317 lines
43 KiB
XML
317 lines
43 KiB
XML
<document id="E15B2F3150D22E871A883B30B3120706" ID-CLB-Dataset="302163" ID-DOI="10.11646/phytotaxa.641.4.3" ID-GBIF-Dataset="6e865334-8fff-48f7-b67f-c403995caa69" ID-ISSN="1179-3163" ID-Zenodo-Dep="13214070" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1722601437600" checkinUser="felipe" docAuthor="Karis, Per Ola & Mckenzie, Robert J." docDate="2024" docId="03910E2F5F70FFC9FF6A1E23FB41F82E" docLanguage="en" docName="phytotaxa.641.4.3.pdf" docOrigin="Phytotaxa 641 (4)" docSource="http://dx.doi.org/10.11646/phytotaxa.641.4.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Stobaea rigida Thunberg 1800" docType="treatment" docVersion="5" lastPageNumber="271" masterDocId="FFA876575F72FFCDFFE21A17FFEFFFDC" masterDocTitle="Second-step lectotypification of Stobaea rigida and reassessment of heterotypic synonyms of Berkheya rigida (Asteraceae, Arctotideae)" masterLastPageNumber="276" masterPageNumber="267" pageNumber="269" updateTime="1725380514620" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="27195295D1DBFC106F884C6E43DC42FB">
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<mods:title id="CE2AE97D97F9AA21E98A75B5E9FA961B">Second-step lectotypification of Stobaea rigida and reassessment of heterotypic synonyms of Berkheya rigida (Asteraceae, Arctotideae)</mods:title>
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</mods:titleInfo>
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<mods:name id="7EB9CC0E42B1F517DAA58A35A63B8CB9" type="personal">
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<mods:roleTerm id="1C3341D84BFCB8E300EFA7A8BB15235B">Author</mods:roleTerm>
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<mods:namePart id="03F802FF6F1D4E4B05B6DD5A9FB70A97">Karis, Per Ola</mods:namePart>
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<mods:nameIdentifier id="2FF13024F1F6E4B0D7F45C1C120815FC" type="ORCID">0000-0001-5840-7071</mods:nameIdentifier>
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<mods:affiliation id="31EB4B43017F57E7520D102FAC2C22CF">Department of Ecology, Environment and Plant Sciences, Stockholm University, SE- 106 91 Stockholm, Sweden</mods:affiliation>
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<mods:nameIdentifier id="7D58CC4AB7A8D791A0D85E04F6242614" type="email">per-ola.karis@su.se</mods:nameIdentifier>
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</mods:name>
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<mods:name id="C12F0976CBEB31184F3706DB54D16697" type="personal">
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<mods:roleTerm id="7D942D9C311A8D11AE4453169807540F">Author</mods:roleTerm>
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<mods:namePart id="E862F3E6CDD7F1CD14A1D1771B4A7028">Mckenzie, Robert J.</mods:namePart>
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<mods:nameIdentifier id="19FC697272703D7B49B6F515A93EC8E0" type="ORCID">0000-0001-7054-3308</mods:nameIdentifier>
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<mods:affiliation id="199177BD237D5C8F2A2B12A0196EB53E">Department of Plant Sciences and Afromontane Research Unit, University of the Free State, Private Bag X 13, Phuthaditjhaba 9866, South Africa</mods:affiliation>
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<mods:nameIdentifier id="10875FDA310FFB0EA618611FAA65BEB4" type="email">robt.mckenzie@gmail.com</mods:nameIdentifier>
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<mods:title id="2196A899ADFF4C601555B5002D2846CF">Phytotaxa</mods:title>
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<mods:part id="5522789BDFADF5983DBEA330ADE52D85">
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<mods:date id="89E171F87772604FE44A3DA8D3E50481">2024</mods:date>
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<mods:number id="78273921FCEC101541536C22D7BA36E9">2024-03-21</mods:number>
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<mods:number id="52F40BB5D3D9982364CB1566769D505F">641</mods:number>
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<mods:number id="80F5CFFA32CEEEED37DFB8E48DBD5DD1">4</mods:number>
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<mods:start id="6519C693C4F046AA34E7311CEF0A6327">267</mods:start>
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<mods:url id="5558E37D117BC1060BBC41C6794D5FB7">http://dx.doi.org/10.11646/phytotaxa.641.4.3</mods:url>
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<treatment id="03910E2F5F70FFC9FF6A1E23FB41F82E" ID-DOI="http://doi.org/10.5281/zenodo.13645995" ID-GBIF-Taxon="240157507" ID-Zenodo-Dep="13645995" LSID="urn:lsid:plazi:treatment:03910E2F5F70FFC9FF6A1E23FB41F82E" httpUri="http://treatment.plazi.org/id/03910E2F5F70FFC9FF6A1E23FB41F82E" lastPageId="4" lastPageNumber="271" pageId="2" pageNumber="269">
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<subSubSection id="C322ECB25F70FFCFFF6A1E23FE68FB92" box="[136,391,1076,1103]" pageId="2" pageNumber="269" type="nomenclature">
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<paragraph id="8B87BF395F70FFCFFF6A1E23FE68FB92" blockId="2.[136,391,1076,1103]" box="[136,391,1076,1103]" pageId="2" pageNumber="269">
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<heading id="D0CF08555F70FFCFFF6A1E23FE68FB92" box="[136,391,1076,1103]" fontSize="11" level="2" pageId="2" pageNumber="269" reason="3">
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<taxonomicName id="4C38C4BA5F70FFCFFF6A1E23FE68FB92" ID-CoL="52SVC" authority="Thunb." authorityName="Thunberg" authorityYear="1800" box="[136,391,1076,1103]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="269" phylum="Tracheophyta" rank="species" species="rigida">
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<emphasis id="B94C632B5F70FFCFFF6A1E23FEC0FB93" bold="true" box="[136,303,1076,1103]" italics="true" pageId="2" pageNumber="269">Stobaea rigida</emphasis>
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Thunb.
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C322ECB25F70FFC9FF6A1E6BFB41F82E" lastPageId="4" lastPageNumber="271" pageId="2" pageNumber="269" type="discussion">
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<paragraph id="8B87BF395F70FFCFFF6A1E6BFD90FADE" blockId="2.[136,1452,1148,2003]" pageId="2" pageNumber="269">
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<bibRefCitation id="EFA9C2C85F70FFCFFF6A1E6BFE6DFB4A" author="Thunberg, C. P." box="[136,386,1148,1174]" pageId="2" pageNumber="269" refId="ref6546" refString="Thunberg, C. P. (1800) Prodromus plantarum capensium: quas in promontorio bonae spei Africes, annis 1772 - 1775, collegit Carol. Pet. Thunberg 2. J. Edman, Upsaliae, 191 pp. https: // doi. org / 10.5962 / bhl. title. 84" type="book" year="1800">Thunberg (1800: 141)</bibRefCitation>
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published the name
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<taxonomicName id="4C38C4BA5F70FFCFFD8F1E6BFCFBFB4A" authorityName="Thunberg" authorityYear="1800" box="[621,788,1148,1174]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="269" phylum="Tracheophyta" rank="species" species="rigida">
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<emphasis id="B94C632B5F70FFCFFD8F1E6BFCFBFB4A" box="[621,788,1148,1174]" italics="true" pageId="2" pageNumber="269">Stobaea rigida</emphasis>
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</taxonomicName>
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accompanied only by the brief diagnostic phrase “
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<emphasis id="B94C632B5F70FFCFFAB41E6BFDA5FB66" italics="true" pageId="2" pageNumber="269">
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<taxonomicName id="4C38C4BA5F70FFCFFAB41E6BFF0CFB66" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="269" phylum="Tracheophyta" rank="subSpecies" species="foliis" subSpecies="cordatis">S. foliis cordatis</taxonomicName>
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pinnatifidis spinosis tomentosis
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</emphasis>
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”. Subsequently,
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<bibRefCitation id="EFA9C2C85F70FFCFFCEA1EB7FC10FB66" author="Thunberg, C. P." box="[776,1023,1184,1210]" pageId="2" pageNumber="269" refId="ref6602" refString="Thunberg, C. P. (1823) Flora capensis, sistens plantas promontorii bonae spei Africes, secundem systema sexuale emendatum redactas ad classes, ordines, genera et species, cum differentiis specificis, synonymis et descriptionibus. J. G. Cottae, Stuttgardtiae, 803 pp. https: // doi. org / 10.5962 / bhl. title. 3649" type="book" year="1823">Thunberg (1823: 621)</bibRefCitation>
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provided a morphological description of
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<taxonomicName id="4C38C4BA5F70FFCFFF471ED3FEEBFB02" authorityName="Thunberg" authorityYear="1800" box="[165,260,1220,1246]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="269" phylum="Tracheophyta" rank="species" species="rigida">
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<emphasis id="B94C632B5F70FFCFFF471ED3FEEBFB02" box="[165,260,1220,1246]" italics="true" pageId="2" pageNumber="269">S. rigida</emphasis>
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</taxonomicName>
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together with the following locality information: “
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<emphasis id="B94C632B5F70FFCFFCD51ED3FD9CFADE" italics="true" pageId="2" pageNumber="269">Crescit in campis extra urbem et supra littus maris, pone Duyvels kopp et alibi. Floret Martio, Aprili.</emphasis>
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”
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</paragraph>
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<paragraph id="8B87BF395F70FFCFFF5F1F1BFC5FF9B7" blockId="2.[136,1452,1148,2003]" pageId="2" pageNumber="269">
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The former personal herbarium of Thunberg, containing material that he collected while resident at the
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<collectingRegion id="49FC71DB5F70FFCFFAB11F1BFA62FAFA" box="[1363,1421,1292,1318]" country="South Africa" name="Western Cape" pageId="2" pageNumber="269">Cape</collectingRegion>
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of Good Hope, is housed in Herb. UPS-THUNB, although specimens collected by Thunberg are found in numerous other herbaria (
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<bibRefCitation id="EFA9C2C85F70FFCFFF101F43FE16FAB2" author="Stafleu, F. A. & Cowan, R. S." box="[242,505,1364,1390]" pageId="2" pageNumber="269" refId="ref6447" refString="Stafleu, F. A. & Cowan, R. S. (1986) Taxonomic literature 6, 2 nd ed. Regnum Vegetabile 115. Bohn, Scheltema & Holkema, Utrecht; Dr. W. Junk, The Hague, 926 pp." type="book" year="1986">Stafleu & Cowan 1986</bibRefCitation>
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). The Herb. UPS-THUNB holdings unequivocally represent original material for the names published by Thunberg.
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<bibRefCitation id="EFA9C2C85F70FFCFFE081F6FFD39FA4E" author="Roessler, H." box="[490,726,1400,1427]" pageId="2" pageNumber="269" pagination="71 - 500" refId="ref6338" refString="Roessler, H. (1959) Revision der Arctotideae - Gorteriinae (Compositae). Mitteilungen der Botanische Staatssammlung Munchen 3: 71 - 500." type="journal article" year="1959">Roessler (1959: 188)</bibRefCitation>
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cited the type for
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<taxonomicName id="4C38C4BA5F70FFCFFC441F6FFBE8FA4E" authorityName="Thunberg" authorityYear="1800" box="[934,1031,1400,1426]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="269" phylum="Tracheophyta" rank="species" species="rigida">
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<emphasis id="B94C632B5F70FFCFFC441F6FFBE8FA4E" box="[934,1031,1400,1426]" italics="true" pageId="2" pageNumber="269">S. rigida</emphasis>
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</taxonomicName>
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as “Typus specie: in Herb. Thunberg (UPS)”. Although Roessler did not cite a collection number or specific herbarium sheet number in Herb. UPS, under ICN Art. 7.11, in using the term ‘typus’,
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<bibRefCitation id="EFA9C2C85F70FFCFFD851FD7FCF3FA07" author="Roessler, H." box="[615,796,1472,1499]" pageId="2" pageNumber="269" pagination="71 - 500" refId="ref6338" refString="Roessler, H. (1959) Revision der Arctotideae - Gorteriinae (Compositae). Mitteilungen der Botanische Staatssammlung Munchen 3: 71 - 500." type="journal article" year="1959">Roessler (1959)</bibRefCitation>
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effectively designated a
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<typeStatus id="5483019B5F70FFCFFBDD1FD7FB49FA06" box="[1087,1190,1472,1498]" pageId="2" pageNumber="269" type="lectotype">lectotype</typeStatus>
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for
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<taxonomicName id="4C38C4BA5F70FFCFFB3B1FD7FAD3FA06" authorityName="Thunberg" authorityYear="1800" box="[1241,1340,1472,1498]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="269" phylum="Tracheophyta" rank="species" species="rigida">
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<emphasis id="B94C632B5F70FFCFFB3B1FD7FAD3FA06" box="[1241,1340,1472,1498]" italics="true" pageId="2" pageNumber="269">S. rigida</emphasis>
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</taxonomicName>
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. A single sheet in Herb. UPS-THUNB (UPS-THUNB 18601;
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<figureCitation id="1303A3BC5F70FFCFFD361FF3FCF8FA22" box="[724,791,1508,1534]" captionStart="FIGURE 2" captionStartId="3.[136,229,1952,1974]" captionTargetBox="[250,1338,190,1928]" captionTargetId="figure-17@3.[250,1338,190,1928]" captionTargetPageId="3" captionText="FIGURE 2. Original material for Stobaea rigida Thunb. (UPS-THUNB 18601). Photograph by R.J. McKenzie." figureDoi="http://doi.org/10.5281/zenodo.13214074" httpUri="https://zenodo.org/record/13214074/files/figure.png" pageId="2" pageNumber="269">Fig. 2</figureCitation>
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) is annotated in Thunberg’s handwriting in the lower-right corner with the name “
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<taxonomicName id="4C38C4BA5F70FFCFFE671C1FFDC5F9FE" authorityName="Thunberg" authorityYear="1800" box="[389,554,1544,1570]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="269" phylum="Tracheophyta" rank="species" species="rigida">
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<emphasis id="B94C632B5F70FFCFFE671C1FFDC5F9FE" box="[389,554,1544,1570]" italics="true" pageId="2" pageNumber="269">Stobaea rigida</emphasis>
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</taxonomicName>
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”. The sheet is annotated on the back with the locality information “
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<emphasis id="B94C632B5F70FFCFFAFA1C1EFDF0F99A" italics="true" pageId="2" pageNumber="269">in arvis extra Knocken supra littus in Cap. B. Spei</emphasis>
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” [
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<emphasis id="B94C632B5F70FFCFFDDE1C3BFCEFF99A" box="[572,768,1580,1606]" italics="true" pageId="2" pageNumber="269">Caput bonae spei</emphasis>
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]. This sheet is interpreted to be original material for
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<taxonomicName id="4C38C4BA5F70FFCFFAB31C3BFF23F9B6" authorityName="Thunberg" authorityYear="1800" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="269" phylum="Tracheophyta" rank="species" species="rigida">
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<emphasis id="B94C632B5F70FFCFFAB31C3BFF23F9B6" italics="true" pageId="2" pageNumber="269">Stobaea rigida</emphasis>
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</taxonomicName>
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and the
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<typeStatus id="5483019B5F70FFCFFECE1C47FE7CF9B6" box="[300,403,1616,1642]" pageId="2" pageNumber="269" type="lectotype">lectotype</typeStatus>
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for that name as designated by
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<bibRefCitation id="EFA9C2C85F70FFCFFD141C47FC44F9B7" author="Roessler, H." box="[758,939,1616,1643]" pageId="2" pageNumber="269" pagination="71 - 500" refId="ref6338" refString="Roessler, H. (1959) Revision der Arctotideae - Gorteriinae (Compositae). Mitteilungen der Botanische Staatssammlung Munchen 3: 71 - 500." type="journal article" year="1959">Roessler (1959)</bibRefCitation>
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||
.
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</paragraph>
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<paragraph id="8B87BF395F70FFCFFF5F1C63FA68F80E" blockId="2.[136,1452,1148,2003]" pageId="2" pageNumber="269">
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Three fertile shoots are mounted on the sheet UPS-THUNB 18601. Each shoot is unbranched, ranging in length from ca.
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<quantity id="4CC012DC5F70FFCFFF111C8FFED4F96F" box="[243,315,1688,1715]" metricMagnitude="-1" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="269" unit="cm" value="23.0">23 cm</quantity>
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(left specimen) to ca.
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<quantity id="4CC012DC5F70FFCFFDA41C8FFD61F96F" box="[582,654,1688,1715]" metricMagnitude="-1" metricUnit="m" metricValue="2.9" pageId="2" pageNumber="269" unit="cm" value="29.0">29 cm</quantity>
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(central specimen), and has a terminal capitulum and one to several capitula on short, lateral, axillary branches at the upper nodes. The underground plant parts are not represented in the specimens and were not described by
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<bibRefCitation id="EFA9C2C85F70FFCFFD8F1CF7FCC7F926" author="Thunberg, C. P." box="[621,808,1760,1786]" pageId="2" pageNumber="269" refId="ref6546" refString="Thunberg, C. P. (1800) Prodromus plantarum capensium: quas in promontorio bonae spei Africes, annis 1772 - 1775, collegit Carol. Pet. Thunberg 2. J. Edman, Upsaliae, 191 pp. https: // doi. org / 10.5962 / bhl. title. 84" type="book" year="1800">Thunberg (1800</bibRefCitation>
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||
,
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<bibRefCitation id="EFA9C2C85F70FFCFFCD41CF7FC80F926" author="Thunberg, C. P." box="[822,879,1760,1786]" pageId="2" pageNumber="269" refId="ref6602" refString="Thunberg, C. P. (1823) Flora capensis, sistens plantas promontorii bonae spei Africes, secundem systema sexuale emendatum redactas ad classes, ordines, genera et species, cum differentiis specificis, synonymis et descriptionibus. J. G. Cottae, Stuttgardtiae, 803 pp. https: // doi. org / 10.5962 / bhl. title. 3649" type="book" year="1823">1823</bibRefCitation>
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||
) in his morphological description of
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<taxonomicName id="4C38C4BA5F70FFCFFAC21CF7FA6CF926" authorityName="Thunberg" authorityYear="1800" box="[1312,1411,1760,1786]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="269" phylum="Tracheophyta" rank="species" species="rigida">
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<emphasis id="B94C632B5F70FFCFFAC21CF7FA6CF926" box="[1312,1411,1760,1786]" italics="true" pageId="2" pageNumber="269">S. rigida</emphasis>
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</taxonomicName>
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. In the central specimen, leaves are absent from the lower nodes and only incomplete leaf remnants are present at the middle nodes on the stem, whereas leaves are present at all nodes of the left and right specimens. The upper leaves are morphologically similar in the
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<specimenCount id="9D3E74B05F70FFCFFDEF1D5BFD2AF8BA" box="[525,709,1868,1894]" count="3" pageId="2" pageNumber="269" type="generic">three specimens</specimenCount>
|
||
, i.e., sessile, pinnatifid, the lateral lobes longer than the undivided portion of the lamina along the midrib, similar in length and width of the lateral lobes, and the apex of all lamina lobes is acute and drawn into a conspicuous spine. In the
|
||
<specimenCount id="9D3E74B05F70FFCFFD321D83FC67F872" box="[720,904,1940,1966]" count="3" pageId="2" pageNumber="269" type="generic">three specimens</specimenCount>
|
||
, the lamina is discolorous, with dense appressed tomentum clothing the adaxial surface, but the abaxial surface is more densely tomentose in the lateral specimens.
|
||
</paragraph>
|
||
<caption id="DF47EFB15F71FFCEFF6A1DB7FB53F86B" ID-DOI="http://doi.org/10.5281/zenodo.13214074" ID-Zenodo-Dep="13214074" box="[136,1212,1952,1975]" httpUri="https://zenodo.org/record/13214074/files/figure.png" pageId="3" pageNumber="270" startId="3.[136,229,1952,1974]" targetBox="[250,1338,190,1928]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8B87BF395F71FFCEFF6A1DB7FB53F86B" blockId="3.[136,1212,1952,1975]" box="[136,1212,1952,1975]" pageId="3" pageNumber="270">
|
||
<emphasis id="B94C632B5F71FFCEFF6A1DB7FF11F869" bold="true" box="[136,254,1952,1974]" pageId="3" pageNumber="270">FIGURE 2.</emphasis>
|
||
Original material for
|
||
<taxonomicName id="4C38C4BA5F71FFCEFE321DB6FD45F86B" authority="Thunb." authorityName="Thunberg" authorityYear="1800" box="[464,682,1953,1975]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="270" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F71FFCEFE321DB6FDB1F86A" box="[464,606,1953,1974]" italics="true" pageId="3" pageNumber="270">Stobaea rigida</emphasis>
|
||
Thunb.
|
||
</taxonomicName>
|
||
(UPS-THUNB 18601). Photograph by R.J. McKenzie.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B87BF395F76FFC9FF5F1A89FB51FE4C" blockId="4.[136,1452,158,401]" pageId="4" pageNumber="271">
|
||
Crucially, the central specimen differs in capitulum
|
||
<typeStatus id="5483019B5F76FFC9FCE21A88FCDFFF65" box="[768,816,159,185]" pageId="4" pageNumber="271">type</typeStatus>
|
||
and medial involucral bract morphology from the lateral specimens on the sheet UPS-THUNB 18601 (
|
||
<figureCitation id="1303A3BC5F76FFC9FD631AD5FD2CFF00" box="[641,707,194,220]" captionStart="FIGURE 3" captionStartId="4.[136,229,894,916]" captionTargetBox="[151,1406,425,870]" captionTargetId="figure-142@4.[151,1435,425,870]" captionTargetPageId="4" captionText="FIGURE 3. Capitula on original material for Stobaea rigida Thunb. (UPS-THUNB 18601). A. Terminal capitula on middle specimen. Scale bar = 1 cm. B. Radiate capitulum on right specimen.Arrows indicate individual ray florets. Photographs by R.J. McKenzie." figureDoi="http://doi.org/10.5281/zenodo.13214076" httpUri="https://zenodo.org/record/13214076/files/figure.png" pageId="4" pageNumber="271">Fig. 3</figureCitation>
|
||
). The lateral specimens have radiate capitula, containing peripheral ray florets with a short limb, whereas the central specimen has discoid capitula that lack ray florets. The medial involucral bracts are notably longer in the lateral specimens (ca.
|
||
<quantity id="4CC012DC5F76FFC9FCAD1B1DFC56FEF9" box="[847,953,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="4" pageNumber="271" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">8–12 mm</quantity>
|
||
in the lateral specimens vs ca.
|
||
<quantity id="4CC012DC5F76FFC9FAEE1B1DFA87FEF9" box="[1292,1384,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="4" pageNumber="271" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">4–8 mm</quantity>
|
||
in the central specimen). Accordingly, the medial bract shape differs (lanceolate in the lateral specimens vs ovate-lanceolate in the central specimen). However, in all
|
||
<specimenCount id="9D3E74B05F76FFC9FDB41B45FCE2FEB0" box="[598,781,338,364]" count="3" pageId="4" pageNumber="271" type="generic">three specimens</specimenCount>
|
||
, the bract margins are indicated to be revolute (conferring a convex shape to the bract lamina) and the medial involucral bract width is similar (ca.
|
||
<quantity id="4CC012DC5F76FFC9FBB31B61FB5DFE4C" box="[1105,1202,374,400]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="4" pageNumber="271" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF47EFB15F76FFC9FF6A1969FA8AFC65" ID-DOI="http://doi.org/10.5281/zenodo.13214076" ID-Zenodo-Dep="13214076" httpUri="https://zenodo.org/record/13214076/files/figure.png" pageId="4" pageNumber="271" startId="4.[136,229,894,916]" targetBox="[151,1406,425,870]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8B87BF395F76FFC9FF6A1969FA8AFC65" blockId="4.[136,1452,894,953]" pageId="4" pageNumber="271">
|
||
<emphasis id="B94C632B5F76FFC9FF6A1969FF10FC4F" bold="true" box="[136,255,894,916]" pageId="4" pageNumber="271">FIGURE 3.</emphasis>
|
||
Capitula on original material for
|
||
<taxonomicName id="4C38C4BA5F76FFC9FDAB1968FCC9FC49" authority="Thunb." authorityName="Thunberg" authorityYear="1800" box="[585,806,895,917]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F76FFC9FDAB1968FD36FC48" box="[585,729,895,916]" italics="true" pageId="4" pageNumber="271">Stobaea rigida</emphasis>
|
||
Thunb.
|
||
</taxonomicName>
|
||
(UPS-THUNB 18601). A. Terminal capitula on middle specimen. Scale bar = 1 cm. B. Radiate capitulum on right specimen. Arrows indicate individual ray florets. Photographs by R.J. McKenzie.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B87BF395F76FFC9FF5F19FFFB87FB05" blockId="4.[136,1452,999,2034]" pageId="4" pageNumber="271">
|
||
<bibRefCitation id="EFA9C2C85F76FFC9FF5F19FFFE8DFBDE" author="Harvey, W. H." box="[189,354,999,1026]" pageId="4" pageNumber="271" pagination="44 - 530" refId="ref5790" refString="Harvey, W. H. (1865) Compositae. In: Harvey, W. H. & Sonder, O. W. (Eds.) Flora capensis: being a systematic description of the plants of the Cape Colony, Caffraria, & Port Natal 3. Hodges, Smith, and Co., Dublin, pp. 44 - 530. https: // doi. org / 10.5962 / bhl. title. 821" type="book chapter" year="1865">Harvey (1865)</bibRefCitation>
|
||
is the only previous author to have noted the heterogeneity of the capitula in the original material for
|
||
<taxonomicName id="4C38C4BA5F76FFC9FF521E1CFEFFFBF9" authorityName="Thunberg" authorityYear="1800" box="[176,272,1035,1061]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F76FFC9FF521E1CFEFFFBF9" box="[176,272,1035,1061]" italics="true" pageId="4" pageNumber="271">S. rigida</emphasis>
|
||
</taxonomicName>
|
||
, but he expressed uncertainty as to its significance.
|
||
<bibRefCitation id="EFA9C2C85F76FFC9FCBB1E1BFBD9FBF9" author="Harvey, W. H." box="[857,1078,1035,1062]" pageId="4" pageNumber="271" pagination="44 - 530" refId="ref5790" refString="Harvey, W. H. (1865) Compositae. In: Harvey, W. H. & Sonder, O. W. (Eds.) Flora capensis: being a systematic description of the plants of the Cape Colony, Caffraria, & Port Natal 3. Hodges, Smith, and Co., Dublin, pp. 44 - 530. https: // doi. org / 10.5962 / bhl. title. 821" type="book chapter" year="1865">Harvey (1865: 499)</bibRefCitation>
|
||
stated, “I have only seen
|
||
<emphasis id="B94C632B5F76FFC9FABA1E1BFA67FBF9" box="[1368,1416,1036,1061]" italics="true" pageId="4" pageNumber="271">rays</emphasis>
|
||
on
|
||
<specimenCount id="9D3E74B05F76FFC9FF6A1E27FF38FB96" box="[136,215,1072,1098]" count="1" pageId="4" pageNumber="271" type="generic">one sp.</specimenCount>
|
||
(Hb. Th.); are they generally present? They scarcely exceed the disc flowers in length.”
|
||
<bibRefCitation id="EFA9C2C85F76FFC9FB4B1E38FA8FFB95" author="Thunberg, C. P." box="[1193,1376,1071,1097]" pageId="4" pageNumber="271" refId="ref6546" refString="Thunberg, C. P. (1800) Prodromus plantarum capensium: quas in promontorio bonae spei Africes, annis 1772 - 1775, collegit Carol. Pet. Thunberg 2. J. Edman, Upsaliae, 191 pp. https: // doi. org / 10.5962 / bhl. title. 84" type="book" year="1800">Thunberg (1800</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFA9C2C85F76FFC9FA881E38FA4CFB95" author="Thunberg, C. P." box="[1386,1443,1071,1097]" pageId="4" pageNumber="271" refId="ref6602" refString="Thunberg, C. P. (1823) Flora capensis, sistens plantas promontorii bonae spei Africes, secundem systema sexuale emendatum redactas ad classes, ordines, genera et species, cum differentiis specificis, synonymis et descriptionibus. J. G. Cottae, Stuttgardtiae, 803 pp. https: // doi. org / 10.5962 / bhl. title. 3649" type="book" year="1823">1823</bibRefCitation>
|
||
) did not mention whether the capitula of
|
||
<taxonomicName id="4C38C4BA5F76FFC9FDAB1E44FD46FBB1" authorityName="Thunberg" authorityYear="1800" box="[585,681,1107,1133]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F76FFC9FDAB1E44FD46FBB1" box="[585,681,1107,1133]" italics="true" pageId="4" pageNumber="271">S. rigida</emphasis>
|
||
</taxonomicName>
|
||
were discoid or radiate.
|
||
<bibRefCitation id="EFA9C2C85F76FFC9FC5C1E44FB7DFBB1" author="Lessing, C. F." box="[958,1170,1107,1133]" pageId="4" pageNumber="271" refId="ref6100" refString="Lessing, C. F. (1832) Synopsis generum compositarum earumque dispositionis novae tentamen, monographis multarum Capensium interjectis. Dunckeri et Humblotii, Berolini, 473 pp. https: // doi. org / 10.5962 / bhl. title. 51470" type="book" year="1832">Lessing (1832: 68)</bibRefCitation>
|
||
described the capitula of
|
||
<taxonomicName id="4C38C4BA5F76FFC9FF6A1E60FF06FB4D" authorityName="Thunberg" authorityYear="1800" box="[136,233,1143,1169]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F76FFC9FF6A1E60FF06FB4D" box="[136,233,1143,1169]" italics="true" pageId="4" pageNumber="271">S. rigida</emphasis>
|
||
</taxonomicName>
|
||
as “
|
||
<emphasis id="B94C632B5F76FFC9FEFC1E60FE3FFB4D" box="[286,464,1143,1169]" italics="true" pageId="4" pageNumber="271">breviter radiata</emphasis>
|
||
” and made no reference to discoid capitula in his concept of
|
||
<taxonomicName id="4C38C4BA5F76FFC9FB6C1E60FB00FB4D" authorityName="Thunberg" authorityYear="1800" box="[1166,1263,1143,1169]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F76FFC9FB6C1E60FB00FB4D" box="[1166,1263,1143,1169]" italics="true" pageId="4" pageNumber="271">S. rigida</emphasis>
|
||
</taxonomicName>
|
||
. While
|
||
<bibRefCitation id="EFA9C2C85F76FFC9FAA91E60FF3CFB6A" author="Roessler, H." pageId="4" pageNumber="271" pagination="71 - 500" refId="ref6338" refString="Roessler, H. (1959) Revision der Arctotideae - Gorteriinae (Compositae). Mitteilungen der Botanische Staatssammlung Munchen 3: 71 - 500." type="journal article" year="1959">Roessler (1959)</bibRefCitation>
|
||
noted the general morphological variability of
|
||
<taxonomicName id="4C38C4BA5F76FFC9FD001E8BFCAAFB69" authorityName="Thunberg" authorityYear="1800" box="[738,837,1179,1205]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F76FFC9FD001E8BFCAAFB69" box="[738,837,1179,1205]" italics="true" pageId="4" pageNumber="271">B. rigida</emphasis>
|
||
</taxonomicName>
|
||
, he described the capitula of the species as discoid and made no comment that the original material for
|
||
<taxonomicName id="4C38C4BA5F76FFC9FD431EA8FCEEFB05" authorityName="Thunberg" authorityYear="1800" box="[673,769,1215,1241]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F76FFC9FD431EA8FCEEFB05" box="[673,769,1215,1241]" italics="true" pageId="4" pageNumber="271">S. rigida</emphasis>
|
||
</taxonomicName>
|
||
was in any way heteromorphic.
|
||
</paragraph>
|
||
<paragraph id="8B87BF395F76FFC9FF5F1EF4FD25F9C1" blockId="4.[136,1452,999,2034]" pageId="4" pageNumber="271">
|
||
The presence of discoid capitula with convex, ovate-lanceolate medial involucral bracts are consistent characters in
|
||
<taxonomicName id="4C38C4BA5F76FFC9FF4A1F1FFEE1FAFD" authorityName="Thunberg" authorityYear="1800" box="[168,270,1287,1313]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F76FFC9FF4A1F1FFEE1FAFD" box="[168,270,1287,1313]" italics="true" pageId="4" pageNumber="271">B. rigida</emphasis>
|
||
</taxonomicName>
|
||
. All other characters accessible on the central specimen on the sheet UPS-THUNB 18601 (i.e., stem, leaf and other involucral bract characters) are congruous with the range of morphological variation in
|
||
<taxonomicName id="4C38C4BA5F76FFC9FAC61F3BFA65FA99" authorityName="Thunberg" authorityYear="1800" box="[1316,1418,1323,1349]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F76FFC9FAC61F3BFA65FA99" box="[1316,1418,1323,1349]" italics="true" pageId="4" pageNumber="271">B. rigida</emphasis>
|
||
</taxonomicName>
|
||
as circumscribed by
|
||
<bibRefCitation id="EFA9C2C85F76FFC9FEB21F58FDEAFAB6" author="Roessler, H." box="[336,517,1359,1386]" pageId="4" pageNumber="271" pagination="71 - 500" refId="ref6338" refString="Roessler, H. (1959) Revision der Arctotideae - Gorteriinae (Compositae). Mitteilungen der Botanische Staatssammlung Munchen 3: 71 - 500." type="journal article" year="1959">Roessler (1959)</bibRefCitation>
|
||
. In contrast, radiate capitula and exclusively lanceolate medial involucral bracts are incompatible with
|
||
<taxonomicName id="4C38C4BA5F76FFC9FEBA1F63FE55FA51" authorityName="Thunberg" authorityYear="1800" box="[344,442,1395,1421]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F76FFC9FEBA1F63FE55FA51" box="[344,442,1395,1421]" italics="true" pageId="4" pageNumber="271">B. rigida</emphasis>
|
||
</taxonomicName>
|
||
. These features, together with all other accessible morphological characters for the left and right specimens on the sheet UPS-THUNB 18601, especially the discolorous pinnatifid leaves with the adaxial surface glabrous and non-spinescent and the abaxial surface covered with dense woolly indumentum, are compatible with certain elements included by
|
||
<bibRefCitation id="EFA9C2C85F76FFC9FE2E1FC8FD92FA26" author="Roessler, H." box="[460,637,1503,1530]" pageId="4" pageNumber="271" pagination="71 - 500" refId="ref6338" refString="Roessler, H. (1959) Revision der Arctotideae - Gorteriinae (Compositae). Mitteilungen der Botanische Staatssammlung Munchen 3: 71 - 500." type="journal article" year="1959">Roessler (1959)</bibRefCitation>
|
||
within a broadly circumscribed, polymorphic
|
||
<taxonomicName id="4C38C4BA5F76FFC9FB621FC8FD29F9C1" authority="(Candolle 1838: 517) Roessler (1959: 193)" authorityName="Roessler" authorityPageNumber="193" authorityYear="1959" baseAuthorityName="Candolle" baseAuthorityPageNumber="517" baseAuthorityYear="1838" class="Magnoliopsida" family="Asteraceae" genus="Berkheya" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="variety" species="heterophylla" variety="radiata">
|
||
<emphasis id="B94C632B5F76FFC9FB621FC8FA93FA25" box="[1152,1404,1503,1529]" italics="true" pageId="4" pageNumber="271">Berkheya heterophylla</emphasis>
|
||
var.
|
||
<emphasis id="B94C632B5F76FFC9FF6A1C14FF35F9C1" box="[136,218,1539,1565]" italics="true" pageId="4" pageNumber="271">radiata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA9C2C85F76FFC9FF0B1C14FE26F9C2" author="Candolle, A. P. de" box="[233,457,1539,1566]" pageId="4" pageNumber="271" refId="ref5318" refString="Candolle, A. P. de. (1838 [dated 1837]) Prodromus systematis naturalis regni vegetabilis, sive enumeratio contracta ordinum, generum, specierumque plantarum hucusque cognitarum, juxta methodi naturalis normas digesta 6. Treuttel & Wurtz, Paris, 687 pp. https: // doi. org / 10.5962 / bhl. title. 286" type="book" year="1838">Candolle 1838: 517</bibRefCitation>
|
||
)
|
||
<bibRefCitation id="EFA9C2C85F76FFC9FE3A1C14FD29F9C1" author="Roessler, H." box="[472,710,1539,1566]" pageId="4" pageNumber="271" pagination="71 - 500" refId="ref6338" refString="Roessler, H. (1959) Revision der Arctotideae - Gorteriinae (Compositae). Mitteilungen der Botanische Staatssammlung Munchen 3: 71 - 500." type="journal article" year="1959">Roessler (1959: 193)</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B87BF395F76FFC9FF5F1C30FCBBF929" blockId="4.[136,1452,999,2034]" pageId="4" pageNumber="271">
|
||
Based on the foregoing interpretation, a second-step lectotypification (under ICN Art. 9.17) is required to fix the application of the name
|
||
<taxonomicName id="4C38C4BA5F76FFC9FE7F1C5CFDACF9B9" authorityName="Thunberg" authorityYear="1800" box="[413,579,1611,1637]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F76FFC9FE7F1C5CFDACF9B9" box="[413,579,1611,1637]" italics="true" pageId="4" pageNumber="271">Stobaea rigida</emphasis>
|
||
</taxonomicName>
|
||
and the combination
|
||
<taxonomicName id="4C38C4BA5F76FFC9FCDF1C5CFC1CF9B9" authorityName="Ewart, Jean White & B. Rees" baseAuthorityName="Thunberg" baseAuthorityYear="1800" box="[829,1011,1611,1637]" class="Magnoliopsida" family="Asteraceae" genus="Berkheya" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F76FFC9FCDF1C5CFC1CF9B9" box="[829,1011,1611,1637]" italics="true" pageId="4" pageNumber="271">Berkheya rigida</emphasis>
|
||
</taxonomicName>
|
||
. When a
|
||
<typeStatus id="5483019B5F76FFC9FB811C5BFB7CF9BA" box="[1123,1171,1612,1638]" pageId="4" pageNumber="271">type</typeStatus>
|
||
comprises elements that belong to more than one taxon (e.g., a gathering of mixed material), the name must remain attached to the specimen (as defined in ICN Art. 8.2) that corresponds most nearly with the original description or diagnosis (ICN Art. 9.14). In addition, in the lectotypification process, all aspects of the protologue and which of the specimens correspond most closely with the protologue must be considered (ICN Art. 9.14).
|
||
</paragraph>
|
||
<paragraph id="8B87BF395F76FFC9FF5F1CE8FB41F82E" blockId="4.[136,1452,999,2034]" pageId="4" pageNumber="271">
|
||
The
|
||
<specimenCount id="9D3E74B05F76FFC9FF101CE8FE48F8C5" box="[242,423,1791,1817]" count="3" pageId="4" pageNumber="271" type="generic">three specimens</specimenCount>
|
||
on the sheet UPS-THUNB 18601 are equally concordant with
|
||
<bibRefCitation id="EFA9C2C85F76FFC9FB9B1CE8FAA3F8C5" author="Thunberg, C. P." box="[1145,1356,1791,1817]" pageId="4" pageNumber="271" refId="ref6546" refString="Thunberg, C. P. (1800) Prodromus plantarum capensium: quas in promontorio bonae spei Africes, annis 1772 - 1775, collegit Carol. Pet. Thunberg 2. J. Edman, Upsaliae, 191 pp. https: // doi. org / 10.5962 / bhl. title. 84" type="book" year="1800">Thunberg’s (1800)</bibRefCitation>
|
||
original diagnosis for
|
||
<taxonomicName id="4C38C4BA5F76FFC9FEC41D34FE66F8E1" authorityName="Ewart, Jean White & B. Rees" baseAuthorityName="Thunberg" baseAuthorityYear="1800" box="[294,393,1827,1853]" class="Magnoliopsida" family="Asteraceae" genus="Berkheya" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F76FFC9FEC41D34FE66F8E1" box="[294,393,1827,1853]" italics="true" pageId="4" pageNumber="271">S. rigida</emphasis>
|
||
</taxonomicName>
|
||
and with
|
||
<bibRefCitation id="EFA9C2C85F76FFC9FDE21D34FD3BF8E1" author="Thunberg, C. P." box="[512,724,1827,1853]" pageId="4" pageNumber="271" refId="ref6602" refString="Thunberg, C. P. (1823) Flora capensis, sistens plantas promontorii bonae spei Africes, secundem systema sexuale emendatum redactas ad classes, ordines, genera et species, cum differentiis specificis, synonymis et descriptionibus. J. G. Cottae, Stuttgardtiae, 803 pp. https: // doi. org / 10.5962 / bhl. title. 3649" type="book" year="1823">Thunberg’s (1823)</bibRefCitation>
|
||
more detailed morphological description of
|
||
<taxonomicName id="4C38C4BA5F76FFC9FB3A1D34FAD4F8E1" authorityName="Ewart, Jean White & B. Rees" baseAuthorityName="Thunberg" baseAuthorityYear="1800" box="[1240,1339,1827,1853]" class="Magnoliopsida" family="Asteraceae" genus="Berkheya" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F76FFC9FB3A1D34FAD4F8E1" box="[1240,1339,1827,1853]" italics="true" pageId="4" pageNumber="271">S. rigida</emphasis>
|
||
</taxonomicName>
|
||
. Notably,
|
||
<bibRefCitation id="EFA9C2C85F76FFC9FF6A1D50FEAEF8BD" author="Thunberg, C. P." box="[136,321,1863,1889]" pageId="4" pageNumber="271" refId="ref6546" refString="Thunberg, C. P. (1800) Prodromus plantarum capensium: quas in promontorio bonae spei Africes, annis 1772 - 1775, collegit Carol. Pet. Thunberg 2. J. Edman, Upsaliae, 191 pp. https: // doi. org / 10.5962 / bhl. title. 84" type="book" year="1800">Thunberg (1800</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFA9C2C85F76FFC9FEAF1D50FE69F8BD" author="Thunberg, C. P." box="[333,390,1863,1889]" pageId="4" pageNumber="271" refId="ref6602" refString="Thunberg, C. P. (1823) Flora capensis, sistens plantas promontorii bonae spei Africes, secundem systema sexuale emendatum redactas ad classes, ordines, genera et species, cum differentiis specificis, synonymis et descriptionibus. J. G. Cottae, Stuttgardtiae, 803 pp. https: // doi. org / 10.5962 / bhl. title. 3649" type="book" year="1823">1823</bibRefCitation>
|
||
) did not specify the capitulum type for
|
||
<taxonomicName id="4C38C4BA5F76FFC9FCAB1D50FC45F8BD" authorityName="Ewart, Jean White & B. Rees" baseAuthorityName="Thunberg" baseAuthorityYear="1800" box="[841,938,1863,1889]" class="Magnoliopsida" family="Asteraceae" genus="Berkheya" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F76FFC9FCAB1D50FC45F8BD" box="[841,938,1863,1889]" italics="true" pageId="4" pageNumber="271">S. rigida</emphasis>
|
||
</taxonomicName>
|
||
. However, designation of either of the left or right specimens of UPS-THUNB 18601 as the second-step
|
||
<typeStatus id="5483019B5F76FFC9FCFE1D7CFC6CF859" box="[796,899,1899,1925]" pageId="4" pageNumber="271" type="lectotype">lectotype</typeStatus>
|
||
would conflict with the predominant usage of the name
|
||
<taxonomicName id="4C38C4BA5F76FFC9FF281D98FE91F875" authorityName="Ewart, Jean White & B. Rees" baseAuthorityName="Thunberg" baseAuthorityYear="1800" box="[202,382,1935,1961]" class="Magnoliopsida" family="Asteraceae" genus="Berkheya" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F76FFC9FF281D98FE91F875" box="[202,382,1935,1961]" italics="true" pageId="4" pageNumber="271">Berkheya rigida</emphasis>
|
||
</taxonomicName>
|
||
for the last century, and require the publication of a new name for the species currently known as
|
||
<taxonomicName id="4C38C4BA5F76FFC9FF6A1DA3FF04F811" authorityName="Ewart, Jean White & B. Rees" baseAuthorityName="Thunberg" baseAuthorityYear="1800" box="[136,235,1971,1997]" class="Magnoliopsida" family="Asteraceae" genus="Berkheya" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F76FFC9FF6A1DA3FF04F811" box="[136,235,1971,1997]" italics="true" pageId="4" pageNumber="271">B. rigida</emphasis>
|
||
</taxonomicName>
|
||
. Therefore, we designate the central specimen of the sheet UPS-THUNB 18601 as the second-step
|
||
<typeStatus id="5483019B5F76FFC9FAA71DA4FA43F811" box="[1349,1452,1971,1997]" pageId="4" pageNumber="271" type="lectotype">lectotype</typeStatus>
|
||
for
|
||
<taxonomicName id="4C38C4BA5F76FFC9FF521DC0FEBAF82D" authorityName="Thunberg" authorityYear="1800" box="[176,341,2007,2033]" class="Magnoliopsida" family="Asteraceae" genus="Stobaea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F76FFC9FF521DC0FEBAF82D" box="[176,341,2007,2033]" italics="true" pageId="4" pageNumber="271">Stobaea rigida</emphasis>
|
||
</taxonomicName>
|
||
, consistent with the predominant usage of the combination
|
||
<taxonomicName id="4C38C4BA5F76FFC9FC101DC0FB48F82D" authorityName="Ewart, Jean White & B. Rees" baseAuthorityName="Thunberg" baseAuthorityYear="1800" box="[1010,1191,2007,2033]" class="Magnoliopsida" family="Asteraceae" genus="Berkheya" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="271" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B94C632B5F76FFC9FC101DC0FB48F82D" box="[1010,1191,2007,2033]" italics="true" pageId="4" pageNumber="271">Berkheya rigida</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |