207 lines
26 KiB
XML
207 lines
26 KiB
XML
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<mods:title id="8716D16127C3F5ADAEAC0A2F6EBDDD73">Harvestmen (Arachnida: Opiliones) in Eocene Rovno amber (Ukraine)</mods:title>
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<mods:namePart id="DF2415022BFDA70C81C3602069AA9060">Mitov, Plamen G.</mods:namePart>
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<mods:namePart id="3295AEBA803A5910A46B41280B2AAE13">Perkovsky, Evgeny E.</mods:namePart>
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<mods:namePart id="A0A1A044476C319274B4912A82D7DA43">Dunlop, Jason A.</mods:namePart>
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<mods:nameIdentifier id="8DF83BF11179F39364161DFEFBBD284B" type="email">jason.dunlop@mfn.berlin</mods:nameIdentifier>
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<mods:date id="A416D54610E20BE47AE87F2D591BB7F9">2021</mods:date>
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<mods:number id="E7881A035962C8984C68C7555CBE7527">2021-06-10</mods:number>
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<treatment id="038106260C79716518C0F9FBFD46415E" ID-DOI="http://doi.org/10.5281/zenodo.4928224" ID-GBIF-Taxon="184587104" ID-Zenodo-Dep="4928224" LSID="urn:lsid:plazi:treatment:038106260C79716518C0F9FBFD46415E" httpUri="http://treatment.plazi.org/id/038106260C79716518C0F9FBFD46415E" lastPageId="18" lastPageNumber="61" pageId="16" pageNumber="59">
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<subSubSection id="C332E4BB0C79716718C0F9FBFDC64421" box="[151,550,1579,1606]" pageId="16" pageNumber="59" type="nomenclature">
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<paragraph id="8B97B7300C79716718C0F9FBFDC64421" blockId="16.[151,550,1579,1606]" box="[151,550,1579,1606]" pageId="16" pageNumber="59">
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<heading id="D0DF005C0C79716718C0F9FBFDC64421" bold="true" box="[151,550,1579,1606]" fontSize="11" level="1" pageId="16" pageNumber="59" reason="1">
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<emphasis id="B95C6B220C79716718C0F9FBFDC64421" bold="true" box="[151,550,1579,1606]" pageId="16" pageNumber="59">
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Genus
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<taxonomicName id="4C28CCB30C79716718B9F9FCFE2C4421" ID-CoL="9X5HX" authority="Mitov & Perkovsky & Dunlop, 2021" authorityName="Mitov & Perkovsky & Dunlop" authorityYear="2021" box="[238,460,1579,1606]" class="Arachnida" family="Nemastomatidae" genus="Parahistricostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="59" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B95C6B220C79716718B9F9FCFE2C4421" bold="true" box="[238,460,1579,1606]" italics="true" pageId="16" pageNumber="59">Parahistricostoma</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26FD6590C7971671984F9FBFDC64421" box="[467,550,1580,1606]" pageId="16" pageNumber="59" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C332E4BB0C79716718C0F9A3FAB345ED" pageId="16" pageNumber="59" type="diagnosis">
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<paragraph id="8B97B7300C79716718C0F9A3FAB345ED" blockId="16.[151,1436,1652,2039]" pageId="16" pageNumber="59">
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<emphasis id="B95C6B220C79716718C0F9A3FEEA44E9" bold="true" box="[151,266,1652,1678]" pageId="16" pageNumber="59">Diagnosis</emphasis>
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. Small, dark fossil nemastomatids (body length range 1.25–1.52, body width ca. 1.0) with ovoid to quadrangular body, dorsally with fine granulation, only anterior corners of scutum magnum with more prominent granules, and with four pairs of slender, pillar-like spines, each slightly bulbous at the top, standing perpendicular to the body surface. In males, apophyses resembling an ‘ice-cream scoop’ (or in dorso/dorso-lateral outline, boxing gloves) present on first cheliceral articles; apophysis approx. low, from 0.11 to ca.
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<quantity id="4CD01AD50C7971671C7CF8D3FB644578" box="[1067,1156,1796,1823]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="59" unit="mm" value="0.2">0.2 mm</quantity>
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. Pedipalps and legs elongate and slender. Pedipalps without apophyses (spurs/thorns/denticles). Palpal tarsus relatively short, in lateral view machete-like – distally rounded and slightly widened; in ventral view – like a baseball bat. Palpal Ta/Ti between 0.44 and
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<quantity id="4CD01AD50C79716718A8F8A7FE8645EC" box="[255,358,1904,1931]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="16" pageNumber="59" unit="mm" value="0.55">0.55 mm</quantity>
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. Microsculpture elements of the leg femurs include loose-textured denticles and granules.
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</paragraph>
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</subSubSection>
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<subSubSection id="C332E4BB0C7971671890F843FB0645C9" box="[199,1254,1940,1967]" pageId="16" pageNumber="59" type="type_taxon">
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<paragraph id="8B97B7300C7971671890F843FB0645C9" blockId="16.[151,1436,1652,2039]" box="[199,1254,1940,1967]" pageId="16" pageNumber="59">
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<emphasis id="B95C6B220C7971671890F843FE8245C9" bold="true" box="[199,354,1940,1966]" pageId="16" pageNumber="59">
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<typeStatus id="549309920C7971671890F843FEE245C9" box="[199,258,1940,1966]" pageId="16" pageNumber="59">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C28CCB30C797167193EF842FC3245C8" ID-CoL="9WXW8" authority="C. L. Koch & Berendt, 1854" authorityName="C. L. Koch & Berendt" authorityYear="1854" box="[361,978,1940,1967]" class="Arachnida" family="Nemastomatidae" genus="Nemastoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="59" phylum="Arthropoda" rank="species" species="tuberculatum">
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<emphasis id="B95C6B220C797167193EF842FD6845C9" box="[361,648,1940,1966]" italics="true" pageId="16" pageNumber="59">Nemastoma tuberculatum</emphasis>
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C. L.
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<bibRefCitation id="EFB9CAC10C7971671A98F843FC3245C8" author="Koch, C. L. & Berendt, G. C." box="[719,978,1940,1967]" pageId="16" pageNumber="59" refId="ref17431" refString="Koch, C. L. & Berendt, G. C. (1854) Die im Bernstein befindlichen Myriapoden, Arachniden und Apteren der Vorwelt. In: Berendt, G. C. (Ed.), Die in Bernstein befindlichen organischen Reste der Vorwelt gesammelt in Verbindung mit mehreren bearbeitetet und herausgegeben 1. Berlin, Nicolai, 124 pp." type="book" year="1854">Koch & Berendt, 1854</bibRefCitation>
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</taxonomicName>
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, by original designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C332E4BB0C7971671890F86FFD944590" pageId="16" pageNumber="59" type="etymology">
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<paragraph id="8B97B7300C7971671890F86FFD944590" blockId="16.[151,1436,1652,2039]" pageId="16" pageNumber="59">
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<emphasis id="B95C6B220C7971671890F86FFEA845B5" bold="true" box="[199,328,1976,2002]" pageId="16" pageNumber="59">Etymology</emphasis>
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. From the Greek ‘para’ (beside, near) and the modern harvestman genus
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<taxonomicName id="4C28CCB30C7971671CD5F86EFAC145B5" box="[1154,1313,1977,2002]" class="Arachnida" family="Nemastomatidae" genus="Histricostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="59" phylum="Arthropoda" rank="genus">
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<emphasis id="B95C6B220C7971671CD5F86EFAC145B5" box="[1154,1313,1977,2002]" italics="true" pageId="16" pageNumber="59">Histricostoma</emphasis>
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</taxonomicName>
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which this material closely resembles. Gender neuter.
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</paragraph>
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||
</subSubSection>
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||
<caption id="DF57E7B80C78716618C0F8F7FD7D458B" ID-DOI="http://doi.org/10.5281/zenodo.4928516" ID-Zenodo-Dep="4928516" httpUri="https://zenodo.org/record/4928516/files/figure.png" pageId="17" pageNumber="60" startId="17.[151,264,1824,1848]" targetBox="[243,1344,184,1799]" targetPageId="17">
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<paragraph id="8B97B7300C78716618C0F8F7FD7D458B" blockId="17.[151,1437,1824,2029]" pageId="17" pageNumber="60">
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<emphasis id="B95C6B220C78716618C0F8F7FEB4455F" bold="true" box="[151,340,1824,1848]" pageId="17" pageNumber="60">FIGURES 18–21.</emphasis>
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<taxonomicName id="4C28CCB30C787166190CF8F6FD4B455F" authority="(Menge, 1854)" baseAuthorityName="Menge" baseAuthorityYear="1854" box="[347,683,1824,1849]" class="Arachnida" family="Sabaconidae" genus="Sabacon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="17" pageNumber="60" phylum="Arthropoda" rank="species" species="claviger">
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<emphasis id="B95C6B220C787166190CF8F6FDEB455E" box="[347,523,1825,1849]" italics="true" pageId="17" pageNumber="60">Sabacon claviger</emphasis>
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(
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<bibRefCitation id="EFB9CAC10C7871661A4EF8F6FD44455F" author="Menge, A." box="[537,676,1824,1849]" pageId="17" pageNumber="60" pagination="1 - 124" refId="ref18617" refString="Menge, A. (1854) Footnotes. In: Koch, C. L. & Berendt, G. C., Die im Bernstein befindlichen Myriapoden, Arachniden und Apteren der Vorwelt. In: Berendt, G. C. (Ed.), Die in Bernstein befindlichen organischen Reste der Vorwelt gesammelt in Verbindung mit mehreren bearbeitetet und herausgegeben 1. Berlin, Nicolai, pp. 1 - 124." type="book chapter" year="1854">Menge, 1854</bibRefCitation>
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)
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</taxonomicName>
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, SIZK K-7898, from Rovno amber (Klesov, Pugach quarry, Ukraine), late Eocene. 18 Detail of the body and limbs, dorsal view. 19 Detail of the body and limbs (view from back), showing the opisthosomal dorsum ornamented by regular rows of black thorns (arrowed). 20 Detail of the left pedipalp in ventro-ectal view, showing tibia and tarsus throughout with dense brush-like setae and the tibial long seta (arrowed). Part of the palpal setae are not visible due to amber aberrations—compare with Fig. 21. 21 Detail of the right pedipalp, showing the tibia and tarsus, closer ectal view. Scale bars: 18–19: 1 mm; 20: 0.5 mm; 21: 0.3 mm.
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</paragraph>
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</caption>
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<subSubSection id="C332E4BB0C7B71651890FF40FD46415E" pageId="18" pageNumber="61" type="discussion">
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<paragraph id="8B97B7300C7B71651890FF40FDE343B6" blockId="18.[151,1437,151,825]" pageId="18" pageNumber="61">
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<emphasis id="B95C6B220C7B71651890FF40FED442D6" bold="true" box="[199,308,151,177]" pageId="18" pageNumber="61">Remarks</emphasis>
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. The extinct genus
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<taxonomicName id="4C28CCB30C7B71651A42FF40FD0542D6" authorityName="Mitov & Perkovsky & Dunlop" authorityYear="2021" box="[533,741,151,177]" class="Arachnida" family="Nemastomatidae" genus="Parahistricostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="18" pageNumber="61" phylum="Arthropoda" rank="genus">
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<emphasis id="B95C6B220C7B71651A42FF40FD0542D6" box="[533,741,151,177]" italics="true" pageId="18" pageNumber="61">Parahistricostoma</emphasis>
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</taxonomicName>
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is morphologically closest to the recent genus
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<taxonomicName id="4C28CCB30C7B71651CA9FF40FEF242B2" authority="Kratochvil" authorityName="Kratochvil" class="Arachnida" family="Nemastomatidae" genus="Histricostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="18" pageNumber="61" phylum="Arthropoda" rank="genus">
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<emphasis id="B95C6B220C7B71651CA9FF40FA7D42D7" box="[1278,1437,151,176]" italics="true" pageId="18" pageNumber="61">Histricostoma</emphasis>
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Kratochvíl
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</taxonomicName>
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, which can be found today from the Alps through to the Balkans,
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<collectingCountry id="F33FF7A00C7B71651BBCFF6CFBDB42B2" box="[1003,1083,187,213]" name="Turkey" pageId="18" pageNumber="61">Turkey</collectingCountry>
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and the Caucasus, and includes eight species (
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<bibRefCitation id="EFB9CAC10C7B71651964FF08FE0D429E" author="Kratochvil, J." box="[307,493,222,249]" pageId="18" pageNumber="61" pagination="523 - 576" refId="ref17382" refString="Kratochvil, J. (1958) Hohlenweberknechte Bulgariens (Palpatores - Nemastomatidae) [Jeskynni sekaci Bulharska (Palpatores - Nemastomatidae)]. Prace Brnenske zakladny Ceskoslovenske akademie ved (Acta Academiae Scientiarum Cechoslovenicae Basis Brunensis), 30 (12), 523 - 576." type="journal article" year="1958">Kratochvíl 1958</bibRefCitation>
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;
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<bibRefCitation id="EFB9CAC10C7B716519AFFF08FD7B429E" author="Starega, W." box="[504,667,223,249]" pageId="18" pageNumber="61" pagination="287 - 433" refId="ref21462" refString="Starega, W. (1976 b) Die Weberknechte (Opiliones, excl. Sironidae) Bulgariens. Annales Zoologici [Polska Akademia Nauk], 33 (18), 287 - 433." type="journal article" year="1976">Staręga 1976b</bibRefCitation>
|
||
;
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||
<bibRefCitation id="EFB9CAC10C7B71651AF2FF08FCA2429E" author="Martens, J." box="[677,834,223,249]" pageId="18" pageNumber="61" refId="ref18365" refString="Martens, J. (1978) Spinnentiere, Arachnida: Weberknechte, Opiliones. Die Tierwelt Deutschlands. Vol. 64. G. Fischer Verlag, Jena. 464 pp." type="book" year="1978">Martens 1978</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFB9CAC10C7B71651B1CFF08FC67429E" author="Martens, J." box="[843,903,223,249]" pageId="18" pageNumber="61" pagination="145 - 210" refId="ref18429" refString="Martens, J. (2006) Weberknechte aus dem Kaukasus (Arachnida, Opiliones, Nemastomatidae). Senckenbergiana biologica, 86, 145 - 210." type="journal article" year="2006">2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB9CAC10C7B71651BC6FF08FB2C429E" author="Snegovaya, N. Y. & Marusik, Y. M." box="[913,1228,223,249]" pageId="18" pageNumber="61" pagination="59 - 70" refId="ref21092" refString="Snegovaya, N. Y. & Marusik, Y. M. (2012) New species and collections of Opiliones (Arachnida) from Turkey. Acta Arachnologica, 61 (2), 59 - 70. https: // doi. org / 10.2476 / asjaa. 61.59" type="journal article" year="2012">Snegovaya & Marusik 2012</bibRefCitation>
|
||
; Schönhofer 2013;
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||
<bibRefCitation id="EFB9CAC10C7B716518C0FED4FE6C437A" author="Iorio, E. & Delfoss, E." box="[151,396,258,285]" pageId="18" pageNumber="61" pagination="33 - 40" refId="ref16944" refString="Iorio, E. & Delfoss, E. (2015) Decouverte d'un opilion remarquable dans l'Ouest de la France: Histricostoma argenteolunulatum (Canestrini, 1875) (Opiliones, Nemastomatidae). Invertebres Armoricains, 12, 33 - 40." type="journal article" year="2015">Iorio & Delfoss 2015</bibRefCitation>
|
||
). The traditional assignment of
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||
<bibRefCitation id="EFB9CAC10C7B71651AB8FED4FBC0437A" author="Koch, C. L. & Berendt, G. C." box="[751,1056,258,285]" pageId="18" pageNumber="61" refId="ref17431" refString="Koch, C. L. & Berendt, G. C. (1854) Die im Bernstein befindlichen Myriapoden, Arachniden und Apteren der Vorwelt. In: Berendt, G. C. (Ed.), Die in Bernstein befindlichen organischen Reste der Vorwelt gesammelt in Verbindung mit mehreren bearbeitetet und herausgegeben 1. Berlin, Nicolai, 124 pp." type="book" year="1854">Koch and Berendt’s (1854)</bibRefCitation>
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amber species
|
||
<taxonomicName id="4C28CCB30C7B71651C9AFED4FF114326" authorityName="C. L. Koch & Berendt" authorityYear="1854" class="Arachnida" family="Nemastomatidae" genus="Nemastoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="18" pageNumber="61" phylum="Arthropoda" rank="species" species="tuberculatum">
|
||
<emphasis id="B95C6B220C7B71651C9AFED4FF114326" italics="true" pageId="18" pageNumber="61">Nemastoma tuberculatum</emphasis>
|
||
</taxonomicName>
|
||
to the genus
|
||
<taxonomicName id="4C28CCB30C7B716519D1FEF0FDC54327" box="[390,549,295,320]" class="Arachnida" family="Nemastomatidae" genus="Histricostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="18" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95C6B220C7B716519D1FEF0FDC54327" box="[390,549,295,320]" italics="true" pageId="18" pageNumber="61">Histricostoma</emphasis>
|
||
</taxonomicName>
|
||
was initially based on the presence of false articulations on the leg femora (see
|
||
<bibRefCitation id="EFB9CAC10C7B716518C0FE9CFE914301" author="Starega, W." box="[151,369,331,358]" pageId="18" pageNumber="61" pagination="601 - 604" refId="ref21542" refString="Starega, W. (2002) Baltic amber harvestmen (Opiliones) from Polish collections. Annales Zoologici [Polska Akademia Nauk], 52, 601 - 604." type="journal article" year="2002">Staręga, 2002: 602</bibRefCitation>
|
||
), and later largely based on its dorsal ornament (
|
||
<bibRefCitation id="EFB9CAC10C7B71651BF7FE9CFBDA4302" author="Dunlop, J. A." box="[928,1082,331,357]" pageId="18" pageNumber="61" pagination="167 - 182" refId="ref15734" refString="Dunlop, J. A. (2006) Baltic amber harvestman types (Arachnida: Opiliones: Eupnoi and Dyspnoi). Fossil Record, 9, 167 - 182. https: // doi. org / 10.1002 / mmng. 200600006" type="journal article" year="2006">Dunlop 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB9CAC10C7B71651C10FE9CFAAC4302" author="Dunlop, J. A. & Mitov, P. G." box="[1095,1356,331,357]" pageId="18" pageNumber="61" pagination="347 - 375" refId="ref15864" refString="Dunlop, J. A. & Mitov, P. G. (2009) Fossil harvestmen (Arachnida, Opiliones) from Bitterfeld amber. ZooKeys, 16, 347 - 375. https: // doi. org / 10.3897 / zookeys. 16.224" type="journal article" year="2009">Dunlop & Mitov 2009</bibRefCitation>
|
||
). As in
|
||
<taxonomicName id="4C28CCB30C7B716518C0FEB8FED643EF" box="[151,310,367,392]" class="Arachnida" family="Nemastomatidae" genus="Histricostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="18" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95C6B220C7B716518C0FEB8FED643EF" box="[151,310,367,392]" italics="true" pageId="18" pageNumber="61">Histricostoma</emphasis>
|
||
</taxonomicName>
|
||
, four pairs of slender, pillar-like spines, each slightly bulbous at the top (“Stabdorne”
|
||
<emphasis id="B95C6B220C7B71651CACFEB8FAD943EF" box="[1275,1337,367,392]" italics="true" pageId="18" pageNumber="61">sensu</emphasis>
|
||
<bibRefCitation id="EFB9CAC10C7B71651D17FEB8FF3043CA" author="Martens, J." pageId="18" pageNumber="61" refId="ref18365" refString="Martens, J. (1978) Spinnentiere, Arachnida: Weberknechte, Opiliones. Die Tierwelt Deutschlands. Vol. 64. G. Fischer Verlag, Jena. 464 pp." type="book" year="1978">Martens 1978</bibRefCitation>
|
||
) are observed. The distal end of the pedipalp femur in both the fossils and living
|
||
<taxonomicName id="4C28CCB30C7B71651C0CFE44FB1A43CB" box="[1115,1274,403,428]" class="Arachnida" family="Nemastomatidae" genus="Histricostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="18" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95C6B220C7B71651C0CFE44FB1A43CB" box="[1115,1274,403,428]" italics="true" pageId="18" pageNumber="61">Histricostoma</emphasis>
|
||
</taxonomicName>
|
||
is similar, and the leg trochanters are spinulate.
|
||
</paragraph>
|
||
<paragraph id="8B97B7300C7B71651890FE0CFD46415E" blockId="18.[151,1437,151,825]" pageId="18" pageNumber="61">
|
||
However, the question remains do these features alone justify placing the fossil in the modern genus? We could identify at least four subtle differences between the
|
||
<taxonomicName id="4C28CCB30C7B71651AB2FE28FC64407F" box="[741,900,511,536]" class="Arachnida" family="Nemastomatidae" genus="Histricostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="18" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95C6B220C7B71651AB2FE28FC64407F" box="[741,900,511,536]" italics="true" pageId="18" pageNumber="61">Histricostoma</emphasis>
|
||
</taxonomicName>
|
||
-like fossils and modern
|
||
<taxonomicName id="4C28CCB30C7B71651CCAFE28FA7D407E" box="[1181,1437,511,537]" class="Arachnida" family="Nemastomatidae" genus="Histricostoma" kingdom="Animalia" order="Opiliones" pageId="18" pageNumber="61" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B95C6B220C7B71651CCAFE28FADC407F" box="[1181,1340,511,536]" italics="true" pageId="18" pageNumber="61">Histricostoma</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
First, the form of the cheliceral apophysis in the fossils differs from that seen in recent species. Second, there is no evidence for a medial spur (thorn/denticle) on the distal end of the male palpal patella, as is seen in all modern
|
||
<taxonomicName id="4C28CCB30C7B716518C0FDBCFED640E3" box="[151,310,619,644]" class="Arachnida" family="Nemastomatidae" genus="Histricostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="18" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95C6B220C7B716518C0FDBCFED640E3" box="[151,310,619,644]" italics="true" pageId="18" pageNumber="61">Histricostoma</emphasis>
|
||
</taxonomicName>
|
||
. Third, the legs and pedipalps of the fossils are somewhat longer and more slender than the appendages observed in living species. Finally, the microsculptural elements of the leg femora are also different compared to those in the
|
||
<typeStatus id="549309920C7B71651916FD64FE9140AA" box="[321,369,691,717]" pageId="18" pageNumber="61">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="4C28CCB30C7B716519A4FD64FD7240AB" box="[499,658,691,716]" class="Arachnida" family="Nemastomatidae" genus="Histricostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="18" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95C6B220C7B716519A4FD64FD7240AB" box="[499,658,691,716]" italics="true" pageId="18" pageNumber="61">Histricostoma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C28CCB30C7B71651AF4FD64FCFF40AA" authorityName="Kratochvil" authorityYear="1958" box="[675,799,691,717]" class="Arachnida" family="Nemastomatidae" genus="Histricostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="18" pageNumber="61" phylum="Arthropoda" rank="species" species="drenskii">
|
||
<emphasis id="B95C6B220C7B71651AF4FD64FCFF40AA" box="[675,799,691,717]" italics="true" pageId="18" pageNumber="61">H. drenskii</emphasis>
|
||
</taxonomicName>
|
||
), as well to those of
|
||
<taxonomicName id="4C28CCB30C7B71651C45FD64FB7640AB" baseAuthorityName="Roewer" baseAuthorityYear="1951" box="[1042,1174,691,716]" class="Arachnida" family="Nemastomatidae" genus="Histricostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="18" pageNumber="61" phylum="Arthropoda" rank="species" species="creticum">
|
||
<emphasis id="B95C6B220C7B71651C45FD64FB7640AB" box="[1042,1174,691,716]" italics="true" pageId="18" pageNumber="61">H. creticum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C28CCB30C7B71651C98FD64FA8940AA" baseAuthorityName="Ausserer" baseAuthorityYear="1867" box="[1231,1385,691,717]" class="Arachnida" family="Nemastomatidae" genus="Histricostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="18" pageNumber="61" phylum="Arthropoda" rank="species" species="dentipalpe">
|
||
<emphasis id="B95C6B220C7B71651C98FD64FA8940AA" box="[1231,1385,691,717]" italics="true" pageId="18" pageNumber="61">H. dentipalpe</emphasis>
|
||
</taxonomicName>
|
||
. We feel that this character combination does merit a separate genus, diagnosed as above. Furthermore, lacking any data on the structure of the penis in this fossil species, we cannot resolve its generic relationships through comparing its genital morphology with that of extant species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |