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<mods:title id="3A3DDA34AE2CF6A75F4A3CEAF1C73576">Systematic review of the Dasymutilla monticola species-group (Hymenoptera: Mutillidae): using phylogenetics to address species-group placement and sex associations</mods:title>
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<treatment id="1F2F8784F974057EFF3B9833FCEAFD60" ID-DOI="http://doi.org/10.5281/zenodo.5674215" ID-GBIF-Taxon="119555435" ID-Zenodo-Dep="5674215" LSID="urn:lsid:plazi:treatment:1F2F8784F974057EFF3B9833FCEAFD60" httpUri="http://treatment.plazi.org/id/1F2F8784F974057EFF3B9833FCEAFD60" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
<subSubSection id="DF9C6519F9740578FF3B9833FDBBF8FC" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="97393692F9740578FF3B9833FDCBF883" blockId="11.[151,630,1813,1872]" box="[151,630,1813,1839]" pageId="11" pageNumber="12">
<heading id="CC7181FEF9740578FF3B9833FDCBF883" bold="true" box="[151,630,1813,1839]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<taxonomicName id="50864D11F9740578FF3B9833FDCBF883" authority="Melander 1903" authorityName="Melander" authorityYear="1903" box="[151,630,1813,1839]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="birkmani">
<emphasis id="A5F2EA80F9740578FF3B9833FDCBF883" bold="true" box="[151,630,1813,1839]" pageId="11" pageNumber="12">
<emphasis id="A5F2EA80F9740578FF3B9833FE1DF883" bold="true" box="[151,416,1813,1839]" italics="true" pageId="11" pageNumber="12">Dasymutilla birkmani</emphasis>
(Melander 1903)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="97393692F9740578FF3B981EFDBBF8FC" blockId="11.[151,630,1813,1872]" box="[151,518,1848,1872]" pageId="11" pageNumber="12">
(
<figureCitation id="0FBD2A17F9740578FF33981EFEBEF8FC" box="[159,259,1848,1872]" captionStart="FIGURES 1 8. 1, 2" captionStartId="8.[151,264,1883,1906]" captionTargetBox="[153,1432,665,1861]" captionTargetId="figure@8.[151,1436,663,1862]" captionTargetPageId="8" captionText="FIGURES 1 8. 1, 2, Habitus, lateral view, Ƥ. 1. Dasymutilla canella; 2. D. spilota. 3, 4, Mesosoma, dorsal view, male. 3. D. saetigera; 4. D. vesta. 5 8, Head, dorsal view, female. 5. D. arenneronea (D. chattahoochei); 6. D. birkmani; 7. D. birkmani (D. corcyra); 8. D. macilenta (D. chattahoochei)." httpUri="https://zenodo.org/record/213055/files/figure.png" pageId="11" pageNumber="12">Figs 6, 7</figureCitation>
,
<figureCitation id="0FBD2A17F9740578FEA3981EFEEBF8FC" box="[271,342,1848,1872]" captionStart="FIGURES 9 23" captionStartId="10.[151,264,1950,1973]" captionTargetBox="[172,1430,183,1933]" captionTargetId="figure@10.[172,1436,183,1936]" captionTargetPageId="10" captionText="FIGURES 9 23. Habitus, dorsal view, Ƥ. 9. Dasymutilla archboldi; 10. D. arenneronea (D. chattahoochei); 11. D. birkmani; 12. D. bonita; 13. D. campanula; 14. D. canella; 15. D. chalcocephala; 16. D. birkmani (D. corcyra); 17. D. eurynome; 18. D. macilenta; 19. D. monticola; 20. D. radkei; 21. D. saetigera; 22. D. spilota; 23. D. vesta." httpUri="https://zenodo.org/record/213056/files/figure.png" pageId="11" pageNumber="12">11, 16</figureCitation>
,
<figureCitation id="0FBD2A17F9740578FECE981EFE16F8FC" box="[354,427,1848,1872]" captionStart="FIGURES 24 35. 24 35" captionStartId="11.[151,264,1660,1683]" captionTargetBox="[152,1432,196,1637]" captionTargetId="figure@11.[151,1436,193,1639]" captionTargetPageId="11" captionText="FIGURES 24 35. 24 35, Habitus, lateral view, 3. 24. Dasymutilla archboldi; 25. D. arenneronea (D. chattahoochei); 26. D. birkmani; 27. D. birkmani (D. arcana); 28. D. bonita; 29. D. canella; 30. D. fasciventris sensu Manley &amp; Pitts; 31. D. gibbosa; 32. D. macilenta; 33. D. monticola; 34. D. saetigera (D. polia); 35. D. vesta." httpUri="https://zenodo.org/record/213057/files/figure.png" pageId="11" pageNumber="12">26, 27</figureCitation>
,
<figureCitation id="0FBD2A17F9740578FE1A981EFE43F8FC" box="[438,510,1848,1872]" captionStart="FIGURES 36 71" captionStartId="13.[151,264,1887,1910]" captionTargetBox="[339,1253,777,1858]" captionTargetId="figure@13.[323,1264,772,1866]" captionTargetPageId="13" captionText="FIGURES 36 71. Male genitalia: dorsal view, ventral view, lateral view, and penial valve. 36 39. Dasymutilla archboldi; 40 43. D. arenneronea; 44 47. D. birkmani; 48 51. D. bonita; 52 55: D. canella; 56 59. D. macilenta; 60 63. D. monticola; 64 67. D. saetigera; 68 71. D. vesta." httpUri="https://zenodo.org/record/213058/files/figure.png" pageId="11" pageNumber="12">4447</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DF9C6519F974057FFF3B98A6FC5EFE84" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="97393692F9740578FF3B98A6FB8FF819" blockId="11.[151,1436,1920,2035]" pageId="11" pageNumber="12">
<taxonomicName id="50864D11F9740578FF3B98A6FE50F83A" authority="Birkmani Melander 1903" authorityName="Birkmani Melander" authorityYear="1903" box="[151,493,1920,1942]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="A5F2EA80F9740578FF3B98A6FEFAF83A" box="[151,327,1920,1942]" italics="true" pageId="11" pageNumber="12">Mutilla Birkmani</emphasis>
Melander 1903
</taxonomicName>
. Trans. Amer. Ent. Soc. 29: 313.
<typeStatus id="483D8830F9740578FCEB98A7FC11F83A" box="[839,940,1921,1942]" pageId="11" pageNumber="12" type="lectotype">Lectotype</typeStatus>
(designated by
<bibRefCitation id="F3174B63F9740578FBFD98A6FB67F83A" author="Mickel" box="[1105,1242,1920,1942]" pageId="11" pageNumber="12" refString="Mickel, C. E. (1928) Biological and taxonomic investigations on the mutillid wasps. United States Museum Bulletin, 143, 1 - 351." type="journal article" year="1928">Mickel, 1928</bibRefCitation>
by referencing the “
<typeStatus id="483D8830F9740578FF7D98B9FF42F818" box="[209,255,1951,1972]" pageId="11" pageNumber="12">type</typeStatus>
” [Art. 74.5,
<bibRefCitation id="F3174B63F9740578FED398B9FE43F819" author="ICZN" box="[383,510,1951,1973]" pageId="11" pageNumber="12" refString="International Commission on Zoological Nomenclature (ICZN). (1999) International Code of Zoological Nomenclature. Fourth Edition. ITZN, London. i - xxx + 1 - 306." type="book chapter" year="1999">ICZN, 1999</bibRefCitation>
]), Ƥ, Fedor, Lee County, Tex[as], G. Birkman (MCZC).
</paragraph>
<paragraph id="97393692F9740578FF3B9898FDCCF85E" blockId="11.[151,1436,1920,2035]" pageId="11" pageNumber="12">
<taxonomicName id="50864D11F9740578FF3B9898FE46F87F" authority="Mickel, 1928" authorityName="Mickel" authorityYear="1928" box="[151,507,1981,2004]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="corcyra">
<emphasis id="A5F2EA80F9740578FF3B9898FED5F878" box="[151,360,1982,2004]" italics="true" pageId="11" pageNumber="12">Dasymutilla corcyra</emphasis>
<bibRefCitation id="F3174B63F9740578FEDC989BFE46F87F" author="Mickel" box="[368,507,1981,2003]" pageId="11" pageNumber="12" refString="Mickel, C. E. (1928) Biological and taxonomic investigations on the mutillid wasps. United States Museum Bulletin, 143, 1 - 351." type="journal article" year="1928">Mickel, 1928</bibRefCitation>
</taxonomicName>
.
<collectingCountry id="EF917602F9740578FDAA9898FD86F87F" box="[518,571,1982,2003]" name="United States of America" pageId="11" pageNumber="12">U. S.</collectingCountry>
Nat. Mus. Bull. 143: 180.
<typeStatus id="483D8830F9740578FCFF989BFC09F87F" box="[851,948,1981,2003]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
, Ƥ, Rock Island, Tex[as], Skull Creek,
<date id="E3381052F9740578FAE7989BFF41F85E" pageId="11" pageNumber="12" value="1922-07-25">July 25, 1922</date>
, Grace O. Wiley (UMSP).,
<emphasis id="A5F2EA80F9740578FDB898FBFDCCF85E" bold="true" box="[532,625,2013,2034]" pageId="11" pageNumber="12">syn. nov.</emphasis>
</paragraph>
<paragraph id="97393692F973057FFF3B9FBEFE3BFF60" blockId="12.[151,1436,152,296]" pageId="12" pageNumber="13">
<taxonomicName id="50864D11F973057FFF3B9FBEFE4FFF01" authority="Mickel, 1928" authorityName="Mickel" authorityYear="1928" box="[151,498,152,174]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="arcana">
<emphasis id="A5F2EA80F973057FFF3B9FBEFEDCFF02" box="[151,353,152,174]" italics="true" pageId="12" pageNumber="13">Dasymutilla arcana</emphasis>
<bibRefCitation id="F3174B63F973057FFECB9FBEFE4FFF01" author="Mickel" box="[359,498,152,174]" pageId="12" pageNumber="13" refString="Mickel, C. E. (1928) Biological and taxonomic investigations on the mutillid wasps. United States Museum Bulletin, 143, 1 - 351." type="journal article" year="1928">Mickel, 1928</bibRefCitation>
</taxonomicName>
.
<collectingCountry id="EF917602F973057FFE509FBFFD8DFF01" box="[508,560,152,174]" name="United States of America" pageId="12" pageNumber="13">U. S.</collectingCountry>
Nat. Mus. Bull. 143: 217.
<typeStatus id="483D8830F973057FFCEF9FBEFC19FF02" box="[835,932,152,174]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
, 3, Lee County, Texas,
<date id="E3381052F973057FFB369FBFFAA3FF02" box="[1178,1310,152,174]" pageId="12" pageNumber="13" value="1906-06-08">June 8, 1906</date>
, J. Bequaert (SEMC),
<emphasis id="A5F2EA80F973057FFE859F91FE3BFF60" bold="true" box="[297,390,183,204]" pageId="12" pageNumber="13">syn. nov.</emphasis>
</paragraph>
<paragraph id="97393692F973057FFF3B9FF3FE99FEA6" blockId="12.[151,1436,152,296]" pageId="12" pageNumber="13">
<taxonomicName id="50864D11F973057FFF3B9FF3FE4FFF46" authority="Mickel, 1928" authorityName="Mickel" authorityYear="1928" box="[151,498,213,235]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="reclusa">
<emphasis id="A5F2EA80F973057FFF3B9FF3FEDFFF47" box="[151,354,213,235]" italics="true" pageId="12" pageNumber="13">Dasymutilla reclusa</emphasis>
<bibRefCitation id="F3174B63F973057FFEC49FF3FE4FFF46" author="Mickel" box="[360,498,213,235]" pageId="12" pageNumber="13" refString="Mickel, C. E. (1928) Biological and taxonomic investigations on the mutillid wasps. United States Museum Bulletin, 143, 1 - 351." type="journal article" year="1928">Mickel, 1928</bibRefCitation>
</taxonomicName>
.
<collectingCountry id="EF917602F973057FFE569FF0FD90FF46" box="[506,557,213,235]" name="United States of America" pageId="12" pageNumber="13">U. S.</collectingCountry>
Nat. Mus. Bull. 143: 219.
<typeStatus id="483D8830F973057FFC959FF3FC27FF47" box="[825,922,213,235]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
, 3, Halsey, Nebr.,
<date id="E3381052F973057FFBFB9FF3FABCFF46" box="[1111,1281,213,234]" pageId="12" pageNumber="13" value="1924-08-29">August 29, 1924</date>
, R. W. Dawson (UMSP).
</paragraph>
<paragraph id="97393692F973057FFF3B9E34FC5EFE84" blockId="12.[151,1436,152,296]" box="[151,995,274,296]" pageId="12" pageNumber="13">
<taxonomicName id="50864D11F973057FFF3B9E34FE5DFE84" authority="Melander" authorityName="Melander" box="[151,480,274,296]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="birkmani">
<emphasis id="A5F2EA80F973057FFF3B9E34FEC9FE84" box="[151,372,274,296]" italics="true" pageId="12" pageNumber="13">Dasymutilla birkmani</emphasis>
Melander
</taxonomicName>
:
<bibRefCitation id="F3174B63F973057FFE459E34FD17FE84" author="Pilgrim" box="[489,682,274,296]" pageId="12" pageNumber="13" refString="Pilgrim, E. M., Williams, K. A. &amp; Pitts, J. P. (2008) Sex association and synonymy in Southwestern U. S. species of Dasymutilla (Hymenoptera: Mutillidae). Pan-Pacific Entomologist, 84, 58 - 69." type="journal article" year="2008">
Pilgrim
<emphasis id="A5F2EA80F973057FFD979E35FDDAFE84" box="[571,615,274,296]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
. 2008
</bibRefCitation>
. Pan-Pacific Ent., 84: 64,
<specimenCount id="8180FD1BF973057FFC1D9E35FC79FE84" box="[945,964,275,296]" pageId="12" pageNumber="13" type="generic">3</specimenCount>
Ƥ.
</paragraph>
</subSubSection>
<subSubSection id="DF9C6519F973057FFF3B9E73FCB2FD1D" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="97393692F973057FFF3B9E73FCB2FD1D" blockId="12.[151,1436,341,2021]" pageId="12" pageNumber="13">
<emphasis id="A5F2EA80F973057FFF3B9E73FEACFEC2" bold="true" box="[151,273,341,366]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
FEMALE. This
<taxonomicName id="50864D11F973057FFE4D9E73FD8FFEC1" box="[481,562,341,365]" class="Insecta" family="Scoliidae" genus="Scolia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species">species</taxonomicName>
varies greatly and can be difficult to separate from
<taxonomicName id="50864D11F973057FFB039E70FAF5FEC1" box="[1199,1352,341,365]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="A5F2EA80F973057FFB039E70FAF5FEC1" box="[1199,1352,341,365]" italics="true" pageId="12" pageNumber="13">D. monticola</emphasis>
</taxonomicName>
. In all examined specimens, the first flagellar segment is only slightly longer than the second and the propodeum has a thick brush of silver or golden setae (
<figureCitation id="0FBD2A17F973057FFDE29EBBFD68FE19" box="[590,725,413,437]" captionStart="FIGURES 9 23" captionStartId="10.[151,264,1950,1973]" captionTargetBox="[172,1430,183,1933]" captionTargetId="figure@10.[172,1436,183,1936]" captionTargetPageId="10" captionText="FIGURES 9 23. Habitus, dorsal view, Ƥ. 9. Dasymutilla archboldi; 10. D. arenneronea (D. chattahoochei); 11. D. birkmani; 12. D. bonita; 13. D. campanula; 14. D. canella; 15. D. chalcocephala; 16. D. birkmani (D. corcyra); 17. D. eurynome; 18. D. macilenta; 19. D. monticola; 20. D. radkei; 21. D. saetigera; 22. D. spilota; 23. D. vesta." httpUri="https://zenodo.org/record/213056/files/figure.png" pageId="12" pageNumber="13">Figs 11, 16</figureCitation>
). Most specimens have the anterior portion of the mesosoma clothed with dark brown or black setae dorsally (
<figureCitation id="0FBD2A17F973057FFD129EE7FCFDFE75" box="[702,832,449,473]" captionStart="FIGURES 9 23" captionStartId="10.[151,264,1950,1973]" captionTargetBox="[172,1430,183,1933]" captionTargetId="figure@10.[172,1436,183,1936]" captionTargetPageId="10" captionText="FIGURES 9 23. Habitus, dorsal view, Ƥ. 9. Dasymutilla archboldi; 10. D. arenneronea (D. chattahoochei); 11. D. birkmani; 12. D. bonita; 13. D. campanula; 14. D. canella; 15. D. chalcocephala; 16. D. birkmani (D. corcyra); 17. D. eurynome; 18. D. macilenta; 19. D. monticola; 20. D. radkei; 21. D. saetigera; 22. D. spilota; 23. D. vesta." httpUri="https://zenodo.org/record/213056/files/figure.png" pageId="12" pageNumber="13">Figs 11, 16</figureCitation>
); these differ from
<taxonomicName id="50864D11F973057FFBB59EE4FB11FE75" box="[1049,1196,449,473]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="A5F2EA80F973057FFBB59EE4FB11FE75" box="[1049,1196,449,473]" italics="true" pageId="12" pageNumber="13">D. monticola</emphasis>
</taxonomicName>
, which has the entire mesosoma clothed with golden setae (
<figureCitation id="0FBD2A17F973057FFDF99EC3FD10FE51" box="[597,685,485,509]" captionStart="FIGURES 9 23" captionStartId="10.[151,264,1950,1973]" captionTargetBox="[172,1430,183,1933]" captionTargetId="figure@10.[172,1436,183,1936]" captionTargetPageId="10" captionText="FIGURES 9 23. Habitus, dorsal view, Ƥ. 9. Dasymutilla archboldi; 10. D. arenneronea (D. chattahoochei); 11. D. birkmani; 12. D. bonita; 13. D. campanula; 14. D. canella; 15. D. chalcocephala; 16. D. birkmani (D. corcyra); 17. D. eurynome; 18. D. macilenta; 19. D. monticola; 20. D. radkei; 21. D. saetigera; 22. D. spilota; 23. D. vesta." httpUri="https://zenodo.org/record/213056/files/figure.png" pageId="12" pageNumber="13">Fig. 19</figureCitation>
). In the few specimens of
<taxonomicName id="50864D11F973057FFC409EC0FBC5FE51" box="[1004,1144,485,509]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="birkmani">
<emphasis id="A5F2EA80F973057FFC409EC0FBC5FE51" box="[1004,1144,485,509]" italics="true" pageId="12" pageNumber="13">D. birkmani</emphasis>
</taxonomicName>
that have the mesosoma clothed entirely with golden setae, the apical fringe of T2 is broadly silver with only a slight spot of black setae medially (as in
<figureCitation id="0FBD2A17F973057FFEE49D0BFE01FDE9" box="[328,444,557,581]" captionStart="FIGURES 9 23" captionStartId="10.[151,264,1950,1973]" captionTargetBox="[172,1430,183,1933]" captionTargetId="figure@10.[172,1436,183,1936]" captionTargetPageId="10" captionText="FIGURES 9 23. Habitus, dorsal view, Ƥ. 9. Dasymutilla archboldi; 10. D. arenneronea (D. chattahoochei); 11. D. birkmani; 12. D. bonita; 13. D. campanula; 14. D. canella; 15. D. chalcocephala; 16. D. birkmani (D. corcyra); 17. D. eurynome; 18. D. macilenta; 19. D. monticola; 20. D. radkei; 21. D. saetigera; 22. D. spilota; 23. D. vesta." httpUri="https://zenodo.org/record/213056/files/figure.png" pageId="12" pageNumber="13">Figs 911</figureCitation>
);
<taxonomicName id="50864D11F973057FFE7D9D08FDD8FDE9" box="[465,613,557,581]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="A5F2EA80F973057FFE7D9D08FDD8FDE9" box="[465,613,557,581]" italics="true" pageId="12" pageNumber="13">D. monticola</emphasis>
</taxonomicName>
has the fringe of T2 clothed mostly with black setae medially (
<figureCitation id="0FBD2A17F973057FFA969D0BFA32FDE9" box="[1338,1423,557,581]" captionStart="FIGURES 9 23" captionStartId="10.[151,264,1950,1973]" captionTargetBox="[172,1430,183,1933]" captionTargetId="figure@10.[172,1436,183,1936]" captionTargetPageId="10" captionText="FIGURES 9 23. Habitus, dorsal view, Ƥ. 9. Dasymutilla archboldi; 10. D. arenneronea (D. chattahoochei); 11. D. birkmani; 12. D. bonita; 13. D. campanula; 14. D. canella; 15. D. chalcocephala; 16. D. birkmani (D. corcyra); 17. D. eurynome; 18. D. macilenta; 19. D. monticola; 20. D. radkei; 21. D. saetigera; 22. D. spilota; 23. D. vesta." httpUri="https://zenodo.org/record/213056/files/figure.png" pageId="12" pageNumber="13">Fig. 19</figureCitation>
). MALE. The male of this
<taxonomicName id="50864D11F973057FFE629D77FDA2FDC5" box="[462,543,593,617]" class="Insecta" family="Scoliidae" genus="Scolia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species">species</taxonomicName>
can be separated from other
<taxonomicName id="50864D11F973057FFC299D77FBB3FDC5" box="[901,1038,593,617]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="A5F2EA80F973057FFC299D77FBB3FDC5" box="[901,1038,593,617]" italics="true" pageId="12" pageNumber="13">Dasymutilla</emphasis>
</taxonomicName>
by the following combination of characters: there is no seta-filled pit on S2, there are scattered silver or gray setae on the apical terga, and the cuspis is more than 0.8x the free paramere length (
<figureCitation id="0FBD2A17F973057FFD2C9DBFFCBFFD1D" box="[640,770,665,689]" captionStart="FIGURES 36 71" captionStartId="13.[151,264,1887,1910]" captionTargetBox="[339,1253,777,1858]" captionTargetId="figure@13.[323,1264,772,1866]" captionTargetPageId="13" captionText="FIGURES 36 71. Male genitalia: dorsal view, ventral view, lateral view, and penial valve. 36 39. Dasymutilla archboldi; 40 43. D. arenneronea; 44 47. D. birkmani; 48 51. D. bonita; 52 55: D. canella; 56 59. D. macilenta; 60 63. D. monticola; 64 67. D. saetigera; 68 71. D. vesta." httpUri="https://zenodo.org/record/213058/files/figure.png" pageId="12" pageNumber="13">Figs 44, 45</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DF9C6519F973057FFF6B9D9BFE3EFBB5" pageId="12" pageNumber="13" type="description">
<paragraph id="97393692F973057FFF6B9D9BFD00FC7D" blockId="12.[151,1436,341,2021]" pageId="12" pageNumber="13">
<emphasis id="A5F2EA80F973057FFF6B9D9BFD9BFD7A" bold="true" box="[199,550,701,726]" pageId="12" pageNumber="13">Description of male genitalia</emphasis>
(
<figureCitation id="0FBD2A17F973057FFD9B9D9BFD00FD79" box="[567,701,701,725]" captionStart="FIGURES 36 71" captionStartId="13.[151,264,1887,1910]" captionTargetBox="[339,1253,777,1858]" captionTargetId="figure@13.[323,1264,772,1866]" captionTargetPageId="13" captionText="FIGURES 36 71. Male genitalia: dorsal view, ventral view, lateral view, and penial valve. 36 39. Dasymutilla archboldi; 40 43. D. arenneronea; 44 47. D. birkmani; 48 51. D. bonita; 52 55: D. canella; 56 59. D. macilenta; 60 63. D. monticola; 64 67. D. saetigera; 68 71. D. vesta." httpUri="https://zenodo.org/record/213058/files/figure.png" pageId="12" pageNumber="13">Figs 4447</figureCitation>
). Free length of paramere dorsally curved apically, with ventral brush of long dense setae basally, remainder having scattered short sparse setae; paramere laterally kinked in basal 0.2, apices noticeably divergent. Cuspis slightly laterally compressed, tapering apically, ~0.85 × free-length of paramere, having sparse short setae throughout, except outer surface having longer denser setae; outer row of setae anteriorly directed, inner row posteriorly directed. Densely setose basal lobe present. Digitus laterally curved internally, tapering slightly at apex, asetose, ~0.4 × free-length of paramere. Penial valve emarginated anterodorsally, ventral margin bidentate apically, teeth separated, unidentate medially; having longitudinal row of setae at apex and subapically on external margin.
</paragraph>
<paragraph id="97393692F973057FFF6B9CFBFD53FC59" blockId="12.[151,1436,341,2021]" box="[199,750,989,1013]" pageId="12" pageNumber="13">
<emphasis id="A5F2EA80F973057FFF6B9CFBFE99FC59" bold="true" box="[199,292,989,1013]" pageId="12" pageNumber="13">Length.</emphasis>
Females: 3.5
<quantity id="507E9B77F973057FFE649CFBFDB6FC59" box="[456,523,989,1013]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="13" unit="mm" value="9.0">9 mm</quantity>
; males: 5.5
<quantity id="507E9B77F973057FFD359CFBFD57FC59" box="[665,746,989,1013]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="13" unit="mm" value="10.0">10 mm</quantity>
.
</paragraph>
<paragraph id="97393692F973057FFF6B9B24FE3EFBB5" blockId="12.[151,1436,341,2021]" box="[199,387,1025,1050]" pageId="12" pageNumber="13">
<emphasis id="A5F2EA80F973057FFF6B9B24FEBBFBB6" bold="true" box="[199,262,1026,1050]" pageId="12" pageNumber="13">Host.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="DF9C6519F973057FFF6B9B03FBE1FB91" box="[199,1116,1061,1086]" pageId="12" pageNumber="13" type="distribution">
<paragraph id="97393692F973057FFF6B9B03FBE1FB91" blockId="12.[151,1436,341,2021]" box="[199,1116,1061,1086]" pageId="12" pageNumber="13">
<emphasis id="A5F2EA80F973057FFF6B9B03FEE4FB92" bold="true" box="[199,345,1061,1086]" pageId="12" pageNumber="13">Distribution</emphasis>
. This
<taxonomicName id="50864D11F973057FFE0C9B03FE4CFB91" box="[416,497,1061,1085]" class="Insecta" family="Scoliidae" genus="Scolia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species">species</taxonomicName>
is widespread in the central and western
<collectingCountry id="EF917602F973057FFC129B03FBEAFB91" box="[958,1111,1061,1085]" name="United States of America" pageId="12" pageNumber="13">United States</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="DF9C6519F973057FFF6B9B6FFACCF8F9" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="97393692F973057FFF6B9B6FFACCF8F9" blockId="12.[151,1436,341,2021]" pageId="12" pageNumber="13">
<emphasis id="A5F2EA80F973057FFF6B9B6FFE10FBCE" bold="true" box="[199,429,1097,1122]" pageId="12" pageNumber="13">Material examined</emphasis>
.
<typeStatus id="483D8830F973057FFE6D9B6CFE4BFBCE" box="[449,502,1098,1122]" pageId="12" pageNumber="13">Type</typeStatus>
material.
<emphasis id="A5F2EA80F973057FFDD89B6FFD5BFBCE" bold="true" box="[628,742,1097,1122]" italics="true" pageId="12" pageNumber="13">
<typeStatus id="483D8830F973057FFDD89B6FFD5BFBCE" box="[628,742,1097,1122]" pageId="12" pageNumber="13" type="holotype">Holotypes</typeStatus>
</emphasis>
:
<taxonomicName id="50864D11F973057FFD579B6FFC58FBCD" box="[763,997,1097,1121]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="corcyra">
<emphasis id="A5F2EA80F973057FFD579B6FFC58FBCD" box="[763,997,1097,1121]" italics="true" pageId="12" pageNumber="13">Dasymutilla corcyra</emphasis>
</taxonomicName>
,
<collectingCountry id="EF917602F973057FFC559B6FFB88FBCD" box="[1017,1077,1097,1121]" name="United States of America" pageId="12" pageNumber="13">USA</collectingCountry>
:
<emphasis id="A5F2EA80F973057FFBE99B6CFB3BFBCE" bold="true" box="[1093,1158,1098,1122]" pageId="12" pageNumber="13">Texas</emphasis>
, [
<emphasis id="A5F2EA80F973057FFB099B6FFAF9FBCD" box="[1189,1348,1097,1121]" italics="true" pageId="12" pageNumber="13">Colorado Co.</emphasis>
], Rock Island, Skull Creek, 1Ƥ,
<date id="E3381052F973057FFE699B4BFDEDFB29" box="[453,592,1133,1157]" pageId="12" pageNumber="13" value="1922-07-25">25.VII.1922</date>
, G.O. Wiley (
<collectionCode id="F197AE57F973057FFD569B4BFCF1FB29" box="[762,844,1133,1157]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="12" pageNumber="13">UMSP</collectionCode>
);
<taxonomicName id="50864D11F973057FFCC99B4BFBF5FB29" box="[869,1096,1133,1157]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="arcana">
<emphasis id="A5F2EA80F973057FFCC99B4BFBF5FB29" box="[869,1096,1133,1157]" italics="true" pageId="12" pageNumber="13">Dasymutilla arcana</emphasis>
</taxonomicName>
,
<collectingCountry id="EF917602F973057FFBF19B4BFB24FB29" box="[1117,1177,1133,1157]" name="United States of America" pageId="12" pageNumber="13">USA</collectingCountry>
:
<emphasis id="A5F2EA80F973057FFB069B4BFB57FB29" bold="true" box="[1194,1258,1133,1157]" pageId="12" pageNumber="13">Texas</emphasis>
,
<emphasis id="A5F2EA80F973057FFAAD9B48FAE3FB29" box="[1281,1374,1134,1157]" italics="true" pageId="12" pageNumber="13">Lee Co.</emphasis>
, 13,
<date id="E3381052F973057FFF3B9BB7FF19FB05" box="[151,164,1169,1193]" pageId="12" pageNumber="13" value="1906-06-08">8</date>
.VI.1906, J. Bequaert (
<collectionCode id="F197AE57F973057FFE0B9BB7FE44FB05" box="[423,505,1169,1193]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="13">SEMC</collectionCode>
);
<taxonomicName id="50864D11F973057FFDA29BB7FD4CFB05" box="[526,753,1169,1193]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="reclusa">
<emphasis id="A5F2EA80F973057FFDA29BB7FD4CFB05" box="[526,753,1169,1193]" italics="true" pageId="12" pageNumber="13">Dasymutilla reclusa</emphasis>
</taxonomicName>
,
<collectingCountry id="EF917602F973057FFCAC9BB7FC81FB05" box="[768,828,1169,1193]" name="United States of America" pageId="12" pageNumber="13">USA</collectingCountry>
:
<emphasis id="A5F2EA80F973057FFCE59BB4FC06FB06" bold="true" box="[841,955,1170,1194]" pageId="12" pageNumber="13">Nebraska</emphasis>
, [
<emphasis id="A5F2EA80F973057FFC789BB7FBE0FB05" box="[980,1117,1169,1193]" italics="true" pageId="12" pageNumber="13">Thomas Co.</emphasis>
], Halsey, 13,
<date id="E3381052F973057FFAAF9BB7FA9DFB05" box="[1283,1312,1169,1193]" pageId="12" pageNumber="13" value="1924-08-29">29</date>
.VIII.1924, R.W. Dawson (
<collectionCode id="F197AE57F973057FFEE49B93FE27FB61" box="[328,410,1205,1229]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="12" pageNumber="13">UMSP</collectionCode>
);
<emphasis id="A5F2EA80F973057FFE1D9B90FD94FB61" bold="true" box="[433,553,1206,1229]" italics="true" pageId="12" pageNumber="13">
<typeStatus id="483D8830F973057FFE1D9B90FD94FB61" box="[433,553,1206,1229]" pageId="12" pageNumber="13" type="lectotype">Lectotypes</typeStatus>
</emphasis>
:
<taxonomicName id="50864D11F973057FFD909B93FD42FB61" authority="Birkmani" authorityName="Birkmani" box="[572,767,1205,1229]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="A5F2EA80F973057FFD909B93FD42FB61" box="[572,767,1205,1229]" italics="true" pageId="12" pageNumber="13">Mutilla Birkmani</emphasis>
</taxonomicName>
,
<collectingCountry id="EF917602F973057FFCBE9B93FCF1FB61" box="[786,844,1205,1229]" name="United States of America" pageId="12" pageNumber="13">USA</collectingCountry>
,
<emphasis id="A5F2EA80F973057FFCF79B93FC21FB61" bold="true" box="[859,924,1205,1229]" pageId="12" pageNumber="13">Texas</emphasis>
, Lee Co., Fedor, 1Ƥ, G. Birkman (
<collectionCode id="F197AE57F973057FFA969B93FA32FB61" box="[1338,1423,1205,1229]" pageId="12" pageNumber="13">MCZC</collectionCode>
);
<emphasis id="A5F2EA80F973057FFF3B9BFFFE83FB5E" bold="true" box="[151,318,1241,1266]" italics="true" pageId="12" pageNumber="13">
<typeStatus id="483D8830F973057FFF3B9BFFFE83FB5E" box="[151,318,1241,1266]" pageId="12" pageNumber="13" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
,
<taxonomicName id="50864D11F973057FFEE29BFFFDADFB5D" authority="Birkmani" authorityName="Birkmani" box="[334,528,1241,1265]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="A5F2EA80F973057FFEE29BFFFDADFB5D" box="[334,528,1241,1265]" italics="true" pageId="12" pageNumber="13">Mutilla Birkmani</emphasis>
</taxonomicName>
,
<collectingCountry id="EF917602F973057FFD8D9BFFFDE1FB5D" box="[545,604,1241,1265]" name="United States of America" pageId="12" pageNumber="13">USA</collectingCountry>
,
<emphasis id="A5F2EA80F973057FFDC59BFCFD17FB5E" bold="true" box="[617,682,1242,1266]" pageId="12" pageNumber="13">Texas</emphasis>
, Lee Co., Fedor, 4Ƥ, G. Birkman (
<collectionCode id="F197AE57F973057FFB939BFFFB29FB5D" box="[1087,1172,1241,1265]" pageId="12" pageNumber="13">MCZC</collectionCode>
). Other material.
<collectingCountry id="EF917602F973057FFAF19BFFFA25FB5D" box="[1373,1432,1241,1265]" name="United States of America" pageId="12" pageNumber="13">USA</collectingCountry>
,
<emphasis id="A5F2EA80F973057FFF3B9BDBFEBBFABA" bold="true" box="[151,262,1277,1302]" pageId="12" pageNumber="13">Colorado</emphasis>
:
<emphasis id="A5F2EA80F973057FFEBA9BDBFE1AFAB9" box="[278,423,1277,1301]" italics="true" pageId="12" pageNumber="13">Sedgwick Co</emphasis>
., 2.5 km S Julesburg,
<date id="E3381052F973057FFD099BDBFCDFFAB9" box="[677,866,1277,1301]" pageId="12" pageNumber="13" value="2001-08-20" valueMax="2001-08-27" valueMin="2001-08-20">2027.VIII.2001</date>
, M.E. Irwin et al. (
<taxonomicName id="50864D11F973057FFB979BD8FB0EFAB9" box="[1083,1203,1278,1301]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="corcyra">
<emphasis id="A5F2EA80F973057FFB979BD8FB0EFAB9" box="[1083,1203,1278,1301]" italics="true" pageId="12" pageNumber="13">D. corcyra</emphasis>
</taxonomicName>
,
<collectionCode id="F197AE57F973057FFB6E9BDBFAA9FAB9" box="[1218,1300,1277,1301]" pageId="12" pageNumber="13">EMUS</collectionCode>
);
<emphasis id="A5F2EA80F973057FFA859BD8FA36FAB9" box="[1321,1419,1278,1301]" italics="true" pageId="12" pageNumber="13">Yuma Co</emphasis>
., Bonny State Park, 3Ƥ53,
<date id="E3381052F973057FFE629A07FE51FA95" box="[462,492,1313,1337]" pageId="12" pageNumber="13" value="2011-06-22">22</date>
.VI.2011, K.A. Williams (
<collectionCode id="F197AE57F973057FFCB59A07FCD1FA95" box="[793,876,1313,1337]" pageId="12" pageNumber="13">EMUS</collectionCode>
);
<emphasis id="A5F2EA80F973057FFC289A04FC66FA96" bold="true" box="[900,987,1314,1338]" pageId="12" pageNumber="13">Kansas</emphasis>
:
<emphasis id="A5F2EA80F973057FFC439A07FB3AFA95" box="[1007,1159,1313,1337]" italics="true" pageId="12" pageNumber="13">Ellsworth Co</emphasis>
., Kanopolis State Park, 2Ƥ43,
<date id="E3381052F973057FFF5C9A63FEB3FAF1" box="[240,270,1349,1373]" pageId="12" pageNumber="13" value="2011-06-22">22</date>
.VI.2011, K.A. Williams (
<collectionCode id="F197AE57F973057FFD919A63FD2DFAF1" box="[573,656,1349,1373]" pageId="12" pageNumber="13">EMUS</collectionCode>
);
<emphasis id="A5F2EA80F973057FFD069A60FC8CFAF1" box="[682,817,1350,1373]" italics="true" pageId="12" pageNumber="13">Morton Co.</emphasis>
, Cimarron National Grassland, 1Ƥ,
<date id="E3381052F973057FFB4D9A63FAE9FAF1" box="[1249,1364,1349,1373]" pageId="12" pageNumber="13" value="2009-06-07">7.VI.2009</date>
, K.A. Williams (
<collectionCode id="F197AE57F973057FFEBF9A4FFEDBFA2D" box="[275,358,1385,1409]" pageId="12" pageNumber="13">EMUS</collectionCode>
);
<emphasis id="A5F2EA80F973057FFE2D9A4FFDA1FA2D" box="[385,540,1385,1409]" italics="true" pageId="12" pageNumber="13">Sheridan Co.</emphasis>
, Sheridan Lake, 13, J.M. Carpenter (
<collectionCode id="F197AE57F973057FFC429A4FFBF9FA2D" box="[1006,1092,1385,1409]" pageId="12" pageNumber="13">MCZC</collectionCode>
);
<emphasis id="A5F2EA80F973057FFBF39A4FFB5BFA2D" box="[1119,1254,1385,1409]" italics="true" pageId="12" pageNumber="13">Wallace Co</emphasis>
., Wallace, 33,
<date id="E3381052F973057FFF3B9AABFF18FA09" box="[151,165,1421,1445]" pageId="12" pageNumber="13" value="1933-07-02">2</date>
.VII.1933 (
<collectionCode id="F197AE57F973057FFE889AABFEC8FA09" box="[292,373,1421,1445]" pageId="12" pageNumber="13">EMUS</collectionCode>
,
<collectionCode id="F197AE57F973057FFE289AABFE67FA09" LSID="urn:lsid:biocol.org:col:34868" box="[388,474,1421,1445]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="12" pageNumber="13">UNSM</collectionCode>
);
<emphasis id="A5F2EA80F973057FFE5D9AABFDDEFA09" bold="true" box="[497,611,1421,1445]" pageId="12" pageNumber="13">Nebraska</emphasis>
:
<emphasis id="A5F2EA80F973057FFDD99AABFD59FA09" box="[629,740,1421,1445]" italics="true" pageId="12" pageNumber="13">Keith Co.</emphasis>
: Lake McConaughy, West Arthur Bay,
<date id="E3381052F973057FFB199AABFA8BFA09" box="[1205,1334,1421,1445]" pageId="12" pageNumber="13" value="2003-06-15">15.VI.2003</date>
, R. &amp; J. Matthews (
<collectionCode id="F197AE57F973057FFEB89A97FEDAFA65" box="[276,359,1457,1481]" pageId="12" pageNumber="13">EMUS</collectionCode>
);
<quantity id="507E9B77F973057FFED79A97FE10FA65" box="[379,429,1457,1481]" metricMagnitude="3" metricUnit="m" metricValue="9.656064" pageId="12" pageNumber="13" unit="mi" value="6.0">6 mi</quantity>
N Paxton, 23,
<date id="E3381052F973057FFDF09A97FDD4FA65" box="[604,617,1457,1481]" pageId="12" pageNumber="13" value="1972-06-05">5</date>
.VI.1972, R.M. Miller (Iowa State);
<emphasis id="A5F2EA80F973057FFBAC9A94FBFCFA66" bold="true" box="[1024,1089,1458,1482]" pageId="12" pageNumber="13">Texas</emphasis>
:
<emphasis id="A5F2EA80F973057FFBFE9A97FB75FA65" box="[1106,1224,1457,1481]" italics="true" pageId="12" pageNumber="13">Austin Co.</emphasis>
, Stephen F. Austin State Park, Sealy, 33,
<date id="E3381052F973057FFE3E9AF3FE13FA41" box="[402,430,1493,1517]" pageId="12" pageNumber="13" value="1968-05-14">14</date>
.V.1968, H. Evans &amp; O. Flint (
<taxonomicName id="50864D11F973057FFCA99AF0FCC8FA41" box="[773,885,1494,1517]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="arcana">
<emphasis id="A5F2EA80F973057FFCA99AF0FCC8FA41" box="[773,885,1494,1517]" italics="true" pageId="12" pageNumber="13">D. arcana</emphasis>
</taxonomicName>
,
<collectionCode id="F197AE57F973057FFC2F9AF3FC6AFA41" box="[899,983,1493,1517]" pageId="12" pageNumber="13">MCZC</collectionCode>
);
<emphasis id="A5F2EA80F973057FFC479AF0FBDDFA41" box="[1003,1120,1494,1517]" italics="true" pageId="12" pageNumber="13">Brazos Co</emphasis>
., Lick Creek Park,
<quantity id="507E9B77F973057FFA979AF3FACEFA41" box="[1339,1395,1493,1517]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="13" unit="km" value="6.0">6 km</quantity>
SE College Station, 3Ƥ,
<date id="E3381052F973057FFE2C9ADFFDBDF9BD" box="[384,512,1529,1553]" pageId="12" pageNumber="13" value="2009-06-11">11.VI.2009</date>
, K.A. Williams (
<collectionCode id="F197AE57F973057FFD169ADFFCB0F9BD" box="[698,781,1529,1553]" pageId="12" pageNumber="13">EMUS</collectionCode>
);
<emphasis id="A5F2EA80F973057FFC8D9ADFFC03F9BD" box="[801,958,1529,1553]" italics="true" pageId="12" pageNumber="13">Comanche Co</emphasis>
., Comanche, 13,
<date id="E3381052F973057FFB249ADFFB1BF9BD" box="[1160,1190,1529,1553]" pageId="12" pageNumber="13" value="1953-07-26">26</date>
.VII.1953, Evans, Lin, &amp; Yoshimoto (
<collectionCode id="F197AE57F973057FFEEC993BFE28F999" box="[320,405,1565,1589]" pageId="12" pageNumber="13">MCZC</collectionCode>
);
<emphasis id="A5F2EA80F973057FFE069938FDBEF999" box="[426,515,1566,1589]" italics="true" pageId="12" pageNumber="13">Lee Co.</emphasis>
, Fedor, 13,
<date id="E3381052F973057FFD3A993BFD0FF999" box="[662,690,1565,1589]" pageId="12" pageNumber="13" value="1909-06-17">17</date>
.VI.1909, Birkman (
<collectionCode id="F197AE57F973057FFC34993BFC50F999" box="[920,1005,1565,1589]" pageId="12" pageNumber="13">MCZC</collectionCode>
);
<emphasis id="A5F2EA80F973057FFBAF9938FB1DF999" box="[1027,1184,1565,1589]" italics="true" pageId="12" pageNumber="13">Jim Wells Co.</emphasis>
,
<quantity id="507E9B77F973057FFB1C993BFB59F999" box="[1200,1252,1565,1589]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="12" pageNumber="13" unit="mi" value="8.0">8 mi</quantity>
W BenBolt, La Copita Research Station, 13,
<date id="E3381052F973057FFE4E9967FDBDF9F5" box="[482,512,1601,1625]" pageId="12" pageNumber="13" value="1987-05-20">20</date>
.V.1987, J.B. Woolley (
<taxonomicName id="50864D11F973057FFCAD9964FCCCF9F5" box="[769,881,1602,1625]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="arcana">
<emphasis id="A5F2EA80F973057FFCAD9964FCCCF9F5" box="[769,881,1602,1625]" italics="true" pageId="12" pageNumber="13">D. arcana</emphasis>
</taxonomicName>
,
<collectionCode id="F197AE57F973057FFCD39967FC69F9F5" box="[895,980,1601,1625]" pageId="12" pageNumber="13">TAMU</collectionCode>
);
<emphasis id="A5F2EA80F973057FFC4A9967FBD0F9F5" box="[998,1133,1601,1625]" italics="true" pageId="12" pageNumber="13">Kleberg Co.</emphasis>
,
<quantity id="507E9B77F973057FFBD79967FB7FF9F5" box="[1147,1218,1601,1625]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="12" pageNumber="13" unit="km" value="32.0">32 km</quantity>
SE Kingsville: 13,
<date id="E3381052F973057FFF3B9943FF09F9D1" box="[151,180,1637,1661]" pageId="12" pageNumber="13" value="1985-04-30">30</date>
.IV.1985, W.J. Pulawski (
<collectionCode id="F197AE57F973057FFE7A9943FD94F9D1" box="[470,553,1637,1661]" pageId="12" pageNumber="13">EMUS</collectionCode>
); 13,
<date id="E3381052F973057FFDD89943FD3CF9D1" box="[628,641,1637,1661]" pageId="12" pageNumber="13" value="1985-05-03">3</date>
.V.1985, W.J. Pulawski (
<taxonomicName id="50864D11F973057FFC329940FBACF9D1" box="[926,1041,1638,1661]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="arcana">
<emphasis id="A5F2EA80F973057FFC329940FBACF9D1" box="[926,1041,1638,1661]" italics="true" pageId="12" pageNumber="13">D. arcana</emphasis>
</taxonomicName>
,
<collectionCode id="F197AE57F973057FFB8F9943FBC8F9D1" box="[1059,1141,1637,1661]" pageId="12" pageNumber="13">EMUS</collectionCode>
); La Salle Co., Chaparral Wildlife Management Area, 1Ƥ,
<date id="E3381052F973057FFDAA99AFFD19F90D" box="[518,676,1673,1697]" pageId="12" pageNumber="13" value="2006-05-19" valueMax="2006-05-21" valueMin="2006-05-19">1921.V.2006</date>
, J.S. Wilson &amp; K.A. Williams (
<collectionCode id="F197AE57F973057FFBAD99AFFBE9F90D" box="[1025,1108,1673,1697]" pageId="12" pageNumber="13">EMUS</collectionCode>
);
<emphasis id="A5F2EA80F973057FFBC499AFFB4CF90D" box="[1128,1265,1673,1697]" italics="true" pageId="12" pageNumber="13">Madison Co</emphasis>
., Trinity River,
<quantity id="507E9B77F973057FFF3B9988FF6DF969" box="[151,208,1709,1733]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="13" unit="km" value="7.0">7 km</quantity>
E
<collectingCountry id="EF917602F973057FFF43998BFEF0F969" box="[239,333,1709,1733]" name="United States Pacific Island Wildlife Refuges" pageId="12" pageNumber="13">Midway</collectingCountry>
, 11Ƥ,
<date id="E3381052F973057FFE3B998BFD97F969" box="[407,554,1709,1733]" pageId="12" pageNumber="13" value="2008-08-23">23.VIII.2008</date>
, T.R. Clifton (
<collectionCode id="F197AE57F973057FFD67998BFCA3F969" box="[715,798,1709,1733]" pageId="12" pageNumber="13">EMUS</collectionCode>
);
<emphasis id="A5F2EA80F973057FFC9E9988FC1AF969" box="[818,935,1710,1733]" italics="true" pageId="12" pageNumber="13">Potter Co.</emphasis>
, Red River on Hwy
<quantity id="507E9B77F973057FFB23998BFAB3F969" box="[1167,1294,1709,1733]" metricMagnitude="5" metricUnit="m" metricValue="2.8725" pageId="12" pageNumber="13" unit="km" value="287.25">287, 25 km</quantity>
N Amarillo, 3Ƥ13,
<date id="E3381052F973057FFF4099F7FF47F945" box="[236,250,1745,1769]" pageId="12" pageNumber="13" value="2009-06-07">7</date>
.VI.2009, K.A. Williams (
<collectionCode id="F197AE57F973057FFD8E99F7FDC8F945" box="[546,629,1745,1769]" pageId="12" pageNumber="13">EMUS</collectionCode>
);
<emphasis id="A5F2EA80F973057FFD2799F4FCFFF945" box="[651,834,1746,1769]" italics="true" pageId="12" pageNumber="13">San Patricio Co</emphasis>
., Arkansas River,
<quantity id="507E9B77F973057FFBBB99F7FBDDF945" box="[1047,1120,1745,1769]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="13" unit="km" value="16.0">16 km</quantity>
NE Sinton, 1Ƥ,
<date id="E3381052F973057FFA8F99F7FA2AF945" box="[1315,1431,1745,1769]" pageId="12" pageNumber="13" value="1985-05-12">12.V.1985</date>
, W.J. Pulawski (
<collectionCode id="F197AE57F973057FFEE399D3FE1FF8A1" box="[335,418,1781,1805]" pageId="12" pageNumber="13">EMUS</collectionCode>
); Somervell Co., Dinosaur Valley State Park, 1Ƥ,
<date id="E3381052F973057FFC5299D3FBCFF8A1" box="[1022,1138,1781,1805]" pageId="12" pageNumber="13" value="1985-05-17">17.V.1985</date>
, W.J. Pulawski (
<collectionCode id="F197AE57F973057FFA9099D3FA32F8A1" box="[1340,1423,1781,1805]" pageId="12" pageNumber="13">EMUS</collectionCode>
);
<emphasis id="A5F2EA80F973057FFF3B983FFEB4F89E" bold="true" box="[151,265,1817,1842]" pageId="12" pageNumber="13">Wyoming</emphasis>
:
<emphasis id="A5F2EA80F973057FFEB4983FFE3CF89D" box="[280,385,1817,1841]" italics="true" pageId="12" pageNumber="13">Platte Co</emphasis>
.,
<collectingCountry id="EF917602F973057FFE3A983FFDBFF89D" box="[406,514,1817,1841]" name="Guernsey" pageId="12" pageNumber="13">Guernsey</collectingCountry>
, 1Ƥ23,
<date id="E3381052F973057FFDF3983FFDD0F89D" box="[607,621,1817,1841]" pageId="12" pageNumber="13" value="2002-08-06">6</date>
.VIII.2002, Williams (
<collectionCode id="F197AE57F973057FFCCF983FFC0BF89D" box="[867,950,1817,1841]" pageId="12" pageNumber="13">EMUS</collectionCode>
). Over 50 additional specimens from across the range of this
<taxonomicName id="50864D11F973057FFEFE981BFE1FF8F9" box="[338,418,1853,1877]" class="Insecta" family="Scoliidae" genus="Scolia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species">species</taxonomicName>
(
<collectingCountry id="EF917602F973057FFE1D981BFE53F8F9" box="[433,494,1853,1877]" name="United States of America" pageId="12" pageNumber="13">USA</collectingCountry>
: CO, KS, NE, OK, TX) were examined (
<collectionCode id="F197AE57F973057FFC19981BFC4AF8F9" box="[949,1015,1853,1877]" pageId="12" pageNumber="13">CISC</collectionCode>
,
<collectionCode id="F197AE57F973057FFBAE981BFBF5F8F9" box="[1026,1096,1853,1877]" httpUri="http://grbio.org/cool/sxrc-6x8s" name="Cornell University Insect Collection" pageId="12" pageNumber="13">CUIC</collectionCode>
,
<collectionCode id="F197AE57F973057FFBFE981BFB1CF8F9" box="[1106,1185,1853,1877]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="13">SEMC</collectionCode>
,
<collectionCode id="F197AE57F973057FFB009818FB42F8FA" box="[1196,1279,1854,1878]" pageId="12" pageNumber="13">TAMU</collectionCode>
,
<collectionCode id="F197AE57F973057FFAA6981BFADBF8F9" box="[1290,1382,1853,1877]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="12" pageNumber="13">UMMZ</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="DF9C6519F973057EFF6B9844FCEAFD60" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="discussion">
<paragraph id="97393692F973057FFF6B9844FA21F849" blockId="12.[151,1436,341,2021]" pageId="12" pageNumber="13">
<emphasis id="A5F2EA80F973057FFF6B9844FE89F8D6" bold="true" box="[199,308,1890,1914]" pageId="12" pageNumber="13">Remarks</emphasis>
. Molecular sequence data from ITS1 and ITS2 were used previously to associate
<taxonomicName id="50864D11F973057FFAA39844FA21F8D5" box="[1295,1436,1889,1913]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="birkmani">
<emphasis id="A5F2EA80F973057FFAA39844FA21F8D5" box="[1295,1436,1889,1913]" italics="true" pageId="12" pageNumber="13">D. birkmani</emphasis>
</taxonomicName>
females with males (formerly
<taxonomicName id="50864D11F973057FFE4A98A0FDE5F831" box="[486,600,1925,1949]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="reclusa">
<emphasis id="A5F2EA80F973057FFE4A98A0FDE5F831" box="[486,600,1925,1949]" italics="true" pageId="12" pageNumber="13">D. reclusa</emphasis>
</taxonomicName>
;
<bibRefCitation id="F3174B63F973057FFDCB98A3FCFEF831" author="Pilgrim" box="[615,835,1925,1949]" pageId="12" pageNumber="13" refString="Pilgrim, E. M., Williams, K. A. &amp; Pitts, J. P. (2008) Sex association and synonymy in Southwestern U. S. species of Dasymutilla (Hymenoptera: Mutillidae). Pan-Pacific Entomologist, 84, 58 - 69." type="journal article" year="2008">
Pilgrim
<emphasis id="A5F2EA80F973057FFD6D98A0FD4FF831" box="[705,754,1925,1949]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
., 2008
</bibRefCitation>
). Our study has found two more
<taxonomicName id="50864D11F973057FFB0198A3FB43F831" box="[1197,1278,1925,1949]" class="Insecta" family="Scoliidae" genus="Scolia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species">species</taxonomicName>
that appear to be conspecific with
<taxonomicName id="50864D11F973057FFED4988CFDBDF86D" box="[376,512,1961,1985]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="birkmani">
<emphasis id="A5F2EA80F973057FFED4988CFDBDF86D" box="[376,512,1961,1985]" italics="true" pageId="12" pageNumber="13">D. birkmani</emphasis>
</taxonomicName>
. Specimens identified as
<taxonomicName id="50864D11F973057FFC8C988FFC43F86D" box="[800,1022,1961,1985]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="arcana">
<emphasis id="A5F2EA80F973057FFC8C988FFC43F86D" box="[800,1022,1961,1985]" italics="true" pageId="12" pageNumber="13">Dasymutilla arcana</emphasis>
</taxonomicName>
and
<taxonomicName id="50864D11F973057FFB99988CFB13F86D" box="[1077,1198,1962,1985]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="corcyra">
<emphasis id="A5F2EA80F973057FFB99988CFB13F86D" box="[1077,1198,1962,1985]" italics="true" pageId="12" pageNumber="13">D. corcyra</emphasis>
</taxonomicName>
are recovered within a clade that includes seven females and one male identified as
<taxonomicName id="50864D11F973057FFCF898E8FC61F849" box="[852,988,1997,2021]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="birkmani">
<emphasis id="A5F2EA80F973057FFCF898E8FC61F849" box="[852,988,1997,2021]" italics="true" pageId="12" pageNumber="13">D. birkmani</emphasis>
</taxonomicName>
(
<figureCitation id="0FBD2A17F973057FFC4098EBFB82F849" box="[1004,1087,1997,2021]" captionStart="FIGURE 73" captionStartId="23.[151,250,1948,1971]" captionTargetBox="[297,1287,647,1911]" captionTargetId="figure@23.[282,1305,625,1927]" captionTargetPageId="23" captionText="FIGURE 73: Bayesian phylogeny reconstructed using ITS 1 and ITS 2 sequences and only including members of the Dasymutilla monticola species-group and three outgroups from other Dasymutilla species-groups." httpUri="https://zenodo.org/record/213060/files/figure.png" pageId="12" pageNumber="13">Fig. 73</figureCitation>
). The relationships between
<emphasis id="A5F2EA80F973057FFA2D98E8FA21F849" box="[1409,1436,1998,2021]" italics="true" pageId="12" pageNumber="13">D.</emphasis>
</paragraph>
<paragraph id="97393692F972057EFF3B9FBFFB79FF78" blockId="13.[151,1437,151,716]" pageId="13" pageNumber="14">
<taxonomicName id="50864D11F972057EFF3B9FBFFF50FF1C" box="[151,237,153,176]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="corcyra">
<emphasis id="A5F2EA80F972057EFF3B9FBFFF50FF1C" box="[151,237,153,176]" italics="true" pageId="13" pageNumber="14">corcyra</emphasis>
</taxonomicName>
,
<taxonomicName id="50864D11F972057EFF579FBFFED6FF1C" box="[251,363,153,176]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="arcana">
<emphasis id="A5F2EA80F972057EFF579FBFFED6FF1C" box="[251,363,153,176]" italics="true" pageId="13" pageNumber="14">D. arcana</emphasis>
</taxonomicName>
, and two clades of
<taxonomicName id="50864D11F972057EFDE89FBFFD71FF1C" box="[580,716,152,176]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="birkmani">
<emphasis id="A5F2EA80F972057EFDE89FBFFD71FF1C" box="[580,716,152,176]" italics="true" pageId="13" pageNumber="14">D. birkmani</emphasis>
</taxonomicName>
are unresolved or poorly supported, but both male specimens of
<taxonomicName id="50864D11F972057EFF3B9F9BFEBAFF78" box="[151,263,189,212]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="arcana">
<emphasis id="A5F2EA80F972057EFF3B9F9BFEBAFF78" box="[151,263,189,212]" italics="true" pageId="13" pageNumber="14">D. arcana</emphasis>
</taxonomicName>
have identical ITS1 and ITS2 sequences to one of the
<taxonomicName id="50864D11F972057EFCC79F9BFC4FFF78" box="[875,1010,188,212]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="birkmani">
<emphasis id="A5F2EA80F972057EFCC79F9BFC4FFF78" box="[875,1010,188,212]" italics="true" pageId="13" pageNumber="14">D. birkmani</emphasis>
</taxonomicName>
females (JP1885).
</paragraph>
<paragraph id="97393692F972057EFF6B9FF9FAEDFEC8" blockId="13.[151,1437,151,716]" pageId="13" pageNumber="14">
The only diagnostic character separating
<taxonomicName id="50864D11F972057EFD339FC7FCAFFF54" box="[671,786,225,248]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="arcana">
<emphasis id="A5F2EA80F972057EFD339FC7FCAFFF54" box="[671,786,225,248]" italics="true" pageId="13" pageNumber="14">D. arcana</emphasis>
</taxonomicName>
from males of
<taxonomicName id="50864D11F972057EFC6A9FC7FBEDFF54" box="[966,1104,224,248]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="birkmani">
<emphasis id="A5F2EA80F972057EFC6A9FC7FBEDFF54" box="[966,1104,224,248]" italics="true" pageId="13" pageNumber="14">D. birkmani</emphasis>
</taxonomicName>
is the metasomal coloration, where
<taxonomicName id="50864D11F972057EFF479E23FED9FEB0" box="[235,356,261,284]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="arcana">
<emphasis id="A5F2EA80F972057EFF479E23FED9FEB0" box="[235,356,261,284]" italics="true" pageId="13" pageNumber="14">D. arcana</emphasis>
</taxonomicName>
has S2 red and
<taxonomicName id="50864D11F972057EFDEE9E23FD6FFEB0" box="[578,722,260,284]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="birkmani">
<emphasis id="A5F2EA80F972057EFDEE9E23FD6FFEB0" box="[578,722,260,284]" italics="true" pageId="13" pageNumber="14">D. birkmani</emphasis>
</taxonomicName>
has S2 black. Multiple
<taxonomicName id="50864D11F972057EFBAB9E22FB2DFEB0" box="[1031,1168,260,284]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="A5F2EA80F972057EFBAB9E22FB2DFEB0" box="[1031,1168,260,284]" italics="true" pageId="13" pageNumber="14">Dasymutilla</emphasis>
</taxonomicName>
<taxonomicName id="50864D11F972057EFB0C9E22FB4EFEB0" box="[1184,1267,260,284]" class="Insecta" family="Scoliidae" genus="Scolia" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species">species</taxonomicName>
, including
<taxonomicName id="50864D11F972057EFA2D9E23FD82FE93" authority="Pilgrim et al., 2009" authorityName="Pilgrim et al." authorityYear="2009" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="quadriguttata">
<emphasis id="A5F2EA80F972057EFA2D9E23FE8CFEEC" italics="true" pageId="13" pageNumber="14">D. quadriguttata</emphasis>
(
<bibRefCitation id="F3174B63F972057EFEEA9E01FD8AFE93" author="Pilgrim" box="[326,567,295,320]" pageId="13" pageNumber="14" refString="Pilgrim, E. M., Williams, K. A., Manley, D. G., &amp; Pitts, J. P. (2009) Addressing the Dasymutilla quadriguttata species-group and species-complex (Hymenoptera: Mutillidae): several distinct species or a single, morphologically variable species? Journal of the Kansas Entomological Society, 82, 231 - 249." type="journal article" year="2009">
Pilgrim
<emphasis id="A5F2EA80F972057EFE049E0FFE5BFEEC" box="[424,486,296,320]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
, 2009
</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="50864D11F972057EFDFC9E0FFBBCFE93" authority="Williams et al., 2011" authorityName="Williams et al." authorityYear="2011" box="[592,1025,295,320]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bioculata">
<emphasis id="A5F2EA80F972057EFDFC9E0FFD5EFEEC" box="[592,739,296,320]" italics="true" pageId="13" pageNumber="14">D. bioculata</emphasis>
(
<bibRefCitation id="F3174B63F972057EFD5B9E01FC44FE93" author="Williams" box="[759,1017,295,320]" pageId="13" pageNumber="14" refString="Williams, K. A., Manley, D. G., Pilgrim, E. M., von Dohlen, C. D., &amp; Pitts, J. P. (2011) Multi-faceted assessment of species validity in the Dasymutilla bioculata species-group (Hymenoptera: Mutillidae). Systematic Entomology, 36, 180 - 191." type="journal article" year="2011">
Williams
<emphasis id="A5F2EA80F972057EFCC09E0FFC17FEEC" box="[876,938,296,320]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
, 2011
</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="50864D11F972057EFBE49E0FFAD5FE93" authority="Mickel, 1928" authorityName="Mickel" authorityYear="1928" box="[1096,1384,295,320]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="vesta">
<emphasis id="A5F2EA80F972057EFBE49E0FFB17FEEC" box="[1096,1194,297,320]" italics="true" pageId="13" pageNumber="14">D. vesta</emphasis>
(
<bibRefCitation id="F3174B63F972057EFB139E01FAE2FE93" author="Mickel" box="[1215,1375,295,319]" pageId="13" pageNumber="14" refString="Mickel, C. E. (1928) Biological and taxonomic investigations on the mutillid wasps. United States Museum Bulletin, 143, 1 - 351." type="journal article" year="1928">Mickel, 1928</bibRefCitation>
)
</taxonomicName>
, are dimorphic for this character. Males of
<taxonomicName id="50864D11F972057EFDE99E6BFD08FEC8" box="[581,693,333,356]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="arcana">
<emphasis id="A5F2EA80F972057EFDE99E6BFD08FEC8" box="[581,693,333,356]" italics="true" pageId="13" pageNumber="14">D. arcana</emphasis>
</taxonomicName>
and
<taxonomicName id="50864D11F972057EFD409E6BFCC9FEC8" box="[748,884,332,356]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="birkmani">
<emphasis id="A5F2EA80F972057EFD409E6BFCC9FEC8" box="[748,884,332,356]" italics="true" pageId="13" pageNumber="14">D. birkmani</emphasis>
</taxonomicName>
also have identical genitalia (
<figureCitation id="0FBD2A17F972057EFB6D9E6AFAF9FEC8" box="[1217,1348,332,356]" captionStart="FIGURES 36 71" captionStartId="13.[151,264,1887,1910]" captionTargetBox="[339,1253,777,1858]" captionTargetId="figure@13.[323,1264,772,1866]" captionTargetPageId="13" captionText="FIGURES 36 71. Male genitalia: dorsal view, ventral view, lateral view, and penial valve. 36 39. Dasymutilla archboldi; 40 43. D. arenneronea; 44 47. D. birkmani; 48 51. D. bonita; 52 55: D. canella; 56 59. D. macilenta; 60 63. D. monticola; 64 67. D. saetigera; 68 71. D. vesta." httpUri="https://zenodo.org/record/213058/files/figure.png" pageId="13" pageNumber="14">Figs 4447</figureCitation>
).
</paragraph>
<paragraph id="97393692F972057EFF6B9E49FD96FDF3" blockId="13.[151,1437,151,716]" pageId="13" pageNumber="14">
Mickel separated
<taxonomicName id="50864D11F972057EFE329E57FDA2FE24" box="[414,543,369,392]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="corcyra">
<emphasis id="A5F2EA80F972057EFE329E57FDA2FE24" box="[414,543,369,392]" italics="true" pageId="13" pageNumber="14">D. corcyra</emphasis>
</taxonomicName>
(
<figureCitation id="0FBD2A17F972057EFD9B9E49FD3FFE2B" box="[567,642,367,391]" captionStart="FIGURES 1 8. 1, 2" captionStartId="8.[151,264,1883,1906]" captionTargetBox="[153,1432,665,1861]" captionTargetId="figure@8.[151,1436,663,1862]" captionTargetPageId="8" captionText="FIGURES 1 8. 1, 2, Habitus, lateral view, Ƥ. 1. Dasymutilla canella; 2. D. spilota. 3, 4, Mesosoma, dorsal view, male. 3. D. saetigera; 4. D. vesta. 5 8, Head, dorsal view, female. 5. D. arenneronea (D. chattahoochei); 6. D. birkmani; 7. D. birkmani (D. corcyra); 8. D. macilenta (D. chattahoochei)." httpUri="https://zenodo.org/record/213055/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
) from
<taxonomicName id="50864D11F972057EFD4C9E57FCD2FE24" box="[736,879,368,392]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="birkmani">
<emphasis id="A5F2EA80F972057EFD4C9E57FCD2FE24" box="[736,879,368,392]" italics="true" pageId="13" pageNumber="14">D. birkmani</emphasis>
</taxonomicName>
females by the presence of tubercles on the posterolateral head margins (
<bibRefCitation id="F3174B63F972057EFE759EB2FDC9FE00" author="Mickel" box="[473,628,404,428]" pageId="13" pageNumber="14" refString="Mickel, C. E. (1928) Biological and taxonomic investigations on the mutillid wasps. United States Museum Bulletin, 143, 1 - 351." type="journal article" year="1928">Mickel, 1928</bibRefCitation>
). All female specimens of
<taxonomicName id="50864D11F972057EFC319EB3FB99FE00" box="[925,1060,404,428]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="birkmani">
<emphasis id="A5F2EA80F972057EFC319EB3FB99FE00" box="[925,1060,404,428]" italics="true" pageId="13" pageNumber="14">D. birkmani</emphasis>
</taxonomicName>
, however, have variably angulate posterolateral head margins and the posterior head margin virtually flat or concave (
<figureCitation id="0FBD2A17F972057EFBDB9E91FB7CFE63" box="[1143,1217,439,463]" captionStart="FIGURES 1 8. 1, 2" captionStartId="8.[151,264,1883,1906]" captionTargetBox="[153,1432,665,1861]" captionTargetId="figure@8.[151,1436,663,1862]" captionTargetPageId="8" captionText="FIGURES 1 8. 1, 2, Habitus, lateral view, Ƥ. 1. Dasymutilla canella; 2. D. spilota. 3, 4, Mesosoma, dorsal view, male. 3. D. saetigera; 4. D. vesta. 5 8, Head, dorsal view, female. 5. D. arenneronea (D. chattahoochei); 6. D. birkmani; 7. D. birkmani (D. corcyra); 8. D. macilenta (D. chattahoochei)." httpUri="https://zenodo.org/record/213055/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
). Specimens of
<taxonomicName id="50864D11F972057EFA2D9E9FFF50FE58" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="corcyra">
<emphasis id="A5F2EA80F972057EFA2D9E9FFF50FE58" italics="true" pageId="13" pageNumber="14">D. corcyra</emphasis>
</taxonomicName>
are typically larger (7.5
<quantity id="507E9B77F972057EFDA59EFAFDF7FE58" box="[521,586,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="14" unit="mm" value="9.0">9 mm</quantity>
, mean=8.3, n=6) than those of
<taxonomicName id="50864D11F972057EFC0B9EFBFB92FE58" box="[935,1071,476,500]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="birkmani">
<emphasis id="A5F2EA80F972057EFC0B9EFBFB92FE58" box="[935,1071,476,500]" italics="true" pageId="13" pageNumber="14">D. birkmani</emphasis>
</taxonomicName>
(3.58.5 mm, mean=5.4, n=40); hypothetically, the magnitude of head tubercles may be correlated with body size and not
<taxonomicName id="50864D11F972057EFB129ED9FAB2FDBB" box="[1214,1295,511,535]" class="Insecta" family="Scoliidae" genus="Scolia" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species">species</taxonomicName>
divergence. Although not to this extent, intraspecific variation in head tubercle structure has also been observed in
<taxonomicName id="50864D11F972057EFA2D9D03FD9AFDF3" authority="Pilgrim et al., 2009" authorityName="Pilgrim et al." authorityYear="2009" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="quadriguttata">
<emphasis id="A5F2EA80F972057EFA2D9D03FE8CFDCC" italics="true" pageId="13" pageNumber="14">D. quadriguttata</emphasis>
(
<bibRefCitation id="F3174B63F972057EFE939D61FDA2FDF3" author="Pilgrim" box="[319,543,583,608]" pageId="13" pageNumber="14" refString="Pilgrim, E. M., Williams, K. A., Manley, D. G., &amp; Pitts, J. P. (2009) Addressing the Dasymutilla quadriguttata species-group and species-complex (Hymenoptera: Mutillidae): several distinct species or a single, morphologically variable species? Journal of the Kansas Entomological Society, 82, 231 - 249." type="journal article" year="2009">
Pilgrim
<emphasis id="A5F2EA80F972057EFE309D6FFE69FDCC" box="[412,468,584,608]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
, 2009
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="97393692F972057EFF6B9D4AFCEAFD60" blockId="13.[151,1437,151,716]" pageId="13" pageNumber="14">
<taxonomicName id="50864D11F972057EFF6B9D4AFE14FD28" box="[199,425,620,644]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="arcana">
<emphasis id="A5F2EA80F972057EFF6B9D4AFE14FD28" box="[199,425,620,644]" italics="true" pageId="13" pageNumber="14">Dasymutilla arcana</emphasis>
</taxonomicName>
,
<taxonomicName id="50864D11F972057EFE179D4BFDFBFD28" box="[443,582,620,644]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="birkmani">
<emphasis id="A5F2EA80F972057EFE179D4BFDFBFD28" box="[443,582,620,644]" italics="true" pageId="13" pageNumber="14">D. birkmani</emphasis>
</taxonomicName>
, and
<taxonomicName id="50864D11F972057EFD209D4BFCB4FD28" box="[652,777,621,644]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="corcyra">
<emphasis id="A5F2EA80F972057EFD209D4BFCB4FD28" box="[652,777,621,644]" italics="true" pageId="13" pageNumber="14">D. corcyra</emphasis>
</taxonomicName>
also have broadly overlapping geographic ranges in the central
<collectingCountry id="EF917602F972057EFF479DA9FE9AFD0B" box="[235,295,655,679]" name="United States of America" pageId="13" pageNumber="14">USA</collectingCountry>
. Based on phylogenetic affinities, morphological similarities, and overlapping distributions, we place
<taxonomicName id="50864D11F972057EFF3B9D93FEBAFD60" box="[151,263,693,716]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="arcana">
<emphasis id="A5F2EA80F972057EFF3B9D93FEBAFD60" box="[151,263,693,716]" italics="true" pageId="13" pageNumber="14">D. arcana</emphasis>
</taxonomicName>
and
<taxonomicName id="50864D11F972057EFE929D93FE0BFD60" box="[318,438,693,716]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="corcyra">
<emphasis id="A5F2EA80F972057EFE929D93FE0BFD60" box="[318,438,693,716]" italics="true" pageId="13" pageNumber="14">D. corcyra</emphasis>
</taxonomicName>
as
<emphasis id="A5F2EA80F972057EFE779D95FD19FD60" bold="true" box="[475,676,691,716]" pageId="13" pageNumber="14">junior synonyms</emphasis>
of
<taxonomicName id="50864D11F972057EFD649D93FCEDFD60" box="[712,848,692,716]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="birkmani">
<emphasis id="A5F2EA80F972057EFD649D93FCEDFD60" box="[712,848,692,716]" italics="true" pageId="13" pageNumber="14">D. birkmani</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>