347 lines
37 KiB
XML
347 lines
37 KiB
XML
<document id="C174680E71FBB33E73C799A89C099D1A" ID-DOI="10.11646/phytotaxa.48.1.7" ID-ISSN="1179-3163" ID-Zenodo-Dep="4776766" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621533780868" checkinUser="felipe" docAuthor="Snow, Neil, Rabenantoandro, Johny, Randriatafika, Faly, Rabehevitra, David, Razafimamonjy, N. Darie & Cable, Stuart" docDate="2012" docId="0381A35C0063FFD46099FD51FD057A06" docLanguage="en" docName="Phytotaxa.48.39-60.pdf" docOrigin="Phytotaxa 48 (1)" docSource="http://biotaxa.org/Phytotaxa/article/view/phytotaxa.48.1.7" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Eugenia stictophylla N. Snow & Razafim. 2012, sp. nov." docType="treatment" docVersion="3" lastPageNumber="51" masterDocId="FFB8DB240069FFD8600EFFAEFF917F10" masterDocTitle="Studies of Malagasy Eugenia (Myrtaceae) - III: Seven new species of high conservation concern from the eastern littoral forests" masterLastPageNumber="60" masterPageNumber="39" pageNumber="49" updateTime="1698924875650" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="1DF522C3851D8024D3829614F6EBAE5C">
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<mods:title id="2664277F09CED937D51A16C4B496FE3F">Studies of Malagasy Eugenia (Myrtaceae) - III: Seven new species of high conservation concern from the eastern littoral forests</mods:title>
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<mods:name id="370E7E376A468016B0405C206106EE0A" type="personal">
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<mods:roleTerm id="43688A2F4ACCF4D7A5533AECF431D6A4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="425D9BFD42EAF74757060E21FB203E61">Snow, Neil</mods:namePart>
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<mods:affiliation id="DB4EBD1245EEBE79B489A71DE2EC5530">Herbarium Pacificum, B. P. Bishop Museum, 1525 Bernice Street, Honolulu, Hawaii 96825, USA</mods:affiliation>
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<mods:nameIdentifier id="E6B4EF223A8EBD25D0B048F6974480FC" type="email">nsnow2@mt.gov</mods:nameIdentifier>
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</mods:name>
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<mods:name id="E6E9ED26AC542C18B6210C30F91EE2EB" type="personal">
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<mods:roleTerm id="DD365E765D64ABD4F0B8D8B507EAAE80">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CE37082AF970D33D72E856992FBF2702">Rabenantoandro, Johny</mods:namePart>
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<mods:affiliation id="E8AA5660636BB313BE1D46612992BDBF">Rio Tinto / QIT Madagascar Minerals S. A., Villa 3 H Lot J, 169 Antananarivo, 101, MADAGASCAR; E-mails: johny. rabenantoandro @ riotinto. com;</mods:affiliation>
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<mods:nameIdentifier id="75D6278DD0D329406B3A1213AD258139" type="email">johny.rabenantoandro@riotinto.com</mods:nameIdentifier>
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<mods:roleTerm id="7637DABEF6D2556B7891FFC8A40F1034">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F2C25052EC1D95256EF7A37D0A4ED0B5">Randriatafika, Faly</mods:namePart>
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<mods:affiliation id="0C6C0B526053F1A99E7CDA3417344D89">Rio Tinto / QIT Madagascar Minerals S. A., Villa 3 H Lot J, 169 Antananarivo, 101, MADAGASCAR; E-mails: johny. rabenantoandro @ riotinto. com;</mods:affiliation>
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<mods:nameIdentifier id="75E2BBDA7368D23BA6DDD659E086F852" type="email">johny.rabenantoandro@riotinto.com</mods:nameIdentifier>
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<mods:roleTerm id="8C119121278AC3759B56F65070153093">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2FCDC47CC0CF34071E11E75914762B52">Rabehevitra, David</mods:namePart>
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<mods:affiliation id="E369C44270DC810B972D9820FE231994">Rio Tinto / QIT Madagascar Minerals S. A., Villa 3 H Lot J, 169 Antananarivo, 101, MADAGASCAR; E-mails: johny. rabenantoandro @ riotinto. com;</mods:affiliation>
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<mods:nameIdentifier id="2B8310797550614475FF72EE0F10C506" type="email">johny.rabenantoandro@riotinto.com</mods:nameIdentifier>
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<mods:roleTerm id="EF0A230572E0FA5A574000CCED265FC1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BF223EC63B57E69DEA40B81F80857BDE">Razafimamonjy, N. Darie</mods:namePart>
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<mods:affiliation id="231F4EFCA969CD9D6883AF51ECBB09C6">Department of Botany, University of Antananarivo, MADAGASCAR; E-mail: dariejosmail @ yahoo. com</mods:affiliation>
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</mods:role>
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<mods:namePart id="5CF671359123BDA66205B0507F285023">Cable, Stuart</mods:namePart>
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<mods:affiliation id="C6F76EE8B4D38F4BAEEC3A363D9A533E">Royal Botanic Gardens, Kew, Richmond, Surrey, UNITED KINGDOM; E-mail: s. cable @ kew. org</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="877B7E7E54E33B708DF55530D2717219">text</mods:typeOfResource>
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<mods:title id="F963BBFFFC959FE72D253151258BC0EA">Phytotaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="21B923FB437F5E3B8B6A84F86EC1952B">
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<mods:date id="3081FA4DA7517A36E4552449AF3F5550">2012</mods:date>
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<mods:number id="E2B2136F8263D587888C46334F8D32CD">2012-03-15</mods:number>
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<mods:detail id="1CE101D5385BF6018E93534CF226DF61" type="volume">
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<mods:number id="236139FF44D154B790B975B08E9D4238">48</mods:number>
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</mods:detail>
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<mods:start id="1E0B2F34E7E1DA1288F0E7DA6A7943C6">39</mods:start>
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<mods:url id="07A0EE8A4075B62B4E82FE3293BBA854">http://biotaxa.org/Phytotaxa/article/view/phytotaxa.48.1.7</mods:url>
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</mods:location>
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<mods:classification id="98672CDE8F07E3BEC8BF42BC027A7D14">journal article</mods:classification>
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<mods:identifier id="49D3C4EDB1D316B50715A7ACE1623B5F" type="DOI">10.11646/phytotaxa.48.1.7</mods:identifier>
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<mods:identifier id="2939D9889792F9CD5B4E502DA1B2F706" type="ISSN">1179-3163</mods:identifier>
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<mods:identifier id="9001CDEF1F2527DA5392ABAA497294A4" type="Zenodo-Dep">4776766</mods:identifier>
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</mods:mods>
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<treatment id="0381A35C0063FFD46099FD51FD057A06" LSID="urn:lsid:plazi:treatment:0381A35C0063FFD46099FD51FD057A06" httpUri="http://treatment.plazi.org/id/0381A35C0063FFD46099FD51FD057A06" lastPageId="12" lastPageNumber="51" pageId="10" pageNumber="49">
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<subSubSection id="C33241C10063FFD26099FD51FC9F7CB1" pageId="10" pageNumber="49" type="nomenclature">
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<paragraph id="8B97124A0063FFD26099FD51FC467C0C" blockId="10.[151,983,767,796]" box="[151,983,767,796]" pageId="10" pageNumber="49">
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<heading id="D0DFA5260063FFD26099FD51FC467C0C" bold="true" box="[151,983,767,796]" fontSize="12" level="1" pageId="10" pageNumber="49" reason="1">
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<emphasis id="B95CCE580063FFD26099FD51FC467C0C" bold="true" box="[151,983,767,796]" pageId="10" pageNumber="49">
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||
<taxonomicName id="4C2869C90063FFD26099FD51FD597C0C" authority="N. Snow & Razafim." authorityName="N. Snow & Razafim." authorityYear="2012" box="[151,712,767,796]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="stictophylla" status="sp. nov.">
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||
<emphasis id="B95CCE580063FFD26099FD51FE377C0B" bold="true" box="[151,422,767,795]" italics="true" pageId="10" pageNumber="49">Eugenia stictophylla</emphasis>
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||
N. Snow & Razafim.
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||
</taxonomicName>
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||
,
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||
<emphasis id="B95CCE580063FFD262DBFD51FCA97C0B" bold="true" box="[725,824,767,795]" italics="true" pageId="10" pageNumber="49">
|
||
<taxonomicNameLabel id="A26F73230063FFD262DBFD51FCA97C0B" box="[725,824,767,795]" pageId="10" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
|
||
(
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||
<figureCitation id="13130ECF0063FFD26344FD51FC347C0C" box="[842,933,767,796]" captionStart="FIGURE 2" captionStartId="2.[151,244,1785,1806]" captionTargetBox="[166,1430,868,1771]" captionTargetId="figure-397@2.[166,1431,868,1771]" captionTargetPageId="2" captionText="FIGURE 2. Distribution of species and selected specimens of Eugenia in southeastern Madagascar based on vouchered herbarium specimens and recently captured (but non-vouchered) digital images. Remaining forested areas (Petriky, Mandena, Sainte-Luce) along the coast are indicated in gray. In cases where the icons largely or completely overlapped based on geo-coordinates they have been separated slightly for easier viewing (e.g., E. ardyceae, E. stictophylla and E. cloiselii along the coast west of Tolagnaro). Many additional specimens of Eugenia from the region await identification or description and cannot be mapped confidently at the present time." figureDoi="http://doi.org/10.5281/zenodo.4776772" httpUri="https://zenodo.org/record/4776772/files/figure.png" pageId="10" pageNumber="49">Figs. 2</figureCitation>
|
||
,
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||
<figureCitation id="13130ECF0063FFD263BAFD51FC577C0C" box="[948,966,767,796]" captionStart="FIGURE 6" captionStartId="11.[151,244,2014,2035]" captionTargetBox="[191,1396,191,1989]" captionTargetId="figure-26@11.[191,1396,191,1989]" captionTargetPageId="11" captionText="FIGURE 6. Isotype of Eugenia stictophylla (Dumetz 1106 & McPherson [BISH])." figureDoi="http://doi.org/10.5281/zenodo.4776786" httpUri="https://zenodo.org/record/4776786/files/figure.png" pageId="10" pageNumber="49">6</figureCitation>
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||
).
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8B97124A0063FFD26099FCE4FC9F7CB1" blockId="10.[151,1436,842,929]" pageId="10" pageNumber="49">
|
||
<emphasis id="B95CCE580063FFD26099FCE4FC9F7CB1" italics="true" pageId="10" pageNumber="49">
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||
A congeneribis madagascariensibus combinatione characterum sequentium distinguitur: arbor
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||
<quantity id="4CD0BFAF0063FFD264AEFCE4FB757C71" box="[1184,1252,842,865]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="10" pageNumber="49" unit="m" value="5.5" valueMax="7.0" valueMin="4.0">4–7 m</quantity>
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||
alta; petioli
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||
<quantity id="4CD0BFAF0063FFD2657CFCE4FEB07C91" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.5" metricValueMin="3.5" pageId="10" pageNumber="49" unit="mm" value="4.5" valueMax="5.5" valueMin="3.5">3.5- 5.5 mm</quantity>
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||
longi, sulcati; folia
|
||
<quantity id="4CD0BFAF0063FFD2620AFCC4FDE67C91" box="[516,631,874,897]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="10" pageNumber="49" unit="mm" value="35.0" valueMax="50.0" valueMin="20.0">20–50 mm</quantity>
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||
longa, 7–12(–15) mm lata, anguste elliptica, basi cuneata, apice obtuso, lamina dense glandulosa; petala alba; stamina 35–70.
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||
</emphasis>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33241C10063FFD46099FC64FCB17E63" lastPageId="12" lastPageNumber="51" pageId="10" pageNumber="49" type="materials_examined">
|
||
<paragraph id="8B97124A0063FFD26099FC64FE9D7B31" blockId="10.[151,1437,970,1057]" pageId="10" pageNumber="49">
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||
<emphasis id="B95CCE580063FFD26099FC64FF497CF2" bold="true" box="[151,216,970,994]" pageId="10" pageNumber="49">Type:</emphasis>
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||
―
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||
<materialsCitation id="3B4018170063FFD260FCFC64FE967B31" collectingDate="1990-12-05" collectorName="de Petriky. & N. Dumetz & G. McPherson" country="Madagascar" latitude="-25.083334" location="District Tolagnaro" longLatPrecision="1248" longitude="46.866665" pageId="10" pageNumber="49" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
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||
<collectingCountry id="F33F52DA0063FFD260FCFC64FE227CF1" box="[242,435,970,993]" name="Madagascar" pageId="10" pageNumber="49">MADAGASCAR</collectingCountry>
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||
. Prov.
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||
<collectingRegion id="49ECDCA80063FFD261F0FC64FDDF7CF1" box="[510,590,970,993]" country="Madagascar" name="Toliara" pageId="10" pageNumber="49">Toliara</collectingRegion>
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||
:
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||
<location id="8EF744910063FFD26257FC64FCB07CF1" LSID="urn:lsid:plazi:treatment:0381A35C0063FFD46099FD51FD057A06:8EF744910063FFD26257FC64FCB07CF1" box="[601,801,970,993]" country="Madagascar" latitude="-25.083334" longLatPrecision="1248" longitude="46.866665" name="District Tolagnaro" pageId="10" pageNumber="49" stateProvince="Toliara">District Tolagnaro</location>
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||
; forêt
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||
<collectorName id="26DD779C0063FFD26366FC64FC4C7CF1" box="[872,989,970,993]" pageId="10" pageNumber="49">de Petriky.</collectorName>
|
||
<geoCoordinate id="EE1C748D0063FFD263EAFC64FBAF7CF1" box="[996,1086,970,993]" degrees="25" direction="south" minutes="05" orientation="latitude" pageId="10" pageNumber="49" precision="925" value="-25.083334">25°05’S</geoCoordinate>
|
||
,
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||
<geoCoordinate id="EE1C748D0063FFD26445FC64FB377CF1" box="[1099,1190,970,993]" degrees="46" direction="east" minutes="52" orientation="longitude" pageId="10" pageNumber="49" precision="925" value="46.866665">46°52’E</geoCoordinate>
|
||
,
|
||
<date id="FF96348A0063FFD264BDFC64FAE77CF1" box="[1203,1398,970,993]" pageId="10" pageNumber="49" value="1990-12-05">
|
||
<collectingDate id="EFD2CD620063FFD264BDFC64FAE77CF1" box="[1203,1398,970,993]" pageId="10" pageNumber="49" value="1990-12-05">5 December 1990</collectingDate>
|
||
</date>
|
||
,
|
||
<emphasis id="B95CCE580063FFD2658AFC64FD8B7B11" italics="true" pageId="10" pageNumber="49">
|
||
<collectorName id="26DD779C0063FFD2658AFC64FE887B11" pageId="10" pageNumber="49">N. Dumetz</collectorName>
|
||
1106 &
|
||
<collectorName id="26DD779C0063FFD26170FC44FD8B7B11" box="[382,538,1002,1025]" pageId="10" pageNumber="49">G. McPherson</collectorName>
|
||
</emphasis>
|
||
(
|
||
<typeStatus id="5493ACE80063FFD26225FC44FD1B7B11" box="[555,650,1002,1025]" pageId="10" pageNumber="49" type="holotype">holotype</typeStatus>
|
||
MO! [sheet no. 3778655]; isotypes BISH! [sheet no. 747508], MO, P, TAN)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B97124A0063FFD46099FBE5FE707E36" blockId="10.[151,1436,1098,2014]" lastBlockId="12.[151,1436,152,1302]" lastPageId="12" lastPageNumber="51" pageId="10" pageNumber="49">
|
||
<emphasis id="B95CCE580063FFD26099FBE5FF467B74" box="[151,215,1099,1124]" italics="true" pageId="10" pageNumber="49">Trees</emphasis>
|
||
4–7 meters. Indumentum (where present) shortly sericeous, trichomes dibrachiate and ferrugineous.
|
||
<emphasis id="B95CCE580063FFD26099FBDFFE867B9A" box="[151,279,1137,1162]" italics="true" pageId="10" pageNumber="49">Branchlets</emphasis>
|
||
laterally compressed and maroon or pinkish on emergence, becoming terete and grayish (dried), smooth, sericeous below the base of (often supra-axillary) pedicels but glabrous or only sparsely sericeous elsewhere; oil glands sparse, indistinct; internodes 1.0–
|
||
<quantity id="4CD0BFAF0063FFD2633EFB10FC137BC8" box="[816,898,1214,1240]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="10" pageNumber="49" unit="cm" value="2.2">2.2 cm</quantity>
|
||
long. Stipules of short, setose hairs (
|
||
<typeStatus id="5493ACE80063FFD2654FFB10FAE77BC8" box="[1345,1398,1214,1240]" pageId="10" pageNumber="49">type</typeStatus>
|
||
A:
|
||
<bibRefCitation id="EFB96FBB0063FFD26099FB4BFEEF7BEF" author="Snow, N. & Guymer, G. P. & Sawvel, G." box="[151,382,1253,1279]" pageId="10" pageNumber="49" pagination="1 - 95" refId="ref12113" refString="Snow, N., Guymer, G. P. & Sawvel, G. (2003) Systematics of Austromyrtus, Lenwebbia, and the Australian species of Gossia (Myrtaceae). Systematic Botany Monographs 65: 1 - 95." type="journal article" year="2003">
|
||
Snow
|
||
<emphasis id="B95CCE580063FFD260EDFB4BFE8E7BEE" box="[227,287,1253,1278]" italics="true" pageId="10" pageNumber="49">et al.</emphasis>
|
||
2003:7
|
||
</bibRefCitation>
|
||
).
|
||
<emphasis id="B95CCE580063FFD2619AFB4BFE777BEE" box="[404,486,1253,1278]" italics="true" pageId="10" pageNumber="49">Leaves</emphasis>
|
||
thinly coriaceous, mostly evenly spaced on branchlets, secondary veins faintly protruding above (dried material), slightly discolorous, somewhat glossy above, matte below; venation (obscurely) brochidodromous.
|
||
<emphasis id="B95CCE580063FFD26206FA9DFDF67A5C" box="[520,615,1331,1356]" italics="true" pageId="10" pageNumber="49">Petioles</emphasis>
|
||
<quantity id="4CD0BFAF0063FFD2627FFA9CFC917A5C" box="[625,768,1330,1356]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.5" metricValueMin="3.5" pageId="10" pageNumber="49" unit="mm" value="4.5" valueMax="5.5" valueMin="3.5">3.5–5.5 mm</quantity>
|
||
long, deeply sulcate adaxially, prominently glabrescent along apex of sulcus, abaxially verrucose.
|
||
<emphasis id="B95CCE580063FFD26293FAF7FCB87A62" box="[669,809,1369,1394]" italics="true" pageId="10" pageNumber="49">Leaf blades</emphasis>
|
||
2.0–5.0 x 0.7–1.2(–1.5) cm, narrowly elliptic, base strongly cuneate, margin flat or drying slightly revolute, obtuse; adaxial surface glabrescent, secondary nerves indistinct and somewhat protruding (dried material), oil glands small but dense (use magnification), midvein sulcate proximally but becoming flush distally; abaxial surface glabrous, oil glands much less prominent than above, secondary veins obscure (sometimes scarcely visible), of 10–14 pairs, intramarginal vein indistinct, closely paralleling margin, ca.
|
||
<quantity id="4CD0BFAF0063FFD2621FF9B4FDFF7924" box="[529,622,1562,1588]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="49" unit="mm" value="0.5">0.5 mm</quantity>
|
||
from margin at midpoint of blade.
|
||
<emphasis id="B95CCE580063FFD2642FF9B5FB527924" box="[1057,1219,1563,1588]" italics="true" pageId="10" pageNumber="49">Inflorescence</emphasis>
|
||
a solitary flower, axillary or (mostly) supra-axillary.
|
||
<emphasis id="B95CCE580063FFD26243F9EFFD5A794A" box="[589,715,1601,1626]" italics="true" pageId="10" pageNumber="49">Peduncles</emphasis>
|
||
7–12(–15) mm long, laterally compressed (dried), mostly ascending, somewhat flexuous, sericeous near base but more sparsely distally, oil glands faint, moderately common.
|
||
<emphasis id="B95CCE580063FFD2611CF921FE0479B8" box="[274,405,1679,1704]" italics="true" pageId="10" pageNumber="49">Bracteoles</emphasis>
|
||
2, 0.5–1.0 mm long x <
|
||
<quantity id="4CD0BFAF0063FFD262C0F920FCA179B8" box="[718,816,1678,1704]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="49" unit="mm" value="0.5">0.5 mm</quantity>
|
||
, ovate to broadly ovate, scale-like, stiffly erect to ascending, stiff, sparsely sericeous, persisting in flower.
|
||
<emphasis id="B95CCE580063FFD2634DF91BFC4779DE" box="[835,982,1717,1742]" italics="true" pageId="10" pageNumber="49">Hypanthium</emphasis>
|
||
1.5–2.0 mm long (dried), ca.
|
||
<quantity id="4CD0BFAF0063FFD2654CF91BFF5479E6" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.4" metricValueMin="1.3" pageId="10" pageNumber="49" unit="mm" value="1.85" valueMax="2.4" valueMin="1.3">1.3–2.4 mm</quantity>
|
||
wide below base of calyx lobes, obconic, terete, sparsely sericeous (especially above base of bracteoles towards calyx lobes), oil glands small but dense. Ovary apex sparsely short-villous along outer edges (i.e., in region adjacent to inner margin of staminal ring).
|
||
<emphasis id="B95CCE580063FFD262E6F887FCE37852" box="[744,882,1833,1858]" italics="true" pageId="10" pageNumber="49">Calyx lobes</emphasis>
|
||
4, 1.5–3.0 mm long, distinct and dimorphic in bud (the inner 2 somewhat longer than the outer 2), orbicular to shortly to broadly ovate, ciliate and sparsely sericeous, densely glandular (glands larger and more protruding than those of hypanthium), persistence in fruit unknown.
|
||
<emphasis id="B95CCE580063FFD26141F833FE0978A6" box="[335,408,1949,1974]" italics="true" pageId="10" pageNumber="49">Petals</emphasis>
|
||
4, 5–
|
||
<quantity id="4CD0BFAF0063FFD261D3F833FDB178A7" box="[477,544,1949,1975]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="49" unit="mm" value="7.0">7 mm</quantity>
|
||
long, (2.5–)3.5–4.0 mm wide, obovate or widely obovate, white, glabrous, oil glands common and of similar size to those of calyx lobes.
|
||
<emphasis id="B95CCE580063FFD2638AF86AFC7778CD" box="[900,998,1988,2013]" italics="true" pageId="10" pageNumber="49">Stamens</emphasis>
|
||
35–70, in several whorls; filaments
|
||
<quantity id="4CD0BFAF0065FFD46099FF36FF6B7FA2" box="[151,250,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="12" pageNumber="51" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3–6 mm</quantity>
|
||
long; anthers
|
||
<quantity id="4CD0BFAF0065FFD461A7FF36FDAB7FA2" box="[425,570,152,178]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="12" pageNumber="51" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.5–0.8 mm</quantity>
|
||
long, globose to subcylindrical, sub-basifixed, with a single prominent apical gland.
|
||
<emphasis id="B95CCE580065FFD4613BFF11FEFC7FC8" box="[309,365,191,216]" italics="true" pageId="12" pageNumber="51">Style</emphasis>
|
||
<quantity id="4CD0BFAF0065FFD4617BFF10FE7D7FC8" box="[373,492,190,216]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="12" pageNumber="51" unit="mm" value="4.75" valueMax="5.0" valueMin="4.5">4.5–5 mm</quantity>
|
||
long, glabrous, tapering to a narrow punctiform stigma. Locules 2; placentation parietal, placenta capitate from middle of locule; ovules 8–13 per ovary, radiating from placenta.
|
||
<emphasis id="B95CCE580065FFD4650AFF4BFAD17FEE" box="[1284,1344,229,254]" italics="true" pageId="12" pageNumber="51">Fruit</emphasis>
|
||
ca. 2–3 x
|
||
<quantity id="4CD0BFAF0065FFD460A3FEA2FE967E36" box="[173,263,268,294]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="12" pageNumber="51" unit="cm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 cm</quantity>
|
||
, yellowish-brown.
|
||
</paragraph>
|
||
<caption id="DF5742C20062FFD36099F870FC2378E3" ID-DOI="http://doi.org/10.5281/zenodo.4776786" ID-Zenodo-Dep="4776786" box="[151,946,2014,2035]" httpUri="https://zenodo.org/record/4776786/files/figure.png" pageId="11" pageNumber="50" startId="11.[151,244,2014,2035]" targetBox="[191,1396,191,1989]" targetPageId="11">
|
||
<paragraph id="8B97124A0062FFD36099F870FC2378E3" blockId="11.[151,946,2014,2035]" box="[151,946,2014,2035]" pageId="11" pageNumber="50">
|
||
<emphasis id="B95CCE580062FFD36099F870FE9D78E3" bold="true" box="[151,268,2014,2035]" pageId="11" pageNumber="50">FIGURE 6.</emphasis>
|
||
Isotype of
|
||
<taxonomicName id="4C2869C90062FFD36176F870FDAF78E3" box="[376,574,2014,2035]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="50" phylum="Arthropoda" rank="species" species="stictophylla">
|
||
<emphasis id="B95CCE580062FFD36176F870FDAF78E3" box="[376,574,2014,2035]" italics="true" pageId="11" pageNumber="50">Eugenia stictophylla</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B95CCE580062FFD36242F870FCC678E3" box="[588,855,2014,2035]" italics="true" pageId="11" pageNumber="50">Dumetz 1106 & McPherson</emphasis>
|
||
[BISH]).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B97124A0065FFD460C8FE9CFCB17E63" blockId="12.[151,1436,152,1302]" pageId="12" pageNumber="51">
|
||
<emphasis id="B95CCE580065FFD460C8FE9CFE0D7E5C" bold="true" box="[198,412,306,332]" pageId="12" pageNumber="51">Field characters:</emphasis>
|
||
―Trees 4–7 meters. Petioles deeply sulcate above; leaves densely glandular on upper surface, narrowly elliptic, apically obtuse. Petals white.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33241C10065FFD460C8FED1FC457EF7" pageId="12" pageNumber="51" type="distribution">
|
||
<paragraph id="8B97124A0065FFD460C8FED1FB247E8A" blockId="12.[151,1436,152,1302]" box="[198,1205,383,410]" pageId="12" pageNumber="51">
|
||
<emphasis id="B95CCE580065FFD460C8FED1FEF87E89" bold="true" box="[198,361,383,409]" pageId="12" pageNumber="51">Distribution:</emphasis>
|
||
―Known only from Petriky and Ambinanibe forests west of Tolagnaro.
|
||
</paragraph>
|
||
<paragraph id="8B97124A0065FFD460C8FE08FBB37ED0" blockId="12.[151,1436,152,1302]" box="[198,1058,422,448]" pageId="12" pageNumber="51">
|
||
<emphasis id="B95CCE580065FFD460C8FE08FE5B7ED0" bold="true" box="[198,458,422,448]" pageId="12" pageNumber="51">Habitat and ecology:</emphasis>
|
||
―Littoral forests on sand; elevation to about
|
||
<specimenCount id="9D2ED9C30065FFD463D1FE08FBB37ED0" box="[991,1058,422,448]" pageId="12" pageNumber="51" type="generic">
|
||
<quantity id="4CD0BFAF0065FFD463D1FE08FB8F7ED0" box="[991,1054,422,448]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="51" unit="m" value="15.0">15 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
<paragraph id="8B97124A0065FFD460C8FE63FC457EF7" blockId="12.[151,1436,152,1302]" box="[198,980,461,487]" pageId="12" pageNumber="51">
|
||
<emphasis id="B95CCE580065FFD460C8FE63FEC37EF7" bold="true" box="[198,338,461,487]" pageId="12" pageNumber="51">Phenology:</emphasis>
|
||
―Flowering December and January; fruits not known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33241C10065FFD460C8FE5DFC5E7C7C" pageId="12" pageNumber="51" type="etymology">
|
||
<paragraph id="8B97124A0065FFD460C8FE5DFE2F7D24" blockId="12.[151,1436,152,1302]" pageId="12" pageNumber="51">
|
||
<emphasis id="B95CCE580065FFD460C8FE5DFEC67D1D" bold="true" box="[198,343,499,525]" pageId="12" pageNumber="51">Etymology:</emphasis>
|
||
―The specific epithet is derived from the Greek elements
|
||
<emphasis id="B95CCE580065FFD4641DFE5AFBD57D1D" box="[1043,1092,500,525]" italics="true" pageId="12" pageNumber="51">stict</emphasis>
|
||
and
|
||
<emphasis id="B95CCE580065FFD4648EFE5AFB2B7D1D" box="[1152,1210,500,525]" italics="true" pageId="12" pageNumber="51">phyll</emphasis>
|
||
in reference to the densely glandular leaves.
|
||
</paragraph>
|
||
<paragraph id="8B97124A0065FFD460C8FDEEFE0C7DB8" blockId="12.[151,1436,152,1302]" pageId="12" pageNumber="51">
|
||
<emphasis id="B95CCE580065FFD460C8FDEEFD487D4A" bold="true" box="[198,729,576,602]" pageId="12" pageNumber="51">
|
||
Additional specimen examined (
|
||
<typeStatus id="5493ACE80065FFD4625AFDEEFD5A7D4A" box="[596,715,576,602]" pageId="12" pageNumber="51" type="paratype">paratype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
―
|
||
<materialsCitation id="3B4018170065FFD462F8FDEFFE087DB8" collectingDate="1990-01-12" collectorName="G. McPherson" country="Madagascar" latitude="-25.083334" location="Petriky" longLatPrecision="1248" longitude="46.866665" municipality="Tolagnaro region" pageId="12" pageNumber="51" specimenCount="1" stateProvince="Toliara">
|
||
<collectingCountry id="F33F52DA0065FFD462F8FDEFFC5A7D4B" box="[758,971,577,603]" name="Madagascar" pageId="12" pageNumber="51">MADAGASCAR</collectingCountry>
|
||
. Prov.
|
||
<collectingRegion id="49ECDCA80065FFD46417FDEFFBE17D4B" box="[1049,1136,577,603]" country="Madagascar" name="Toliara" pageId="12" pageNumber="51">Toliara</collectingRegion>
|
||
:
|
||
<collectingMunicipality id="6BF388300065FFD46474FDEFFAD47D4B" box="[1146,1349,577,603]" pageId="12" pageNumber="51">Tolagnaro region</collectingMunicipality>
|
||
; SW of town in forest called
|
||
<location id="8EF744910065FFD46180FDC6FE757D92" LSID="urn:lsid:plazi:treatment:0381A35C0063FFD46099FD51FD057A06:8EF744910065FFD46180FDC6FE757D92" box="[398,484,616,642]" country="Madagascar" latitude="-25.083334" longLatPrecision="1248" longitude="46.866665" municipality="Tolagnaro region" name="Petriky" pageId="12" pageNumber="51" stateProvince="Toliara">Petriky</location>
|
||
, along QIT road,
|
||
<geoCoordinate id="EE1C748D0065FFD462BCFDC6FC857D92" box="[690,788,616,642]" degrees="25" direction="south" minutes="05" orientation="latitude" pageId="12" pageNumber="51" precision="925" value="-25.083334">25°05’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1C748D0065FFD4632FFDC6FC147D92" box="[801,901,616,642]" degrees="46" direction="east" minutes="52" orientation="longitude" pageId="12" pageNumber="51" precision="925" value="46.866665">46°52’E</geoCoordinate>
|
||
,
|
||
<date id="FF96348A0065FFD4639CFDC6FBC77D92" box="[914,1110,616,642]" pageId="12" pageNumber="51" value="1990-01-12">
|
||
<collectingDate id="EFD2CD620065FFD4639CFDC6FBC77D92" box="[914,1110,616,642]" pageId="12" pageNumber="51" value="1990-01-12">12 January 1990</collectingDate>
|
||
</date>
|
||
,
|
||
<emphasis id="B95CCE580065FFD4646CFDC6FA0D7D91" box="[1122,1436,616,641]" italics="true" pageId="12" pageNumber="51">
|
||
<collectorName id="26DD779C0065FFD4646CFDC6FA9B7D91" box="[1122,1290,616,641]" pageId="12" pageNumber="51">G. McPherson</collectorName>
|
||
14774 et al.
|
||
</emphasis>
|
||
(G, K!, MO!, P, TAN)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B97124A0065FFD460C8FD1BFC5E7C7C" blockId="12.[151,1436,152,1302]" pageId="12" pageNumber="51">
|
||
<emphasis id="B95CCE580065FFD460C8FD1BFE5C7DDF" bold="true" box="[198,461,693,719]" pageId="12" pageNumber="51">Conservation status:</emphasis>
|
||
―The imprecise geo-coordinates preclude knowing whether the
|
||
<specimenCount id="9D2ED9C30065FFD464E4FD1BFA0D7DDF" box="[1258,1436,693,719]" pageId="12" pageNumber="51" type="generic">two specimens</specimenCount>
|
||
occur(ed) in conservation zone P1 of Petriky Forest (
|
||
<figureCitation id="13130ECF0065FFD46321FD72FCEC7DE6" box="[815,893,732,758]" captionStart="FIGURE 2" captionStartId="2.[151,244,1785,1806]" captionTargetBox="[166,1430,868,1771]" captionTargetId="figure-397@2.[166,1431,868,1771]" captionTargetPageId="2" captionText="FIGURE 2. Distribution of species and selected specimens of Eugenia in southeastern Madagascar based on vouchered herbarium specimens and recently captured (but non-vouchered) digital images. Remaining forested areas (Petriky, Mandena, Sainte-Luce) along the coast are indicated in gray. In cases where the icons largely or completely overlapped based on geo-coordinates they have been separated slightly for easier viewing (e.g., E. ardyceae, E. stictophylla and E. cloiselii along the coast west of Tolagnaro). Many additional specimens of Eugenia from the region await identification or description and cannot be mapped confidently at the present time." figureDoi="http://doi.org/10.5281/zenodo.4776772" httpUri="https://zenodo.org/record/4776772/files/figure.png" pageId="12" pageNumber="51">Fig. 2</figureCitation>
|
||
; see also
|
||
<bibRefCitation id="EFB96FBB0065FFD463F9FD72FB4C7DE6" author="Lowry, P. P. II & Randriatafika, F. & Rabenantoandro, J." box="[1015,1245,732,758]" pageId="12" pageNumber="51" pagination="55 - 63" refId="ref11601" refString="Lowry, P. P. II, Randriatafika, F., & Rabenantoandro, J. (2008) Conservation status of vascular plant species from the QMM / Rio Tinto mining area at Mandena, Tolagnaro (Fort Dauphin) region, southeast Madagascar. Madagascar Conservation and Development 3: 55 - 63." type="journal article" year="2008">
|
||
Lowry
|
||
<emphasis id="B95CCE580065FFD4645CFD72FB197DE5" box="[1106,1160,732,757]" italics="true" pageId="12" pageNumber="51">et al</emphasis>
|
||
. 2008
|
||
</bibRefCitation>
|
||
). With just two collections from a restricted forest, an AOO of
|
||
<quantity id="4CD0BFAF0065FFD462B3FCABFD667C0F" box="[701,759,773,799]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="51" unit="km" value="9.0">9 km</quantity>
|
||
<superScript id="7C5DBF020065FFD462F9FCACFD6E7C00" attach="left" box="[759,767,770,784]" fontSize="6" pageId="12" pageNumber="51">2</superScript>
|
||
, and a single subpopulation constituting a single location with an estimated number of less than 250 individuals, near or within a protected area (Petriky),
|
||
<taxonomicName id="4C2869C90065FFD4658FFC82FEB17C7C" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="51" phylum="Arthropoda" rank="species" species="stictophylla">
|
||
<emphasis id="B95CCE580065FFD4658FFC82FA027C55" box="[1409,1427,812,837]" italics="true" pageId="12" pageNumber="51">E</emphasis>
|
||
.
|
||
<emphasis id="B95CCE580065FFD46099FCFDFEB17C7C" box="[151,288,851,876]" italics="true" pageId="12" pageNumber="51">stictophylla</emphasis>
|
||
</taxonomicName>
|
||
is assigned a preliminary status of “Endangered” (EN: D).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33241C10065FFD460C8FCD7FD057A06" pageId="12" pageNumber="51" type="discussion">
|
||
<paragraph id="8B97124A0065FFD460C8FCD7FC347B6B" blockId="12.[151,1436,152,1302]" pageId="12" pageNumber="51">
|
||
<emphasis id="B95CCE580065FFD460C8FCD7FEC57C83" bold="true" box="[198,340,889,915]" pageId="12" pageNumber="51">Discussion:</emphasis>
|
||
―Our reports of fruiting material are taken from the field and from information provided by people locally who know the species. Among other species of
|
||
<taxonomicName id="4C2869C90065FFD46391FC0EFB957CA9" box="[927,1028,928,953]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="51" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95CCE580065FFD46391FC0EFB957CA9" box="[927,1028,928,953]" italics="true" pageId="12" pageNumber="51">Eugenia</emphasis>
|
||
</taxonomicName>
|
||
in the vicinity of Petriky Forest,
|
||
<taxonomicName id="4C2869C90065FFD46099FC69FE1B7CF0" box="[151,394,967,992]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="51" phylum="Arthropoda" rank="species" species="stictophylla">
|
||
<emphasis id="B95CCE580065FFD46099FC69FE1B7CF0" box="[151,394,967,992]" italics="true" pageId="12" pageNumber="51">Eugenia stictophylla</emphasis>
|
||
</taxonomicName>
|
||
most closely resembles
|
||
<taxonomicName id="4C2869C90065FFD462A0FC69FBCA7CF1" authority="Lam. However" authorityName="Lam. However" box="[686,1115,967,993]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="51" phylum="Arthropoda" rank="species" species="cassinoides">
|
||
<emphasis id="B95CCE580065FFD462A0FC69FC317CF0" box="[686,928,967,992]" italics="true" pageId="12" pageNumber="51">Eugenia cassinoides</emphasis>
|
||
Lam. However
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2869C90065FFD46466FC69FA837CF0" box="[1128,1298,967,992]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="51" phylum="Arthropoda" rank="species" species="stictophylla">
|
||
<emphasis id="B95CCE580065FFD46466FC69FA837CF0" box="[1128,1298,967,992]" italics="true" pageId="12" pageNumber="51">E. stictophylla</emphasis>
|
||
</taxonomicName>
|
||
has a more deeply sulcate petiole, more narrowly elliptic leaf blades, denser and much darker laminar oil glands (living and dried material), and more flexible pedicles. The leaf blades of the new species tend to dry significantly greener than those of
|
||
<taxonomicName id="4C2869C90065FFD46195FB95FDD47B44" box="[411,581,1083,1108]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="51" phylum="Arthropoda" rank="species" species="cassinoides">
|
||
<emphasis id="B95CCE580065FFD46195FB95FDD47B44" box="[411,581,1083,1108]" italics="true" pageId="12" pageNumber="51">E. cassinoides</emphasis>
|
||
</taxonomicName>
|
||
, the latter of which typically dry darkish brown adaxially, although the dried color may be an artifact of drying temperatures and duration.
|
||
</paragraph>
|
||
<paragraph id="8B97124A0065FFD46099FB26FD057A06" blockId="12.[151,1436,152,1302]" pageId="12" pageNumber="51">
|
||
The
|
||
<typeStatus id="5493ACE80065FFD460C1FB26FEA87BB2" box="[207,313,1160,1186]" pageId="12" pageNumber="51" type="paratype">paratype</typeStatus>
|
||
specimen of
|
||
<taxonomicName id="4C2869C90065FFD461EDFB26FD067BB1" box="[483,663,1160,1185]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="51" phylum="Arthropoda" rank="species" species="stictophylla">
|
||
<emphasis id="B95CCE580065FFD461EDFB26FD067BB1" box="[483,663,1160,1185]" italics="true" pageId="12" pageNumber="51">E. stictophylla</emphasis>
|
||
</taxonomicName>
|
||
may be slightly introgressed with the much more common
|
||
<taxonomicName id="4C2869C90065FFD4658CFB26FE8E7BD8" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="51" phylum="Arthropoda" rank="species" species="cassinoides">
|
||
<emphasis id="B95CCE580065FFD4658CFB26FE8E7BD8" italics="true" pageId="12" pageNumber="51">E. cassinoides</emphasis>
|
||
</taxonomicName>
|
||
, although genetic studies would be necessary to test this hypothesis. Differences of the
|
||
<typeStatus id="5493ACE80065FFD46538FB01FA0A7BD9" box="[1334,1435,1199,1225]" pageId="12" pageNumber="51" type="paratype">paratype</typeStatus>
|
||
from the type gathering include fewer stamens (ca. 35–45 v. 60–70), subglobular (
|
||
<taxonomicName id="4C2869C90065FFD46468FB7BFA937BFF" box="[1126,1282,1237,1263]" pageId="12" pageNumber="51" rank="variety" variety="cylindrical">v. cylindrical</taxonomicName>
|
||
) anther sacs, and fewer (8–9 v. 11–13) ovules per locule.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |