treatments-xml/data/03/C4/7D/03C47D45C749E724A0CACA2BFBB6B8BC.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.5825403" ID-GBIF-Dataset="54629b9a-8031-4e3b-bae4-79622c6be2a2" ID-GBIF-Taxon="191913329" ID-Zenodo-Dep="5825403" LSID="urn:lsid:zoobank.org:act:0D7EA76E-5268-47D2-882B-866E9FA8BAE4" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1641480009623" checkinUser="carolina" docAuthor="Weigmann, Simon, Ebert, David A. &amp; Séret, Bernard" docDate="2021" docId="03C47D45C749E724A0CACA2BFBB6B8BC" docLanguage="en" docName="MarBiodivers.51.58.1-30.pdf" docOrigin="Marine Biodiversity 51" docTitle="Acroteriobatus andysabini Weigmann &amp; Ebert &amp; Séret 2021, sp. nov." docType="treatment" docUuid="0D7EA76E-5268-47D2-882B-866E9FA8BAE4" docUuidSource="ZooBank" docVersion="8" lastPageId="9" lastPageNumber="10" masterDocId="FFFD053DC74BE72DA042C952FF93BB78" masterDocTitle="Resolution of the Acroteriobatus leucospilus species complex, with a redescription of A. leucospilus (Norman, 1926) and descriptions of two new western Indian Ocean species of Acroteriobatus (Rhinopristiformes, Rhinobatidae)" masterLastPageNumber="30" masterPageNumber="1" pageId="2" pageNumber="3" updateTime="1641489698393" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title>Resolution of the Acroteriobatus leucospilus species complex, with a redescription of A. leucospilus (Norman, 1926) and descriptions of two new western Indian Ocean species of Acroteriobatus (Rhinopristiformes, Rhinobatidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Weigmann, Simon</mods:namePart>
<mods:affiliation>Elasmo-Lab, Elasmobranch Research Laboratory, Georg-Bonne-Str. 5 83, 22609 Hamburg, Germany &amp; Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
<mods:nameIdentifier type="email">simon.weigmann@elasmo-lab.de &amp; simon.weigmann@unihamburg.de</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ebert, David A.</mods:namePart>
<mods:affiliation>Pacific Shark Research Center, Moss Landing Marine Laboratories, Moss Landing, CA 95039, USA &amp; South African Institute for Aquatic Biodiversity, Private Bag 1015, Grahamstown 6140, South Africa &amp; Department of Ichthyology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Séret, Bernard</mods:namePart>
<mods:affiliation>IchtyoConsult, 6 bis rue du Centre, 91430, Igny, France</mods:affiliation>
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<mods:title>Marine Biodiversity</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2021-06-21</mods:number>
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<mods:number>51</mods:number>
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<mods:identifier type="DOI">10.1007/s12526-021-01208-6</mods:identifier>
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<taxonomicName authority="Weigmann &amp; Ebert &amp; Séret, 2021" authorityName="Weigmann &amp; Ebert &amp; Séret" authorityYear="2021" box="[136,404,889,913]" class="Elasmobranchii" family="Rhinobatidae" genus="Acroteriobatus" kingdom="Animalia" order="Rhinopristiformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="andysabini" status="sp. nov.">
<emphasis bold="true" box="[136,404,889,913]" italics="true" pageId="2" pageNumber="3">Acroteriobatus andysabini</emphasis>
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<taxonomicNameLabel box="[409,490,890,913]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
Weigmann, Ebert &amp; Séret
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http://zoobank.org/ 
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<paragraph blockId="2.[136,771,889,1577]" box="[136,493,956,979]" pageId="2" pageNumber="3">(Malagasy blue-spotted guitarfish)</paragraph>
</subSubSection>
<subSubSection box="[166,763,989,1012]" pageId="2" pageNumber="3" type="description">
<paragraph blockId="2.[136,771,889,1577]" box="[166,763,989,1012]" pageId="2" pageNumber="3">
(
<figureCitation box="[174,246,989,1012]" captionStart="Fig" captionStartId="2.[136,165,1835,1855]" captionTargetBox="[137,770,1616,1817]" captionTargetId="figure-507@2.[136,771,1615,1818]" captionTargetPageId="2" captionText="Fig. 1 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 565 mm TL fresh, in total dorsal view taken directly after catching. Photograph by Elaine Heemstra, NRF-SAIAB" figureDoi="http://doi.org/10.5281/zenodo.5825449" httpUri="https://zenodo.org/record/5825449/files/figure.png" pageId="2" pageNumber="3">Figs. 1</figureCitation>
, 
<figureCitation box="[259,272,989,1012]" captionStart="Fig" captionStartId="2.[816,845,585,605]" captionTargetBox="[819,1448,146,568]" captionTargetId="figure-536@2.[818,1449,144,569]" captionTargetPageId="2" captionText="Fig. 2 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, in total dorsal view. Scale bar: 5 cm. Photograph courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825447" httpUri="https://zenodo.org/record/5825447/files/figure.png" pageId="2" pageNumber="3">2</figureCitation>
, 
<figureCitation box="[286,299,989,1012]" captionStart="Fig" captionStartId="2.[816,845,1835,1855]" captionTargetBox="[827,1448,1400,1818]" captionTargetId="figure-564@2.[818,1449,1399,1819]" captionTargetPageId="2" captionText="Fig. 3 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, in total ventral view. Scale bar: 5 cm. Photograph courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825451" httpUri="https://zenodo.org/record/5825451/files/figure.png" pageId="2" pageNumber="3">3</figureCitation>
, 
<figureCitation box="[312,325,989,1012]" captionStart="Fig" captionStartId="3.[136,165,554,574]" captionTargetBox="[136,789,126,543]" captionTargetId="figure-318@3.[138,769,144,538]" captionTargetPageId="3" captionText="Fig. 4 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 565 mm TL fresh, head in dorsal view taken directly after catching. Photograph by Elaine Heemstra, NRF-SAIAB" figureDoi="http://doi.org/10.5281/zenodo.5825453" httpUri="https://zenodo.org/record/5825453/files/figure.png" pageId="2" pageNumber="3">4</figureCitation>
, 
<figureCitation box="[339,352,989,1012]" captionStart="Fig" captionStartId="3.[136,165,1835,1855]" captionTargetBox="[139,768,1268,1818]" captionTargetId="figure-347@3.[138,769,1267,1819]" captionTargetPageId="3" captionText="Fig. 5 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, head in dorsal view. Photograph courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825457" httpUri="https://zenodo.org/record/5825457/files/figure.png" pageId="2" pageNumber="3">5</figureCitation>
, 
<figureCitation box="[366,379,989,1012]" captionStart="Fig" captionStartId="3.[816,845,581,601]" captionTargetBox="[807,1451,123,566]" captionTargetId="figure-372@3.[818,1449,144,565]" captionText="Fig. 6 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, close-up of orbital and spiracular region. Photograph courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825455" httpUri="https://zenodo.org/record/5825455/files/figure.png" pageId="2" pageNumber="3">6</figureCitation>
, 
<figureCitation box="[392,405,989,1012]" captionStart="Fig" captionStartId="3.[816,845,1835,1855]" captionTargetBox="[819,1448,1441,1817]" captionTargetId="figure-398@3.[818,1449,1440,1818]" captionTargetPageId="3" captionText="Fig. 7 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 565 mm TL fresh, head in ventral view taken directly after catching. Photograph by Elaine Heemstra, NRF-SAIAB" figureDoi="http://doi.org/10.5281/zenodo.5825459" httpUri="https://zenodo.org/record/5825459/files/figure.png" pageId="2" pageNumber="3">7</figureCitation>
, 
<figureCitation box="[419,432,989,1012]" captionStart="Fig" captionStartId="4.[136,165,747,767]" captionTargetBox="[139,768,145,729]" captionTargetId="figure-222@4.[138,769,144,730]" captionTargetPageId="4" captionText="Fig. 8 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, head in ventral view. Photograph courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825461" httpUri="https://zenodo.org/record/5825461/files/figure.png" pageId="2" pageNumber="3">8</figureCitation>
, 
<figureCitation box="[445,458,989,1012]" captionStart="Fig" captionStartId="4.[136,165,1835,1855]" captionTargetBox="[139,768,1399,1818]" captionTargetId="figure-246@4.[138,769,1398,1819]" captionTargetPageId="4" captionText="Fig. 9 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, close-up of mouth-nasal region. Photograph courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825466" httpUri="https://zenodo.org/record/5825466/files/figure.png" pageId="2" pageNumber="3">9</figureCitation>
, 
<figureCitation box="[472,498,989,1012]" captionStart="Fig" captionStartId="4.[816,845,1425,1445]" captionTargetBox="[819,1448,145,1337]" captionTargetId="figure-272@4.[818,1449,144,1409]" captionTargetPageId="4" captionText="Fig. 10 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL; first dorsal (a), second dorsal (b), and caudal (c) fins in lateral views (c image reversed). Photographs courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825463" httpUri="https://zenodo.org/record/5825463/files/figure.png" pageId="2" pageNumber="3">10</figureCitation>
, 
<figureCitation box="[512,538,989,1012]" captionStart="Fig" captionStartId="5.[136,165,607,627]" captionTargetBox="[139,768,145,589]" captionTargetId="figure-686@5.[138,769,144,590]" captionTargetPageId="5" captionText="Fig. 11 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, radiograph of cranium and snout in dorsal view. The radiograph was taken and kindly provided by Jon Fong" figureDoi="http://doi.org/10.5281/zenodo.5825468" httpUri="https://zenodo.org/record/5825468/files/figure.png" pageId="2" pageNumber="3">11</figureCitation>
, 
<figureCitation box="[551,577,989,1012]" captionStart="Fig" captionStartId="5.[136,168,1834,1854]" captionTargetBox="[139,768,1345,1817]" captionTargetId="figure-720@5.[138,769,1344,1818]" captionTargetPageId="5" captionText="Fig. 12 Acroteriobatus andysabini sp. nov., Malagasy artisanal fisherman with large (~1000 mm TL) specimen. The photograph was taken and kindly provided by Kerry Sink" figureDoi="http://doi.org/10.5281/zenodo.5825470" httpUri="https://zenodo.org/record/5825470/files/figure.png" pageId="2" pageNumber="3">12</figureCitation>
, and 13, 
<tableCitation box="[675,754,989,1012]" captionStart="Table 1" captionStartId="7.[136,184,155,175]" captionTargetBox="[136,1452,205,1897]" captionText="Table 1 Acroteriobatus andysabini sp. nov., morphometrics and meristics Acroteriobatus Acroteriobatus Minimum Maximum Mean SD andysabini sp. andysabini sp. (n = 9) (n = 9) (n = 11) nov., juvenile nov., juvenile male holotype, female paratype, SAIAB 97396 SAIAB 189815" pageId="2" pageNumber="3">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="reference_group">
<paragraph blockId="2.[136,771,889,1577]" box="[166,665,1022,1046]" pageId="2" pageNumber="3">
<treatmentCitationGroup box="[166,665,1022,1046]" pageId="2" pageNumber="3">
<taxonomicName authorityName="Smith" box="[166,393,1023,1046]" class="Insecta" family="Curculionidae" genus="Rhinobatus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="annulatus">
<emphasis box="[166,393,1023,1046]" italics="true" pageId="2" pageNumber="3">Rhinobatus annulatus</emphasis>
</taxonomicName>
― 
<treatmentCitation author="Fourmanoir P" box="[420,661,1023,1046]" page="35" pageId="2" pageNumber="3" year="1963">
<bibRefCitation author="Fourmanoir P" box="[420,661,1023,1046]" journalOrPublisher="Cah ORSTOM Oceanogr" pageId="2" pageNumber="3" pagination="33 - 58" part="6" refId="ref22501" refString="Fourmanoir P (1963) Raies et requins-scie de la cote ouest de Madagascar (Ordre des Batoidei). Cah ORSTOM Oceanogr 1963 (6): 33 - 58" title="Raies et requins-scie de la cote ouest de Madagascar (Ordre des Batoidei)" type="journal article" year="1963">Fourmanoir (1963: 35)</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph blockId="2.[136,771,889,1577]" box="[166,628,1055,1079]" pageId="2" pageNumber="3">
<treatmentCitationGroup box="[166,628,1055,1079]" pageId="2" pageNumber="3">
<taxonomicName authorityName="Muller &amp; Henle" authorityYear="1841" box="[166,393,1056,1079]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rhinopristiformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="annulatus">
<emphasis box="[166,393,1056,1079]" italics="true" pageId="2" pageNumber="3">Rhinobatos annulatus</emphasis>
</taxonomicName>
― 
<treatmentCitation author="Mauge AL" box="[420,625,1056,1079]" page="217" pageId="2" pageNumber="3" year="1967">
<bibRefCitation author="Mauge AL" box="[420,625,1056,1079]" journalOrPublisher="Ann Fac Sci Univ Madagascar" pageId="2" pageNumber="3" pagination="215 - 246" part="5" refId="ref23365" refString="Mauge AL (1967) Contribution preliminaire a l' inventaire ichtyologique de la region de Tulear. Ann Fac Sci Univ Madagascar 5: 215 - 246" title="Contribution preliminaire a l' inventaire ichtyologique de la region de Tulear" type="journal article" year="1967">Maugé (1967: 217)</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph blockId="2.[136,771,889,1577]" box="[166,716,1088,1112]" pageId="2" pageNumber="3">
<treatmentCitationGroup box="[166,716,1088,1112]" pageId="2" pageNumber="3">
<taxonomicName baseAuthorityName="Norman" baseAuthorityYear="1926" box="[166,444,1089,1112]" class="Elasmobranchii" family="Rhinobatidae" genus="Acroteriobatus" kingdom="Animalia" order="Rhinopristiformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="leucospilus">
<emphasis box="[166,444,1089,1112]" italics="true" pageId="2" pageNumber="3">Acroteriobatus leucospilus</emphasis>
</taxonomicName>
― 
<treatmentCitation author="Fricke R &amp; Mahafina J &amp; Behivoke F &amp; Jaonalison H &amp; Leopold M &amp; Ponton D" box="[470,712,1089,1112]" page="23" pageId="2" pageNumber="3" year="2018">
<bibRefCitation author="Fricke R &amp; Mahafina J &amp; Behivoke F &amp; Jaonalison H &amp; Leopold M &amp; Ponton D" box="[470,712,1089,1112]" journalOrPublisher="FishTaxa" pageId="2" pageNumber="3" pagination="1 - 432" part="3" refId="ref22596" refString="Fricke R, Mahafina J, Behivoke F, Jaonalison H, Leopold M, Ponton D (2018) Annotated checklist of the fishes of Madagascar, southwestern Indian Ocean, with 158 new records. FishTaxa 3 (1): 1 - 432" title="Annotated checklist of the fishes of Madagascar, southwestern Indian Ocean, with 158 new records" type="journal article" year="2018">Fricke et al. (2018: 23)</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph blockId="2.[136,771,889,1577]" box="[166,696,1122,1146]" pageId="2" pageNumber="3">
<treatmentCitationGroup box="[166,696,1122,1146]" pageId="2" pageNumber="3">
<taxonomicName baseAuthorityName="Norman" baseAuthorityYear="1926" box="[166,444,1122,1145]" class="Elasmobranchii" family="Rhinobatidae" genus="Acroteriobatus" kingdom="Animalia" order="Rhinopristiformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="leucospilus">
<emphasis box="[166,444,1122,1145]" italics="true" pageId="2" pageNumber="3">Acroteriobatus leucospilus</emphasis>
</taxonomicName>
― 
<treatmentCitation author="Ghilardi et al." box="[470,696,1122,1145]" pageId="2" pageNumber="3" year="2019">
<bibRefCitation author="Ghilardi M &amp; Emmanuel P &amp; Gibbons E" box="[470,692,1122,1145]" journalOrPublisher="University of Mauritius, Reduit" pageId="2" pageNumber="3" refId="ref22642" refString="Ghilardi M, Emmanuel P, Gibbons E (2019) Guitarfish fishing in southwest Madagascar: an assessment of the status of grayspotted guitarfish (Acroteriobatus leucospilus). WIOMSA 11 th scientific symposium, July 1 - 6, 2019, University of Mauritius, Reduit" title="Guitarfish fishing in southwest Madagascar: an assessment of the status of grayspotted guitarfish (Acroteriobatus leucospilus)" type="journal volume" year="2019">Ghilardi et al. (2019)</bibRefCitation>
.
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<paragraph blockId="2.[136,771,889,1577]" pageId="2" pageNumber="3">
The 
<typeStatus box="[212,299,1155,1178]" pageId="2" pageNumber="3">holotype</typeStatus>
and 
<specimenCount box="[348,505,1155,1178]" pageId="2" pageNumber="3" type="generic" typeStatus="paratypes">seven paratypes</specimenCount>
are deposited in the South African Institute for Aquatic Biodiversity (
<collectionCode box="[611,696,1189,1212]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="2" pageNumber="3" type="Other type of research institution/biorepository">SAIAB</collectionCode>
); 
<specimenCount pageId="2" pageNumber="3" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
in the Muséum national d’ Histoire naturelle, Paris (
<collectionCode box="[143,226,1255,1278]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="3">MNHN</collectionCode>
); 
<specimenCount box="[245,386,1255,1278]" pageId="2" pageNumber="3" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
in the American Museum of Natural History, 
<collectingRegion box="[224,332,1288,1311]" country="United States of America" name="New York" pageId="2" pageNumber="3">New York</collectingRegion>
(
<collectionCode box="[347,430,1288,1311]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="3">AMNH</collectionCode>
); and 
<specimenCount box="[493,622,1288,1311]" pageId="2" pageNumber="3" type="generic" typeStatus="paratype">one paratype</specimenCount>
in each of the 
<collectingRegion box="[136,235,1322,1345]" country="United States of America" name="California" pageId="2" pageNumber="3">California</collectingRegion>
Academy of Sciences (
<collectionCode box="[464,515,1322,1345]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="2" pageNumber="3" type="Museum">CAS</collectionCode>
) and National Museum of Natural History, Smithsonian Institution, 
<collectingRegion box="[584,771,1355,1378]" country="United States of America" name="District of Columbia" pageId="2" pageNumber="3">Washington D.C.</collectingRegion>
(
<collectionCode box="[143,221,1388,1411]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="3" type="Museum">USNM</collectionCode>
).
</paragraph>
<paragraph blockId="2.[136,771,889,1577]" pageId="2" pageNumber="3">
<materialsCitation ID-GBIF-Occurrence="3427235332" collectingDate="2010-04-27" collectionCode="SAIAB" collectorName="E. Heemstra &amp; T. Razafimanambina &amp; P. C. Heemstra" country="Madagascar" latitude="-25.027517" location="Douane Beach" longLatPrecision="1" longitude="46.9956" pageId="2" pageNumber="3" specimenCode="SAIAB 97396" specimenCount="1" typeStatus="holotype">
<typeStatus box="[166,263,1421,1444]" pageId="2" pageNumber="3">Holotype</typeStatus>
<emphasis bold="true" box="[269,420,1421,1444]" pageId="2" pageNumber="3">
<specimenCode box="[269,420,1421,1444]" pageId="2" pageNumber="3">
SAIAB 
<quantity box="[355,420,1421,1444]" metricMagnitude="4" metricUnit="m" metricValue="9.7396565" pageId="2" pageNumber="3" unit="mm" value="9.7396565E7">97396</quantity>
</specimenCode>
</emphasis>
, 565 mm TL (fresh), 
<quantity box="[646,733,1421,1444]" metricMagnitude="-1" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="3" unit="mm" value="550.0">550 mm</quantity>
TL (after preservation), 
<specimenCount box="[370,527,1454,1477]" pageId="2" pageNumber="3" type="juvenile">juvenile male</specimenCount>
, 
<location LSID="urn:lsid:plazi:treatment:03C47D45C749E724A0CACA2BFBB6B8BC:8EB29A88C749E72FA25CCCFCFD56BEBD" box="[542,709,1454,1477]" country="Madagascar" latitude="-25.027517" longLatPrecision="1" longitude="46.9956" name="Douane Beach" pageId="2" pageNumber="3">Douane Beach</location>
, 
<location LSID="urn:lsid:plazi:treatment:03C47D45C749E724A0CACA2BFBB6B8BC:8EB29A88C749E72FA297CCFCFF74BE9F" country="Madagascar" latitude="-25.027517" longLatPrecision="1" longitude="46.9956" name="Fort Dauphin" pageId="2" pageNumber="3">Fort Dauphin</location>
, Anosy, southeastern 
<collectingCountry box="[477,609,1488,1511]" name="Madagascar" pageId="2" pageNumber="3">Madagascar</collectingCountry>
, 
<geoCoordinate box="[624,766,1487,1511]" degrees="25" direction="south" minutes="1.651" orientation="latitude" pageId="2" pageNumber="3" precision="1" value="-25.027517">25° 1.651′ S</geoCoordinate>
, 
<geoCoordinate box="[136,283,1520,1544]" degrees="46" direction="east" minutes="59.736" orientation="longitude" pageId="2" pageNumber="3" precision="1" value="46.9956">46° 59.736′ E</geoCoordinate>
, collectors 
<collectorName box="[403,534,1521,1544]" pageId="2" pageNumber="3">E. Heemstra</collectorName>
, 
<collectorName box="[546,768,1521,1544]" pageId="2" pageNumber="3">T. Razafimanambina</collectorName>
, and 
<collectorName box="[180,333,1554,1577]" pageId="2" pageNumber="3">P.C. Heemstra</collectorName>
, 
<date box="[344,495,1554,1577]" pageId="2" pageNumber="3" value="2010-04-27">
<collectingDate box="[344,495,1554,1577]" pageId="2" pageNumber="3" value="2010-04-27">27 April 2010</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph blockId="2.[816,1452,703,1357]" lastBlockId="3.[816,1452,698,887]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<typeStatus box="[846,942,703,726]" pageId="2" pageNumber="3">Paratypes</typeStatus>
(14) 
<materialsCitation ID-GBIF-Occurrence="3427235314" collectingDate="2010-05-08" collectionCode="NOSY" country="Madagascar" elevation="70" latitude="-25.091667" location="Fort Dauphin" longLatPrecision="124" longitude="47.091667" pageId="2" pageNumber="3" specimenCode="SAIAB 189672" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1">
<emphasis bold="true" box="[997,1157,703,726]" pageId="2" pageNumber="3">
<specimenCode box="[997,1157,703,726]" pageId="2" pageNumber="3">
SAIAB 
<quantity box="[1082,1157,703,726]" metricMagnitude="5" metricUnit="m" metricValue="1.896722" pageId="2" pageNumber="3" unit="mm" value="1.896722E8">189672</quantity>
</specimenCode>
</emphasis>
, 200 mm TL (after preservation), 
<specimenCount box="[881,964,736,759]" pageId="2" pageNumber="3" type="juvenile">juvenile</specimenCount>
<specimenCount box="[974,1027,736,759]" pageId="2" pageNumber="3" type="male">male</specimenCount>
, Secteur de 
<location LSID="urn:lsid:plazi:treatment:03C47D45C749E724A0CACA2BFBB6B8BC:8EB29A88C749E72FA4C9CBB2FA8FB98F" box="[1163,1308,736,759]" country="Madagascar" latitude="-25.091667" longLatPrecision="124" longitude="47.091667" name="Fort Dauphin" pageId="2" pageNumber="3">Fort Dauphin</location>
, southeastern 
<collectingCountry box="[816,938,769,792]" name="Madagascar" pageId="2" pageNumber="3">Madagascar</collectingCountry>
, 
<geoCoordinate box="[950,1053,769,793]" degrees="25" direction="south" minutes="5.5" orientation="latitude" pageId="2" pageNumber="3" precision="92" value="-25.091667">25° 5.5′ S</geoCoordinate>
, 
<geoCoordinate box="[1064,1169,769,793]" degrees="47" direction="east" minutes="5.5" orientation="longitude" pageId="2" pageNumber="3" precision="92" value="47.091667">47° 5.5′ E</geoCoordinate>
, 
<quantity box="[1181,1272,769,793]" metricMagnitude="1" metricUnit="m" metricValue="6.95" metricValueMax="7.0" metricValueMin="6.9" pageId="2" pageNumber="3" unit="m" value="69.5" valueMax="70.0" valueMin="69.0">
<elevation box="[1181,1272,769,793]" metricMagnitude="1" metricUnit="m" metricValue="6.95" metricValueMax="7.0" metricValueMin="6.9" pageId="2" pageNumber="3" unit="m" value="69.5" valueMax="70.0" valueMin="69.0">69–70 m</elevation>
</quantity>
depth, FV 
<emphasis italics="true" pageId="2" pageNumber="3">
<collectionCode box="[1389,1451,769,792]" pageId="2" pageNumber="3">NOSY</collectionCode>
BE 11
</emphasis>
, station CP 3569, beam trawl, 
<date box="[1222,1358,802,825]" pageId="2" pageNumber="3" value="2010-05-08">
<collectingDate box="[1222,1358,802,825]" pageId="2" pageNumber="3" value="2010-05-08">8 May 2010</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="3427235329" collectingDate="2010-05-08" collectionCode="NOSY" country="Madagascar" elevation="65" latitude="-25.078333" location="Fort Dauphin" longLatPrecision="124" longitude="47.056667" pageId="2" pageNumber="3" specimenCode="SAIAB 189683" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1">
<emphasis bold="true" pageId="2" pageNumber="3">
<specimenCode pageId="2" pageNumber="3">
SAIAB 
<quantity box="[816,897,836,859]" metricMagnitude="5" metricUnit="m" metricValue="1.89683178" pageId="2" pageNumber="3" unit="mm" value="1.89683178E8">189683</quantity>
</specimenCode>
</emphasis>
, 178 mm TL (after preservation), 
<specimenCount box="[1294,1381,836,859]" pageId="2" pageNumber="3" type="juvenile">juvenile</specimenCount>
<specimenCount box="[1392,1447,836,859]" pageId="2" pageNumber="3" type="male">male</specimenCount>
, Secteur de 
<location LSID="urn:lsid:plazi:treatment:03C47D45C749E724A0CACA2BFBB6B8BC:8EB29A88C749E72FA3E0CA37FBBFB804" box="[930,1068,869,892]" country="Madagascar" latitude="-25.078333" longLatPrecision="124" longitude="47.056667" name="Fort Dauphin" pageId="2" pageNumber="3">Fort Dauphin</location>
, southeastern 
<collectingCountry box="[1210,1332,869,892]" name="Madagascar" pageId="2" pageNumber="3">Madagascar</collectingCountry>
, 
<geoCoordinate box="[1343,1447,868,892]" degrees="25" direction="south" minutes="4.7" orientation="latitude" pageId="2" pageNumber="3" precision="92" value="-25.078333">25° 4.7′ S</geoCoordinate>
, 
<geoCoordinate box="[816,920,901,925]" degrees="47" direction="east" minutes="3.4" orientation="longitude" pageId="2" pageNumber="3" precision="92" value="47.056667">47° 3.4′ E</geoCoordinate>
, 
<quantity box="[931,1022,901,925]" metricMagnitude="1" metricUnit="m" metricValue="6.45" metricValueMax="6.5" metricValueMin="6.4" pageId="2" pageNumber="3" unit="m" value="64.5" valueMax="65.0" valueMin="64.0">
<elevation box="[931,1022,901,925]" metricMagnitude="1" metricUnit="m" metricValue="6.45" metricValueMax="6.5" metricValueMin="6.4" pageId="2" pageNumber="3" unit="m" value="64.5" valueMax="65.0" valueMin="64.0">64–65 m</elevation>
</quantity>
depth, FV 
<emphasis box="[1137,1270,902,925]" italics="true" pageId="2" pageNumber="3">
<collectionCode box="[1137,1199,902,925]" pageId="2" pageNumber="3">NOSY</collectionCode>
BE 11
</emphasis>
, station CP 3568, beam trawl, 
<date box="[956,1096,935,958]" pageId="2" pageNumber="3" value="2010-05-08">
<collectingDate box="[956,1096,935,958]" pageId="2" pageNumber="3" value="2010-05-08">8 May 2010</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3427235319" collectingDate="2010-04-29" collectionCode="NOSY" country="Madagascar" elevation="80" latitude="-25.243334" location="Fort Dauphin" longLatPrecision="124" longitude="47.15167" pageId="2" pageNumber="3" specimenCode="SAIAB 189815" specimenCount="6" specimenCount-female="1" specimenCount-juvenile="2" specimenCount-male="1">
<emphasis bold="true" box="[1110,1282,935,958]" pageId="2" pageNumber="3">
<specimenCode box="[1110,1282,935,958]" pageId="2" pageNumber="3">SAIAB 189815</specimenCode>
</emphasis>
(
<specimenCount box="[1301,1440,935,958]" pageId="2" pageNumber="3" type="generic">2 specimens</specimenCount>
), 
<quantity box="[816,904,969,992]" metricMagnitude="-1" metricUnit="m" metricValue="2.15" pageId="2" pageNumber="3" unit="mm" value="215.0">215 mm</quantity>
TL (after preservation), 
<specimenCount box="[1166,1247,969,992]" pageId="2" pageNumber="3" type="juvenile">juvenile</specimenCount>
<specimenCount box="[1257,1306,969,992]" pageId="2" pageNumber="3" type="male">male</specimenCount>
and 
<quantity box="[1362,1451,969,992]" metricMagnitude="-1" metricUnit="m" metricValue="3.45" pageId="2" pageNumber="3" unit="mm" value="345.0">345 mm</quantity>
TL (after preservation), 
<specimenCount box="[1089,1176,1002,1025]" pageId="2" pageNumber="3" type="juvenile">juvenile</specimenCount>
<specimenCount box="[1187,1263,1002,1025]" pageId="2" pageNumber="3" type="female">female</specimenCount>
, Secteur de 
<location LSID="urn:lsid:plazi:treatment:03C47D45C749E724A0CACA2BFBB6B8BC:8EB29A88C749E72FA53CCAB8FC18BF5A" country="Madagascar" latitude="-25.243334" longLatPrecision="124" longitude="47.15167" name="Fort Dauphin" pageId="2" pageNumber="3">Fort Dauphin</location>
, southeastern 
<collectingCountry box="[1058,1184,1035,1058]" name="Madagascar" pageId="2" pageNumber="3">Madagascar</collectingCountry>
, 
<geoCoordinate box="[1197,1320,1034,1058]" degrees="25" direction="south" minutes="14.6" orientation="latitude" pageId="2" pageNumber="3" precision="92" value="-25.243334">25° 14.6′ S</geoCoordinate>
, 
<geoCoordinate box="[1335,1447,1034,1058]" degrees="47" direction="east" minutes="9.1" orientation="longitude" pageId="2" pageNumber="3" precision="92" value="47.15167">47° 9.1′ E</geoCoordinate>
, 
<quantity box="[816,906,1068,1092]" metricMagnitude="1" metricUnit="m" metricValue="7.95" metricValueMax="8.0" metricValueMin="7.9" pageId="2" pageNumber="3" unit="m" value="79.5" valueMax="80.0" valueMin="79.0">
<elevation box="[816,906,1068,1092]" metricMagnitude="1" metricUnit="m" metricValue="7.95" metricValueMax="8.0" metricValueMin="7.9" pageId="2" pageNumber="3" unit="m" value="79.5" valueMax="80.0" valueMin="79.0">79–80 m</elevation>
</quantity>
depth, FV 
<emphasis box="[1019,1149,1068,1091]" italics="true" pageId="2" pageNumber="3">
<collectionCode box="[1019,1081,1068,1091]" pageId="2" pageNumber="3">NOSY</collectionCode>
BE 11
</emphasis>
, station CP 3510, beam trawl, 
<date box="[816,956,1101,1124]" pageId="2" pageNumber="3" value="2010-04-29">
<collectingDate box="[816,956,1101,1124]" pageId="2" pageNumber="3" value="2010-04-29">29 April 2010</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3427235326" collectingDate="2010-05-04" collectionCode="NOSY" country="Madagascar" elevation="54" latitude="-25.281666" location="Madagascar" longLatPrecision="124" longitude="46.521667" pageId="2" pageNumber="3" specimenCode="SAIAB 189816" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1">
<emphasis bold="true" box="[965,1122,1101,1124]" pageId="2" pageNumber="3">
<specimenCode box="[965,1122,1101,1124]" pageId="2" pageNumber="3">
SAIAB 
<quantity box="[1047,1122,1101,1124]" metricMagnitude="5" metricUnit="m" metricValue="1.89816602" pageId="2" pageNumber="3" unit="mm" value="1.89816602E8">189816</quantity>
</specimenCode>
</emphasis>
, 602 mm TL (fresh), 
<quantity box="[1332,1415,1101,1124]" metricMagnitude="-1" metricUnit="m" metricValue="5.95" pageId="2" pageNumber="3" unit="mm" value="595.0">595 mm</quantity>
TL (after preservation), 
<specimenCount box="[1024,1104,1135,1158]" pageId="2" pageNumber="3" type="juvenile">juvenile</specimenCount>
<specimenCount box="[1113,1164,1135,1158]" pageId="2" pageNumber="3" type="male">male</specimenCount>
, Sud-Ouest Cap Andavaka, southeastern 
<collectingCountry box="[950,1074,1168,1191]" name="Madagascar" pageId="2" pageNumber="3">Madagascar</collectingCountry>
, 
<geoCoordinate box="[1087,1207,1167,1191]" degrees="25" direction="south" minutes="16.9" orientation="latitude" pageId="2" pageNumber="3" precision="92" value="-25.281666">25° 16.9′ S</geoCoordinate>
, 
<geoCoordinate box="[1221,1344,1167,1191]" degrees="46" direction="east" minutes="31.3" orientation="longitude" pageId="2" pageNumber="3" precision="92" value="46.521667">46° 31.3′ E</geoCoordinate>
, 
<quantity box="[1357,1451,1167,1191]" metricMagnitude="1" metricUnit="m" metricValue="5.35" metricValueMax="5.4" metricValueMin="5.3" pageId="2" pageNumber="3" unit="m" value="53.5" valueMax="54.0" valueMin="53.0">
<elevation box="[1357,1451,1167,1191]" metricMagnitude="1" metricUnit="m" metricValue="5.35" metricValueMax="5.4" metricValueMin="5.3" pageId="2" pageNumber="3" unit="m" value="53.5" valueMax="54.0" valueMin="53.0">53–54 m</elevation>
</quantity>
depth, FV 
<emphasis box="[938,1084,1201,1224]" italics="true" pageId="2" pageNumber="3">
<collectionCode box="[938,1003,1201,1224]" pageId="2" pageNumber="3">NOSY</collectionCode>
BE 11
</emphasis>
, station CP 3549, beam trawl, 
<date pageId="2" pageNumber="3" value="2010-05-04">
<collectingDate pageId="2" pageNumber="3" value="2010-05-04">4 May 2010</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3427235331" collectingDate="2010-05-08" collectionCode="NOSY" country="Madagascar" elevation="75" latitude="-25.11" location="Fort Dauphin" longLatPrecision="124" longitude="47.115" pageId="2" pageNumber="3" specimenCode="SAIAB 189943" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1">
<emphasis bold="true" box="[939,1101,1234,1257]" pageId="2" pageNumber="3">
<specimenCode box="[939,1101,1234,1257]" pageId="2" pageNumber="3">
SAIAB 
<quantity box="[1026,1101,1234,1257]" metricMagnitude="5" metricUnit="m" metricValue="1.8994319499999999" pageId="2" pageNumber="3" unit="mm" value="1.89943195E8">189943</quantity>
</specimenCode>
</emphasis>
, 195 mm TL (after preservation), 
<specimenCount box="[816,908,1267,1290]" pageId="2" pageNumber="3" type="juvenile">juvenile</specimenCount>
<specimenCount box="[921,979,1267,1290]" pageId="2" pageNumber="3" type="male">male</specimenCount>
, Secteur de 
<location LSID="urn:lsid:plazi:treatment:03C47D45C749E724A0CACA2BFBB6B8BC:8EB29A88C749E72FA42ECDA1FA9FBE72" box="[1132,1292,1267,1290]" country="Madagascar" latitude="-25.11" longLatPrecision="124" longitude="47.115" name="Fort Dauphin" pageId="2" pageNumber="3">Fort Dauphin</location>
, southeastern 
<collectingCountry box="[816,938,1301,1324]" name="Madagascar" pageId="2" pageNumber="3">Madagascar</collectingCountry>
, 
<geoCoordinate box="[949,1052,1300,1324]" degrees="25" direction="south" minutes="6.6" orientation="latitude" pageId="2" pageNumber="3" precision="92" value="-25.11">25° 6.6′ S</geoCoordinate>
, 
<geoCoordinate box="[1064,1169,1300,1324]" degrees="47" direction="east" minutes="6.9" orientation="longitude" pageId="2" pageNumber="3" precision="92" value="47.115">47° 6.9′ E</geoCoordinate>
, 
<quantity box="[1180,1233,1301,1324]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="3" unit="m" value="75.0">
<elevation box="[1180,1233,1301,1324]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="3" unit="m" value="75.0">75 m</elevation>
</quantity>
depth, FV 
<emphasis italics="true" pageId="2" pageNumber="3">
<collectionCode box="[1350,1412,1301,1324]" pageId="2" pageNumber="3">NOSY</collectionCode>
BE 11
</emphasis>
, station CP 3570, beam trawl, 
<date box="[1150,1275,1334,1357]" pageId="2" pageNumber="3" value="2010-05-08">
<collectingDate box="[1150,1275,1334,1357]" pageId="2" pageNumber="3" value="2010-05-08">8 May 2010</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3427235325" collectingDate="2010-05-08" collectionCode="NOSY" country="Madagascar" elevation="77" lastPageId="3" lastPageNumber="4" latitude="-25.138334" location="Fort Dauphin" longLatPrecision="124" longitude="47.15167" pageId="2" pageNumber="3" specimenCode="SAIAB 205733" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1">
<emphasis bold="true" box="[1285,1444,1334,1357]" pageId="2" pageNumber="3">
<specimenCode box="[1285,1444,1334,1357]" pageId="2" pageNumber="3">
SAIAB 
<quantity box="[1369,1444,1334,1357]" metricMagnitude="5" metricUnit="m" metricValue="2.05733191" pageId="2" pageNumber="3" unit="mm" value="2.05733191E8">205733</quantity>
</specimenCode>
</emphasis>
, 191 mm TL (after preservation), 
<specimenCount box="[469,548,691,714]" pageId="3" pageNumber="4" type="juvenile">juvenile</specimenCount>
<specimenCount box="[555,606,691,714]" pageId="3" pageNumber="4" type="male">male</specimenCount>
, Secteur de 
<location LSID="urn:lsid:plazi:treatment:03C47D45C749E724A0CACA2BFBB6B8BC:8EB29A88C748E72EA29BCBE1FF72B993" country="Madagascar" latitude="-25.138334" longLatPrecision="124" longitude="47.15167" name="Fort Dauphin" pageId="3" pageNumber="4">Fort Dauphin</location>
, southeastern 
<collectingCountry box="[364,486,724,747]" name="Madagascar" pageId="3" pageNumber="4">Madagascar</collectingCountry>
, 
<geoCoordinate box="[495,596,723,747]" degrees="25" direction="south" minutes="8.3" orientation="latitude" pageId="3" pageNumber="4" precision="92" value="-25.138334">25° 8.3′ S</geoCoordinate>
, 
<geoCoordinate box="[606,709,723,747]" degrees="47" direction="east" minutes="9.1" orientation="longitude" pageId="3" pageNumber="4" precision="92" value="47.15167">47° 9.1′ E</geoCoordinate>
, 
<quantity box="[719,770,724,747]" metricMagnitude="1" metricUnit="m" metricValue="7.7" pageId="3" pageNumber="4" unit="m" value="77.0">
<elevation box="[719,770,724,747]" metricMagnitude="1" metricUnit="m" metricValue="7.7" pageId="3" pageNumber="4" unit="m" value="77.0">77 m</elevation>
</quantity>
depth, FV 
<emphasis box="[258,404,757,780]" italics="true" pageId="3" pageNumber="4">
<collectionCode box="[258,323,757,780]" pageId="3" pageNumber="4">NOSY</collectionCode>
BE 11
</emphasis>
, station CP 3571, beam trawl, 
<date pageId="3" pageNumber="4" value="2010-05-08">
<collectingDate pageId="3" pageNumber="4" value="2010-05-08">8 May 2010</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3427235312" collectingDate="2008-09-08" collectionCode="SAIAB, ACEP" collectorName="J. Escobar" country="Madagascar" elevation="69" latitude="-25.052668" location="Madagascar" longLatPrecision="12" longitude="47.0735" pageId="3" pageNumber="4" specimenCode="SAIAB 192245, ACEP08" specimenCount="3" specimenCount-juvenile="1" specimenCount-male="1">
<emphasis bold="true" box="[258,668,790,813]" pageId="3" pageNumber="4">
CAS-ICH 247251 (
<specimenCount box="[465,490,790,813]" pageId="3" pageNumber="4" type="generic">ex</specimenCount>
<specimenCode box="[499,661,790,813]" pageId="3" pageNumber="4">SAIAB 192245</specimenCode>
)
</emphasis>
, 
<quantity box="[684,771,790,813]" metricMagnitude="-1" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="4" unit="mm" value="340.0">340 mm</quantity>
TL (fresh), 
<quantity box="[254,341,823,846]" metricMagnitude="-1" metricUnit="m" metricValue="3.25" pageId="3" pageNumber="4" unit="mm" value="325.0">325 mm</quantity>
TL (after preservation), 
<specimenCount box="[592,671,823,846]" pageId="3" pageNumber="4" type="juvenile">juvenile</specimenCount>
<specimenCount box="[679,730,823,846]" pageId="3" pageNumber="4" type="male">male</specimenCount>
, off southeastern 
<collectingCountry box="[269,392,857,880]" name="Madagascar" pageId="3" pageNumber="4">Madagascar</collectingCountry>
, 
<geoCoordinate box="[405,524,856,880]" degrees="25" direction="south" minutes="3.16" orientation="latitude" pageId="3" pageNumber="4" precision="9" value="-25.052668">25° 3.16′ S</geoCoordinate>
, 
<geoCoordinate box="[538,659,856,880]" degrees="47" direction="east" minutes="4.41" orientation="longitude" pageId="3" pageNumber="4" precision="9" value="47.0735">47° 4.41′ E</geoCoordinate>
, 
<specimenCode box="[672,765,857,880]" pageId="3" pageNumber="4">ACEP08</specimenCode>
- 352/ASCLME survey, trawl 13, collector 
<collectorName box="[566,673,890,913]" pageId="3" pageNumber="4">J. Escobar</collectorName>
, depth of capture 
<quantity box="[217,272,923,946]" metricMagnitude="1" metricUnit="m" metricValue="6.9" pageId="3" pageNumber="4" unit="m" value="69.0">
<elevation box="[217,272,923,946]" metricMagnitude="1" metricUnit="m" metricValue="6.9" pageId="3" pageNumber="4" unit="m" value="69.0">69 m</elevation>
</quantity>
, 
<date box="[285,473,923,946]" pageId="3" pageNumber="4" value="2008-09-08">
<collectingDate box="[285,473,923,946]" pageId="3" pageNumber="4" value="2008-09-08">8 September 2008</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3427235330" collectionCode="MNHN" collectorName="G. Petit" country="Madagascar" location="Toliara" pageId="3" pageNumber="4" specimenCode="MNHN 1929" specimenCount="2" specimenCount-female="1" specimenCount-juvenile="1" stateProvince="Toliara">
<emphasis bold="true" box="[485,669,923,946]" pageId="3" pageNumber="4">
<specimenCode box="[485,625,923,946]" pageId="3" pageNumber="4">
MNHN 
<quantity box="[575,625,923,946]" metricMagnitude="2" metricUnit="m" metricValue="1.12766" metricValueMax="2.23603" metricValueMin="0.019289999999999998" pageId="3" pageNumber="4" unit="mm" value="112766.0" valueMax="223603.0" valueMin="1929.0">1929</quantity>
</specimenCode>
-223
</emphasis>
, 603 mm TL, 
<specimenCount box="[184,266,956,979]" pageId="3" pageNumber="4" type="juvenile">juvenile</specimenCount>
<specimenCount box="[276,348,956,979]" pageId="3" pageNumber="4" type="female">female</specimenCount>
, collected between Tuléar (
<collectingRegion box="[641,717,956,979]" country="Madagascar" name="Toliara" pageId="3" pageNumber="4">Toliara</collectingRegion>
) and Ankilibe, southwestern 
<collectingCountry box="[374,496,989,1012]" name="Madagascar" pageId="3" pageNumber="4">Madagascar</collectingCountry>
, collector 
<collectorName box="[599,679,989,1012]" pageId="3" pageNumber="4">G. Petit</collectorName>
;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3427235307" collectionCode="MNHN" country="Madagascar" location="Toliara" pageId="3" pageNumber="4" specimenCode="MNHN 1985" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1" stateProvince="Toliara">
<emphasis bold="true" pageId="3" pageNumber="4">
<specimenCode pageId="3" pageNumber="4">
MNHN 
<quantity box="[136,188,1023,1046]" metricMagnitude="2" metricUnit="m" metricValue="1.166515" metricValueMax="2.31318" metricValueMin="0.01985" pageId="3" pageNumber="4" unit="mm" value="116651.5" valueMax="231318.0" valueMin="1985.0">1985</quantity>
</specimenCode>
-231
</emphasis>
, 318 mm TL, 
<specimenCount box="[395,476,1023,1046]" pageId="3" pageNumber="4" type="juvenile">juvenile</specimenCount>
<specimenCount box="[485,537,1023,1046]" pageId="3" pageNumber="4" type="male">male</specimenCount>
, 
<collectingRegion box="[550,625,1023,1046]" country="Madagascar" name="Toliara" pageId="3" pageNumber="4">Toliara</collectingRegion>
, southwestern 
<collectingCountry box="[136,261,1056,1079]" name="Madagascar" pageId="3" pageNumber="4">Madagascar</collectingCountry>
;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3427235309" collectionCode="MNHN" country="Madagascar" location="Fort Dauphin" pageId="3" pageNumber="4" specimenCode="MNHN 1989" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1">
<emphasis bold="true" box="[275,463,1056,1079]" pageId="3" pageNumber="4">
<specimenCode box="[275,418,1056,1079]" pageId="3" pageNumber="4">
MNHN 
<quantity box="[366,418,1056,1079]" metricMagnitude="2" metricUnit="m" metricValue="2.777275" metricValueMax="5.53466" metricValueMin="0.01989" pageId="3" pageNumber="4" unit="mm" value="277727.5" valueMax="553466.0" valueMin="1989.0">1989</quantity>
</specimenCode>
-553
</emphasis>
, 466 mm TL, 
<specimenCount box="[625,706,1056,1079]" pageId="3" pageNumber="4" type="juvenile">juvenile</specimenCount>
<specimenCount box="[716,768,1056,1079]" pageId="3" pageNumber="4" type="male">male</specimenCount>
, 
<location LSID="urn:lsid:plazi:treatment:03C47D45C749E724A0CACA2BFBB6B8BC:8EB29A88C748E72EA0CACD13FE86BF20" box="[136,277,1089,1112]" country="Madagascar" name="Fort Dauphin" pageId="3" pageNumber="4">Fort Dauphin</location>
(Tôlanaro), southeastern 
<collectingCountry box="[548,675,1089,1112]" name="Madagascar" pageId="3" pageNumber="4">Madagascar</collectingCountry>
;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3427235311" collectingDate="2000-12-11" collectionCode="MAD" collectorName="P. Doukakis &amp; Mananjo" country="Madagascar" latitude="-16.0765" location="Ambodipaka" longLatPrecision="889" longitude="49.616665" municipality="Antongil Bay" pageId="3" pageNumber="4" specimenCode="AMNH 232484" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1">
<emphasis bold="true" pageId="3" pageNumber="4">
<specimenCode pageId="3" pageNumber="4">
AMNH 
<quantity box="[136,211,1122,1145]" metricMagnitude="5" metricUnit="m" metricValue="2.32484265" pageId="3" pageNumber="4" unit="mm" value="2.32484265E8">232484</quantity>
</specimenCode>
</emphasis>
, 265 mm TL, 
<specimenCount box="[367,446,1122,1145]" pageId="3" pageNumber="4" type="juvenile">juvenile</specimenCount>
<specimenCount box="[455,506,1122,1145]" pageId="3" pageNumber="4" type="male">male</specimenCount>
, field number PD-00-XII- 11 
<collectionCode box="[176,243,1155,1178]" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="3" pageNumber="4">MAD</collectionCode>
, off northeastern 
<collectingCountry box="[454,592,1155,1178]" name="Madagascar" pageId="3" pageNumber="4">Madagascar</collectingCountry>
, 
<collectingMunicipality box="[608,766,1155,1178]" pageId="3" pageNumber="4">Antongil Bay</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:03C47D45C749E724A0CACA2BFBB6B8BC:8EB29A88C748E72EA0CACDF7FE86BFC4" box="[136,277,1189,1212]" country="Madagascar" latitude="-16.0765" longLatPrecision="889" longitude="49.616665" municipality="Antongil Bay" name="Ambodipaka" pageId="3" pageNumber="4">Ambodipaka</location>
, 
<geoCoordinate box="[292,420,1188,1212]" degrees="16" direction="south" minutes="4.59" orientation="latitude" pageId="3" pageNumber="4" precision="9" value="-16.0765">16° 4.59′ S</geoCoordinate>
, 
<geoCoordinate box="[437,547,1188,1212]" degrees="49" direction="east" minutes="37" orientation="longitude" pageId="3" pageNumber="4" precision="925" value="49.616665">49° 37′ E</geoCoordinate>
, local fishermen, 
<collectorName box="[750,1046,698,1212]" pageId="3" pageNumber="4">P. Doukakis &amp; Mananjo</collectorName>
, 
<date box="[1060,1263,698,721]" pageId="3" pageNumber="4" value="2000-12-11">
<collectingDate box="[1060,1263,698,721]" pageId="3" pageNumber="4" value="2000-12-11">11 December 2000</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3427235323" collectingDate="2001-03-27" collectionCode="MAD" collectorName="P. Doukakis" country="Madagascar" latitude="-15.643167" location="Rantohely" longLatPrecision="891" longitude="49.65" municipality="Antongil Bay" pageId="3" pageNumber="4" specimenCode="AMNH 232511" specimenCount="2" specimenCount-female="1" specimenCount-juvenile="1">
<emphasis bold="true" box="[1277,1445,698,721]" pageId="3" pageNumber="4">
<specimenCode box="[1277,1445,698,721]" pageId="3" pageNumber="4">
AMNH 
<quantity box="[1368,1445,698,721]" metricMagnitude="5" metricUnit="m" metricValue="2.32511361" pageId="3" pageNumber="4" unit="mm" value="2.32511361E8">232511</quantity>
</specimenCode>
</emphasis>
, 361 mm TL, 
<specimenCount box="[970,1056,731,754]" pageId="3" pageNumber="4" type="juvenile">juvenile</specimenCount>
<specimenCount box="[1068,1143,731,754]" pageId="3" pageNumber="4" type="female">female</specimenCount>
, field number PD-01-III-27 
<collectionCode box="[816,878,765,788]" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="3" pageNumber="4">MAD</collectionCode>
, off northeastern 
<collectingCountry box="[1057,1179,765,788]" name="Madagascar" pageId="3" pageNumber="4">Madagascar</collectingCountry>
, 
<collectingMunicipality box="[1192,1331,765,788]" pageId="3" pageNumber="4">Antongil Bay</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:03C47D45C749E724A0CACA2BFBB6B8BC:8EB29A88C748E72EA57DCBAFFA3BB86C" box="[1343,1448,765,788]" country="Madagascar" latitude="-15.643167" longLatPrecision="891" longitude="49.65" municipality="Antongil Bay" name="Rantohely" pageId="3" pageNumber="4">Rantohely</location>
, 
<geoCoordinate box="[816,946,797,821]" degrees="15" direction="south" minutes="38.59" orientation="latitude" pageId="3" pageNumber="4" precision="9" value="-15.643167">15°38.59′ S</geoCoordinate>
, 
<geoCoordinate box="[963,1072,797,821]" degrees="49" direction="east" minutes="39" orientation="longitude" pageId="3" pageNumber="4" precision="925" value="49.65">49° 39′ E</geoCoordinate>
, local fishermen, 
<collectorName box="[1273,1410,798,821]" pageId="3" pageNumber="4">P. Doukakis</collectorName>
, 
<date pageId="3" pageNumber="4" value="2001-03-27">
<collectingDate pageId="3" pageNumber="4" value="2001-03-27">27 March 2001</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3427235317" collectingDate="1955-03" collectionCode="NOSY" collectorName="J. Millot" country="Madagascar" location="Madagascar" pageId="3" pageNumber="4" specimenCode="USNM 171058" specimenCount="1" specimenCount-male="1">
<emphasis bold="true" box="[963,1128,831,854]" pageId="3" pageNumber="4">
<specimenCode box="[963,1128,831,854]" pageId="3" pageNumber="4">USNM 171058</specimenCode>
</emphasis>
, ~ 
<quantity box="[1158,1248,831,854]" metricMagnitude="-1" metricUnit="m" metricValue="1.77" pageId="3" pageNumber="4" unit="mm" value="177.0">177 mm</quantity>
TL, 
<specimenCount box="[1306,1356,831,854]" pageId="3" pageNumber="4" type="male">male</specimenCount>
embryo, 
<collectingCountry box="[816,938,864,887]" name="Madagascar" pageId="3" pageNumber="4">Madagascar</collectingCountry>
, FV 
<emphasis box="[988,1088,864,887]" italics="true" pageId="3" pageNumber="4">
<collectionCode box="[988,1050,864,887]" pageId="3" pageNumber="4">NOSY</collectionCode>
BE
</emphasis>
, collector 
<collectorName box="[1192,1279,864,887]" pageId="3" pageNumber="4">J. Millot</collectorName>
, 
<date box="[1289,1414,864,887]" pageId="3" pageNumber="4" value="1955-03">
<collectingDate box="[1289,1414,864,887]" pageId="3" pageNumber="4" value="1955-03">March 1955</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="diagnosis">
<paragraph blockId="3.[816,917,945,969]" box="[816,917,945,969]" pageId="3" pageNumber="4">
<heading bold="true" box="[816,917,945,969]" fontSize="10" level="3" pageId="3" pageNumber="4" reason="2">
<emphasis bold="true" box="[816,917,945,969]" pageId="3" pageNumber="4">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="3.[816,1452,1012,1400]" lastBlockId="4.[136,771,864,1020]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Alarge 
<taxonomicName box="[898,1132,1012,1035]" pageId="3" pageNumber="4">
<emphasis box="[898,1053,1012,1035]" italics="true" pageId="3" pageNumber="4">Acroteriobatus</emphasis>
species
</taxonomicName>
distinguished by the following combination of characters: dorsal surface smooth, without prominent thorns or tubercles, except for enlarged granular denticles partially around orbital rims and irregularly distributed along midline from nape to somewhat anterior to first dorsal-fin origin; absent between dorsal fins and upper caudal fin. Snout semi-translucent with elongated bluish-gray spots giving “stripe-nosed” appearance, numerous small bluishgray spots covering disc and pectoral, pelvic, dorsal, and caudal fins on a brown background with larger, slightly darker, indistinct brown spots covering the tail and fin bases; outer edges of pelvic-fin margins white, lateral tail folds striped orange and white; ventral surface white, except for a V-shape patternof faint to dark speckledblack spots onsnout tip. Nasal lamellae 42–48; upper jaw tooth row count ~80–87; 179–185 post-synarcual centra; 195–200 total vertebral segments; 64– 68 total pectoral skeleton radials.
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="10" pageId="4" pageNumber="5" type="description">
<paragraph blockId="4.[136,424,1081,1105]" box="[136,424,1081,1105]" pageId="4" pageNumber="5">
<heading bold="true" box="[136,424,1081,1105]" fontSize="10" level="3" pageId="4" pageNumber="5" reason="2">
<emphasis bold="true" box="[136,424,1081,1105]" pageId="4" pageNumber="5">
Description of the 
<typeStatus box="[331,424,1081,1105]" pageId="4" pageNumber="5">holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="4.[136,771,1148,1370]" pageId="4" pageNumber="5">
Values of the 
<specimenCount box="[283,441,1148,1171]" pageId="4" pageNumber="5" type="generic" typeStatus="paratypes">eight paratypes</specimenCount>
at SAIAB and CAS, as well as the two larger 
<typeStatus box="[284,383,1181,1204]" pageId="4" pageNumber="5">paratypes</typeStatus>
at MNHN in parentheses, more complex differences are described separately. Where relevant, ratios are based on horizontal measurements unless otherwise stated. Detailed morphometric measurements and meristics of the 
<typeStatus box="[173,266,1314,1337]" pageId="4" pageNumber="5">holotype</typeStatus>
, 
<specimenCount box="[276,432,1314,1337]" pageId="4" pageNumber="5" type="generic" typeStatus="paratypes">eight paratypes</specimenCount>
at SAIAB and CAS, and the two larger 
<typeStatus box="[203,302,1347,1370]" pageId="4" pageNumber="5">paratypes</typeStatus>
at MNHN are given in 
<tableCitation box="[551,630,1347,1370]" captionStart="Table 1" captionStartId="7.[136,184,155,175]" captionTargetBox="[136,1452,205,1897]" captionText="Table 1 Acroteriobatus andysabini sp. nov., morphometrics and meristics Acroteriobatus Acroteriobatus Minimum Maximum Mean SD andysabini sp. andysabini sp. (n = 9) (n = 9) (n = 11) nov., juvenile nov., juvenile male holotype, female paratype, SAIAB 97396 SAIAB 189815" pageId="4" pageNumber="5">Table 1</tableCitation>
.
</paragraph>
<paragraph blockId="4.[816,1452,1620,1909]" lastBlockId="5.[136,771,724,1278]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<emphasis bold="true" box="[816,1398,1620,1644]" pageId="4" pageNumber="5">
External morphology (
<figureCitation box="[1047,1114,1620,1644]" captionStart="Fig" captionStartId="2.[136,165,1835,1855]" captionTargetBox="[137,770,1616,1817]" captionTargetId="figure-507@2.[136,771,1615,1818]" captionTargetPageId="2" captionText="Fig. 1 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 565 mm TL fresh, in total dorsal view taken directly after catching. Photograph by Elaine Heemstra, NRF-SAIAB" figureDoi="http://doi.org/10.5281/zenodo.5825449" httpUri="https://zenodo.org/record/5825449/files/figure.png" pageId="4" pageNumber="5">Figs. 1</figureCitation>
, 
<figureCitation box="[1125,1138,1620,1644]" captionStart="Fig" captionStartId="2.[816,845,585,605]" captionTargetBox="[819,1448,146,568]" captionTargetId="figure-536@2.[818,1449,144,569]" captionTargetPageId="2" captionText="Fig. 2 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, in total dorsal view. Scale bar: 5 cm. Photograph courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825447" httpUri="https://zenodo.org/record/5825447/files/figure.png" pageId="4" pageNumber="5">2</figureCitation>
, 
<figureCitation box="[1149,1162,1620,1644]" captionStart="Fig" captionStartId="2.[816,845,1835,1855]" captionTargetBox="[827,1448,1400,1818]" captionTargetId="figure-564@2.[818,1449,1399,1819]" captionTargetPageId="2" captionText="Fig. 3 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, in total ventral view. Scale bar: 5 cm. Photograph courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825451" httpUri="https://zenodo.org/record/5825451/files/figure.png" pageId="4" pageNumber="5">3</figureCitation>
, 
<figureCitation box="[1174,1187,1620,1644]" captionStart="Fig" captionStartId="3.[136,165,554,574]" captionTargetBox="[136,789,126,543]" captionTargetId="figure-318@3.[138,769,144,538]" captionTargetPageId="3" captionText="Fig. 4 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 565 mm TL fresh, head in dorsal view taken directly after catching. Photograph by Elaine Heemstra, NRF-SAIAB" figureDoi="http://doi.org/10.5281/zenodo.5825453" httpUri="https://zenodo.org/record/5825453/files/figure.png" pageId="4" pageNumber="5">4</figureCitation>
, 
<figureCitation box="[1198,1211,1620,1644]" captionStart="Fig" captionStartId="3.[136,165,1835,1855]" captionTargetBox="[139,768,1268,1818]" captionTargetId="figure-347@3.[138,769,1267,1819]" captionTargetPageId="3" captionText="Fig. 5 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, head in dorsal view. Photograph courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825457" httpUri="https://zenodo.org/record/5825457/files/figure.png" pageId="4" pageNumber="5">5</figureCitation>
, 
<figureCitation box="[1222,1235,1620,1644]" captionStart="Fig" captionStartId="3.[816,845,581,601]" captionTargetBox="[807,1451,123,566]" captionTargetId="figure-372@3.[818,1449,144,565]" captionText="Fig. 6 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, close-up of orbital and spiracular region. Photograph courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825455" httpUri="https://zenodo.org/record/5825455/files/figure.png" pageId="4" pageNumber="5">6</figureCitation>
, 
<figureCitation box="[1247,1260,1620,1644]" captionStart="Fig" captionStartId="3.[816,845,1835,1855]" captionTargetBox="[819,1448,1441,1817]" captionTargetId="figure-398@3.[818,1449,1440,1818]" captionTargetPageId="3" captionText="Fig. 7 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 565 mm TL fresh, head in ventral view taken directly after catching. Photograph by Elaine Heemstra, NRF-SAIAB" figureDoi="http://doi.org/10.5281/zenodo.5825459" httpUri="https://zenodo.org/record/5825459/files/figure.png" pageId="4" pageNumber="5">7</figureCitation>
, 
<figureCitation box="[1271,1284,1620,1644]" captionStart="Fig" captionStartId="4.[136,165,747,767]" captionTargetBox="[139,768,145,729]" captionTargetId="figure-222@4.[138,769,144,730]" captionTargetPageId="4" captionText="Fig. 8 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, head in ventral view. Photograph courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825461" httpUri="https://zenodo.org/record/5825461/files/figure.png" pageId="4" pageNumber="5">8</figureCitation>
, 
<figureCitation box="[1296,1309,1620,1644]" captionStart="Fig" captionStartId="4.[136,165,1835,1855]" captionTargetBox="[139,768,1399,1818]" captionTargetId="figure-246@4.[138,769,1398,1819]" captionTargetPageId="4" captionText="Fig. 9 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, close-up of mouth-nasal region. Photograph courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825466" httpUri="https://zenodo.org/record/5825466/files/figure.png" pageId="4" pageNumber="5">9</figureCitation>
, and 10)
</emphasis>
Disc wedge-shaped, narrowly angular anteriorly, snout angle in front of eyes 79.1° (76–84.6°); anteriormargin undulate, outer pectoral margin broadly rounded, posterior margin convex, rear tip broadly rounded; disc length 1.22 (1.13–1.24) times its width. Pelvic fins elongate, short-based, base length 1.82 (0.82–1.76) times inner margin length; pelvic-fin total length 1.55 (1.43–2.13) times base length, 2.29 (1.95–3.96) times width; anterior margin weakly concave anteriorly, becoming convex posterior to constriction at the level of first propterygials, turning to weakly concave posterior to the level of posterior margin of spiracles and again becoming weakly convex towards broadly rounded apex, posterior margin weakly convex, free rear tip narrowly rounded. Tail elongate, becoming slender, tapering weakly to caudal; depressed, in cross section rounded above, nearly flat below; tail length from anterior cloaca 1.47 (1.29–1.81) times precloacal length, 1.43 (1.25–1.64) timesdisc length, and 5.95 (4.28–6.42) times body width at pelvic-fin insertions; body width 2.2 (2.25– 3.22) times depth at pelvic-fin insertions, 2.41 (2.5–3.17) at first dorsal-fin origin, 2.23 (1.58–2.5) at second dorsal-fin origin. Dermal fold lateral along tail, its origin just posterior to pelvic-fin insertion, extending to just behind the ventral origin of caudal-fin; fold well developed, maximum width at interdorsal space slightly more than the width of posterior nasal flap (
<figureCitation box="[250,322,1255,1278]" captionStart="Fig" captionStartId="2.[136,165,1835,1855]" captionTargetBox="[137,770,1616,1817]" captionTargetId="figure-507@2.[136,771,1615,1818]" captionTargetPageId="2" captionText="Fig. 1 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 565 mm TL fresh, in total dorsal view taken directly after catching. Photograph by Elaine Heemstra, NRF-SAIAB" figureDoi="http://doi.org/10.5281/zenodo.5825449" httpUri="https://zenodo.org/record/5825449/files/figure.png" pageId="5" pageNumber="6">Figs. 1</figureCitation>
, 
<figureCitation box="[336,349,1255,1278]" captionStart="Fig" captionStartId="2.[816,845,585,605]" captionTargetBox="[819,1448,146,568]" captionTargetId="figure-536@2.[818,1449,144,569]" captionTargetPageId="2" captionText="Fig. 2 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, in total dorsal view. Scale bar: 5 cm. Photograph courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825447" httpUri="https://zenodo.org/record/5825447/files/figure.png" pageId="5" pageNumber="6">2</figureCitation>
, and 3).
</paragraph>
<paragraph blockId="5.[816,1452,159,1909]" pageId="5" pageNumber="6">
Head moderately elongate, ventral length 25.3 (25.2– 29.8)% TL, dorsal length 24.2 (25.7–33.5)% TL; snout rather short, bluntly pointed, preoral length 2.44 (2.00–2.62) times mouth width, 4.88 (3.75–4.89) times internarial distance, 1.15 (1.05–2.27) times dorsal caudal-fin margin, 2.86 (2.27–2.9) times distance from nostril to margin of disc; direct preorbital snout length 2.38 (1.92–2.39) times interspiracular distance, 4.06 (2.9–4.09) times orbit diameter, 3.63 (3.09–3.63) times skeletal interorbital space; interorbital space weakly concave, relatively broad; eyes moderately large, slightly elevated, but not protruding, orbit diameter 1.7 (0.92–2.5) times spiracle length, 0.89 (0.85–1.13) times skeletal interorbital distance. Spiracles relatively large, narrowly bean-shaped; two prominent and strongly compressed folds on upper posterior margin, length of inner spiracular fold 0.7 (0.5–0.7) times that of outer fold; distance between bases of folds 0.5 (0.3–0.7) times length of shortest fold (
<figureCitation box="[1060,1131,691,714]" captionStart="Fig" captionStartId="3.[136,165,554,574]" captionTargetBox="[136,789,126,543]" captionTargetId="figure-318@3.[138,769,144,538]" captionTargetPageId="3" captionText="Fig. 4 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 565 mm TL fresh, head in dorsal view taken directly after catching. Photograph by Elaine Heemstra, NRF-SAIAB" figureDoi="http://doi.org/10.5281/zenodo.5825453" httpUri="https://zenodo.org/record/5825453/files/figure.png" pageId="5" pageNumber="6">Figs. 4</figureCitation>
, 
<figureCitation box="[1145,1158,691,714]" captionStart="Fig" captionStartId="3.[136,165,1835,1855]" captionTargetBox="[139,768,1268,1818]" captionTargetId="figure-347@3.[138,769,1267,1819]" captionTargetPageId="3" captionText="Fig. 5 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, head in dorsal view. Photograph courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825457" httpUri="https://zenodo.org/record/5825457/files/figure.png" pageId="5" pageNumber="6">5</figureCitation>
, and 6).
</paragraph>
<paragraph blockId="5.[816,1452,159,1909]" pageId="5" pageNumber="6">
Nostril moderate large, oblique, all nasal flaps well developed; anterior aperture subcircular; nostril length 3.14 (2.6– 3.83) times anterior aperture width, 1.22 (0.95–1.6) times anterior nasal-flap base length, 0.76 (0.53–0.77) times distance from nostril to edge of disc, 1.29 (1.0–1.28) times internarial distance. Anterior nasal-flap base greatly extended, penetrating horizontally into internarial space, mesial edges barely separated (by about width of posterior nasal flap); innermost extension of flap not narrow; flap base length 1.80 (1.6–2.6) times its width at process, 2.57 (1.67–3.28) times anterior aperture width; minimum distance between mesial insertions of flaps 7.29 (4.8–13.0) in greatest distance across nostrils anteriorly, 2.43 (1.6–4.5) in minimum internarial distance; process of flap two or more times as long as wide, narrowing distally to blunt pointed tip, and adjoining with posterolateral nasal flap and anterior aperture posterior margin. Posterolateral nasal flap well developed, lobe-like, broadest medially, length 3.0 (1.22–3.5) times its width; origin slightly posterior to lateral extremity of anterior nasal aperture, with free fold posteriorly extending nearly to the level of anterior nasal flap insertion. Posterior nasal flap well developed, lobelike, base length 1.86 (1.67–3.0) times its width, its inner edge nearly reaching innermost margin of nostril; width 1.0 (1.0– 1.67) times anterior aperture width, 1.4 (1.0–2.0) times posterolateral nasal-flap width (
<figureCitation box="[1108,1180,1521,1544]" captionStart="Fig" captionStartId="3.[816,845,1835,1855]" captionTargetBox="[819,1448,1441,1817]" captionTargetId="figure-398@3.[818,1449,1440,1818]" captionTargetPageId="3" captionText="Fig. 7 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 565 mm TL fresh, head in ventral view taken directly after catching. Photograph by Elaine Heemstra, NRF-SAIAB" figureDoi="http://doi.org/10.5281/zenodo.5825459" httpUri="https://zenodo.org/record/5825459/files/figure.png" pageId="5" pageNumber="6">Figs. 7</figureCitation>
, 
<figureCitation box="[1193,1206,1521,1544]" captionStart="Fig" captionStartId="4.[136,165,747,767]" captionTargetBox="[139,768,145,729]" captionTargetId="figure-222@4.[138,769,144,730]" captionTargetPageId="4" captionText="Fig. 8 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, head in ventral view. Photograph courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825461" httpUri="https://zenodo.org/record/5825461/files/figure.png" pageId="5" pageNumber="6">8</figureCitation>
, and 9).
</paragraph>
<paragraph blockId="5.[816,1452,159,1909]" lastBlockId="6.[136,772,911,1897]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
Mouth width 1.55 (1.44–2.0) times nostril length, 6.56 (5.13–6.71) in precloacal length; positioned beneath posterior margin of orbit. Upper jaw slightly convex, upper lip broadly arched; lower lip not pronounced, not separated from oral groove by ridges of strongly corrugated skin; corners of mouth with several strong, short lateral grooves. Teeth arranged in quincunx, small, close-set, rhombic, broad-based with short, bluntly rounded cusp; upper and lower teeth similar in shape and size; ~84 (~80–87) tooth rows in upper and ~79 (~80–87) in lower jaws. Gill openings strongly wavy; length of third gill slit 2.75 (1.93–3.0) in nostril length, 5.5 (4.75–6.67) in distance between fifth gill slits; distance between first gill slits 1.48 (1.3–1.58) times distance between fifth gill slits; distance between fifth gill slits 2.59 (2.22– 2.73) times internarial distance, 1.29 (1.12–1.54) times mouth width, 0.32 (0.29–0.37) of ventral head length (
<figureCitation box="[631,693,1044,1067]" captionStart="Fig" captionStartId="4.[136,165,1835,1855]" captionTargetBox="[139,768,1399,1818]" captionTargetId="figure-246@4.[138,769,1398,1819]" captionTargetPageId="4" captionText="Fig. 9 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, close-up of mouth-nasal region. Photograph courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825466" httpUri="https://zenodo.org/record/5825466/files/figure.png" pageId="6" pageNumber="7">Fig. 9</figureCitation>
).
</paragraph>
<paragraph blockId="6.[136,772,911,1897]" pageId="6" pageNumber="7">
Dorsal fins erect, relatively tall, dissimilar in size and shape; first dorsal-fin anterior margin weakly convex, curving rearwards towards rounded apex; posterior margin weakly convex near tip, then becoming weakly concave to nearly straight; second dorsal-fin anterior margin weakly convex, curving rearwards towards pointed apex; posterior margin nearly straight; free rear tips rounded of both dorsal fins, almost forming right angle, not produced; first dorsal fin height subequal to second, length of first 0.93 (0.92–1.44) times height, its base length 2.15 (1.17–2.5) times inner margin length; second dorsal-fin length 1.0 (0.93–1.42) times its height, base length 1.47 (1.34–2.21) times inner margin length (
<figureCitation box="[144,254,1476,1499]" captionStart="Fig" captionStartId="4.[816,845,1425,1445]" captionTargetBox="[819,1448,145,1337]" captionTargetId="figure-272@4.[818,1449,144,1409]" captionTargetPageId="4" captionText="Fig. 10 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL; first dorsal (a), second dorsal (b), and caudal (c) fins in lateral views (c image reversed). Photographs courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825463" httpUri="https://zenodo.org/record/5825463/files/figure.png" pageId="6" pageNumber="7">Fig. 10a, b</figureCitation>
). First dorsal-fin origin well posterior to pelvic-fin free rear tips, interspace 1.29 (1.0–1.15) times interdorsal distance; interdorsal space relatively short, 1.48 (1.16–2.09) times second dorsal-fin height, 2.11 (1.88–2.88) times base of first dorsal fin, 1.11 (0.71–1.25) times tail width at origin of first dorsal fin, 1.59 (1.11–1.9) times interspace between second dorsal-fin insertion and upper origin of caudal fin. Caudal fin relatively small; dorsal caudal margin weakly convex, length 1.6 (1.53–2.08) times preventral margin length; ventral lobe broadly rounded; posterior margin weakly concave; fin tip acutely pointed (
<figureCitation box="[440,527,1808,1831]" captionStart="Fig" captionStartId="4.[816,845,1425,1445]" captionTargetBox="[819,1448,145,1337]" captionTargetId="figure-272@4.[818,1449,144,1409]" captionTargetPageId="4" captionText="Fig. 10 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL; first dorsal (a), second dorsal (b), and caudal (c) fins in lateral views (c image reversed). Photographs courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825463" httpUri="https://zenodo.org/record/5825463/files/figure.png" pageId="6" pageNumber="7">Fig. 10c</figureCitation>
).
</paragraph>
<paragraph blockId="6.[136,772,911,1897]" lastBlockId="6.[816,1452,911,1898]" pageId="6" pageNumber="7">
Dermal denticles mostly minute, close-set, covering entire body and fins; surfaces mostly smooth to the touch, except for a continuous row of ~15 to 20 somewhat enlarged granular and smaller denticles around the anterior and inner rim of orbits, extending to the anterior edge of inner spiracle; enlarged, coarse denticles patchily distributed along midline from nape to ~ 
<quantity box="[967,1041,1044,1067]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="7" unit="mm" value="50.0">50 mm</quantity>
anterior to first dorsal-fin origin, absent along midline between dorsal fins and upper caudal origin; enlarged denticles absent on snout tip, anterior and lateral margins of disc, pelvic fins, and bases of dorsal fins; nasal flaps and lamellae naked. Crowns of denticles mostly oval, except largest denticles with slightly erect, bluntly acute cusp, base stellate. Thorns absent. Prebranchial sensory pore patch distinct, extending to just behind the level of third gill slit. Post-scapular sensory canal rather indistinct, weakly undulated anteriorly, terminating near pectoral-fin insertions; sensory pores minute, canal not forming a shallow groove (
<figureCitation box="[1349,1410,1376,1399]" captionStart="Fig" captionStartId="2.[816,845,1835,1855]" captionTargetBox="[827,1448,1400,1818]" captionTargetId="figure-564@2.[818,1449,1399,1819]" captionTargetPageId="2" captionText="Fig. 3 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, in total ventral view. Scale bar: 5 cm. Photograph courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825451" httpUri="https://zenodo.org/record/5825451/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
).
</paragraph>
<paragraph blockId="6.[816,1452,911,1898]" lastBlockId="8.[136,772,1355,1910]" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
Rostral cartilage broad, its shaft just slightly increasing in width posteriorly from rostral node; rostral node rounded at apex, not angular, broadly expanded, relatively short, axis at maximum node width 34.1 (33.6–39.3)% of length of rostral cartilage from tip; anterior fontanelle relatively broad posteriorly, tapering gradually towards rostral node, dorsolateral edges of cartilage surrounding fontanelle (rostral ridges on surface of snout) well separated posteriorly, not constricting medially; rostral cartilage 57.5 (52.6–57.1)% of neurocranium length, ventral edges of rostral cartilage united; nasal capsules large, with their transverse axes directed anterolaterally; width of cranium across nasal capsules 1.25 (1.17–1.27) times nasobasal length (base of rostrum to occipital condyles); width of nasal capsule 1.79 (1.56–1.87) times its length; basal plate relatively broad, its minimum width 3.37 (3.15–3.75) times in nasobasal length; cranial roof with small, oval-shaped fenestra, with its anterior edge located behind precerebral cavity by a distance ~1.5 times its length; anterior cartilage triangular, narrow, posteriorly wedge-shaped, without an anterior lobe extending past nasal capsules; preorbital processes well developed; postorbital processes moderately large, not bifurcate; greatest width across processes 1.91 (1.79–1.99) times in nasobasal length (
<figureCitation box="[323,397,1587,1610]" captionStart="Fig" captionStartId="5.[136,165,607,627]" captionTargetBox="[139,768,145,589]" captionTargetId="figure-686@5.[138,769,144,590]" captionTargetPageId="5" captionText="Fig. 11 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, radiograph of cranium and snout in dorsal view. The radiograph was taken and kindly provided by Jon Fong" figureDoi="http://doi.org/10.5281/zenodo.5825468" httpUri="https://zenodo.org/record/5825468/files/figure.png" pageId="8" pageNumber="9">Fig. 11</figureCitation>
).
</paragraph>
<paragraph blockId="8.[136,772,1355,1910]" lastBlockId="8.[816,1452,1354,1411]" pageId="8" pageNumber="9">Nasal lamellae 45 (42–48). Pectoral skeleton with 32 (30– 32) propterygial, 9 (8–10) mesopterygial, 2 (1–2) neopterygial, 25 (24–25) metapterygial, amounting to 68 (64–68) total radials. Total pelvic-fin radials 25 (24–27). Total vertebral segment (synarcual and free) counts 196 (195–200); post-synarcual centra 180 (179–185); precaudal centra (excluding synarcual centra) 139 (137–144); synarcual segments 16 (14–17); monospondylous precaudal centra 39 (37–42); diplosondylous precaudal centra 100 (98–104); diplosondylous caudal centra 41 (40–42). Monospondylous to diplosondylous centra transition posterior to pelvic girdle.</paragraph>
<paragraph blockId="8.[816,925,1454,1478]" box="[816,925,1454,1478]" pageId="8" pageNumber="9">
<heading bold="true" box="[816,925,1454,1478]" fontSize="10" level="3" pageId="8" pageNumber="9" reason="2">
<emphasis bold="true" box="[816,925,1454,1478]" pageId="8" pageNumber="9">Coloration</emphasis>
</heading>
</paragraph>
<paragraph blockId="8.[816,1452,1521,1909]" lastBlockId="9.[136,772,543,997]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis box="[816,1039,1521,1544]" italics="true" pageId="8" pageNumber="9">Prior to preservation</emphasis>
: dorsal surface light to medium brown with an orangish tinge, an organized pattern of smaller bluishgray circular spots, with a darker ring encircling a lighter bluish-gray center, and indistinct larger brownish spots (
<figureCitation box="[824,894,1654,1677]" captionStart="Fig" captionStartId="2.[136,165,1835,1855]" captionTargetBox="[137,770,1616,1817]" captionTargetId="figure-507@2.[136,771,1615,1818]" captionTargetPageId="2" captionText="Fig. 1 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 565 mm TL fresh, in total dorsal view taken directly after catching. Photograph by Elaine Heemstra, NRF-SAIAB" figureDoi="http://doi.org/10.5281/zenodo.5825449" httpUri="https://zenodo.org/record/5825449/files/figure.png" pageId="8" pageNumber="9">Figs. 1</figureCitation>
and 
<figureCitation box="[942,955,1654,1677]" captionStart="Fig" captionStartId="3.[136,165,554,574]" captionTargetBox="[136,789,126,543]" captionTargetId="figure-318@3.[138,769,144,538]" captionTargetPageId="3" captionText="Fig. 4 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 565 mm TL fresh, head in dorsal view taken directly after catching. Photograph by Elaine Heemstra, NRF-SAIAB" figureDoi="http://doi.org/10.5281/zenodo.5825453" httpUri="https://zenodo.org/record/5825453/files/figure.png" pageId="8" pageNumber="9">4</figureCitation>
); smaller bluish-gray spots cover thesnout, disc, pectoral, pelvic, dorsal, and caudal fins; larger brownish spots mostly coverthetail andfin bases, but not extending onto fins; snout semi-translucent with somewhat elongated bluish-gray spots giving it a stripe-nosed appearance (
<figureCitation box="[1253,1325,1786,1809]" captionStart="Fig" captionStartId="2.[136,165,1835,1855]" captionTargetBox="[137,770,1616,1817]" captionTargetId="figure-507@2.[136,771,1615,1818]" captionTargetPageId="2" captionText="Fig. 1 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 565 mm TL fresh, in total dorsal view taken directly after catching. Photograph by Elaine Heemstra, NRF-SAIAB" figureDoi="http://doi.org/10.5281/zenodo.5825449" httpUri="https://zenodo.org/record/5825449/files/figure.png" pageId="8" pageNumber="9">Figs. 1</figureCitation>
, 
<figureCitation box="[1339,1352,1786,1809]" captionStart="Fig" captionStartId="3.[136,165,554,574]" captionTargetBox="[136,789,126,543]" captionTargetId="figure-318@3.[138,769,144,538]" captionTargetPageId="3" captionText="Fig. 4 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 565 mm TL fresh, head in dorsal view taken directly after catching. Photograph by Elaine Heemstra, NRF-SAIAB" figureDoi="http://doi.org/10.5281/zenodo.5825453" httpUri="https://zenodo.org/record/5825453/files/figure.png" pageId="8" pageNumber="9">4</figureCitation>
, and 12). Outer edges of pelvic-fin margins white, outer edges of posterior dorsal-fin margins translucent whitish, lateral tail folds striped orange and white. Ventral surface mostly white, except for faint to dark speckled black spots forming a V-shape on rostral snout tip; spots becoming darker and more numerous with growth (
<figureCitation box="[277,338,609,632]" captionStart="Fig" captionStartId="3.[816,845,1835,1855]" captionTargetBox="[819,1448,1441,1817]" captionTargetId="figure-398@3.[818,1449,1440,1818]" captionTargetPageId="3" captionText="Fig. 7 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 565 mm TL fresh, head in ventral view taken directly after catching. Photograph by Elaine Heemstra, NRF-SAIAB" figureDoi="http://doi.org/10.5281/zenodo.5825459" httpUri="https://zenodo.org/record/5825459/files/figure.png" pageId="9" pageNumber="10">Fig. 7</figureCitation>
); in small (≤ 
<quantity box="[472,559,609,632]" metricMagnitude="-1" metricUnit="m" metricValue="2.15" pageId="9" pageNumber="10" unit="mm" value="215.0">215 mm</quantity>
TL), newborn specimens short V-shaped black lines with faint dark spots at base extending from tip to about one-third snout length to midpoint of internostril space; larger individuals (&gt; 
<quantity box="[633,723,709,732]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="10" unit="mm" value="300.0">300 mm</quantity>
TL) with prominent dark to black spots, becoming more numerous and with spots extending to about the midpoint of rostrum; pectoral and pelvic fins mostly white, except fin edges brownish (
<figureCitation box="[185,251,842,865]" captionStart="Fig" captionStartId="3.[816,845,1835,1855]" captionTargetBox="[819,1448,1441,1817]" captionTargetId="figure-398@3.[818,1449,1440,1818]" captionTargetPageId="3" captionText="Fig. 7 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 565 mm TL fresh, head in ventral view taken directly after catching. Photograph by Elaine Heemstra, NRF-SAIAB" figureDoi="http://doi.org/10.5281/zenodo.5825459" httpUri="https://zenodo.org/record/5825459/files/figure.png" pageId="9" pageNumber="10">Fig. 7</figureCitation>
). 
<emphasis box="[277,474,842,865]" italics="true" pageId="9" pageNumber="10">After preservation</emphasis>
: dorsal surface similar but bluish-gray spots less pronounced and large brownish spots mostly faded and not evident (
<figureCitation box="[468,533,908,931]" captionStart="Fig" captionStartId="2.[816,845,585,605]" captionTargetBox="[819,1448,146,568]" captionTargetId="figure-536@2.[818,1449,144,569]" captionTargetPageId="2" captionText="Fig. 2 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, in total dorsal view. Scale bar: 5 cm. Photograph courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825447" httpUri="https://zenodo.org/record/5825447/files/figure.png" pageId="9" pageNumber="10">Fig. 2</figureCitation>
). Ventral surface also similar, except edges of pectoral and pelvic-fin margins fade to a lighter brown (
<figureCitation box="[337,398,974,997]" captionStart="Fig" captionStartId="2.[816,845,1835,1855]" captionTargetBox="[827,1448,1400,1818]" captionTargetId="figure-564@2.[818,1449,1399,1819]" captionTargetPageId="2" captionText="Fig. 3 Acroteriobatus andysabini sp. nov., SAIAB 97396, juvenile male holotype, 550 mm TL, in total ventral view. Scale bar: 5 cm. Photograph courtesy Marsha Englebrecht ©" figureDoi="http://doi.org/10.5281/zenodo.5825451" httpUri="https://zenodo.org/record/5825451/files/figure.png" pageId="9" pageNumber="10">Fig. 3</figureCitation>
).
</paragraph>
<paragraph blockId="9.[136,178,1041,1065]" box="[136,178,1041,1065]" pageId="9" pageNumber="10">
<heading bold="true" box="[136,178,1041,1065]" fontSize="10" level="3" pageId="9" pageNumber="10" reason="2">
<emphasis bold="true" box="[136,178,1041,1065]" pageId="9" pageNumber="10">Size</emphasis>
</heading>
</paragraph>
<paragraph blockId="9.[136,772,1108,1297]" pageId="9" pageNumber="10">
The new species is described from 
<specimenCount box="[511,654,1108,1131]" pageId="9" pageNumber="10" type="generic">15 specimens</specimenCount>
(
<specimenCount box="[669,771,1108,1131]" pageId="9" pageNumber="10" type="male">ten males</specimenCount>
and 
<specimenCount box="[179,308,1141,1164]" pageId="9" pageNumber="10" type="female">five females</specimenCount>
), ranging in size from 
<quantity box="[534,690,1141,1164]" metricMagnitude="-1" metricUnit="m" metricValue="3.9" metricValueMax="6.029999999999999" metricValueMin="1.77" pageId="9" pageNumber="10" unit="mm" value="390.0" valueMax="603.0" valueMin="177.0">177 to 603 mm</quantity>
TL. All 
<typeStatus box="[136,191,1174,1197]" pageId="9" pageNumber="10">types</typeStatus>
are juvenile, the size at birth is ~ 
<quantity box="[535,623,1174,1197]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="10" unit="mm" value="180.0">180 mm</quantity>
TL. Based on images of unretained specimens (
<figureCitation box="[505,586,1207,1230]" captionStart="Fig" captionStartId="5.[136,168,1834,1854]" captionTargetBox="[139,768,1345,1817]" captionTargetId="figure-720@5.[138,769,1344,1818]" captionTargetPageId="5" captionText="Fig. 12 Acroteriobatus andysabini sp. nov., Malagasy artisanal fisherman with large (~1000 mm TL) specimen. The photograph was taken and kindly provided by Kerry Sink" figureDoi="http://doi.org/10.5281/zenodo.5825470" httpUri="https://zenodo.org/record/5825470/files/figure.png" pageId="9" pageNumber="10">Fig. 12</figureCitation>
), 
<taxonomicName authority="Weigmann &amp; Ebert &amp; Séret, 2021" authorityName="Weigmann &amp; Ebert &amp; Séret" authorityYear="2021" class="Elasmobranchii" family="Rhinobatidae" genus="Acroteriobatus" kingdom="Animalia" order="Rhinopristiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="andysabini" status="sp. nov.">
<emphasis italics="true" pageId="9" pageNumber="10">Acroteriobatus andysabini</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[256,338,1241,1264]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
is a large species of 
<taxonomicName authorityName="Minimum Maximum Mean SD" box="[548,703,1241,1264]" class="Elasmobranchii" family="Rhinobatidae" genus="Acroteriobatus" kingdom="Animalia" order="Rhinopristiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis box="[548,703,1241,1264]" italics="true" pageId="9" pageNumber="10">Acroteriobatus</emphasis>
</taxonomicName>
growing to more than 
<quantity box="[315,415,1274,1297]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="10" unit="mm" value="1000.0">1000 mm</quantity>
TL.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="distribution">
<paragraph blockId="9.[136,260,1341,1365]" box="[136,260,1341,1365]" pageId="9" pageNumber="10">
<heading bold="true" box="[136,260,1341,1365]" fontSize="10" level="3" pageId="9" pageNumber="10" reason="2">
<emphasis bold="true" box="[136,260,1341,1365]" pageId="9" pageNumber="10">Distribution</emphasis>
</heading>
</paragraph>
<paragraph blockId="9.[136,771,1407,1563]" lastBlockId="9.[816,1453,543,732]" pageId="9" pageNumber="10">
<taxonomicName authority="Weigmann &amp; Ebert &amp; Séret, 2021" authorityName="Weigmann &amp; Ebert &amp; Séret" authorityYear="2021" box="[136,414,1407,1430]" class="Elasmobranchii" family="Rhinobatidae" genus="Acroteriobatus" kingdom="Animalia" order="Rhinopristiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="andysabini" status="sp. nov.">
<emphasis box="[136,414,1407,1430]" italics="true" pageId="9" pageNumber="10">Acroteriobatus andysabini</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[422,508,1407,1430]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
is presently only known from the coastal waters surrounding 
<collectingCountry box="[517,642,1441,1464]" name="Madagascar" pageId="9" pageNumber="10">Madagascar</collectingCountry>
and may be endemic to this region (
<figureCitation box="[416,499,1474,1497]" captionStart="Fig" captionStartId="6.[136,165,152,172]" captionTargetBox="[476,1451,147,843]" captionTargetId="figure-569@6.[475,1452,146,844]" captionTargetPageId="6" captionText="Fig. 13 Map of the southwestern Indian Ocean depicting the catch locations of the examined specimens of all three species of the Acroteriobatus leucospilus species complex. Holotype (white star) and paratypes (white circles) of A. andysabini sp. nov., holotype (gray star) and paratypes (gray star and gray circle) of A. stehmanni sp. nov., and syntypes (black triangles) and other specimens (black squares) of A. leucospilus" figureDoi="http://doi.org/10.5281/zenodo.5825472" httpUri="https://zenodo.org/record/5825472/files/figure.png" pageId="9" pageNumber="10">Fig. 13</figureCitation>
). Records are from off Tôlanaro, Anosy region, and Cape Andavaka (southeastern 
<collectingCountry box="[136,289,1540,1563]" name="Madagascar" pageId="9" pageNumber="10">Madagascar</collectingCountry>
), 
<collectingRegion box="[316,404,1540,1563]" country="Madagascar" name="Toliara" pageId="9" pageNumber="10">Toliara</collectingRegion>
and Ankilibe (southwestern 
<collectingCountry box="[816,945,543,566]" name="Madagascar" pageId="9" pageNumber="10">Madagascar</collectingCountry>
), and Antongil Bay (northeastern 
<collectingCountry box="[1312,1442,543,566]" name="Madagascar" pageId="9" pageNumber="10">Madagascar</collectingCountry>
). Its depth range is from shallow coastal waters to at least 
<specimenCount pageId="9" pageNumber="10" type="generic">
<quantity metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="10" unit="m" value="80.0">80 m</quantity>
.
</specimenCount>
<bibRefCitation author="Fricke R &amp; Mahafina J &amp; Behivoke F &amp; Jaonalison H &amp; Leopold M &amp; Ponton D" box="[848,1042,609,632]" journalOrPublisher="FishTaxa" pageId="9" pageNumber="10" pagination="1 - 432" part="3" refId="ref22596" refString="Fricke R, Mahafina J, Behivoke F, Jaonalison H, Leopold M, Ponton D (2018) Annotated checklist of the fishes of Madagascar, southwestern Indian Ocean, with 158 new records. FishTaxa 3 (1): 1 - 432" title="Annotated checklist of the fishes of Madagascar, southwestern Indian Ocean, with 158 new records" type="journal article" year="2018">Fricke et al. (2018)</bibRefCitation>
reported 
<taxonomicName baseAuthorityName="Norman" baseAuthorityYear="1926" box="[1137,1280,609,632]" class="Elasmobranchii" family="Rhinobatidae" genus="Acroteriobatus" kingdom="Animalia" order="Rhinopristiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="leucospilus">
<emphasis box="[1137,1280,609,632]" italics="true" pageId="9" pageNumber="10">A. leucospilus</emphasis>
</taxonomicName>
as anew species record for 
<collectingCountry box="[931,1056,642,665]" name="Madagascar" pageId="9" pageNumber="10">Madagascar</collectingCountry>
but commented that the species had been misidentified previously as 
<taxonomicName authorityName="Linck" authorityYear="1790" box="[1266,1409,675,698]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rhinopristiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis box="[1266,1409,675,698]" italics="true" pageId="9" pageNumber="10">Rhinobatos</emphasis>
</taxonomicName>
(= 
<taxonomicName authorityName="Minimum Maximum Mean SD" box="[816,971,709,732]" class="Elasmobranchii" family="Rhinobatidae" genus="Acroteriobatus" kingdom="Animalia" order="Rhinopristiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis box="[816,971,709,732]" italics="true" pageId="9" pageNumber="10">Acroteriobatus</emphasis>
</taxonomicName>
) 
<emphasis box="[986,1102,709,732]" italics="true" pageId="9" pageNumber="10">annuulatus</emphasis>
by several earlier authors.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="etymology">
<paragraph blockId="9.[816,927,808,832]" box="[816,927,808,832]" pageId="9" pageNumber="10">
<heading bold="true" box="[816,927,808,832]" fontSize="10" level="3" pageId="9" pageNumber="10" reason="2">
<emphasis bold="true" box="[816,927,808,832]" pageId="9" pageNumber="10">Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="9.[816,1451,875,964]" pageId="9" pageNumber="10">
The species name 
<taxonomicName authorityName="Weigmann &amp; Ebert &amp; Séret" authorityYear="2021" box="[1012,1126,875,898]" class="Elasmobranchii" family="Rhinobatidae" genus="Acroteriobatus" kingdom="Animalia" order="Rhinopristiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="andysabini">
<emphasis box="[1012,1126,875,898]" italics="true" pageId="9" pageNumber="10">andysabini</emphasis>
</taxonomicName>
is in honor of Andy Sabin for his gracious support of the Lost Sharks project at the Pacific Shark Research Center.
</paragraph>
</subSubSection>
</treatment>
</document>