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<document id="AABB10AF7A9616879825861868F07CDE" ID-CLB-Dataset="20797" ID-DOI="10.11646/phytotaxa.532.3.2" ID-GBIF-Dataset="3e51d403-8d7a-4533-93fa-d9d8abe9574c" ID-ISSN="1179-3163" ID-Zenodo-Dep="5964614" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643787883339" checkinUser="plazi" docAuthor="Henderson, Andrew &amp; D ựng, Nguy ễn Qu ốc" docDate="2022" docId="58558799FF81FFE4FF0996C38F09744F" docLanguage="en" docName="phytotaxa.532.3.2.pdf" docOrigin="Phytotaxa 532 (3)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Lanonia tenuisecta Henderson &amp; Nguy" docType="treatment" docVersion="7" lastPageNumber="239" masterDocId="A46CFFE1FF9FFFFAFF8195518D347336" masterDocTitle="A revision of Lanonia (Arecaceae, Coryphoideae, Livistoninae)" masterLastPageNumber="245" masterPageNumber="209" pageNumber="239" updateTime="1699305980494" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="5ACA45D5DB8CD4149FE3BB6AF76187C6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0B9D3457F9C94F50DB97545E86BF26B9">
<mods:title id="38F070600FB0776A6C86AE3089E57C2F">A revision of Lanonia (Arecaceae, Coryphoideae, Livistoninae)</mods:title>
</mods:titleInfo>
<mods:name id="265BF6B701F59348DFB25629FEE23A0B" type="personal">
<mods:role id="EC5274EDC3CEFB16950058C03E9BC173">
<mods:roleTerm id="F9B1A14944D413A7078353A5DAFC70B4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="73A4F621DD3BD9E9FAD8FDF4F748E5F7">Henderson, Andrew</mods:namePart>
<mods:affiliation id="93E03D6A6D2A27C76BB147068A9A4CDE">The New York Botanical Garden, Bronx, NY 10458 - 5126, U. S. A.</mods:affiliation>
</mods:name>
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<mods:role id="F6B9AA2DF6B85319BCFA4E74817D88D8">
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</mods:role>
<mods:namePart id="4DC8037E1EECEE8AF70A308836E36D9C">D ựng, Nguy ễn Qu ốc</mods:namePart>
<mods:affiliation id="F86AD43FD7B8ED08555F6B357F331A4E">Forest Inventory and Planning Institute, Thanh Tri, Hanoi, Vietnam</mods:affiliation>
</mods:name>
<mods:typeOfResource id="47A35793A7563A60C4C14064CEDA2517">text</mods:typeOfResource>
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<mods:titleInfo id="A360E7BF62E6DD86B6D9AE58F68F5D16">
<mods:title id="A7743D7F250F31A8266A4EEFFD3667A0">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="432A718CC6E61755ADE8921FBCAACFF3">
<mods:date id="4A7468E6AB39F0ECD072D615CFC50FA4">2022</mods:date>
<mods:detail id="55013D2BF8C428C618D53BAD6EA687F2" type="pubDate">
<mods:number id="33574D65AAA1A862B14D8383CDB56D17">2022-02-02</mods:number>
</mods:detail>
<mods:detail id="6297EE43118FBA20E8A49795C434F24B" type="volume">
<mods:number id="0CBF614F7EA4179A0C7208AD2A2D313A">532</mods:number>
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<mods:detail id="59575405904942175AB0B402E19B16F8" type="issue">
<mods:number id="90E3A8DC2288C5BB2EFD1AE04B0C17C4">3</mods:number>
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<mods:start id="31AD73A728D7703405599D7ADAB6510E">209</mods:start>
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<mods:classification id="3B74A9FC67EFBF57B1CF05301F641D7E">journal article</mods:classification>
<mods:identifier id="42AC39CC20CAEF4D49E09E49EB330E6C" type="CLB-Dataset">20797</mods:identifier>
<mods:identifier id="636046F434DAA5F424F7616F00EF4F1E" type="DOI">10.11646/phytotaxa.532.3.2</mods:identifier>
<mods:identifier id="E2EE431FB32D23E1FFFD7C5B5F240DFF" type="GBIF-Dataset">3e51d403-8d7a-4533-93fa-d9d8abe9574c</mods:identifier>
<mods:identifier id="F272DC6D456A10E0F914B8CB662CA66B" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="B78B64ED97692DF3069355E3B9EDFFD4" type="Zenodo-Dep">5964614</mods:identifier>
</mods:mods>
<treatment id="58558799FF81FFE4FF0996C38F09744F" ID-DOI="http://doi.org/10.5281/zenodo.6301146" ID-GBIF-Taxon="193276896" ID-Zenodo-Dep="6301146" LSID="urn:lsid:plazi:treatment:58558799FF81FFE4FF0996C38F09744F" httpUri="http://treatment.plazi.org/id/58558799FF81FFE4FF0996C38F09744F" lastPageNumber="239" pageId="30" pageNumber="239">
<subSubSection id="98E66504FF81FFE4FF0996C38E4270C2" pageId="30" pageNumber="239" type="nomenclature">
<paragraph id="D043368FFF81FFE4FF0996C38E4270C2" blockId="30.[136,1452,914,1012]" pageId="30" pageNumber="239">
<emphasis id="E288EA9DFF81FFE4FF0996C38EBF709B" bold="true" box="[136,907,914,943]" pageId="30" pageNumber="239">
18.
<taxonomicName id="17FC4D0CFF81FFE4FF3396C38FB47099" authority="Henderson &amp; Nguy" authorityName="Henderson &amp; Nguy" box="[178,640,914,943]" class="Liliopsida" family="Arecaceae" genus="Lanonia" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="239" phylum="Tracheophyta" rank="species" species="tenuisecta">
<emphasis id="E288EA9DFF81FFE4FF3396C38CA5709A" bold="true" box="[178,401,914,940]" italics="true" pageId="30" pageNumber="239">Lanonia tenuisecta</emphasis>
Henderson &amp; Nguy
</taxonomicName>
ễn Quốc Dựng
<emphasis id="E288EA9DFF81FFE4FCB596C28EBF709B" bold="true" box="[820,907,915,941]" italics="true" pageId="30" pageNumber="239">sp. nov.</emphasis>
</emphasis>
<materialsCitation id="60943CD2FF81FFE4FC1396C38E4670C2" ID-GBIF-Occurrence="3462293303" collectingDate="2010-07-11" collectionCode="A, HN, NY" collectorName="Henderson &amp; Bui Van Thanh &amp; Nguyen Thi Bich Ngoc" country="Vietnam" county="Type" elevation="1290" latitude="12.116" location="Hon Ba Nature Reserve" longLatPrecision="77" longitude="108.95" municipality="Dien Khanh district" pageId="30" pageNumber="239" specimenCount="1" stateProvince="Khanh Hoa" typeStatus="holotype">
<collectingCounty id="39224E03FF81FFE4FC1396C38EF8709A" box="[914,972,914,940]" pageId="30" pageNumber="239">Type</collectingCounty>
:—
<collectingCountry id="A8EB761FFF81FFE4FC6C96C38946709A" box="[1005,1138,914,940]" name="Vietnam" pageId="30" pageNumber="239">VIETNAM</collectingCountry>
.
<collectingRegion id="1238F86DFF81FFE4FBFC96C3885E709A" box="[1149,1386,914,941]" country="Vietnam" name="Khanh Hoa" pageId="30" pageNumber="239">Khanh Hoa province</collectingRegion>
,
<collectingMunicipality id="3027ACF5FF81FFE4FAF496C38C1F70E6" pageId="30" pageNumber="239">Dien Khanh district</collectingMunicipality>
,
<location id="D5236054FF81FFE4FEB696E68F7270E7" LSID="urn:lsid:plazi:treatment:58558799FF81FFE4FF0996C38F09744F:D5236054FF81FFE4FEB696E68F7270E7" box="[311,582,951,977]" country="Vietnam" county="Type" latitude="12.116" longLatPrecision="77" longitude="108.95" municipality="Dien Khanh district" name="Hon Ba Nature Reserve" pageId="30" pageNumber="239" stateProvince="Khanh Hoa">Hon Ba Nature Reserve</location>
,
<geoCoordinate id="B5C85048FF81FFE4FDD096E78F8570E6" box="[593,689,950,976]" degrees="12.116" direction="north" orientation="latitude" pageId="30" pageNumber="239" precision="55" value="12.116">12.116N</geoCoordinate>
<geoCoordinate id="B5C85048FF81FFE4FD3896E78E1C70E7" box="[697,808,950,977]" degrees="108.950" direction="east" orientation="longitude" pageId="30" pageNumber="239" precision="55" value="108.95">108.950E</geoCoordinate>
,
<quantity id="17049B6AFF81FFE4FCB596E78EA670E7" box="[820,914,950,977]" metricMagnitude="3" metricUnit="m" metricValue="1.29" pageId="30" pageNumber="239" unit="m" value="1290.0">
<elevation id="5BD1D1BCFF81FFE4FCB596E78EA670E7" box="[820,914,950,977]" metricMagnitude="3" metricUnit="m" metricValue="1.29" pageId="30" pageNumber="239" unit="m" value="1290.0">1,290 m</elevation>
</quantity>
,
<date id="A442104FFF81FFE4FC1F96E7890670E6" box="[926,1074,950,976]" pageId="30" pageNumber="239" value="2010-07-11">
<collectingDate id="B406E9A7FF81FFE4FC1F96E7890670E6" box="[926,1074,950,976]" pageId="30" pageNumber="239" value="2010-07-11">11 July 2010</collectingDate>
</date>
,
<emphasis id="E288EA9DFF81FFE4FBBF96E68CF170C2" italics="true" pageId="30" pageNumber="239">
<collectionCode id="B6EDAE4AFF81FFE4FBBF96E6896670E7" box="[1086,1106,951,977]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="30" pageNumber="239" type="Herbarium">A</collectionCode>
.
<collectorName id="7D095359FF81FFE4FBDC96E789EF70E6" box="[1117,1243,950,976]" pageId="30" pageNumber="239">Henderson</collectorName>
,
<collectorName id="7D095359FF81FFE4FB6696E688BA70E6" box="[1255,1422,950,976]" pageId="30" pageNumber="239">Bui Van Thanh</collectorName>
&amp;
<collectorName id="7D095359FF81FFE4FF09968A8CB270C2" box="[136,390,986,1012]" pageId="30" pageNumber="239">Nguyen Thi Bich Ngoc</collectorName>
3707
</emphasis>
(
<typeStatus id="0F47882DFF81FFE4FE55968B8F0C70C2" box="[468,568,986,1012]" pageId="30" pageNumber="239" type="holotype">holotype</typeStatus>
<collectionCode id="B6EDAE4AFF81FFE4FDBE968B8F5B70C2" box="[575,623,986,1012]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="30" pageNumber="239" type="Herbarium">HN</collectionCode>
!, isotype
<collectionCode id="B6EDAE4AFF81FFE4FD57968B8E3370C2" box="[726,775,986,1012]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="30" pageNumber="239" type="Herbarium">NY</collectionCode>
!).
<figureCitation id="48C72A0AFF81FFE4FC9E968B8E4670C2" box="[799,882,986,1012]" captionStart="FIGURE 18" captionStartId="31.[136,229,1900,1922]" captionTargetBox="[159,1428,193,1873]" captionTargetId="figure-17@31.[158,1430,191,1874]" captionTargetPageId="31" captionText="FIGURE 18. Type of L. tenuisecta (Henderson et al. 3707)." figureDoi="http://doi.org/10.5281/zenodo.5964676" httpUri="https://zenodo.org/record/5964676/files/figure.png" pageId="30" pageNumber="239">Fig. 18</figureCitation>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="98E66504FF81FFE4FF0991738916754A" pageId="30" pageNumber="239" type="description">
<paragraph id="D043368FFF81FFE4FF0991738916754A" blockId="30.[136,1452,1058,1913]" pageId="30" pageNumber="239">
<emphasis id="E288EA9DFF81FFE4FF0991738DE4770A" bold="true" box="[136,208,1058,1084]" pageId="30" pageNumber="239">Stems</emphasis>
1.4(1.0–2.8) m long, 2.2(1.8–2.5) cm diameter, solitary.
<emphasis id="E288EA9DFF81FFE4FCC191738EA6770A" bold="true" box="[832,914,1058,1084]" pageId="30" pageNumber="239">Leaves</emphasis>
12; ligules 32.9(29.5–40.0) cm long, papery, not disintegrating into fibers; sheaths and proximalmost part of petioles with dense, flat, black scales; petioles 42.1(35.0– 60.0) cm long, 0.3(0.3–0.4) cm wide at the apex; petiole thorns usually stout, reddish-brown or dark brown, some bifid or often some in pairs, on proximalmost part of petiole; hastulas flat, rounded, not infolded; leaf blades 53.3(44.0– 70.0) cm wide; costas 2.9(1.8–4.0) cm long, triangular, with a pulvinus at the apex abaxially, with the numerous (rarely few) segments free almost to the base; segments 13(10–15) per leaf, not mottled, with minute, reddish-brown scales abaxially, with (on dried specimens) ruffled margins; middle segments 31.0(24.5–37.5) cm long, 2.2(1.4–3.0) cm wide at the apex; apices of middle segments with adaxial splits much deeper than abaxial ones.
<emphasis id="E288EA9DFF81FFE4FB1D904F8871760E" bold="true" box="[1180,1349,1310,1336]" pageId="30" pageNumber="239">Inflorescences</emphasis>
85.0 cm long; prophylls 26.5(12.0-37.0) cm long; peduncles 57.0(44.5–70.0) cm long; rachis bracts narrowly tubular, splitting apically, not or sparsely tomentose; rachillae filiform, sparsely to moderately covered with short, mostly simple, brownish hairs;
<emphasis id="E288EA9DFF81FFE4FEBB90DB8C9B7692" bold="true" box="[314,431,1418,1444]" pageId="30" pageNumber="239">staminate</emphasis>
rachises absent; staminate partial inflorescences 1, branched to 3 orders; staminate rachillae 34, 6.0 cm long,
<quantity id="17049B6AFF81FFE4FED190FF8C9D76FF" box="[336,425,1454,1481]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="239" unit="mm" value="0.3">0.3 mm</quantity>
diameter; staminate flowers
<quantity id="17049B6AFF81FFE4FD7990FF8E6576FF" box="[760,849,1454,1481]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="30" pageNumber="239" unit="mm" value="1.7">1.7 mm</quantity>
long; sepals not pedicellate at the base, not densely hairy; petals rounded at the apex; stamens with alternately long and short filaments and small, almost square anthers; filaments not elongate; pistillodes undivided;
<emphasis id="E288EA9DFF81FFE4FD0890A78FC47526" bold="true" box="[649,752,1526,1552]" pageId="30" pageNumber="239">pistillate</emphasis>
rachises absent; pistillate partial inflorescences 1, branched to 2 orders; pistillate rachillae on proximalmost partial inflorescence 3(2–4), 13.1(10.0–16.2) cm long, 1.0(0.8–1.1) mm diameter; pistillate flowers
<quantity id="17049B6AFF81FFE4FE41936F8F23756F" box="[448,535,1598,1625]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="30" pageNumber="239" unit="mm" value="2.5">2.5 mm</quantity>
long, with short styles;
<emphasis id="E288EA9DFF81FFE4FCAD936F8E59756E" bold="true" box="[812,877,1598,1624]" pageId="30" pageNumber="239">fruits</emphasis>
7.6(7.4–7.7) mm long, 7.3(7.1–7.4) mm diameter, globose, white, with rough, pebbled surfaces; seeds with a straight basal intrusion.
</paragraph>
</subSubSection>
<subSubSection id="98E66504FF81FFE4FF3C93D78F09744F" pageId="30" pageNumber="239" type="distribution">
<paragraph id="D043368FFF81FFE4FF3C93D78E5975F2" blockId="30.[136,1452,1058,1913]" pageId="30" pageNumber="239">
<emphasis id="E288EA9DFF81FFE4FF3C93D78CE97596" bold="true" box="[189,477,1670,1696]" pageId="30" pageNumber="239">Distribution and habitat</emphasis>
:—Southern
<collectingCountry id="A8EB761FFF81FFE4FDEB93D78FFD7596" box="[618,713,1670,1696]" name="Vietnam" pageId="30" pageNumber="239">Vietnam</collectingCountry>
in
<collectingRegion id="1238F86DFF81FFE4FD6893D78EE67596" box="[745,978,1670,1697]" country="Vietnam" name="Khanh Hoa" pageId="30" pageNumber="239">Khanh Hoa province</collectingRegion>
, known only from Hon Ba Nature Reserve, in montane rainforest at 1,449(
<geoCoordinate id="B5C85048FF81FFE4FE6393FB8F1775F3" box="[482,547,1706,1733]" degrees="1,250" direction="north" orientation="latitude" pageId="30" pageNumber="239" precision="55" value="1.25">1,250</geoCoordinate>
<geoCoordinate id="B5C85048FF81FFE4FDA293FB8F5F75F3" box="[547,619,1706,1733]" degrees="1,578" direction="west" orientation="longitude" pageId="30" pageNumber="239" precision="55" value="-1.578">-1,578</geoCoordinate>
) m elevation (
<figureCitation id="48C72A0AFF81FFE4FC8C93FB8E5475F2" box="[781,864,1706,1732]" captionStart="FIGURE 17" captionStartId="29.[136,229,1425,1447]" captionTargetBox="[243,1346,229,1338]" captionTargetId="figure-17@29.[222,1365,212,1355]" captionTargetPageId="29" captionText="FIGURE 17. Distribution maps of L. nuichuaensis, L. poilanei, L. tenuisecta, and L. verrucosa." figureDoi="http://doi.org/10.5281/zenodo.5964672" httpUri="https://zenodo.org/record/5964672/files/figure.png" pageId="30" pageNumber="239">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="D043368FFF81FFE4FF3C939F8F09744F" blockId="30.[136,1452,1058,1913]" pageId="30" pageNumber="239">
<emphasis id="E288EA9DFF81FFE4FF3C939F8C9F75DE" bold="true" box="[189,427,1742,1768]" pageId="30" pageNumber="239">Taxonomic notes:—</emphasis>
Six
<emphasis id="E288EA9DFF81FFE4FE57939F8CD575DE" bold="true" box="[470,481,1742,1768]" pageId="30" pageNumber="239">s</emphasis>
pecimens formerly included in
<taxonomicName id="17FC4D0CFF81FFE4FCBA939E892975DE" authorityName="Henderson &amp; Bacon" authorityYear="2011" baseAuthorityName="Gagnepain" box="[827,1053,1742,1768]" class="Liliopsida" family="Arecaceae" genus="Lanonia" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="239" phylum="Tracheophyta" rank="species" species="hexasepala">
<emphasis id="E288EA9DFF81FFE4FCBA939E892975DE" box="[827,1053,1742,1768]" italics="true" pageId="30" pageNumber="239">Lanonia hexasepala</emphasis>
</taxonomicName>
(see notes under that species) differ from others by their narrow segments with ruffled margins (
<emphasis id="E288EA9DFF81FFE4FCBE93A28EB3743A" box="[831,903,1779,1804]" italics="true" pageId="30" pageNumber="239">versus</emphasis>
wider segments with straight margins), and are here recognized as a separate species,
<taxonomicName id="17FC4D0CFF81FFE4FDB492468FF07406" box="[565,708,1815,1840]" class="Liliopsida" family="Arecaceae" genus="Lanonia" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="239" phylum="Tracheophyta" rank="species" species="tenuisecta">
<emphasis id="E288EA9DFF81FFE4FDB492468FF07406" box="[565,708,1815,1840]" italics="true" pageId="30" pageNumber="239">L. tenuisecta</emphasis>
</taxonomicName>
. Quantitatively
<taxonomicName id="17FC4D0CFF81FFE4FCF89246893D7406" box="[889,1033,1815,1840]" class="Liliopsida" family="Arecaceae" genus="Lanonia" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="239" phylum="Tracheophyta" rank="species" species="tenuisecta">
<emphasis id="E288EA9DFF81FFE4FCF89246893D7406" box="[889,1033,1815,1840]" italics="true" pageId="30" pageNumber="239">L. tenuisecta</emphasis>
</taxonomicName>
differs from
<taxonomicName id="17FC4D0CFF81FFE4FB1C9246880C7406" authorityName="Henderson &amp; Bacon" authorityYear="2011" baseAuthorityName="Gagnepain" box="[1181,1336,1814,1840]" class="Liliopsida" family="Arecaceae" genus="Lanonia" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="239" phylum="Tracheophyta" rank="species" species="hexasepala">
<emphasis id="E288EA9DFF81FFE4FB1C9246880C7406" box="[1181,1336,1814,1840]" italics="true" pageId="30" pageNumber="239">L. hexasepala</emphasis>
</taxonomicName>
in several variables, notably its exceptionally long ligules (mean length
<quantity id="17049B6AFF81FFE4FCC9926B8E907463" box="[840,932,1850,1877]" metricMagnitude="-1" metricUnit="m" metricValue="3.29" pageId="30" pageNumber="239" unit="cm" value="32.9">32.9 cm</quantity>
<emphasis id="E288EA9DFF81FFE4FC2C926A8EC17462" box="[941,1013,1851,1876]" italics="true" pageId="30" pageNumber="239">versus</emphasis>
<quantity id="17049B6AFF81FFE4FC7C926B896F7463" box="[1021,1115,1850,1877]" metricMagnitude="-1" metricUnit="m" metricValue="1.04" pageId="30" pageNumber="239" unit="cm" value="10.4">10.4 cm</quantity>
) and longer middle segments (mean length 31.0 cm
<emphasis id="E288EA9DFF81FFE4FE02920E8CFF744E" box="[387,459,1887,1912]" italics="true" pageId="30" pageNumber="239">versus</emphasis>
20.0 cm).
</paragraph>
</subSubSection>
</treatment>
</document>
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