377 lines
33 KiB
XML
377 lines
33 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e90570" ID-Pensoft-Pub="1864-8312-80-649" ID-Pensoft-UUID="FAE6DFEF69735531A4CBA275E078320D" ID-ZooBank="AD0B1CC262414460B6B888E6B95F20D7" ModsDocID="1864-8312-80-649" checkinTime="1671114540108" checkinUser="pensoft" docAuthor="Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S. & Harms, Danilo" docDate="2022" docId="BA67104534735E24B141557B9823E2F6" docLanguage="en" docName="ArthropodSystPhyl 80: 649-691" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-12-14" docSource="http://dx.doi.org/10.3897/asp.80.e90570" docTitle="Mahajanganella Lorenz & Loria & Harvey & Harms 2022, gen. nov." docType="treatment" docVersion="1" id="FAE6DFEF69735531A4CBA275E078320D" lastPageNumber="649" masterDocId="FAE6DFEF69735531A4CBA275E078320D" masterDocTitle="The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots" masterLastPageNumber="691" masterPageNumber="649" pageNumber="649" updateTime="1671114540108" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lorenz, Michelle</mods:namePart>
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<mods:affiliation>Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany & University of Hamburg, Biology Department, Ohnhorststrasse 18, 22609 Hamburg, Germany</mods:affiliation>
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<mods:nameIdentifier type="email">michellelorenz@gmx.net</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Loria, Stephanie F.</mods:namePart>
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<mods:affiliation>Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Harvey, Mark S.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1482-0109</mods:nameIdentifier>
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<mods:affiliation>Western Australian Museum, Collections & Research, 49 Kew Street, Welshpool, WA 6106, Australia & University of Western Australia, School of Biological Sciences, Crawley WA 6009, Australia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Harms, Danilo</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7189-5345</mods:nameIdentifier>
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<mods:affiliation>Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-12-14</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>80</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>649</mods:start>
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<mods:end>691</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/asp.80.e90570</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e90570</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1864-8312-80-649</mods:identifier>
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<mods:identifier type="ZooBank">AD0B1CC262414460B6B888E6B95F20D7</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FAE6DFEF69735531A4CBA275E078320D</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:NomEnClAturAlACts/CACF3D4C-1557-46AB-AB1D-F2C9829023A4" httpUri="http://treatment.plazi.org/id/BA67104534735E24B141557B9823E2F6" lastPageNumber="649" pageId="0" pageNumber="649">
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<subSubSection pageId="0" pageNumber="649" type="nomenclature">
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<paragraph pageId="0" pageNumber="649">
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<taxonomicName LSID="https://zoobank.org/NomenclaturalActs/CACF3D4C-1557-46AB-AB1D-F2C9829023A4" authority="Lorenz & Loria & Harvey & Harms, 2022" authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Mahajanganella" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Mahajanganella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus" status="gen. nov.">Mahajanganella</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="649">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="649" type="type species">
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<paragraph pageId="0" pageNumber="649">Type species.</paragraph>
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<paragraph pageId="0" pageNumber="649">
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<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Mahajanganella" kingdom="Animalia" lsidName="Mahajanganella heraclis" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="species" species="heraclis">
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<emphasis italics="true" pageId="0" pageNumber="649">Mahajanganella heraclis</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="649" type="diagnosis">
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<paragraph pageId="0" pageNumber="649">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="649">
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<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Mahajanganella" kingdom="Animalia" lsidName="Mahajanganella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="649">Mahajanganella</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
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is morphologically unique by having the following characters, which differ from other feaelloid members in Madagascar: 5 specialized setae on the retrolateral face of the movable chelal finger arranged in a group between trichobothrium
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<emphasis italics="true" pageId="0" pageNumber="649">b</emphasis>
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and terminal teeth (arranged in a row in
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<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Antsirananaella" kingdom="Animalia" lsidName="Antsirananaella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="649">Antsirananaella</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
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); smaller overall size than
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<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Antsirananaella" kingdom="Animalia" lsidName="Antsirananaella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="649">Antsirananaella</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
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; stronger granulate cuticle than
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<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Antsirananaella" kingdom="Animalia" lsidName="Antsirananaella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="649">Antsirananaella</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
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; all four anterior lobes of carapace with the same distance to each other (smaller distance between anteriomedial lobes than to anteriolateral ones in
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<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Antsirananaella" kingdom="Animalia" lsidName="Antsirananaella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="649">Antsirananaella</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
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). Like
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<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Antsirananaella" kingdom="Animalia" lsidName="Antsirananaella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="649">Antsirananaella</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
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and
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<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Toliaranella" kingdom="Animalia" lsidName="Toliaranella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="649">Toliaranella</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
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, it differs from
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<taxonomicName authorityName="Judson" authorityYear="2017" class="Arachnida" family="Feaellidae" genus="Cybella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybella" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="649">Cybella</emphasis>
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</taxonomicName>
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by having platelets on the pleural membrane (absent in
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<taxonomicName authorityName="Judson" authorityYear="2017" class="Arachnida" family="Feaellidae" genus="Cybella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybella" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="649">Cybella</emphasis>
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</taxonomicName>
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), from
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<taxonomicName authorityName="Harvey, Andrade & Pinto-da-Rocha" authorityYear="2016" class="Arachnida" family="Feaellidae" genus="Iporangella" kingdom="Animalia" lsidName="Iporangella" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="649">Iporangella</emphasis>
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</taxonomicName>
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by the presence of specialized setae on the movable chelal finger (absent in
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<taxonomicName authorityName="Harvey, Andrade & Pinto-da-Rocha" authorityYear="2016" class="Arachnida" family="Feaellidae" genus="Iporangella" kingdom="Animalia" lsidName="Iporangella" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="649">Iporangella</emphasis>
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</taxonomicName>
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), from
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<taxonomicName authority="" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (Difeaella)" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="subGenus" subGenus="Difeaella">Feaella (Difeaella)</taxonomicName>
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and
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<taxonomicName authority="(Feaella)" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (Feaella)" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="subGenus" subGenus="Feaella">Feaella (Feaella)</taxonomicName>
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by the presence of four anterior carapaceal lobes (two and six, respectively), from
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<taxonomicName authority="" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (Tetrafeaella)" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="subGenus" subGenus="Tetrafeaella">Feaella (Tetrafeaella)</taxonomicName>
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in continental Africa by having a less pronounced depression on the base of coxa I and on top of coxa II (distinctly more pronounced in
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<taxonomicName authorityName="Tullgren" authorityYear="1907" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (T.) mucronata" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="mucronata" subGenus="T.">Feaella (T.) cf. mucronata</taxonomicName>
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), and from the Australian species presently attributed to
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<taxonomicName authority="" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (Tetrafeaella)" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="subGenus" subGenus="Tetrafeaella">Feaella (Tetrafeaella)</taxonomicName>
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by having fewer coxal spines (one pair versus three or four in the Australian taxa).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="649" type="etymology">
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<paragraph pageId="0" pageNumber="649">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="649">The genus is named after the former Mahajanga Province, where many of these feaellids occur. The gender is feminine.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="649" type="description">
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<paragraph pageId="0" pageNumber="649">Description.</paragraph>
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<paragraph pageId="0" pageNumber="649">
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The following description is based on holotype and allotype of
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<taxonomicName lsidName="M. heraclis" pageId="0" pageNumber="649" rank="species" species="heraclis">
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<emphasis italics="true" pageId="0" pageNumber="649">M. heraclis</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
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-
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<emphasis italics="true" pageId="0" pageNumber="649">Carapace</emphasis>
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(Figs
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Mahajanganella gen. nov., scanning electron micrographs. Specimens used for imaging: A M. fridakahloae sp. nov. female paratype (ZMH-A 0016709); B, C M. schwarzeneggeri sp. nov. male paratype (ZMH-A 0016717). A carapace, dorsal; B sternum ventral; C coxal spines (highlighted in red). Scale bars: 0.2 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/asp.80.e90570.figure12" httpUri="https://binary.pensoft.net/fig/782004" pageId="0" pageNumber="649">12A</figureCitation>
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,
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Drawings of Mahajanganella heraclis sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure14" httpUri="https://binary.pensoft.net/fig/782006" pageId="0" pageNumber="649">14A</figureCitation>
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,
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Drawings of Mahajanganella fridakahloae sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure15" httpUri="https://binary.pensoft.net/fig/782007" pageId="0" pageNumber="649">15A</figureCitation>
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,
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Drawings of Mahajanganella schwarzeneggeri sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure16" httpUri="https://binary.pensoft.net/fig/782008" pageId="0" pageNumber="649">16A</figureCitation>
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): Moderate granulate and all four anterior lobes with same distance to each other. -
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<emphasis italics="true" pageId="0" pageNumber="649">Pedipalp</emphasis>
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(Figs
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Mahajanganella gen. nov., habitus. Specimen used for imaging: M. schwarzeneggeri sp. nov. allotype (ZMH-A 0016716). A dorsal; B ventral; C lateral. Scale bar: 0.2 mm." figureDoi="10.3897/asp.80.e90570.figure11" httpUri="https://binary.pensoft.net/fig/782003" pageId="0" pageNumber="649">11A, B</figureCitation>
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,
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Mahajanganella gen. nov., scanning electron micrographs. Specimens used for imaging: A M. heraclis sp. nov. male paratype (ZMH-A 0016701); B M. fridakahloae sp. nov. male paratype (ZMH-A 0016715); C M. schwarzeneggeri sp. nov. male paratype (ZMH-A 0016717). A right chela from ventral with specialized setae in a group in detail (highlighted in red); B left chela with double trichobothrium dt in detail (highlighted in red); C left chela from medial with terminal teeth in a compact group in detail. Scale bar: 0.2 mm." figureDoi="10.3897/asp.80.e90570.figure13" httpUri="https://binary.pensoft.net/fig/782005" pageId="0" pageNumber="649">13A, B, C</figureCitation>
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,
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Drawings of Mahajanganella heraclis sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure14" httpUri="https://binary.pensoft.net/fig/782006" pageId="0" pageNumber="649">14C, D</figureCitation>
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,
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Drawings of Mahajanganella fridakahloae sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure15" httpUri="https://binary.pensoft.net/fig/782007" pageId="0" pageNumber="649">15C, D</figureCitation>
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,
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Drawings of Mahajanganella schwarzeneggeri sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure16" httpUri="https://binary.pensoft.net/fig/782008" pageId="0" pageNumber="649">16C, D</figureCitation>
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): Trichobothrial pattern:
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<emphasis italics="true" pageId="0" pageNumber="649">esb</emphasis>
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between
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<emphasis italics="true" pageId="0" pageNumber="649">st</emphasis>
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and
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<emphasis italics="true" pageId="0" pageNumber="649">ist</emphasis>
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on the sagittal level with
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<emphasis italics="true" pageId="0" pageNumber="649">ist</emphasis>
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slightly more distal and
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<emphasis italics="true" pageId="0" pageNumber="649">st</emphasis>
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slightly more proximal than
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<emphasis italics="true" pageId="0" pageNumber="649">esb</emphasis>
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;
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<emphasis italics="true" pageId="0" pageNumber="649">t</emphasis>
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distinctly closer to
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<emphasis italics="true" pageId="0" pageNumber="649">sb</emphasis>
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than to
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<emphasis italics="true" pageId="0" pageNumber="649">b</emphasis>
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;
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<emphasis italics="true" pageId="0" pageNumber="649">sb</emphasis>
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distinctly closer to
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<emphasis italics="true" pageId="0" pageNumber="649">t</emphasis>
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than to
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<emphasis italics="true" pageId="0" pageNumber="649">st</emphasis>
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;
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<emphasis italics="true" pageId="0" pageNumber="649">it</emphasis>
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on the fixed finger between
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<emphasis italics="true" pageId="0" pageNumber="649">eb</emphasis>
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and
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<emphasis italics="true" pageId="0" pageNumber="649">et</emphasis>
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distinctly closer to
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<emphasis italics="true" pageId="0" pageNumber="649">eb</emphasis>
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than to
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<emphasis italics="true" pageId="0" pageNumber="649">et.</emphasis>
|
|
Movable finger with five specialized setae on the ventral face, arranged in a group and slightly more distally situated than
|
|
<emphasis italics="true" pageId="0" pageNumber="649">b</emphasis>
|
|
(Fig.
|
|
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Mahajanganella gen. nov., scanning electron micrographs. Specimens used for imaging: A M. heraclis sp. nov. male paratype (ZMH-A 0016701); B M. fridakahloae sp. nov. male paratype (ZMH-A 0016715); C M. schwarzeneggeri sp. nov. male paratype (ZMH-A 0016717). A right chela from ventral with specialized setae in a group in detail (highlighted in red); B left chela with double trichobothrium dt in detail (highlighted in red); C left chela from medial with terminal teeth in a compact group in detail. Scale bar: 0.2 mm." figureDoi="10.3897/asp.80.e90570.figure13" httpUri="https://binary.pensoft.net/fig/782005" pageId="0" pageNumber="649">13A</figureCitation>
|
|
). Terminal teeth varying in number, both in fixed and the movable fingers each with 7-8 terminal teeth (one large tooth included on fixed finger). -
|
|
<emphasis italics="true" pageId="0" pageNumber="649">Chelicera</emphasis>
|
|
: Most specimens were too dirty to get secure information about characters.
|
|
<emphasis italics="true" pageId="0" pageNumber="649">Coxal region</emphasis>
|
|
(Fig.
|
|
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Mahajanganella gen. nov., scanning electron micrographs. Specimens used for imaging: A M. fridakahloae sp. nov. female paratype (ZMH-A 0016709); B, C M. schwarzeneggeri sp. nov. male paratype (ZMH-A 0016717). A carapace, dorsal; B sternum ventral; C coxal spines (highlighted in red). Scale bars: 0.2 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/asp.80.e90570.figure12" httpUri="https://binary.pensoft.net/fig/782004" pageId="0" pageNumber="649">12B, C</figureCitation>
|
|
),
|
|
<emphasis italics="true" pageId="0" pageNumber="649">legs</emphasis>
|
|
(Figs
|
|
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Mahajanganella gen. nov., habitus. Specimen used for imaging: M. schwarzeneggeri sp. nov. allotype (ZMH-A 0016716). A dorsal; B ventral; C lateral. Scale bar: 0.2 mm." figureDoi="10.3897/asp.80.e90570.figure11" httpUri="https://binary.pensoft.net/fig/782003" pageId="0" pageNumber="649">11B</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Drawings of Mahajanganella heraclis sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure14" httpUri="https://binary.pensoft.net/fig/782006" pageId="0" pageNumber="649">14B</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Drawings of Mahajanganella fridakahloae sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure15" httpUri="https://binary.pensoft.net/fig/782007" pageId="0" pageNumber="649">15B</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Drawings of Mahajanganella schwarzeneggeri sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure16" httpUri="https://binary.pensoft.net/fig/782008" pageId="0" pageNumber="649">16B</figureCitation>
|
|
),
|
|
<emphasis italics="true" pageId="0" pageNumber="649">abdomen</emphasis>
|
|
(Fig.
|
|
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Mahajanganella gen. nov., habitus. Specimen used for imaging: M. schwarzeneggeri sp. nov. allotype (ZMH-A 0016716). A dorsal; B ventral; C lateral. Scale bar: 0.2 mm." figureDoi="10.3897/asp.80.e90570.figure11" httpUri="https://binary.pensoft.net/fig/782003" pageId="0" pageNumber="649">11A, B, C</figureCitation>
|
|
) and
|
|
<emphasis italics="true" pageId="0" pageNumber="649">genital region</emphasis>
|
|
(Fig.
|
|
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Mahajanganella gen. nov., scanning electron micrographs. Specimens used for imaging: A M. fridakahloae sp. nov. female paratype (ZMH-A 0016709); B, C M. schwarzeneggeri sp. nov. male paratype (ZMH-A 0016717). A carapace, dorsal; B sternum ventral; C coxal spines (highlighted in red). Scale bars: 0.2 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/asp.80.e90570.figure12" httpUri="https://binary.pensoft.net/fig/782004" pageId="0" pageNumber="649">12B</figureCitation>
|
|
) with the same characters as
|
|
<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Antsirananaella" kingdom="Animalia" lsidName="Antsirananaella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">Antsirananaella</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
|
|
</paragraph>
|
|
<caption doi="10.3897/asp.80.e90570.figure11" httpUri="https://binary.pensoft.net/fig/782003" pageId="0" pageNumber="649" start="Figure 11" startId="F11">
|
|
<paragraph pageId="0" pageNumber="649">
|
|
<emphasis bold="true" pageId="0" pageNumber="649">Figure 11.</emphasis>
|
|
<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Mahajanganella" kingdom="Animalia" lsidName="Mahajanganella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">Mahajanganella</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
|
|
, habitus. Specimen used for imaging:
|
|
<taxonomicName lsidName="M. schwarzeneggeri" pageId="0" pageNumber="649" rank="species" species="schwarzeneggeri">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">M. schwarzeneggeri</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
allotype (ZMH-A0016716).
|
|
<emphasis bold="true" pageId="0" pageNumber="649">A</emphasis>
|
|
dorsal;
|
|
<emphasis bold="true" pageId="0" pageNumber="649">B</emphasis>
|
|
ventral;
|
|
<emphasis bold="true" pageId="0" pageNumber="649">C</emphasis>
|
|
lateral. Scale bar: 0.2 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/asp.80.e90570.figure12" httpUri="https://binary.pensoft.net/fig/782004" pageId="0" pageNumber="649" start="Figure 12" startId="F12">
|
|
<paragraph pageId="0" pageNumber="649">
|
|
<emphasis bold="true" pageId="0" pageNumber="649">Figure 12.</emphasis>
|
|
<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Mahajanganella" kingdom="Animalia" lsidName="Mahajanganella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">Mahajanganella</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
|
|
, scanning electron micrographs. Specimens used for imaging:
|
|
<emphasis bold="true" pageId="0" pageNumber="649">A</emphasis>
|
|
<taxonomicName lsidName="M. fridakahloae" pageId="0" pageNumber="649" rank="species" species="fridakahloae">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">M. fridakahloae</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
female paratype (ZMH-A0016709);
|
|
<emphasis bold="true" pageId="0" pageNumber="649">B</emphasis>
|
|
,
|
|
<emphasis bold="true" pageId="0" pageNumber="649">C</emphasis>
|
|
<taxonomicName lsidName="M. schwarzeneggeri" pageId="0" pageNumber="649" rank="species" species="schwarzeneggeri">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">M. schwarzeneggeri</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
male paratype (ZMH-A0016717).
|
|
<emphasis bold="true" pageId="0" pageNumber="649">A</emphasis>
|
|
carapace, dorsal;
|
|
<emphasis bold="true" pageId="0" pageNumber="649">B</emphasis>
|
|
sternum ventral;
|
|
<emphasis bold="true" pageId="0" pageNumber="649">C</emphasis>
|
|
coxal spines (highlighted in red). Scale bars: 0.2 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="649">A</emphasis>
|
|
,
|
|
<emphasis bold="true" pageId="0" pageNumber="649">B</emphasis>
|
|
); 0.1 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="649">C</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/asp.80.e90570.figure13" httpUri="https://binary.pensoft.net/fig/782005" pageId="0" pageNumber="649" start="Figure 13" startId="F13">
|
|
<paragraph pageId="0" pageNumber="649">
|
|
<emphasis bold="true" pageId="0" pageNumber="649">Figure 13.</emphasis>
|
|
<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Mahajanganella" kingdom="Animalia" lsidName="Mahajanganella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">Mahajanganella</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
|
|
, scanning electron micrographs. Specimens used for imaging:
|
|
<emphasis bold="true" pageId="0" pageNumber="649">A</emphasis>
|
|
<taxonomicName lsidName="M. heraclis" pageId="0" pageNumber="649" rank="species" species="heraclis">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">M. heraclis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
male paratype (ZMH-A0016701);
|
|
<emphasis bold="true" pageId="0" pageNumber="649">B</emphasis>
|
|
<taxonomicName lsidName="M. fridakahloae" pageId="0" pageNumber="649" rank="species" species="fridakahloae">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">M. fridakahloae</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
male paratype (ZMH-A0016715);
|
|
<emphasis bold="true" pageId="0" pageNumber="649">C</emphasis>
|
|
<taxonomicName lsidName="M. schwarzeneggeri" pageId="0" pageNumber="649" rank="species" species="schwarzeneggeri">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">M. schwarzeneggeri</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
male paratype (ZMH-A0016717).
|
|
<emphasis bold="true" pageId="0" pageNumber="649">A</emphasis>
|
|
right chela from ventral with specialized setae in a group in detail (highlighted in red);
|
|
<emphasis bold="true" pageId="0" pageNumber="649">B</emphasis>
|
|
left chela with double trichobothrium
|
|
<emphasis italics="true" pageId="0" pageNumber="649">dt</emphasis>
|
|
in detail (highlighted in red);
|
|
<emphasis bold="true" pageId="0" pageNumber="649">C</emphasis>
|
|
left chela from medial with terminal teeth in a compact group in detail. Scale bar: 0.2 mm.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="649" type="included species">
|
|
<paragraph pageId="0" pageNumber="649">Included species.</paragraph>
|
|
<paragraph pageId="0" pageNumber="649">
|
|
<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Mahajanganella" kingdom="Animalia" lsidName="Mahajanganella fridakahloae" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="species" species="fridakahloae">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">Mahajanganella fridakahloae</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
;
|
|
<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Mahajanganella" kingdom="Animalia" lsidName="Mahajanganella heraclis" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="species" species="heraclis">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">Mahajanganella heraclis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
; and
|
|
<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Mahajanganella" kingdom="Animalia" lsidName="Mahajanganella schwarzeneggeri" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="species" species="schwarzeneggeri">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">Mahajanganella schwarzeneggeri</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="649" type="distribution">
|
|
<paragraph pageId="0" pageNumber="649">Distribution.</paragraph>
|
|
<paragraph pageId="0" pageNumber="649">
|
|
Specimens of
|
|
<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Mahajanganella" kingdom="Animalia" lsidName="Mahajanganella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">Mahajanganella</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
|
|
have been found in dry localities (dry tropical forests) in the Boeny and Menabe Regions (formerly Mahajanga and Toliara Provinces) along the western coast of Madagascar.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |