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<mods:title id="34A2FDB7E7F68C6CCD075960F921F66D">Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records</mods:title>
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<taxonomicName id="C649122FF2A84D9483B1991BE8C47F17" ID-CoL="53FGV" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" class="Insecta" family="Scarabaeidae" genus="Sulcophanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sulcophanaeus faunus" order="Coleoptera" pageId="229" pageNumber="230" phylum="Arthropoda" rank="species" species="faunus">Sulcophanaeus faunus (Fabricius, 1775)</taxonomicName>
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<paragraph id="306DDBF66D65EA2CBE1B4DCB763164F3" pageId="229" pageNumber="230">
<taxonomicName id="8B3A7C04DB2EB4DE84FC6D26AEDABAFB" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scarabaeus faunus" order="Coleoptera" pageId="229" pageNumber="230" phylum="Arthropoda" rank="species" species="faunus">Scarabaeus faunus</taxonomicName>
Fabricius, 1775: 23 (original description. Type locality: Cayennae).
</paragraph>
<paragraph id="A7FBB0662D12474F9E71DCEA651940E2" pageId="229" pageNumber="230">
<taxonomicName id="07235E6B549E7756FA87CD715B62F8B5" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scarabaeus faunus" order="Coleoptera" pageId="229" pageNumber="230" phylum="Arthropoda" rank="species" species="faunus">Scarabaeus faunus</taxonomicName>
:
<bibRefCitation id="A8CDE1A621236B51F1EA6276A266169F" author="Olivier, AG" journalOrPublisher="Baudouin, Paris" pageId="274" pageNumber="275" pagination="1 - 190" title="Entomologie, ou histoire naturelle des insectes, avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Coleopteres." volume="1" year="1789">Olivier 1789</bibRefCitation>
: 103 (redescription).
</paragraph>
<paragraph id="F3C934B45A0BC7A074217DB9FAEA4C98" pageId="229" pageNumber="230">
<taxonomicName id="3F6BF6417121DD40A64D24D8F6B365C3" class="Insecta" family="Scarabaeidae" genus="Copris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copris faunus" order="Coleoptera" pageId="229" pageNumber="230" phylum="Arthropoda" rank="species" species="faunus">Copris faunus</taxonomicName>
:
<bibRefCitation id="41B6C5660B2CFA385B328A663449C347" author="Olivier, AG" journalOrPublisher="Paris, 5" pageId="274" pageNumber="275" title="Encyclopedie methodique. Dictionnaire des Insectes. Pankouke." year="1790">Olivier 1790</bibRefCitation>
: 154 (new combination under the genus
<taxonomicName id="C6F60F6B402E307FFB81BEE1FA6E3365" class="Insecta" family="Scarabaeidae" genus="Copris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copris" order="Coleoptera" pageId="229" pageNumber="230" phylum="Arthropoda" rank="genus">Copris</taxonomicName>
<normalizedToken id="F64266ED0ACC617E31168BD52DDDF8F6" originalValue="“Müller">&quot;Mueller</normalizedToken>
, 1764&quot;, redescription);
<bibRefCitation id="B9A054A2B1FF4159381E69760F45408B" author="Sturm, J" journalOrPublisher="Nuernberg" pageId="278" pageNumber="279" pagination="1 - 136" title="Abbildungen zu Karl Illiger's Uebersetzung von Olivier's Entomologie, oder, Naturgeschichte der Insecten: mit ihren Gattungs-und Artmerkmalen ihrer Beschreibung und Synonymie." url="https://doi.org/10.5962/bhl.title.12457" volume="1" year="1802">Sturm 1802</bibRefCitation>
: 62 (redescription).
</paragraph>
<paragraph id="D0579DE9FB6DED6F0AEC07EC6B741EE7" pageId="229" pageNumber="230">
<taxonomicName id="B37C8B4A547B5BCAE6BBD9C8965E795A" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus faunus" order="Coleoptera" pageId="229" pageNumber="230" phylum="Arthropoda" rank="species" species="faunus">Phanaeus faunus</taxonomicName>
:
<bibRefCitation id="5138C7CB092CD2AEDB8B300A40BB8859" pageId="229" pageNumber="230">Gemminger and Harold 1869</bibRefCitation>
: 1017 (list, distribution, cited as new combination and mentioned as
<taxonomicName id="9C140045DE48DE40CEBCCEB03B509F65" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus Faunus" order="Coleoptera" pageId="229" pageNumber="230" phylum="Arthropoda" rank="species" species="Faunus">Phanaeus Faunus</taxonomicName>
Fabr);
<bibRefCitation id="6B47BDFA750237B76D94BF0B500E8080" author="Nevinson, BG" journalOrPublisher="Published privately, London" pageId="274" pageNumber="275" title="Revised synonymic list of species in the genera Phanaeus Macleay and Oxysternon Castelnau." url="https://doi.org/10.5962/bhl.title.9416" year="1892">Nevinson 1892</bibRefCitation>
: 4 (list of species of the genus
<taxonomicName id="2806957667D857AA075C3299777B639B" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus" order="Coleoptera" pageId="229" pageNumber="230" phylum="Arthropoda" rank="genus">Phanaeus</taxonomicName>
, mentioned as
<taxonomicName id="8182DE273DF60CFE1B569BEEECB49B44" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus Faunus" order="Coleoptera" pageId="229" pageNumber="230" phylum="Arthropoda" rank="species" species="Faunus">Phanaeus Faunus</taxonomicName>
Fabricius);
<bibRefCitation id="6675361E9F7FA72A557115FA3F573ACB" author="Gillet, JJE" journalOrPublisher="IW Junk, Berlin, Germany" pageId="268" pageNumber="269" title="Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae." year="1911 a">Gillet 1911a</bibRefCitation>
: 83 (complete list of species);
<normalizedToken id="9AB2593F1903638F9DF1421460986948" originalValue="dOlsoufieff">d'Olsoufieff</normalizedToken>
, 1924: 32 (characters in key), 80 (diagnosis), 146 (distribution);
<bibRefCitation id="396D812FD7B2D2C05713924292853DBF" author="Blackwelder, RE" journalOrPublisher="National Museum Bulletin" pageId="262" pageNumber="263" pagination="189 - 341" title="Checklist of the coleopterous insects of Mexico, Central America, West Indies and South America. Part 2." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
: 209 (list of species from Latin America).
</paragraph>
<paragraph id="650CB16F9FCC7D5944E27F19B2233FEF" pageId="229" pageNumber="230">
<taxonomicName id="8B8FEFE690F52941BEDD96339D00B5E0" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus (Phanaeus) faunus" order="Coleoptera" pageId="229" pageNumber="230" phylum="Arthropoda" rank="species" species="faunus" subGenus="Phanaeus">Phanaeus (Phanaeus) faunus</taxonomicName>
:
<bibRefCitation id="8FED461A286587837C2E026B3F2CBAE4" author="Pessoa, SB" journalOrPublisher="Annales Facultad Medical Sao Paulo" pageId="275" pageNumber="276" pagination="279 - 314" title="Contribuicao para o conhecimento das especies brasileiras do genero Phanaeus." volume="10" year="1934">
<normalizedToken id="63C3EB93414DB6B8E2A49FD501C4A039" originalValue="Pessôa">Pessoa</normalizedToken>
1934
</bibRefCitation>
: 304 (characters in key) 305 (redescription);
<bibRefCitation id="4B953D34DB731F0AA9C8857743F80DEE" author="Vulcano, MA" journalOrPublisher="Atas do Simposio sobre a Biota Amazonica" pageId="279" pageNumber="280" pagination="533 - 603" title="Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera)." volume="5" year="1967">Vulcano and Pereira 1967</bibRefCitation>
: 572 (characters in key).
</paragraph>
<paragraph id="A180D51CCAAB0A9770770F382CBBFCB5" pageId="229" pageNumber="230">
<taxonomicName id="1C2E5307602282A92633F4CA3CF4D3B4" class="Insecta" family="Scarabaeidae" genus="Sulcophanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sulcophanaeus faunus" order="Coleoptera" pageId="229" pageNumber="230" phylum="Arthropoda" rank="species" species="faunus">Sulcophanaeus faunus</taxonomicName>
:
<bibRefCitation id="CF91C41FC36E80C7F66903036DCBA122" author="Edmonds, WD" journalOrPublisher="Folia Herovskyana Supplementum" pageId="266" pageNumber="267" pagination="1 - 60" title="Revision of the Neotropical dung beetle genus Sulcophanaeus." volume="6" year="2000">Edmonds 2000</bibRefCitation>
: 8 (cited as new combination, characters in key), 9 (diagnosis);
<bibRefCitation id="ABDF8AD6B3ED5C53DC936722C37EA203" author="Vaz-de-Mello, FZ" editor="Martin-Piera, F" journalOrPublisher="Sociedad Entomologica Aragonesa" pageId="278" pageNumber="279" pagination="183 - 195" title="Estado atual de conhecimentos dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil." volume="1" volumeTitle="Hacia un proyecto CYTED para el inventario y estimacion de la diversidad entomologica en Iberoamerica: PRIBES- 2000. Monografias Tercer Milenio m 3 m." year="2000">Vaz-de-Mello 2000</bibRefCitation>
: 195 (cited for Brazil);
<bibRefCitation id="FBDEE1BFEA8B62AB10E5298EFFB9B100" author="Vitolo, AL" journalOrPublisher="Revista de la Academia Colombiana de Ciencias Exactas Fisicas y Naturales" pageId="278" pageNumber="279" pagination="591 - 601" title="Clave para la identificacion de los generos y especies Phanaeinas de Colombia." volume="24" year="2000">Vitolo 2000</bibRefCitation>
: 595 (characters in key);
<bibRefCitation id="9647425EAD323DBC09FA26257B872E90" author="Medina, CA" journalOrPublisher="Biota Colombiana" pageId="274" pageNumber="275" pagination="131 - 144" title="Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia." volume="2" year="2001">Medina et al. 2001</bibRefCitation>
: 140 (cited for Colombia);
<bibRefCitation id="64361B0B2709E042ED8484D6DF465B49" author="Arnaud, P" journalOrPublisher="Hillside Books, Canterbury, England" pageId="261" pageNumber="262" title="Les Coleopteres du Monde, Vol. 28. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus." year="2002 a">Arnaud 2002a</bibRefCitation>
: 131 (diagnosis);
<bibRefCitation id="154BE3B58DAE2171FB7A458825928243" author="Vitolo, A" editor="Fernandez, F" journalOrPublisher="Facultad de Ciencias, Universidad Nacional de Colombia, Bogota" pageId="278" pageNumber="279" pagination="277 - 318" title="Escarabajos estercoleros de la tribu Phanaeini de Colombia (Coleoptera: Scarabaeoidea: Scarabaeidae)." volume="3" volumeTitle="Insectos de Colombia." year="2004">Vitolo 2004</bibRefCitation>
: 280 (diagnosis);
<bibRefCitation id="59202A184D04D3535513BDE5633CC0A7" author="Hamel-Leigue, A" journalOrPublisher="Revista Boliviana de Ecologia y Conservacion Ambiental" pageId="268" pageNumber="269" pagination="1 - 18" title="Hacia un inventario de los escarabajos peloteros (Coleoptera: Scarabaeinae) de Bolivia: primera compilacion de los generos y especies registrados para el pais." volume="20" year="2006">Hamel-Leigue et al. 2006</bibRefCitation>
: 18 (cited for Bolivia);
<bibRefCitation id="9B075A6C069968D3BC956E22C1591F81" author="Hamel-Leigue, A" journalOrPublisher="Kempffiana" pageId="268" pageNumber="269" pagination="43 - 95" title="Distribucion e historia natural de escarabajos coprofagos de la tribu Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae) en Bolivia." volume="5" year="2009">Hamel-Leigue et al. 2009</bibRefCitation>
: 67 (comment);
<bibRefCitation id="7D2F497E719D80872AFBD1BA5662F4BB" author="Carvajal, V" journalOrPublisher="Quito, Ecuado" pageId="264" pageNumber="265" title="Escarabajos del Ecuador. Principales generos. Instituto de Ciencias Biologicas. Escuela Politecnica Nacional. Serie Entomologia, Nro. 1." year="2011">Carvajal et al. 2011</bibRefCitation>
: 320-321 (cited for Ecuador);
<bibRefCitation id="900AC549686EE926586E1115CB6DC04F" pageId="229" pageNumber="230">Krajcik 2012</bibRefCitation>
: 204 (cited as species of the genus
<taxonomicName id="0318B0FCB0B7F4B39492B932FA75FAAE" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus" order="Coleoptera" pageId="229" pageNumber="230" phylum="Arthropoda" rank="genus">Phanaeus</taxonomicName>
Macleay, 1819);
<bibRefCitation id="00CDAF2B57AECF1ED57D4DF21022CEB5" author="Boilly, O" journalOrPublisher="ACOREP-France" pageId="263" pageNumber="264" pagination="103 - 112" title="Les Scarabaeinae de Guyane: cle illustree des genres." volume="7" year="2013">Boilly and Vaz-de-Mello 2013</bibRefCitation>
: 105 (figure 21);
<bibRefCitation id="6313B29DDB6D70ED6B3DB76F12E78A04" author="Figueroa, L" journalOrPublisher="Revista Peruana de Biologia" pageId="267" pageNumber="268" pagination="125 - 138" title="La tribu Phanaeini (Coleoptera, Scarabaeidae, Scarabaeinae) en el Peru." url="https://doi.org/10.15381/rpb.v21i2.9815" volume="21" year="2014">Figueroa et al. 2014</bibRefCitation>
: 132 (distributional records from Peru);
<bibRefCitation id="5799E01FF513F8CF6105EC506404FCA2" author="Ratcliffe, BC" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="276" pageNumber="277" pagination="186 - 207" title="Beetles (Coleoptera) of Peru: A Survey of the Families." url="https://doi.org/10.2317/kent-88-02-186-207.1" volume="88" year="2015">Ratcliffe et al. 2015</bibRefCitation>
: 195 (cited for Peru);
<bibRefCitation id="C8F3182A4CE8A980805CA13DF3FD096D" author="Boilly, O" journalOrPublisher="ACOREP-France" pageId="263" pageNumber="264" pagination="86 - 97" title="Les Phanaeini de Guyane: liste commentee, cles et iconographie (Coleoptera, Scarabaeidae)." volume="10" year="2016">Boilly et al. 2016</bibRefCitation>
: 89 (figures 7a, 7b and 7c); 95 (cited for Guyana);
<bibRefCitation id="B1350119CD65C8D5A072523E60984365" author="Chamorro, W" journalOrPublisher="Revista Colombiana de Entomologi ́ a" pageId="264" pageNumber="265" pagination="72 - 100" title="Listado de especies y clave de ge ́ neros y subge ́ neros de escarabajos estercoleros (Coleoptera: Scarabaei- dae: Scarabaeinae) presentes y presuntos para Ecuador." url="https://doi.org/10.25100/socolen.v44i1.6545" volume="44" year="2018">Chamorro et al. 2018</bibRefCitation>
: 98 (cited for Ecuador).
</paragraph>
</subSubSection>
<subSubSection id="48007F5EF16165D7D93F30054EB5207C" pageId="229" pageNumber="230" type="type specimens">
<paragraph id="11E6E12934CC1590EE683B33253AC59F" pageId="229" pageNumber="230">Type specimens.</paragraph>
<paragraph id="CE2082847509BB898F06F28A960B20EC" pageId="229" pageNumber="230">
<taxonomicName id="7F99EEE04DD2B2FF473B3917A2204726" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scarabaeus faunus" order="Coleoptera" pageId="229" pageNumber="230" phylum="Arthropoda" rank="species" species="faunus">Scarabaeus faunus</taxonomicName>
Fabricius, 1775. Type material not examined.
</paragraph>
</subSubSection>
<subSubSection id="F7229CFCFB063F4802659B14757A462B" pageId="229" pageNumber="230" type="distribution">
<paragraph id="1F7F07EE622814BB4870ABF464A749C5" pageId="229" pageNumber="230">Distribution.</paragraph>
<paragraph id="7E8F98883DC270D04368161D71FCFFE1" pageId="229" pageNumber="230">Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Paraguay, and Peru.</paragraph>
</subSubSection>
<subSubSection id="6BAB9ED0E501DBAB2B2B6B24E0B95877" pageId="230" pageNumber="231" type="records examined">
<paragraph id="425B61CF5BF16D4026DEAEBE19E035DA" pageId="230" pageNumber="231">
<pageBreakToken id="D9FEE0C26900F02BE1273085DCE557BD" pageId="230" pageNumber="231" start="start">Records</pageBreakToken>
examined.
</paragraph>
<paragraph id="7DCA7EBB5C24A237D32EAAB64072667F" pageId="230" pageNumber="231">UNDETERMINED PROVINCE: without specific locality (1 specimen MECN.</paragraph>
</subSubSection>
<subSubSection id="22201DBC1480679CA382CAE3B0439398" pageId="230" pageNumber="231" type="temporal data">
<paragraph id="D4326E3D11AF85EC35C3899855148449" pageId="230" pageNumber="231">Temporal data.</paragraph>
<paragraph id="A0FD782F9D9A3BBA482BF5F593A07B79" pageId="230" pageNumber="231">It is not known when this species was collected.</paragraph>
</subSubSection>
<subSubSection id="7099A2705BE2C9F1BBE35A3B8FBAF7C6" pageId="230" pageNumber="231" type="remarks">
<paragraph id="92DBBDAFCBE1C3E1E537DE4AE0763ACF" pageId="230" pageNumber="231">Remarks.</paragraph>
<paragraph id="FA4DD7C98A3FBAFF808D0D1DE2656EA9" pageId="230" pageNumber="231">Ecological needs unknown, it may be found in Amazonian lowland forests. The collection method is unknown.</paragraph>
</subSubSection>
</treatment>
</document>