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<document id="3436621F988D804A8AB3F2CC8B2705B0" ID-CLB-Dataset="4939" ID-DOI="10.11646/zootaxa.4984.1.6" ID-GBIF-Dataset="723d2e5b-aa7b-4d6f-84ad-bc45531fa175" ID-ISSN="1175-5326" ID-Zenodo-Dep="4928494" ID-ZooBank="5F2D2119-40F9-4DCE-B4AE-CDE87E44C0EA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623434454951" checkinUser="plazi" docAuthor="Mitov, Plamen G., Perkovsky, Evgeny E. &amp; Dunlop, Jason A." docDate="2021" docId="038106260C7D716318C0FC79FB5F47FA" docLanguage="en" docName="zootaxa.4984.1.6.pdf" docOrigin="Zootaxa 4984 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parahistricostoma tuberculatum Mitov &amp; Perkovsky &amp; Dunlop 2021" docType="treatment" docVersion="7" lastPageNumber="63" masterDocId="FFB87E5E0C6971771857FFD7FFE04267" masterDocTitle="Harvestmen (Arachnida: Opiliones) in Eocene Rovno amber (Ukraine)" masterLastPageNumber="72" masterPageNumber="43" pageNumber="63" updateTime="1699012069377" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FDFAD0152912EAF9EE94E61B6CD23856">Harvestmen (Arachnida: Opiliones) in Eocene Rovno amber (Ukraine)</mods:title>
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<mods:namePart id="C5C710299E95C6ADA9A508582FDA2787">Mitov, Plamen G.</mods:namePart>
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<mods:namePart id="1D6DBE61A638B6BAA1B68F550FCA177B">Perkovsky, Evgeny E.</mods:namePart>
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<mods:namePart id="7C810A0F430F8EBA96042D1CD9B2D4BA">Dunlop, Jason A.</mods:namePart>
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<treatment id="038106260C7D716318C0FC79FB5F47FA" ID-DOI="http://doi.org/10.5281/zenodo.5217786" ID-GBIF-Taxon="184587106" ID-Zenodo-Dep="5217786" LSID="urn:lsid:plazi:treatment:038106260C7D716318C0FC79FB5F47FA" httpUri="http://treatment.plazi.org/id/038106260C7D716318C0FC79FB5F47FA" lastPageNumber="63" pageId="20" pageNumber="63">
<subSubSection id="C332E4BB0C7D716318C0FC79FC5D41AF" box="[151,957,942,969]" pageId="20" pageNumber="63" type="nomenclature">
<paragraph id="8B97B7300C7D716318C0FC79FC5D41AF" blockId="20.[151,957,942,1005]" box="[151,957,942,969]" pageId="20" pageNumber="63">
<heading id="D0DF005C0C7D716318C0FC79FC5D41AF" bold="true" box="[151,957,942,969]" fontSize="11" level="1" pageId="20" pageNumber="63" reason="1">
<emphasis id="B95C6B220C7D716318C0FC79FC5D41AF" bold="true" box="[151,957,942,969]" pageId="20" pageNumber="63">
cf.
<taxonomicName id="4C28CCB30C7D716318EAFC79FC5D41AF" ID-CoL="9X5HY" authority="(C. L. Koch &amp; Berendt, 1854)" authorityName="Mitov &amp; Perkovsky &amp; Dunlop" authorityYear="2021" baseAuthorityName="C. L. Koch &amp; Berendt" baseAuthorityYear="1854" box="[189,957,942,969]" class="Arachnida" family="Nemastomatidae" genus="Parahistricostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="20" pageNumber="63" phylum="Arthropoda" rank="species" species="tuberculatum">
<emphasis id="B95C6B220C7D716318EAFC79FDA541AE" bold="true" box="[189,581,942,969]" italics="true" pageId="20" pageNumber="63">Parahistricostoma tuberculatum</emphasis>
(C. L.
<bibRefCitation id="EFB9CAC10C7D71631ACBFC78FC5541AF" author="Koch, C. L. &amp; Berendt, G. C." box="[668,949,942,969]" pageId="20" pageNumber="63" refId="ref17431" refString="Koch, C. L. &amp; Berendt, G. C. (1854) Die im Bernstein befindlichen Myriapoden, Arachniden und Apteren der Vorwelt. In: Berendt, G. C. (Ed.), Die in Bernstein befindlichen organischen Reste der Vorwelt gesammelt in Verbindung mit mehreren bearbeitetet und herausgegeben 1. Berlin, Nicolai, 124 pp." type="book" year="1854">Koch &amp; Berendt, 1854</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C332E4BB0C7D716318C0FC04FEF5418A" box="[151,277,978,1005]" pageId="20" pageNumber="63" type="description">
<paragraph id="8B97B7300C7D716318C0FC04FEF5418A" blockId="20.[151,957,942,1005]" box="[151,277,978,1005]" pageId="20" pageNumber="63">
<figureCitation id="1313ABB50C7D716318C0FC04FEF5418A" box="[151,277,978,1005]" captionStart="FIGURES 2527" captionStartId="21.[151,264,1790,1814]" captionTargetBox="[151,1435,345,1767]" captionTargetId="figure-95@21.[151,1435,345,1767]" captionTargetPageId="21" captionText="FIGURES 2527. cf. Parahistricostoma tuberculatum (C. L. Koch &amp; Berendt, 1854), juvenile, SIZK UA-1565, from Rovno amber (precise locality unclear), Ukraine; late Eocene. 25 Dorso-lateral overview of the complete animal. 26 Detail of 25, showing anterior prosoma and pedipalps with clavate setae (arrowed), dorso-lateral view. 27 Detail of 26 close up showing palpal segments with clavate setae; note also the head of clavate seta (arrowed), lateral view. Scale bars: 25: 1 mm; 26: 0.2 mm; 27: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4928520" httpUri="https://zenodo.org/record/4928520/files/figure.png" pageId="20" pageNumber="63">Figs 2527</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C332E4BB0C7D716318C0FBCDFEFE463E" pageId="20" pageNumber="63" type="materials_examined">
<paragraph id="8B97B7300C7D716318C0FBCDFEFE463E" blockId="20.[151,1436,1050,1437]" pageId="20" pageNumber="63">
<emphasis id="B95C6B220C7D716318C0FBCDFDC44653" bold="true" box="[151,548,1050,1077]" pageId="20" pageNumber="63">
Material examined..
<collectingCountry id="F33FF7A00C7D716319C1FBCDFDFF4653" box="[406,543,1050,1076]" name="Ukraine" pageId="20" pageNumber="63">UKRAINE</collectingCountry>
:
</emphasis>
<collectionCode id="ED392FF50C7D71631A7CFBCCFD894652" box="[555,617,1051,1077]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="20" pageNumber="63">SIZK</collectionCode>
<collectionCode id="ED392FF50C7D71631A27FBCDFD434652" box="[624,675,1050,1077]" pageId="20" pageNumber="63">UA-</collectionCode>
1565,
<collectingRegion id="49EC79D20C7D71631ABFFBCCFCD34652" box="[744,819,1051,1077]" country="Ukraine" name="Rivne" pageId="20" pageNumber="63">Rovno</collectingRegion>
amber, precise locality unclear; late Eocene. Syninclusions: none.
</paragraph>
</subSubSection>
<subSubSection id="C332E4BB0C7D71631890FBB5FB4D476A" pageId="20" pageNumber="63" type="description">
<paragraph id="8B97B7300C7D71631890FBB5FB4D476A" blockId="20.[151,1436,1050,1437]" pageId="20" pageNumber="63">
<emphasis id="B95C6B220C7D71631890FBB5FEB9461B" bold="true" box="[199,345,1122,1148]" pageId="20" pageNumber="63">Description.</emphasis>
A juvenile covered by a white emulsion concealing the dorsal granulation and the armature of the scutum (
<figureCitation id="1313ABB50C7D716318AFFB50FEAD46C6" box="[248,333,1158,1185]" captionStart="FIGURES 2527" captionStartId="21.[151,264,1790,1814]" captionTargetBox="[151,1435,345,1767]" captionTargetId="figure-95@21.[151,1435,345,1767]" captionTargetPageId="21" captionText="FIGURES 2527. cf. Parahistricostoma tuberculatum (C. L. Koch &amp; Berendt, 1854), juvenile, SIZK UA-1565, from Rovno amber (precise locality unclear), Ukraine; late Eocene. 25 Dorso-lateral overview of the complete animal. 26 Detail of 25, showing anterior prosoma and pedipalps with clavate setae (arrowed), dorso-lateral view. 27 Detail of 26 close up showing palpal segments with clavate setae; note also the head of clavate seta (arrowed), lateral view. Scale bars: 25: 1 mm; 26: 0.2 mm; 27: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4928520" httpUri="https://zenodo.org/record/4928520/files/figure.png" pageId="20" pageNumber="63">Fig. 25</figureCitation>
). With L: 0.50; prosoma width: 0.27; abdomen width: 0.28; ocular tubercle elliptic (
<figureCitation id="1313ABB50C7D71631D55FB50FAB546C5" box="[1282,1365,1159,1186]" captionStart="FIGURES 2527" captionStartId="21.[151,264,1790,1814]" captionTargetBox="[151,1435,345,1767]" captionTargetId="figure-95@21.[151,1435,345,1767]" captionTargetPageId="21" captionText="FIGURES 2527. cf. Parahistricostoma tuberculatum (C. L. Koch &amp; Berendt, 1854), juvenile, SIZK UA-1565, from Rovno amber (precise locality unclear), Ukraine; late Eocene. 25 Dorso-lateral overview of the complete animal. 26 Detail of 25, showing anterior prosoma and pedipalps with clavate setae (arrowed), dorso-lateral view. 27 Detail of 26 close up showing palpal segments with clavate setae; note also the head of clavate seta (arrowed), lateral view. Scale bars: 25: 1 mm; 26: 0.2 mm; 27: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4928520" httpUri="https://zenodo.org/record/4928520/files/figure.png" pageId="20" pageNumber="63">Fig. 26</figureCitation>
), with length: 0.06 and width: 0.08; diameter of each lens 0.03; Palps with article lengths: Fe: 0.308; Pa: 0.238; Ti: 0.238; Ta: 0.14. Palpal articles (except trochanters) ventrally with clavate setae (L: 0.056) (see
<figureCitation id="1313ABB50C7D71631C22FB18FB17468D" box="[1141,1271,1231,1258]" captionStart="FIGURES 2527" captionStartId="21.[151,264,1790,1814]" captionTargetBox="[151,1435,345,1767]" captionTargetId="figure-95@21.[151,1435,345,1767]" captionTargetPageId="21" captionText="FIGURES 2527. cf. Parahistricostoma tuberculatum (C. L. Koch &amp; Berendt, 1854), juvenile, SIZK UA-1565, from Rovno amber (precise locality unclear), Ukraine; late Eocene. 25 Dorso-lateral overview of the complete animal. 26 Detail of 25, showing anterior prosoma and pedipalps with clavate setae (arrowed), dorso-lateral view. 27 Detail of 26 close up showing palpal segments with clavate setae; note also the head of clavate seta (arrowed), lateral view. Scale bars: 25: 1 mm; 26: 0.2 mm; 27: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4928520" httpUri="https://zenodo.org/record/4928520/files/figure.png" pageId="20" pageNumber="63">Figs 2627</figureCitation>
). The tarsus of each leg is one-segmented (
<figureCitation id="1313ABB50C7D7163199AFB24FDFF476A" box="[461,543,1266,1293]" captionStart="FIGURES 2527" captionStartId="21.[151,264,1790,1814]" captionTargetBox="[151,1435,345,1767]" captionTargetId="figure-95@21.[151,1435,345,1767]" captionTargetPageId="21" captionText="FIGURES 2527. cf. Parahistricostoma tuberculatum (C. L. Koch &amp; Berendt, 1854), juvenile, SIZK UA-1565, from Rovno amber (precise locality unclear), Ukraine; late Eocene. 25 Dorso-lateral overview of the complete animal. 26 Detail of 25, showing anterior prosoma and pedipalps with clavate setae (arrowed), dorso-lateral view. 27 Detail of 26 close up showing palpal segments with clavate setae; note also the head of clavate seta (arrowed), lateral view. Scale bars: 25: 1 mm; 26: 0.2 mm; 27: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4928520" httpUri="https://zenodo.org/record/4928520/files/figure.png" pageId="20" pageNumber="63">Fig. 25</figureCitation>
) with a length of: Ta I: 0.28; Ta II.IV.: 0.25, respectively.
</paragraph>
</subSubSection>
<subSubSection id="C332E4BB0C7D71631890FAC0FB5F47FA" pageId="20" pageNumber="63" type="discussion">
<paragraph id="8B97B7300C7D71631890FAC0FB5F47FA" blockId="20.[151,1436,1050,1437]" pageId="20" pageNumber="63">
<emphasis id="B95C6B220C7D71631890FAC0FEDB4756" bold="true" box="[199,315,1303,1329]" pageId="20" pageNumber="63">Remarks.</emphasis>
Given the shape of the ocular tubercle, the proportions of the pedipalps, and the shape of their segments, this juvenile most probably belongs to
<taxonomicName id="4C28CCB30C7D71631AF9FAECFBFC4732" authorityName="Mitov &amp; Perkovsky &amp; Dunlop" authorityYear="2021" baseAuthorityName="C. L. Koch &amp; Berendt" baseAuthorityYear="1854" box="[686,1052,1339,1365]" class="Arachnida" family="Nemastomatidae" genus="Parahistricostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="20" pageNumber="63" phylum="Arthropoda" rank="species" species="tuberculatum">
<emphasis id="B95C6B220C7D71631AF9FAECFBFC4732" box="[686,1052,1339,1365]" italics="true" pageId="20" pageNumber="63">Parahistricostoma tuberculatum</emphasis>
</taxonomicName>
, too. The length of the body, the single tarsal segments and their length, indicate that this juvenile nemastomatid, by analogy with other extant nemastomatid species, could be the first or the second juvenile stage (cf.
<bibRefCitation id="EFB9CAC10C7D71631BF8FA54FBDB47FA" author="Immel, V." box="[943,1083,1410,1437]" pageId="20" pageNumber="63" pagination="129 - 184" refId="ref16911" refString="Immel, V. (1954) Zur Biologie und Physiologie von Nemastoma quadripunctatum (Opiliones, Dyspnoi). Zoologisches Jahrbuch, Abteilung fur Systematik, 83, 129 - 184." type="journal article" year="1954">Immel 1954</bibRefCitation>
: figs 8, 10).
</paragraph>
</subSubSection>
</treatment>
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