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<document id="DE5B1C4831465F07DAB7445889C361ED" ID-DOI="10.11646/zootaxa.1161.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5058330" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625190197324" checkinUser="felipe" docAuthor="Hippa, Heikki" docDate="2006" docId="5D4C8785284EFFA55C1A0F10FD24FA36" docLanguage="en" docName="zt01161p049.pdf" docOrigin="Zootaxa 1161 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1161.1.1" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Manota spadix Hippa 2006, sp. n." docType="treatment" docVersion="4" lastPageNumber="11" masterDocId="A175FFFD2844FFAF5D120E76FFC5FFE7" masterDocTitle="Diversity of Manota Williston (Diptera: Mycetophilidae) in a Malaysian rainforest: description of twenty-seven new sympatric species" masterLastPageNumber="49" masterPageNumber="1" pageNumber="11" updateTime="1698983105622" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="F7D59D0EABE5A42C60FCD4CEDED88DA0">Diversity of Manota Williston (Diptera: Mycetophilidae) in a Malaysian rainforest: description of twenty-seven new sympatric species</mods:title>
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<mods:date id="8920BAD8197F60727E63079C294051F6">2006</mods:date>
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<mods:number id="EAF41B20EB5E06C2FF561E386B640186">2006-03-30</mods:number>
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<mods:number id="CDF44ABA0A943CFFA5526C672FC72526">1161</mods:number>
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<mods:classification id="C74D55ADF3384B174517AA41FDC14979">journal article</mods:classification>
<mods:identifier id="322044388D3547AE0D2EB02B248E2750" type="DOI">10.11646/zootaxa.1161.1.1</mods:identifier>
<mods:identifier id="1133758F6C5BDD17C1322D0F63C953A4" type="ISSN">1175­5334</mods:identifier>
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<treatment id="5D4C8785284EFFA55C1A0F10FD24FA36" ID-DOI="http://doi.org/10.5281/zenodo.5059282" ID-Zenodo-Dep="5059282" LSID="urn:lsid:plazi:treatment:5D4C8785284EFFA55C1A0F10FD24FA36" httpUri="http://treatment.plazi.org/id/5D4C8785284EFFA55C1A0F10FD24FA36" lastPageNumber="11" pageId="10" pageNumber="11">
<subSubSection id="9DFF6518284EFFA55C1A0F10FDC0FE67" box="[264,517,358,384]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="D55A3693284EFFA55C1A0F10FDC0FE67" blockId="10.[264,517,358,419]" box="[264,517,358,384]" pageId="10" pageNumber="11">
<heading id="8E1281FF284EFFA55C1A0F10FDC0FE67" bold="true" box="[264,517,358,384]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="E791EA81284EFFA55C1A0F10FDC0FE67" bold="true" box="[264,517,358,384]" pageId="10" pageNumber="11">
<taxonomicName id="12E54D10284EFFA55C1A0F10FE7EFE67" ID-CoL="3XVYM" authority="Hippa, 2006" authorityName="Hippa" authorityYear="2006" box="[264,443,358,384]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spadix" status="sp. nov.">
<emphasis id="E791EA81284EFFA55C1A0F10FE7EFE67" bold="true" box="[264,443,358,384]" italics="true" pageId="10" pageNumber="11">Manota spadix</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FCA257FA284EFFA55CD10F10FDC0FE67" box="[451,517,358,384]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="9DFF6518284EFFA55C1A0FFFFD24FA36" pageId="10" pageNumber="11" type="description">
<paragraph id="D55A3693284EFFA55C1A0FFFFE13FE44" blockId="10.[264,517,358,419]" box="[264,470,393,419]" pageId="10" pageNumber="11">
(
<figureCitation id="4DDE2A16284EFFA55C030FFFFEBCFE44" box="[273,377,393,419]" captionStart="FIGURE 1" captionStartId="4.[264,368,1691,1715]" captionTargetBox="[300,1268,323,1675]" captionTargetId="figure-116@4.[285,1310,298,1688]" captionTargetPageId="4" captionText="FIGURE 1. Left palp, dorsal view (A); apical part of palpomere 3, lateral view (B, C, D); and apical part of tibia 1 with basal part of tarsomere 1, prolateral view (E). A, C. Manota heptacantha sp. n (paratypes). B. Manota pappi sp. n. (paratype). D. Manota spadix sp. n. (paratype). E. Manota angustata sp. n. (paratype). Scale 0.1 mm. 1 = palpomere 1, 2 = palpomere 2, 3 = palpomere 3, 4 = palpomere 4, 5 = palpomere 5, 6 = sensory pit, 7 = curved sensillum, 8 = bluntended sensillum, 9 = seta, 10 = parasegment." figureDoi="http://doi.org/10.5281/zenodo.5058332" httpUri="https://zenodo.org/record/5058332/files/figure.png" pageId="10" pageNumber="11">Figs. 1D</figureCitation>
;
<figureCitation id="4DDE2A16284EFFA55C940FFFFE09FE44" box="[390,460,393,419]" captionStart="FIGURE 3" captionStartId="11.[264,367,1691,1715]" captionTargetBox="[295,1277,328,1647]" captionTargetId="figure-74@11.[281,1306,301,1664]" captionTargetPageId="11" captionText="FIGURE 3. Antennal flagellomere 4, lateral view (A, C); hypopygium ventral (B) and dorsal (C) view; and cerci, dorsal view (E). A, B. Manota spadix sp. n. (holotype). C, D, E. Manota roslii sp. n. (holotype). Scale for A and C 0.05 mm; for B, D, and E 0.10 mm. For abbreviations, see under Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5058338" httpUri="https://zenodo.org/record/5058338/files/figure.png" pageId="10" pageNumber="11">3A, B</figureCitation>
)
</paragraph>
<paragraph id="D55A3693284EFFA55C1A0FAFFDD6FE14" blockId="10.[264,1323,473,1099]" box="[264,531,473,499]" pageId="10" pageNumber="11">
A small­sized
<taxonomicName id="12E54D10284EFFA55CA30FAFFDC9FE15" authorityName="Williston" authorityYear="1896" box="[433,524,473,498]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="E791EA81284EFFA55CA30FAFFDC9FE15" box="[433,524,473,498]" italics="true" pageId="10" pageNumber="11">Manota</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="D55A3693284EFFA55C2A0C77FC48FBC4" blockId="10.[264,1323,473,1099]" pageId="10" pageNumber="11">
Male.
<emphasis id="E791EA81284EFFA55C970C77FE02FDFC" bold="true" box="[389,455,513,539]" pageId="10" pageNumber="11">Head</emphasis>
. Flagellomere 4,
<figureCitation id="4DDE2A16284EFFA55F8F0C77FD38FDFC" box="[669,765,513,539]" captionStart="FIGURE 3" captionStartId="11.[264,367,1691,1715]" captionTargetBox="[295,1277,328,1647]" captionTargetId="figure-74@11.[281,1306,301,1664]" captionTargetPageId="11" captionText="FIGURE 3. Antennal flagellomere 4, lateral view (A, C); hypopygium ventral (B) and dorsal (C) view; and cerci, dorsal view (E). A, B. Manota spadix sp. n. (holotype). C, D, E. Manota roslii sp. n. (holotype). Scale for A and C 0.05 mm; for B, D, and E 0.10 mm. For abbreviations, see under Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5058338" httpUri="https://zenodo.org/record/5058338/files/figure.png" pageId="10" pageNumber="11">Fig. 3A</figureCitation>
. Maxillary palpus with palpomere 3 bearing 4 apically expanded curved sensilla; palpomere 4 with parasegment.
<emphasis id="E791EA81284EFFA5590B0C5FFBB3FDA4" bold="true" box="[1049,1142,553,579]" pageId="10" pageNumber="11">Thorax</emphasis>
. Anepisternum setose. Anterior basalare non­setose. Preepisternum 2 setose. Laterotergite setose. Episternum 3 with few setae.
<emphasis id="E791EA81284EFFA55F6A0C0FFD79FD74" bold="true" box="[632,700,633,659]" pageId="10" pageNumber="11">Wing</emphasis>
. Length
<quantity id="121D9B76284EFFA55E3E0C0FFC05FD74" box="[812,960,633,659]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="10" pageNumber="11" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.5–1.7 mm</quantity>
. Membrane with few setae at posterior margin. Vein 1A indicated by few setae. Sc distally of h non­setose.
<emphasis id="E791EA81284EFFA55C1A0CBFFE77FD04" bold="true" box="[264,434,713,739]" pageId="10" pageNumber="11">Hypopygium.</emphasis>
(
<figureCitation id="4DDE2A16284EFFA55CD00CBFFDD8FD04" box="[450,541,713,739]" captionStart="FIGURE 3" captionStartId="11.[264,367,1691,1715]" captionTargetBox="[295,1277,328,1647]" captionTargetId="figure-74@11.[281,1306,301,1664]" captionTargetPageId="11" captionText="FIGURE 3. Antennal flagellomere 4, lateral view (A, C); hypopygium ventral (B) and dorsal (C) view; and cerci, dorsal view (E). A, B. Manota spadix sp. n. (holotype). C, D, E. Manota roslii sp. n. (holotype). Scale for A and C 0.05 mm; for B, D, and E 0.10 mm. For abbreviations, see under Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5058338" httpUri="https://zenodo.org/record/5058338/files/figure.png" pageId="10" pageNumber="11">Fig. 3B</figureCitation>
). Sternite 9 large, posteriorly extending nearly as far as ventral part of gonocoxa, laterally sharply separated from gonocoxa, apically deeply notched so that sternite is almost divided into 2 separate halves, setae similar to ventral setae of gonocoxa. Gonocoxa in position III with 2 stout curved and apically flattened megasetae, dorsally from these there is a finger­like lobe (not visible in
<figureCitation id="4DDE2A16284EFFA55E800D1FFC2AFC64" box="[914,1007,873,899]" captionStart="FIGURE 3" captionStartId="11.[264,367,1691,1715]" captionTargetBox="[295,1277,328,1647]" captionTargetId="figure-74@11.[281,1306,301,1664]" captionTargetPageId="11" captionText="FIGURE 3. Antennal flagellomere 4, lateral view (A, C); hypopygium ventral (B) and dorsal (C) view; and cerci, dorsal view (E). A, B. Manota spadix sp. n. (holotype). C, D, E. Manota roslii sp. n. (holotype). Scale for A and C 0.05 mm; for B, D, and E 0.10 mm. For abbreviations, see under Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5058338" httpUri="https://zenodo.org/record/5058338/files/figure.png" pageId="10" pageNumber="11">Fig. 3B</figureCitation>
) with long setae at apex. Gonostylus short, strongly widening toward apex, the apex with a concave excision. Parastylar lobe absent. Tegmen unusual, with short broad basal part which beyond the very strong lateral shoulders forms a long and narrow, apically expanded apical part, and which posteriorly extends beyond other parts of hypopygium.
</paragraph>
<paragraph id="D55A3693284EFFA55C2A0A47FCDAFBAC" blockId="10.[264,1323,473,1099]" box="[312,799,1073,1099]" pageId="10" pageNumber="11">Female and preimaginal stages unknown.</paragraph>
<paragraph id="D55A3693284EFFA55C1A0A0AFE4CFB72" blockId="10.[264,1322,1148,1254]" box="[264,393,1148,1173]" pageId="10" pageNumber="11">
<emphasis id="E791EA81284EFFA55C1A0A0AFE4CFB72" box="[264,393,1148,1173]" italics="true" pageId="10" pageNumber="11">Discussion</emphasis>
</paragraph>
<paragraph id="D55A3693284EFFA55C2A0AD2FC3AFB01" blockId="10.[264,1322,1148,1254]" pageId="10" pageNumber="11">
<taxonomicName id="12E54D10284EFFA55C2A0AD2FE29FB5A" authorityName="Hippa" authorityYear="2006" box="[312,492,1188,1213]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spadix">
<emphasis id="E791EA81284EFFA55C2A0AD2FE29FB5A" box="[312,492,1188,1213]" italics="true" pageId="10" pageNumber="11">Manota spadix</emphasis>
</taxonomicName>
is highly dissimilar to all other known
<taxonomicName id="12E54D10284EFFA55EFA0AD2FB86FB5A" authorityName="Williston" authorityYear="1896" box="[1000,1091,1188,1213]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="E791EA81284EFFA55EFA0AD2FB86FB5A" box="[1000,1091,1188,1213]" italics="true" pageId="10" pageNumber="11">Manota</emphasis>
</taxonomicName>
. It is immediately distinguished by the exceptional length and shape of the tegmen.
</paragraph>
<paragraph id="D55A3693284EFFA55C1A0B61FE8FFAD7" blockId="10.[264,1323,1303,1489]" box="[264,330,1303,1328]" pageId="10" pageNumber="11">
<emphasis id="E791EA81284EFFA55C1A0B61FE8FFAD7" box="[264,330,1303,1328]" italics="true" pageId="10" pageNumber="11">Types</emphasis>
</paragraph>
<paragraph id="D55A3693284EFFA55C2A0B49FD24FA36" blockId="10.[264,1323,1303,1489]" pageId="10" pageNumber="11">
<materialsCitation id="658D3CCE284EFFA55C2A0B49FE04FA4E" collectingDate="1997-02-22" collectingDateMax="1997-03-21" collectingDateMin="1997-02-22" collectorName="H. Hippa &amp; M. Jaschhof &amp; B. Viklund" country="Malaysia" county="Male" elevation="244" location="Ulu Gombak" municipality="West" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Selangor" typeStatus="holotype">
<emphasis id="E791EA81284EFFA55C2A0B49FE66FABF" box="[312,419,1343,1368]" italics="true" pageId="10" pageNumber="11">
<typeStatus id="0A5E8831284EFFA55C2A0B49FE66FABF" box="[312,419,1343,1368]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectingCounty id="3C3B4E1F284EFFA55CA60B49FE36FABE" box="[436,499,1343,1369]" pageId="10" pageNumber="11">Male</collectingCounty>
,
<collectingMunicipality id="353EACE9284EFFA55F100B49FDF9FABE" box="[514,572,1343,1369]" pageId="10" pageNumber="11">West</collectingMunicipality>
<collectingCountry id="ADF27603284EFFA55F570B49FD76FABE" box="[581,691,1343,1369]" name="Malaysia" pageId="10" pageNumber="11">Malaysia</collectingCountry>
,
<collectingRegion id="1721F871284EFFA55FD30B49FCEEFABE" box="[705,811,1343,1369]" country="Malaysia" name="Selangor" pageId="10" pageNumber="11">Selangor</collectingRegion>
,
<location id="D03A6048284EFFA55E280B49FC12FABE" LSID="urn:lsid:plazi:treatment:5D4C8785284EFFA55C1A0F10FD24FA36:D03A6048284EFFA55E280B49FC12FABE" box="[826,983,1343,1369]" country="Malaysia" county="Male" municipality="West" name="Ulu Gombak" pageId="10" pageNumber="11" stateProvince="Selangor">Ulu Gombak</location>
,
<location id="D03A6048284EFFA55EF70B49FE60FA66" LSID="urn:lsid:plazi:treatment:5D4C8785284EFFA55C1A0F10FD24FA36:D03A6048284EFFA55EF70B49FE60FA66" country="Malaysia" county="Male" municipality="West" name="University of Malaya Field Study Centre" pageId="10" pageNumber="11" stateProvince="Selangor">University of Malaya Field Study Centre</location>
,
<quantity id="121D9B76284EFFA55CA00B11FE3EFA66" box="[434,507,1383,1409]" metricMagnitude="2" metricUnit="m" metricValue="2.4384" pageId="10" pageNumber="11" unit="ft" value="800.0">
<elevation id="5EC8D1A0284EFFA55CA00B11FE3EFA66" box="[434,507,1383,1409]" metricMagnitude="2" metricUnit="m" metricValue="2.4384" pageId="10" pageNumber="11" unit="ft" value="800.0">800 ft</elevation>
</quantity>
, Malaise in jungle,
<date id="A15B1053284EFFA55FF60B11FBD4FA66" box="[740,1041,1383,1409]" pageId="10" pageNumber="11" value="1997-02-22" valueMax="1997-03-21" valueMin="1997-02-22">
<collectingDate id="B11FE9BB284EFFA55FF60B11FBD4FA66" box="[740,1041,1383,1409]" pageId="10" pageNumber="11" value="1997-02-22" valueMax="1997-03-21" valueMin="1997-02-22">22. Feb.–21. March 1997</collectingDate>
</date>
,
<collectorName id="78105345284EFFA5590D0B11FB4BFA66" box="[1055,1166,1383,1409]" pageId="10" pageNumber="11">H. Hippa</collectorName>
,
<collectorName id="78105345284EFFA559880B11FAEFFA66" box="[1178,1322,1383,1409]" pageId="10" pageNumber="11">M. Jaschhof</collectorName>
and
<collectorName id="78105345284EFFA55C290BF9FE04FA4E" box="[315,449,1423,1449]" pageId="10" pageNumber="11">B. Viklund</collectorName>
</materialsCitation>
.
<materialsCitation id="658D3CCE284EFFA55CDF0BF9FC35FA4E" box="[461,1008,1423,1449]" collectingDate="1997-02-22" collectingDateMax="1997-03-21" collectingDateMin="1997-02-22" collectorName="H. Hippa &amp; M. Jaschhof &amp; B. Viklund" country="Malaysia" county="Male" elevation="244" location="Ulu Gombak" municipality="West" pageId="10" pageNumber="11" specimenCount="1" specimenCount-male="1" stateProvince="Selangor" typeStatus="paratype">
<typeStatus id="0A5E8831284EFFA55CDF0BF9FD86FA4E" box="[461,579,1423,1449]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="C3E3FD1A284EFFA55F5D0BF9FD65FA4E" box="[591,672,1423,1449]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
with same data as holotype
</materialsCitation>
;
<materialsCitation id="658D3CCE284EFFA55EEF0BF9FEABFA36" collectingDate="1997-02-22" collectingDateMax="1997-03-21" collectingDateMin="1997-02-22" collectorName="H. Hippa &amp; M. Jaschhof &amp; B. Viklund" country="Malaysia" county="Male" elevation="244" location="Ulu Gombak" municipality="West" pageId="10" pageNumber="11" specimenCount="1" specimenCount-male="1" stateProvince="Selangor" typeStatus="paratype">
<specimenCount id="C3E3FD1A284EFFA55EEF0BF9FB88FA4E" box="[1021,1101,1423,1449]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
with same data as holotype
</materialsCitation>
except for 24. Feb.–21. March.
</paragraph>
</subSubSection>
</treatment>
</document>
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