264 lines
20 KiB
XML
264 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.13.8824" ID-GBIF-Dataset="2ffb2201-36d7-4721-adb2-7e8790c2556e" ID-GBIF-Taxon="127889337" ID-Pensoft-Pub="1314-4049-13-35" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-4049-13-35" ModsDocOrigin="MycoKeys 13" ModsDocTitle="Psoraaltotibetica (Psoraceae, Lecanorales), a new lichen species from the Tibetan part of the Himalayas" checkinTime="1463149516296" checkinUser="pensoft" docAuthor="Timdal, Einar, Obermayer, Walter & Bendiksby, Mika" docDate="2016" docId="34D39C2F7363AAE34BEC39F10CDE2536" docLanguage="en" docName="MycoKeys 13: 35-48" docOrigin="MycoKeys 13" docSource="http://dx.doi.org/10.3897/mycokeys.13.8824" docTitle="Psora altotibetica Timdal, Obermayer & Bendiksby, sp. nov." docType="treatment" docVersion="3" lastPageId="8" lastPageNumber="43" masterDocId="FFFFFFF96161FFA4FFAE012CFF90322C" masterDocTitle="Psoraaltotibetica (Psoraceae, Lecanorales), a new lichen species from the Tibetan part of the Himalayas" masterLastPageNumber="48" masterPageNumber="35" pageId="5" pageNumber="40" updateTime="1643495094047" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Psoraaltotibetica (Psoraceae, Lecanorales), a new lichen species from the Tibetan part of the Himalayas</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Timdal, Einar</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Obermayer, Walter</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bendiksby, Mika</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>MycoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="volume">
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<mods:number>13</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>35</mods:start>
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<mods:end>48</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/mycokeys.13.8824</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.13.8824</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-4049-13-35</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="127889337" LSID="urn:lsid:plazi:treatment:34D39C2F7363AAE34BEC39F10CDE2536" httpUri="http://treatment.plazi.org/id/34D39C2F7363AAE34BEC39F10CDE2536" lastPageId="8" lastPageNumber="43" pageId="5" pageNumber="40">
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<subSubSection pageId="5" pageNumber="40" type="multiple">
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<paragraph pageId="5" pageNumber="40">Taxon classification Fungi Lecanorales Psoraceae</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="40" type="nomenclature">
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<paragraph pageId="5" pageNumber="40">
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<taxonomicName LSID="Mycobank: MB 816840" authority="Timdal, Obermayer & Bendiksby" class="Lecanoromycetes" family="Psoraceae" genus="Psora" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Psora altotibetica" order="Lecanorales" pageId="5" pageNumber="40" phylum="Ascomycota" rank="species" species="altotibetica">Psora altotibetica Timdal, Obermayer & Bendiksby</taxonomicName>
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<taxonomicNameLabel pageId="5" pageNumber="40">sp. nov.</taxonomicNameLabel>
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Fig. 2
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="40" type="diagnosis">
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<paragraph pageId="5" pageNumber="40">Diagnosis.</paragraph>
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<paragraph pageId="5" pageNumber="40">
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Similar to
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<taxonomicName class="Lecanoromycetes" family="Psoraceae" genus="Psora" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Psora indigirkae" order="Lecanorales" pageId="5" pageNumber="40" phylum="Ascomycota" rank="species" species="indigirkae">Psora indigirkae</taxonomicName>
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, but apothecia black, ascospores shorter, and bourgeanic acid absent from the thallus.
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</paragraph>
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<caption pageId="5" pageNumber="40">
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<paragraph pageId="5" pageNumber="40">
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Figure 2.
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<taxonomicName class="Lecanoromycetes" family="Psoraceae" genus="Psora" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Psora altotibetica" order="Lecanorales" pageId="5" pageNumber="40" phylum="Ascomycota" rank="species" species="altotibetica">Psora altotibetica</taxonomicName>
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, part of holotype. Scale bar = 1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="41" type="materials_examined">
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<paragraph pageId="6" pageNumber="41">
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<pageBreakToken pageId="6" pageNumber="41" start="start">Type</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="6" pageNumber="41">
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CHINA. Xizang: Himalaya Range, 165 km SSE of Lhasa, 40 km W of
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<normalizedToken originalValue="Lhünze">Lhuenze</normalizedToken>
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, little village on way to Nera Tso (=Ni La Hu),
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="28.383333">28°23'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="92.083336">92°05'E</geoCoordinate>
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, 4300-4400 m alt., dry-valley, N-exposed dry slopes, on the ground, 1 Aug 1994, W.Obermayer 5282 (holotype: GZU!).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="41" type="description">
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<paragraph pageId="6" pageNumber="41">Description.</paragraph>
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<paragraph pageId="6" pageNumber="41">
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Thallus squamulose; squamules up to 3 mm wide, rounded, adnate, dispersed to adjacent, weakly concave to plane; upper surface medium brown, dull, becoming moderately to densely pruinose, smooth when young, later with fissures in the cortex; margin concolorous with upper side or partly white, straight to slightly down-turned or slightly up-turned, entire; upper cortex 40-60
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<normalizedToken originalValue="μm">μm</normalizedToken>
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thick, composed of pale brown, thin-walled hyphae with rounded lumina, containing remnants of algae throughout (Chlor-zinc-iodine!), containing crystals of gyrophoric acid (assumedly, dissolving in K) and (in pruinose squamules) crystals of calcium oxalate; epinecral layer hardly developed. Medulla not amyloid, containing both lichen substances (dissolving in K) and calcium oxalate; lower cortex poorly developed, composed of mainly periclinally oriented, pale brown hyphae, not containing crystals; lower surface brown. Apothecia up to 1.2 mm diam., marginal or submarginal on the squamules, plane and indistinctly marginate when young, soon becoming convex and immarginate, black, epruinose or faintly white pruinose at the margin. Hypothecium colourless, containing crystals of calcium oxalate; epihymenium yellowish brown, containing orange crystals dissolving in K, K+ purple. Ascus clavate, with a well-developed, amyloid tholus containing a deeper amyloid tube, lacking an ocular chamber (
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<taxonomicName class="Lecanoromycetes" family="Lecideaceae" genus="Porpidia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Porpidia" order="Lecideales" pageId="6" pageNumber="41" phylum="Ascomycota" rank="genus">Porpidia</taxonomicName>
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-type); ascospores ellipsoid, simple, hyaline, 9-14
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<normalizedToken originalValue="×">x</normalizedToken>
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5-7
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(n = 30). Conidiomata not seen.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="42" type="chemistry">
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<paragraph pageId="7" pageNumber="42">
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<pageBreakToken pageId="7" pageNumber="42" start="start">Chemistry</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="7" pageNumber="42">
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Gyrophoric acid (by TLC); upper cortex and medulla
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<normalizedToken originalValue="K–">K-</normalizedToken>
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, C+ faintly red, KC+ faintly red,
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<normalizedToken originalValue="P–">P-</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="42" type="habitat">
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<paragraph pageId="7" pageNumber="42">Habitat and distribution.</paragraph>
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<paragraph pageId="7" pageNumber="42">The species is terricolous and known from nine localities in China (Tibet) and Nepal at altitudes between 4230 and 5000 m (Fig. 3).</paragraph>
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<caption pageId="7" pageNumber="42">
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<paragraph pageId="7" pageNumber="42">
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Figure 3.
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<taxonomicName class="Lecanoromycetes" family="Psoraceae" genus="Psora" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Psora altotibetica" order="Lecanorales" pageId="7" pageNumber="42" phylum="Ascomycota" rank="species" species="altotibetica">Psora altotibetica</taxonomicName>
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, known distribution. Open circle = holotype locality.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="42" type="etymology">
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<paragraph pageId="7" pageNumber="42">Etymology.</paragraph>
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<paragraph pageId="7" pageNumber="42">The name refers to its occurrence at high altitude in the Tibetan part of the Himalayas.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="43" pageId="7" pageNumber="42" type="materials_examined">
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<paragraph pageId="7" pageNumber="42">Other specimens examined.</paragraph>
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<paragraph lastPageId="8" lastPageNumber="43" pageId="7" pageNumber="42">
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<taxonomicName class="Lecanoromycetes" family="Psoraceae" genus="Psora" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Psora altotibetica" order="Lecanorales" pageId="7" pageNumber="42" phylum="Ascomycota" rank="species" species="altotibetica">Psora altotibetica</taxonomicName>
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. CHINA. Xizang: Shegar, Rongbuk Valley, S of Rongpu-si, 5000 m alt., moraine hill, 7 Jul 1993, T.
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<normalizedToken originalValue="Engelskjøn">Engelskjon</normalizedToken>
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T-030b (TROM L-42812); 5000 m alt., rock base, patchwise on silt, 7 Jul 1993, T.
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<normalizedToken originalValue="Engelskjøn">Engelskjon</normalizedToken>
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T-036 (TROM L-42813); Upper Tsangpo basin, N of Saga,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="29.566668">29°34'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="85.25">85°15'E</geoCoordinate>
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, alt. 4760 m, 28°W, grazed stony relicts of
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<taxonomicName class="Pinopsida" family="Cupressaceae" genus="Juniperus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Juniperus" order="Pinales" pageId="7" pageNumber="42" phylum="Tracheophyta" rank="genus">Juniperus</taxonomicName>
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dwarf-scrub, on ground, 26 Aug 1993, G.Miehe. & S.Miehe 9573/23/02 (GZU) & 9573/23/03 (GZU); 120 km SSW of Quamdo (=Changtu), 10 km S of Bamda,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="30.15">30°09'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="97.28333">97°17'E</geoCoordinate>
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, 4500-4700 m alt., on mosses, overhang, NNW-exp., 6 Aug 1994, W.Obermayer 3967 (GZU); Himalaya Range, 160 km S of Lhasa, dry valley of Kuru river, 10 km NW Lhozag,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="28.4">28°24'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="90.65">90°39'E</geoCoordinate>
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, 4230 m alt., N-exposed steep rocks in a glen, on soil, 17 Jul 1994, W.Obermayer 4350 (GZU) & 4365 (GZU); Himalaya Range, 170 km S of Lhasa, between Lhozhag and Lhakhang Dzong, W-facing slopes of Dhalari mountain,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="28.333334">28°20'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="90.96667">90°58'E</geoCoordinate>
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, 4300 m alt., NNW-exposed,
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<normalizedToken originalValue="±">+/-</normalizedToken>
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underhang, 20 Jul 1994, W.Obermayer 4485 (GZU) & 4502 (GZU); Himalaya Range, 170 km SE of Lhasa, 110 km SSE of Tsetang (Nedong),
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="28.583334">28°35'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="92.38333">92°23'E</geoCoordinate>
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, 4700 m alt., alpine meadows with
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<taxonomicName authority="Willdenow, 1805" authorityName="Willdenow" authorityYear="1805" class="Magnoliopsida" family="Cyperaceae" genus="Kobresia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Kobresia pygmaea" order="Poales" pageId="7" pageNumber="42" phylum="Tracheophyta" rank="species" species="pygmaea">Kobresia pygmaea</taxonomicName>
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, ground with
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<taxonomicName class="Mammalia" family="Ochotonidae" genus="Ochotona" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ochotona" order="Lagomorpha" pageId="7" pageNumber="42" phylum="Chordata" rank="genus">Ochotona</taxonomicName>
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-burrows, 26 Jul 1994, W.Obermayer 4981 (GZU); Himalaya Range, 130 km SE of Lhasa, 50 km SSE of Tsetang (Nedong), on way to the pass Putrang La,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="28.866667">28°52'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="92.1">92°06'E</geoCoordinate>
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, 4400 m alt., dry slope, on soil, 2 Aug 1994,
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<pageBreakToken pageId="8" pageNumber="43" start="start">W</pageBreakToken>
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.Obermayer 5223 (GZU). NEPAL. Mahalangur Himal, Khumbu,
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<normalizedToken originalValue="Moränen">Moraenen</normalizedToken>
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des Khumbu-Gletschers bei Lobuche, 4950-5000 m alt., Sep 1962, J.Poelt 1138 (GZU).
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</paragraph>
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<paragraph pageId="8" pageNumber="43">
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<taxonomicName class="Lecanoromycetes" family="Psoraceae" genus="Psora" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Psora himalayana" order="Lecanorales" pageId="8" pageNumber="43" phylum="Ascomycota" rank="species" species="himalayana">Psora himalayana</taxonomicName>
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. PAKISTAN. Karakorum, Naz Bar (Yasin), 36°17-25'N, 73°0-17'E, 3400-3470 m alt., subalpine
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<taxonomicName class="Pinopsida" family="Cupressaceae" genus="Juniperus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Juniperus" order="Pinales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="genus">Juniperus</taxonomicName>
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communis dwarf-scrub with
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<taxonomicName class="Pinopsida" family="Cupressaceae" genus="Juniperus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Juniperus" order="Pinales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="genus">Juniperus</taxonomicName>
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macropoda trees in Seriphidium maritimum steppe; on silt in rock crevices, 11 Sep 1990, G.Miehe & S.Miehe 3529 (GZU); Karakorum, Baltistan, Haramosh Range,
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<normalizedToken originalValue="“Alm”">"Alm"</normalizedToken>
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Pakora SE Ganto La,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="35.683334">35°41'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="75.35">75°21'E</geoCoordinate>
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, 3600-3800 m alt., pasture and rocks around the alm, rocky slopes, 3 Jul 1991, J.Poelt K91-416 (GZU).
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</paragraph>
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<paragraph pageId="8" pageNumber="43">
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<taxonomicName class="Lecanoromycetes" family="Psoraceae" genus="Psora" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Psora indigirkae" order="Lecanorales" pageId="8" pageNumber="43" phylum="Ascomycota" rank="species" species="indigirkae">Psora indigirkae</taxonomicName>
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. RUSSIA. Sakha Republic: Momskii region, along the river Indigirka, c. 48 km NNW of Tyubelyakh,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="65.8">65°48'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="142.88333">142°53'E</geoCoordinate>
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, 200-300 m alt., on calcareous soil in limestone cliffs, 20 Jul 1992, R.Haugan & E.Timdal YAK17/24 (O L-19086); c. 54 km N-NNW of Tyubelyakh,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="65.85">65°51'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="143.01666">143°01'E</geoCoordinate>
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, 200-300 m alt., 20 Jul 1992, M.P.Zhurbenko 92185 (O L-118686).
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</paragraph>
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<paragraph pageId="8" pageNumber="43">
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<taxonomicName class="Lecanoromycetes" family="Psoraceae" genus="Psora" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Psora tenuifolia" order="Lecanorales" pageId="8" pageNumber="43" phylum="Ascomycota" rank="species" species="tenuifolia">Psora tenuifolia</taxonomicName>
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. CHINA. Sichuan: Tibetan fringe mountains (=Hengduan Shan), Shaluli Shan, on the outskirts of Yajiang, 200 m E of the river Yalong Jiang,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="30.039444">
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30°02
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<normalizedToken originalValue="’22”">'22"</normalizedToken>
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N
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</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="101.00445">
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101°00
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<normalizedToken originalValue="’16”">'16"</normalizedToken>
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E
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</geoCoordinate>
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, 2610 m alt., NE-exposed dry slopes with schist outcrops, on thin soil crust (over schist), 12 Aug 2000, W.Obermayer 9791 (GZU); Xizang: Himalaya Range, 170 km S of Lhasa, between Lhozhag and Lhakhang Dzong, W-facing slopes of Dhalari mountain,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="28.333334">28°20'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="90.96667">90°58'E</geoCoordinate>
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, 4300 m alt., NNW-exposed,
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<normalizedToken originalValue="±">+/-</normalizedToken>
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underhang, 20 Jul 1994, W.Obermayer 4487 (GZU); Himalaya Range, 175-180 km S of Lhasa, between Lhozhag and Lhakhang Dzong, Kuru river valley, pass,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="28.2">28°12'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="91.0">91°00'E</geoCoordinate>
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, 3600 m alt., on soil, 21 Jul 1994, W.Obermayer 4525 (GZU); Himalaya Range, 210 km SE of Lhasa, 15 km ESE of
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<normalizedToken originalValue="Lhünze">Lhuenze</normalizedToken>
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, way to
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<normalizedToken originalValue="Qayü">Qayue</normalizedToken>
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, dry-valley of Subansiri,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="28.4">28°24'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="92.61667">92°37'E</geoCoordinate>
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, 4100-4200 m alt., on soil (+ mosses), 31 Jul 1994, W.Obermayer 5236 (GZU).
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</paragraph>
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<paragraph pageId="8" pageNumber="43">
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<taxonomicName class="Lecanoromycetes" family="Psoraceae" genus="Psora" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Psora vallesiaca" order="Lecanorales" pageId="8" pageNumber="43" phylum="Ascomycota" rank="species" species="vallesiaca">Psora vallesiaca</taxonomicName>
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. CHINA. Sichuan: Shalui Shan Mts, 30 km NE Batang, S Yidun,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="30.266666">30°16'N</geoCoordinate>
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,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="925" value="99.416664">99°25'E</geoCoordinate>
|
||
, 3750-3800 m alt., on marble outcrops, soil, 25 Jun 1994, W.Obermayer 3227 (GZU); Xizang: Himalaya Range, 165 km SSE of Lhasa, 40 km W of
|
||
<normalizedToken originalValue="Lhünze">Lhuenze</normalizedToken>
|
||
, little village on way to Nera Tso (=Ni La Hu),
|
||
<geoCoordinate direction="north" orientation="latitude" precision="925" value="28.383333">28°23'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="925" value="93.583336">92°95'E</geoCoordinate>
|
||
, 4300-4400 m alt., dry-valley, N-exposed dry slopes, on the ground, 1 Aug 1994, W.Obermayer 5279 (GZU); Himalaya Range, 170 km S of Lhasa, between Lhozhag and Lhakhang Dzong, W-facing slopes of Dhalari mountain,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="925" value="28.333334">28°20'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="925" value="90.96667">90°58'E</geoCoordinate>
|
||
, 4300 m alt., NNW-exposed,
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
underhang, 20 Jul 1994, W.Obermayer 4482 (GZU); PAKISTAN. Karakorum, Baltistan, Haramosh Range, between
|
||
<normalizedToken originalValue="“Alm”">"Alm"</normalizedToken>
|
||
Matumdus (3620 m,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="925" value="35.7">35°42'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="925" value="75.38333">75°23'E</geoCoordinate>
|
||
) and Hemasil in the Basna Valley, 3100-3200 m alt., 7 Jul 1991, J.Poelt K91-705 (GZU); Karakorum, Baltistan, Basna valley, Basnald, 2500 m alt., 10 Jul 1991, J.Poelt s.n. (GZU).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |