402 lines
40 KiB
XML
402 lines
40 KiB
XML
<document id="DB8084E64221B76D51F4F138F13A5824" ID-CLB-Dataset="298601" ID-DOI="10.3897/zse.100.120207" ID-GBIF-Dataset="6f7f44bf-fab1-47dd-aa0a-d5ad482b207e" ID-ZooBank="55DBF63A-85CF-42C0-8218-15F310FB177A" ID-publisher-id="120207" URI-arpha="123D4283-0C8F-5E2E-9E65-4D0700BDBA20" URI-zoobank="http://zoobank.org/55DBF63A-85CF-42C0-8218-15F310FB177A" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1718641467983" checkinUser="pensoft" docAuthor="Villarreal, Osvaldo, Ahumada-C., Daniela & Navas-S., Gabriel R." docDate="2024" docId="4B688B0A7C9D5F86AA3513FEBAB43BB3" docLanguage="en" docName="Zoosystematics and Evolution 100(3): 803-820" docOrigin="Zoosystematics and Evolution 100 (3)" docSource="https://zse.pensoft.net/article/120207/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Eutimesius canoabo Villarreal" docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="820" masterDocId="123D42830C8F5E2E9E654D0700BDBA20" masterDocTitle="Exploring the diversity of Eutimesius Roewer, 1913: new species and records from Colombia and Venezuela (Opiliones, Gonyleptoidea, Stygnidae)" masterLastPageNumber="820" masterPageNumber="803" pageNumber="803" updateTime="1718651764718" updateUser="ExternalLinkService">
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<mods:title id="5C734F8AD10213E8426DB08B67D521E0">Exploring the diversity of Eutimesius Roewer, 1913: new species and records from Colombia and Venezuela (Opiliones, Gonyleptoidea, Stygnidae)</mods:title>
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<mods:namePart id="4D25C9F44C0BA37B0A57149D03FB1332">Villarreal, Osvaldo</mods:namePart>
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<mods:affiliation id="39E6BF192C58AF137632710DD34F8DD6">Centro de Ecología, Instituto Venezolano de Investigaciones Científicas (IVIC), km 11 carretera Panamericana, Altos de Pipe, edo. Miranda 1204 - A, Venezuela & Instituto y Museo del Instituto de Zoología Agrícola, Facultad de Agronomía, Universidad Central de Venezuela, Apartado 4579, Maracay 2101, Aragua, Venezuela & Departamento de Invertebrados, Museu Nacional / UFRJ, Quinta da Boa Vista, São Cristóvão, 20.940 - 040, Rio de Janeiro – RJ, Brazil</mods:affiliation>
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<mods:namePart id="C6DCFC20FC498E1583CEF1EDAEAA7F52">Ahumada-C., Daniela</mods:namePart>
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<mods:nameIdentifier id="3C0983F0452C4F9BB867F18C6C7161D3" type="ORCID">0000-0002-4182-5143</mods:nameIdentifier>
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<mods:affiliation id="08CEE241B31A6CAC4A7D63C39DC05BE9">Grupo de Investigación Biología Descriptiva y Aplicada, Programa de Biología, Universidad de Cartagena, Zaragocilla Cra. 50 # 24 - 120, Cartagena de Indias, Bolívar, Colombia</mods:affiliation>
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<mods:namePart id="9EBDFE7DA3DA507BF8DD94A3B69CA0A5">Navas-S., Gabriel R.</mods:namePart>
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<mods:nameIdentifier id="892243BC2BF50C359C7889622CB4D897" type="ORCID">0000-0001-9554-6345</mods:nameIdentifier>
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<mods:affiliation id="528CC986FB09B8B31AADB2A7DD5CA889">Grupo de Investigación Biología Descriptiva y Aplicada, Programa de Biología, Universidad de Cartagena, Zaragocilla Cra. 50 # 24 - 120, Cartagena de Indias, Bolívar, Colombia</mods:affiliation>
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<mods:title id="7E75480A1336F7E77E20D4308058056B">Zoosystematics and Evolution</mods:title>
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<mods:part id="A233CB33FDD280682F33AAC7A65CD8AB">
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<mods:date id="7B58FE2A4F86A8164DF58423C84893F3">2024</mods:date>
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<mods:number id="E9785F8B0D031865805E937059F55FAB">2024-06-17</mods:number>
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<mods:number id="147993C8846B47E4B607FB5E17AB59D3">100</mods:number>
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<mods:classification id="6E9547E628E0CD218B5E8F60E9648345">journal article</mods:classification>
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<mods:identifier id="A96CA842B51169D707228D261D421155" type="CLB-Dataset">298601</mods:identifier>
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<mods:identifier id="BCFC9E5CEB0846BB269EF0340A8669B7" type="DOI">10.3897/zse.100.120207</mods:identifier>
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<treatment id="4B688B0A7C9D5F86AA3513FEBAB43BB3" ID-DOI="http://doi.org/10.5281/zenodo.11975133" ID-GBIF-Taxon="233216501" ID-Zenodo-Dep="11975133" ID-arpha="4B688B0A-7C9D-5F86-AA35-13FEBAB43BB3" ID-zoobank="https://zoobank.org/D9A7D2ED-CF81-4446-B721-B4402141826B" LSID="urn:lsid:plazi:treatment:4B688B0A7C9D5F86AA3513FEBAB43BB3" LSID-ZBK="urn:lsid:zoobank.org:act:D9A7D2ED-CF81-4446-B721-B4402141826B" httpUri="http://treatment.plazi.org/id/4B688B0A7C9D5F86AA3513FEBAB43BB3">
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||
<subSubSection id="6AE25CDA0F9832451E135DD82942AB31" type="nomenclature">
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||
<paragraph id="AC4D13A48790E6FE8EC6039F017135ED">
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||
<taxonomicName id="64A78C9CB86480F20DFE82F4022F870C" authority="Villarreal" authorityName="Villarreal" class="Arachnida" family="Stygnidae" genus="Eutimesius" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="canoabo">
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||
<emphasis id="53FBF065D3C6A0DA1F432A14E9A5A2D1" italics="true">Eutimesius canoabo</emphasis>
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||
Villarreal
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||
</taxonomicName>
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& Ahumada-C. sp. nov.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="6A3D0C87115ABA63D344AD073AEACD84" type="description">
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||
<paragraph id="1948E5DCEE31580EFBEA350650CCC6A5">
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||
<figureCitation id="85C88EA741D038D4982C6F642B037E43" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Eutimesius canoabo sp. nov., A, B. Male habitus, dorsal and lateral views (MIZA 0105945); C, D. Female habitus, dorsal view (MIZA 0105946). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure6" httpUri="https://binary.pensoft.net/fig/1072634">Figs 6</figureCitation>
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||
,
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<figureCitation id="13425D9A606C26DC5C51979996ABE156" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Eutimesius canoabo sp. nov., male (MIZA 0105945): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure7" httpUri="https://binary.pensoft.net/fig/1072635">7</figureCitation>
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||
,
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||
<figureCitation id="D7E785F57CA640DE28FCFEC5C060EF5E" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Eutimesius canoabo sp. nov. A – C. From Carabobo State, Venezuela." figureDoi="10.3897/zse.100.120207.figure8" httpUri="https://binary.pensoft.net/fig/1072636">8</figureCitation>
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,
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<figureCitation id="37A9C53A0F3D9A11BEBF7D22EBABD312" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eutimesius spp., penis, distal portion in dorsal, ventral, and lateral view: A – C. E. aroa sp. nov., (MIZA 0105934); D – F. E. canoabo sp. nov., (MIZA 0105945); G – I. E. ephippiatus, (ICN-Ao - 1492); J – L. E. guaichia sp. nov., (MIZA 0105935); M – O. E. simoni, (ICN-Ao - 328). A – I. Unscaled." figureDoi="10.3897/zse.100.120207.figure14" httpUri="https://binary.pensoft.net/fig/1072642">14 D – F</figureCitation>
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,
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<figureCitation id="F04310BE7195DABD175A8902DE1BF99A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Map showing the distribution of Eutimesius." figureDoi="10.3897/zse.100.120207.figure2" httpUri="https://binary.pensoft.net/fig/1072630">2</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="SECID0EOJAG" type="type material">
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||
<paragraph id="7789124204503EAFF10FE62B739C0787">
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||
<heading id="3B0D965141872CA3E797F721DD4090A0" reason="title">Type material.</heading>
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||
</paragraph>
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||
<paragraph id="2B036667950B0759BBA231353B15E819">
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||
<materialsCitation id="87DF37B8DAF9064E333EBA44D1E7DD3D" ID-GBIF-Occurrence="4879930303" collectedFrom="on vegetation abbott 1.5 - 2 m above the ground" collectingDate="2008-03-19" collectionCode="MIZA" collectorName="Villarreal O. & Pereira M. P." country="Venezuela" county="El Santuario" elevation="1220" latitude="10.3132" location="Canoabo" longLatPrecision="7" longitude="-68.2232" municipality="Posada Ecologica Casa Maria" specimenCode="MIZA 0105945" specimenCount="1" specimenCount-male="1" stateProvince="Carabobo" typeStatus="holotype">
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||
<collectingCountry id="03E68476BCE1020EEBE10BDEFB95C8DB" name="Venezuela">Venezuela</collectingCountry>
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||
•
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||
<specimenCount id="B9018878A5CB30E318C2410C374E21D6" count="1" type="male">♂</specimenCount>
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||
<emphasis id="3204658E7B7E11654A5FC069BC117757" bold="true" italics="true">
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||
<typeStatus id="3744F37191DB9F11CDA33B90950415DE" type="holotype">holotype</typeStatus>
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||
</emphasis>
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||
;
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||
<collectingRegion id="9137B5F09132727A1FBDD6206AEAD129" country="Venezuela" name="Carabobo">Carabobo</collectingRegion>
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||
,
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||
<collectingCounty id="6D862701A63B34A06413BCAD6AA3A28C">El Santuario</collectingCounty>
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||
,
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<collectingMunicipality id="9F1EDBD1CD168B16B0D23A6D17138FEF">Posada Ecológica Casa María</collectingMunicipality>
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||
, near
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||
<location id="9D246788E8CD7A4D39717F8C1D41ACD2" LSID="urn:lsid:plazi:treatment:4B688B0A7C9D5F86AA3513FEBAB43BB3:9D246788E8CD7A4D39717F8C1D41ACD2" country="Venezuela" county="El Santuario" latitude="10.3132" longLatPrecision="7" longitude="-68.2232" municipality="Posada Ecologica Casa Maria" name="Canoabo" stateProvince="Carabobo">Canoabo</location>
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||
, (
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||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1JAG" specific-use="{"type":"Point","coordinates":[-68.223200,10.313200]}">
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||
<geoCoordinate id="FF0A8203ADF931525F96AB35930B2692" degrees="10.3132" direction="north" orientation="latitude" precision="5" value="10.3132">10.3132 ° N</geoCoordinate>
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||
,
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<geoCoordinate id="38A1FD30524D34316219C60BED4097F0" degrees="68.2232" direction="west" orientation="longitude" precision="5" value="-68.2232">68.2232 ° W</geoCoordinate>
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||
</named-content>
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||
);
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||
<elevation id="DA0E8F8F300DCE9E1A6944D451711DAE" metricMagnitude="3" metricUnit="m" metricValue="1.22" unit="m" value="1220.0">
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||
<quantity id="2C55C455A4DF73A33FB2BCCEEE3E39E6" metricMagnitude="3" metricUnit="m" metricValue="1.22" unit="m" value="1220.0">1,220 m</quantity>
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a. s. l.
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</elevation>
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||
;
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<date id="053303917B8790F17625DC7A24182771" value="2008-03-19">
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<collectingDate id="86C3350A7D43CCCF1CB1A6FE6ACBCCE4" value="2008-03-19">19 Mar. 2008</collectingDate>
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||
</date>
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; (
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<collectorName id="45BD9E8D15A3671FEB061F4943FD6B03">Villarreal O.</collectorName>
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,
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<collectorName id="DE37619BB7F2906BDC28523258B8BC93">Pereira M. P.</collectorName>
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leg.);
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<collectedFrom id="BAC68C5F7652023EF5D620C5008C55E0">
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on vegetation abbott
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<quantity id="09E14896566B07825C08DD5488413B20" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" unit="m" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 m</quantity>
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above the ground
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</collectedFrom>
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(
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<specimenCode id="78545EA07D6F4B6CC019B7997FC77B70" collectionCode="MIZA" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" type="Museum">
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||
<named-content id="43E886AE077D09FFBC63D10A4422A983" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-del-instituto-de-zoologia-agricola-francisco-fernandez-yepez" xlink_title="Museo del Instituto de Zoología Agrícola “Francisco Fernández Yépez”, Maracay, Venezuela">MIZA</named-content>
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0105945
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||
</specimenCode>
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||
)
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||
</materialsCitation>
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||
.
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<materialsCitation id="A91F172676F379EF11E30464740F75C4" ID-GBIF-Occurrence="4879930321" collectedFrom="on vegetation abbott 1.5 - 2 m above the ground" collectingDate="2008-03-19" collectionCode="MIZA" collectorName="Villarreal O. & Pereira M. P." country="Venezuela" county="El Santuario" elevation="1220" latitude="10.3132" location="Canoabo" longLatPrecision="7" longitude="-68.2232" municipality="Posada Ecologica Casa Maria" specimenCode="MIZA 0105946" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Carabobo" typeStatus="paratype">
|
||
<emphasis id="BB33739FD7DF5BFBE07C2629872C3CCB" bold="true" italics="true">
|
||
<typeStatus id="A0AB717D0E90E3C5FDC46C4626D4A8B0" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
•
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||
<specimenCount id="E1055F1000D1DFCEDE962E05B4A42916" count="2" type="female">2 ♀</specimenCount>
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||
,
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||
<specimenCount id="EEBC73F48958C2916E95411C5CBC86F2" count="1" type="male">1 ♂</specimenCount>
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||
; same as the holotype; (
|
||
<specimenCode id="2EFFFB6465EA697E773A94A7AD7A52C7" collectionCode="MIZA" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" type="Museum">
|
||
<named-content id="65666B1149BC6F6B10828AD667E5DB71" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-del-instituto-de-zoologia-agricola-francisco-fernandez-yepez" xlink_title="Museo del Instituto de Zoología Agrícola “Francisco Fernández Yépez”, Maracay, Venezuela">MIZA</named-content>
|
||
0105946
|
||
</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EMKAG" type="diagnosis">
|
||
<paragraph id="B03A751FCEAAAFF69FB11C916464FE40">
|
||
<heading id="E8EC31CBF30381D6A523DE74F54819C4" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="2C53685CFD399CEB00F9AD0E419DA1BD">
|
||
It is distinguishable from all other species in the genus by the pattern of dry white spots, occupying lateral zones of the scutal areas II and III, the medial zone of the scutal areas I and II, dispersal spots on the medial zone of the carapace, and part of the lateral margins of the dorsal scutum (Figs
|
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<figureCitation id="5C49A5466EA7300F85E0A16B8F4863ED" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Eutimesius canoabo sp. nov., A, B. Male habitus, dorsal and lateral views (MIZA 0105945); C, D. Female habitus, dorsal view (MIZA 0105946). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure6" httpUri="https://binary.pensoft.net/fig/1072634">6 A, B</figureCitation>
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||
,
|
||
<figureCitation id="D0AEB385E4DF1BC01147572BC8506ABF" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Eutimesius canoabo sp. nov., male (MIZA 0105945): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure7" httpUri="https://binary.pensoft.net/fig/1072635">7 A</figureCitation>
|
||
,
|
||
<figureCitation id="F12552D77FB7770C2BF2DF9CDBA0878F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Eutimesius canoabo sp. nov. A – C. From Carabobo State, Venezuela." figureDoi="10.3897/zse.100.120207.figure8" httpUri="https://binary.pensoft.net/fig/1072636">8 A</figureCitation>
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||
); and by the ornamentation of the male femur IV, with large tubercles on both complete ventral rows (Fig.
|
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<figureCitation id="E2F2A1556C7F9661577652F1D77900F5" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Eutimesius canoabo sp. nov., male (MIZA 0105945): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure7" httpUri="https://binary.pensoft.net/fig/1072635">7 G</figureCitation>
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||
).
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||
</paragraph>
|
||
<caption id="87C3840F5855AEE8ADD5B56AA6CAE1BF" ID-DOI="10.3897/zse.100.120207.figure6" ID-arpha="DFD12474-59C8-578F-82C2-19A880158DAE" httpUri="https://binary.pensoft.net/fig/1072634" startId="F6">
|
||
<paragraph id="8503F232154CCF89CC227CA560B6C037">
|
||
<label id="A93F5AA74DFCFD785EBAA3AD46B8FAE4">Figure 6.</label>
|
||
</paragraph>
|
||
<paragraph id="3A4D2E74C1795F200A10A1B7F04507FF">
|
||
<taxonomicName id="30D08A04FA8D27F352633E08AC5612D2" authority="Villarreal & Ahumada-C. & Navas-S., 2024" authorityName="Villarreal & Ahumada-C. & Navas-S." authorityYear="2024" class="Arachnida" family="Stygnidae" genus="Eutimesius" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="canoabo" status="sp. nov.">
|
||
<emphasis id="DBB4BB5096D36383C4B9BA1E342DFEE5" italics="true">Eutimesius canoabo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8CD3376D0C27A2FF9684CF3D342D2DBE" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="2697D9936144B77FBF40582D66158B7A" bold="true">A, B.</emphasis>
|
||
Male habitus, dorsal and lateral views (
|
||
<specimenCode id="B1E028E3BB4775040509EADD10ADDE64">
|
||
<named-content id="3F307CB403CC2766091D797282F87D85" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-del-instituto-de-zoologia-agricola-francisco-fernandez-yepez" xlink_title="Museo del Instituto de Zoología Agrícola “Francisco Fernández Yépez”, Maracay, Venezuela">MIZA</named-content>
|
||
0105945
|
||
</specimenCode>
|
||
);
|
||
<emphasis id="69A96744F26808E09EAA5787EE6CC5D2" bold="true">C, D.</emphasis>
|
||
Female habitus, dorsal view (
|
||
<specimenCode id="7A6669834BDADCFBAC24AF11480BC013">
|
||
<named-content id="92D4608D5C7063766542117645A92CFC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-del-instituto-de-zoologia-agricola-francisco-fernandez-yepez" xlink_title="Museo del Instituto de Zoología Agrícola “Francisco Fernández Yépez”, Maracay, Venezuela">MIZA</named-content>
|
||
0105946
|
||
</specimenCode>
|
||
). Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="44D81E008EA5A8714169C3BE12FC3A39" ID-DOI="10.3897/zse.100.120207.figure7" ID-arpha="EDC4B4D2-F052-57A9-90B3-0F55C28453C4" httpUri="https://binary.pensoft.net/fig/1072635" startId="F7">
|
||
<paragraph id="4FE70B202520A768F182AB458673042E">
|
||
<label id="BDD66F772B4ACCC19DACDC77B30BC852">Figure 7.</label>
|
||
</paragraph>
|
||
<paragraph id="6937857FA4B7AE0E30041F8723CC2323">
|
||
<taxonomicName id="840E41960B1C2E691A0C63B0114483AF" authority="Villarreal & Ahumada-C. & Navas-S., 2024" authorityName="Villarreal & Ahumada-C. & Navas-S." authorityYear="2024" class="Arachnida" family="Stygnidae" genus="Eutimesius" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="canoabo" status="sp. nov.">
|
||
<emphasis id="A093A5CF2FDA6A19FDB6E2B88F9E6BF2" italics="true">Eutimesius canoabo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="35C2F6E58958FAE07067F5D8039012ED" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male (
|
||
<specimenCode id="EDB33D822BA704F675FE718D0453BEBD">
|
||
<named-content id="7FFA5672EC204DB481BBC51B02751A80" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-del-instituto-de-zoologia-agricola-francisco-fernandez-yepez" xlink_title="Museo del Instituto de Zoología Agrícola “Francisco Fernández Yépez”, Maracay, Venezuela">MIZA</named-content>
|
||
0105945
|
||
</specimenCode>
|
||
):
|
||
<emphasis id="2B95CCB0766045436F0F4116041EE1FF" bold="true">A.</emphasis>
|
||
Habitus, dorsal view;
|
||
<emphasis id="269DB0D90A0EF6A88AA759DAB78661D5" bold="true">B.</emphasis>
|
||
Ditto, lateral view;
|
||
<emphasis id="CEC806A65B0F9B0FBB4A6A57B475EDF7" bold="true">C.</emphasis>
|
||
Right chelicera, frontal view;
|
||
<emphasis id="E9F255F71A9245603077732CCE5CAC4D" bold="true">D.</emphasis>
|
||
Right pedipalp, tibia, and tarsus, ectal view;
|
||
<emphasis id="4F92863418E40104DCF11F03C4C5089A" bold="true">E.</emphasis>
|
||
Ditto, mesal view;
|
||
<emphasis id="D7E4211EAC7FF5603A2DC012B0609319" bold="true">F.</emphasis>
|
||
Right leg IV, dorsal view;
|
||
<emphasis id="7E9A28EAEE3A90850824C4619DE77A3A" bold="true">G.</emphasis>
|
||
Ditto, ventral view. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="3129188EEAE1386BE6E7F1A5FDA24CB2" ID-DOI="10.3897/zse.100.120207.figure8" ID-arpha="1E2BD392-0667-5E92-9C01-A261F6CE1022" httpUri="https://binary.pensoft.net/fig/1072636" startId="F8">
|
||
<paragraph id="2D084997FE5C55DBBEB2B41146E6DA72">
|
||
<label id="E53744D31A2E17F9EE98A53F90CFA6CF">Figure 8.</label>
|
||
</paragraph>
|
||
<paragraph id="FA160F983587BD068F912C2E70A0ED60">
|
||
<taxonomicName id="8C722292169A45B0F87EC0B2798D1F5D" authority="Villarreal & Ahumada-C. & Navas-S., 2024" authorityName="Villarreal & Ahumada-C. & Navas-S." authorityYear="2024" class="Arachnida" family="Stygnidae" genus="Eutimesius" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="canoabo" status="sp. nov.">
|
||
<emphasis id="E0238CA5B6547CAE2A49FCED55A998D0" italics="true">Eutimesius canoabo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2529048E7B51F676A44A9DD6924AC0AC" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="5C31B1F65B9ECE417530600818771AAB" bold="true">A – C.</emphasis>
|
||
From Carabobo State, Venezuela.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ENOAG" type="etymology">
|
||
<paragraph id="59FCEC9B641E00039FC3DF69794E633F">
|
||
<heading id="85233BE033D9E33B7C9588AE3C8F802F" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="901FCD55168F37A2314C3F2EA230AD1E">
|
||
Canoabo is an indigenous word of Arawako origin that means “ village next to fresh water. ” The species name refers to the
|
||
<typeStatus id="C71B9351659204F1C165D2F43C336318">type</typeStatus>
|
||
locality, a forest near Canoabo, a town and river of the Cordillera de la Costa in
|
||
<collectingRegion id="49ADFD67C4CDDFEF9CB845993883E004" country="Venezuela" name="Carabobo">Carabobo State</collectingRegion>
|
||
,
|
||
<collectingCountry id="50814CC5C2401C6D4EB7CDC2B1A7E266" name="Venezuela">Venezuela</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ESOAG" type="description">
|
||
<paragraph id="8FAF8A9A5AAF8DD7E5BD02427D2AD9DF">
|
||
<heading id="908D9B9F319EA0380CDB49A5B5204B24" reason="title">Description.</heading>
|
||
</paragraph>
|
||
<paragraph id="5D0525154222452387599194F4053FF3">
|
||
<emphasis id="F11F892629A13CE7027814034E044394" bold="true">
|
||
Male.
|
||
<emphasis id="4A7A91A0FC311DDD43C3E5443293A225" bold="true" italics="true">Measurements</emphasis>
|
||
.
|
||
</emphasis>
|
||
<abbrev id="ABBRID0E4OAG" xlink_title="dorsal scutum length">DSL</abbrev>
|
||
3.4;
|
||
<abbrev id="ABBRID0EBPAG" xlink_title="dorsal scutum width">DSW</abbrev>
|
||
3.1;
|
||
<abbrev id="ABBRID0EFPAG" xlink_title="maximum abdominal scutum length">AL</abbrev>
|
||
1.8;
|
||
<abbrev id="ABBRID0EJPAG" xlink_title="maximum abdominal scutum width">AW</abbrev>
|
||
3.1;
|
||
<abbrev id="ABBRID0ENPAG" xlink_title="interocular distance">IOD</abbrev>
|
||
1.9; pedipalp:
|
||
<abbrev id="ABBRID0ERPAG" xlink_title="pedipalpal coxa">CoPp</abbrev>
|
||
0.5, TrPp 0.9,
|
||
<abbrev id="ABBRID0EVPAG" xlink_title="pedipalpal femur">FePp</abbrev>
|
||
3.3,
|
||
<abbrev id="ABBRID0EZPAG" xlink_title="pedipalpal patella">PaPp</abbrev>
|
||
1.5,
|
||
<abbrev id="ABBRID0E4PAG" xlink_title="pedipalpal tibia">TiPp</abbrev>
|
||
1.2,
|
||
<abbrev id="ABBRID0EBQAG" xlink_title="pedipalpal tarsus">TaPp</abbrev>
|
||
1.1,
|
||
<abbrev id="ABBRID0EFQAG" xlink_title="pedipalp claw">ClPp</abbrev>
|
||
1.1; total 9.6; leg IV:
|
||
<abbrev id="ABBRID0EJQAG" xlink_title="femur length">FeL</abbrev>
|
||
7.4,
|
||
<abbrev id="ABBRID0ENQAG" xlink_title="tibia length">TiL</abbrev>
|
||
3.6.
|
||
<emphasis id="5CD13514A08469DE9DFC644533491057" bold="true" italics="true">Dorsum</emphasis>
|
||
(Figs
|
||
<figureCitation id="8E65FFB07579144D161E812683A23096" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Eutimesius canoabo sp. nov., A, B. Male habitus, dorsal and lateral views (MIZA 0105945); C, D. Female habitus, dorsal view (MIZA 0105946). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure6" httpUri="https://binary.pensoft.net/fig/1072634">6 A</figureCitation>
|
||
,
|
||
<figureCitation id="E86A9C5FB2F42A87AB826A279748228E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Eutimesius canoabo sp. nov., male (MIZA 0105945): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure7" httpUri="https://binary.pensoft.net/fig/1072635">7 A</figureCitation>
|
||
,
|
||
<figureCitation id="2D607CA1C7D063FC4FF3EA9FF075E366" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Eutimesius canoabo sp. nov. A – C. From Carabobo State, Venezuela." figureDoi="10.3897/zse.100.120207.figure8" httpUri="https://binary.pensoft.net/fig/1072636">8 A, C</figureCitation>
|
||
).
|
||
<abbrev id="ABBRID0EARAG" xlink_title="dorsal scutum">DS</abbrev>
|
||
outline Epsilon
|
||
<typeStatus id="097D29F7C0B8F03E5466BD026C2A071B">type</typeStatus>
|
||
. Anterior margin of
|
||
<abbrev id="ABBRID0EERAG" xlink_title="dorsal scutum">DS</abbrev>
|
||
with two anterolateral tubercles. Anteromedial process of the cheliceral sockets shorter than lateral processes. Eyes separated into two small smooth mounds, placed posteriorly on the carapace. Interocular region with one central elevated mound with small granules that terminates in four short spines. Lateral margins smooth. Mesotergum divided into four areas, III – IV fused: I divided medially into two triangular halves, with one conspicuous tubercle on each side; II entire, with four conspicuous tubercles; III – IV with a lateral pair of large tubercles; one pair of paramedian large spines with granulated base, with two posterior tubercles. Posterior margin and free tergites smooth, with a pair of paramedian acute granules.
|
||
<emphasis id="E6D73C6CF8163D338F0B59FBE2E80393" bold="true" italics="true">Venter</emphasis>
|
||
(Figs
|
||
<figureCitation id="FC40F36892DD0ADF327B14409EC96F7D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Eutimesius canoabo sp. nov., A, B. Male habitus, dorsal and lateral views (MIZA 0105945); C, D. Female habitus, dorsal view (MIZA 0105946). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure6" httpUri="https://binary.pensoft.net/fig/1072634">6 B</figureCitation>
|
||
,
|
||
<figureCitation id="DE4A43625781CC94000AB7756C0CDE1C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Eutimesius canoabo sp. nov., male (MIZA 0105945): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure7" httpUri="https://binary.pensoft.net/fig/1072635">7 B</figureCitation>
|
||
). Coxa I with a cluster of three mesal granules and about 7–8 dispersed granules; II with five intercoxal tubercles, with a median row of eight tubercles and two distal tubercles; III with five-six intercoxal tubercles, 14–15 six tubercles, medially aligned and distally irregularly distributed; IV with seven anterior tubercles and about 18–19 posterior tubercles not aligned. Genital operculum with scattered granules. Stigmatic area with a posterior row of minute granules on the posterior border. Free sternites with a row of small granules.
|
||
<emphasis id="F298369D7025A9FCC6ACFEEC77BCD342" bold="true" italics="true">Chelicerae</emphasis>
|
||
(Figs
|
||
<figureCitation id="D4F471FBE06DCF2A26466848E99C249A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Eutimesius canoabo sp. nov., A, B. Male habitus, dorsal and lateral views (MIZA 0105945); C, D. Female habitus, dorsal view (MIZA 0105946). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure6" httpUri="https://binary.pensoft.net/fig/1072634">6 A, B</figureCitation>
|
||
,
|
||
<figureCitation id="AE8591208F9E798D2177D1EC5A42AF8B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Eutimesius canoabo sp. nov., male (MIZA 0105945): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure7" httpUri="https://binary.pensoft.net/fig/1072635">7 A – C</figureCitation>
|
||
). Segment I smooth with well-defined bulla, with three or four ectoproximal tubercles and one ectodistal tubercle. Segment II swollen, fixed finger with a proximal wide laminar tooth, followed by one medial tooth and one small denticle subdistal; mobile finger with one subproximal large truncated tooth, one medial pyramidal tooth, and two small subdistal teeth.
|
||
<emphasis id="30747B2AD8A8DCE04BCF65B5BA28FB79" bold="true" italics="true">Pedipalps</emphasis>
|
||
(Fig.
|
||
<figureCitation id="8B1082EF47598B6C0A924360A2DB9744" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Eutimesius canoabo sp. nov., male (MIZA 0105945): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure7" httpUri="https://binary.pensoft.net/fig/1072635">7 D, E</figureCitation>
|
||
). Coxa with a group of about nine ventral tubercles and three dorsal tubercles. Trochanter with two ventral tubercles and two dorsal tubercles. Femur with a ventroectal row of six-eight small granules and one-two ventromesal granules, and dorsally with a row of minute granules. Patella smooth, distally swollen. Tibia dorsal smooth, ventrally with some proximal minute granules; mesal IIiIi; ectal IIiIii. Tarsus dorsally smooth, ventrally with two rows of minute granules, mesal IiIiIi; ectal iiIiiiIii.
|
||
<emphasis id="0F1134AC5CFFE592C4B83177E77BADDA" bold="true" italics="true">Legs</emphasis>
|
||
(Fig.
|
||
<figureCitation id="47DE7E33A0C7BCA6F24352BF800C0AEA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Eutimesius canoabo sp. nov., male (MIZA 0105945): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure7" httpUri="https://binary.pensoft.net/fig/1072635">7 F, G</figureCitation>
|
||
). Coxae I – II with two dorsal tubercles; III – IV connected by one intercoxal tubercle; IV with four dorsodistal tubercles and scattered small lateral granules and tubercles. Trochanter I dorsally smooth, ventrally with three tubercles; II with one dorsal and three ventral tubercles; III with one retrolateral and five ventral tubercles; IV with one prolateral and one retrolateral tubercle and seven ventral tubercles. Femora I – II smooth; III with longitudinal rows of tubercles and granules, the ventrodistal larger, and one retrodorsal distal large tubercle; IV with the one proventral and one retroventral rows of tubercles increasing in size distally, and with two dorsoapical tubercles. Patella III dorsally granulated and with a proventral tubercle; IV with one large ventral tubercle; and pro and retrodorsal distal large tubercles and with sparse minute granules. Tibia III slightly increased distally, with two rows of ventral granules in the distal portion; IV with the proventral row of tubercles increasing in size distally; and retrovental row of tubercles, with the distal-most tubercle larger than the others. Basitarsus I slightly swollen. Tarsal process and scopula present. Tarsal claws III and IV opposites and pectinated. Tarsal counts: 7 (2) - 7 (3) / 19 (3) - 18 (3) / 8 / 9-10.
|
||
<emphasis id="001B9AA57F030C2D2B65DC62AD2D29F4" bold="true" italics="true">Penis</emphasis>
|
||
(Fig.
|
||
<figureCitation id="198BD0DDBB4C1EB0AE5A93E8E4761CAE" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eutimesius spp., penis, distal portion in dorsal, ventral, and lateral view: A – C. E. aroa sp. nov., (MIZA 0105934); D – F. E. canoabo sp. nov., (MIZA 0105945); G – I. E. ephippiatus, (ICN-Ao - 1492); J – L. E. guaichia sp. nov., (MIZA 0105935); M – O. E. simoni, (ICN-Ao - 328). A – I. Unscaled." figureDoi="10.3897/zse.100.120207.figure14" httpUri="https://binary.pensoft.net/fig/1072642">14 D – F</figureCitation>
|
||
).
|
||
<taxonomicName id="A354EFA786B86834AF237FF6CF358293" authorityName="Roewer" authorityYear="1913" class="Arachnida" family="Stygnidae" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="subFamily" subFamily="Heterostygninae">Heterostygninae</taxonomicName>
|
||
general pattern, as described in
|
||
<bibRefCitation id="C98388CB392E5AE051DD28370E2AC440" DOI="10.1590/1678-4766e2019008" author="Villarreal O & Sanchéz N & De Ascenção A" issue="1" journalOrPublisher="Série Zoologia" pagination="1-6" refId="B18" refString="Villarreal O, Sanchéz N, De Ascenção A (2019 b) New generic assignment to the harvestman Metaphareus punctatus (Opiliones: Stygnidae) and observations about it reproductive behavior. Iheringia. Série Zoologia 109 (1): 1–6. https://doi.org/10.1590/1678-4766e2019008" title="New generic assignment to the harvestman Metaphareus punctatus (Opiliones: Stygnidae) and observations about it reproductive behavior. Iheringia." volume="109">Villarreal et al. (2019 b</bibRefCitation>
|
||
). Truncus with the malleus swollen.
|
||
<emphasis id="39F87910189C878EB70D85A987BC1B29" italics="true">Lamina parva</emphasis>
|
||
(
|
||
<abbrev id="ABBRID0E5SAG" xlink_title="lamina parva">LP</abbrev>
|
||
) with a shallow neck and a deep distal cleft.
|
||
<abbrev id="ABBRID0ECTAG" xlink_title="macrosetae of penis">MS</abbrev>
|
||
- A 1 - A 2 located on the malleus, one pair laterally and one pair more ventrally, with duplication in the more ventral on the left side;
|
||
<abbrev id="ABBRID0EGTAG" xlink_title="macrosetae of penis">MS</abbrev>
|
||
- B pair ventrally located, distally to the
|
||
<abbrev id="ABBRID0EKTAG" xlink_title="macrosetae of penis">MS</abbrev>
|
||
- A; two pairs of
|
||
<abbrev id="ABBRID0EOTAG" xlink_title="macrosetae of penis">MS</abbrev>
|
||
- C located medially on the
|
||
<abbrev id="ABBRID0ESTAG" xlink_title="lamina parva">LP</abbrev>
|
||
, dorsally to the neck;
|
||
<abbrev id="ABBRID0EWTAG" xlink_title="macrosetae of penis">MS</abbrev>
|
||
- D 1 located slightly proximal to between
|
||
<abbrev id="ABBRID0E1TAG" xlink_title="macrosetae of penis">MS</abbrev>
|
||
- E and
|
||
<abbrev id="ABBRID0E5TAG" xlink_title="macrosetae of penis">MS</abbrev>
|
||
- C;
|
||
<abbrev id="ABBRID0ECUAG" xlink_title="macrosetae of penis">MS</abbrev>
|
||
- D 2 basally located, near the base of the gland;
|
||
<abbrev id="ABBRID0EGUAG" xlink_title="macrosetae of penis">MS</abbrev>
|
||
- E with only one pair of short setae visible. Gland globose, with short and dorsally curved stylus with inconspicuous dorsal process.
|
||
<emphasis id="BA65D3B1A53CE0785AF4609337286A0B" bold="true" italics="true">Color</emphasis>
|
||
(Figs
|
||
<figureCitation id="9F6E2C4660A103F9A02C9D60077C5655" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Eutimesius canoabo sp. nov., A, B. Male habitus, dorsal and lateral views (MIZA 0105945); C, D. Female habitus, dorsal view (MIZA 0105946). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure6" httpUri="https://binary.pensoft.net/fig/1072634">6</figureCitation>
|
||
,
|
||
<figureCitation id="B7999A0D979E8B2347B07158DB04CA37" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Eutimesius canoabo sp. nov. A – C. From Carabobo State, Venezuela." figureDoi="10.3897/zse.100.120207.figure8" httpUri="https://binary.pensoft.net/fig/1072636">8</figureCitation>
|
||
).
|
||
<abbrev id="ABBRID0EVUAG" xlink_title="dorsal scutum">DS</abbrev>
|
||
mottled on dark yellowish brown (78). Spots on the carapace Brilliant orange (49); tubercles of area I – IV; margins of
|
||
<abbrev id="ABBRID0EZUAG" xlink_title="dorsal scutum">DS</abbrev>
|
||
; margins of free tergites; and edge of coxae I – IV; dorsally light olive brown (94). Trochanters I – IV, with the same colors as
|
||
<abbrev id="ABBRID0E4UAG" xlink_title="dorsal scutum">DS</abbrev>
|
||
. Femora I – II reticulated dark yellowish brown (78) on a background of light olive brown (94); III – IV with the same colors as
|
||
<abbrev id="ABBRID0EBVAG" xlink_title="dorsal scutum">DS</abbrev>
|
||
. Chelicerae reticulated in the same colors as
|
||
<abbrev id="ABBRID0EFVAG" xlink_title="dorsal scutum">DS</abbrev>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="2406B7123DCB560D14B8D315A7136EDD">
|
||
<emphasis id="43C1C1F968BB3DD7DB89EC3C30BB04DE" bold="true">
|
||
Female.
|
||
<emphasis id="A10DBB539563F19F3D147F158210DE23" bold="true" italics="true">Measurements</emphasis>
|
||
.
|
||
</emphasis>
|
||
Dorsal scutum length 3.9; dorsal scutum width 3.5; abdominal scutum length 1.9; abdominal scutum width 2.9; interocular distance 1.5; pedipalp: coxa 0.8, trochanter 0.7, femur 2.8, patella 1.2, tibia 1.8, tarsus 1.6, claw 1.3; total 10.2; leg IV: femur 8.0, tibia 3.4.
|
||
<emphasis id="BC8023DF37159C02E158E119B6E06F7B" bold="true" italics="true">Description</emphasis>
|
||
(Fig.
|
||
<figureCitation id="F4AAC0113CE0E35BD1D2CBE6CA3B3A58" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Eutimesius canoabo sp. nov., A, B. Male habitus, dorsal and lateral views (MIZA 0105945); C, D. Female habitus, dorsal view (MIZA 0105946). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure6" httpUri="https://binary.pensoft.net/fig/1072634">6 C, D</figureCitation>
|
||
). Similar to male, except by abdominal scutum Epsilon
|
||
<typeStatus id="352FD16D4AF29DE9C308056739753F3B">type</typeStatus>
|
||
; chelicerae not swollen; interocular projection lower and slightly forward; ornamentation of leg IV conspicuously less developed; basitarsus I not swollen. Chelicerae and legs lighter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EXVAG" type="distribution">
|
||
<paragraph id="61A052EB2091E3312360F158A4D9A02F">
|
||
<heading id="9BE1FAF978B1F865011C8E050839A39C" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="73CFAA6501B1B7E526FB59DC202A20D3">
|
||
<collectingCountry id="9532C04FB5278EBD507C363CDB956D09" name="Venezuela">Venezuela</collectingCountry>
|
||
,
|
||
<collectingRegion id="251124FB01B614CBCB6CE408D51BB832" country="Venezuela" name="Yaracuy">Yaracuy</collectingRegion>
|
||
. Only known from the
|
||
<typeStatus id="0112E52310270600702EF02EE3A1FAB7">type</typeStatus>
|
||
locality (Fig.
|
||
<figureCitation id="62369EF8BEDA8810CE4C02F33F6F29E2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Map showing the distribution of Eutimesius." figureDoi="10.3897/zse.100.120207.figure2" httpUri="https://binary.pensoft.net/fig/1072630">2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |