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<document id="F8D26D7E40D408657DA71BD354EBDE22" ID-DOI="10.5281/zenodo.173679" ID-GBIF-Dataset="b3ebcdfb-29ea-4c1d-86a0-4a163165d5c0" ID-ISSN="1175­5326" ID-Zenodo-Dep="173679" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459782727286" checkinUser="plazi" docAuthor="Ahyong, Shane T. &amp; Dawson, Elliot W." docDate="2006" docId="A96987DC2628FFF72001F963FB07FD47" docLanguage="en" docName="zt01303p068.pdf" docOrigin="Zootaxa 1303" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Neolithodes yaldwyni Ahyong &amp; Dawson, 2006, sp. nov." docType="treatment" docVersion="9" lastPageNumber="55" masterDocId="5550FFA42629FFFD2109FF92FFF6FFD9" masterDocTitle="Lithodidae from the Ross Sea, Antarctica, with descriptions of two new species (Crustacea: Decapoda: Anomura)" masterLastPageNumber="68" masterPageNumber="45" pageNumber="46" updateTime="1698218021192" updateUser="plazi">
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<mods:title id="9147A5D1B97C942AD219A98EC4FC7E5E">Lithodidae from the Ross Sea, Antarctica, with descriptions of two new species (Crustacea: Decapoda: Anomura)</mods:title>
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<mods:namePart id="802DA0B1A3D94C60E90CAEB77F6612B2">Ahyong, Shane T.</mods:namePart>
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<subSubSection id="69DA65412628FFFC2001F963FD0FF8D2" box="[264,761,1777,1803]" pageId="1" pageNumber="46" type="nomenclature">
<paragraph id="217F36CA2628FFFC2001F963FD0FF8D2" blockId="1.[264,761,1777,1803]" box="[264,761,1777,1803]" pageId="1" pageNumber="46">
<heading id="7A3781A62628FFFC2001F963FD0FF8D2" bold="true" box="[264,761,1777,1803]" fontSize="11" level="1" pageId="1" pageNumber="46" reason="1">
<emphasis id="13B4EAD82628FFFC2001F963FD0FF8D2" bold="true" box="[264,761,1777,1803]" pageId="1" pageNumber="46">
<taxonomicName id="E6C04D492628FFFC2001F963FDFFF8D2" ID-CoL="75f1870c-a4aa-418b-9055-7b223eee038f" box="[264,521,1777,1803]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="yaldwyni" status="sp. nov.">
<emphasis id="13B4EAD82628FFFC2001F963FDFFF8D2" bold="true" box="[264,521,1777,1803]" italics="true" pageId="1" pageNumber="46">Neolithodes yaldwyni</emphasis>
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<taxonomicNameLabel id="088757A32628FFFC231BF963FD99F8D2" box="[530,623,1777,1803]" pageId="1" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="B9FB2A4F2628FFFC2376F963FD19F8D2" box="[639,751,1777,1803]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[264,367,1845,1869]" captionStartId-1="4.[264,367,1536,1560]" captionStartId-2="5.[264,367,1767,1791]" captionStartId-3="6.[264,367,1809,1833]" captionTargetBox-0="[264,1323,329,1817]" captionTargetBox-1="[264,1323,315,1512]" captionTargetBox-2="[264,1323,591,1755]" captionTargetBox-3="[264,1323,326,1785]" captionTargetId-0="figure@3.[264,1323,329,1818]" captionTargetId-1="figure@4.[264,1323,315,1512]" captionTargetId-2="figure@5.[264,1323,591,1758]" captionTargetId-3="figure@6.[264,1323,326,1785]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Neolithodes yaldwyni sp. nov., male holotype (cl 141.5 mm, cw 124.1 mm, NIWA 3434). A, overall dorsal view. B, carapace, dorsal view. C, anterior carapace, right lateral view." captionText-1="FIGURE 2. Neolithodes yaldwyni sp. nov., male holotype (cl 141.5 mm, cw 124.1 mm, NIWA 3434). A, carapace, right lateral view. B, left antenna, dorsal view. C, ventral surface. D, posterior carapace and abdomen, posterior view. E, coxa and basis of left third ambulatory leg, ventral view. F, rostrum and anterior carapace, dorsal view." captionText-2="FIGURE 3. Neolithodes yaldwyni sp. nov. A D, male holotype (cl 141.5 mm, cw 124.1 mm, NIWA 3434). E F, male paratype (cl 120.0 mm, cw 108.7 mm, NIWA 3433). A, left cheliped, dorsal view. B, left chela, outer surface. C, right cheliped, dorsal view. D, right chela, outer surface. E, left chela, outer surface. F, right chela, outer surface." captionText-3="FIGURE 4. Neolithodes yaldwyni sp. nov., overall view, dorsal. A, male paratype (cl 101.7 mm, cw 91.6 mm, NMNZ CR 11002). B, male paratype (cl 136.6 mm, cw 131.1 mm, NMNZ CR 11003)." httpUri-0="https://zenodo.org/record/173680/files/figure.png" httpUri-1="https://zenodo.org/record/173681/files/figure.png" httpUri-2="https://zenodo.org/record/173682/files/figure.png" httpUri-3="https://zenodo.org/record/173683/files/figure.png" pageId="1" pageNumber="46">Figs. 14</figureCitation>
)
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<paragraph id="217F36CA2628FFFC2001F8A5FD64F8B4" blockId="1.[264,1322,1846,1902]" pageId="1" pageNumber="46">
<treatmentCitationGroup id="01D011E42628FFFC2001F8A5FD64F8B4" pageId="1" pageNumber="46">
<taxonomicName id="E6C04D492628FFFC2001F8A5FE2CF897" ID-CoL="e63447e8-fa4b-4c65-af30-a7c90fd6231f" box="[264,474,1847,1870]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="brodiei">
<emphasis id="13B4EAD82628FFFC2001F8A5FE2CF897" box="[264,474,1847,1870]" italics="true" pageId="1" pageNumber="46">Neolithodes brodiei</emphasis>
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.—
<treatmentCitation id="A06110DB2628FFFC20F0F8A4FCFBF894" author="Thatje" box="[505,781,1846,1869]" page="335" pageId="1" pageNumber="46" year="2005">
<bibRefCitation id="45514B3B2628FFFC20F0F8A4FD2DF894" author="Thatje" box="[505,731,1846,1869]" pageId="1" pageNumber="68" refString="Thatje, S. &amp; Lorz, A. - N. (2005) First record of lithodid crabs from Antarctic waters off the Balleny Islands. Polar Biology, 28, 334 - 337." type="journal article" year="2005">Thatje &amp; Lörz, 2005</bibRefCitation>
: 335
</treatmentCitation>
336, fig. 2A [NIWA 3433, 3434 only; not
<taxonomicName id="E6C04D492628FFFC25D7F8A5FD7EF8B4" ID-CoL="e63447e8-fa4b-4c65-af30-a7c90fd6231f" authority="Dawson &amp; Yaldwyn, 1970" authorityName="Dawson &amp; Yaldwyn" authorityYear="1970" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="brodiei">
<emphasis id="13B4EAD82628FFFC25D7F8A5FE96F8B7" italics="true" pageId="1" pageNumber="46">N. brodiei</emphasis>
<bibRefCitation id="45514B3B2628FFFC206EF8C4FD7EF8B4" author="Dawson" box="[359,648,1878,1901]" pageId="1" pageNumber="68" refString="Dawson E. W. &amp; Yaldwyn, J. C. (1970) Diagnosis of a new species of Neolithodes (Crustacea: Anomura: Lithodidae) from New Zealand. New Zealand Journal of Marine and Freshwater Research 4, 227 - 228." type="journal article" year="1970">Dawson &amp; Yaldwyn, 1970</bibRefCitation>
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].
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<paragraph id="217F36CA262BFFFF2001FE8EFE46FE8A" blockId="2.[264,1323,284,371]" pageId="2" pageNumber="47">
<treatmentCitationGroup id="01D011E4262BFFFF2001FE8EFE46FE8A" pageId="2" pageNumber="47">
Not
<taxonomicName id="E6C04D49262BFFFF203EFE8EFDFFFEEA" ID-CoL="e63447e8-fa4b-4c65-af30-a7c90fd6231f" box="[311,521,284,307]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="brodiei">
<emphasis id="13B4EAD8262BFFFF203EFE8EFDFFFEEA" box="[311,521,284,307]" italics="true" pageId="2" pageNumber="47">Neolithodes brodiei</emphasis>
</taxonomicName>
.—
<treatmentCitation id="A06110DB262BFFFF2321FE8EFCC8FEEA" author="Thatje" box="[552,830,284,307]" page="335" pageId="2" pageNumber="47" year="2005">
<bibRefCitation id="45514B3B262BFFFF2321FE8EFCFDFEEA" author="Thatje" box="[552,779,284,307]" pageId="2" pageNumber="68" refString="Thatje, S. &amp; Lorz, A. - N. (2005) First record of lithodid crabs from Antarctic waters off the Balleny Islands. Polar Biology, 28, 334 - 337." type="journal article" year="2005">Thatje &amp; Lörz, 2005</bibRefCitation>
: 335
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[NIWA 3432 =
<taxonomicName id="E6C04D49262BFFFF22E4FE8EFE53FE8A" ID-CoL="4e79ccd5-3742-43da-a05e-8566dcca7097" authority="Macpherson, 1988" authorityName="Macpherson" authorityYear="1988" class="Malacostraca" family="Lithodidae" genus="Paralomis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="birsteini">
<emphasis id="13B4EAD8262BFFFF22E4FE8EFB4AFEEA" box="[1005,1212,284,307]" italics="true" pageId="2" pageNumber="47">Paralomis birsteini</emphasis>
Macpherson, 1988
</taxonomicName>
].
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</paragraph>
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<subSubSection id="69DA6541262BFFFF2001FECEFAE1FEAA" box="[264,1303,348,371]" pageId="2" pageNumber="47" type="description">
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<treatmentCitationGroup id="01D011E4262BFFFF2001FECEFAE1FEAA" box="[264,1303,348,371]" pageId="2" pageNumber="47">
Not
<taxonomicName id="E6C04D49262BFFFF203EFECEFDFFFEAA" ID-CoL="e63447e8-fa4b-4c65-af30-a7c90fd6231f" box="[311,521,348,371]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="brodiei">
<emphasis id="13B4EAD8262BFFFF203EFECEFDFFFEAA" box="[311,521,348,371]" italics="true" pageId="2" pageNumber="47">Neolithodes brodiei</emphasis>
</taxonomicName>
.—
<treatmentCitation id="A06110DB262BFFFF232EFECEFCCDFEAA" author="Thatje" box="[551,827,348,371]" page="335" pageId="2" pageNumber="47" year="2005">
<bibRefCitation id="45514B3B262BFFFF232EFECEFCFFFEAA" author="Thatje" box="[551,777,348,371]" pageId="2" pageNumber="68" refString="Thatje, S. &amp; Lorz, A. - N. (2005) First record of lithodid crabs from Antarctic waters off the Balleny Islands. Polar Biology, 28, 334 - 337." type="journal article" year="2005">Thatje &amp; Lörz, 2005</bibRefCitation>
: 335
</treatmentCitation>
[NIWA 3435 =
<taxonomicName id="E6C04D49262BFFFF22E0FECEFB44FEAA" box="[1001,1202,348,371]" class="Malacostraca" family="Lithodidae" genus="Paralomis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="stevensi" status="sp. nov.">
<emphasis id="13B4EAD8262BFFFF22E0FECEFB44FEAA" box="[1001,1202,348,371]" italics="true" pageId="2" pageNumber="47">Paralomis stevensi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="088757A3262BFFFF25B3FECEFAFDFEAA" box="[1210,1291,348,371]" pageId="2" pageNumber="47" rank="species">sp. nov</taxonomicNameLabel>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
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material.
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<typeStatus id="FE7B8868262BFFFF20C4FE36FD9AFE67" box="[461,620,420,446]" pageId="2" pageNumber="47" type="holotype">HOLOTYPE</typeStatus>
:
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3434, male (cl
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, cw
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, tcl
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), N of Sturge I., Balleny Is., 67°13.2211.97S, 164°17.7814.87E,
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, TAN0402/226, RV
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<emphasis id="13B4EAD8262BFFFF20FBFE66FD94FDD4" box="[498,610,500,525]" italics="true" pageId="2" pageNumber="47">Tangaroa</emphasis>
</taxonomicName>
,
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3
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</date>
.
<typeStatus id="FE7B8868262BFFFF2218FE66FC4EFDD7" box="[785,952,500,526]" pageId="2" pageNumber="47" type="paratype">PARATYPES</typeStatus>
:
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3433,
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(cl 120.0 mm, cw
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, tcl 144.0 mm), seamount W of Sturge I., Balleny Is., 67°26.0926.37S, 163°52.9851.79E,
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, TAN0402/235, RV
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<emphasis id="13B4EAD8262BFFFF256BFDD6FB24FD84" box="[1122,1234,580,605]" italics="true" pageId="2" pageNumber="47">Tangaroa</emphasis>
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,
<date id="557E100A262BFFFF25EBFDD6FEB0FD5F" pageId="2" pageNumber="47" value="2004-03-04">
4
<bibRefCitation id="45514B3B262BFFFF25F3FDD6FEB0FD5F" author="Mar" pageId="2" pageNumber="68" refString="Macpherson, E. (2004) A new species and new records of lithodid crabs (Crustacea: Decapoda: Lithodidae) from the Crozet and Kerguelen Islands area (Subantarctica). Polar Biology, 27, 418 - 422." type="journal article" year="2004">Mar 2004</bibRefCitation>
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;
<collectionCode id="47D1AE0F262BFFFF205FFDFEFE5AFD5F" box="[342,428,620,646]" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="2" pageNumber="47">NMNZ</collectionCode>
CR11002,
<specimenCount id="37C6FD43262BFFFF2330FDFEFD61FD5F" box="[569,663,620,646]" pageId="2" pageNumber="47" type="male">2 males</specimenCount>
(cl 101.7103.6, cw 91.695.2, tcl
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), between Scott and Balleny Is., 67°0806S, 170°5447E,
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, RV
<emphasis id="13B4EAD8262BFFFF2586FD06FAD5FD74" box="[1167,1315,660,685]" italics="true" pageId="2" pageNumber="47">San Aotea II</emphasis>
, trip 2010, haul 180, E. Winslade,
<date id="557E100A262BFFFF23ADFD2EFCB6FD0F" box="[676,832,700,726]" pageId="2" pageNumber="47" value="2005-02-14">14 Feb 2005</date>
;
<collectionCode id="47D1AE0F262BFFFF2247FD2EFC53FD0F" box="[846,933,700,726]" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="2" pageNumber="47">NMNZ</collectionCode>
CR11003,
<specimenCount id="37C6FD43262BFFFF2539FD2EFB74FD0F" box="[1072,1154,700,726]" pageId="2" pageNumber="47" type="male">1 male</specimenCount>
(cl 136.6, cw 131.1, tcl
<quantity id="E6389B2F262BFFFF208CFD76FDF3FD27" box="[389,517,740,766]" metricMagnitude="-1" metricUnit="m" metricValue="1.833" pageId="2" pageNumber="47" unit="mm" value="183.3">183.3 mm</quantity>
), near Scott I., 68°2321S, 179°5357E, 1337
<quantity id="E6389B2F262BFFFF2554FD76FB4BFD27" box="[1117,1213,740,766]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="2" pageNumber="47" unit="m" value="1050.0">1050 m</quantity>
, RV
<emphasis id="13B4EAD8262BFFFF25F7FD76FE9EFCFC" italics="true" pageId="2" pageNumber="47">San Aotea II</emphasis>
, trip 2010, haul 137B, on hook, B, E. Winslade,
<date id="557E100A262BFFFF22ADFC9EFBC3FCFF" box="[932,1077,780,806]" pageId="2" pageNumber="47" value="2005-01-31">31 Jan 2005</date>
.
</paragraph>
</subSubSection>
<subSubSection id="69DA6541262BFFFF2031FCA6FB3FFB57" pageId="2" pageNumber="47" type="diagnosis">
<paragraph id="217F36CA262BFFFF2031FCA6FB3FFB57" blockId="2.[264,1324,419,1846]" pageId="2" pageNumber="47">
<emphasis id="13B4EAD8262BFFFF2031FCA6FDDAFC97" bold="true" box="[312,556,820,846]" pageId="2" pageNumber="47">Diagnosis of adults.</emphasis>
Carapace dorsal surface with thick, conical spines and scattered, widely separated granules or minute secondary spines; longest spine not exceeding 0.2 times carapace length. Rostrum 0.10.3 times carapace length; ventral surface smooth. Posterior orbital margin near vertical; outer orbital spine slender, reaching to but not beyond cornea. Scaphocerite shorter than fourth antennal segment. Major cheliped palm with prominent dorsal and lateral spines; dactylus with convex dorsal margin, slightly longer than dorsal margin of palm. Ambulatory legs spinose, secondary spines absent or sparsely distributed; meri ovate in cross­section, shorter than carapace; propodi subcylindrical in cross­section; dactyli distinctly longer than half propodal length.
</paragraph>
</subSubSection>
<subSubSection id="69DA6541262BFFFA2031FB0EFDA8F9D7" lastPageId="7" lastPageNumber="52" pageId="2" pageNumber="47" type="description">
<paragraph id="217F36CA262BFFFF2031FB0EFBCEF9B7" blockId="2.[264,1324,419,1846]" pageId="2" pageNumber="47">
<emphasis id="13B4EAD8262BFFFF2031FB0EFE39FB6F" bold="true" box="[312,463,1180,1206]" pageId="2" pageNumber="47">Description.</emphasis>
<emphasis id="13B4EAD8262BFFFF20D2FB0EFDB8FB6C" box="[475,590,1180,1205]" italics="true" pageId="2" pageNumber="47">Carapace</emphasis>
: Pyriform, about 1.1 times longer than wide; dorsal surface armed with thick, conical spines and scattered, widely separated granules or minute secondary spines, surface otherwise smooth; longest spine shorter than 0.2 carapace length. Gastric region bearing 6 large spines forming transverse hexagon; with 2 near transverse rows of 3 smaller spines, first across centre of hexagon and second slightly anterior to hexagon. Hepatic spine strong, anterolaterally directed; margin between outer orbital spine and hepatic spine with 2 or 3 spines. Branchial regions with 8 or 9 large dorsal spines in addition to scattered smaller secondary spines; margins with 13 large spines in addition to several smaller spines. Cardiac region with 4 equal spines of similar size to gastric spines and pair of smaller spines near intestinal region. Intestinal region with 2 upright spines in transverse row and 2 smaller posterior directed spines. Pterygostomian region spinose or tuberculate, with small distal spine.
</paragraph>
<paragraph id="217F36CA262BFFFF2031F9EEFDB6F8EF" blockId="2.[264,1324,419,1846]" pageId="2" pageNumber="47">
<emphasis id="13B4EAD8262BFFFF2031F9EEFDEBF94C" box="[312,541,1660,1685]" italics="true" pageId="2" pageNumber="47">Rostrum and orbit</emphasis>
: Rostrum 0.10.3 times carapace length; median spine nearly horizontal, with pair of divergent dorsal spines and smaller pair of basal spines; ventral surface smooth. Posterior orbital margin near vertical; outer orbital spine slender, reaching to but not beyond cornea (when eyes directed forwards). Anterolateral spine about half as long as outer orbital spine.
</paragraph>
<caption id="75BF6642262AFFFE2001F8A7FB0AF8B6" httpUri="https://zenodo.org/record/173680/files/figure.png" pageId="3" pageNumber="48" targetBox="[264,1323,329,1817]" targetPageId="3">
<paragraph id="217F36CA262AFFFE2001F8A7FB0AF8B6" blockId="3.[264,1322,1845,1903]" pageId="3" pageNumber="48">
<emphasis id="13B4EAD8262AFFFE2001F8A7FE7BF894" bold="true" box="[264,397,1845,1869]" pageId="3" pageNumber="48">FIGURE 1.</emphasis>
<taxonomicName id="E6C04D49262AFFFE209EF8A7FD76F895" box="[407,640,1845,1868]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="yaldwyni" status="sp. nov.">
<emphasis id="13B4EAD8262AFFFE209EF8A7FD76F895" box="[407,640,1845,1868]" italics="true" pageId="3" pageNumber="48">Neolithodes yaldwyni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="088757A3262AFFFE2383F8A7FD14F894" box="[650,738,1845,1869]" pageId="3" pageNumber="48" rank="species">
<emphasis id="13B4EAD8262AFFFE2383F8A7FD14F894" bold="true" box="[650,738,1845,1869]" pageId="3" pageNumber="48">sp. nov.</emphasis>
</taxonomicNameLabel>
, male holotype (cl 141.5 mm, cw 124.1 mm, NIWA 3434). A, overall dorsal view. B, carapace, dorsal view. C, anterior carapace, right lateral view.
</paragraph>
</caption>
<caption id="75BF6642262DFFF92001F992FD1BF9A6" httpUri="https://zenodo.org/record/173681/files/figure.png" pageId="4" pageNumber="49" targetBox="[264,1323,315,1512]" targetPageId="4">
<paragraph id="217F36CA262DFFF92001F992FD1BF9A6" blockId="4.[264,1322,1536,1663]" pageId="4" pageNumber="49">
<emphasis id="13B4EAD8262DFFF92001F992FE7BF9C1" bold="true" box="[264,397,1536,1560]" pageId="4" pageNumber="49">FIGURE 2.</emphasis>
<taxonomicName id="E6C04D49262DFFF9209EF993FD76F9C1" box="[407,640,1537,1560]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="yaldwyni" status="sp. nov.">
<emphasis id="13B4EAD8262DFFF9209EF993FD76F9C1" box="[407,640,1537,1560]" italics="true" pageId="4" pageNumber="49">Neolithodes yaldwyni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="088757A3262DFFF92383F992FD14F9C1" box="[650,738,1536,1560]" pageId="4" pageNumber="49" rank="species">
<emphasis id="13B4EAD8262DFFF92383F992FD14F9C1" bold="true" box="[650,738,1536,1560]" pageId="4" pageNumber="49">sp. nov.</emphasis>
</taxonomicNameLabel>
, male holotype (cl 141.5 mm, cw 124.1 mm, NIWA 3434). A, carapace, right lateral view. B, left antenna, dorsal view. C, ventral surface. D, posterior carapace and abdomen, posterior view. E, coxa and basis of left third ambulatory leg, ventral view. F, rostrum and anterior carapace, dorsal view.
</paragraph>
</caption>
<paragraph id="217F36CA262DFFF92031F93CFCA5F8C1" blockId="4.[264,1323,1710,1896]" pageId="4" pageNumber="49">
<emphasis id="13B4EAD8262DFFF92031F93CFE0AF91E" box="[312,508,1710,1735]" italics="true" pageId="4" pageNumber="49">Ocular peduncle</emphasis>
: longer than cornea; with scattered dorsal granules or spinules.
<emphasis id="13B4EAD8262DFFF92031F944FE59F936" box="[312,431,1750,1775]" italics="true" pageId="4" pageNumber="49">Antennule</emphasis>
: peduncle unarmed, reaching anteriorly beyond antennal peduncle by about half­length of distal antennular peduncle segment.
</paragraph>
<paragraph id="217F36CA262DFFF82031F8B4FC74FE5F" blockId="4.[264,1323,1710,1896]" lastBlockId="5.[264,1323,284,550]" lastPageId="5" lastPageNumber="50" pageId="4" pageNumber="49">
<emphasis id="13B4EAD8262DFFF92031F8B4FE6FF8E6" box="[312,409,1830,1855]" italics="true" pageId="4" pageNumber="49">Antenna</emphasis>
: basal antennal segment with small anterolateral spine; outer margin of second segment with small basal spine and long slender spine that reaches beyond end of fourth segment but not beyond midlength; third segment with sharp to angular inner distal tooth; scaphocerite minute, shorter than fourth segment, apex sharp to blunt; fourth segment unarmed, about half as long as fifth segment.
</paragraph>
<paragraph id="217F36CA262CFFF82031FE06FAFCFDFF" blockId="5.[264,1323,284,550]" pageId="5" pageNumber="50">
<emphasis id="13B4EAD8262CFFF82031FE06FE50FE74" box="[312,422,404,429]" italics="true" pageId="5" pageNumber="50">Abdomen</emphasis>
: Second abdominal somite with 10 long spines and several smaller spines on median plate; submedian plate with 4 or 5 long spines on posterior border and 24 shorter spines on surface; lateral border of lateral plates with 68 teeth; surface and margins of remaining segments multispinose or nodulose; telson and penultimate segment spinose.
</paragraph>
<caption id="75BF6642262CFFF82001F975FCA1F8BF" httpUri="https://zenodo.org/record/173682/files/figure.png" pageId="5" pageNumber="50" targetBox="[264,1323,591,1755]" targetPageId="5">
<paragraph id="217F36CA262CFFF82001F975FCA1F8BF" blockId="5.[264,1323,1767,1894]" pageId="5" pageNumber="50">
<emphasis id="13B4EAD8262CFFF82001F975FE78F926" bold="true" box="[264,398,1767,1791]" pageId="5" pageNumber="50">FIGURE 3.</emphasis>
<taxonomicName id="E6C04D49262CFFF82093F97AFD72F926" box="[410,644,1768,1791]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="yaldwyni" status="sp. nov.">
<emphasis id="13B4EAD8262CFFF82093F97AFD72F926" box="[410,644,1768,1791]" italics="true" pageId="5" pageNumber="50">Neolithodes yaldwyni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="088757A3262CFFF82386F975FD1FF926" box="[655,745,1767,1791]" pageId="5" pageNumber="50" rank="species">
<emphasis id="13B4EAD8262CFFF82386F975FD1FF926" bold="true" box="[655,745,1767,1791]" pageId="5" pageNumber="50">sp. nov.</emphasis>
</taxonomicNameLabel>
AD, male holotype (cl 141.5 mm, cw 124.1 mm, NIWA 3434). EF, male paratype (cl 120.0 mm, cw 108.7 mm, NIWA 3433). A, left cheliped, dorsal view. B, left chela, outer surface. C, right cheliped, dorsal view. D, right chela, outer surface. E, left chela, outer surface. F, right chela, outer surface.
</paragraph>
</caption>
<caption id="75BF6642262FFFFB2001F883FADDF892" httpUri="https://zenodo.org/record/173683/files/figure.png" pageId="6" pageNumber="51" targetBox="[264,1323,326,1785]" targetPageId="6">
<paragraph id="217F36CA262FFFFB2001F883FADDF892" blockId="6.[264,1323,1809,1867]" pageId="6" pageNumber="51">
<emphasis id="13B4EAD8262FFFFB2001F883FE7DF8F0" bold="true" box="[264,395,1809,1833]" pageId="6" pageNumber="51">FIGURE 4.</emphasis>
<taxonomicName id="E6C04D49262FFFFB209DF883FD8AF8F1" box="[404,636,1809,1832]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="yaldwyni" status="sp. nov.">
<emphasis id="13B4EAD8262FFFFB209DF883FD8AF8F1" box="[404,636,1809,1832]" italics="true" pageId="6" pageNumber="51">Neolithodes yaldwyni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="088757A3262FFFFB238CF883FD2AF8F0" box="[645,732,1809,1833]" pageId="6" pageNumber="51" rank="species">
<emphasis id="13B4EAD8262FFFFB238CF883FD2AF8F0" bold="true" box="[645,732,1809,1833]" pageId="6" pageNumber="51">sp. nov.</emphasis>
</taxonomicNameLabel>
, overall view, dorsal. A, male paratype (cl 101.7 mm, cw 91.6 mm, NMNZ CR11002). B, male paratype (cl 136.6 mm, cw 131.1 mm, NMNZ CR11003).
</paragraph>
</caption>
<paragraph id="217F36CA262EFFFA2031FE8EFB4AFC07" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="52">
<emphasis id="13B4EAD8262EFFFA2031FE8EFE5BFEEC" box="[312,429,284,309]" italics="true" pageId="7" pageNumber="52">Chelipeds</emphasis>
: Major cheliped 1.31.6 times carapace length; minor cheliped 1.21.4 times carapace length. Coxa with blunt tubercles and tufts of setae, unarmed; ischiobasis with 4 or 5 stout ventral spines. Merus inner margin smooth or granular, with stout subdistal spine; ventral margin with two rows of stout spines, mesial row with 2 spines, lateral row with 4 spines; dorsal and lateral surface spinose, spines largest distally. Carpus with prominent spines on dorsal and lateral surfaces; dorsal margin with row of 4 or 5 stout spines; lateral margin with 2 or 3 rows of 4 or 5 stout spines of similar size to dorsal row; ventral margin small scattered acute tubercles. Upper palm length 1.11.2 times height (major chela), 1.31.4 times height (minor chela); prominently spinose on dorsal, lateral and ventral surfaces, inner surface with acute tubercles; dorsal margin with row of about 5 prominent conical spines; midlateral surface with 2 rows of 4 or 5 spines of similar size to dorsal row; ventral surface with 2 rows of about 6 or 7 spines, smaller than lateral and dorsal spines. Fingers with 2 or 3 basal spines and rows of tufts of golden setae. Fingers of major chela with occlusal margins corneous for slightly less than distal half, proximally with 3 calcareous nodules; dactylus dorsal margin convex, 1.21.4 times longer than dorsal margin of palm (apex broken in
<typeStatus id="FE7B8868262EFFFA226AFCE6FC39FC57" box="[867,975,884,910]" pageId="7" pageNumber="52" type="holotype">holotype</typeStatus>
). Fingers of minor chela with occlusal margins corneous for slightly more than distal half, proximally crenulate; dactylus dorsal margin convex, 1.51.8 times longer than dorsal margin of palm.
</paragraph>
<paragraph id="217F36CA262EFFFA2031FC7EFAE7FA3F" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="52">
<emphasis id="13B4EAD8262EFFFA2031FC7EFE0BFBDC" box="[312,509,1004,1029]" italics="true" pageId="7" pageNumber="52">Ambulatory legs</emphasis>
: Segments spinose, surface between major spines smooth or with few, minute, sparsely distributed spines. Third ambulatory leg longest, 2.83.0 times carapace length. Distal margins of coxae with short spines, longest on third coxa. Ischiobasis with 3 or 4 distal spines and 2 or 3 smaller ventral spines. Merus ovate in cross section, shorter than carapace; 5.87.3 times longer than high; extensor margin with 8 or 9 spines in addition to paired distal spines; dorsal surface with row of 46 spines of similar size to extensor spines; flexor margin with 2 rows of 57 spines. Carpus slightly longer than half merus length; extensor margin with 5 or 6 spines, distal and second proximal spines longest, about twice as long as other spines; dorsal surface with 4 or 5 spines. Propodus ovate to subcircular in cross­section; about 0.9 merus length, 8.911.9 times longer than high; with 79 spines on extensor margin and 810 similar spines on dorsal surface; flexor margin with 610 smaller spines. Dactylus curved, rounded in cross section, with 46 small proximal spines, apex corneous; exceeding 0.6 propodus length.
</paragraph>
<paragraph id="217F36CA262EFFFA2031FA66FDA8F9D7" blockId="7.[264,1324,284,1870]" box="[312,606,1524,1550]" pageId="7" pageNumber="52">
<emphasis id="13B4EAD8262EFFFA2031FA66FE13F9D7" bold="true" box="[312,485,1524,1550]" pageId="7" pageNumber="52">Colour in life.</emphasis>
Deep red.
</paragraph>
</subSubSection>
<subSubSection id="69DA6541262EFFFA2031F98EFD91F95F" pageId="7" pageNumber="52" type="etymology">
<paragraph id="217F36CA262EFFFA2031F98EFD91F95F" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="52">
<emphasis id="13B4EAD8262EFFFA2031F98EFE32F9EF" bold="true" box="[312,452,1564,1590]" pageId="7" pageNumber="52">Etymology.</emphasis>
Named in honour of the late John Yaldwyn, in recognition of his important contributions to
<collectingCountry id="59D7765A262EFFFA2360F9D6FCE5F987" box="[617,787,1604,1630]" name="New Zealand" pageId="7" pageNumber="52">New Zealand</collectingCountry>
and subantarctic zoology, including the systematics of the
<taxonomicName id="E6C04D49262EFFFA20E8F9FEFD92F95F" box="[481,612,1644,1670]" class="Malacostraca" family="Lithodidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="52" phylum="Arthropoda" rank="family">Lithodidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="69DA6541262EFFF72031F906FD0BFDAF" lastPageId="10" lastPageNumber="55" pageId="7" pageNumber="52" type="discussion">
<paragraph id="217F36CA262EFFF52031F906FDD4FA9F" blockId="7.[264,1324,284,1870]" lastBlockId="8.[264,1325,284,1870]" lastPageId="8" lastPageNumber="53" pageId="7" pageNumber="52">
<emphasis id="13B4EAD8262EFFFA2031F906FE47F977" bold="true" box="[312,433,1684,1710]" pageId="7" pageNumber="52">Remarks.</emphasis>
<taxonomicName id="E6C04D49262EFFFA20B0F906FD41F974" box="[441,695,1684,1709]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="yaldwyni" status="sp. nov.">
<emphasis id="13B4EAD8262EFFFA20B0F906FD41F974" box="[441,695,1684,1709]" italics="true" pageId="7" pageNumber="52">Neolithodes yaldwyni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="088757A3262EFFFA23B7F906FCEBF977" box="[702,797,1684,1710]" pageId="7" pageNumber="52" rank="species">
<emphasis id="13B4EAD8262EFFFA23B7F906FCEBF977" bold="true" box="[702,797,1684,1710]" pageId="7" pageNumber="52">sp. nov.</emphasis>
</taxonomicNameLabel>
belongs to the group of species in the genus in which the carapace and ambulatory legs at most bear scattered, minute secondary spines or tubercles amongst the primary dorsal spines. These species are:
<taxonomicName id="E6C04D49262EFFFA2576F976FE39F8FF" authority="Benedict, 1895" authorityName="Benedict" authorityYear="1895" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="diomedeae">
<emphasis id="13B4EAD8262EFFFA2576F976FADCF924" box="[1151,1322,1764,1789]" italics="true" pageId="7" pageNumber="52">N. diomedeae</emphasis>
(
<bibRefCitation id="45514B3B262EFFFA2019F89EFE30F8FF" author="Benedict" box="[272,454,1804,1830]" pageId="7" pageNumber="67" refString="Benedict, J. E. (1895) Scientific results of explorations by the U. S. Fish Commission steamer Albatross. No. XXXI. Descriptions of new genera and species of crabs of the family Lithodidae, with notes on the young of Lithodes camtschaticus and Lithodes brevipes. Proceedings of the Unites States National Museum, 17, 479 - 488." type="journal article" year="1895">Benedict, 1895</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="E6C04D49262EFFFA20D5F89EFCEBF8FF" authority="Stebbing, 1905" authorityName="Stebbing" authorityYear="1905" box="[476,797,1804,1830]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="13B4EAD8262EFFFA20D5F89EFD95F8FC" box="[476,611,1804,1829]" italics="true" pageId="7" pageNumber="52">N. capensis</emphasis>
<bibRefCitation id="45514B3B262EFFFA2363F89EFCEBF8FF" author="Stebbing" box="[618,797,1804,1830]" pageId="7" pageNumber="68" refString="Stebbing, T. R. R. (1905) South African Crustacea. Part 3. Marine Investigations in South Africa, 4, 21 - 123, pls. 1 - 10." type="journal article" year="1905">Stebbing, 1905</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="E6C04D49262EFFFA2220F89EFEBAF897" authority="A. Milne Edwards &amp; Bouvier, 1894" authorityName="A. Milne Edwards &amp; Bouvier" authorityYear="1894" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="grimaldii">
<emphasis id="13B4EAD8262EFFFA2220F89EFC4EF8FC" box="[809,952,1804,1829]" italics="true" pageId="7" pageNumber="52">N. grimaldii</emphasis>
(A.
<bibRefCitation id="45514B3B262EFFFA22E4F89EFEB2F897" author="Milne" pageId="7" pageNumber="68" refString="Milne Edwards, A. &amp; Bouvier, E. L. (1894) Brachyures et Anomures. In: Crustaces provenant des campagnes du yacht l'Hirondelle (1886, 1887, 1888), Premiere Partie. Resultats des Campagnes Scientifiques accomplies sur son yacht par Albert I er, Prince Souverain de Monaco, 7, 1 - 112, pl. 1 - 11." type="journal article" year="1894">Milne Edwards &amp; Bouvier, 1894</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="E6C04D49262EFFFA2087F8A6FCD8F897" authority="Macpherson, 1988" authorityName="Macpherson" authorityYear="1988" box="[398,814,1844,1870]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="vinogradovi">
<emphasis id="13B4EAD8262EFFFA2087F8A6FDB4F894" box="[398,578,1844,1869]" italics="true" pageId="7" pageNumber="52">N. vinogradovi</emphasis>
<bibRefCitation id="45514B3B262EFFFA2344F8A6FCD8F897" author="Macpherson" box="[589,814,1844,1870]" pageId="7" pageNumber="68" refString="Macpherson, E. (1988 b) Revision of the family Lithodidae Samouelle, 1819 (Crustacea, Decapoda, Anomura) in the Atlantic Ocean. Monografias de Zoologia Marina, 2, 9 - 153." type="journal article" year="1988" yearSuffix="b">Macpherson, 1988</bibRefCitation>
</taxonomicName>
b.
<taxonomicName id="E6C04D49262EFFFA2244F8A6FB92F894" box="[845,1124,1844,1869]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="diomedeae">
<emphasis id="13B4EAD8262EFFFA2244F8A6FB92F894" box="[845,1124,1844,1869]" italics="true" pageId="7" pageNumber="52">Neolithodes diomedeae</emphasis>
</taxonomicName>
is distinguished from all others in the group by the shape of the cheliped fingers, which are triangular instead of rounded in cross­section and dorsally flattened or concave instead of convex.
<taxonomicName id="E6C04D492621FFF52001FEFEFDFFFE5C" box="[264,521,364,389]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="grimaldii">
<emphasis id="13B4EAD82621FFF52001FEFEFDFFFE5C" box="[264,521,364,389]" italics="true" pageId="8" pageNumber="53">Neolithodes grimaldii</emphasis>
</taxonomicName>
and
<taxonomicName id="E6C04D492621FFF5234CFEFEFD00FE5C" box="[581,758,364,389]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="vinogradovi">
<emphasis id="13B4EAD82621FFF5234CFEFEFD00FE5C" box="[581,758,364,389]" italics="true" pageId="8" pageNumber="53">N. vinogradovi</emphasis>
</taxonomicName>
are readily distinguished from
<taxonomicName id="E6C04D492621FFF52562FEFEFB0EFE5C" box="[1131,1272,364,389]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82621FFF52562FEFEFB0EFE5C" box="[1131,1272,364,389]" italics="true" pageId="8" pageNumber="53">N. yaldwyni</emphasis>
</taxonomicName>
and
<taxonomicName id="E6C04D492621FFF52001FE06FE66FE74" box="[264,400,404,429]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="13B4EAD82621FFF52001FE06FE66FE74" box="[264,400,404,429]" italics="true" pageId="8" pageNumber="53">N. capensis</emphasis>
</taxonomicName>
by the considerably longer dorsal spines in adults, many of which exceed half the carapace length. Thus,
<taxonomicName id="E6C04D492621FFF52336FE2EFD3DFE0C" box="[575,715,444,469]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82621FFF52336FE2EFD3DFE0C" box="[575,715,444,469]" italics="true" pageId="8" pageNumber="53">N. yaldwyni</emphasis>
</taxonomicName>
is closest to
<taxonomicName id="E6C04D492621FFF52256FE2EFC13FE0C" box="[863,997,444,469]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="13B4EAD82621FFF52256FE2EFC13FE0C" box="[863,997,444,469]" italics="true" pageId="8" pageNumber="53">N. capensis</emphasis>
</taxonomicName>
, which it resembles in most respects. The strong resemblance between
<taxonomicName id="E6C04D492621FFF5220CFE76FC78FE24" box="[773,910,484,509]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="13B4EAD82621FFF5220CFE76FC78FE24" box="[773,910,484,509]" italics="true" pageId="8" pageNumber="53">N. capensis</emphasis>
</taxonomicName>
and
<taxonomicName id="E6C04D492621FFF522C3FE76FBAEFE24" box="[970,1112,484,509]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82621FFF522C3FE76FBAEFE24" box="[970,1112,484,509]" italics="true" pageId="8" pageNumber="53">N. yaldwyni</emphasis>
</taxonomicName>
gave some pause to referring the latter to a new species, but the morphological differences between the two forms are consistent. Adult
<taxonomicName id="E6C04D492621FFF52357FDA6FD18FD94" box="[606,750,564,589]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82621FFF52357FDA6FD18FD94" box="[606,750,564,589]" italics="true" pageId="8" pageNumber="53">N. yaldwyni</emphasis>
</taxonomicName>
differ most obviously from
<taxonomicName id="E6C04D492621FFF52544FDA6FB2FFD94" box="[1101,1241,564,589]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="13B4EAD82621FFF52544FDA6FB2FFD94" box="[1101,1241,564,589]" italics="true" pageId="8" pageNumber="53">N. capensis</emphasis>
</taxonomicName>
in the proportional lengths of the dactyli of the ambulatory legs, being distinctly more than half the propodus length (exceeding 0.6) instead of less than half (about 0.4). Although the proportional lengths of the dactyli of the ambulatory legs vary allometrically in
<taxonomicName id="E6C04D492621FFF52001FD46FE62FD34" box="[264,404,724,749]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="13B4EAD82621FFF52001FD46FE62FD34" box="[264,404,724,749]" italics="true" pageId="8" pageNumber="53">Neolithodes</emphasis>
</taxonomicName>
, the dactyli become proportionally shorter with increasing body size. Thus, the morphometric differences in dactyl length between
<taxonomicName id="E6C04D492621FFF522ADFD6EFBC2FCCC" box="[932,1076,764,789]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82621FFF522ADFD6EFBC2FCCC" box="[932,1076,764,789]" italics="true" pageId="8" pageNumber="53">N. yaldwyni</emphasis>
</taxonomicName>
and
<taxonomicName id="E6C04D492621FFF5257BFD6EFB0AFCCC" box="[1138,1276,764,789]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="13B4EAD82621FFF5257BFD6EFB0AFCCC" box="[1138,1276,764,789]" italics="true" pageId="8" pageNumber="53">N. capensis</emphasis>
</taxonomicName>
are not attributable to allometry. The largest specimen of
<taxonomicName id="E6C04D492621FFF5228DFCB6FBE4FCE4" box="[900,1042,804,829]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82621FFF5228DFCB6FBE4FCE4" box="[900,1042,804,829]" italics="true" pageId="8" pageNumber="53">N. yaldwyni</emphasis>
</taxonomicName>
(cl
<quantity id="E6389B2F2621FFF52548FCB6FB4DFCE7" box="[1089,1211,804,830]" metricMagnitude="-1" metricUnit="m" metricValue="1.415" pageId="8" pageNumber="53" unit="mm" value="141.5">141.5 mm</quantity>
) is larger than the largest reported specimen of
<taxonomicName id="E6C04D492621FFF523CEFCDEFCA6FCBC" box="[711,848,844,869]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="13B4EAD82621FFF523CEFCDEFCA6FCBC" box="[711,848,844,869]" italics="true" pageId="8" pageNumber="53">N. capensis</emphasis>
</taxonomicName>
(cl
<quantity id="E6389B2F2621FFF52276FCDEFC12FCBF" box="[895,996,844,870]" metricMagnitude="-1" metricUnit="m" metricValue="1.31" pageId="8" pageNumber="53" unit="mm" value="131.0">131 mm</quantity>
,
<bibRefCitation id="45514B3B2621FFF522F8FCDEFB2EFCBF" author="Macpherson" box="[1009,1240,844,870]" pageId="8" pageNumber="68" refString="Macpherson, E. (1988 b) Revision of the family Lithodidae Samouelle, 1819 (Crustacea, Decapoda, Anomura) in the Atlantic Ocean. Monografias de Zoologia Marina, 2, 9 - 153." type="journal article" year="1988" yearSuffix="b">Macpherson 1988b</bibRefCitation>
: 40) in which the dactylus measured about 0.43 propodus length (based on
<bibRefCitation id="45514B3B2621FFF52534FCE6FAD3FC57" author="Macpherson" box="[1085,1317,884,910]" pageId="8" pageNumber="68" refString="Macpherson, E. (1988 b) Revision of the family Lithodidae Samouelle, 1819 (Crustacea, Decapoda, Anomura) in the Atlantic Ocean. Monografias de Zoologia Marina, 2, 9 - 153." type="journal article" year="1988" yearSuffix="b">Macpherson 1988b</bibRefCitation>
: fig. 17H). Thus, proportional differences in dactyl/propodal lengths can be expected to be even greater in size­matched individuals of
<taxonomicName id="E6C04D492621FFF52207FC56FC6DFC04" box="[782,923,964,989]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82621FFF52207FC56FC6DFC04" box="[782,923,964,989]" italics="true" pageId="8" pageNumber="53">N. yaldwyni</emphasis>
</taxonomicName>
and
<taxonomicName id="E6C04D492621FFF522DFFC56FBA8FC04" box="[982,1118,964,989]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="13B4EAD82621FFF522DFFC56FBA8FC04" box="[982,1118,964,989]" italics="true" pageId="8" pageNumber="53">N. capensis</emphasis>
</taxonomicName>
. Similar dactylar proportions (0.45) were reported by
<bibRefCitation id="45514B3B2621FFF523B9FC7EFC9AFBDF" author="Stebbing" box="[688,876,1004,1030]" pageId="8" pageNumber="68" refString="Stebbing, T. R. R. (1905) South African Crustacea. Part 3. Marine Investigations in South Africa, 4, 21 - 123, pls. 1 - 10." type="journal article" year="1905">Stebbing (1905)</bibRefCitation>
in the
<typeStatus id="FE7B88682621FFF522B2FC7EFC18FBDF" box="[955,1006,1004,1030]" pageId="8" pageNumber="53">type</typeStatus>
description of
<taxonomicName id="E6C04D492621FFF52595FC7EFAD5FBDC" box="[1180,1315,1004,1029]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="13B4EAD82621FFF52595FC7EFAD5FBDC" box="[1180,1315,1004,1029]" italics="true" pageId="8" pageNumber="53">N. capensis</emphasis>
</taxonomicName>
, and figured for the female
<typeStatus id="FE7B88682621FFF5234AFB86FD68FBF7" box="[579,670,1044,1070]" pageId="8" pageNumber="53" type="syntype">syntype</typeStatus>
(tcl
<quantity id="E6389B2F2621FFF523DDFB86FCCFFBF7" box="[724,825,1044,1070]" metricMagnitude="-1" metricUnit="m" metricValue="1.48" pageId="8" pageNumber="53" unit="mm" value="148.0">148 mm</quantity>
, cl
<quantity id="E6389B2F2621FFF52268FB86FC3EFBF7" box="[865,968,1044,1070]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="53" unit="mm" value="120.0">120 mm</quantity>
).
<bibRefCitation id="45514B3B2621FFF522D5FB86FADCFBF7" author="Macpherson" box="[988,1322,1044,1070]" pageId="8" pageNumber="68" refString="Macpherson, E. (1988 b) Revision of the family Lithodidae Samouelle, 1819 (Crustacea, Decapoda, Anomura) in the Atlantic Ocean. Monografias de Zoologia Marina, 2, 9 - 153." type="journal article" year="1988" yearSuffix="b">Macpherson (1988b: pl. 5A)</bibRefCitation>
also figured an
<quantity id="E6389B2F2621FFF520B5FBAEFDFBFB8F" box="[444,525,1084,1110]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="8" pageNumber="53" unit="mm" value="82.0">82 mm</quantity>
specimen of
<taxonomicName id="E6C04D492621FFF523A3FBAEFCC7FB8C" box="[682,817,1084,1109]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="13B4EAD82621FFF523A3FBAEFCC7FB8C" box="[682,817,1084,1109]" italics="true" pageId="8" pageNumber="53">N. capensis</emphasis>
</taxonomicName>
in which the dactyli of the ambulatory legs are half the length of their respective propodi. Thus, even at
<quantity id="E6389B2F2621FFF522C7FBF6FBD7FBA7" box="[974,1057,1124,1150]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="8" pageNumber="53" unit="mm" value="82.0">82 mm</quantity>
cl, the two species are distinguishable based on the relative length of the ambulatory leg dactyli. The dactylus/ propodus proportions of the last ambulatory leg of the
<quantity id="E6389B2F2621FFF522ACFB26FC0DFB17" box="[933,1019,1204,1230]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="8" pageNumber="53" unit="mm" value="65.0">65 mm</quantity>
specimen of
<taxonomicName id="E6C04D492621FFF525A9FB26FADDFB14" box="[1184,1323,1204,1229]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="13B4EAD82621FFF525A9FB26FADDFB14" box="[1184,1323,1204,1229]" italics="true" pageId="8" pageNumber="53">N. capensis</emphasis>
</taxonomicName>
figured by
<bibRefCitation id="45514B3B2621FFF52082FB4EFD00FB2F" author="Macpherson" box="[395,758,1244,1270]" pageId="8" pageNumber="68" refString="Macpherson, E. (1988 b) Revision of the family Lithodidae Samouelle, 1819 (Crustacea, Decapoda, Anomura) in the Atlantic Ocean. Monografias de Zoologia Marina, 2, 9 - 153." type="journal article" year="1988" yearSuffix="b">Macpherson (1988b: fig. 17G)</bibRefCitation>
approaches that of the much larger (+
<quantity id="E6389B2F2621FFF525B4FB4EFAD5FB2F" box="[1213,1315,1244,1270]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="53" unit="mm" value="100.0">100 mm</quantity>
)
<typeStatus id="FE7B88682621FFF52001FA96FECCFAC7" box="[264,314,1284,1310]" pageId="8" pageNumber="53">type</typeStatus>
specimens of
<taxonomicName id="E6C04D492621FFF520EFFA96FD84FAC4" box="[486,626,1284,1309]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82621FFF520EFFA96FD84FAC4" box="[486,626,1284,1309]" italics="true" pageId="8" pageNumber="53">N. yaldwyni</emphasis>
</taxonomicName>
, and as such, small specimens of the two species might be difficult to differentiate.
</paragraph>
<paragraph id="217F36CA2621FFF52031FAC6FB51FA3F" blockId="8.[264,1325,284,1870]" pageId="8" pageNumber="53">
<bibRefCitation id="45514B3B2621FFF52031FAC6FE18FAB7" author="Barnard" box="[312,494,1364,1390]" pageId="8" pageNumber="67" refString="Barnard, K. H. (1950) Descriptive catalogue of South African decapod Crustacea. Annals of the South African Museum, 38, 1 - 837." type="journal article" year="1950">Barnard (1950)</bibRefCitation>
noted that the relative lengths of the cheliped fingers do not change appreciably with size in
<taxonomicName id="E6C04D492621FFF5232CFAEEFD47FA4C" box="[549,689,1404,1429]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="13B4EAD82621FFF5232CFAEEFD47FA4C" box="[549,689,1404,1429]" italics="true" pageId="8" pageNumber="53">Neolithodes</emphasis>
</taxonomicName>
. Thus,
<taxonomicName id="E6C04D492621FFF52201FAEEFC62FA4C" box="[776,916,1404,1429]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82621FFF52201FAEEFC62FA4C" box="[776,916,1404,1429]" italics="true" pageId="8" pageNumber="53">N. yaldwyni</emphasis>
</taxonomicName>
may also be distinguished from
<taxonomicName id="E6C04D492621FFF52419FAEEFE9BFA64" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="13B4EAD82621FFF52419FAEEFE9BFA64" italics="true" pageId="8" pageNumber="53">N. capensis</emphasis>
</taxonomicName>
in having a proportionally longer dactylus on the major cheliped, being longer than instead of as long as or slightly shorter than the dorsal margin of the palm.
</paragraph>
<paragraph id="217F36CA2621FFF52031FA66FB62F927" blockId="8.[264,1325,284,1870]" pageId="8" pageNumber="53">
Spine development in lithodids is inversely related to body size and evident in the
<typeStatus id="FE7B88682621FFF525FFFA66FADEF9D7" box="[1270,1320,1524,1550]" pageId="8" pageNumber="53">type</typeStatus>
series of
<taxonomicName id="E6C04D492621FFF52072F98EFDFDF9EC" box="[379,523,1564,1589]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82621FFF52072F98EFDFDF9EC" box="[379,523,1564,1589]" italics="true" pageId="8" pageNumber="53">N. yaldwyni</emphasis>
</taxonomicName>
. Even so, the overall spination of
<taxonomicName id="E6C04D492621FFF522B3F98EFBBDF9EC" box="[954,1099,1564,1589]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82621FFF522B3F98EFBBDF9EC" box="[954,1099,1564,1589]" italics="true" pageId="8" pageNumber="53">N. yaldwyni</emphasis>
</taxonomicName>
is distinctly more pronounced than in size­matched
<taxonomicName id="E6C04D492621FFF523DCF9D6FC9BF984" box="[725,877,1604,1629]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="13B4EAD82621FFF523DCF9D6FC9BF984" box="[725,877,1604,1629]" italics="true" pageId="8" pageNumber="53">N. capensis</emphasis>
</taxonomicName>
: the dorsal carapace spines are proportionally longer (compare
<figureCitation id="B9FB2A4F2621FFF52389F9FEFD34F95F" box="[640,706,1644,1670]" captionStart="FIGURE 1" captionStartId="3.[264,367,1845,1869]" captionTargetBox="[264,1323,329,1817]" captionTargetId="figure@3.[264,1323,329,1818]" captionTargetPageId="3" captionText="FIGURE 1. Neolithodes yaldwyni sp. nov., male holotype (cl 141.5 mm, cw 124.1 mm, NIWA 3434). A, overall dorsal view. B, carapace, dorsal view. C, anterior carapace, right lateral view." httpUri="https://zenodo.org/record/173680/files/figure.png" pageId="8" pageNumber="53">Fig. 1</figureCitation>
B, 4 with
<bibRefCitation id="45514B3B2621FFF52230F9FEFBD6F95F" author="Macpherson" box="[825,1056,1644,1670]" pageId="8" pageNumber="68" refString="Macpherson, E. (1988 b) Revision of the family Lithodidae Samouelle, 1819 (Crustacea, Decapoda, Anomura) in the Atlantic Ocean. Monografias de Zoologia Marina, 2, 9 - 153." type="journal article" year="1988" yearSuffix="b">Macpherson 1988b</bibRefCitation>
: pl. 5B) and the spines of the cheliped palm are well developed. In
<taxonomicName id="E6C04D492621FFF52205F906FC6EF974" box="[780,920,1684,1709]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82621FFF52205F906FC6EF974" box="[780,920,1684,1709]" italics="true" pageId="8" pageNumber="53">N. yaldwyni</emphasis>
</taxonomicName>
, the spines on the outer surface of the palm are of similar size to the dorsal spines, whereas in
<taxonomicName id="E6C04D492621FFF522C5F92EFBA3F90C" box="[972,1109,1724,1749]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="13B4EAD82621FFF522C5F92EFBA3F90C" box="[972,1109,1724,1749]" italics="true" pageId="8" pageNumber="53">N. capensis</emphasis>
</taxonomicName>
, the spines on the outer palm are poorly developed and distinctly smaller than the dorsal spines.
</paragraph>
<paragraph id="217F36CA2621FFF42031F89EFCC3FE0F" blockId="8.[264,1325,284,1870]" lastBlockId="9.[264,1324,284,1870]" lastPageId="9" lastPageNumber="54" pageId="8" pageNumber="53">
The development of the outer orbital spine might also prove useful in separating large specimens of
<taxonomicName id="E6C04D492621FFF520A4F8A6FDCDF894" box="[429,571,1844,1869]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82621FFF520A4F8A6FDCDF894" box="[429,571,1844,1869]" italics="true" pageId="8" pageNumber="53">N. yaldwyni</emphasis>
</taxonomicName>
from
<taxonomicName id="E6C04D492621FFF5238FF8A6FCE6F894" box="[646,784,1844,1869]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="13B4EAD82621FFF5238FF8A6FCE6F894" box="[646,784,1844,1869]" italics="true" pageId="8" pageNumber="53">N. capensis</emphasis>
</taxonomicName>
. The outer orbital spine is well­developed in all specimens of
<emphasis id="13B4EAD82620FFF420DDFE8EFD9FFEEC" box="[468,617,284,309]" italics="true" pageId="9" pageNumber="54">
<taxonomicName id="E6C04D492620FFF420DDFE8EFD93FEEC" box="[468,613,284,309]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="yaldwyni">N. yaldwyni</taxonomicName>
,
</emphasis>
reaching to, but not beyond the cornea (when the eyes are directed forward), whereas in the l
<quantity id="E6389B2F2620FFF423DCFED6FCC2FE87" box="[725,820,324,350]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="9" pageNumber="54" unit="mm" value="31.0">31 mm</quantity>
specimen of
<taxonomicName id="E6C04D492620FFF422FAFED6FB70FE84" box="[1011,1158,324,349]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="13B4EAD82620FFF422FAFED6FB70FE84" box="[1011,1158,324,349]" italics="true" pageId="9" pageNumber="54">N. capensis</emphasis>
</taxonomicName>
reported by
<bibRefCitation id="45514B3B2620FFF42001FEFEFDF2FE5F" author="Macpherson" box="[264,516,364,390]" pageId="9" pageNumber="68" refString="Macpherson, E. (1988 b) Revision of the family Lithodidae Samouelle, 1819 (Crustacea, Decapoda, Anomura) in the Atlantic Ocean. Monografias de Zoologia Marina, 2, 9 - 153." type="journal article" year="1988" yearSuffix="b">Macpherson (1988b)</bibRefCitation>
, the outer orbital spine is reduced to a short triangular tooth, not reaching the midlength of the ocular peduncle. Further studies are required to assess the degree of allometric and intraspecific variation.
</paragraph>
<paragraph id="217F36CA2620FFF42031FE76FAEBFB8F" blockId="9.[264,1324,284,1870]" pageId="9" pageNumber="54">
Aside from size­related differences, the
<typeStatus id="FE7B88682620FFF42269FE76FC65FE27" box="[864,915,484,510]" pageId="9" pageNumber="54">type</typeStatus>
material is in most respects morphologically uniform. Two
<typeStatus id="FE7B88682620FFF42389FD9EFD02FDFF" box="[640,756,524,550]" pageId="9" pageNumber="54" type="paratype">paratypes</typeStatus>
, however, are noteworthy: a
<quantity id="E6389B2F2620FFF4255BFD9EFB3DFDFF" box="[1106,1227,524,550]" metricMagnitude="-1" metricUnit="m" metricValue="1.366" pageId="9" pageNumber="54" unit="mm" value="136.6">136.6 mm</quantity>
cl male (NMNZ CR11003) (
<figureCitation id="B9FB2A4F2620FFF420F3FDA6FDB4FD97" box="[506,578,564,590]" captionStart="FIGURE 4" captionStartId="6.[264,367,1809,1833]" captionTargetBox="[264,1323,326,1785]" captionTargetId="figure@6.[264,1323,326,1785]" captionTargetPageId="6" captionText="FIGURE 4. Neolithodes yaldwyni sp. nov., overall view, dorsal. A, male paratype (cl 101.7 mm, cw 91.6 mm, NMNZ CR 11002). B, male paratype (cl 136.6 mm, cw 131.1 mm, NMNZ CR 11003)." httpUri="https://zenodo.org/record/173683/files/figure.png" pageId="9" pageNumber="54">Fig. 4</figureCitation>
B) and a 120.0 mm cl male (NIWA 3433) (
<figureCitation id="B9FB2A4F2620FFF42549FDA6FB70FD97" box="[1088,1158,564,590]" captionStart="FIGURE 3" captionStartId="5.[264,367,1767,1791]" captionTargetBox="[264,1323,591,1755]" captionTargetId="figure@5.[264,1323,591,1758]" captionTargetPageId="5" captionText="FIGURE 3. Neolithodes yaldwyni sp. nov. A D, male holotype (cl 141.5 mm, cw 124.1 mm, NIWA 3434). E F, male paratype (cl 120.0 mm, cw 108.7 mm, NIWA 3433). A, left cheliped, dorsal view. B, left chela, outer surface. C, right cheliped, dorsal view. D, right chela, outer surface. E, left chela, outer surface. F, right chela, outer surface." httpUri="https://zenodo.org/record/173682/files/figure.png" pageId="9" pageNumber="54">Fig. 3</figureCitation>
E, F). Though the dorsal spine development and body size are inversely related in most specimens, spination of the
<quantity id="E6389B2F2620FFF420CDFD16FDCDFD47" box="[452,571,644,670]" metricMagnitude="-1" metricUnit="m" metricValue="1.366" pageId="9" pageNumber="54" unit="mm" value="136.6">136.6 mm</quantity>
cl
<typeStatus id="FE7B88682620FFF42354FD16FD34FD47" box="[605,706,644,670]" pageId="9" pageNumber="54" type="paratype">paratype</typeStatus>
is more pronounced than other specimens, including the smallest
<typeStatus id="FE7B88682620FFF42090FD3EFE08FD1F" box="[409,510,684,710]" pageId="9" pageNumber="54" type="paratype">paratype</typeStatus>
(cl
<quantity id="E6389B2F2620FFF42323FD3EFD52FD1F" box="[554,676,684,710]" metricMagnitude="-1" metricUnit="m" metricValue="1.017" pageId="9" pageNumber="54" unit="mm" value="101.7">101.7 mm</quantity>
,
<figureCitation id="B9FB2A4F2620FFF423A6FD3EFD02FD1F" box="[687,756,684,710]" captionStart="FIGURE 4" captionStartId="6.[264,367,1809,1833]" captionTargetBox="[264,1323,326,1785]" captionTargetId="figure@6.[264,1323,326,1785]" captionTargetPageId="6" captionText="FIGURE 4. Neolithodes yaldwyni sp. nov., overall view, dorsal. A, male paratype (cl 101.7 mm, cw 91.6 mm, NMNZ CR 11002). B, male paratype (cl 136.6 mm, cw 131.1 mm, NMNZ CR 11003)." httpUri="https://zenodo.org/record/173683/files/figure.png" pageId="9" pageNumber="54">Fig. 4</figureCitation>
A). The longest spine on the
<quantity id="E6389B2F2620FFF4254AFD3EFB52FD1F" box="[1091,1188,684,710]" metricMagnitude="-1" metricUnit="m" metricValue="1.3599999999999999" pageId="9" pageNumber="54" unit="mm" value="136.0">136 mm</quantity>
cl
<typeStatus id="FE7B88682620FFF425CFFD3EFADDFD1F" box="[1222,1323,684,710]" pageId="9" pageNumber="54" type="paratype">paratype</typeStatus>
measures almost 0.2 cl in contrast to 0.1 cl in the smallest
<typeStatus id="FE7B88682620FFF422C5FD46FBC3FD37" box="[972,1077,724,750]" pageId="9" pageNumber="54" type="paratype">paratype</typeStatus>
. In other specimens, the carapace spines are less than 0.1 cl. Similarly, the spines on the ambulatory legs of
<quantity id="E6389B2F2620FFF425F7FD6EFEC0FCE7" metricMagnitude="-1" metricUnit="m" metricValue="1.3599999999999999" pageId="9" pageNumber="54" unit="mm" value="136.0">136 mm</quantity>
cl
<typeStatus id="FE7B88682620FFF4206DFCB6FE3FFCE7" box="[356,457,804,830]" pageId="9" pageNumber="54" type="paratype">paratype</typeStatus>
are noticeably longer than in other specimens, with the length of the longest spines of the carpi and propodi exceeding, rather than at most equaling, the respective segment heights. Aside from spine length, the more spinous
<typeStatus id="FE7B88682620FFF42579FCE6FB23FC57" box="[1136,1237,884,910]" pageId="9" pageNumber="54" type="paratype">paratype</typeStatus>
agrees well with other specimens of
<taxonomicName id="E6C04D492620FFF42371FC0EFCFFFC6C" box="[632,777,924,949]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82620FFF42371FC0EFCFFFC6C" box="[632,777,924,949]" italics="true" pageId="9" pageNumber="54">N. yaldwyni</emphasis>
</taxonomicName>
. One might speculate that the more spinous
<typeStatus id="FE7B88682620FFF42001FC56FE9BFC07" box="[264,365,964,990]" pageId="9" pageNumber="54" type="paratype">paratype</typeStatus>
of
<taxonomicName id="E6C04D492620FFF4209DFC56FDD7FC04" box="[404,545,964,989]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82620FFF4209DFC56FDD7FC04" box="[404,545,964,989]" italics="true" pageId="9" pageNumber="54">N. yaldwyni</emphasis>
</taxonomicName>
represents a large specimen of
<taxonomicName id="E6C04D492620FFF4229BFC56FAD1FC07" authority="Macpherson, 1988" authorityName="Macpherson" authorityYear="1988" box="[914,1319,964,990]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="vinogradovi">
<emphasis id="13B4EAD82620FFF4229BFC56FBB4FC04" box="[914,1090,964,989]" italics="true" pageId="9" pageNumber="54">N. vinogradovi</emphasis>
Macpherson, 1988
</taxonomicName>
. Aside from the much longer dorsal spines of
<taxonomicName id="E6C04D492620FFF4224CFC7EFC0DFBDC" box="[837,1019,1004,1029]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="vinogradovi">
<emphasis id="13B4EAD82620FFF4224CFC7EFC0DFBDC" box="[837,1019,1004,1029]" italics="true" pageId="9" pageNumber="54">N. vinogradovi</emphasis>
</taxonomicName>
, however, Macphersons species is immediately excluded by the length of the merus of the third ambulatory leg in males—longer than the carapace in
<taxonomicName id="E6C04D492620FFF423A4FBAEFCAAFB8C" box="[685,860,1084,1109]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="vinogradovi">
<emphasis id="13B4EAD82620FFF423A4FBAEFCAAFB8C" box="[685,860,1084,1109]" italics="true" pageId="9" pageNumber="54">N. vinogradovi</emphasis>
</taxonomicName>
; shorter than carapace in
<taxonomicName id="E6C04D492620FFF42583FBAEFAE0FB8C" box="[1162,1302,1084,1109]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82620FFF42583FBAEFAE0FB8C" box="[1162,1302,1084,1109]" italics="true" pageId="9" pageNumber="54">N. yaldwyni</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="217F36CA2620FFF42031FBF6FB8EFB2F" blockId="9.[264,1324,284,1870]" pageId="9" pageNumber="54">
The
<quantity id="E6389B2F2620FFF4207CFBF6FE29FBA7" box="[373,479,1124,1150]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="54" unit="mm" value="120.0">120 mm</quantity>
cl
<typeStatus id="FE7B88682620FFF4231AFBF6FD8EFBA7" box="[531,632,1124,1150]" pageId="9" pageNumber="54" type="paratype">paratype</typeStatus>
(NIWA 3433,
<figureCitation id="B9FB2A4F2620FFF42236FBF6FC7DFBA7" box="[831,907,1124,1150]" captionStart="FIGURE 3" captionStartId="5.[264,367,1767,1791]" captionTargetBox="[264,1323,591,1755]" captionTargetId="figure@5.[264,1323,591,1758]" captionTargetPageId="5" captionText="FIGURE 3. Neolithodes yaldwyni sp. nov. A D, male holotype (cl 141.5 mm, cw 124.1 mm, NIWA 3434). E F, male paratype (cl 120.0 mm, cw 108.7 mm, NIWA 3433). A, left cheliped, dorsal view. B, left chela, outer surface. C, right cheliped, dorsal view. D, right chela, outer surface. E, left chela, outer surface. F, right chela, outer surface." httpUri="https://zenodo.org/record/173682/files/figure.png" pageId="9" pageNumber="54">Fig. 3</figureCitation>
E, F) is unusual in the reversed asymmetry of the chelipeds whereby the left chela is larger than the right, forming the crushing claw. The right chela is usually the larger in
<taxonomicName id="E6C04D492620FFF422FAFB26FB83FB17" box="[1011,1141,1204,1230]" class="Malacostraca" family="Lithodidae" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="family">Lithodidae</taxonomicName>
, and reversed asymmetry is perhaps a result of autotomy and replacement (
<bibRefCitation id="45514B3B2620FFF422DBFB4EFB9AFB2F" author="Zaklan" box="[978,1132,1244,1270]" pageId="9" pageNumber="68" refString="Zaklan, S. (2000) A case of reversed asymmetry in Lithodes maja (Linnaeus, 1758) (Decapoda, Anomura, Lithodidae). Crustaceana, 73 (8), 1019 - 1022." type="journal article" year="2000">Zaklan 2000</bibRefCitation>
).
</paragraph>
<paragraph id="217F36CA2620FFF72031FA96FD68FE5F" blockId="9.[264,1324,284,1870]" lastBlockId="10.[264,1323,284,670]" lastPageId="10" lastPageNumber="55" pageId="9" pageNumber="54">
Though
<bibRefCitation id="45514B3B2620FFF42095FA96FD6BFAC7" author="Thatje" box="[412,669,1284,1310]" pageId="9" pageNumber="68" refString="Thatje, S. &amp; Lorz, A. - N. (2005) First record of lithodid crabs from Antarctic waters off the Balleny Islands. Polar Biology, 28, 334 - 337." type="journal article" year="2005">Thatje &amp; Lörz (2005)</bibRefCitation>
confused
<taxonomicName id="E6C04D492620FFF42210FA96FC51FAC4" box="[793,935,1284,1309]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82620FFF42210FA96FC51FAC4" box="[793,935,1284,1309]" italics="true" pageId="9" pageNumber="54">N. yaldwyni</emphasis>
</taxonomicName>
with
<taxonomicName id="E6C04D492620FFF422E5FA96FB92FAC4" box="[1004,1124,1284,1309]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="brodiei">
<emphasis id="13B4EAD82620FFF422E5FA96FB92FAC4" box="[1004,1124,1284,1309]" italics="true" pageId="9" pageNumber="54">N. brodiei</emphasis>
</taxonomicName>
, the two species are readily separated.
<bibRefCitation id="45514B3B2620FFF42311FABEFCD2FA9F" author="Thatje" box="[536,804,1324,1350]" pageId="9" pageNumber="68" refString="Thatje, S. &amp; Lorz, A. - N. (2005) First record of lithodid crabs from Antarctic waters off the Balleny Islands. Polar Biology, 28, 334 - 337." type="journal article" year="2005">Thatje &amp; Lörz (2005)</bibRefCitation>
noted the differences in secondary dorsal spinulation between typical
<taxonomicName id="E6C04D492620FFF42357FAC6FD2EFAB4" box="[606,728,1364,1389]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="brodiei">
<emphasis id="13B4EAD82620FFF42357FAC6FD2EFAB4" box="[606,728,1364,1389]" italics="true" pageId="9" pageNumber="54">N. brodiei</emphasis>
</taxonomicName>
and the Ross Sea specimens, which is the most obvious distinguishing feature. Secondary spinules or tubercles are only sparsely distributed on the dorsum of
<taxonomicName id="E6C04D492620FFF4237CFA36FCF0FA64" box="[629,774,1444,1469]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82620FFF4237CFA36FCF0FA64" box="[629,774,1444,1469]" italics="true" pageId="9" pageNumber="54">N. yaldwyni</emphasis>
</taxonomicName>
but densely and uniformly distributed in
<taxonomicName id="E6C04D492620FFF42419FA36FEADFA3C" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="brodiei">
<emphasis id="13B4EAD82620FFF42419FA36FEADFA3C" italics="true" pageId="9" pageNumber="54">N. brodiei</emphasis>
</taxonomicName>
. The differences in dorsal spinulation between
<taxonomicName id="E6C04D492620FFF42280FA5EFBE0FA3C" box="[905,1046,1484,1509]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82620FFF42280FA5EFBE0FA3C" box="[905,1046,1484,1509]" italics="true" pageId="9" pageNumber="54">N. yaldwyni</emphasis>
</taxonomicName>
and
<taxonomicName id="E6C04D492620FFF42546FA5EFB33FA3C" box="[1103,1221,1484,1509]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="brodiei">
<emphasis id="13B4EAD82620FFF42546FA5EFB33FA3C" box="[1103,1221,1484,1509]" italics="true" pageId="9" pageNumber="54">N. brodiei</emphasis>
</taxonomicName>
(both as
<taxonomicName id="E6C04D492620FFF42001FA66FE89F9D4" box="[264,383,1524,1549]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="brodiei">
<emphasis id="13B4EAD82620FFF42001FA66FE89F9D4" box="[264,383,1524,1549]" italics="true" pageId="9" pageNumber="54">N. brodiei</emphasis>
</taxonomicName>
) were well illustrated by
<bibRefCitation id="45514B3B2620FFF423B1FA66FBBDF9D7" author="Thatje" box="[696,1099,1524,1550]" pageId="9" pageNumber="68" refString="Thatje, S. &amp; Lorz, A. - N. (2005) First record of lithodid crabs from Antarctic waters off the Balleny Islands. Polar Biology, 28, 334 - 337." type="journal article" year="2005">Thatje &amp; Lörz (2005: fig. 2A, B)</bibRefCitation>
. Although the text discussion attributes the figures correctly, Thatje &amp; Lörzs (2005) caption for figures 2A &amp; B were inadvertently reversed: figure 2A depicts a Balleny Island specimen (the
<typeStatus id="FE7B88682620FFF425CDF9D6FADDF987" box="[1220,1323,1604,1630]" pageId="9" pageNumber="54" type="holotype">holotype</typeStatus>
of
<taxonomicName id="E6C04D492620FFF4202EF9FEFE44F95C" box="[295,434,1644,1669]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82620FFF4202EF9FEFE44F95C" box="[295,434,1644,1669]" italics="true" pageId="9" pageNumber="54">N. yaldwyni</emphasis>
</taxonomicName>
) and figure 2B the
<typeStatus id="FE7B88682620FFF4239CF9FEFD0DF95F" box="[661,763,1644,1670]" pageId="9" pageNumber="54" type="holotype">holotype</typeStatus>
of
<taxonomicName id="E6C04D492620FFF4222BF9FEFC6EF95C" box="[802,920,1644,1669]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="brodiei">
<emphasis id="13B4EAD82620FFF4222BF9FEFC6EF95C" box="[802,920,1644,1669]" italics="true" pageId="9" pageNumber="54">N. brodiei</emphasis>
</taxonomicName>
. Aside from dorsal spinulation,
<taxonomicName id="E6C04D492620FFF42419F9FEFE84F974" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="13B4EAD82620FFF42419F9FEFE84F974" italics="true" pageId="9" pageNumber="54">N. yaldwyni</emphasis>
</taxonomicName>
also differs from
<taxonomicName id="E6C04D492620FFF42349F906FD43F974" box="[576,693,1684,1709]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="brodiei">
<emphasis id="13B4EAD82620FFF42349F906FD43F974" box="[576,693,1684,1709]" italics="true" pageId="9" pageNumber="54">N. brodiei</emphasis>
</taxonomicName>
in numerous other features including the near straight and vertical instead of distinctly concave posterior orbital margin, the smooth instead of minutely spinulose ventral proximal surface of the rostrum, and the subcylindrical instead of dorsoventrally flattened propodi of the ambulatory legs. The smallest specimen reported by
<bibRefCitation id="45514B3B2620FFF4203BF8A6FDC8F897" author="Thatje" box="[306,574,1844,1870]" pageId="9" pageNumber="68" refString="Thatje, S. &amp; Lorz, A. - N. (2005) First record of lithodid crabs from Antarctic waters off the Balleny Islands. Polar Biology, 28, 334 - 337." type="journal article" year="2005">Thatje &amp; Lörz (2005)</bibRefCitation>
as
<taxonomicName id="E6C04D492620FFF42366F8A6FD1CF894" box="[623,746,1844,1869]" class="Malacostraca" family="Lithodidae" genus="Neolithodes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="brodiei">
<emphasis id="13B4EAD82620FFF42366F8A6FD1CF894" box="[623,746,1844,1869]" italics="true" pageId="9" pageNumber="54">N. brodiei</emphasis>
</taxonomicName>
, a
<quantity id="E6389B2F2620FFF4221EF8A6FC99F897" box="[791,879,1844,1870]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="9" pageNumber="54" unit="mm" value="75.0">75 mm</quantity>
cl ovigerous female, is referable to
<taxonomicName id="E6C04D492623FFF72001FE8EFD13FEEF" authority="Macpherson, 1988" authorityName="Macpherson" authorityYear="1988" box="[264,741,284,310]" class="Malacostraca" family="Lithodidae" genus="Paralomis" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="55" phylum="Arthropoda" rank="species" species="birsteini">
<emphasis id="13B4EAD82623FFF72001FE8EFE04FEEC" box="[264,498,284,309]" italics="true" pageId="10" pageNumber="55">Paralomis birsteini</emphasis>
Macpherson, 1988
</taxonomicName>
. The fourth specimen reported
<bibRefCitation id="45514B3B2623FFF7257CFE8EFEAFFE87" author="Thatje" pageId="10" pageNumber="68" refString="Thatje, S. &amp; Lorz, A. - N. (2005) First record of lithodid crabs from Antarctic waters off the Balleny Islands. Polar Biology, 28, 334 - 337." type="journal article" year="2005">Thatje &amp; Lörz (2005)</bibRefCitation>
, a “severely damaged” ovigerous female (NIWA 3435) is referable to
<taxonomicName id="E6C04D492623FFF725B9FED6FE93FE5C" class="Malacostraca" family="Lithodidae" genus="Paralomis" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="55" phylum="Arthropoda" rank="species" species="stevensi" status="sp. nov.">
<emphasis id="13B4EAD82623FFF725B9FED6FE93FE5C" italics="true" pageId="10" pageNumber="55">Paralomis stevensi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="088757A32623FFF72065FEF9FE3CFE5C" box="[364,458,363,389]" pageId="10" pageNumber="55" rank="species">
<emphasis id="13B4EAD82623FFF72065FEF9FE3CFE5C" bold="true" box="[364,458,363,389]" pageId="10" pageNumber="55">sp. nov.</emphasis>
</taxonomicNameLabel>
, described below.
</paragraph>
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<bibRef id="F860A23C2623FFF72031FE06FD0BFDAF" pageId="10" pageNumber="55" type="journal article">
<author id="F97DDE2F2623FFF72031FE06FE3BFE74" box="[312,461,404,430]" fns="I" npo="LF" pageId="10" pageNumber="55">
Presently,
<emphasis id="13B4EAD82623FFF720BBFE06FE3BFE74" box="[434,461,404,429]" italics="true" pageId="10" pageNumber="55">N.</emphasis>
</author>
yaldwyni is known only from the vicinity of Scott Island and the Balleny Islands, Ross Sea (see
<bibRefCitation id="45514B3B2623FFF72323FE2EFD25FE0F" author="Dawson" box="[554,723,444,470]" lastPageId="10" lastPageNumber="68" pageId="10" pageNumber="67" refString="Dawson E. W. (1969) Sturge Island provisional bathymetry. Island Chart Series 1: 200 0 0 0, Bathymetry (New Zealand Oceanographic Institute, Wellington)." type="book chapter" year="1969">
Dawson
<year id="556877992623FFF72391FE2EFD25FE0F" box="[664,723,444,470]" pageId="10" pageNumber="55">1969</year>
</bibRefCitation>
<title id="53C4DDFC2623FFF723E9FE2EFC6CFE0F" box="[736,922,444,470]" jopScore="0.16945253" pageId="10" pageNumber="55" type="_" typeScore="44">for bathymetry</title>
).
<journalOrPublisher id="FB013C062623FFF722BAFE2EFCDEFDFF" jopScore="0.36035272" pageId="10" pageNumber="55">
<emphasis id="13B4EAD82623FFF722BAFE2EFB44FE0C" box="[947,1202,444,469]" italics="true" pageId="10" pageNumber="55">Neolithodes capensis</emphasis>
has been reported from
<collectingCountry id="59D7765A2623FFF720B7FE76FDA8FE27" box="[446,606,484,510]" name="South Africa" pageId="10" pageNumber="55">South Africa</collectingCountry>
, the subantarctic (Banzare Bank, Kerguelen Islands), and
<collectingCountry id="59D7765A2623FFF72001FD9EFE74FDFF" box="[264,386,524,550]" name="Antarctica" pageId="10" pageNumber="55">Antarctica</collectingCountry>
(Bellingshausen Sea) (
<bibRefCitation id="45514B3B2623FFF7239CFD9EFCDEFDFF" author="Macpherson" box="[661,808,524,550]" pageId="10" pageNumber="68" refString="Macpherson, E. (2004) A new species and new records of lithodid crabs (Crustacea: Decapoda: Lithodidae) from the Crozet and Kerguelen Islands area (Subantarctica). Polar Biology, 27, 418 - 422." type="journal article" year="2004">Macpherson</bibRefCitation>
</journalOrPublisher>
<part id="55649C572623FFF72238FD9EFC99FDFF" box="[817,879,524,550]" pageId="10" pageNumber="55" type="volume">2004</part>
;
<bibRefCitation id="45514B3B2623FFF72275FD9EFB6CFDFF" author="Garcia" box="[892,1178,524,550]" pageId="10" pageNumber="68" refString="Garcia Raso, J. E., Manjon-Cabeza, M. E., Ramos, A. &amp; Ilaso, I. (2005) New record of Lithodidae (Crustacea, Decapoda, Anomura) from the Antarctic (Bellingshausen Sea). Polar Biology, 28, 642 - 646." type="journal article" year="2005">
García Raso
<emphasis id="13B4EAD82623FFF7251FFD9EFBBDFDFC" box="[1046,1099,524,549]" italics="true" pageId="10" pageNumber="55">et al</emphasis>
.
<pagination id="1809C83E2623FFF72552FD9EFB6CFDFF" box="[1115,1178,524,550]" pageId="10" pageNumber="55" type="pagination">2005</pagination>
</bibRefCitation>
). In view of the strong similarities between
<emphasis id="13B4EAD82623FFF7237DFDA6FCF6FD94" box="[628,768,564,589]" italics="true" pageId="10" pageNumber="55">N. yaldwyni</emphasis>
and
<emphasis id="13B4EAD82623FFF72231FDA6FC49FD94" box="[824,959,564,589]" italics="true" pageId="10" pageNumber="55">N. capensis</emphasis>
, the
<collectingCountry id="59D7765A2623FFF722FEFDA6FB92FD97" box="[1015,1124,564,590]" name="Antarctica" pageId="10" pageNumber="55">Antarctic</collectingCountry>
and subantarctic records of
<emphasis id="13B4EAD82623FFF7208CFDCEFDFBFDAC" box="[389,525,604,629]" italics="true" pageId="10" pageNumber="55">N. capensis</emphasis>
require verification.
</bibRef>
</paragraph>
</subSubSection>
<subSubSection id="69DA65412623FFF72031FD16FB07FD47" box="[312,1265,644,670]" pageId="10" pageNumber="55" type="distribution">
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<emphasis id="13B4EAD82623FFF72031FD16FE2EFD47" bold="true" box="[312,472,644,670]" pageId="10" pageNumber="55">Distribution.</emphasis>
Presently known only from the Ross Sea,
<collectingCountry id="59D7765A2623FFF722C5FD16FBBCFD47" box="[972,1098,644,670]" name="Antarctica" pageId="10" pageNumber="55">Antarctica</collectingCountry>
;
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.
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