treatments-xml/data/5B/1D/26/5B1D2668FFA7950EC3F9F9C2FCBFFC7B.xml

<document id="66A6EE0D5641FF0FFA144C7DDF9EB859" ID-CLB-Dataset="29237" ID-DOI="10.11646/zootaxa.4496.1.8" ID-GBIF-Dataset="6a8d52af-2897-436e-b03b-a66a0809b89a" ID-ISSN="1175-5326" ID-Zenodo-Dep="1447053" ID-ZooBank="AEBC3EB5-DD2A-49BB-898F-EA792AEBBC46" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538748490746" checkinUser="plazi" docAuthor="Nagy, Hajnalka, Felföldi, Tamás &amp; Dózsa-Farkas, Klára" docDate="2018" docId="5B1D2668FFA7950EC3F9F9C2FCBFFC7B" docLanguage="en" docName="zootaxa.4496.1.8.pdf" docOrigin="Zootaxa 4496 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fridericia ventrochaetosa Nagy &amp; Felföldi &amp; Dózsa-Farkas 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="117" masterDocId="A7245E10FFA69509C36EFFF6FF87FFBC" masterDocTitle="Morphological and molecular distinction of two Fridericia species (Clitellata, Enchytraeidae) having same spermatheca type" masterLastPageNumber="123" masterPageNumber="111" pageNumber="112" updateTime="1698567014823" updateUser="ExternalLinkService">
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<mods:title id="89F80F3895FCB320F8FB30066F3B41BB">Morphological and molecular distinction of two Fridericia species (Clitellata, Enchytraeidae) having same spermatheca type</mods:title>
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<mods:namePart id="57635F6934AA8FA2D1B52F78E9948D44">Nagy, Hajnalka</mods:namePart>
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<mods:namePart id="BBC903100EA0A72C9AE5195B26870DD7">Felföldi, Tamás</mods:namePart>
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<mods:namePart id="EBBA8A082C4EA72B13BDC18537F733E7">Dózsa-Farkas, Klára</mods:namePart>
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<treatment id="5B1D2668FFA7950EC3F9F9C2FCBFFC7B" ID-DOI="http://doi.org/10.5281/zenodo.5951820" ID-GBIF-Taxon="148640632" ID-Zenodo-Dep="5951820" LSID="urn:lsid:plazi:treatment:5B1D2668FFA7950EC3F9F9C2FCBFFC7B" httpUri="http://treatment.plazi.org/id/5B1D2668FFA7950EC3F9F9C2FCBFFC7B" lastPageId="7" lastPageNumber="117" pageId="1" pageNumber="112">
<subSubSection id="9BAEC4F5FFA79508C3F9F9C2FEA9F9D3" pageId="1" pageNumber="112" type="nomenclature">
<paragraph id="D30B977EFFA79508C3F9F9C2FD9FF9F2" blockId="1.[151,536,1588,1647]" box="[151,536,1588,1614]" pageId="1" pageNumber="112">
<heading id="88432012FFA79508C3F9F9C2FD9FF9F2" bold="true" box="[151,536,1588,1614]" fontSize="11" level="1" pageId="1" pageNumber="112" reason="1">
<emphasis id="E1C04B6CFFA79508C3F9F9C2FD9FF9F2" bold="true" box="[151,536,1588,1614]" pageId="1" pageNumber="112">
<taxonomicName id="14B4ECFDFFA79508C3F9F9C2FE49F9F2" authority="Nagy &amp; Felföldi &amp; Dózsa-Farkas, 2018" authorityName="Nagy &amp; Felföldi &amp; Dózsa-Farkas" authorityYear="2018" box="[151,462,1588,1614]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="112" phylum="Magnoliopsida" rank="species" species="ventrochaetosa" status="sp. nov.">
<emphasis id="E1C04B6CFFA79508C3F9F9C2FE49F9F2" bold="true" box="[151,462,1588,1614]" italics="true" pageId="1" pageNumber="112">Fridericia ventrochaetosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAF3F617FFA79508C2B8F9C2FD9FF9F2" box="[470,536,1588,1614]" pageId="1" pageNumber="112" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D30B977EFFA79508C3F9F9A0FEA9F9D3" blockId="1.[151,536,1588,1647]" box="[151,302,1622,1647]" pageId="1" pageNumber="112">
(
<figureCitation id="4B8F8BFBFFA79508C3F0F9A0FEA2F9D3" box="[158,293,1622,1647]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1802,1824]" captionStartId-1="4.[151,250,1802,1824]" captionStartId-2="5.[151,250,1618,1640]" captionTargetBox-0="[158,1372,193,1745]" captionTargetBox-1="[158,1376,193,1743]" captionTargetBox-2="[178,1398,288,1505]" captionTargetId-0="figure@3.[158,1428,193,1780]" captionTargetId-1="figure@4.[158,1428,193,1780]" captionTargetId-2="figure@5.[178,1408,288,1524]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Micrographs of Fridericia ventrochaetosa sp. n. A. Brain (DNA 1109). B. Brain (holotype). C. Epidermal glands. D. Thick body wall and cuticle (marked with arrow). E. Strong septa, 5/6– 9/10 (holotype). F–G. Head region (the thick epithelium of prostomium marked with arrow). H. Clitellar glands, lateral view. I. Clitellum of a dissected species (above the dorsal gap without glands, below the clitellar glands in transverse rows). J–K. Coelomocytes (J, paratype P.112.1.2, No. 2219=DNA 978). L. Nephridium of a juvenile specimen, in the middle part of body; M. Preclitellar nephridium (paratype P.112.1.2, No. 2219=DNA 978). A, C, H, J, L–M in vivo, B, E fixed, not stained, D, F–G, I, K. fixed, stained. Scale bars 50 µm." captionText-1="FIGURE 2. Micrographs of Fridericia ventrochaetosa sp. n. A. End of oesophageal appendage with multiple branches (arrows). B. Male copulatory organs (arrows); clitellar gland cells visible laterally and ventrally posterior to copulatory organs. C. Midgut pars tumida (arrows) in XXXI–XXXIII (holotype). D. Chylus cells in XIII–XIV. E. Male copulatory organs, ventral view, holotype. Strong bursal muscle marked with arrow. Asterisk: region between the two organs without clitellar glands. F– G. Bursal slits (marked with arrow in G). H. Male copulatory organ, lateral view (paratype P.112.1.2, No. 2219=DNA 978). I,J. Sperm funnels (paratype P.112.1.2, No. 2219=DNA 978). A, F, G, I in vivo, B, D, H, J fixed, stained, C, E fixed, not stained. Scale bars 50 µm." captionText-2="FIGURE 3. Micrograph of Fridericia ventrochaetosa sp. n. A. Sperm funnels (paratype P.112.1.2, No. 2219=DNA 978). B, D. Spermathecae (paratype P.112.2.6, No. 2345=DNA 1116). C. Spermathecae (holotype). E. Spermatheca (paratype P.112.1.2, No. 2219=DNA 978). F. Dissected spermatheca (paratype P.112.2.1, No. 2266). G. Epidermal gland cells around the opening of spermatheca (opening marked with arrow). A, B, D, G in vivo, E, F fixed, stained, C fixed, not stained. Scale bars in A 20 µm, in the rest 50 µm." httpUri-0="https://zenodo.org/record/1447055/files/figure.png" httpUri-1="https://zenodo.org/record/1447057/files/figure.png" httpUri-2="https://zenodo.org/record/1447059/files/figure.png" pageId="1" pageNumber="112">Figures 1–3</figureCitation>
)
</paragraph>
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<paragraph id="D30B977EFFA79508C3F9F968FCA5F908" blockId="1.[151,802,1694,1716]" box="[151,802,1694,1716]" pageId="1" pageNumber="112">
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<taxonomicName id="14B4ECFDFFA79508C3F9F968FF78F908" box="[151,255,1694,1716]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="112" phylum="Magnoliopsida" rank="genus">
<emphasis id="E1C04B6CFFA79508C3F9F968FF78F908" box="[151,255,1694,1716]" italics="true" pageId="1" pageNumber="112">Fridericia</emphasis>
</taxonomicName>
sp. 
<treatmentCitation id="5215B16FFFA79508C246F968FD04F908" author="" box="[296,643,1694,1716]" page="52" pageId="1" pageNumber="112">
<bibRefCitation id="B725EA8FFFA79508C246F968FDE7F908" box="[296,608,1694,1716]" pageId="1" pageNumber="112" refId="ref5046" refString="Dozsa-Farkas, K. &amp; Felfoldi, T. (2017) On the enchytraeid fauna of Koszeg Mountains with description of a new Fridericia species (Clitellata, Enchytraeidae). Opuscula Zoologica, Budapest, 48 (1), 41 - 54. https: // doi. org / 10.18348 / opzool. 2017.1.41." type="journal article">Dózsa-Farkas &amp; Felföldi 2017</bibRefCitation>
: 52
</treatmentCitation>
<emphasis id="E1C04B6CFFA79508C1EAF969FD16F908" bold="true" box="[644,657,1695,1716]" pageId="1" pageNumber="112">–</emphasis>
53, 2018: 4 
<emphasis id="E1C04B6CFFA79508C06DF969FC88F908" bold="true" box="[771,783,1695,1716]" pageId="1" pageNumber="112">–</emphasis>
5.
</treatmentCitationGroup>
</paragraph>
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<paragraph id="D30B977EFFA79508C3F9F917FC75F946" blockId="1.[151,1437,1761,2009]" box="[151,1010,1761,1786]" pageId="1" pageNumber="112">
<emphasis id="E1C04B6CFFA79508C3F9F917FE83F946" bold="true" box="[151,260,1761,1786]" pageId="1" pageNumber="112">Holotype</emphasis>
. 
<emphasis id="E1C04B6CFFA79508C27CF917FECFF946" bold="true" box="[274,328,1761,1786]" pageId="1" pageNumber="112">F. 29</emphasis>
, slide 
<emphasis id="E1C04B6CFFA79508C2FAF917FE7BF946" bold="true" box="[404,508,1761,1786]" pageId="1" pageNumber="112">No. 2267</emphasis>
, adult, unstained, whole-mounted specimen.
</paragraph>
<paragraph id="D30B977EFFA79508C3A9F8FEFBB4F8FB" blockId="1.[151,1437,1761,2009]" pageId="1" pageNumber="112">
<materialsCitation id="63DC9D23FFA79508C3A9F8FEFBB4F8FB" ID-GBIF-Occurrence="1914266229" collectingDate="2016-10-24" collectorName="K. Dozsa-Farkas &amp; Farkas &amp; Pobozsny" country="Hungary" elevation="667" latitude="47.370018" location="Koszeg Mts." longLatPrecision="1" longitude="16.467417" municipality="Steirer Houses" pageId="1" pageNumber="112" specimenCount="1" typeStatus="holotype">
<emphasis id="E1C04B6CFFA79508C3A9F8FEFEE1F89D" bold="true" box="[199,358,1800,1825]" pageId="1" pageNumber="112">
<typeStatus id="0C0F29DCFFA79508C3A9F8FEFE85F89D" box="[199,258,1800,1825]" pageId="1" pageNumber="112">Type</typeStatus>
locality
</emphasis>
. 
<location id="D66BC1A5FFA79508C213F8FEFD93F89C" LSID="urn:lsid:plazi:treatment:5B1D2668FFA7950EC3F9F9C2FCBFFC7B:D66BC1A5FFA79508C213F8FEFD93F89C" box="[381,532,1800,1825]" country="Hungary" latitude="47.370018" longLatPrecision="1" longitude="16.467417" municipality="Steirer Houses" name="Koszeg Mts." pageId="1" pageNumber="112">Kȏszeg Mts.</location>
(
<collectingCountry id="ABA3D7EEFFA79508C145F8FEFD14F89C" box="[555,659,1800,1824]" name="Hungary" pageId="1" pageNumber="112">Hungary</collectingCountry>
), 
<emphasis id="E1C04B6CFFA79508C1C1F8FEFC63F89C" box="[687,996,1800,1824]" italics="true" pageId="1" pageNumber="112">
<collectingMunicipality id="336F0D04FFA79508C1C1F8FEFCD9F89C" box="[687,862,1800,1824]" pageId="1" pageNumber="112">Steirer Houses</collectingMunicipality>
, mesophile
</emphasis>
montane hay meadow, 
<geoCoordinate id="B680F1B9FFA79508C67EF8FEFA1FF89C" box="[1296,1432,1797,1824]" direction="north" orientation="latitude" pageId="1" pageNumber="112" precision="1" value="47.370018">47o22.201N</geoCoordinate>
, 
<geoCoordinate id="B680F1B9FFA79508C3F9F8D8FE9CF8FB" box="[151,283,1835,1863]" direction="east" orientation="longitude" pageId="1" pageNumber="112" precision="1" value="16.467417">16o28.045E</geoCoordinate>
, 
<elevation id="5899704DFFA79508C249F8D8FE10F8FB" box="[295,407,1838,1863]" metricMagnitude="2" metricUnit="m" metricValue="6.67" pageId="1" pageNumber="112" unit="m" value="667.0">
<quantity id="144C3A9BFFA79508C249F8D8FEE9F8FB" box="[295,366,1838,1863]" metricMagnitude="2" metricUnit="m" metricValue="6.67" pageId="1" pageNumber="112" unit="m" value="667.0">667 m</quantity>
asl
</elevation>
, leg. 
<collectorName id="7E41F2A8FFA79508C2BCF8C6FD16F8FB" box="[466,657,1838,1863]" pageId="1" pageNumber="112">K. Dózsa-Farkas</collectorName>
, J. 
<collectorName id="7E41F2A8FFA79508C1DBF8D8FC84F8FB" box="[693,771,1838,1863]" pageId="1" pageNumber="112">Farkas</collectorName>
, M. 
<collectorName id="7E41F2A8FFA79508C05AF8D8FC25F8FB" box="[820,930,1838,1863]" pageId="1" pageNumber="112">Pobozsny</collectorName>
, 
<collectingDate id="B74E4856FFA79508C0C3F8D8FBA8F8FB" box="[941,1071,1838,1863]" pageId="1" pageNumber="112" value="2016-10-24">
<date id="A70AB1BEFFA79508C0C3F8D8FBA8F8FB" box="[941,1071,1838,1863]" pageId="1" pageNumber="112" value="2016-10-24">24.10.2016</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph id="D30B977EFFA79508C3A9F8A5FA1AF865" blockId="1.[151,1437,1761,2009]" pageId="1" pageNumber="112">
<emphasis id="E1C04B6CFFA79508C3A9F8A5FEB8F8D0" bold="true" box="[199,319,1875,1900]" pageId="1" pageNumber="112">Paratypes</emphasis>
(in total 14 specimens). 
<emphasis id="E1C04B6CFFA79508C10BF8A5FCABF8D0" bold="true" box="[613,812,1875,1900]" pageId="1" pageNumber="112">P.112.1.1–112.1.4</emphasis>
, slides 
<emphasis id="E1C04B6CFFA79508C0E7F8A5FBB9F8D0" bold="true" box="[905,1086,1875,1900]" pageId="1" pageNumber="112">No. 2218, 2219</emphasis>
(DNS 978), 
<emphasis id="E1C04B6CFFA79508C7B4F8A5FADBF8D0" bold="true" box="[1242,1372,1875,1900]" pageId="1" pageNumber="112">2225, 2226</emphasis>
, four adult, stained specimens from Rax Mts., mixed forest (
<taxonomicName id="14B4ECFDFFA79508C072F88CFC26F82D" box="[796,929,1913,1937]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="1" pageNumber="112" phylum="Tracheophyta" rank="species" species="abies">
<emphasis id="E1C04B6CFFA79508C072F88CFC26F82D" box="[796,929,1913,1937]" italics="true" pageId="1" pageNumber="112">Picea abies</emphasis>
</taxonomicName>
, 
<taxonomicName id="14B4ECFDFFA79508C0DDF88FFBE6F82D" authorityName="Roth" authorityYear="1788" box="[947,1121,1913,1937]" class="Magnoliopsida" family="Betulaceae" genus="Betula" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="112" phylum="Tracheophyta" rank="species" species="pendula">
<emphasis id="E1C04B6CFFA79508C0DDF88FFBE6F82D" box="[947,1121,1913,1937]" italics="true" pageId="1" pageNumber="112">Betula pendula</emphasis>
</taxonomicName>
), 47o40.538N, 15o43.140E, 
<quantity id="144C3A9BFFA79508C3F9F86BFF69F809" box="[151,238,1949,1974]" metricMagnitude="3" metricUnit="m" metricValue="1.116" pageId="1" pageNumber="112" unit="m" value="1116.0">1116 m</quantity>
asl, leg. J. Farkas, 
<date id="A70AB1BEFFA79508C2B9F86BFDDEF80A" box="[471,601,1949,1974]" pageId="1" pageNumber="112" value="2016-05-13">13.05.2016</date>
. 
<emphasis id="E1C04B6CFFA79508C109F86BFCB7F80A" bold="true" box="[615,816,1949,1974]" pageId="1" pageNumber="112">P.112.2.1–112.2.7</emphasis>
, slides 
<emphasis id="E1C04B6CFFA79508C0E2F86BFB2AF80A" bold="true" box="[908,1197,1949,1974]" pageId="1" pageNumber="112">
No. 2266a–c– 
<date id="A70AB1BEFFA79508C740F86BFB2AF80A" box="[1070,1197,1949,1974]" pageId="1" pageNumber="112" value="2268" valueMax="2269">2268–2269</date>
</emphasis>
(DNS 1102), 
<date id="A70AB1BEFFA79508C638F86BFF48F865" pageId="1" pageNumber="112" value="2273" valueMax="2274">
<emphasis id="E1C04B6CFFA79508C638F86BFF48F865" bold="true" pageId="1" pageNumber="112">2273– 2274</emphasis>
</date>
, 
<emphasis id="E1C04B6CFFA79508C3B3F836FE92F865" bold="true" box="[221,277,1984,2009]" pageId="1" pageNumber="112">2345</emphasis>
(DNS 1116), 
<emphasis id="E1C04B6CFFA79508C2DDF836FE6CF865" bold="true" box="[435,491,1984,2009]" pageId="1" pageNumber="112">2346</emphasis>
, 7 adult specimens, stained (except P.112.2.4 No. 2273), whole-mounted specimens
</paragraph>
<paragraph id="D30B977EFFA4950BC3F9FF61FC1AFF68" blockId="2.[151,1437,151,2035]" pageId="2" pageNumber="113">
from the type locality, leg. K. Dózsa-Farkas, J. Farkas, M. Pobozsny, 
<date id="A70AB1BEFFA4950BC0DBFF61FBB0FF0C" box="[949,1079,151,176]" pageId="2" pageNumber="113" value="2016-10-24">24.10.2016</date>
. 
<emphasis id="E1C04B6CFFA4950BC72DFF61FB13FF0C" bold="true" box="[1091,1172,151,176]" pageId="2" pageNumber="113">P.112.3</emphasis>
, one specimen in 70% ethanol (body end was processed for molecular analysis, DNS 1114).
</paragraph>
<paragraph id="D30B977EFFA4950BC3A9FF29FD64FEA0" blockId="2.[151,1437,151,2035]" pageId="2" pageNumber="113">
<emphasis id="E1C04B6CFFA4950BC3A9FF29FD8AFF44" bold="true" box="[199,525,223,248]" pageId="2" pageNumber="113">Further material examined</emphasis>
. Four juvenile and two adult specimens 
<emphasis id="E1C04B6CFFA4950BC0B9FF16FBA4FF4B" box="[983,1059,224,247]" italics="true" pageId="2" pageNumber="113">in vivo</emphasis>
(the whole adult specimens were processed for molecular analysis, 
<collectionCode id="B5A50FBBFFA4950BC17DFEF5FDD7FEA7" box="[531,592,259,283]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="2" pageNumber="113">DNA</collectionCode>
1009, 1115).
</paragraph>
</subSubSection>
<subSubSection id="9BAEC4F5FFA4950BC3A9FED0FA97FE83" box="[199,1296,294,319]" pageId="2" pageNumber="113" type="etymology">
<paragraph id="D30B977EFFA4950BC3A9FED0FA97FE83" blockId="2.[151,1437,151,2035]" box="[199,1296,294,319]" pageId="2" pageNumber="113">
<emphasis id="E1C04B6CFFA4950BC3A9FED0FEC9FE83" bold="true" box="[199,334,294,319]" pageId="2" pageNumber="113">Etymology.</emphasis>
Named after the chaetae, which are present only in the ventro-lateral chaetal bundles.
</paragraph>
</subSubSection>
<subSubSection id="9BAEC4F5FFA4950BC3A9FEBCFAE2FD3D" pageId="2" pageNumber="113" type="diagnosis">
<paragraph id="D30B977EFFA4950BC3A9FEBCFAE2FD3D" blockId="2.[151,1437,151,2035]" pageId="2" pageNumber="113">
<emphasis id="E1C04B6CFFA4950BC3A9FEBCFEC6FEDF" bold="true" box="[199,321,330,355]" pageId="2" pageNumber="113">Diagnosis.</emphasis>
The new species can be recognized by the following combination of characters: (1) large size (body length 
<quantity id="144C3A9BFFA4950BC389FE98FEE3FE3A" box="[231,356,366,391]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.4" metricValueMin="1.5" pageId="2" pageNumber="113" unit="mm" value="19.5" valueMax="24.0" valueMin="15.0">15–24 mm</quantity>
<emphasis id="E1C04B6CFFA4950BC200FE99FE3AFE3A" box="[366,445,367,390]" italics="true" pageId="2" pageNumber="113">in vivo</emphasis>
), segments (43) 48–60; (2) lateral chaetae absent, ventral maximum 3–4 chaetae per bundle; (3) clitellum widely interrupted dorsally and the hyalocytes and granulocytes arranged in dense rows laterally and after the bursal opening; (4) body wall strong and cuticle thick (8–14 µm); (5) five preclitellar pairs of nephridia; (6) coelomo-mucocytes b-type, lenticytes large, 8–12 µm long; (7) blood light pink; (8) chylus cells in XII–XV, occupying 2–3 segments; (9) bursal slit longitudinal with transverse extensions; (10) seminal vesicle large; (11) subneural glands absent; (12) sperm funnel approximately as long as body diameter or somewhat longer, collar narrower than funnel diameter, spermatozoa long, difficult to measure, sperm heads 150–250 µm long (
<emphasis id="E1C04B6CFFA4950BC6E8FDB3FF41FD3C" italics="true" pageId="2" pageNumber="113">in vivo</emphasis>
); (13) spermatheca with 4–6 stalked diverticula, long ectal duct without ectal glands and separate entally.
</paragraph>
</subSubSection>
<subSubSection id="9BAEC4F5FFA4950BC3A9FD7AFDFFF817" pageId="2" pageNumber="113" type="description">
<paragraph id="D30B977EFFA4950BC3A9FD7AFDFFF817" blockId="2.[151,1437,151,2035]" pageId="2" pageNumber="113">
<emphasis id="E1C04B6CFFA4950BC3A9FD7AFEDEFD19" bold="true" box="[199,345,652,677]" pageId="2" pageNumber="113">Description.</emphasis>
Large, whitish, stiff worms. 
<typeStatus id="0C0F29DCFFA4950BC1C7FD7AFC96FD19" box="[681,785,652,677]" pageId="2" pageNumber="113" type="holotype">Holotype</typeStatus>
<quantity id="144C3A9BFFA4950BC075FD7BFC05FD18" box="[795,898,653,677]" metricMagnitude="-2" metricUnit="m" metricValue="2.33" pageId="2" pageNumber="113" unit="mm" value="23.3">23.3 mm</quantity>
long, 530 µm wide at VIII and 630 µm at the clitellum (
<emphasis id="E1C04B6CFFA4950BC265FD46FED1FD7B" box="[267,342,688,711]" italics="true" pageId="2" pageNumber="113">in vivo</emphasis>
). The fixed worm 19.0 mm long, 700 µm wide at VIII and 750 µm at the clitellum, 56 segments. Body length of the 
<typeStatus id="0C0F29DCFFA4950BC217FD22FE63FD50" box="[377,484,724,748]" pageId="2" pageNumber="113" type="paratype">paratypes</typeStatus>
<quantity id="144C3A9BFFA4950BC280FD25FDECFD57" box="[494,619,723,748]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.4" metricValueMin="1.5" pageId="2" pageNumber="113" unit="mm" value="19.5" valueMax="24.0" valueMin="15.0">15–24 mm</quantity>
, width 500–620 µm at VIII and 600–730 µm at the clitellum (
<emphasis id="E1C04B6CFFA4950BC650FD22FA0BFD57" box="[1342,1420,724,747]" italics="true" pageId="2" pageNumber="113">in vivo</emphasis>
). Length of fixed specimens 
<quantity id="144C3A9BFFA4950BC2A5FD01FDF7FCB3" box="[459,624,759,784]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.85" metricValueMin="1.15" pageId="2" pageNumber="113" unit="mm" value="15.0" valueMax="18.5" valueMin="11.5">11.5–18.5 mm</quantity>
, width 520–700 µm at VIII and 600–770 µm at the clitellum. Segments (42), 48–60. Chaetae only in ventro-lateral bundles, except in one specimen (P. 112.2.5. slide No. 2273), here 1 chaeta in 1 lateral bundle of IV. Chaetal formula: 0 – 0: 1,2,3,(4) – (3),2,1,0,1. Inner chaetae shorter and thinner than outer: 58– 
<date id="A70AB1BEFFA4950BC22EFC94FE24FCC7" box="[320,419,866,891]" pageId="2" pageNumber="113" value="1970-10-05" valueMax="1970-10-06" valueMin="1970-10-05">70 x 5–6</date>
µm against 70– 
<date id="A70AB1BEFFA4950BC139FC95FD1AFCC7" box="[599,669,867,891]" pageId="2" pageNumber="113" value="1980-10-09">80 x 9</date>
µm and 34– 
<date id="A70AB1BEFFA4950BC047FC95FCE8FCC7" box="[809,879,867,891]" pageId="2" pageNumber="113" value="1942-10-04">42 x 4</date>
µm (in preclitellar bundles). Behind clitellum the two chaetae in a bundle of different size, at body end often only 1 chaeta per bundle, size about 90–110 
<date id="A70AB1BEFFA4950BC62EFC71FA1BFC23" box="[1344,1436,903,927]" pageId="2" pageNumber="113" value="2010-10-12">x 10–12</date>
µm. Head pore at 0/I. Dorsal pores from VII. Epidermal gland cells 5–6 transverse rows per segments (
<figureCitation id="4B8F8BFBFFA4950BC65BFC5CFA09FC7E" box="[1333,1422,938,963]" captionStart="FIGURE 1" captionStartId="3.[151,250,1802,1824]" captionTargetBox="[158,1372,193,1745]" captionTargetId="figure@3.[158,1428,193,1780]" captionTargetPageId="3" captionText="FIGURE 1. Micrographs of Fridericia ventrochaetosa sp. n. A. Brain (DNA 1109). B. Brain (holotype). C. Epidermal glands. D. Thick body wall and cuticle (marked with arrow). E. Strong septa, 5/6– 9/10 (holotype). F–G. Head region (the thick epithelium of prostomium marked with arrow). H. Clitellar glands, lateral view. I. Clitellum of a dissected species (above the dorsal gap without glands, below the clitellar glands in transverse rows). J–K. Coelomocytes (J, paratype P.112.1.2, No. 2219=DNA 978). L. Nephridium of a juvenile specimen, in the middle part of body; M. Preclitellar nephridium (paratype P.112.1.2, No. 2219=DNA 978). A, C, H, J, L–M in vivo, B, E fixed, not stained, D, F–G, I, K. fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447055/files/figure.png" pageId="2" pageNumber="113">Fig. 1C</figureCitation>
). Epidermis glandular around spermathecal pore (
<figureCitation id="4B8F8BFBFFA4950BC1D1FC3BFC9BFC5A" box="[703,796,973,998]" captionStart="FIGURE 3" captionStartId="5.[151,250,1618,1640]" captionTargetBox="[178,1398,288,1505]" captionTargetId="figure@5.[178,1408,288,1524]" captionTargetPageId="5" captionText="FIGURE 3. Micrograph of Fridericia ventrochaetosa sp. n. A. Sperm funnels (paratype P.112.1.2, No. 2219=DNA 978). B, D. Spermathecae (paratype P.112.2.6, No. 2345=DNA 1116). C. Spermathecae (holotype). E. Spermatheca (paratype P.112.1.2, No. 2219=DNA 978). F. Dissected spermatheca (paratype P.112.2.1, No. 2266). G. Epidermal gland cells around the opening of spermatheca (opening marked with arrow). A, B, D, G in vivo, E, F fixed, stained, C fixed, not stained. Scale bars in A 20 µm, in the rest 50 µm." httpUri="https://zenodo.org/record/1447059/files/figure.png" pageId="2" pageNumber="113">Fig. 3G</figureCitation>
). Clitellum in 
<date id="A70AB1BEFFA4950BC0ABFC38FB80FC5A" box="[965,1031,974,998]" bridgedPair="-" pageId="2" pageNumber="113">XII–1</date>
/2XIII. Glands dorsally absent, gap about 200–400 µm wide (
<figureCitation id="4B8F8BFBFFA4950BC2D7FC07FD82FBB6" box="[441,517,1009,1034]" captionStart="FIGURE 1" captionStartId="3.[151,250,1802,1824]" captionTargetBox="[158,1372,193,1745]" captionTargetId="figure@3.[158,1428,193,1780]" captionTargetPageId="3" captionText="FIGURE 1. Micrographs of Fridericia ventrochaetosa sp. n. A. Brain (DNA 1109). B. Brain (holotype). C. Epidermal glands. D. Thick body wall and cuticle (marked with arrow). E. Strong septa, 5/6– 9/10 (holotype). F–G. Head region (the thick epithelium of prostomium marked with arrow). H. Clitellar glands, lateral view. I. Clitellum of a dissected species (above the dorsal gap without glands, below the clitellar glands in transverse rows). J–K. Coelomocytes (J, paratype P.112.1.2, No. 2219=DNA 978). L. Nephridium of a juvenile specimen, in the middle part of body; M. Preclitellar nephridium (paratype P.112.1.2, No. 2219=DNA 978). A, C, H, J, L–M in vivo, B, E fixed, not stained, D, F–G, I, K. fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447055/files/figure.png" pageId="2" pageNumber="113">
Fig. 
<date id="A70AB1BEFFA4950BC280FC04FD82FBB6" box="[494,517,1010,1034]" pageId="2" pageNumber="113">1I</date>
</figureCitation>
), ventrally absent between and anterior to male apparatus, but present posteriorly (
<figureCitation id="4B8F8BFBFFA4950BC3F1FBE3FF7DFB92" box="[159,250,1045,1070]" captionStart="FIGURE 2" captionStartId="4.[151,250,1802,1824]" captionTargetBox="[158,1376,193,1743]" captionTargetId="figure@4.[158,1428,193,1780]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Fridericia ventrochaetosa sp. n. A. End of oesophageal appendage with multiple branches (arrows). B. Male copulatory organs (arrows); clitellar gland cells visible laterally and ventrally posterior to copulatory organs. C. Midgut pars tumida (arrows) in XXXI–XXXIII (holotype). D. Chylus cells in XIII–XIV. E. Male copulatory organs, ventral view, holotype. Strong bursal muscle marked with arrow. Asterisk: region between the two organs without clitellar glands. F– G. Bursal slits (marked with arrow in G). H. Male copulatory organ, lateral view (paratype P.112.1.2, No. 2219=DNA 978). I,J. Sperm funnels (paratype P.112.1.2, No. 2219=DNA 978). A, F, G, I in vivo, B, D, H, J fixed, stained, C, E fixed, not stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447057/files/figure.png" pageId="2" pageNumber="113">Fig. 2B</figureCitation>
), hyalocytes larger than granulocytes, arranged in dense transverse rows (
<figureCitation id="4B8F8BFBFFA4950BC736FBE3FB32FB92" box="[1112,1205,1045,1070]" captionStart="FIGURE 1" captionStartId="3.[151,250,1802,1824]" captionTargetBox="[158,1372,193,1745]" captionTargetId="figure@3.[158,1428,193,1780]" captionTargetPageId="3" captionText="FIGURE 1. Micrographs of Fridericia ventrochaetosa sp. n. A. Brain (DNA 1109). B. Brain (holotype). C. Epidermal glands. D. Thick body wall and cuticle (marked with arrow). E. Strong septa, 5/6– 9/10 (holotype). F–G. Head region (the thick epithelium of prostomium marked with arrow). H. Clitellar glands, lateral view. I. Clitellum of a dissected species (above the dorsal gap without glands, below the clitellar glands in transverse rows). J–K. Coelomocytes (J, paratype P.112.1.2, No. 2219=DNA 978). L. Nephridium of a juvenile specimen, in the middle part of body; M. Preclitellar nephridium (paratype P.112.1.2, No. 2219=DNA 978). A, C, H, J, L–M in vivo, B, E fixed, not stained, D, F–G, I, K. fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447055/files/figure.png" pageId="2" pageNumber="113">Fig. 1H</figureCitation>
). Body wall strong, thickness about 40–70 µm, cuticle thick, about 8–14 µm 
<emphasis id="E1C04B6CFFA4950BC073FBCCFCEDFBED" box="[797,874,1082,1105]" italics="true" pageId="2" pageNumber="113">in vivo</emphasis>
and fixed (
<figureCitation id="4B8F8BFBFFA4950BC082FBCFFBC2FBED" box="[1004,1093,1081,1106]" captionStart="FIGURE 1" captionStartId="3.[151,250,1802,1824]" captionTargetBox="[158,1372,193,1745]" captionTargetId="figure@3.[158,1428,193,1780]" captionTargetPageId="3" captionText="FIGURE 1. Micrographs of Fridericia ventrochaetosa sp. n. A. Brain (DNA 1109). B. Brain (holotype). C. Epidermal glands. D. Thick body wall and cuticle (marked with arrow). E. Strong septa, 5/6– 9/10 (holotype). F–G. Head region (the thick epithelium of prostomium marked with arrow). H. Clitellar glands, lateral view. I. Clitellum of a dissected species (above the dorsal gap without glands, below the clitellar glands in transverse rows). J–K. Coelomocytes (J, paratype P.112.1.2, No. 2219=DNA 978). L. Nephridium of a juvenile specimen, in the middle part of body; M. Preclitellar nephridium (paratype P.112.1.2, No. 2219=DNA 978). A, C, H, J, L–M in vivo, B, E fixed, not stained, D, F–G, I, K. fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447055/files/figure.png" pageId="2" pageNumber="113">Fig. 1D</figureCitation>
), (in forepart slightly stronger than at the body end). Brain egg-shaped, about 200–240 µm long, about two times longer than wide 
<emphasis id="E1C04B6CFFA4950BC67FFBABFAD9FBC8" box="[1297,1374,1117,1140]" italics="true" pageId="2" pageNumber="113">in vivo</emphasis>
(
<figureCitation id="4B8F8BFBFFA4950BC600FBAAFF3EFB25" captionStart="FIGURE 1" captionStartId="3.[151,250,1802,1824]" captionTargetBox="[158,1372,193,1745]" captionTargetId="figure@3.[158,1428,193,1780]" captionTargetPageId="3" captionText="FIGURE 1. Micrographs of Fridericia ventrochaetosa sp. n. A. Brain (DNA 1109). B. Brain (holotype). C. Epidermal glands. D. Thick body wall and cuticle (marked with arrow). E. Strong septa, 5/6– 9/10 (holotype). F–G. Head region (the thick epithelium of prostomium marked with arrow). H. Clitellar glands, lateral view. I. Clitellum of a dissected species (above the dorsal gap without glands, below the clitellar glands in transverse rows). J–K. Coelomocytes (J, paratype P.112.1.2, No. 2219=DNA 978). L. Nephridium of a juvenile specimen, in the middle part of body; M. Preclitellar nephridium (paratype P.112.1.2, No. 2219=DNA 978). A, C, H, J, L–M in vivo, B, E fixed, not stained, D, F–G, I, K. fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447055/files/figure.png" pageId="2" pageNumber="113">Fig. 1A</figureCitation>
) and 170–203 µm long and 1.4–1.8 times longer than wide in the fixed specimens (
<figureCitation id="4B8F8BFBFFA4950BC7F6FB76FB75FB25" box="[1176,1266,1152,1177]" captionStart="FIGURE 1" captionStartId="3.[151,250,1802,1824]" captionTargetBox="[158,1372,193,1745]" captionTargetId="figure@3.[158,1428,193,1780]" captionTargetPageId="3" captionText="FIGURE 1. Micrographs of Fridericia ventrochaetosa sp. n. A. Brain (DNA 1109). B. Brain (holotype). C. Epidermal glands. D. Thick body wall and cuticle (marked with arrow). E. Strong septa, 5/6– 9/10 (holotype). F–G. Head region (the thick epithelium of prostomium marked with arrow). H. Clitellar glands, lateral view. I. Clitellum of a dissected species (above the dorsal gap without glands, below the clitellar glands in transverse rows). J–K. Coelomocytes (J, paratype P.112.1.2, No. 2219=DNA 978). L. Nephridium of a juvenile specimen, in the middle part of body; M. Preclitellar nephridium (paratype P.112.1.2, No. 2219=DNA 978). A, C, H, J, L–M in vivo, B, E fixed, not stained, D, F–G, I, K. fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447055/files/figure.png" pageId="2" pageNumber="113">Fig. 1B</figureCitation>
). Oesophageal appendages long with numerous, elongate branches at the end in V–VI (
<figureCitation id="4B8F8BFBFFA4950BC084FB52FBCEFB01" box="[1002,1097,1188,1213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1802,1824]" captionTargetBox="[158,1376,193,1743]" captionTargetId="figure@4.[158,1428,193,1780]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Fridericia ventrochaetosa sp. n. A. End of oesophageal appendage with multiple branches (arrows). B. Male copulatory organs (arrows); clitellar gland cells visible laterally and ventrally posterior to copulatory organs. C. Midgut pars tumida (arrows) in XXXI–XXXIII (holotype). D. Chylus cells in XIII–XIV. E. Male copulatory organs, ventral view, holotype. Strong bursal muscle marked with arrow. Asterisk: region between the two organs without clitellar glands. F– G. Bursal slits (marked with arrow in G). H. Male copulatory organ, lateral view (paratype P.112.1.2, No. 2219=DNA 978). I,J. Sperm funnels (paratype P.112.1.2, No. 2219=DNA 978). A, F, G, I in vivo, B, D, H, J fixed, stained, C, E fixed, not stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447057/files/figure.png" pageId="2" pageNumber="113">Fig. 2A</figureCitation>
). All pharyngeal glands with ventral lobes, those in 4/5 mostly united dorsally, those in 5/6 and 6/7 weakly united dorsally, occasionally all three pairs unconnected dorsally. Septa of V–X thickened (
<figureCitation id="4B8F8BFBFFA4950BC07EFB1DFCEDFAB8" box="[784,874,1259,1284]" captionStart="FIGURE 1" captionStartId="3.[151,250,1802,1824]" captionTargetBox="[158,1372,193,1745]" captionTargetId="figure@3.[158,1428,193,1780]" captionTargetPageId="3" captionText="FIGURE 1. Micrographs of Fridericia ventrochaetosa sp. n. A. Brain (DNA 1109). B. Brain (holotype). C. Epidermal glands. D. Thick body wall and cuticle (marked with arrow). E. Strong septa, 5/6– 9/10 (holotype). F–G. Head region (the thick epithelium of prostomium marked with arrow). H. Clitellar glands, lateral view. I. Clitellum of a dissected species (above the dorsal gap without glands, below the clitellar glands in transverse rows). J–K. Coelomocytes (J, paratype P.112.1.2, No. 2219=DNA 978). L. Nephridium of a juvenile specimen, in the middle part of body; M. Preclitellar nephridium (paratype P.112.1.2, No. 2219=DNA 978). A, C, H, J, L–M in vivo, B, E fixed, not stained, D, F–G, I, K. fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447055/files/figure.png" pageId="2" pageNumber="113">Fig. 1E</figureCitation>
). Chloragocytes from V, 15–25 µm long (fixed specimens). Dorsal vessel from XX–XXIII, blood light pink. Midgut pars tumida in XXX–XXXVIII occupying 5– 7 segments (
<figureCitation id="4B8F8BFBFFA4950BC24DFAC5FEFCFAF7" box="[291,379,1331,1356]" captionStart="FIGURE 2" captionStartId="4.[151,250,1802,1824]" captionTargetBox="[158,1376,193,1743]" captionTargetId="figure@4.[158,1428,193,1780]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Fridericia ventrochaetosa sp. n. A. End of oesophageal appendage with multiple branches (arrows). B. Male copulatory organs (arrows); clitellar gland cells visible laterally and ventrally posterior to copulatory organs. C. Midgut pars tumida (arrows) in XXXI–XXXIII (holotype). D. Chylus cells in XIII–XIV. E. Male copulatory organs, ventral view, holotype. Strong bursal muscle marked with arrow. Asterisk: region between the two organs without clitellar glands. F– G. Bursal slits (marked with arrow in G). H. Male copulatory organ, lateral view (paratype P.112.1.2, No. 2219=DNA 978). I,J. Sperm funnels (paratype P.112.1.2, No. 2219=DNA 978). A, F, G, I in vivo, B, D, H, J fixed, stained, C, E fixed, not stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447057/files/figure.png" pageId="2" pageNumber="113">Fig. 2C</figureCitation>
). Five pairs of preclitellar nephridia from 
<date id="A70AB1BEFFA4950BC03EFAC5FC5FFAF7" box="[848,984,1331,1356]" pageId="2" pageNumber="113" value="2011-07-06" valueMax="2011-10-06" valueMin="2011-07-06">6/7 to 10/11</date>
, length ratio anteseptale: postseptale 1: 1.5–2, adseptal origin of the long efferent duct. The nephrostome not embedded in the anterior part, oriented horizontally (infrequent in 
<taxonomicName id="14B4ECFDFFA4950BC2B4FA8DFDCAFA2F" box="[474,589,1403,1427]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="113" phylum="Magnoliopsida" rank="genus">
<emphasis id="E1C04B6CFFA4950BC2B4FA8DFDCAFA2F" box="[474,589,1403,1427]" italics="true" pageId="2" pageNumber="113">Fridericia</emphasis>
</taxonomicName>
species) (
<figureCitation id="4B8F8BFBFFA4950BC1A5FA8CFCDCFA2F" box="[715,859,1402,1427]" captionStart="FIGURE 1" captionStartId="3.[151,250,1802,1824]" captionTargetBox="[158,1372,193,1745]" captionTargetId="figure@3.[158,1428,193,1780]" captionTargetPageId="3" captionText="FIGURE 1. Micrographs of Fridericia ventrochaetosa sp. n. A. Brain (DNA 1109). B. Brain (holotype). C. Epidermal glands. D. Thick body wall and cuticle (marked with arrow). E. Strong septa, 5/6– 9/10 (holotype). F–G. Head region (the thick epithelium of prostomium marked with arrow). H. Clitellar glands, lateral view. I. Clitellum of a dissected species (above the dorsal gap without glands, below the clitellar glands in transverse rows). J–K. Coelomocytes (J, paratype P.112.1.2, No. 2219=DNA 978). L. Nephridium of a juvenile specimen, in the middle part of body; M. Preclitellar nephridium (paratype P.112.1.2, No. 2219=DNA 978). A, C, H, J, L–M in vivo, B, E fixed, not stained, D, F–G, I, K. fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447055/files/figure.png" pageId="2" pageNumber="113">Figs. 1L–M</figureCitation>
). Coelomo-mucocytes b-type, 
<emphasis id="E1C04B6CFFA4950BC7A9FA8DFB6DFA2E" box="[1223,1258,1403,1426]" italics="true" pageId="2" pageNumber="113">i.e.</emphasis>
with refractile vesicles at cell periphery, length 34–53 µm 
<emphasis id="E1C04B6CFFA4950BC1E4FA69FD5FFA0A" box="[650,728,1439,1462]" italics="true" pageId="2" pageNumber="113">in vivo</emphasis>
(
<figureCitation id="4B8F8BFBFFA4950BC187FA68FCBFFA0B" box="[745,824,1438,1463]" captionStart="FIGURE 1" captionStartId="3.[151,250,1802,1824]" captionTargetBox="[158,1372,193,1745]" captionTargetId="figure@3.[158,1428,193,1780]" captionTargetPageId="3" captionText="FIGURE 1. Micrographs of Fridericia ventrochaetosa sp. n. A. Brain (DNA 1109). B. Brain (holotype). C. Epidermal glands. D. Thick body wall and cuticle (marked with arrow). E. Strong septa, 5/6– 9/10 (holotype). F–G. Head region (the thick epithelium of prostomium marked with arrow). H. Clitellar glands, lateral view. I. Clitellum of a dissected species (above the dorsal gap without glands, below the clitellar glands in transverse rows). J–K. Coelomocytes (J, paratype P.112.1.2, No. 2219=DNA 978). L. Nephridium of a juvenile specimen, in the middle part of body; M. Preclitellar nephridium (paratype P.112.1.2, No. 2219=DNA 978). A, C, H, J, L–M in vivo, B, E fixed, not stained, D, F–G, I, K. fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447055/files/figure.png" pageId="2" pageNumber="113">Fig. 1J</figureCitation>
), 20–35 µm in the fixed worms (
<figureCitation id="4B8F8BFBFFA4950BC7D8FA68FA96FA0B" box="[1206,1297,1438,1463]" captionStart="FIGURE 1" captionStartId="3.[151,250,1802,1824]" captionTargetBox="[158,1372,193,1745]" captionTargetId="figure@3.[158,1428,193,1780]" captionTargetPageId="3" captionText="FIGURE 1. Micrographs of Fridericia ventrochaetosa sp. n. A. Brain (DNA 1109). B. Brain (holotype). C. Epidermal glands. D. Thick body wall and cuticle (marked with arrow). E. Strong septa, 5/6– 9/10 (holotype). F–G. Head region (the thick epithelium of prostomium marked with arrow). H. Clitellar glands, lateral view. I. Clitellum of a dissected species (above the dorsal gap without glands, below the clitellar glands in transverse rows). J–K. Coelomocytes (J, paratype P.112.1.2, No. 2219=DNA 978). L. Nephridium of a juvenile specimen, in the middle part of body; M. Preclitellar nephridium (paratype P.112.1.2, No. 2219=DNA 978). A, C, H, J, L–M in vivo, B, E fixed, not stained, D, F–G, I, K. fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447055/files/figure.png" pageId="2" pageNumber="113">Fig. 1K</figureCitation>
). Lenticytes scarce, large, 8–12 µm long. Chylus cells in XII–XV, occupying 2–3 segments (
<figureCitation id="4B8F8BFBFFA4950BC777FA34FBF4FA66" box="[1049,1139,1474,1499]" captionStart="FIGURE 2" captionStartId="4.[151,250,1802,1824]" captionTargetBox="[158,1376,193,1743]" captionTargetId="figure@4.[158,1428,193,1780]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Fridericia ventrochaetosa sp. n. A. End of oesophageal appendage with multiple branches (arrows). B. Male copulatory organs (arrows); clitellar gland cells visible laterally and ventrally posterior to copulatory organs. C. Midgut pars tumida (arrows) in XXXI–XXXIII (holotype). D. Chylus cells in XIII–XIV. E. Male copulatory organs, ventral view, holotype. Strong bursal muscle marked with arrow. Asterisk: region between the two organs without clitellar glands. F– G. Bursal slits (marked with arrow in G). H. Male copulatory organ, lateral view (paratype P.112.1.2, No. 2219=DNA 978). I,J. Sperm funnels (paratype P.112.1.2, No. 2219=DNA 978). A, F, G, I in vivo, B, D, H, J fixed, stained, C, E fixed, not stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447057/files/figure.png" pageId="2" pageNumber="113">Fig. 2D</figureCitation>
). Seminal vesicle in IX–X or XI–XII. Sperm funnels cylindrical, mostly tapering distad (
<figureCitation id="4B8F8BFBFFA4950BC00FFA13FC36FA42" box="[865,945,1509,1534]" captionStart="FIGURE 2" captionStartId="4.[151,250,1802,1824]" captionTargetBox="[158,1376,193,1743]" captionTargetId="figure@4.[158,1428,193,1780]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Fridericia ventrochaetosa sp. n. A. End of oesophageal appendage with multiple branches (arrows). B. Male copulatory organs (arrows); clitellar gland cells visible laterally and ventrally posterior to copulatory organs. C. Midgut pars tumida (arrows) in XXXI–XXXIII (holotype). D. Chylus cells in XIII–XIV. E. Male copulatory organs, ventral view, holotype. Strong bursal muscle marked with arrow. Asterisk: region between the two organs without clitellar glands. F– G. Bursal slits (marked with arrow in G). H. Male copulatory organ, lateral view (paratype P.112.1.2, No. 2219=DNA 978). I,J. Sperm funnels (paratype P.112.1.2, No. 2219=DNA 978). A, F, G, I in vivo, B, D, H, J fixed, stained, C, E fixed, not stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447057/files/figure.png" pageId="2" pageNumber="113">
Fig. 
<date id="A70AB1BEFFA4950BC0F6FA10FC36FA42" box="[920,945,1510,1534]" pageId="2" pageNumber="113">2I</date>
</figureCitation>
), about 600–860 µm long and 2.4–3 times longer than wide (in one specimen 4 times longer than wide) (
<emphasis id="E1C04B6CFFA4950BC039F9FCFC24F99D" box="[855,931,1546,1569]" italics="true" pageId="2" pageNumber="113">in vivo</emphasis>
). Funnel length in fixed specimens 350–550 µm, funnel 1.5–2.5 times longer than wide (
<figureCitation id="4B8F8BFBFFA4950BC1E1F9DBFD6DF9FA" box="[655,746,1581,1606]" captionStart="FIGURE 2" captionStartId="4.[151,250,1802,1824]" captionTargetBox="[158,1376,193,1743]" captionTargetId="figure@4.[158,1428,193,1780]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Fridericia ventrochaetosa sp. n. A. End of oesophageal appendage with multiple branches (arrows). B. Male copulatory organs (arrows); clitellar gland cells visible laterally and ventrally posterior to copulatory organs. C. Midgut pars tumida (arrows) in XXXI–XXXIII (holotype). D. Chylus cells in XIII–XIV. E. Male copulatory organs, ventral view, holotype. Strong bursal muscle marked with arrow. Asterisk: region between the two organs without clitellar glands. F– G. Bursal slits (marked with arrow in G). H. Male copulatory organ, lateral view (paratype P.112.1.2, No. 2219=DNA 978). I,J. Sperm funnels (paratype P.112.1.2, No. 2219=DNA 978). A, F, G, I in vivo, B, D, H, J fixed, stained, C, E fixed, not stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447057/files/figure.png" pageId="2" pageNumber="113">Figs. 2J</figureCitation>
, 
<figureCitation id="4B8F8BFBFFA4950BC196F9D8FC9AF9FA" box="[760,797,1582,1606]" captionStart="FIGURE 3" captionStartId="5.[151,250,1618,1640]" captionTargetBox="[178,1398,288,1505]" captionTargetId="figure@5.[178,1408,288,1524]" captionTargetPageId="5" captionText="FIGURE 3. Micrograph of Fridericia ventrochaetosa sp. n. A. Sperm funnels (paratype P.112.1.2, No. 2219=DNA 978). B, D. Spermathecae (paratype P.112.2.6, No. 2345=DNA 1116). C. Spermathecae (holotype). E. Spermatheca (paratype P.112.1.2, No. 2219=DNA 978). F. Dissected spermatheca (paratype P.112.2.1, No. 2266). G. Epidermal gland cells around the opening of spermatheca (opening marked with arrow). A, B, D, G in vivo, E, F fixed, stained, C fixed, not stained. Scale bars in A 20 µm, in the rest 50 µm." httpUri="https://zenodo.org/record/1447059/files/figure.png" pageId="2" pageNumber="113">3A</figureCitation>
); collar narrower than funnel body. The entire length of spermatozoa is difficult to measure 
<emphasis id="E1C04B6CFFA4950BC146F9A4FDF3F9D5" box="[552,628,1618,1641]" italics="true" pageId="2" pageNumber="113">in vivo</emphasis>
, heads 150–250 µm, in fixed specimens spermatozoa 350–700 µm long and sperm heads 150–230 µm. Diameter of sperm ducts 9–10 µm (fixed). Male copulatory organs large, 260–350 µm long, 130–160 µm wide and 110–125 µm high (fixed), bursa large (
<figureCitation id="4B8F8BFBFFA4950BC0DEF96EFBC1F90D" box="[944,1094,1688,1713]" captionStart="FIGURE 2" captionStartId="4.[151,250,1802,1824]" captionTargetBox="[158,1376,193,1743]" captionTargetId="figure@4.[158,1428,193,1780]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Fridericia ventrochaetosa sp. n. A. End of oesophageal appendage with multiple branches (arrows). B. Male copulatory organs (arrows); clitellar gland cells visible laterally and ventrally posterior to copulatory organs. C. Midgut pars tumida (arrows) in XXXI–XXXIII (holotype). D. Chylus cells in XIII–XIV. E. Male copulatory organs, ventral view, holotype. Strong bursal muscle marked with arrow. Asterisk: region between the two organs without clitellar glands. F– G. Bursal slits (marked with arrow in G). H. Male copulatory organ, lateral view (paratype P.112.1.2, No. 2219=DNA 978). I,J. Sperm funnels (paratype P.112.1.2, No. 2219=DNA 978). A, F, G, I in vivo, B, D, H, J fixed, stained, C, E fixed, not stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447057/files/figure.png" pageId="2" pageNumber="113">Figs. 2B,E,H</figureCitation>
). Bursal slits longitudinal with additional transverse extensions (
<figureCitation id="4B8F8BFBFFA4950BC161F94AFD13F969" box="[527,660,1724,1749]" captionStart="FIGURE 2" captionStartId="4.[151,250,1802,1824]" captionTargetBox="[158,1376,193,1743]" captionTargetId="figure@4.[158,1428,193,1780]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Fridericia ventrochaetosa sp. n. A. End of oesophageal appendage with multiple branches (arrows). B. Male copulatory organs (arrows); clitellar gland cells visible laterally and ventrally posterior to copulatory organs. C. Midgut pars tumida (arrows) in XXXI–XXXIII (holotype). D. Chylus cells in XIII–XIV. E. Male copulatory organs, ventral view, holotype. Strong bursal muscle marked with arrow. Asterisk: region between the two organs without clitellar glands. F– G. Bursal slits (marked with arrow in G). H. Male copulatory organ, lateral view (paratype P.112.1.2, No. 2219=DNA 978). I,J. Sperm funnels (paratype P.112.1.2, No. 2219=DNA 978). A, F, G, I in vivo, B, D, H, J fixed, stained, C, E fixed, not stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447057/files/figure.png" pageId="2" pageNumber="113">Figs. 2F–G</figureCitation>
). Subneural glands absent. Spermathecae (
<figureCitation id="4B8F8BFBFFA4950BC718F94AFB71F969" box="[1142,1270,1724,1749]" captionStart="FIGURE 3" captionStartId="5.[151,250,1618,1640]" captionTargetBox="[178,1398,288,1505]" captionTargetId="figure@5.[178,1408,288,1524]" captionTargetPageId="5" captionText="FIGURE 3. Micrograph of Fridericia ventrochaetosa sp. n. A. Sperm funnels (paratype P.112.1.2, No. 2219=DNA 978). B, D. Spermathecae (paratype P.112.2.6, No. 2345=DNA 1116). C. Spermathecae (holotype). E. Spermatheca (paratype P.112.1.2, No. 2219=DNA 978). F. Dissected spermatheca (paratype P.112.2.1, No. 2266). G. Epidermal gland cells around the opening of spermatheca (opening marked with arrow). A, B, D, G in vivo, E, F fixed, stained, C fixed, not stained. Scale bars in A 20 µm, in the rest 50 µm." httpUri="https://zenodo.org/record/1447059/files/figure.png" pageId="2" pageNumber="113">Figs. 3B–F</figureCitation>
) similar to the spermathecae of 
<taxonomicName id="14B4ECFDFFA4950BC237F917FE31F944" baseAuthorityName="Hoffmeister" baseAuthorityYear="1843" box="[345,438,1760,1784]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="113" phylum="Magnoliopsida" rank="species" species="galba">
<emphasis id="E1C04B6CFFA4950BC237F917FE31F944" box="[345,438,1760,1784]" italics="true" pageId="2" pageNumber="113">F. galba</emphasis>
</taxonomicName>
(
<figureCitation id="4B8F8BFBFFA4950BC2A6F929FDD7F944" box="[456,592,1759,1784]" captionStart="FIGURE 5" captionStartId="7.[151,250,1919,1941]" captionTargetBox="[176,1391,1086,1861]" captionTargetId="figure@7.[176,1410,1086,1877]" captionTargetPageId="7" captionText="FIGURE 5. Micrographs of Fridericia galba. A. Bursal slit (arrow). B. Male copulatory organs, ventral view. C. Sperm funnels. D, E. Spermathecae. A, C–E in vivo, B fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447063/files/figure.png" pageId="2" pageNumber="113">Figs. 5D–E</figureCitation>
): no ectal gland, ectal ducts very long, about 650–840 µm and 32–35 µm wide, canal about 8 µm wide 
<emphasis id="E1C04B6CFFA4950BC28DF8F2FDA8F8A7" box="[483,559,1796,1819]" italics="true" pageId="2" pageNumber="113">in vivo</emphasis>
(520–640 µm long, 25–30 µm wide, canal 5 µm, fixed), not widening entally, projecting into ampulla, ental bulbs wide, about 70–90 µm. Ampullae surrounded distally by 4–6 stalked diverticula of various size: diameter of diverticula 45–90 µm, stalks about as wide and long as diverticula, with ciliated subchamber, ampullae 100–130 µm wide and not considerably set off from distal part, separate openings into oesophagus. 1–3 mature egg at a time.
</paragraph>
</subSubSection>
<subSubSection id="9BAEC4F5FFA4950BC3A9F840FDB6F84E" pageId="2" pageNumber="113" type="distribution">
<paragraph id="D30B977EFFA4950BC3A9F840FDB6F84E" blockId="2.[151,1437,151,2035]" pageId="2" pageNumber="113">
<emphasis id="E1C04B6CFFA4950BC3A9F840FE6BF873" bold="true" box="[199,492,1974,1999]" pageId="2" pageNumber="113">Distribution and habitat</emphasis>
. In Kȏszeg Mts., in 
<emphasis id="E1C04B6CFFA4950BC1B4F840FCCBF872" box="[730,844,1974,1998]" italics="true" pageId="2" pageNumber="113">mesophile</emphasis>
montane hay meadow, and in Rax Mts., in a mixed forest (
<taxonomicName id="14B4ECFDFFA4950BC389F82DFEEEF84E" box="[231,361,2010,2034]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="113" phylum="Tracheophyta" rank="species" species="abies">
<emphasis id="E1C04B6CFFA4950BC389F82DFEEEF84E" box="[231,361,2010,2034]" italics="true" pageId="2" pageNumber="113">Picea abies</emphasis>
</taxonomicName>
, 
<taxonomicName id="14B4ECFDFFA4950BC219F82CFDA6F84E" authorityName="Roth" authorityYear="1788" box="[375,545,2010,2034]" class="Magnoliopsida" family="Betulaceae" genus="Betula" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="113" phylum="Tracheophyta" rank="species" species="pendula">
<emphasis id="E1C04B6CFFA4950BC219F82CFDA6F84E" box="[375,545,2010,2034]" italics="true" pageId="2" pageNumber="113">Betula pendula</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="87CBC7F6FFA5950AC3F9F8FCFF38F864" httpUri="https://zenodo.org/record/1447055/files/figure.png" pageId="3" pageNumber="114" startId="3.[151,250,1802,1824]" targetBox="[158,1372,193,1745]" targetPageId="3">
<paragraph id="D30B977EFFA5950AC3F9F8FCFF38F864" blockId="3.[151,1437,1802,2008]" pageId="3" pageNumber="114">
<emphasis id="E1C04B6CFFA5950AC3F9F8FCFE94F8A3" bold="true" box="[151,275,1802,1824]" pageId="3" pageNumber="114">FIGURE 1.</emphasis>
Micrographs of 
<taxonomicName id="14B4ECFDFFA5950AC2D2F8FCFD43F89C" authority="Nagy &amp; Felföldi &amp; Dózsa-Farkas, 2018" authorityName="Nagy &amp; Felföldi &amp; Dózsa-Farkas" authorityYear="2018" box="[444,708,1802,1824]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="114" phylum="Magnoliopsida" rank="species" species="ventrochaetosa" status="sp. nov.">
<emphasis id="E1C04B6CFFA5950AC2D2F8FCFD43F89C" box="[444,708,1802,1824]" italics="true" pageId="3" pageNumber="114">Fridericia ventrochaetosa</emphasis>
</taxonomicName>
<emphasis id="E1C04B6CFFA5950AC1A4F8FCFCA5F8A3" bold="true" box="[714,802,1802,1823]" pageId="3" pageNumber="114">
<taxonomicNameLabel id="FAF3F617FFA5950AC1A4F8FCFC84F8A3" box="[714,771,1802,1823]" pageId="3" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
A.
</emphasis>
Brain (DNA 1109). 
<emphasis id="E1C04B6CFFA5950AC09DF8FCFB8DF8A3" bold="true" box="[1011,1034,1802,1823]" pageId="3" pageNumber="114">B.</emphasis>
Brain (holotype). 
<emphasis id="E1C04B6CFFA5950AC7ABF8FCFB5AF89C" bold="true" box="[1221,1245,1802,1824]" pageId="3" pageNumber="114">C.</emphasis>
Epidermal glands. 
<emphasis id="E1C04B6CFFA5950AC3F9F8DFFF2EF882" bold="true" box="[151,169,1833,1854]" pageId="3" pageNumber="114">D</emphasis>
. Thick body wall and cuticle (marked with arrow). 
<emphasis id="E1C04B6CFFA5950AC1B8F8DFFD6AF882" bold="true" box="[726,749,1833,1854]" pageId="3" pageNumber="114">E.</emphasis>
Strong septa, 5/6– 9/10 (holotype). 
<emphasis id="E1C04B6CFFA5950AC71FF8DFFB26F882" bold="true" box="[1137,1185,1832,1854]" pageId="3" pageNumber="114">F–G.</emphasis>
Head region (the thick epithelium of prostomium marked with arrow). 
<emphasis id="E1C04B6CFFA5950AC1EFF8B1FD1CF8E0" bold="true" box="[641,667,1863,1884]" pageId="3" pageNumber="114">H.</emphasis>
Clitellar glands, lateral view. 
<emphasis id="E1C04B6CFFA5950AC0A1F8B1FC5EF8E0" bold="true" box="[975,985,1863,1884]" pageId="3" pageNumber="114">I</emphasis>
. Clitellum of a dissected species (above the dorsal gap without glands, below the clitellar glands in transverse rows). 
<emphasis id="E1C04B6CFFA5950AC0C5F890FC59F8C7" bold="true" box="[939,990,1894,1915]" pageId="3" pageNumber="114">J–K.</emphasis>
Coelomocytes (J, paratype P.112.1.2, No. 2219=DNA 978). 
<emphasis id="E1C04B6CFFA5950AC238F873FEEAF826" bold="true" box="[342,365,1925,1946]" pageId="3" pageNumber="114">L.</emphasis>
Nephridium of a juvenile specimen, in the middle part of body; 
<emphasis id="E1C04B6CFFA5950AC770F873FBBAF826" bold="true" box="[1054,1085,1925,1946]" pageId="3" pageNumber="114">M.</emphasis>
Preclitellar nephridium (paratype P.112.1.2, No. 2219=DNA 978). 
<emphasis id="E1C04B6CFFA5950AC287F855FD1DF804" bold="true" box="[489,666,1955,1977]" pageId="3" pageNumber="114">A, C, H, J, L–M</emphasis>
<emphasis id="E1C04B6CFFA5950AC1CFF852FD61F805" box="[673,742,1956,1977]" italics="true" pageId="3" pageNumber="114">in vivo</emphasis>
, 
<emphasis id="E1C04B6CFFA5950AC19AF855FCA4F804" bold="true" box="[756,803,1955,1976]" pageId="3" pageNumber="114">B, E</emphasis>
fixed, not stained, 
<emphasis id="E1C04B6CFFA5950AC087F855FBF1F804" bold="true" box="[1001,1142,1955,1977]" pageId="3" pageNumber="114">D, F–G, I, K.</emphasis>
fixed, stained. Scale bars 50 µm.
</paragraph>
</caption>
<caption id="87CBC7F6FFA2950DC3F9F8FCFECAF864" httpUri="https://zenodo.org/record/1447057/files/figure.png" pageId="4" pageNumber="115" startId="4.[151,250,1802,1824]" targetBox="[158,1376,193,1743]" targetPageId="4">
<paragraph id="D30B977EFFA2950DC3F9F8FCFECAF864" blockId="4.[151,1437,1802,2008]" pageId="4" pageNumber="115">
<emphasis id="E1C04B6CFFA2950DC3F9F8FCFE90F8A3" bold="true" box="[151,279,1802,1824]" pageId="4" pageNumber="115">FIGURE 2.</emphasis>
Micrographs of 
<taxonomicName id="14B4ECFDFFA2950DC2BEF8FCFD5AF89C" authority="Nagy &amp; Felföldi &amp; Dózsa-Farkas, 2018" authorityName="Nagy &amp; Felföldi &amp; Dózsa-Farkas" authorityYear="2018" box="[464,733,1802,1824]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="115" phylum="Magnoliopsida" rank="species" species="ventrochaetosa" status="sp. nov.">
<emphasis id="E1C04B6CFFA2950DC2BEF8FCFD5AF89C" box="[464,733,1802,1824]" italics="true" pageId="4" pageNumber="115">Fridericia ventrochaetosa</emphasis>
</taxonomicName>
<emphasis id="E1C04B6CFFA2950DC186F8FCFCCDF8A3" bold="true" box="[744,842,1802,1823]" pageId="4" pageNumber="115">
<taxonomicNameLabel id="FAF3F617FFA2950DC186F8FCFCA1F8A3" box="[744,806,1802,1823]" pageId="4" pageNumber="115" rank="species">sp. n.</taxonomicNameLabel>
A.
</emphasis>
End of oesophageal appendage with multiple branches (arrows). 
<emphasis id="E1C04B6CFFA2950DC397F8DFFE97F882" bold="true" box="[249,272,1833,1854]" pageId="4" pageNumber="115">B.</emphasis>
Male copulatory organs (arrows); clitellar gland cells visible laterally and ventrally posterior to copulatory organs. 
<emphasis id="E1C04B6CFFA2950DC3F9F8B1FF28F8E1" bold="true" box="[151,175,1863,1885]" pageId="4" pageNumber="115">C.</emphasis>
Midgut pars tumida (arrows) in XXXI–XXXIII (holotype). 
<emphasis id="E1C04B6CFFA2950DC07CF8B1FCADF8E0" bold="true" box="[786,810,1863,1884]" pageId="4" pageNumber="115">D.</emphasis>
Chylus cells in XIII–XIV. 
<emphasis id="E1C04B6CFFA2950DC752F8B1FBD5F8E0" bold="true" box="[1084,1106,1863,1884]" pageId="4" pageNumber="115">E.</emphasis>
Male copulatory organs, ventral view, holotype. Strong bursal muscle marked with arrow. Asterisk: region between the two organs without clitellar glands. 
<emphasis id="E1C04B6CFFA2950DC6EEF890FF2CF826" bold="true" pageId="4" pageNumber="115">F– G.</emphasis>
Bursal slits (marked with arrow in G). 
<emphasis id="E1C04B6CFFA2950DC157F873FDD4F826" bold="true" box="[569,595,1925,1946]" pageId="4" pageNumber="115">H.</emphasis>
Male copulatory organ, lateral view (paratype P.112.1.2, No. 2219=DNA 978). 
<emphasis id="E1C04B6CFFA2950DC617F873FA04F826" bold="true" box="[1401,1411,1925,1946]" pageId="4" pageNumber="115">I</emphasis>
, 
<emphasis id="E1C04B6CFFA2950DC6E7F873FA1BF826" bold="true" box="[1417,1436,1925,1946]" pageId="4" pageNumber="115">J.</emphasis>
Sperm funnels (paratype P.112.1.2, No. 2219=DNA 978). 
<emphasis id="E1C04B6CFFA2950DC187F855FD7CF804" bold="true" box="[745,763,1955,1976]" pageId="4" pageNumber="115">A</emphasis>
, 
<emphasis id="E1C04B6CFFA2950DC066F855FC92F804" bold="true" box="[776,789,1955,1976]" pageId="4" pageNumber="115">F</emphasis>
, 
<emphasis id="E1C04B6CFFA2950DC04DF855FCB3F805" bold="true" box="[803,820,1955,1977]" pageId="4" pageNumber="115">G</emphasis>
, 
<emphasis id="E1C04B6CFFA2950DC02FF855FCCBF804" bold="true" box="[833,844,1955,1976]" pageId="4" pageNumber="115">I</emphasis>
<emphasis id="E1C04B6CFFA2950DC03CF852FC10F805" box="[850,919,1956,1977]" italics="true" pageId="4" pageNumber="115">in vivo</emphasis>
, 
<emphasis id="E1C04B6CFFA2950DC0CAF855FC32F804" bold="true" box="[932,949,1955,1976]" pageId="4" pageNumber="115">B</emphasis>
, 
<emphasis id="E1C04B6CFFA2950DC0ADF855FC52F804" bold="true" box="[963,981,1955,1976]" pageId="4" pageNumber="115">D</emphasis>
, 
<emphasis id="E1C04B6CFFA2950DC08CF855FC71F804" bold="true" box="[994,1014,1955,1976]" pageId="4" pageNumber="115">H</emphasis>
, 
<emphasis id="E1C04B6CFFA2950DC76DF855FB97F804" bold="true" box="[1027,1040,1955,1976]" pageId="4" pageNumber="115">J</emphasis>
fixed, stained, 
<emphasis id="E1C04B6CFFA2950DC7C3F855FB38F805" bold="true" box="[1197,1215,1955,1977]" pageId="4" pageNumber="115">C</emphasis>
, 
<emphasis id="E1C04B6CFFA2950DC7A3F855FB58F804" bold="true" box="[1229,1247,1955,1976]" pageId="4" pageNumber="115">E</emphasis>
fixed, not stained. Scale bars 50 µm.
</paragraph>
</caption>
<caption id="87CBC7F6FFA3950CC3F9F9A4FECEF95F" httpUri="https://zenodo.org/record/1447059/files/figure.png" pageId="5" pageNumber="116" startId="5.[151,250,1618,1640]" targetBox="[178,1398,288,1505]" targetPageId="5">
<paragraph id="D30B977EFFA3950CC3F9F9A4FECEF95F" blockId="5.[151,1436,1618,1763]" pageId="5" pageNumber="116">
<emphasis id="E1C04B6CFFA3950CC3F9F9A4FE8BF9DB" bold="true" box="[151,268,1618,1640]" pageId="5" pageNumber="116">FIGURE 3</emphasis>
. Micrograph of 
<taxonomicName id="14B4ECFDFFA3950CC2DFF9A4FD3EF9D4" authority="Nagy &amp; Felföldi &amp; Dózsa-Farkas, 2018" authorityName="Nagy &amp; Felföldi &amp; Dózsa-Farkas" authorityYear="2018" box="[433,697,1618,1640]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="116" phylum="Magnoliopsida" rank="species" species="ventrochaetosa" status="sp. nov.">
<emphasis id="E1C04B6CFFA3950CC2DFF9A4FD3EF9D4" box="[433,697,1618,1640]" italics="true" pageId="5" pageNumber="116">Fridericia ventrochaetosa</emphasis>
</taxonomicName>
<emphasis id="E1C04B6CFFA3950CC1D0F9A5FC91F9DB" bold="true" box="[702,790,1618,1640]" pageId="5" pageNumber="116">
<taxonomicNameLabel id="FAF3F617FFA3950CC1D0F9A5FD70F9D4" box="[702,759,1619,1640]" pageId="5" pageNumber="116" rank="species">sp. n.</taxonomicNameLabel>
A.
</emphasis>
Sperm funnels (paratype P.112.1.2, No. 2219=DNA 978). 
<emphasis id="E1C04B6CFFA3950CC608F9A5FAF0F9D4" bold="true" box="[1382,1399,1619,1640]" pageId="5" pageNumber="116">B</emphasis>
, 
<emphasis id="E1C04B6CFFA3950CC6EDF9A5FA1BF9D4" bold="true" box="[1411,1436,1619,1640]" pageId="5" pageNumber="116">D.</emphasis>
Spermathecae (paratype P.112.2.6, No. 2345=DNA 1116). 
<emphasis id="E1C04B6CFFA3950CC198F987FC88F93B" bold="true" box="[758,783,1649,1671]" pageId="5" pageNumber="116">C.</emphasis>
Spermathecae (holotype). 
<emphasis id="E1C04B6CFFA3950CC748F987FBBAF93A" bold="true" box="[1062,1085,1649,1670]" pageId="5" pageNumber="116">E.</emphasis>
Spermatheca (paratype P.112.1.2, No. 2219=DNA 978). 
<emphasis id="E1C04B6CFFA3950CC214F966FE0CF919" bold="true" box="[378,395,1680,1701]" pageId="5" pageNumber="116">F.</emphasis>
Dissected spermatheca (paratype P.112.2.1, No. 2266). 
<emphasis id="E1C04B6CFFA3950CC0AEF966FC52F91A" bold="true" box="[960,981,1680,1702]" pageId="5" pageNumber="116">G.</emphasis>
Epidermal gland cells around the opening of spermatheca (opening marked with arrow). 
<emphasis id="E1C04B6CFFA3950CC13CF958FD05F978" bold="true" box="[594,642,1710,1732]" pageId="5" pageNumber="116">A, B</emphasis>
, 
<emphasis id="E1C04B6CFFA3950CC1E0F959FD27F978" bold="true" box="[654,672,1711,1732]" pageId="5" pageNumber="116">D</emphasis>
, 
<emphasis id="E1C04B6CFFA3950CC1C3F958FD46F978" bold="true" box="[685,705,1710,1732]" pageId="5" pageNumber="116">G</emphasis>
<emphasis id="E1C04B6CFFA3950CC1A9F946FC8CF979" box="[711,779,1712,1733]" italics="true" pageId="5" pageNumber="116">in vivo</emphasis>
, 
<emphasis id="E1C04B6CFFA3950CC076F959FCAEF978" bold="true" box="[792,809,1711,1732]" pageId="5" pageNumber="116">E</emphasis>
, 
<emphasis id="E1C04B6CFFA3950CC05BF959FCC3F978" bold="true" box="[821,836,1711,1732]" pageId="5" pageNumber="116">F</emphasis>
fixed, stained, 
<emphasis id="E1C04B6CFFA3950CC08EF958FC75F978" bold="true" box="[992,1010,1710,1732]" pageId="5" pageNumber="116">C</emphasis>
fixed, not stained. Scale bars in 
<emphasis id="E1C04B6CFFA3950CC652F958FAC9F97F" bold="true" box="[1340,1358,1710,1731]" pageId="5" pageNumber="116">A</emphasis>
20 µm, in the rest 50 µm.
</paragraph>
</caption>
<subSubSection id="9BAEC4F5FFA3950EC3A9F8F9FCBFFC7B" lastPageId="7" lastPageNumber="118" pageId="5" pageNumber="116" type="diagnosis">
<paragraph id="D30B977EFFA3950CC3A9F8F9FAB8F860" blockId="5.[151,1437,1807,2013]" pageId="5" pageNumber="116">
<emphasis id="E1C04B6CFFA3950CC3A9F8F9FE57F894" bold="true" box="[199,464,1807,1832]" pageId="5" pageNumber="116">Differential diagnosis.</emphasis>
There are only two species of 
<taxonomicName id="14B4ECFDFFA3950CC059F8E6FC2DF894" box="[823,938,1808,1832]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="116" phylum="Magnoliopsida" rank="genus">
<emphasis id="E1C04B6CFFA3950CC059F8E6FC2DF894" box="[823,938,1808,1832]" italics="true" pageId="5" pageNumber="116">Fridericia</emphasis>
</taxonomicName>
, 
<taxonomicName id="14B4ECFDFFA3950CC0D5F8E7FAA9F894" authority="Xie et al. 2000" authorityName="Xie et al." authorityYear="2000" box="[955,1326,1807,1832]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="116" phylum="Magnoliopsida" rank="species" species="paraunisetosa">
<emphasis id="E1C04B6CFFA3950CC0D5F8E7FBFFF89B" box="[955,1144,1808,1832]" italics="true" pageId="5" pageNumber="116">F. paraunisetosa</emphasis>
<bibRefCitation id="B725EA8FFFA3950CC7EEF8F9FAA9F894" box="[1152,1326,1807,1832]" pageId="5" pageNumber="116" refId="ref5797" refString="Xie, Z., Liang, Y. &amp; Wang, H. (2000) Two new species of Fridericia (Enchytraeidae, Oligochaeta) from Changbaishan Mountain, Jili Province, China. Species Diversity, 5, 53 - 58. https: // doi. org / 10.12782 / specdiv. 5.53" type="journal article">
Xie 
<emphasis id="E1C04B6CFFA3950CC7DCF8E7FB6AF894" box="[1202,1261,1808,1832]" italics="true" pageId="5" pageNumber="116">et al.</emphasis>
2000
</bibRefCitation>
</taxonomicName>
from NE 
<collectingCountry id="ABA3D7EEFFA3950CC3F9F8C2FF5DF8F1" box="[151,218,1844,1869]" name="China" pageId="5" pageNumber="116">China</collectingCountry>
and the new species, with multiple spermathecal diverticula plus laterally completely absent chaetae. 
<taxonomicName id="14B4ECFDFFA3950CC6E9F8C3FEB1F8D3" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="116" phylum="Magnoliopsida" rank="species" species="paraunisetosa">
<emphasis id="E1C04B6CFFA3950CC6E9F8C3FEB1F8D3" italics="true" pageId="5" pageNumber="116">F. paraunisetosa</emphasis>
</taxonomicName>
can easily be distinguished from 
<taxonomicName id="14B4ECFDFFA3950CC1C1F8AFFCF2F8CC" authority="Nagy &amp; Felföldi &amp; Dózsa-Farkas, 2018" authorityName="Nagy &amp; Felföldi &amp; Dózsa-Farkas" authorityYear="2018" box="[687,885,1880,1904]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="116" phylum="Magnoliopsida" rank="species" species="ventrochaetosa" status="sp. nov.">
<emphasis id="E1C04B6CFFA3950CC1C1F8AFFCF2F8CC" box="[687,885,1880,1904]" italics="true" pageId="5" pageNumber="116">F. ventrochaetosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAF3F617FFA3950CC015F8A1FC3CF8D3" box="[891,955,1879,1904]" pageId="5" pageNumber="116" rank="species">
<emphasis id="E1C04B6CFFA3950CC015F8A1FC34F8CC" bold="true" box="[891,947,1879,1904]" pageId="5" pageNumber="116">sp. n</emphasis>
.
</taxonomicNameLabel>
based on the following characters: smaller size (5.0– 
<quantity id="144C3A9BFFA3950CC26BF88AFEDAF828" box="[261,349,1916,1941]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="5" pageNumber="116" unit="mm" value="7.8">7.8 mm</quantity>
<emphasis id="E1C04B6CFFA3950CC20AF88BFE04F828" box="[356,387,1917,1940]" italics="true" pageId="5" pageNumber="116">vs.</emphasis>
<quantity id="144C3A9BFFA3950CC2E5F88AFDB6F828" box="[395,561,1916,1941]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.85" metricValueMin="1.15" pageId="5" pageNumber="116" unit="mm" value="15.0" valueMax="18.5" valueMin="11.5">11.5–18.5 mm</quantity>
, fixed), in ventral bundles only one chaeta per bundle (
<emphasis id="E1C04B6CFFA3950CC7C7F88BFB47F828" box="[1193,1216,1917,1940]" italics="true" pageId="5" pageNumber="116">vs</emphasis>
. 2–4), dorsal pores only from XVIII (
<emphasis id="E1C04B6CFFA3950CC217F857FE17F804" box="[377,400,1953,1976]" italics="true" pageId="5" pageNumber="116">vs</emphasis>
. from VII), brain incised anteriorly (
<emphasis id="E1C04B6CFFA3950CC032F857FCF4F804" box="[860,883,1953,1976]" italics="true" pageId="5" pageNumber="116">vs</emphasis>
. concave), oesophageal appendages stout and unbranched (
<emphasis id="E1C04B6CFFA3950CC244F833FEC6F860" box="[298,321,1989,2012]" italics="true" pageId="5" pageNumber="116">vs</emphasis>
. many branches) and spermatheca with sessile diverticula (
<emphasis id="E1C04B6CFFA3950CC0B5F833FC75F860" box="[987,1010,1989,2012]" italics="true" pageId="5" pageNumber="116">vs</emphasis>
. with stalk) (
<bibRefCitation id="B725EA8FFFA3950CC7E8F832FAB4F860" box="[1158,1331,1988,2013]" pageId="5" pageNumber="116" refId="ref5797" refString="Xie, Z., Liang, Y. &amp; Wang, H. (2000) Two new species of Fridericia (Enchytraeidae, Oligochaeta) from Changbaishan Mountain, Jili Province, China. Species Diversity, 5, 53 - 58. https: // doi. org / 10.12782 / specdiv. 5.53" type="journal article">
Xie 
<emphasis id="E1C04B6CFFA3950CC7D9F833FB77F860" box="[1207,1264,1988,2012]" italics="true" pageId="5" pageNumber="116">et al.</emphasis>
2000
</bibRefCitation>
).
</paragraph>
<caption id="87CBC7F6FFA0950FC3F9FA8FFC49F994" httpUri="https://zenodo.org/record/1447061/files/figure.png" pageId="6" pageNumber="117" startId="6.[151,250,1401,1423]" targetBox="[174,1397,193,1360]" targetPageId="6">
<paragraph id="D30B977EFFA0950FC3F9FA8FFC49F994" blockId="6.[151,1436,1401,1576]" pageId="6" pageNumber="117">
<emphasis id="E1C04B6CFFA0950FC3F9FA8FFE94FA32" bold="true" box="[151,275,1401,1423]" pageId="6" pageNumber="117">FIGURE 4.</emphasis>
Micrographs of 
<taxonomicName id="14B4ECFDFFA0950FC2D2FA8FFCBFFA33" authority="(Hoffmeister, 1843)" authorityName="Hoffmeister" authorityYear="1843" baseAuthorityName="Hoffmeister" baseAuthorityYear="1843" box="[444,824,1401,1423]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="117" phylum="Magnoliopsida" rank="species" species="galba">
<emphasis id="E1C04B6CFFA0950FC2D2FA8FFDE2FA33" box="[444,613,1401,1423]" italics="true" pageId="6" pageNumber="117">Fridericia galba</emphasis>
(Hoffmeister, 1843)
</taxonomicName>
. 
<emphasis id="E1C04B6CFFA0950FC02CFA8FFCD3FA32" bold="true" box="[834,852,1401,1422]" pageId="6" pageNumber="117">A</emphasis>
, 
<emphasis id="E1C04B6CFFA0950FC00FFA8FFCFFFA32" bold="true" box="[865,888,1401,1422]" pageId="6" pageNumber="117">B.</emphasis>
Brain. 
<emphasis id="E1C04B6CFFA0950FC0AAFA8FFC5BFA33" bold="true" box="[964,988,1401,1423]" pageId="6" pageNumber="117">C.</emphasis>
Brown epidermal glands, dorsal view; head pore marked with arrow. 
<emphasis id="E1C04B6CFFA0950FC2F8FA6EFE29FA11" bold="true" box="[406,430,1432,1453]" pageId="6" pageNumber="117">D.</emphasis>
Thick body wall. Thin cuticle marked with arrow. 
<emphasis id="E1C04B6CFFA0950FC0DAFA6EFC4CFA11" bold="true" box="[948,971,1432,1453]" pageId="6" pageNumber="117">E.</emphasis>
Chaetae and brown epidermal glands, ventral view. 
<emphasis id="E1C04B6CFFA0950FC3BAFA41FF62FA70" bold="true" box="[212,229,1463,1484]" pageId="6" pageNumber="117">F.</emphasis>
Head region; thin epithelium of prostomium marked with arrow. 
<emphasis id="E1C04B6CFFA0950FC0E8FA40FC1DFA70" bold="true" box="[902,922,1462,1484]" pageId="6" pageNumber="117">G.</emphasis>
Clitellar glands, dorso-lateral view. 
<emphasis id="E1C04B6CFFA0950FC67FFA41FAABFA70" bold="true" box="[1297,1324,1463,1484]" pageId="6" pageNumber="117">H.</emphasis>
Accessory sexual gland in V. 
<emphasis id="E1C04B6CFFA0950FC23FFA23FEE6FA56" bold="true" box="[337,353,1493,1514]" pageId="6" pageNumber="117">I.</emphasis>
Accessory sexual gland in XI (marked with black arrow, male copulatory organ marked with white arrow), lateral view. 
<emphasis id="E1C04B6CFFA0950FC274FA02FEAAF9B5" bold="true" box="[282,301,1524,1545]" pageId="6" pageNumber="117">J.</emphasis>
Accessory sexual gland in XIII (marked with black arrow, male copulatory organs marked with white arrows), ventral view. 
<emphasis id="E1C04B6CFFA0950FC24EF9E4FEB5F99B" bold="true" box="[288,306,1554,1575]" pageId="6" pageNumber="117">A</emphasis>
, 
<emphasis id="E1C04B6CFFA0950FC250F9E4FED7F994" bold="true" box="[318,336,1554,1576]" pageId="6" pageNumber="117">C</emphasis>
, 
<emphasis id="E1C04B6CFFA0950FC233F9E5FEE9F994" bold="true" box="[349,366,1555,1576]" pageId="6" pageNumber="117">E</emphasis>
, 
<emphasis id="E1C04B6CFFA0950FC215F9E4FE0BF994" bold="true" box="[379,396,1554,1576]" pageId="6" pageNumber="117">G</emphasis>
, 
<emphasis id="E1C04B6CFFA0950FC2F6F9E5FE25F994" bold="true" box="[408,418,1555,1576]" pageId="6" pageNumber="117">I</emphasis>
<emphasis id="E1C04B6CFFA0950FC2C6F9E5FE6BF994" box="[424,492,1555,1576]" italics="true" pageId="6" pageNumber="117">in vivo</emphasis>
; 
<emphasis id="E1C04B6CFFA0950FC294F9E5FD8CF994" bold="true" box="[506,523,1555,1576]" pageId="6" pageNumber="117">B</emphasis>
, 
<emphasis id="E1C04B6CFFA0950FC176F9E5FDADF994" bold="true" box="[536,554,1555,1576]" pageId="6" pageNumber="117">D</emphasis>
, 
<emphasis id="E1C04B6CFFA0950FC158F9E5FDC3F994" bold="true" box="[566,580,1555,1576]" pageId="6" pageNumber="117">F</emphasis>
, 
<emphasis id="E1C04B6CFFA0950FC13FF9E5FDE2F994" bold="true" box="[593,613,1555,1576]" pageId="6" pageNumber="117">H</emphasis>
, 
<emphasis id="E1C04B6CFFA0950FC11FF9E5FDF9F994" bold="true" box="[625,638,1555,1576]" pageId="6" pageNumber="117">J</emphasis>
fixed, stained. Scale bars 50 µm.
</paragraph>
</caption>
<paragraph id="D30B977EFFA0950EC3A9F9A0FCBFFC7B" blockId="6.[151,1437,1621,2006]" lastBlockId="7.[151,1437,151,968]" lastPageId="7" lastPageNumber="118" pageId="6" pageNumber="117">
Apart from the absence of the lateral chaetae, the new species is highly similar morphologically to 
<emphasis id="E1C04B6CFFA0950FC647F9A3FF59F92D" italics="true" pageId="6" pageNumber="117">
<taxonomicName id="14B4ECFDFFA0950FC647F9A3FF5CF92D" baseAuthorityName="Hoffmeister" baseAuthorityYear="1843" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="117" phylum="Magnoliopsida" rank="species" species="galba">Fridericia galba</taxonomicName>
.
</emphasis>
To compare the two species, we studied specimens of 
<taxonomicName id="14B4ECFDFFA0950FC051F98CFC10F92D" baseAuthorityName="Hoffmeister" baseAuthorityYear="1843" box="[831,919,1657,1681]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="117" phylum="Magnoliopsida" rank="species" species="galba">
<emphasis id="E1C04B6CFFA0950FC051F98CFC10F92D" box="[831,919,1657,1681]" italics="true" pageId="6" pageNumber="117">F. galba</emphasis>
</taxonomicName>
sampled from the same two mountains where the new species occurred, and some additional specimens from other sites in 
<collectingCountry id="ABA3D7EEFFA0950FC765F968FBE8F90A" box="[1035,1135,1694,1718]" name="Hungary" pageId="6" pageNumber="117">Hungary</collectingCountry>
. All these specimens of 
<taxonomicName id="14B4ECFDFFA0950FC6E9F968FF50F965" baseAuthorityName="Hoffmeister" baseAuthorityYear="1843" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="117" phylum="Magnoliopsida" rank="species" species="galba">
<emphasis id="E1C04B6CFFA0950FC6E9F968FF50F965" italics="true" pageId="6" pageNumber="117">F. galba</emphasis>
</taxonomicName>
had spermathecae with more than 5 spermathecal diverticula. The comparison of the most important traits of the two species is given in 
<tableCitation id="9E36A2C5FFA0950FC2BDF913FDAEF942" box="[467,553,1765,1790]" captionStart="TABLE 1" captionStartId="8.[151,239,152,175]" captionText="TABLE 1. Comparison the morphological characters of Fridericia ventrochaetosa sp. n. and Fridericia galba (Hoffmeister, 1843) based on our investigation. Data recorded in vivo are marked with bold, while others were measured in whole-mounted specimens on slides." pageId="6" pageNumber="117">Table 1</tableCitation>
. 
<tableCitation id="9E36A2C5FFA0950FC156F913FD0AF942" box="[568,653,1765,1790]" captionStart="TABLE 1" captionStartId="8.[151,239,152,175]" captionText="TABLE 1. Comparison the morphological characters of Fridericia ventrochaetosa sp. n. and Fridericia galba (Hoffmeister, 1843) based on our investigation. Data recorded in vivo are marked with bold, while others were measured in whole-mounted specimens on slides." pageId="6" pageNumber="117">Table 1</tableCitation>
shows that the two species are broadly the same in size, number of segments, oesophageal appendages, origin of dorsal vessel, sperm funnel (comp. 
<figureCitation id="4B8F8BFBFFA0950FC742F8FFFB1AF89D" box="[1068,1181,1801,1826]" captionStart="FIGURE 2" captionStartId="4.[151,250,1802,1824]" captionTargetBox="[158,1376,193,1743]" captionTargetId="figure@4.[158,1428,193,1780]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Fridericia ventrochaetosa sp. n. A. End of oesophageal appendage with multiple branches (arrows). B. Male copulatory organs (arrows); clitellar gland cells visible laterally and ventrally posterior to copulatory organs. C. Midgut pars tumida (arrows) in XXXI–XXXIII (holotype). D. Chylus cells in XIII–XIV. E. Male copulatory organs, ventral view, holotype. Strong bursal muscle marked with arrow. Asterisk: region between the two organs without clitellar glands. F– G. Bursal slits (marked with arrow in G). H. Male copulatory organ, lateral view (paratype P.112.1.2, No. 2219=DNA 978). I,J. Sperm funnels (paratype P.112.1.2, No. 2219=DNA 978). A, F, G, I in vivo, B, D, H, J fixed, stained, C, E fixed, not stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447057/files/figure.png" pageId="6" pageNumber="117">
Figs. 
<date id="A70AB1BEFFA0950FC705F8FFFB05F89D" box="[1131,1154,1801,1825]" pageId="6" pageNumber="117">2I</date>
–J
</figureCitation>
, 
<figureCitation id="4B8F8BFBFFA0950FC7C4F8FFFB4BF89D" box="[1194,1228,1801,1825]" captionStart="FIGURE 3" captionStartId="5.[151,250,1618,1640]" captionTargetBox="[178,1398,288,1505]" captionTargetId="figure@5.[178,1408,288,1524]" captionTargetPageId="5" captionText="FIGURE 3. Micrograph of Fridericia ventrochaetosa sp. n. A. Sperm funnels (paratype P.112.1.2, No. 2219=DNA 978). B, D. Spermathecae (paratype P.112.2.6, No. 2345=DNA 1116). C. Spermathecae (holotype). E. Spermatheca (paratype P.112.1.2, No. 2219=DNA 978). F. Dissected spermatheca (paratype P.112.2.1, No. 2266). G. Epidermal gland cells around the opening of spermatheca (opening marked with arrow). A, B, D, G in vivo, E, F fixed, stained, C fixed, not stained. Scale bars in A 20 µm, in the rest 50 µm." httpUri="https://zenodo.org/record/1447059/files/figure.png" pageId="6" pageNumber="117">3A</figureCitation>
with 
<figureCitation id="4B8F8BFBFFA0950FC663F8FFFAE3F89E" box="[1293,1380,1801,1826]" captionStart="FIGURE 5" captionStartId="7.[151,250,1919,1941]" captionTargetBox="[176,1391,1086,1861]" captionTargetId="figure@7.[176,1410,1086,1877]" captionTargetPageId="7" captionText="FIGURE 5. Micrographs of Fridericia galba. A. Bursal slit (arrow). B. Male copulatory organs, ventral view. C. Sperm funnels. D, E. Spermathecae. A, C–E in vivo, B fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447063/files/figure.png" pageId="6" pageNumber="117">Fig. 5C</figureCitation>
), the male apparatus (comp. 
<figureCitation id="4B8F8BFBFFA0950FC2CEF8DBFD9DF8FA" box="[416,538,1837,1862]" captionStart="FIGURE 2" captionStartId="4.[151,250,1802,1824]" captionTargetBox="[158,1376,193,1743]" captionTargetId="figure@4.[158,1428,193,1780]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Fridericia ventrochaetosa sp. n. A. End of oesophageal appendage with multiple branches (arrows). B. Male copulatory organs (arrows); clitellar gland cells visible laterally and ventrally posterior to copulatory organs. C. Midgut pars tumida (arrows) in XXXI–XXXIII (holotype). D. Chylus cells in XIII–XIV. E. Male copulatory organs, ventral view, holotype. Strong bursal muscle marked with arrow. Asterisk: region between the two organs without clitellar glands. F– G. Bursal slits (marked with arrow in G). H. Male copulatory organ, lateral view (paratype P.112.1.2, No. 2219=DNA 978). I,J. Sperm funnels (paratype P.112.1.2, No. 2219=DNA 978). A, F, G, I in vivo, B, D, H, J fixed, stained, C, E fixed, not stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447057/files/figure.png" pageId="6" pageNumber="117">Figs. 2E,H</figureCitation>
with 
<figureCitation id="4B8F8BFBFFA0950FC132F8DBFD29F8FA" box="[604,686,1837,1862]" captionStart="FIGURE 4" captionStartId="6.[151,250,1401,1423]" captionTargetBox="[174,1397,193,1360]" captionTargetId="figure@6.[174,1412,193,1380]" captionTargetPageId="6" captionText="FIGURE 4. Micrographs of Fridericia galba (Hoffmeister, 1843). A, B. Brain. C. Brown epidermal glands, dorsal view; head pore marked with arrow. D. Thick body wall. Thin cuticle marked with arrow. E. Chaetae and brown epidermal glands, ventral view. F. Head region; thin epithelium of prostomium marked with arrow. G. Clitellar glands, dorso-lateral view. H. Accessory sexual gland in V. I. Accessory sexual gland in XI (marked with black arrow, male copulatory organ marked with white arrow), lateral view. J. Accessory sexual gland in XIII (marked with black arrow, male copulatory organs marked with white arrows), ventral view. A, C, E, G, I in vivo; B, D, F, H, J fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447061/files/figure.png" pageId="6" pageNumber="117">Fig. 4J</figureCitation>
). Spermathecae are also almost the same: ectal duct very long, no ectal gland, more than 2 diverticula with stalks (comp. 
<figureCitation id="4B8F8BFBFFA0950FC194F8A7FCF2F8D5" box="[762,885,1873,1898]" captionStart="FIGURE 3" captionStartId="5.[151,250,1618,1640]" captionTargetBox="[178,1398,288,1505]" captionTargetId="figure@5.[178,1408,288,1524]" captionTargetPageId="5" captionText="FIGURE 3. Micrograph of Fridericia ventrochaetosa sp. n. A. Sperm funnels (paratype P.112.1.2, No. 2219=DNA 978). B, D. Spermathecae (paratype P.112.2.6, No. 2345=DNA 1116). C. Spermathecae (holotype). E. Spermatheca (paratype P.112.1.2, No. 2219=DNA 978). F. Dissected spermatheca (paratype P.112.2.1, No. 2266). G. Epidermal gland cells around the opening of spermatheca (opening marked with arrow). A, B, D, G in vivo, E, F fixed, stained, C fixed, not stained. Scale bars in A 20 µm, in the rest 50 µm." httpUri="https://zenodo.org/record/1447059/files/figure.png" pageId="6" pageNumber="117">Figs. 3B–F</figureCitation>
with 5D–E). The number of diverticula is mostly the same (6), but it varies in the new species between 4 and 6 and in the individuals of 
<taxonomicName id="14B4ECFDFFA0950FC7FAF880FB74F831" baseAuthorityName="Hoffmeister" baseAuthorityYear="1843" box="[1172,1267,1909,1933]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="117" phylum="Magnoliopsida" rank="species" species="galba">
<emphasis id="E1C04B6CFFA0950FC7FAF880FB74F831" box="[1172,1267,1909,1933]" italics="true" pageId="6" pageNumber="117">F. galba</emphasis>
</taxonomicName>
studied by us, between 5–9. The shape of the diverticula and the subchamber and ampullae are almost the same, only the stalks are slightly shorter (
<tableCitation id="9E36A2C5FFA0950FC214F84BFE49F86A" box="[378,462,1981,2006]" captionStart="TABLE 1" captionStartId="8.[151,239,152,175]" captionText="TABLE 1. Comparison the morphological characters of Fridericia ventrochaetosa sp. n. and Fridericia galba (Hoffmeister, 1843) based on our investigation. Data recorded in vivo are marked with bold, while others were measured in whole-mounted specimens on slides." pageId="6" pageNumber="117">Table 1</tableCitation>
). However, differences can also be observed. The main differences are the absence of the lateral chaetae and the lower number of ventral chaetae in the bundles in the case of the new species (maximum of 3–4; in 
<taxonomicName id="14B4ECFDFFA1950EC24DFF4BFE05FF68" baseAuthorityName="Hoffmeister" baseAuthorityYear="1843" box="[291,386,188,212]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="118" phylum="Magnoliopsida" rank="species" species="galba">
<emphasis id="E1C04B6CFFA1950EC24DFF4BFE05FF68" box="[291,386,188,212]" italics="true" pageId="7" pageNumber="118">F. galba</emphasis>
</taxonomicName>
4–7, 
<figureCitation id="4B8F8BFBFFA1950EC2B8FF4AFDB2FF68" box="[470,565,188,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,1401,1423]" captionTargetBox="[174,1397,193,1360]" captionTargetId="figure@6.[174,1412,193,1380]" captionTargetPageId="6" captionText="FIGURE 4. Micrographs of Fridericia galba (Hoffmeister, 1843). A, B. Brain. C. Brown epidermal glands, dorsal view; head pore marked with arrow. D. Thick body wall. Thin cuticle marked with arrow. E. Chaetae and brown epidermal glands, ventral view. F. Head region; thin epithelium of prostomium marked with arrow. G. Clitellar glands, dorso-lateral view. H. Accessory sexual gland in V. I. Accessory sexual gland in XI (marked with black arrow, male copulatory organ marked with white arrow), lateral view. J. Accessory sexual gland in XIII (marked with black arrow, male copulatory organs marked with white arrows), ventral view. A, C, E, G, I in vivo; B, D, F, H, J fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447061/files/figure.png" pageId="7" pageNumber="118">Fig. 4E</figureCitation>
). Furthermore, both species have thick body wall but the cuticle in 
<taxonomicName id="14B4ECFDFFA1950EC6E9FF4BFEC6FF44" authority="Nagy &amp; Felföldi &amp; Dózsa-Farkas, 2018" authorityName="Nagy &amp; Felföldi &amp; Dózsa-Farkas" authorityYear="2018" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="118" phylum="Magnoliopsida" rank="species" species="ventrochaetosa" status="sp. nov.">
<emphasis id="E1C04B6CFFA1950EC6E9FF4BFEC6FF44" italics="true" pageId="7" pageNumber="118">F. ventrochaetosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAF3F617FFA1950EC226FF29FE0FFF44" box="[328,392,223,248]" pageId="7" pageNumber="118" rank="species">
<emphasis id="E1C04B6CFFA1950EC226FF29FE0FFF44" bold="true" box="[328,392,223,248]" pageId="7" pageNumber="118">sp. n.</emphasis>
</taxonomicNameLabel>
is much thicker (8–14 µm; in 
<taxonomicName id="14B4ECFDFFA1950EC1B1FF17FCBCFF44" baseAuthorityName="Hoffmeister" baseAuthorityYear="1843" box="[735,827,224,248]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="118" phylum="Magnoliopsida" rank="species" species="galba">
<emphasis id="E1C04B6CFFA1950EC1B1FF17FCBCFF44" box="[735,827,224,248]" italics="true" pageId="7" pageNumber="118">F. galba</emphasis>
</taxonomicName>
1–1.5 µm) (
<tableCitation id="9E36A2C5FFA1950EC0A8FF29FB9BFF44" box="[966,1052,223,248]" captionStart="TABLE 1" captionStartId="8.[151,239,152,175]" captionText="TABLE 1. Comparison the morphological characters of Fridericia ventrochaetosa sp. n. and Fridericia galba (Hoffmeister, 1843) based on our investigation. Data recorded in vivo are marked with bold, while others were measured in whole-mounted specimens on slides." pageId="7" pageNumber="118">Table 1</tableCitation>
, 
<figureCitation id="4B8F8BFBFFA1950EC744FF29FB0CFF44" box="[1066,1163,223,248]" captionStart="FIGURE 1" captionStartId="3.[151,250,1802,1824]" captionTargetBox="[158,1372,193,1745]" captionTargetId="figure@3.[158,1428,193,1780]" captionTargetPageId="3" captionText="FIGURE 1. Micrographs of Fridericia ventrochaetosa sp. n. A. Brain (DNA 1109). B. Brain (holotype). C. Epidermal glands. D. Thick body wall and cuticle (marked with arrow). E. Strong septa, 5/6– 9/10 (holotype). F–G. Head region (the thick epithelium of prostomium marked with arrow). H. Clitellar glands, lateral view. I. Clitellum of a dissected species (above the dorsal gap without glands, below the clitellar glands in transverse rows). J–K. Coelomocytes (J, paratype P.112.1.2, No. 2219=DNA 978). L. Nephridium of a juvenile specimen, in the middle part of body; M. Preclitellar nephridium (paratype P.112.1.2, No. 2219=DNA 978). A, C, H, J, L–M in vivo, B, E fixed, not stained, D, F–G, I, K. fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447055/files/figure.png" pageId="7" pageNumber="118">Figs. 1D</figureCitation>
<emphasis id="E1C04B6CFFA1950EC7FDFF17FB2DFF44" box="[1171,1194,225,248]" italics="true" pageId="7" pageNumber="118">vs</emphasis>
. 
<figureCitation id="4B8F8BFBFFA1950EC7D7FF29FA93FF44" box="[1209,1300,223,248]" captionStart="FIGURE 4" captionStartId="6.[151,250,1401,1423]" captionTargetBox="[174,1397,193,1360]" captionTargetId="figure@6.[174,1412,193,1380]" captionTargetPageId="6" captionText="FIGURE 4. Micrographs of Fridericia galba (Hoffmeister, 1843). A, B. Brain. C. Brown epidermal glands, dorsal view; head pore marked with arrow. D. Thick body wall. Thin cuticle marked with arrow. E. Chaetae and brown epidermal glands, ventral view. F. Head region; thin epithelium of prostomium marked with arrow. G. Clitellar glands, dorso-lateral view. H. Accessory sexual gland in V. I. Accessory sexual gland in XI (marked with black arrow, male copulatory organ marked with white arrow), lateral view. J. Accessory sexual gland in XIII (marked with black arrow, male copulatory organs marked with white arrows), ventral view. A, C, E, G, I in vivo; B, D, F, H, J fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447061/files/figure.png" pageId="7" pageNumber="118">Fig. 4D</figureCitation>
). Epidermal glands are well-visible, but in 
<taxonomicName id="14B4ECFDFFA1950EC284FEF3FDC6FEA0" baseAuthorityName="Hoffmeister" baseAuthorityYear="1843" box="[490,577,260,284]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="118" phylum="Magnoliopsida" rank="species" species="galba">
<emphasis id="E1C04B6CFFA1950EC284FEF3FDC6FEA0" box="[490,577,260,284]" italics="true" pageId="7" pageNumber="118">F. galba</emphasis>
</taxonomicName>
often occur conspicuous brown glands, too (
<figureCitation id="4B8F8BFBFFA1950EC753FEF2FB32FEA0" box="[1085,1205,260,285]" captionStart="FIGURE 4" captionStartId="6.[151,250,1401,1423]" captionTargetBox="[174,1397,193,1360]" captionTargetId="figure@6.[174,1412,193,1380]" captionTargetPageId="6" captionText="FIGURE 4. Micrographs of Fridericia galba (Hoffmeister, 1843). A, B. Brain. C. Brown epidermal glands, dorsal view; head pore marked with arrow. D. Thick body wall. Thin cuticle marked with arrow. E. Chaetae and brown epidermal glands, ventral view. F. Head region; thin epithelium of prostomium marked with arrow. G. Clitellar glands, dorso-lateral view. H. Accessory sexual gland in V. I. Accessory sexual gland in XI (marked with black arrow, male copulatory organ marked with white arrow), lateral view. J. Accessory sexual gland in XIII (marked with black arrow, male copulatory organs marked with white arrows), ventral view. A, C, E, G, I in vivo; B, D, F, H, J fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447061/files/figure.png" pageId="7" pageNumber="118">Fig. 4C, E</figureCitation>
). The mucocytes are about the same length and the matrix may be filled with refractile vesicles in 
<taxonomicName id="14B4ECFDFFA1950EC765FEDFFBE0FEFC" baseAuthorityName="Hoffmeister" baseAuthorityYear="1843" box="[1035,1127,296,320]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="118" phylum="Magnoliopsida" rank="species" species="galba">
<emphasis id="E1C04B6CFFA1950EC765FEDFFBE0FEFC" box="[1035,1127,296,320]" italics="true" pageId="7" pageNumber="118">F. galba</emphasis>
</taxonomicName>
, but this granulation in the new species is much more prominent (
<figureCitation id="4B8F8BFBFFA1950EC134FEBAFD2DFED8" box="[602,682,332,357]" captionStart="FIGURE 1" captionStartId="3.[151,250,1802,1824]" captionTargetBox="[158,1372,193,1745]" captionTargetId="figure@3.[158,1428,193,1780]" captionTargetPageId="3" captionText="FIGURE 1. Micrographs of Fridericia ventrochaetosa sp. n. A. Brain (DNA 1109). B. Brain (holotype). C. Epidermal glands. D. Thick body wall and cuticle (marked with arrow). E. Strong septa, 5/6– 9/10 (holotype). F–G. Head region (the thick epithelium of prostomium marked with arrow). H. Clitellar glands, lateral view. I. Clitellum of a dissected species (above the dorsal gap without glands, below the clitellar glands in transverse rows). J–K. Coelomocytes (J, paratype P.112.1.2, No. 2219=DNA 978). L. Nephridium of a juvenile specimen, in the middle part of body; M. Preclitellar nephridium (paratype P.112.1.2, No. 2219=DNA 978). A, C, H, J, L–M in vivo, B, E fixed, not stained, D, F–G, I, K. fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447055/files/figure.png" pageId="7" pageNumber="118">Fig. 1J</figureCitation>
) and the lenticytes are also larger (
<tableCitation id="9E36A2C5FFA1950EC727FEBAFB27FED8" box="[1097,1184,332,357]" captionStart="TABLE 1" captionStartId="8.[151,239,152,175]" captionText="TABLE 1. Comparison the morphological characters of Fridericia ventrochaetosa sp. n. and Fridericia galba (Hoffmeister, 1843) based on our investigation. Data recorded in vivo are marked with bold, while others were measured in whole-mounted specimens on slides." pageId="7" pageNumber="118">Table 1</tableCitation>
). Further differences: chylus cells and midgut pars tumida are further back in 
<taxonomicName id="14B4ECFDFFA1950EC06FFE87FCDBFE34" baseAuthorityName="Hoffmeister" baseAuthorityYear="1843" box="[769,860,368,392]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="118" phylum="Magnoliopsida" rank="species" species="galba">
<emphasis id="E1C04B6CFFA1950EC06FFE87FCDBFE34" box="[769,860,368,392]" italics="true" pageId="7" pageNumber="118">F. galba</emphasis>
</taxonomicName>
, and the blood is light pink in 
<taxonomicName id="14B4ECFDFFA1950EC7C1FE87FAF2FE34" authority="Nagy &amp; Felföldi &amp; Dózsa-Farkas, 2018" authorityName="Nagy &amp; Felföldi &amp; Dózsa-Farkas" authorityYear="2018" box="[1199,1397,368,392]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="118" phylum="Magnoliopsida" rank="species" species="ventrochaetosa" status="sp. nov.">
<emphasis id="E1C04B6CFFA1950EC7C1FE87FAF2FE34" box="[1199,1397,368,392]" italics="true" pageId="7" pageNumber="118">F. ventrochaetosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAF3F617FFA1950EC614FE99FF29FE11" pageId="7" pageNumber="118" rank="species">
<emphasis id="E1C04B6CFFA1950EC614FE99FF29FE11" bold="true" pageId="7" pageNumber="118">sp. n.</emphasis>
</taxonomicNameLabel>
but colorless in 
<taxonomicName id="14B4ECFDFFA1950EC21EFE63FE49FE10" baseAuthorityName="Hoffmeister" baseAuthorityYear="1843" box="[368,462,404,428]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="118" phylum="Magnoliopsida" rank="species" species="galba">
<emphasis id="E1C04B6CFFA1950EC21EFE63FE49FE10" box="[368,462,404,428]" italics="true" pageId="7" pageNumber="118">F. galba</emphasis>
</taxonomicName>
(
<tableCitation id="9E36A2C5FFA1950EC2B0FE62FDB3FE10" box="[478,564,404,429]" captionStart="TABLE 1" captionStartId="8.[151,239,152,175]" captionText="TABLE 1. Comparison the morphological characters of Fridericia ventrochaetosa sp. n. and Fridericia galba (Hoffmeister, 1843) based on our investigation. Data recorded in vivo are marked with bold, while others were measured in whole-mounted specimens on slides." pageId="7" pageNumber="118">Table 1</tableCitation>
), and the epithelium of the prostomium of the new species is thicker, more conspicuous (
<figureCitation id="4B8F8BFBFFA1950EC25DFE41FE2AFE6C" box="[307,429,439,464]" captionStart="FIGURE 1" captionStartId="3.[151,250,1802,1824]" captionTargetBox="[158,1372,193,1745]" captionTargetId="figure@3.[158,1428,193,1780]" captionTargetPageId="3" captionText="FIGURE 1. Micrographs of Fridericia ventrochaetosa sp. n. A. Brain (DNA 1109). B. Brain (holotype). C. Epidermal glands. D. Thick body wall and cuticle (marked with arrow). E. Strong septa, 5/6– 9/10 (holotype). F–G. Head region (the thick epithelium of prostomium marked with arrow). H. Clitellar glands, lateral view. I. Clitellum of a dissected species (above the dorsal gap without glands, below the clitellar glands in transverse rows). J–K. Coelomocytes (J, paratype P.112.1.2, No. 2219=DNA 978). L. Nephridium of a juvenile specimen, in the middle part of body; M. Preclitellar nephridium (paratype P.112.1.2, No. 2219=DNA 978). A, C, H, J, L–M in vivo, B, E fixed, not stained, D, F–G, I, K. fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447055/files/figure.png" pageId="7" pageNumber="118">Fig. 1F–G</figureCitation>
), than in 
<taxonomicName id="14B4ECFDFFA1950EC175FE4FFDFEFE6C" baseAuthorityName="Hoffmeister" baseAuthorityYear="1843" box="[539,633,440,464]" class="Magnoliopsida" family="Bignoniaceae" genus="Fridericia" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="118" phylum="Magnoliopsida" rank="species" species="galba">
<emphasis id="E1C04B6CFFA1950EC175FE4FFDFEFE6C" box="[539,633,440,464]" italics="true" pageId="7" pageNumber="118">F. galba</emphasis>
</taxonomicName>
(
<figureCitation id="4B8F8BFBFFA1950EC1E4FE41FD65FE6C" box="[650,738,439,464]" captionStart="FIGURE 4" captionStartId="6.[151,250,1401,1423]" captionTargetBox="[174,1397,193,1360]" captionTargetId="figure@6.[174,1412,193,1380]" captionTargetPageId="6" captionText="FIGURE 4. Micrographs of Fridericia galba (Hoffmeister, 1843). A, B. Brain. C. Brown epidermal glands, dorsal view; head pore marked with arrow. D. Thick body wall. Thin cuticle marked with arrow. E. Chaetae and brown epidermal glands, ventral view. F. Head region; thin epithelium of prostomium marked with arrow. G. Clitellar glands, dorso-lateral view. H. Accessory sexual gland in V. I. Accessory sexual gland in XI (marked with black arrow, male copulatory organ marked with white arrow), lateral view. J. Accessory sexual gland in XIII (marked with black arrow, male copulatory organs marked with white arrows), ventral view. A, C, E, G, I in vivo; B, D, F, H, J fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447061/files/figure.png" pageId="7" pageNumber="118">Fig. 4F</figureCitation>
). Last, but not least, individuals of 
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often have large accessory sexual glands, mostly only one side in V, XI, XIII (
<figureCitation id="4B8F8BFBFFA1950EC03BFE2AFC53FE48" box="[853,980,476,501]" captionStart="FIGURE 4" captionStartId="6.[151,250,1401,1423]" captionTargetBox="[174,1397,193,1360]" captionTargetId="figure@6.[174,1412,193,1380]" captionTargetPageId="6" captionText="FIGURE 4. Micrographs of Fridericia galba (Hoffmeister, 1843). A, B. Brain. C. Brown epidermal glands, dorsal view; head pore marked with arrow. D. Thick body wall. Thin cuticle marked with arrow. E. Chaetae and brown epidermal glands, ventral view. F. Head region; thin epithelium of prostomium marked with arrow. G. Clitellar glands, dorso-lateral view. H. Accessory sexual gland in V. I. Accessory sexual gland in XI (marked with black arrow, male copulatory organ marked with white arrow), lateral view. J. Accessory sexual gland in XIII (marked with black arrow, male copulatory organs marked with white arrows), ventral view. A, C, E, G, I in vivo; B, D, F, H, J fixed, stained. Scale bars 50 µm." httpUri="https://zenodo.org/record/1447061/files/figure.png" pageId="7" pageNumber="118">Figs. 4H–J</figureCitation>
), but these organs are absent in the new species (
<tableCitation id="9E36A2C5FFA1950EC393FE09FED0FDA4" box="[253,343,511,536]" captionStart="TABLE 1" captionStartId="8.[151,239,152,175]" captionText="TABLE 1. Comparison the morphological characters of Fridericia ventrochaetosa sp. n. and Fridericia galba (Hoffmeister, 1843) based on our investigation. Data recorded in vivo are marked with bold, while others were measured in whole-mounted specimens on slides." pageId="7" pageNumber="118">Table 1</tableCitation>
). After surveying references describing 
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<emphasis id="E1C04B6CFFA1950EC053FDF7FC18FDA4" box="[829,927,512,536]" italics="true" pageId="7" pageNumber="118">F. galba</emphasis>
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previously (for complete bibliography see 
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), it could be stated that the features of specimens studied by us fall within the range of variation reported earlier. However, since we provided more morphological characters and measurement data (
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), a more detailed comparison was possible with the new species. It should be noted that in some descriptions, few traits of 
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are more variable as in the case of our specimens; e.g. segment number 40–45, length 
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in 
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, (48)–64–71–(81) and 
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in 
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, 55–61 segments and 
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in 
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or 60–70 segments and 
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length in 
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, respectively. Difference could be found in the number of spermathecal diverticula; e.g. 
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reported 3–5, 
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, 
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, and 
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reported 2–8. In the case of other characters we recorded generally the same as in previous references, although most of the previous descriptions lack measured values. Other authors including us conceive that the presence of accessory sexual glands is a characteristic feature of this species, even if it is absent in some specimens. Despite the above-mentioned variation of morphological characters, the new species could be easily distinguished from 
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.
</paragraph>
</subSubSection>
</treatment>
</document>