" species="sulcata">
|
||
<emphasis id="B6E0EAC6223BE37CFEC727ADB3FCFA02" box="[351,458,1409,1431]" italics="true" pageId="18" pageNumber="369">H. sulcata</emphasis>
|
||
(Thunberg, 1783)
|
||
</taxonomicName>
|
||
—male and female genitalia. 84, phallus (dorsal); 85, phallus (ventral); 86, phallus (lateral); 87, spermatheca; 88, female terminalia (before digestion); 89, female terminalia (after digestion); 90,valvulae VIII.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="842B36D4223BE37DFF57242EB151FEF0" blockId="18.[151,1437,1537,2030]" lastBlockId="19.[151,1436,151,357]" lastPageId="19" lastPageNumber="370" pageId="18" pageNumber="369">
|
||
There are quite a few species (see the details below) described from
|
||
<collectingCountry id="FC837644223BE37CFC42242EB60FF98F" box="[986,1081,1537,1562]" name="Pakistan" pageId="18" pageNumber="369">Pakistan</collectingCountry>
|
||
by various authors but with no valid characters to prove their distinctness from the already described species, as most of the characters are homoplasious and subject of strong intraspecific variation. Based on the original descriptions of
|
||
<taxonomicName id="43944D57223BE37CFB422465B767F9F7" authorityName="Memon, Meier & Manan" authorityYear="2006" box="[1242,1361,1609,1634]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="18" pageNumber="369" phylum="Chordata" rank="species" species="sindilla">
|
||
<emphasis id="B6E0EAC6223BE37CFB422465B767F9F7" box="[1242,1361,1609,1634]" italics="true" pageId="18" pageNumber="369">H. sindilla</emphasis>
|
||
</taxonomicName>
|
||
(as
|
||
<taxonomicName id="43944D57223BE37CFA192465B2C0F913" authorityName="Memon, Meier & Manan" authorityYear="2006" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="18" pageNumber="369" phylum="Chordata" rank="species" species="sindillus">
|
||
<emphasis id="B6E0EAC6223BE37CFA192465B2C0F913" italics="true" pageId="18" pageNumber="369">H. sindillus</emphasis>
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName id="43944D57223BE37CFEAE2441B398F910" authorityName="Shaikh, Memon & Shah" authorityYear="2011" box="[310,430,1646,1669]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="18" pageNumber="369" phylum="Chordata" rank="species" species="spinosa">
|
||
<emphasis id="B6E0EAC6223BE37CFEAE2441B398F910" box="[310,430,1646,1669]" italics="true" pageId="18" pageNumber="369">H. spinosa</emphasis>
|
||
</taxonomicName>
|
||
(as
|
||
<taxonomicName id="43944D57223BE37CFE462441B057F910" authorityName="Shaikh, Memon & Shah" authorityYear="2011" box="[478,609,1646,1669]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="18" pageNumber="369" phylum="Chordata" rank="species" species="spinosus">
|
||
<emphasis id="B6E0EAC6223BE37CFE462441B057F910" box="[478,609,1646,1669]" italics="true" pageId="18" pageNumber="369">H. spinosus</emphasis>
|
||
</taxonomicName>
|
||
) and based on the study of the specimens of
|
||
<taxonomicName id="43944D57223BE37CFBC52441B6E7F913" baseAuthorityName="Thunberg" baseAuthorityYear="1783" box="[1117,1233,1645,1670]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="18" pageNumber="369" phylum="Chordata" rank="species" species="sulcata">
|
||
<emphasis id="B6E0EAC6223BE37CFBC52441B6E7F913" box="[1117,1233,1645,1670]" italics="true" pageId="18" pageNumber="369">H. sulcata</emphasis>
|
||
</taxonomicName>
|
||
, it was found that there are no valid characters to separate them from
|
||
<taxonomicName id="43944D57223BE37CFD7B24BDB16EF93F" baseAuthorityName="Thunberg" baseAuthorityYear="1783" box="[739,856,1681,1706]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="18" pageNumber="369" phylum="Chordata" rank="species" species="sulcata">
|
||
<emphasis id="B6E0EAC6223BE37CFD7B24BDB16EF93F" box="[739,856,1681,1706]" italics="true" pageId="18" pageNumber="369">H. sulcata</emphasis>
|
||
</taxonomicName>
|
||
. The characters listed by
|
||
<bibRefCitation id="E0054B25223BE37CFBE724BDB75DF93F" author="Memon, N. & Meier, R. & Manan, A. & Su, K. F. - Y." box="[1151,1387,1681,1706]" pageId="18" pageNumber="369" pagination="703 - 710" refId="ref13758" refString="Memon, N., Meier, R., Manan, A. & Su, K. F. - Y. (2006) On the use of DNA sequences for determining the species limits of a polymorphic new species in the stink bug genus Halys (Heteroptera: Pentatomidae) from Pakistan. Systematic Entomology, 31 (4), 703 - 710. https: // doi. org / 10.1111 / j. 1365 - 3113.2006.00350. x" type="journal article" year="2006">
|
||
Memon
|
||
<emphasis id="B6E0EAC6223BE37CFB4624BDB72EF93F" box="[1246,1304,1681,1706]" italics="true" pageId="18" pageNumber="369">et al.</emphasis>
|
||
(2006)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="E0054B25223BE37CFF0F249AB34CF95B" author="Shaikh, A. M. & Memon, N. & Shah, M. A." box="[151,378,1717,1742]" pageId="18" pageNumber="369" pagination="157 - 160" refId="ref14285" refString="Shaikh, A. M., Memon, N. & Shah, M. A. (2011) A new halyine species of Halys Fabricius (Hemiptera, Pentatomidae, Pentatominae) from Miani forest, Sindh, Pakistan. Sindh University Research Journal, Science Series, 43 (2), 157 - 160." type="journal article" year="2011">
|
||
Shaikh
|
||
<emphasis id="B6E0EAC6223BE37CFF752499B31EF95B" box="[237,296,1717,1742]" italics="true" pageId="18" pageNumber="369">et al.</emphasis>
|
||
(2011)
|
||
</bibRefCitation>
|
||
such as tooth on lateral margin of head, length of paraclypei with respect to tylus, length of antennal segments, length of hemelytra with respect to abdomen, peritreme shape, shape of apical spine and apical margin of parameral crown, the extension of apical margin of paramere with respect to apical spine, shape of inner margin of paramere, shape of valvifers I in females are regarded as miner intraspecific variations. Similarly,
|
||
<emphasis id="B6E0EAC6223BE37CFA19250DB011F8CB" italics="true" pageId="18" pageNumber="369">
|
||
<taxonomicName id="43944D57223BE37CFA19250DB307F8CB" authorityName="Memon, Parveen, Ahmad & Shaikh" authorityYear="2016" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="18" pageNumber="369" phylum="Chordata" rank="species" species="mulberriensis">H. mulberriensis</taxonomicName>
|
||
and
|
||
<taxonomicName id="43944D57223BE37CFEF32569B015F8CB" authorityName="Memon, Parveen, Ahmad & Shah" authorityYear="2017" box="[363,547,1861,1886]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="18" pageNumber="369" phylum="Chordata" rank="species" species="noakoatensis">H. noakoatensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
are recognized as conspecific and also identical with
|
||
<taxonomicName id="43944D57223BE37CFB132569B6C9F8CB" baseAuthorityName="Thunberg" baseAuthorityYear="1783" box="[1163,1279,1861,1886]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="18" pageNumber="369" phylum="Chordata" rank="species" species="sulcata">
|
||
<emphasis id="B6E0EAC6223BE37CFB132569B6C9F8CB" box="[1163,1279,1861,1886]" italics="true" pageId="18" pageNumber="369">H. sulcata</emphasis>
|
||
</taxonomicName>
|
||
, based on the original descriptions and illustrations except for the weird characters like “very thin and spine-like” apical portions of dorsal conjunctival processes for
|
||
<taxonomicName id="43944D57223BE37CFDB125A1B0D3F833" authorityName="Memon, Parveen, Ahmad & Shaikh" authorityYear="2016" box="[553,741,1933,1958]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="18" pageNumber="369" phylum="Chordata" rank="species" species="mulberriensis">
|
||
<emphasis id="B6E0EAC6223BE37CFDB125A1B0D3F833" box="[553,741,1933,1958]" italics="true" pageId="18" pageNumber="369">H. mulberriensis</emphasis>
|
||
</taxonomicName>
|
||
and the particularly elongate dorsal membranous conjunctival processes with its thin and “sickle-shaped” apical end for
|
||
<taxonomicName id="43944D57223BE37CFCBF259DB1EBF85F" authorityName="Memon, Parveen, Ahmad & Shah" authorityYear="2017" box="[807,989,1969,1994]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="18" pageNumber="369" phylum="Chordata" rank="species" species="noakoatensis">
|
||
<emphasis id="B6E0EAC6223BE37CFCBF259DB1EBF85F" box="[807,989,1969,1994]" italics="true" pageId="18" pageNumber="369">H. noakoatensis</emphasis>
|
||
</taxonomicName>
|
||
. This single diagnostic character of the conjunctival processes appears to be merely an artifact, probably a result of an incomplete inflation or subsequent collapse of the conjunctiva. Other characters listed by
|
||
<bibRefCitation id="E0054B25223AE37DFC8A22B7B1CCFF25" author="Memon, N. & Parveen, R. & Ahmad, I. & Shaikh, A. M." box="[786,1018,151,176]" pageId="19" pageNumber="370" pagination="977 - 980" refId="ref13843" refString="Memon, N., Parveen, R., Ahmad, I. & Shaikh, A. M. (2016) Description of a new species of Oriental genus Halys (Pentatomoidae, Pentatominae, Halyini) from Lahore, Punjab, Pakistan. Pakistan Journal of Zoology, 48 (4), 977 - 980." type="journal article" year="2016">
|
||
Memon
|
||
<emphasis id="B6E0EAC6223AE37DFCEB22B6B19EFF25" box="[883,936,151,176]" italics="true" pageId="19" pageNumber="370">et al</emphasis>
|
||
. (2016
|
||
</bibRefCitation>
|
||
&
|
||
<bibRefCitation id="E0054B25223AE37DFBBC22B7B668FF25" author="Memon, N. & Parveen, R. & Ahmad, I. & Shah, N. A." box="[1060,1118,152,176]" pageId="19" pageNumber="370" pagination="1301 - 1306" refId="ref13904" refString="Memon, N., Parveen, R., Ahmad, I. & Shah, N. A. (2017) Description of two new species of Oriental genus Halys (Pentatomoidae, Pentatominae, Halyini) from Sindh, Pakistan with a key of its nine Asian species. Pakistan Journal of Zoology, 49 (4), 1301 - 1306. https: // doi. org / 10.17582 / journal. pjz / 2017.49.4.1301.1306" type="journal article" year="2017">2017</bibRefCitation>
|
||
) (length of paraclypei with respect to tylus, shape of inner spine of paramere, number of finger-like processes on spermathecal pump) are regarded merely as minor intraspecific variations. Therefore the following synonymies are proposed:
|
||
<taxonomicName id="43944D57223AE37DFABE22CEB36BFE89" authority="(Thunberg, 1783)" baseAuthorityName="Thunberg" baseAuthorityYear="1783" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="370" phylum="Chordata" rank="species" species="sulcata">
|
||
<emphasis id="B6E0EAC6223AE37DFABE22CEB7AAFF6D" box="[1318,1436,223,248]" italics="true" pageId="19" pageNumber="370">H. sulcata</emphasis>
|
||
(
|
||
<bibRefCitation id="E0054B25223AE37DFF06232BB363FE89" author="Thunberg, C. P." box="[158,341,260,285]" pageId="19" pageNumber="370" pagination="29 - 52" refId="ref14448" refString="Thunberg, C. P. (1783) Dissertatio entomologica novas insectorum species, sistens, cujus partem secundam, cons. exper. facult. med. upsal. publice ventilandam exhibent Johan. Edman, Upsala, 22 pp. [pp. 29 - 52]" type="book chapter" year="1783">Thunberg, 1783</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
=
|
||
<taxonomicName id="43944D57223AE37DFEE6232BB1BAFE88" authority="Memon, Meier & Manan, 2006" authorityName="Memon, Meier & Manan" authorityYear="2006" box="[382,908,260,285]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="370" phylum="Chordata" rank="species" species="sindilla">
|
||
<emphasis id="B6E0EAC6223AE37DFEE6232BB02DFE88" box="[382,539,260,285]" italics="true" pageId="19" pageNumber="370">Halys sindilla</emphasis>
|
||
<bibRefCitation id="E0054B25223AE37DFDBC232AB1BAFE88" author="Memon, N. & Meier, R. & Manan, A. & Su, K. F. - Y." box="[548,908,260,285]" pageId="19" pageNumber="370" pagination="703 - 710" refId="ref13758" refString="Memon, N., Meier, R., Manan, A. & Su, K. F. - Y. (2006) On the use of DNA sequences for determining the species limits of a polymorphic new species in the stink bug genus Halys (Heteroptera: Pentatomidae) from Pakistan. Systematic Entomology, 31 (4), 703 - 710. https: // doi. org / 10.1111 / j. 1365 - 3113.2006.00350. x" type="journal article" year="2006">Memon, Meier & Manan, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="ADD357BD223AE37DFC00232BB637FE88" box="[920,1025,260,285]" pageId="19" pageNumber="370">
|
||
<emphasis id="B6E0EAC6223AE37DFC00232BB637FE88" bold="true" box="[920,1025,260,285]" pageId="19" pageNumber="370">syn. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
=
|
||
<taxonomicName id="43944D57223AE37DFBBA232BB32AFED5" authority="Shaikh, Memon & Shah, 2011" authorityName="Shaikh, Memon & Shah" authorityYear="2011" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="370" phylum="Chordata" rank="species" species="spinosa">
|
||
<emphasis id="B6E0EAC6223AE37DFBBA232BB6F7FE89" box="[1058,1217,260,285]" italics="true" pageId="19" pageNumber="370">Halys spinosa</emphasis>
|
||
<bibRefCitation id="E0054B25223AE37DFB51232BB32AFED5" author="Shaikh, A. M. & Memon, N. & Shah, M. A." pageId="19" pageNumber="370" pagination="157 - 160" refId="ref14285" refString="Shaikh, A. M., Memon, N. & Shah, M. A. (2011) A new halyine species of Halys Fabricius (Hemiptera, Pentatomidae, Pentatominae) from Miani forest, Sindh, Pakistan. Sindh University Research Journal, Science Series, 43 (2), 157 - 160." type="journal article" year="2011">Shaikh, Memon & Shah, 2011</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="ADD357BD223AE37DFEB62308B3ADFED5" box="[302,411,295,320]" pageId="19" pageNumber="370">
|
||
<emphasis id="B6E0EAC6223AE37DFEB62308B3ADFED5" bold="true" box="[302,411,295,320]" pageId="19" pageNumber="370">syn. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
=
|
||
<taxonomicName id="43944D57223AE37DFE5D2308B684FED5" authority="Memon, Parveen, Ahmad & Shaikh, 2016" authorityName="Memon, Parveen, Ahmad & Shaikh" authorityYear="2016" box="[453,1202,295,320]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="370" phylum="Chordata" rank="species" species="mulberriensis">
|
||
<emphasis id="B6E0EAC6223AE37DFE5D2308B09BFED5" box="[453,685,295,320]" italics="true" pageId="19" pageNumber="370">Halys mulberriensis</emphasis>
|
||
<bibRefCitation id="E0054B25223AE37DFD222307B684FED5" author="Memon, N. & Parveen, R. & Ahmad, I. & Shaikh, A. M." box="[698,1202,295,320]" pageId="19" pageNumber="370" pagination="977 - 980" refId="ref13843" refString="Memon, N., Parveen, R., Ahmad, I. & Shaikh, A. M. (2016) Description of a new species of Oriental genus Halys (Pentatomoidae, Pentatominae, Halyini) from Lahore, Punjab, Pakistan. Pakistan Journal of Zoology, 48 (4), 977 - 980." type="journal article" year="2016">Memon, Parveen, Ahmad & Shaikh, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="ADD357BD223AE37DFB5B2308B707FED5" box="[1219,1329,295,320]" pageId="19" pageNumber="370">
|
||
<emphasis id="B6E0EAC6223AE37DFB5B2308B707FED5" bold="true" box="[1219,1329,295,320]" pageId="19" pageNumber="370">syn. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
=
|
||
<taxonomicName id="43944D57223AE37DFAC32308B0C2FEF1" authority="Memon, Parveen, Ahmad & Shah, 2017" authorityName="Memon, Parveen, Ahmad & Shah" authorityYear="2017" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="370" phylum="Chordata" rank="species" species="noakoatensis">
|
||
<emphasis id="B6E0EAC6223AE37DFAC32308B31FFEF0" italics="true" pageId="19" pageNumber="370">Halys noakoatensis</emphasis>
|
||
<bibRefCitation id="E0054B25223AE37DFEA82362B0C2FEF1" author="Memon, N. & Parveen, R. & Ahmad, I. & Shah, N. A." box="[304,756,332,357]" pageId="19" pageNumber="370" pagination="1301 - 1306" refId="ref13904" refString="Memon, N., Parveen, R., Ahmad, I. & Shah, N. A. (2017) Description of two new species of Oriental genus Halys (Pentatomoidae, Pentatominae, Halyini) from Sindh, Pakistan with a key of its nine Asian species. Pakistan Journal of Zoology, 49 (4), 1301 - 1306. https: // doi. org / 10.17582 / journal. pjz / 2017.49.4.1301.1306" type="journal article" year="2017">Memon, Parveen, Ahmad & Shah, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="ADD357BD223AE37DFD672363B151FEF0" box="[767,871,332,357]" pageId="19" pageNumber="370">
|
||
<emphasis id="B6E0EAC6223AE37DFD672363B151FEF0" bold="true" box="[767,871,332,357]" pageId="19" pageNumber="370">syn. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |