243 lines
38 KiB
XML
243 lines
38 KiB
XML
<document id="EBAC5CE178EEA48940C23420BF5DD7F0" ID-DOI="10.26107/RBZ-2020-0074" ID-ISSN="2345-7600" ID-Zenodo-Dep="5345178" ID-ZooBank="3DA09D0A-DD6D-42FA-8D06-0899FB9D94BE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630199701236" checkinUser="diego" docAuthor="Nguyen, Lien Thi Phuong" docDate="2020" docId="03F0DC55AD59FFD4D68CFBCC39DAFE1D" docLanguage="en" docName="RBZ-2020-0074.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Pareumenes (Nortonia) caoduong Nguyen 2020, new species" docType="treatment" docVersion="3" lastPageNumber="592" masterDocId="FFC9A42DAD5BFFD0D625FFF23D2FFF88" masterDocTitle="A new Vietnamese species and a key to the Oriental species of Pareumenes (Nortonia) de Saussure, 1855 (Hymenoptera: Vespidae: Eumeninae)" masterLastPageNumber="594" masterPageNumber="588" pageNumber="590" updateTime="1699183355997" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="27FE721E1E4E591DCC0B5DDB71E1F9EB">A new Vietnamese species and a key to the Oriental species of Pareumenes (Nortonia) de Saussure, 1855 (Hymenoptera: Vespidae: Eumeninae)</mods:title>
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</mods:titleInfo>
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<mods:name id="B9C9CD53E6217A9A3BF5AA6E90D1F66C" type="personal">
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<mods:role id="727CAFBAFBFAAB22ACF27C01540E23C2">
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<mods:roleTerm id="73B38F73434D9392CF7AE8FE3CEA03C2">Author</mods:roleTerm>
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<mods:namePart id="37404678C2F4A822C0E9B915BB3B2563">Nguyen, Lien Thi Phuong</mods:namePart>
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<mods:typeOfResource id="77D4FDF9088FF4C1DDF308726209A731">text</mods:typeOfResource>
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<mods:title id="268260A97428DE9DD64088BF3066B67B">Raffles Bulletin of Zoology</mods:title>
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</mods:titleInfo>
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<mods:part id="EED31F1FB8877D32F0E160DC0C50B326">
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<mods:date id="2E677222289803B89050E09120E41787">2020</mods:date>
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<mods:detail id="AAE583621EE95F9E4410D51872AE5FE1" type="pubDate">
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<mods:number id="C5ADDCBD5808A9047E32BDCC245164DD">2020-06-29</mods:number>
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</mods:detail>
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<mods:detail id="FFB7FA8649F079746A24CC4E50A261A1" type="volume">
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<mods:number id="56A6E58749B7DC366129B030D9090E37">68</mods:number>
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</mods:relatedItem>
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<mods:classification id="53E37F0B582484442A89A458EFAE772B">journal article</mods:classification>
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<mods:identifier id="8BE15E2CC3AA6C317E74C1C7E2282210" type="DOI">10.26107/RBZ-2020-0074</mods:identifier>
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<mods:identifier id="F6C3E4A3A3BC58F793C5C91F341431D3" type="ISSN">2345-7600</mods:identifier>
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</mods:mods>
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<treatment id="03F0DC55AD59FFD4D68CFBCC39DAFE1D" LSID="urn:lsid:plazi:treatment:03F0DC55AD59FFD4D68CFBCC39DAFE1D" httpUri="http://treatment.plazi.org/id/03F0DC55AD59FFD4D68CFBCC39DAFE1D" lastPageId="4" lastPageNumber="592" pageId="2" pageNumber="590">
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<subSubSection id="C3433EC8AD59FFD2D68CFBCC3F97FBDD" box="[169,696,1085,1109]" pageId="2" pageNumber="590" type="nomenclature">
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<paragraph id="8BE66D43AD59FFD2D68CFBCC3F97FBDD" blockId="2.[169,696,1085,1141]" box="[169,696,1085,1109]" pageId="2" pageNumber="590">
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<heading id="D0AEDA2FAD59FFD2D68CFBCC3F97FBDD" bold="true" box="[169,696,1085,1109]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="590" reason="2">
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<emphasis id="B92DB151AD59FFD2D68CFBCC3F97FBDD" bold="true" box="[169,696,1085,1109]" pageId="2" pageNumber="590">
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<taxonomicName id="4C5916C0AD59FFD2D68CFBCC3F0CFBDD" authority="Nguyen, 2020" authorityName="Nguyen" authorityYear="2020" box="[169,547,1085,1109]" class="Insecta" family="Vespidae" genus="Pareumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="species" species="caoduong" status="sp. nov." subGenus="Nortonia">
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<emphasis id="B92DB151AD59FFD2D68CFBCC3C1EFBDD" bold="true" box="[169,305,1086,1109]" italics="true" pageId="2" pageNumber="590">Pareumenes</emphasis>
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(
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<emphasis id="B92DB151AD59FFD2D766FBCF3C89FBDD" bold="true" box="[323,422,1085,1109]" italics="true" pageId="2" pageNumber="590">Nortonia</emphasis>
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)
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<emphasis id="B92DB151AD59FFD2D792FBCF3F0CFBDD" bold="true" box="[439,547,1085,1109]" italics="true" pageId="2" pageNumber="590">caoduong</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21E0C2AAD59FFD2D416FBCF3F97FBDD" box="[563,696,1085,1109]" pageId="2" pageNumber="590" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3433EC8AD59FFD2D750FBAF3CC3FBFD" box="[373,492,1117,1141]" pageId="2" pageNumber="590" type="description">
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<paragraph id="8BE66D43AD59FFD2D750FBAF3CC3FBFD" blockId="2.[169,696,1085,1141]" box="[373,492,1117,1141]" pageId="2" pageNumber="590">
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(
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<figureCitation id="136271C6AD59FFD2D758FBAF3CCCFBFD" box="[381,483,1117,1141]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="1.[136,180,1922,1944]" captionStartId-1="3.[136,180,2022,2044]" captionTargetBox-0="[136,1474,300,1906]" captionTargetBox-1="[136,1474,231,2007]" captionTargetId-0="figure-7@1.[136,1474,300,1906]" captionTargetId-1="figure-47@3.[136,1474,231,2007]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionText-0="Figs. 1–6. Pareumenes caoduong, new species, holotype, female. 1, Head in frontal view; 2, Vertex in dorsal view; 3, Mesosoma in dorsal view; 4, Mesosoma in lateral view; 5, Propodeum in posterior view; 6, Tergum I in dorsal view. Scale = 1 mm." captionText-1="Figs. 7–12. Pareumenes caoduong, new species, holotype, female (7–9). 7, Metasomal terga in lateral view; 8, Forewing; 9, Habitus. Pareumenes quadrispinosus acutus Liu, 1941, female (10–12). 10, Head in frontal view; 11, Forewing; 12, Habitus. Scale = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5345182" figureDoi-1="http://doi.org/10.5281/zenodo.5345186" httpUri-0="https://zenodo.org/record/5345182/files/figure.png" httpUri-1="https://zenodo.org/record/5345186/files/figure.png" pageId="2" pageNumber="590">Figs. 1–9</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3433EC8AD59FFD2D654FB6F3DDAFAFD" pageId="2" pageNumber="590" type="materials_examined">
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<paragraph id="8BE66D43AD59FFD2D654FB6F3DDAFAFD" blockId="2.[113,752,1181,1397]" pageId="2" pageNumber="590">
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<emphasis id="B92DB151AD59FFD2D654FB6F3C7BFB3D" bold="true" box="[113,340,1181,1205]" pageId="2" pageNumber="590">Material examined.</emphasis>
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<materialsCitation id="3B31671EAD59FFD2D77BFB6F3D9EFADD" collectingDate="2019-05-15" collectionCode="IEBR, T, V" collectorName="P. Lien & Truong X. Lam & Nguyen Q. Cuong" country="Vietnam" county="Cao Duong" latitude="22.29236" location="Ham Yen" longLatPrecision="14" longitude="104.991104" municipality="Yen Thuan" pageId="2" pageNumber="590" specimenCount="1" stateProvince="Tuyen Quang" typeStatus="holotype">
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<typeStatus id="54E2D3E1AD59FFD2D77BFB6F3CEAFB3D" box="[350,453,1181,1205]" pageId="2" pageNumber="590" type="holotype">Holotype</typeStatus>
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, female, pinned (deposited in
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<collectionCode id="ED48F586AD59FFD2D6B5FB4F3DFEFB5D" box="[144,209,1213,1237]" pageId="2" pageNumber="590">IEBR</collectionCode>
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), labelled “
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<collectingCountry id="F34E2DD3AD59FFD2D775FB4F3CFDFB5D" box="[336,466,1213,1237]" name="Vietnam" pageId="2" pageNumber="590">VIETNAM</collectingCountry>
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:
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<collectingCounty id="628715CFAD59FFD2D7FAFB4F3F4FFB5E" box="[479,608,1213,1238]" pageId="2" pageNumber="590">Cao Duong</collectingCounty>
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,
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<collectingMunicipality id="6B82F739AD59FFD2D448FB4C3FC4FB5D" box="[621,747,1213,1237]" pageId="2" pageNumber="590">Yen Thuan</collectingMunicipality>
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,
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,
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<collectingRegion id="499DA3A1AD59FFD2D6CCFB2C3C50FB7D" box="[233,383,1245,1270]" country="Vietnam" name="Tuyen Quang" pageId="2" pageNumber="590">Tuyen Quang</collectingRegion>
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, Cham Chu NP, way to waterfall,
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<geoCoordinate id="EE6D0B84AD59FFD2D654FB0F3C24FA9D" box="[113,267,1276,1301]" degrees="22" direction="north" minutes="17" orientation="latitude" pageId="2" pageNumber="590" precision="1" seconds="32.5" value="22.29236">22°17′32.5″N</geoCoordinate>
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<geoCoordinate id="EE6D0B84AD59FFD2D732FB0E3C86FA9D" box="[279,425,1276,1301]" degrees="104" direction="east" minutes="59" orientation="longitude" pageId="2" pageNumber="590" precision="15" seconds="28" value="104.991104">104°59′28″E</geoCoordinate>
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,
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<date id="FFE74B83AD59FFD2D79CFB0F3F04FA9D" box="[441,555,1277,1301]" pageId="2" pageNumber="590" value="2019-05-15">
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<collectingDate id="EFA3B26BAD59FFD2D79CFB0F3F04FA9D" box="[441,555,1277,1301]" pageId="2" pageNumber="590" value="2019-05-15">
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15.v.
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<quantity id="4CA1C0A6AD59FFD2D7CEFB0F3F04FA9D" box="[491,555,1277,1301]" metricMagnitude="6" metricUnit="m" metricValue="2.019643" pageId="2" pageNumber="590" unit="m" value="2019643.0">2019</quantity>
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</collectingDate>
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</date>
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, 643 m, Nguyen
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<collectionCode id="ED48F586AD59FFD2D657FAEC3DA9FABD" box="[114,134,1310,1333]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="590">T</collectionCode>
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.
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<collectorName id="26AC0895AD59FFD2D6ACFAEC3DF2FABD" box="[137,221,1309,1333]" pageId="2" pageNumber="590">P. Lien</collectorName>
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,
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<collectorName id="26AC0895AD59FFD2D6CEFAEC3CB2FABD" box="[235,413,1310,1334]" pageId="2" pageNumber="590">Truong X. Lam</collectorName>
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,
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<collectorName id="26AC0895AD59FFD2D78EFAEC3F58FABD" box="[427,631,1309,1334]" pageId="2" pageNumber="590">Nguyen Q. Cuong</collectorName>
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||
& Mai
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<collectionCode id="ED48F586AD59FFD2D4F3FAEC3FC3FABD" box="[726,748,1310,1333]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="590" type="Museum">V</collectionCode>
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||
. Thai”
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||
</materialsCitation>
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.
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||
<materialsCitation id="3B31671EAD59FFD2D69BFACC3DDEFAFD" collectingDate="2019-05-15" collectionCode="IEBR" collectorName="P. Lien & Truong X. Lam & Nguyen Q. Cuong" country="Vietnam" county="Cao Duong" latitude="22.29236" location="Ham Yen" longLatPrecision="14" longitude="104.991104" municipality="Yen Thuan" pageId="2" pageNumber="590" specimenCount="1" specimenCount-female="1" stateProvince="Tuyen Quang" typeStatus="paratype">
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<typeStatus id="54E2D3E1AD59FFD2D69BFACC3C34FADE" box="[190,283,1342,1366]" pageId="2" pageNumber="590" type="paratype">Paratype</typeStatus>
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(deposited in
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<collectionCode id="ED48F586AD59FFD2D799FACF3CD1FADD" box="[444,510,1341,1365]" pageId="2" pageNumber="590">IEBR</collectionCode>
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):
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<specimenCount id="9D5FA6CAAD59FFD2D436FACF3F5AFADD" box="[531,629,1341,1365]" pageId="2" pageNumber="590" type="female">1 female</specimenCount>
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, same data as holotype
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||
</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3433EC8AD59FFD4D654FA6F3C08FA5D" lastPageId="4" lastPageNumber="592" pageId="2" pageNumber="590" type="diagnosis">
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<paragraph id="8BE66D43AD59FFD2D654FA6F3EC3FBBD" blockId="2.[112,753,1437,2070]" lastBlockId="2.[811,1452,189,1077]" pageId="2" pageNumber="590">
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<emphasis id="B92DB151AD59FFD2D654FA6F3DC6FA3D" bold="true" box="[113,233,1437,1461]" pageId="2" pageNumber="590">Diagnosis.</emphasis>
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This species can be distinguished from other species of subgenus
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<taxonomicName id="4C5916C0AD59FFD2D776FA4C3C9CFA5D" authorityName="de Saussure" authorityYear="1869" box="[339,435,1470,1493]" class="Insecta" family="Vespidae" genus="Pareumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="subGenus" subGenus="Nortonia">
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<emphasis id="B92DB151AD59FFD2D776FA4C3C9CFA5D" box="[339,435,1470,1493]" italics="true" pageId="2" pageNumber="590">Nortonia</emphasis>
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||
</taxonomicName>
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||
of the genus
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<taxonomicName id="4C5916C0AD59FFD2D46FFA4C3FE1FA5D" authorityName="de Saussure" authorityYear="1855" box="[586,718,1470,1493]" class="Insecta" family="Vespidae" genus="Pareumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="genus">
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<emphasis id="B92DB151AD59FFD2D46FFA4C3FE1FA5D" box="[586,718,1470,1493]" italics="true" pageId="2" pageNumber="590">Pareumenes</emphasis>
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||
</taxonomicName>
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||
by the following combination of characters: Head in frontal view slightly wider than high. Body with coarse punctures; clypeus flat in the middle part, in lateral view convex from basal half, then straight to apical margin, apical emargination not so deep; vertex with depression behind posterior ocelli, with two foveae placed inside V-shaped carina; distance from posterior ocelli to apical margin of vertex 2.4 times distance from posterior ocelli to inner eye margin; lateral sides of propodeum shallowly emarginate at apex; metasoma petiolate in dorsal view with strongly projecting spiracles at about one-third of the length of the petiole basally, in lateral view abruptly produced near base, then depressed and gradually convex to near apical margin; apical margins of all terga not raised; forewing with parastigma slightly longer than stigma measured along ventral part. Clypeus with irregular yellow mark in the middle; tergum I with two large yellow spots at lateral side near apex, terga II–VI black except a short narrow yellow band at apical margin of tergum III medially.
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<emphasis id="B92DB151AD59FFD2D508FF4F3E93FF5D" bold="true" box="[813,956,189,213]" pageId="2" pageNumber="590">Description.</emphasis>
|
||
Female (
|
||
<figureCitation id="136271C6AD59FFD2D20CFF4F395FFF5D" box="[1065,1136,189,213]" captionStart="Figs" captionStartId="3.[136,180,2022,2044]" captionTargetBox="[136,1474,231,2007]" captionTargetId="figure-47@3.[136,1474,231,2007]" captionTargetPageId="3" captionText="Figs. 7–12. Pareumenes caoduong, new species, holotype, female (7–9). 7, Metasomal terga in lateral view; 8, Forewing; 9, Habitus. Pareumenes quadrispinosus acutus Liu, 1941, female (10–12). 10, Head in frontal view; 11, Forewing; 12, Habitus. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5345186" httpUri="https://zenodo.org/record/5345186/files/figure.png" pageId="2" pageNumber="590">Fig. 9</figureCitation>
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||
). Body length
|
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(
|
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<typeStatus id="54E2D3E1AD59FFD2D511FF2F3EBEFF7D" box="[820,913,221,245]" pageId="2" pageNumber="590" type="holotype">holotype</typeStatus>
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<quantity id="4CA1C0A6AD59FFD2D5B0FF2F3EDAFF7D" box="[917,1013,221,245]" metricMagnitude="-2" metricUnit="m" metricValue="1.98" pageId="2" pageNumber="590" unit="mm" value="19.8">19.8 mm</quantity>
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); forewing length
|
||
<quantity id="4CA1C0A6AD59FFD2D28BFF2F386CFF7D" box="[1198,1347,221,245]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" metricValueMax="1.8699999999999999" metricValueMin="1.85" pageId="2" pageNumber="590" unit="mm" value="18.6" valueMax="18.7" valueMin="18.5">18.5–18.7 mm</quantity>
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(
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<typeStatus id="54E2D3E1AD59FFD2D36AFF2F3883FF7D" box="[1359,1452,221,245]" pageId="2" pageNumber="590" type="holotype">holotype</typeStatus>
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<quantity id="4CA1C0A6AD59FFD2D509FF0F3EBDFE9D" box="[812,914,253,277]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="2" pageNumber="590" unit="mm" value="18.5">18.5 mm</quantity>
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||
). Head in frontal view subcircular, slightly wider than high, about 1.1 times as wide as high (
|
||
<figureCitation id="136271C6AD59FFD2D32FFEEF3864FEBD" box="[1290,1355,285,309]" captionStart="Figs" captionStartId="1.[136,180,1922,1944]" captionTargetBox="[136,1474,300,1906]" captionTargetId="figure-7@1.[136,1474,300,1906]" captionTargetPageId="1" captionText="Figs. 1–6. Pareumenes caoduong, new species, holotype, female. 1, Head in frontal view; 2, Vertex in dorsal view; 3, Mesosoma in dorsal view; 4, Mesosoma in lateral view; 5, Propodeum in posterior view; 6, Tergum I in dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5345182" httpUri="https://zenodo.org/record/5345182/files/figure.png" pageId="2" pageNumber="590">Fig. 1</figureCitation>
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||
). Vertex with slightly swollen area around ocellus, with depression behind posterior ocellus (
|
||
<figureCitation id="136271C6AD59FFD2D264FEAF39ABFEFD" box="[1089,1156,349,373]" captionStart="Figs" captionStartId="1.[136,180,1922,1944]" captionTargetBox="[136,1474,300,1906]" captionTargetId="figure-7@1.[136,1474,300,1906]" captionTargetPageId="1" captionText="Figs. 1–6. Pareumenes caoduong, new species, holotype, female. 1, Head in frontal view; 2, Vertex in dorsal view; 3, Mesosoma in dorsal view; 4, Mesosoma in lateral view; 5, Propodeum in posterior view; 6, Tergum I in dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5345182" httpUri="https://zenodo.org/record/5345182/files/figure.png" pageId="2" pageNumber="590">Fig. 2</figureCitation>
|
||
), with two cephalic foveae placed inside a V-shaped carina, close to each other. Distance from posterior ocelli to apical margin of vertex about 2.4 times distance from posterior ocelli to inner eye margin (
|
||
<figureCitation id="136271C6AD59FFD2D3A5FE4F3E12FE7D" captionStart="Figs" captionStartId="1.[136,180,1922,1944]" captionTargetBox="[136,1474,300,1906]" captionTargetId="figure-7@1.[136,1474,300,1906]" captionTargetPageId="1" captionText="Figs. 1–6. Pareumenes caoduong, new species, holotype, female. 1, Head in frontal view; 2, Vertex in dorsal view; 3, Mesosoma in dorsal view; 4, Mesosoma in lateral view; 5, Propodeum in posterior view; 6, Tergum I in dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5345182" httpUri="https://zenodo.org/record/5345182/files/figure.png" pageId="2" pageNumber="590">Fig. 2</figureCitation>
|
||
). Gena thick, wider than eye, in lateral view 1.2 times as wide as eye at ocular sinus. Occipital carina complete, present along entire length of gena. Inner eye margins slightly convergent ventrally, in frontal view nearly equal from each other at vertex as at clypeus. Clypeus flat in the middle part, disc of clypeus in lateral view weakly convex at basal half, then almost straight to apical margin, in frontal view nearly 1.2 times as high as wide (
|
||
<figureCitation id="136271C6AD59FFD2D265FD4F3953FD5D" box="[1088,1148,701,725]" captionStart="Figs" captionStartId="1.[136,180,1922,1944]" captionTargetBox="[136,1474,300,1906]" captionTargetId="figure-7@1.[136,1474,300,1906]" captionTargetPageId="1" captionText="Figs. 1–6. Pareumenes caoduong, new species, holotype, female. 1, Head in frontal view; 2, Vertex in dorsal view; 3, Mesosoma in dorsal view; 4, Mesosoma in lateral view; 5, Propodeum in posterior view; 6, Tergum I in dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5345182" httpUri="https://zenodo.org/record/5345182/files/figure.png" pageId="2" pageNumber="590">Fig. 1</figureCitation>
|
||
), with basal margin strongly convex medially and almost touching antennal sockets; apical margin emarginate medially, forming sharp tooth on each lateral side (
|
||
<figureCitation id="136271C6AD59FFD2D591FCEF3EDBFCBD" box="[948,1012,797,821]" captionStart="Figs" captionStartId="1.[136,180,1922,1944]" captionTargetBox="[136,1474,300,1906]" captionTargetId="figure-7@1.[136,1474,300,1906]" captionTargetPageId="1" captionText="Figs. 1–6. Pareumenes caoduong, new species, holotype, female. 1, Head in frontal view; 2, Vertex in dorsal view; 3, Mesosoma in dorsal view; 4, Mesosoma in lateral view; 5, Propodeum in posterior view; 6, Tergum I in dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5345182" httpUri="https://zenodo.org/record/5345182/files/figure.png" pageId="2" pageNumber="590">Fig. 1</figureCitation>
|
||
); width of emargination much less than ⅓ width of clypeus between inner eye margins. Mandible with prominent five teeth, the first and second teeth from base denticle at inner margin. Antennal scape long, about 4.5 times as long as its maximum width; flagellomere I 2.0 times as long as wide, flagellomeres II–IV slightly longer than wide, flagellomeres V–VIII as long as wide, flagellomere IX wider than long, terminal flagellomere bullet-shaped, longer than wide.
|
||
</paragraph>
|
||
<paragraph id="8BE66D43AD59FFD2D508FBAC386CF9DD" blockId="2.[812,1452,1117,1621]" pageId="2" pageNumber="590">
|
||
Mesosoma longer than wide in dorsal view (
|
||
<figureCitation id="136271C6AD59FFD2D2DBFBAF3811FBFD" box="[1278,1342,1117,1141]" captionStart="Figs" captionStartId="1.[136,180,1922,1944]" captionTargetBox="[136,1474,300,1906]" captionTargetId="figure-7@1.[136,1474,300,1906]" captionTargetPageId="1" captionText="Figs. 1–6. Pareumenes caoduong, new species, holotype, female. 1, Head in frontal view; 2, Vertex in dorsal view; 3, Mesosoma in dorsal view; 4, Mesosoma in lateral view; 5, Propodeum in posterior view; 6, Tergum I in dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5345182" httpUri="https://zenodo.org/record/5345182/files/figure.png" pageId="2" pageNumber="590">Fig. 3</figureCitation>
|
||
). Pronotal carina strongly raised throughout, reaching ventral corner of pronotum. Mesoscutum in lateral view weakly convex (
|
||
<figureCitation id="136271C6AD59FFD2D3A5FB6F3E12FB5D" captionStart="Figs" captionStartId="1.[136,180,1922,1944]" captionTargetBox="[136,1474,300,1906]" captionTargetId="figure-7@1.[136,1474,300,1906]" captionTargetPageId="1" captionText="Figs. 1–6. Pareumenes caoduong, new species, holotype, female. 1, Head in frontal view; 2, Vertex in dorsal view; 3, Mesosoma in dorsal view; 4, Mesosoma in lateral view; 5, Propodeum in posterior view; 6, Tergum I in dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5345182" httpUri="https://zenodo.org/record/5345182/files/figure.png" pageId="2" pageNumber="590">Fig. 4</figureCitation>
|
||
), in dorsal view almost as long as wide between tegulae, with prescutal furrows near apical margin (
|
||
<figureCitation id="136271C6AD59FFD2D326FB2F3869FB7D" box="[1283,1350,1245,1269]" captionStart="Figs" captionStartId="1.[136,180,1922,1944]" captionTargetBox="[136,1474,300,1906]" captionTargetId="figure-7@1.[136,1474,300,1906]" captionTargetPageId="1" captionText="Figs. 1–6. Pareumenes caoduong, new species, holotype, female. 1, Head in frontal view; 2, Vertex in dorsal view; 3, Mesosoma in dorsal view; 4, Mesosoma in lateral view; 5, Propodeum in posterior view; 6, Tergum I in dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5345182" httpUri="https://zenodo.org/record/5345182/files/figure.png" pageId="2" pageNumber="590">Fig. 3</figureCitation>
|
||
). Disc of scutellum slightly convex, with a short medial longitudinal depression at base, in lateral view at the same level of apical margin of mesoscutum. Metanotum convex, sloping down to apical margin. Propodeum deeply excavated medially at apex, with the excavation wide, about one-third the width of propodeum, basal triangular area with longitudinal basal fovea, with median carina runs from half of the length of propodeum to apical margin (
|
||
<figureCitation id="136271C6AD59FFD2D24BFA2F399FFA7D" box="[1134,1200,1501,1525]" captionStart="Figs" captionStartId="1.[136,180,1922,1944]" captionTargetBox="[136,1474,300,1906]" captionTargetId="figure-7@1.[136,1474,300,1906]" captionTargetPageId="1" captionText="Figs. 1–6. Pareumenes caoduong, new species, holotype, female. 1, Head in frontal view; 2, Vertex in dorsal view; 3, Mesosoma in dorsal view; 4, Mesosoma in lateral view; 5, Propodeum in posterior view; 6, Tergum I in dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5345182" httpUri="https://zenodo.org/record/5345182/files/figure.png" pageId="2" pageNumber="590">Fig. 5</figureCitation>
|
||
), lateral side shallowly emarginate at apex; border between dorsal and lateral faces sharply angled. Forewing with parastigma slightly longer than stigma measured along ventral part (
|
||
<figureCitation id="136271C6AD59FFD2D2D6F9CF381AF9DD" box="[1267,1333,1597,1621]" captionStart="Figs" captionStartId="3.[136,180,2022,2044]" captionTargetBox="[136,1474,231,2007]" captionTargetId="figure-47@3.[136,1474,231,2007]" captionTargetPageId="3" captionText="Figs. 7–12. Pareumenes caoduong, new species, holotype, female (7–9). 7, Metasomal terga in lateral view; 8, Forewing; 9, Habitus. Pareumenes quadrispinosus acutus Liu, 1941, female (10–12). 10, Head in frontal view; 11, Forewing; 12, Habitus. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5345186" httpUri="https://zenodo.org/record/5345186/files/figure.png" pageId="2" pageNumber="590">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE66D43AD59FFD2D508F98F39B7F81D" blockId="2.[811,1453,1661,1941]" pageId="2" pageNumber="590">
|
||
Metasomal tergum I in dorsal view 1.6 times as long as wide, with the strongly projecting spiracles at about one-third of the length of the petiole basally (
|
||
<figureCitation id="136271C6AD59FFD2D291F94F39D8F95D" box="[1204,1271,1725,1749]" captionStart="Figs" captionStartId="1.[136,180,1922,1944]" captionTargetBox="[136,1474,300,1906]" captionTargetId="figure-7@1.[136,1474,300,1906]" captionTargetPageId="1" captionText="Figs. 1–6. Pareumenes caoduong, new species, holotype, female. 1, Head in frontal view; 2, Vertex in dorsal view; 3, Mesosoma in dorsal view; 4, Mesosoma in lateral view; 5, Propodeum in posterior view; 6, Tergum I in dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5345182" httpUri="https://zenodo.org/record/5345182/files/figure.png" pageId="2" pageNumber="590">Fig. 6</figureCitation>
|
||
), in lateral view abruptly produced near base, then depressed and gradually convex to near apical margin (
|
||
<figureCitation id="136271C6AD59FFD2D25EF90F3992F89D" box="[1147,1213,1789,1813]" captionStart="Figs" captionStartId="3.[136,180,2022,2044]" captionTargetBox="[136,1474,231,2007]" captionTargetId="figure-47@3.[136,1474,231,2007]" captionTargetPageId="3" captionText="Figs. 7–12. Pareumenes caoduong, new species, holotype, female (7–9). 7, Metasomal terga in lateral view; 8, Forewing; 9, Habitus. Pareumenes quadrispinosus acutus Liu, 1941, female (10–12). 10, Head in frontal view; 11, Forewing; 12, Habitus. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5345186" httpUri="https://zenodo.org/record/5345186/files/figure.png" pageId="2" pageNumber="590">Fig. 7</figureCitation>
|
||
), much narrower than tergum II, with a fovea near apical margin; tergum II in dorsal view nearly as long as wide; sternum II in lateral view gradually and slightly convex at basal two-third, then straight to apical margin (
|
||
<figureCitation id="136271C6AD59FFD2D262F88F39A4F81D" box="[1095,1163,1917,1941]" captionStart="Figs" captionStartId="3.[136,180,2022,2044]" captionTargetBox="[136,1474,231,2007]" captionTargetId="figure-47@3.[136,1474,231,2007]" captionTargetPageId="3" captionText="Figs. 7–12. Pareumenes caoduong, new species, holotype, female (7–9). 7, Metasomal terga in lateral view; 8, Forewing; 9, Habitus. Pareumenes quadrispinosus acutus Liu, 1941, female (10–12). 10, Head in frontal view; 11, Forewing; 12, Habitus. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5345186" httpUri="https://zenodo.org/record/5345186/files/figure.png" pageId="2" pageNumber="590">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE66D43AD59FFD4D508F84F3FC9FBDD" blockId="2.[811,1451,1981,2069]" lastBlockId="4.[111,753,189,1109]" lastPageId="4" lastPageNumber="592" pageId="2" pageNumber="590">
|
||
Body covered with golden setae. Clypeus covered with sparse and strong punctures, each bearing a short seta, punctures at centre larger than at sides, space between punctures larger than puncture diameter except some punctures at middle with space between punctures smaller than puncture diameter, punctures near apical margin smaller but denser. Mandible with several shallow small punctures. Frons densely covered with coarse punctures. Vertex and gena with strong punctures, space between punctures smooth. Pronotum with punctures coarser than those on frons, spaces between punctures not raised, anterior face smooth. Mesoscutum densely covered with strong and flat-bottomed punctures except area between furrow with sparse punctures (
|
||
<figureCitation id="136271C6AD5FFFD4D4BAFE2F3FCEFE7D" box="[671,737,477,501]" captionStart="Figs" captionStartId="1.[136,180,1922,1944]" captionTargetBox="[136,1474,300,1906]" captionTargetId="figure-7@1.[136,1474,300,1906]" captionTargetPageId="1" captionText="Figs. 1–6. Pareumenes caoduong, new species, holotype, female. 1, Head in frontal view; 2, Vertex in dorsal view; 3, Mesosoma in dorsal view; 4, Mesosoma in lateral view; 5, Propodeum in posterior view; 6, Tergum I in dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5345182" httpUri="https://zenodo.org/record/5345182/files/figure.png" pageId="4" pageNumber="592">Fig. 3</figureCitation>
|
||
), punctures at margins sparser. Punctures on scutellum similar to those on mesoscutum. Punctures on metanotum larger than those on scutellum. Mesepisternum with punctures on upper dorsal part very coarse, space between punctures strongly raised to form reticulation, with strong and large punctures posterodorsally, several shallow and small punctures anteroventrally; border between posterodorsal and anteroventral parts indistinct. Metapleuron almost smooth. Propodeum with strong punctures and some large smooth area on dorsal surface, posterior surface with coarse punctures in between strong punctures, some transverse striations in the middle half at apex; lateral surface with striations and sparse punctures. Metasomal tergum I densely covered with dense and coarse punctures but punctures smaller than those on pronotum, punctures on tergum II much sparser and weaker than those on tergum I, punctures on terga III–V much smaller and weaker than those on tergum II; tergum VI with minute punctures; punctures on sternum II stronger and larger than those on sterna III–V, sternum VI with minute punctures.
|
||
</paragraph>
|
||
<caption id="DF263DCBAD58FFD3D6ADF81438B7F791" ID-DOI="http://doi.org/10.5281/zenodo.5345186" ID-Zenodo-Dep="5345186" httpUri="https://zenodo.org/record/5345186/files/figure.png" pageId="3" pageNumber="591" startId="3.[136,180,2022,2044]" targetBox="[136,1474,231,2007]" targetPageId="3">
|
||
<paragraph id="8BE66D43AD58FFD3D6ADF81438B7F791" blockId="3.[136,1475,2022,2073]" pageId="3" pageNumber="591">
|
||
Figs. 7–12.
|
||
<taxonomicName id="4C5916C0AD58FFD3D725F8153CF0F873" authority="Nguyen, 2020" authorityName="Nguyen" authorityYear="2020" box="[256,479,2022,2044]" class="Insecta" family="Vespidae" genus="Pareumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="caoduong" status="sp. nov.">
|
||
<emphasis id="B92DB151AD58FFD3D725F8153CF0F873" box="[256,479,2022,2044]" italics="true" pageId="3" pageNumber="591">Pareumenes caoduong</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A21E0C2AAD58FFD3D7CBF8153F48F874" box="[494,615,2022,2044]" pageId="3" pageNumber="591" rank="species">new species</taxonomicNameLabel>
|
||
, holotype, female (7–9). 7, Metasomal terga in lateral view; 8, Forewing; 9, Habitus.
|
||
<taxonomicName id="4C5916C0AD58FFD3D6ADF7F63F65F790" authority="Liu, 1941" authorityName="Liu" authorityYear="1941" box="[136,586,2051,2073]" class="Insecta" family="Vespidae" genus="Pareumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="591" phylum="Arthropoda" rank="subSpecies" species="quadrispinosus" subSpecies="acutus">
|
||
<emphasis id="B92DB151AD58FFD3D6ADF7F63CCFF791" box="[136,480,2051,2073]" italics="true" pageId="3" pageNumber="591">Pareumenes quadrispinosus acutus</emphasis>
|
||
<bibRefCitation id="EFC810B2AD58FFD3D7CCF7F13F65F790" author="Liu CL" box="[489,586,2051,2073]" pageId="3" pageNumber="591" pagination="245 - 289" refId="ref4862" refString="Liu CL (1941) Revisional studies of the Vespidae of China. I. The genus Pareumenes Saussure, with description of six new species (Hemenoptera: Eumeninae). Notes d'Entomologie Chinoise, 8 (6): 245 - 289, 2 pls." type="journal article" year="1941">Liu, 1941</bibRefCitation>
|
||
</taxonomicName>
|
||
, female (10–12). 10, Head in frontal view; 11, Forewing; 12, Habitus. Scale = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE66D43AD5FFFD4D654FB8F3C1AFA1D" blockId="4.[113,753,1149,1429]" pageId="4" pageNumber="592">Colour. Black; following parts yellow: an irregular mark at the middle of clypeus, large spot between antennal sockets, antennal scape beneath, two thin short bands at dorsal part of pronotum near pronotal carina (almost touching each other), apical margin of valvulae, two large spots at lateral sides near apex of tergum I, a short apical band on tergum III medially. Legs almost black except some brown parts at apical half of middle and hind tibiae. Wings infuscate, veins dark brown.</paragraph>
|
||
<paragraph id="8BE66D43AD5FFFD4D654FA4F3C08FA5D" blockId="4.[113,295,1469,1493]" box="[113,295,1469,1493]" pageId="4" pageNumber="592">Male. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3433EC8AD5FFFD4D654FA0F3F34F99D" box="[113,539,1533,1557]" pageId="4" pageNumber="592" type="distribution">
|
||
<paragraph id="8BE66D43AD5FFFD4D654FA0F3F34F99D" blockId="4.[113,539,1533,1557]" box="[113,539,1533,1557]" pageId="4" pageNumber="592">
|
||
<emphasis id="B92DB151AD5FFFD4D654FA0F3C2CF99D" bold="true" box="[113,259,1533,1557]" pageId="4" pageNumber="592">Distribution.</emphasis>
|
||
<collectingCountry id="F34E2DD3AD5FFFD4D72EFA0F3C48F99D" box="[267,359,1533,1557]" name="Vietnam" pageId="4" pageNumber="592">Vietnam</collectingCountry>
|
||
(Northern part).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3433EC8AD5FFFD4D654F9CF3CE6F91D" pageId="4" pageNumber="592" type="etymology">
|
||
<paragraph id="8BE66D43AD5FFFD4D654F9CF3CE6F91D" blockId="4.[113,751,1597,1685]" pageId="4" pageNumber="592">
|
||
<emphasis id="B92DB151AD5FFFD4D654F9CF3DDCF9DD" bold="true" box="[113,243,1597,1621]" pageId="4" pageNumber="592">Etymology.</emphasis>
|
||
The specific name refers to the
|
||
<typeStatus id="54E2D3E1AD5FFFD4D445F9CC3FA1F9DE" box="[608,654,1598,1622]" pageId="4" pageNumber="592">type</typeStatus>
|
||
locality, Cao Duong,
|
||
<collectingRegion id="499DA3A1AD5FFFD4D6D9F9AC3CDAF9FD" box="[252,501,1629,1654]" country="Vietnam" name="Tuyen Quang" pageId="4" pageNumber="592">Tuyen Quang province</collectingRegion>
|
||
of
|
||
<collectingCountry id="F34E2DD3AD5FFFD4D439F9AF3F54F9FD" box="[540,635,1629,1653]" name="Vietnam" pageId="4" pageNumber="592">Vietnam</collectingCountry>
|
||
; it is to be treated as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3433EC8AD5FFFD4D654F94F39DAFE1D" pageId="4" pageNumber="592" type="discussion">
|
||
<paragraph id="8BE66D43AD5FFFD4D654F94F39DAFE1D" blockId="4.[112,752,1725,2069]" lastBlockId="4.[813,1452,189,405]" pageId="4" pageNumber="592">
|
||
<emphasis id="B92DB151AD5FFFD4D654F94F3DF0F95D" bold="true" box="[113,223,1725,1749]" pageId="4" pageNumber="592">Remarks.</emphasis>
|
||
This species comes close to
|
||
<emphasis id="B92DB151AD5FFFD4D423F94C3FC1F95D" box="[518,750,1726,1749]" italics="true" pageId="4" pageNumber="592">
|
||
<taxonomicName id="4C5916C0AD5FFFD4D423F94C3FA6F95D" authorityName="de Saussure" authorityYear="1855" box="[518,649,1726,1749]" class="Insecta" family="Vespidae" genus="Pareumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="592" phylum="Arthropoda" rank="genus">Pareumenes</taxonomicName>
|
||
taiwanus
|
||
</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC810B2AD5FFFD4D65DF92F3C27F97D" author="Sonan J" box="[120,264,1757,1781]" pageId="4" pageNumber="592" pagination="14 - 16" refId="ref5185" refString="Sonan J (1937) On two Formosan species of the genus Montezumia Saussure. Transactions of the Natural History Society Formosa, 27: 14 - 16." type="journal article" year="1937">Sonan, 1937</bibRefCitation>
|
||
) in having coarse punctures on mesosoma and vertex, tergum I coarsely punctate, petiolate, gradually widening to apex, but can be distinguished from the latter by having clypeus with sparse and strong punctures, border between punctures smooth (vs. clypeus with coarse and dense punctures, border between punctures raised to form longitudinal reticulation in
|
||
<emphasis id="B92DB151AD5FFFD4D7AEF86C3F28F83D" box="[395,519,1950,1973]" italics="true" pageId="4" pageNumber="592">P. taiwanus</emphasis>
|
||
), frontal view slightly wider than high (vs. as wide as high in
|
||
<emphasis id="B92DB151AD5FFFD4D432F84C3FBAF85D" box="[535,661,1982,2005]" italics="true" pageId="4" pageNumber="592">P. taiwanus</emphasis>
|
||
); vertex with two foveae close to each other (vs. vertex with a median fovea in
|
||
<emphasis id="B92DB151AD5FFFD4D6F6F80C3C7EF79D" box="[211,337,2046,2069]" italics="true" pageId="4" pageNumber="592">P. taiwanus</emphasis>
|
||
); lateral side of propodeum shallowly emarginate at apex (vs. lateral side of propodeum deeply emarginate at apex in
|
||
<emphasis id="B92DB151AD5FFFD4D231FF2C39BEFF7D" box="[1044,1169,222,245]" italics="true" pageId="4" pageNumber="592">P. taiwanus</emphasis>
|
||
); and colouration: clypeus black with irregular yellow mark in the middle (vs. clypeus entirely yellow in
|
||
<emphasis id="B92DB151AD5FFFD4D5D2FEEC3958FEBD" box="[1015,1143,286,309]" italics="true" pageId="4" pageNumber="592">P. taiwanus</emphasis>
|
||
); two short and thin yellow bands on pronotum (vs. two large spots on pronotum in
|
||
<emphasis id="B92DB151AD5FFFD4D3B1FECC3EA0FEFD" italics="true" pageId="4" pageNumber="592">P. taiwanus</emphasis>
|
||
); tergum II entirely black (vs. tergum II with a broad apical yellow band in
|
||
<emphasis id="B92DB151AD5FFFD4D242FE8C39CAFE1D" box="[1127,1253,382,405]" italics="true" pageId="4" pageNumber="592">P. taiwanus</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |