342 lines
46 KiB
XML
342 lines
46 KiB
XML
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<mods:title id="BAA0E753669B7F1CAF50379CC19C1324">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
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<mods:namePart id="5FEA25C383F42E097892997D3610A045">Shi, Hongliang</mods:namePart>
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<mods:namePart id="0C1A2B377C45A7546D80A1635EA09371">Liang, Hongbin</mods:namePart>
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<mods:title id="523CDE762DF8610B5B180EA61F49B2A0">Zootaxa</mods:title>
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<mods:date id="FADBECF5BE1FF82DB70D92CE9A5EBD1A">2023</mods:date>
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<treatment id="038776236279FFA22DEFB2ECFF075C1B" ID-DOI="http://doi.org/10.5281/zenodo.7963751" ID-GBIF-Taxon="209629025" ID-Zenodo-Dep="7963751" LSID="urn:lsid:plazi:treatment:038776236279FFA22DEFB2ECFF075C1B" httpUri="http://treatment.plazi.org/id/038776236279FFA22DEFB2ECFF075C1B" lastPageId="65" lastPageNumber="66" pageId="63" pageNumber="64">
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<subSubSection id="C33494BE6279FFDC2DEFB2ECFD325DEB" box="[151,725,151,178]" pageId="63" pageNumber="64" type="nomenclature">
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<paragraph id="8B91C7356279FFDC2DEFB2ECFD325DEB" blockId="63.[151,1025,151,213]" box="[151,725,151,178]" pageId="63" pageNumber="64">
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<emphasis id="B95A1B276279FFDC2DEFB2ECFD325DEB" bold="true" box="[151,725,151,178]" pageId="63" pageNumber="64">
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[15]
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<taxonomicName id="4C2EBCB66279FFDC2DB7B2ECFD325DEB" authority="Hua, 2002" authorityName="Hua. A" authorityYear="2002" box="[207,725,151,178]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="taiwana" subGenus="Bothynoptera">
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<emphasis id="B95A1B276279FFDC2DB7B2ECFEC05DEB" bold="true" box="[207,295,151,177]" italics="true" pageId="63" pageNumber="64">Parena</emphasis>
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(
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<emphasis id="B95A1B276279FFDC2C40B2ECFE3A5DE8" bold="true" box="[312,477,151,178]" italics="true" pageId="63" pageNumber="64">Bothynoptera</emphasis>
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)
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<emphasis id="B95A1B276279FFDC2C96B2ECFDA95DEB" bold="true" box="[494,590,151,177]" italics="true" pageId="63" pageNumber="64">taiwana</emphasis>
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<bibRefCitation id="EFBFBAC46279FFDC2F2EB2ECFD325DEB" author="Hua, L. Z." box="[598,725,151,177]" pageId="63" pageNumber="64" refId="ref111121" refString="Hua, L. Z. (2002) List of Chinese Insects vol. II. Guangzhou: Zhongshan (Sun Yat-sen) University Press, 612 pp." type="book" year="2002">Hua, 2002</bibRefCitation>
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</taxonomicName>
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||
</emphasis>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C33494BE6279FFDC2DEFB2C0FBE65D8F" box="[151,1025,187,213]" pageId="63" pageNumber="64" type="description">
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<paragraph id="8B91C7356279FFDC2DEFB2C0FBE65D8F" blockId="63.[151,1025,151,213]" box="[151,1025,187,213]" pageId="63" pageNumber="64">
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Habitus:
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<figureCitation id="1315DBB06279FFDC2D85B2C0FEA85D8F" box="[253,335,187,213]" captionStart="FIGURE 36" captionStartId="63.[151,250,1698,1722]" captionTargetBox="[155,1434,977,1670]" captionTargetId="figure-354@63.[151,1436,972,1674]" captionTargetPageId="63" captionText="FIGURE 36. Habitus of Parena (Bothynoptera) taiwana Hua. A Male from Guangxi. B Female from Taiwan. C Male from Taiwan. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959011" httpUri="https://zenodo.org/record/7959011/files/figure.png" pageId="63" pageNumber="64">Fig. 36</figureCitation>
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. Male genitalia:
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<figureCitation id="1315DBB06279FFDC2F72B2C0FDBA5D8F" box="[522,605,187,213]" captionStart="FIGURE 37" captionStartId="64.[151,250,1147,1171]" captionTargetBox="[159,1428,197,1113]" captionTargetId="figure-28@64.[151,1436,181,1124]" captionTargetPageId="64" captionText="FIGURE 37. Median lobe of male genitalia for Parena (Bothynoptera) taiwana Hua, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, Taiwan. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959013" httpUri="https://zenodo.org/record/7959013/files/figure.png" pageId="63" pageNumber="64">Fig. 37</figureCitation>
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. Gonocoxites of ovipositor:
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<figureCitation id="1315DBB06279FFDC2EE3B2C0FC1A5D8F" box="[923,1021,187,213]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="63" pageNumber="64">Fig. 11E</figureCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33494BE6279FFDC2DEFB37BFE9D5C2E" pageId="63" pageNumber="64" type="reference_group">
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<paragraph id="8B91C7356279FFDC2DEFB37BFC8B5C42" blockId="63.[151,1436,256,372]" box="[151,876,256,280]" pageId="63" pageNumber="64">
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<bibRefCitation id="EFBFBAC46279FFDC2DEFB37BFECD5C42" author="Hua, L. Z." box="[151,298,256,280]" pageId="63" pageNumber="64" refId="ref111121" refString="Hua, L. Z. (2002) List of Chinese Insects vol. II. Guangzhou: Zhongshan (Sun Yat-sen) University Press, 612 pp." type="book" year="2002">Hua, 2002: 27</bibRefCitation>
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. replacement name for
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<taxonomicName id="4C2EBCB66279FFDC2F6FB37BFC8E5C42" ID-CoL="4DKPF" authority="Ohkura, 1978" authorityName="Ohkura" authorityYear="1978" box="[535,873,256,280]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="formosana">
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<emphasis id="B95A1B276279FFDC2F6FB37BFD345C42" box="[535,723,256,280]" italics="true" pageId="63" pageNumber="64">Parena formosana</emphasis>
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||
<bibRefCitation id="EFBFBAC46279FFDC2FA2B37BFC8E5C42" author="Ohkura, M." box="[730,873,256,280]" pageId="63" pageNumber="64" pagination="29 - 30" refId="ref112451" refString="Ohkura, M. (1978) A new Parena species from Taiwan (Coleoptera, Harpalidae). Entomological Review of Japan, 32 (1 - 2), 29 - 30." type="journal article" year="1978">Ohkura, 1978</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8B91C7356279FFDC2DEFB364FE9D5C2E" blockId="63.[151,1436,256,372]" pageId="63" pageNumber="64">
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||
<taxonomicName id="4C2EBCB66279FFDC2DEFB364FDF05C6C" ID-CoL="4DKPF" authority="Ohkura, 1978: 29" authorityName="Ohkura" authorityPageNumber="29" authorityYear="1978" box="[151,535,286,310]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="formosana">
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<emphasis id="B95A1B276279FFDC2DEFB364FEBB5C6C" bold="true" box="[151,348,286,310]" italics="true" pageId="63" pageNumber="64">Parena formosana</emphasis>
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||
<materialsCitation id="3B46CD686279FFDC2C1BB365FDF05C6C" ID-GBIF-Occurrence="4115871365" box="[355,535,286,310]" country="Taiwan" location="Kashihara City Museum of Insects" pageId="63" pageNumber="64" specimenCount="1" stateProvince="Nantou" typeStatus="holotype">
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<bibRefCitation id="EFBFBAC46279FFDC2C1BB365FDF05C6C" author="Ohkura, M." box="[355,535,286,310]" pageId="63" pageNumber="64" pagination="29 - 30" refId="ref112451" refString="Ohkura, M. (1978) A new Parena species from Taiwan (Coleoptera, Harpalidae). Entomological Review of Japan, 32 (1 - 2), 29 - 30." type="journal article" year="1978">Ohkura, 1978: 29</bibRefCitation>
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</materialsCitation>
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</taxonomicName>
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(type locality:
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<collectingCountry id="F33987A56279FFDC2FCCB365FCE55C6C" box="[692,770,286,310]" name="Taiwan" pageId="63" pageNumber="64">Taiwan</collectingCountry>
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||
:
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<collectingRegion id="49EA09D76279FFDC2E75B364FCBC5C6C" box="[781,859,287,310]" country="Taiwan" name="Nantou" pageId="63" pageNumber="64">Nantou</collectingRegion>
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;
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<typeStatus id="549579976279FFDC2E1EB365FC585C6C" box="[870,959,286,310]" pageId="63" pageNumber="64" type="holotype">holotype</typeStatus>
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in KCMI:
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<location id="8EF191EE6279FFDC294AB365FA7E5C6C" LSID="urn:lsid:plazi:treatment:038776236279FFA22DEFB2ECFF075C1B:8EF191EE6279FFDC294AB365FA7E5C6C" box="[1074,1433,286,310]" country="Taiwan" name="Kashihara City Museum of Insects" pageId="63" pageNumber="64" stateProvince="Nantou">Kashihara City Museum of Insects</location>
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, Kashihara,
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<collectingCountry id="F33987A56279FFDC2C42B346FE9F5C0F" box="[314,376,317,341]" name="Japan" pageId="63" pageNumber="64">Japan</collectingCountry>
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);
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<bibRefCitation id="EFBFBAC46279FFDC2CF1B346FDCF5C0F" author="Habu, A." box="[393,552,317,341]" pageId="63" pageNumber="64" pagination="51 - 59" refId="ref110845" refString="Habu, A. (1979) Formosan Lebiini collected by Professor T. Shirozu in 1965 (Coleoptera, Carabidae). The Entomological Review of Japan, 33, 51 - 59." type="journal article" year="1979">Habu, 1979: 58</bibRefCitation>
|
||
;
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||
<bibRefCitation id="EFBFBAC46279FFDC2F4AB346FD115C0F" author="Shpeley, D." box="[562,758,317,341]" pageId="63" pageNumber="64" pagination="261 - 349" refId="ref112949" refString="Shpeley, D. (1986) Genera of the subtribe Metallicina and classification, reconstructed phylogeny and geographical history of the species of Euproctinus Leng and Mutchler (Coleoptera: Carabidae: Lebiini). Quaestiones Entomologicae, 22 (4), 261 - 349." type="journal article" year="1986">Shpeley, 1986: 273</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBFBAC46279FFDC2E78B346FC1F5C0F" author="Kirschenhofer, E." box="[768,1016,317,341]" pageId="63" pageNumber="64" pagination="87 - 103" refId="ref111612" refString="Kirschenhofer, E. (2006) Neue Arten der Gattung Parena Motschulsky aus Japan, China Und Indonesien (Sulawesi) (Coleoptera: Carabidae). Mitteilungen des Internationalen Entomologischen Vereins E. V. Frankfurt, 31 (3 - 4), 87 - 103." type="journal article" year="2006">Kirschenhofer, 2006: 88</bibRefCitation>
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||
. Primary homonym of
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||
<taxonomicName id="4C2EBCB66279FFDC2998B345FE915C2E" ID-CoL="4DKPF" authority="Jedlicka, 1946: 9" authorityName="Jedlicka" authorityPageNumber="9" authorityYear="1946" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="formosana">
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||
<emphasis id="B95A1B276279FFDC2998B345FA7B5C0F" box="[1248,1436,317,341]" italics="true" pageId="63" pageNumber="64">Parena formosana</emphasis>
|
||
<bibRefCitation id="EFBFBAC46279FFDC2DBFB327FE915C2E" author="Jedlicka, A." box="[199,374,347,372]" pageId="63" pageNumber="64" refId="ref111358" refString="Jedlicka, A. (1946) Description des Carabides nouveaux de 1 ' Asie orient. Praha: A. Jedlicka, 16 pp." type="book" year="1946">Jedlička, 1946: 9</bibRefCitation>
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||
</taxonomicName>
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||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33494BE6279FFDC2DEFB3D9FB345EEE" pageId="63" pageNumber="64" type="materials_examined">
|
||
<paragraph id="8B91C7356279FFDC2DEFB3D9FB345EEE" blockId="63.[151,1437,417,949]" pageId="63" pageNumber="64">
|
||
<emphasis id="B95A1B276279FFDC2DEFB3D9FE115CE1" bold="true" box="[151,502,417,444]" pageId="63" pageNumber="64">Non-type material examined</emphasis>
|
||
.
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||
<emphasis id="B95A1B276279FFDC2F71B3DAFDBC5CE1" bold="true" box="[521,603,417,443]" pageId="63" pageNumber="64">
|
||
<collectingCountry id="F33987A56279FFDC2F71B3DAFDB15CE1" box="[521,598,417,443]" name="China" pageId="63" pageNumber="64">China</collectingCountry>
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||
:
|
||
</emphasis>
|
||
<specimenCount id="9D280CBC6279FFDC2F1FB3D9FD505CE6" box="[615,695,418,444]" pageId="63" pageNumber="64" type="male">1 male</specimenCount>
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||
(IZAS), "
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||
<collectingRegion id="49EA09D76279FFDC2E57B3D9FC675CE6" box="[815,896,418,444]" country="China" name="Hunan" pageId="63" pageNumber="64">Hunan</collectingRegion>
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||
, Yizhang, Mangshan forestry park, Guizizhai,
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||
<geoCoordinate id="EE1AA1F26279FFDC2DEFB3BDFEFF5CBB" box="[151,280,454,481]" degrees="24.95187" direction="north" orientation="latitude" pageId="63" pageNumber="64" precision="1" value="24.95187">N24.95187</geoCoordinate>
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||
,
|
||
<geoCoordinate id="EE1AA1F26279FFDC2C5BB3BDFE4A5CBA" box="[291,429,454,480]" degrees="112.93079" direction="east" orientation="longitude" pageId="63" pageNumber="64" precision="1" value="112.93079">E112.93079</geoCoordinate>
|
||
,
|
||
<quantity id="4CD66AD06279FFDC2CC1B3BDFDF75CBA" box="[441,528,454,480]" metricMagnitude="3" metricUnit="m" metricValue="1.271" pageId="63" pageNumber="64" unit="m" value="1271.0">1271 m</quantity>
|
||
,
|
||
<date id="FF90E1F56279FFDC2F65B3BDFD405CBA" box="[541,679,454,480]" pageId="63" pageNumber="64" value="2008-07-14">2008.VII.14</date>
|
||
d, Ding Liang lgt.".
|
||
<specimenCount id="9D280CBC6279FFDC2EF2B3BDFC315CBA" box="[906,982,454,480]" pageId="63" pageNumber="64" type="male">1 male</specimenCount>
|
||
(IZAS), "
|
||
<collectingRegion id="49EA09D76279FFDC293DB3BDFB715CBA" box="[1093,1174,454,480]" country="China" name="Hunan" pageId="63" pageNumber="64">Hunan</collectingRegion>
|
||
, Mangshan,
|
||
<date id="FF90E1F56279FFDC285DB3BDFA7F5CBA" box="[1317,1432,454,480]" pageId="63" pageNumber="64" value="2008-07-14">2008.7.14</date>
|
||
, Ding Liang lgt.".
|
||
<specimenCount id="9D280CBC6279FFDC2C16B391FE595F5E" box="[366,446,490,516]" pageId="63" pageNumber="64" type="male">1 male</specimenCount>
|
||
(IZAS), "
|
||
<collectingRegion id="49EA09D76279FFDC2F43B391FD265F5E" box="[571,705,490,516]" country="China" name="Guangdong" pageId="63" pageNumber="64">Guangdong</collectingRegion>
|
||
, Ruyuan county, Nanling res., Ruyang station, on vegetation,
|
||
<geoCoordinate id="EE1AA1F26279FFDC2DEFB075FEF05F72" box="[151,279,526,552]" degrees="24.93730" direction="north" orientation="latitude" pageId="63" pageNumber="64" precision="1" value="24.9373">N24.93730</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1AA1F26279FFDC2C5CB075FE4A5F72" box="[292,429,526,552]" degrees="113.02110" direction="east" orientation="longitude" pageId="63" pageNumber="64" precision="1" value="113.0211">E113.02110</geoCoordinate>
|
||
,
|
||
<quantity id="4CD66AD06279FFDC2CC3B075FDF45F72" box="[443,531,526,552]" metricMagnitude="3" metricUnit="m" metricValue="1.148" pageId="63" pageNumber="64" unit="m" value="1148.0">1148 m</quantity>
|
||
,
|
||
<date id="FF90E1F56279FFDC2F58B075FD4C5F72" box="[544,683,526,552]" pageId="63" pageNumber="64" value="2008-07-19">2008.VII.19</date>
|
||
d, Liang Hongbin lgt.".
|
||
<specimenCount id="9D280CBC6279FFDC2EC5B075FBC75F72" box="[957,1056,526,552]" pageId="63" pageNumber="64" type="female">1 female</specimenCount>
|
||
(ZWWC), "
|
||
<collectingRegion id="49EA09D76279FFDC29D4B075FAC85F72" box="[1196,1327,526,552]" country="China" name="Chongqing" pageId="63" pageNumber="64">Chongqing</collectingRegion>
|
||
, Jiangjin, Simianshan,
|
||
<date id="FF90E1F56279FFDC2C51B049FE2F5F16" box="[297,456,562,588]" pageId="63" pageNumber="64" value="2007-10-23" valueMax="2007-10-28" valueMin="2007-10-23">2007.X.23-28</date>
|
||
, Zhang, Fu & Zeng lgt.".
|
||
<specimenCount id="9D280CBC6279FFDC2F89B049FCA75F16" box="[753,832,562,588]" pageId="63" pageNumber="64" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="9D280CBC6279FFDC2E35B049FC5C5F16" box="[845,955,562,588]" pageId="63" pageNumber="64" type="female">2 females</specimenCount>
|
||
(IZAS), "
|
||
<collectingRegion id="49EA09D76279FFDC2957B049FB755F16" box="[1071,1170,562,588]" country="China" name="Guangxi" pageId="63" pageNumber="64">Guangxi</collectingRegion>
|
||
, Wuming, Damingshan mt.,
|
||
<quantity id="4CD66AD06279FFDC2DB5B02DFEFD5F2A" box="[205,282,598,625]" metricMagnitude="2" metricUnit="m" metricValue="5.18" pageId="63" pageNumber="64" unit="m" value="518.0">518 m</quantity>
|
||
,
|
||
<date id="FF90E1F56279FFDC2C52B02DFE4C5F2A" box="[298,427,598,624]" pageId="63" pageNumber="64" value="2014-06-08">2014.VI.08</date>
|
||
, Lu Yanquan leg. On flower" <
|
||
<figureCitation id="1315DBB06279FFDC2E58B02DFC6C5F2A" box="[800,907,598,624]" captionStart="FIGURE 36" captionStartId="63.[151,250,1698,1722]" captionTargetBox="[155,1434,977,1670]" captionTargetId="figure-354@63.[151,1436,972,1674]" captionTargetPageId="63" captionText="FIGURE 36. Habitus of Parena (Bothynoptera) taiwana Hua. A Male from Guangxi. B Female from Taiwan. C Male from Taiwan. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959011" httpUri="https://zenodo.org/record/7959011/files/figure.png" pageId="63" pageNumber="64">Fig. 36A</figureCitation>
|
||
>. 6 ex (CCCC), "Taiwan, Nantou, Ren'ai, N. Dongyanshan,
|
||
<quantity id="4CD66AD06279FFDC2C46B001FE725FCE" box="[318,405,634,660]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="63" pageNumber="64" unit="m" value="1800.0">1800 m</quantity>
|
||
,
|
||
<date id="FF90E1F56279FFDC2CD8B001FDFA5FCE" box="[416,541,634,660]" pageId="63" pageNumber="64" value="2010-04-22">2010.IV.22</date>
|
||
, Chou Wen-I leg." <
|
||
<figureCitation id="1315DBB06279FFDC2E79B001FC435FCE" box="[769,932,634,660]" captionStart="FIGURE 36" captionStartId="63.[151,250,1698,1722]" captionTargetBox="[155,1434,977,1670]" captionTargetId="figure-354@63.[151,1436,972,1674]" captionTargetPageId="63" captionText="FIGURE 36. Habitus of Parena (Bothynoptera) taiwana Hua. A Male from Guangxi. B Female from Taiwan. C Male from Taiwan. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959011" httpUri="https://zenodo.org/record/7959011/files/figure.png" pageId="63" pageNumber="64">Figs 36B, 36C</figureCitation>
|
||
,
|
||
<figureCitation id="1315DBB06279FFDC2ED7B001FC2A5FCE" box="[943,973,634,660]" captionStart="FIGURE 37" captionStartId="64.[151,250,1147,1171]" captionTargetBox="[159,1428,197,1113]" captionTargetId="figure-28@64.[151,1436,181,1124]" captionTargetPageId="64" captionText="FIGURE 37. Median lobe of male genitalia for Parena (Bothynoptera) taiwana Hua, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, Taiwan. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959013" httpUri="https://zenodo.org/record/7959013/files/figure.png" pageId="63" pageNumber="64">37</figureCitation>
|
||
>. 1 ex (CCCC), "Taiwan, Nantou county, Ren'ai, Sungkang,
|
||
<date id="FF90E1F56279FFDC2C12B0E5FE185FE2" box="[362,511,670,696]" pageId="63" pageNumber="64" value="1994-08-24">1994.VIII.24</date>
|
||
, Chen Changchin lgt.". 1 ex (CCCC), " Taiwan, Nantou county, Ren'ai, Sungkang,
|
||
<quantity id="4CD66AD06279FFDC2DEFB0B9FF085F86" box="[151,239,706,732]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="63" pageNumber="64" unit="m" value="2000.0">2000 m</quantity>
|
||
,
|
||
<date id="FF90E1F56279FFDC2D83B0B9FE775F86" box="[251,400,706,733]" pageId="63" pageNumber="64" value="1995-08-12">1995.VIII.12</date>
|
||
, Lo Chinchi lgt.". 1 ex (CCCC), "Taiwan, Nantou county, Ren'ai, Sungkang,
|
||
<quantity id="4CD66AD06279FFDC2981B0B9FAB65F86" box="[1273,1361,706,732]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="63" pageNumber="64" unit="m" value="2000.0">2000 m</quantity>
|
||
,
|
||
<date id="FF90E1F56279FFDC2825B0B9FF3E5E5A" pageId="63" pageNumber="64" value="2007-07-03">2007. VII.3</date>
|
||
". 1 ex (CCCC), "Taiwan, Nantou county, Ren'ai, Meifeng,
|
||
<date id="FF90E1F56279FFDC2E01B09DFC1E5E5A" box="[889,1017,742,768]" pageId="63" pageNumber="64" value="1994-06-11">1994.VI.11</date>
|
||
, Chen Changchin lgt.". 4 ex (IZAS), "Taiwan, Ilan county, Taiping Shan, Beech Path 38 k,
|
||
<date id="FF90E1F56279FFDC2E75B171FC685E7E" box="[781,911,778,804]" pageId="63" pageNumber="64" value="2020-06-10">2020.VI.10</date>
|
||
, Huang Jialong lgt." <
|
||
<figureCitation id="1315DBB06279FFDC29EEB171FB125E7E" box="[1174,1269,778,804]" captionStart="FIGURE 2" captionStartId="9.[151,250,1853,1877]" captionTargetBox="[158,1424,204,1811]" captionTargetId="figure-19@9.[151,1436,181,1830]" captionTargetPageId="9" captionText="FIGURE 2. Mandibles of Parena spp.: left mandible, dorsum (ld); right mandible, dorsum (rd); left mandible, venter (lv); right mandible, venter (rv). A P. (Crossoglossa) cavipennis (Bates), Shaanxi. B P. (Parena) latecincta (Bates), Zhejiang. C P. (Bothynoptera) dorsigera (Schaum), Guangxi. D P. (Bothynoptera) taiwana Hua, Taiwan. E P. (Bothynoptera) tripunctata (Bates), Beijing. Scale bar = 0.5mm. Abbreviations: rr retinacular ridge of right mandible; vg ventral groove." figureDoi="http://doi.org/10.5281/zenodo.7958941" httpUri="https://zenodo.org/record/7958941/files/figure.png" pageId="63" pageNumber="64">Figs 2D</figureCitation>
|
||
,
|
||
<figureCitation id="1315DBB06279FFDC287BB171FAC35E7E" box="[1283,1316,778,804]" captionStart="FIGURE 3" captionStartId="10.[151,250,881,905]" captionTargetBox="[161,1430,189,857]" captionTargetId="figure-28@10.[151,1436,181,857]" captionTargetPageId="10" captionText="FIGURE 3. A–D Labia of Parena spp.: A P. (Crossoglossa) cavipennis (Bates), Beijing. B P. (Parena) latecincta (Bates), Zhejiang. C P. (Bothynoptera) dorsigera (Schaum), Guangxi. D P. (Bothynoptera) tripunctata (Bates), Beijing. E–F Heads of Parena spp. in lateral view: E P. (Bothynoptera) taiwana Hua, Taiwan, suborbital setae present. F P. (Parena) latecincta (Bates), Yunnan, suborbital seta absent. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7958943" httpUri="https://zenodo.org/record/7958943/files/figure.png" pageId="63" pageNumber="64">3E</figureCitation>
|
||
>.
|
||
<specimenCount id="9D280CBC6279FFDC283BB171FA7B5E7E" box="[1347,1436,778,804]" pageId="63" pageNumber="64" type="male">2 males</specimenCount>
|
||
(CCCC), "Taiwan, Hualien county, Sioulin, Bilyu Scared Tree,
|
||
<date id="FF90E1F56279FFDC2E1FB155FC2A5E12" box="[871,973,814,841]" pageId="63" pageNumber="64" value="1995-05-02">1995.V.2</date>
|
||
, Chen Changchin lgt." <
|
||
<figureCitation id="1315DBB06279FFDC2990B155FAA85E12" box="[1256,1359,814,840]" captionStart="FIGURE 10" captionStartId="16.[151,250,1909,1933]" captionTargetBox="[163,1429,1250,1879]" captionTargetId="figure-182@16.[152,1436,1241,1885]" captionTargetPageId="16" captionText="FIGURE 10. Parameres of male genitalia for Parena spp. A P. (Crossoglossa) cavipennis (Bates), Beijing. B P. (Crossoglossa) testacea (Chaudoir), Shanghai. C P. (Bothynoptera) dorsigera (Schaum), Zhejiang. D P. (Bothynoptera) taiwana Hua, Taiwan. E P. (Bothynoptera) tripunctata (Bates), Shaanxi. F P. (Parena) rubripicta Andrewes, Xizang. G P. (Parena) fasciata (Chaudoir), Taiwan. H P. (Parena) nigrolineata (Chaudoir), Taiwan. I P. (Parena) latecincta (Bates), Yunnan. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958957" httpUri="https://zenodo.org/record/7958957/files/figure.png" pageId="63" pageNumber="64">Fig. 10D</figureCitation>
|
||
>. 1 ex (CCCC), "Taiwan, Hualien county, Sioulin, Bilyu Scared Tree,
|
||
<date id="FF90E1F56279FFDC2E20B129FC335E36" box="[856,980,850,876]" pageId="63" pageNumber="64" value="2006-10-10">2006.X.10</date>
|
||
". 1 ex (CCCC), "Taiwan, Pingtung county, Dahanshan,
|
||
<date id="FF90E1F56279FFDC2C3DB10DFE525ECA" box="[325,437,886,912]" pageId="63" pageNumber="64" value="2008-04-06">2008.IV.6</date>
|
||
, Lin Wensin lgt.". 2 ex (CCCC), "Taiwan, Pintung county, Wutai,
|
||
<date id="FF90E1F56279FFDC29DDB10DFAF35ECA" box="[1189,1300,886,913]" pageId="63" pageNumber="64" value="1995-04-07">1995.IV.7</date>
|
||
, Li Chunlin lgt.".
|
||
<specimenCount id="9D280CBC6279FFDC2DADB1E1FEA65EEE" box="[213,321,922,948]" pageId="63" pageNumber="64" type="female">2 females</specimenCount>
|
||
(CCCC), "Taiwan, Pingtung county, Tai24 Road,
|
||
<quantity id="4CD66AD06279FFDC2E0BB1E1FC2D5EEE" box="[883,970,922,949]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="63" pageNumber="64" unit="m" value="1500.0">1500 m</quantity>
|
||
,
|
||
<date id="FF90E1F56279FFDC2EADB1E1FBDC5EEE" box="[981,1083,922,948]" pageId="63" pageNumber="64" value="1998-05-02">1998.V.2</date>
|
||
" <
|
||
<figureCitation id="1315DBB06279FFDC2924B1E1FB595EEE" box="[1116,1214,922,948]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="63" pageNumber="64">Fig. 11E</figureCitation>
|
||
>.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5197BD6279FFDC2DEFB4D9FE555B87" ID-DOI="http://doi.org/10.5281/zenodo.7959011" ID-Zenodo-Dep="7959011" httpUri="https://zenodo.org/record/7959011/files/figure.png" pageId="63" pageNumber="64" startId="63.[151,250,1698,1722]" targetBox="[155,1434,977,1670]" targetPageId="63" targetType="figure">
|
||
<paragraph id="8B91C7356279FFDC2DEFB4D9FE555B87" blockId="63.[151,1436,1698,1758]" pageId="63" pageNumber="64">
|
||
<emphasis id="B95A1B276279FFDC2DEFB4D9FEFC5BE0" bold="true" box="[151,283,1698,1722]" pageId="63" pageNumber="64">FIGURE 36</emphasis>
|
||
. Habitus of
|
||
<taxonomicName id="4C2EBCB66279FFDC2CE5B4D8FCD25BE0" authority="Hua. A" authorityName="Hua. A" authorityYear="2002" box="[413,821,1698,1722]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="taiwana" subGenus="Bothynoptera">
|
||
<emphasis id="B95A1B276279FFDC2CE5B4D8FE005BE0" box="[413,487,1699,1722]" italics="true" pageId="63" pageNumber="64">Parena</emphasis>
|
||
(
|
||
<emphasis id="B95A1B276279FFDC2C8FB4D9FD665BE0" box="[503,641,1698,1722]" italics="true" pageId="63" pageNumber="64">Bothynoptera</emphasis>
|
||
)
|
||
<emphasis id="B95A1B276279FFDC2FE9B4D8FD045BE0" box="[657,739,1699,1722]" italics="true" pageId="63" pageNumber="64">taiwana</emphasis>
|
||
Hua.
|
||
<emphasis id="B95A1B276279FFDC2E5BB4D9FCD25BE0" bold="true" box="[803,821,1698,1722]" pageId="63" pageNumber="64">A</emphasis>
|
||
</taxonomicName>
|
||
Male from Guangxi.
|
||
<emphasis id="B95A1B276279FFDC2960B4D9FBCE5BE0" bold="true" box="[1048,1065,1698,1722]" pageId="63" pageNumber="64">B</emphasis>
|
||
Female from Taiwan.
|
||
<emphasis id="B95A1B276279FFDC286DB4D9FAC05BE0" bold="true" box="[1301,1319,1698,1722]" pageId="63" pageNumber="64">C</emphasis>
|
||
Male from Taiwan. Scale bars = 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33494BE6279FFDC2DBFB57DFDB05AA3" pageId="63" pageNumber="64" type="diagnosis">
|
||
<paragraph id="8B91C7356279FFDC2DBFB57DFDB05AA3" blockId="63.[151,1437,1798,2041]" pageId="63" pageNumber="64">
|
||
<emphasis id="B95A1B276279FFDC2DBFB57DFE815A7A" bold="true" box="[199,358,1798,1824]" pageId="63" pageNumber="64">Comparisons</emphasis>
|
||
.
|
||
<taxonomicName id="4C2EBCB66279FFDC2C0EB57CFE0A5A7A" authorityName="Hua. A" authorityYear="2002" box="[374,493,1799,1824]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="taiwana">
|
||
<emphasis id="B95A1B276279FFDC2C0EB57CFE0A5A7A" box="[374,493,1799,1824]" italics="true" pageId="63" pageNumber="64">P. taiwana</emphasis>
|
||
</taxonomicName>
|
||
is different from all other species of subgenus
|
||
<taxonomicName id="4C2EBCB66279FFDC2975B57CFB425A7B" authority="Schaum, 1863" box="[1037,1189,1799,1825]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B276279FFDC2975B57CFB425A7B" box="[1037,1189,1799,1825]" italics="true" pageId="63" pageNumber="64">Bothynoptera</emphasis>
|
||
</taxonomicName>
|
||
in the following four characters: male mesotarsomere 1 with biseriate adhesive setae ventrally; antennae extended beyond elytra base; intervals strongly punctate, distance between interval punctures one to two times as that of strial punctures; median lobe of aedeagus very slender, AL/AW 5.5–6.0. Individuals without elytra spots are somewhat similar to
|
||
<taxonomicName id="4C2EBCB66279FFDC2855B508FF3A5AEA" authorityName="Xie & Yu. B" authorityYear="1993" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="shapingensis">
|
||
<emphasis id="B95A1B276279FFDC2855B508FF3A5AEA" italics="true" pageId="63" pageNumber="64">P. shapingensis</emphasis>
|
||
</taxonomicName>
|
||
, but the latter species has a vague pale patch on the elytra. Individuals with four elytra spots are somewhat similar to
|
||
<taxonomicName id="4C2EBCB66279FFDC2C74B5C0FE4E5A8E" authority="Shi & Liang, 2023" authorityName="Shi & Liang" authorityYear="2023" box="[268,425,1979,2004]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="emarginata" status="sp. nov.">
|
||
<emphasis id="B95A1B276279FFDC2C74B5C0FE4E5A8E" box="[268,425,1979,2004]" italics="true" pageId="63" pageNumber="64">P. emarginata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95A1B276279FFDC2CC9B5C0FE155A8F" bold="true" box="[433,498,1979,2005]" pageId="63" pageNumber="64">
|
||
<taxonomicNameLabel id="A269A65C6279FFDC2CC9B5C0FE155A8F" box="[433,498,1979,2005]" pageId="63" pageNumber="64" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, but the latter species has a pair of black stripes on the pronotum, and apex of the labrum dilated and strongly emarginate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5197BD6206FFA32DEFB600FD7159ED" ID-DOI="http://doi.org/10.5281/zenodo.7959013" ID-Zenodo-Dep="7959013" httpUri="https://zenodo.org/record/7959013/files/figure.png" pageId="64" pageNumber="65" startId="64.[151,250,1147,1171]" targetBox="[159,1428,197,1113]" targetPageId="64" targetType="figure">
|
||
<paragraph id="8B91C7356206FFA32DEFB600FD7159ED" blockId="64.[151,1437,1147,1208]" pageId="64" pageNumber="65">
|
||
<emphasis id="B95A1B276206FFA32DEFB600FEFF59CE" bold="true" box="[151,280,1147,1172]" pageId="64" pageNumber="65">FIGURE 37</emphasis>
|
||
. Median lobe of male genitalia for
|
||
<taxonomicName id="4C2EBCB66206FFA32F02B607FC0959C9" authority="Hua" authorityName="Hua. A" authorityYear="2002" box="[634,1006,1148,1172]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="taiwana" subGenus="Bothynoptera">
|
||
<emphasis id="B95A1B276206FFA32F02B607FD2359C9" box="[634,708,1148,1171]" italics="true" pageId="64" pageNumber="65">Parena</emphasis>
|
||
(
|
||
<emphasis id="B95A1B276206FFA32FAAB607FCBB59CE" box="[722,860,1148,1172]" italics="true" pageId="64" pageNumber="65">Bothynoptera</emphasis>
|
||
)
|
||
<emphasis id="B95A1B276206FFA32E12B607FC5B59C9" box="[874,956,1148,1171]" italics="true" pageId="64" pageNumber="65">taiwana</emphasis>
|
||
Hua
|
||
</taxonomicName>
|
||
, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, Taiwan. Scale bar = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33494BE6206FFA32DBFB69DFDF75A8A" pageId="64" pageNumber="65" type="description">
|
||
<paragraph id="8B91C7356206FFA32DBFB69DFDF75A8A" blockId="64.[151,1437,1254,2000]" pageId="64" pageNumber="65">
|
||
<emphasis id="B95A1B276206FFA32DBFB69DFEB6585A" bold="true" box="[199,337,1254,1280]" pageId="64" pageNumber="65">Description</emphasis>
|
||
. Body length 7.1–9.0 mm. Dorsum yellowish brown to pale yellow; elytra with a pair of dark spots centered behind middle of interval 4 one to three intervals wide (
|
||
<figureCitation id="1315DBB06206FFA32E09B771FC30587E" box="[881,983,1290,1316]" captionStart="FIGURE 36" captionStartId="63.[151,250,1698,1722]" captionTargetBox="[155,1434,977,1670]" captionTargetId="figure-354@63.[151,1436,972,1674]" captionTargetPageId="63" captionText="FIGURE 36. Habitus of Parena (Bothynoptera) taiwana Hua. A Male from Guangxi. B Female from Taiwan. C Male from Taiwan. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959011" httpUri="https://zenodo.org/record/7959011/files/figure.png" pageId="64" pageNumber="65">Fig. 36A</figureCitation>
|
||
) in most specimens, without these spots (
|
||
<figureCitation id="1315DBB06206FFA32DE7B755FEE25812" box="[159,261,1326,1352]" captionStart="FIGURE 36" captionStartId="63.[151,250,1698,1722]" captionTargetBox="[155,1434,977,1670]" captionTargetId="figure-354@63.[151,1436,972,1674]" captionTargetPageId="63" captionText="FIGURE 36. Habitus of Parena (Bothynoptera) taiwana Hua. A Male from Guangxi. B Female from Taiwan. C Male from Taiwan. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959011" httpUri="https://zenodo.org/record/7959011/files/figure.png" pageId="64" pageNumber="65">Fig. 36B</figureCitation>
|
||
) or with a pair of additional spots near elytral base (
|
||
<figureCitation id="1315DBB06206FFA32E1CB755FC2D5812" box="[868,970,1326,1352]" captionStart="FIGURE 36" captionStartId="63.[151,250,1698,1722]" captionTargetBox="[155,1434,977,1670]" captionTargetId="figure-354@63.[151,1436,972,1674]" captionTargetPageId="63" captionText="FIGURE 36. Habitus of Parena (Bothynoptera) taiwana Hua. A Male from Guangxi. B Female from Taiwan. C Male from Taiwan. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959011" httpUri="https://zenodo.org/record/7959011/files/figure.png" pageId="64" pageNumber="65">Fig. 36C</figureCitation>
|
||
) in some specimens; legs, antennae, and mouthparts same color as body. Tempora short, abruptly narrowed behind eyes; antennae extended beyond elytra base, to level of first pore on interval 3; antennomere 4 with length approximately two times its width; labrum quadrate, apex straight. Pronotum nearly quadrate, PW/PL = 1.21–1.31, nearly same width as head, PW/HW = 0.95–1.04; lateral explanations narrow; widest at anterior third, weakly rounded at anterior half, and then weakly sinuate before posterior angles; posterior angles nearly rectangular with rounded apex. Elytra slightly dilated to apex, surface without microsculpture or with very faint isodiametric microsculpture; striae shallowly incised, with fine puncture rows in striae; intervals weakly convex, strongly punctate, distance between interval punctures one to two times as that of stria punctures; interval 3 with three setigerous pores; apical truncation indistinct, evenly rounded, sutural angles indistinct. Apex of abdominal sternite VII with two setae on each side in both sexes. Males with biseriate adhesive setae on venter of mesotarsomeres 1–3; adhesive setae present more than three-fourths length of mesotarsomere 1. Median lobe of aedeagus very slender (AL/AW = 5.5–6.0), ventral margin sinuate, apical lamella thin, LL slightly greater than LW, apex rounded. Endophallus with flared basal expansion of primary sclerite very narrow, apical flagellum very short, ending at basal third of median lobe; apical sclerite elongate and well defined, basal core linear, placed near middle of median lobe; basal sheath reduced to very few scales surrounding base of flared expansion, apical sheath large and well scaled; squamate sac not divided, small and triangular, slightly basal to middle of median lobe, right to squamate sheath. Gonocoxite II of ovipositor nearly ovate, length slightly less than basal width, apex projected near inner angle, with six ensiform setae, two on each angle, and two near middle, on outer side of apical projection.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33494BE6207FFA22DBFB2EDFB475DEB" box="[199,1184,150,177]" pageId="65" pageNumber="66" type="distribution">
|
||
<paragraph id="8B91C7356207FFA22DBFB2EDFB475DEB" blockId="65.[151,1437,150,321]" box="[199,1184,150,177]" pageId="65" pageNumber="66">
|
||
<emphasis id="B95A1B276207FFA22DBFB2EDFEBE5DEA" bold="true" box="[199,345,150,176]" pageId="65" pageNumber="66">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="1315DBB06207FFA22C10B2ECFE535DEB" box="[360,436,150,177]" captionStart="MAP 5" captionStartId="47.[152,211,1220,1244]" captionTargetBox="[170,1418,417,1196]" captionTargetId="figure-130@47.[170,1418,417,1196]" captionTargetPageId="47" captionText="MAP 5. Distributions for species in the Parena (Bothynoptera) heteronycha group and Parena (Bothynoptera) taiwana group." figureDoi="http://doi.org/10.5281/zenodo.7963667" httpUri="https://zenodo.org/record/7963667/files/figure.png" pageId="65" pageNumber="66">Map 5</figureCitation>
|
||
, blue).
|
||
<collectingCountry id="F33987A56207FFA22F70B2ECFDAC5DEB" box="[520,587,151,177]" name="China" pageId="65" pageNumber="66">China</collectingCountry>
|
||
(
|
||
<collectingCountry id="F33987A56207FFA22F21B2ECFD575DEB" box="[601,688,151,177]" name="Taiwan" pageId="65" pageNumber="66">Taiwan</collectingCountry>
|
||
, Guangdong, Guangxi, Hunan, Chongqing).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33494BE6207FFA22DBFB2C0FF075C1B" pageId="65" pageNumber="66" type="discussion">
|
||
<paragraph id="8B91C7356207FFA22DBFB2C0FF075C1B" blockId="65.[151,1437,150,321]" pageId="65" pageNumber="66">
|
||
<emphasis id="B95A1B276207FFA22DBFB2C0FED35D8F" bold="true" box="[199,308,187,213]" pageId="65" pageNumber="66">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C2EBCB66207FFA22C39B2C0FD8C5D8F" authority="Ohkura" authorityName="Ohkura" authorityYear="1978" box="[321,619,187,213]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="formosana">
|
||
<emphasis id="B95A1B276207FFA22C39B2C0FDF65D8F" box="[321,529,187,213]" italics="true" pageId="65" pageNumber="66">Parena formosana</emphasis>
|
||
Ohkura
|
||
</taxonomicName>
|
||
is a primary homonym of
|
||
<taxonomicName id="4C2EBCB66207FFA22EEBB2C0FB2F5D8F" authority="Jedlicka" authorityName="Jedlicka" authorityYear="1946" box="[915,1224,187,213]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="formosana">
|
||
<emphasis id="B95A1B276207FFA22EEBB2C0FB855D8F" box="[915,1122,187,213]" italics="true" pageId="65" pageNumber="66">Parena formosana</emphasis>
|
||
Jedlička
|
||
</taxonomicName>
|
||
, although the latter species belongs to the genus
|
||
<taxonomicName id="4C2EBCB66207FFA22CAFB2A4FDB45DA3" box="[471,595,223,249]" class="Insecta" family="Carabidae" genus="Paraphaea" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B276207FFA22CAFB2A4FDB45DA3" box="[471,595,223,249]" italics="true" pageId="65" pageNumber="66">Paraphaea</emphasis>
|
||
</taxonomicName>
|
||
now (
|
||
<bibRefCitation id="EFBFBAC46207FFA22FE0B2A4FCA15DA3" author="Shi, H. & Zhou, H. Z. & Liang, H. B." box="[664,838,223,249]" pageId="65" pageNumber="66" pagination="1 - 129" refId="ref112847" refString="Shi, H. L, Zhou, H. Z. & Liang, H. B. (2013) Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera. ZooKeys, 284, 1 - 129. https: // doi. org / 10.3897 / zookeys. 284.3983" type="journal article" year="2013">
|
||
Shi
|
||
<emphasis id="B95A1B276207FFA22FBDB2A4FD1B5DA3" box="[709,764,223,249]" italics="true" pageId="65" pageNumber="66">et al.</emphasis>
|
||
, 2013
|
||
</bibRefCitation>
|
||
). A replacement name was proposed by
|
||
<bibRefCitation id="EFBFBAC46207FFA22879B2A4FA665DA3" author="Hua, L. Z." box="[1281,1409,223,249]" pageId="65" pageNumber="66" refId="ref111121" refString="Hua, L. Z. (2002) List of Chinese Insects vol. II. Guangzhou: Zhongshan (Sun Yat-sen) University Press, 612 pp." type="book" year="2002">Hua (2002)</bibRefCitation>
|
||
in the List of Chinese Insects. However, this literature was ignored by recent catalogues (
|
||
<bibRefCitation id="EFBFBAC46207FFA2291BB378FB1C5C47" author="Lorenz, W." box="[1123,1275,258,285]" pageId="65" pageNumber="66" refId="ref112064" refString="Lorenz, W. (2005) Systematic list of extant ground beetles of the world (Insecta Coleoptera Geadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition. Tutzing: Wolfgang Lorenz, 530 pp." type="book" year="2005">Lorenz, 2005</bibRefCitation>
|
||
; L̂bl & L̂bl, 2017).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |