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<mods:title id="EEC91F3D0CF03468797CD575C585594E">Revision of the Aspistomella group of genera (Diptera: Ulidiidae: Pterocallinae: Lipsanini)</mods:title>
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<mods:namePart id="AD074F2AEA788F86E4302D84830E50B2">Kovac, Damir</mods:namePart>
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<mods:namePart id="D71B5C69A756C2B7776A753557F32BD5">Kameneva, Elena P.</mods:namePart>
<mods:affiliation id="C1FBEF9A4FFBFF192129D3DA60664102">I. I. Schmalhausen Institute of Zoology National Academy of Sciences of Ukraine, Bogdan Chmielnicki St. 15, 01054 Kyiv, Ukraine https: // orcid. org / 0000 - 0002 - 7986 - 9942 &amp; Museum für Naturkunde, Invalidenstr. 43, Berlin, 10115 Germany Corresponding author: valery. korneyev @ gmail. com; https: // orcid. org / 0000 - 0001 - 9631 - 1038</mods:affiliation>
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<mods:namePart id="231508A5EEE46E97786BB4DBF158F37E">Korneyev, Valery A.</mods:namePart>
<mods:affiliation id="64ABCAC56614B9A0E35CFA81E5D5D285">I. I. Schmalhausen Institute of Zoology National Academy of Sciences of Ukraine, Bogdan Chmielnicki St. 15, 01054 Kyiv, Ukraine https: // orcid. org / 0000 - 0002 - 7986 - 9942 &amp; Museum für Naturkunde, Invalidenstr. 43, Berlin, 10115 Germany Corresponding author: valery. korneyev @ gmail. com; https: // orcid. org / 0000 - 0001 - 9631 - 1038</mods:affiliation>
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<emphasis id="B92D442C1E01FF95FF215606FE7FF8A6" bold="true" box="[151,421,1867,1894]" italics="true" pageId="97" pageNumber="98">Ulivellia pseudinsolita</emphasis>
Kameneva &amp; V. Korneyev
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<figureCitation id="136284BB1E01FF95FF2156D9FF2DF86E" box="[151,247,1940,1966]" captionStart="FIGURE 4" captionStartId="11.[151,250,1686,1710]" captionTargetBox="[247,1341,187,1657]" captionTargetId="figure-20@11.[241,1346,181,1662]" captionTargetPageId="11" captionText="FIGURE 4. Ulivellia, Polyteloptera and similar genera, wing. A, U. amnoni sp. nov.; B, U. inversa Speiser; C, U. laetitiae sp. nov.; DE, U. arcuata sp. nov. (D—male, E—female); F, U. tenoris sp. nov.; G, U. pseudinsolita sp. nov.; H, P. apotropa Hendel; I, “Euxesta” insolita Hendel; J, Gen. sp. near insolita; K, Axiologina ferrumequinum Hendel. Abbreviations. Veins and cells: bm—basal medial cell; br—basal radial cell; c—costal cell; C—costal vein; cua—anterior cubital cell; CuA+CuP— anterior branch of cubital vein + posterior branch of cubital vein; dm—discal medial cell; dm-m—discal medial crossvein; h—humeral crossvein; M1—first branch of media; m1—first medial cell; M4—fourth branch of media; m4—fourth medial cell; ptstg—pterostigma; R1—anterior branch of radius; r1—first radial cell; r2+3—second + third radial cell; R2+3—second branch of radius; R 4+5 —third branch of radius; r 4+5 —fourth + fifth radial cell; r-m—radial-medial crossvein; Sc—subcostal vein. Wing pattern: DB = discal band; HB = humeral band; AAB = anterior apical band; SAB = subapical band; SCB = subcostal band. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023159" httpUri="https://zenodo.org/record/14023159/files/figure.png" pageId="97" pageNumber="98">Figs 4 G</figureCitation>
,
<figureCitation id="136284BB1E01FF95FEB056DEFE96F86E" box="[262,332,1939,1966]" captionStart-0="FIGURE 55" captionStart-1="FIGURE 56" captionStartId-0="99.[151,250,1375,1399]" captionStartId-1="100.[151,250,1759,1783]" captionTargetBox-0="[151,1434,181,1345]" captionTargetBox-1="[158,1425,188,1728]" captionTargetId-0="figure-20@99.[151,1435,181,1351]" captionTargetId-1="figure-30@100.[151,1436,181,1735]" captionTargetPageId-0="99" captionTargetPageId-1="100" captionText-0="FIGURE 55. Ulivellia pseudinsolita sp. nov. holotype ♂. A, habitus, left and dorsal; B, same, dorsal; C, head, anterior; D, same, left; E, mesonotum, dorsal; F, wing, anterobasal part; G, labels. Scale bar 2 mm." captionText-1="FIGURE 56. Ulivellia pseudinsolita sp. nov. holotype ♂. A, postabdomen, posterior; B C, same, ventral; D, same, ventrolateral; E, surstyli and hypandrium, ventral; F, preabdomen, ventral. Abbreviations: cerc—cerci; ej apod—ejaculatory apodeme; epand—epandrium; ph g—phallic guide; ph—phallus; pregt—pregonite; p l sur—posterior lobe of surstylus; v l sur—ventral lobe of surstylus; st 2— second sternite; st 3—third sternite; st 4—fourth sternite; st 5—fifth sternite." figureDoi-0="http://doi.org/10.5281/zenodo.14023310" figureDoi-1="http://doi.org/10.5281/zenodo.14023312" httpUri-0="https://zenodo.org/record/14023310/files/figure.png" httpUri-1="https://zenodo.org/record/14023312/files/figure.png" pageId="97" pageNumber="98">5556</figureCitation>
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<emphasis id="B92D442C1E02FF96FF2151DBFE1DFF71" bold="true" box="[151,455,150,177]" pageId="98" pageNumber="99">Material examined. Type.</emphasis>
<materialsCitation id="3B3192631E02FF96FE7851DAFE1DFF39" collectingDate="1945-02" collectionCode="AMNH" collectorName="S. Cath. &amp; A. Maler" country="Brazil" location="Santa Catarina" pageId="98" pageNumber="99" specimenCount="1" specimenCount-male="1" stateProvince="Santa Catarina" typeStatus="holotype">
<typeStatus id="54E2269C1E02FF96FE7851DAFDECFF71" box="[462,566,151,177]" pageId="98" pageNumber="99" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="F34ED8AE1E02FF96FDD651DBFD73FF70" box="[608,681,150,176]" name="Brazil" pageId="98" pageNumber="99">Brazil</collectingCountry>
</emphasis>
: “Corupa [Corupá],
<collectorName id="26ACFDE81E02FF96FC3951DAFC32FF71" box="[911,1000,151,177]" pageId="98" pageNumber="99">S. Cath.</collectorName>
[
<collectingRegion id="499D56DC1E02FF96FC5851DAFB42FF71" box="[1006,1176,151,177]" country="Brazil" name="Santa Catarina" pageId="98" pageNumber="99">Santa Catarina</collectingRegion>
] / (Hansa Humbolt) /
<date id="FFE7BEFE1E02FF96FA3C51DAFF09FF15" pageId="98" pageNumber="99" value="1945-02">
<collectingDate id="EFA347161E02FF96FA3C51DAFF09FF15" pageId="98" pageNumber="99" value="1945-02">II 1945</collectingDate>
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”, “
<collectorName id="26ACFDE81E02FF96FF4051F6FE85FF15" box="[246,351,187,213]" pageId="98" pageNumber="99">A. Maler</collectorName>
, Coll. / Frank Johnson / Donor” (abdomen dissected and attached in a plastic tube with glycerin under specimen) (
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).
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</paragraph>
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<emphasis id="B92D442C1E02FF96FF71504FFEE0FEDC" bold="true" box="[199,314,258,284]" pageId="98" pageNumber="99">Diagnosis</emphasis>
.
<taxonomicName id="4C59E3BD1E02FF96FEFC504EFD91FEDD" authorityName="Kovac &amp; Kameneva &amp; Korneyev &amp; Araújo &amp; Savaris &amp; Smit &amp; Schneider &amp; Schreiber &amp; Korneyev" authorityYear="2024" box="[330,587,259,285]" class="Insecta" family="Ulidiidae" genus="Ulivellia" kingdom="Animalia" order="Diptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="pseudinsolita">
<emphasis id="B92D442C1E02FF96FEFC504EFD91FEDD" box="[330,587,259,285]" italics="true" pageId="98" pageNumber="99">Ulivellia pseudinsolita</emphasis>
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differs from other species of the genus
<taxonomicName id="4C59E3BD1E02FF96FBAE504EFBA0FEDD" authorityName="Speiser" authorityYear="1929" box="[1048,1146,259,285]" class="Insecta" family="Ulidiidae" genus="Ulivellia" kingdom="Animalia" order="Diptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="B92D442C1E02FF96FBAE504EFBA0FEDD" box="[1048,1146,259,285]" italics="true" pageId="98" pageNumber="99">Ulivellia</emphasis>
</taxonomicName>
in having a wing pattern with oblique discal crossband, prescutellar acrostichal setae present, halter and legs entirely black; it most closely resembles
<taxonomicName id="4C59E3BD1E02FF96FEB85006FEA5FEA4" authorityName="Kovac &amp; Kameneva &amp; Korneyev &amp; Araújo &amp; Savaris &amp; Smit &amp; Schneider &amp; Schreiber &amp; Korneyev" authorityYear="2024" box="[270,383,331,356]" class="Insecta" family="Ulidiidae" genus="Ulivellia" kingdom="Animalia" order="Diptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="tenoris">
<emphasis id="B92D442C1E02FF96FEB85006FEA5FEA4" box="[270,383,331,356]" italics="true" pageId="98" pageNumber="99">U. tenoris</emphasis>
</taxonomicName>
in having wing pattern with four narrow brown crossbands, of which discal and preapical are oblique and clearly diverging posteriorly, differing from it by the wider wing and the hyaline interspace between discal and preapical crossbands narrower, colouration of legs and halters and presence / absence of acrostichal setae (see the key). Both species have a wing pattern similar to that of
<taxonomicName id="4C59E3BD1E02FF96FC3750FAFB0BFE11" authority="Hendel 1909" authorityName="Hendel" authorityYear="1909" box="[897,1233,439,465]" class="Insecta" family="Ulidiidae" genus="Euxesta" kingdom="Animalia" order="Diptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="insolita">
<emphasis id="B92D442C1E02FF96FC3750FAFBEDFE11" box="[897,1079,439,465]" italics="true" pageId="98" pageNumber="99">Euxesta insolita</emphasis>
Hendel 1909
</taxonomicName>
(
<figureCitation id="136284BB1E02FF96FB5750FAFAE3FE11" box="[1249,1337,439,465]" captionStart="FIGURE 4" captionStartId="11.[151,250,1686,1710]" captionTargetBox="[247,1341,187,1657]" captionTargetId="figure-20@11.[241,1346,181,1662]" captionTargetPageId="11" captionText="FIGURE 4. Ulivellia, Polyteloptera and similar genera, wing. A, U. amnoni sp. nov.; B, U. inversa Speiser; C, U. laetitiae sp. nov.; DE, U. arcuata sp. nov. (D—male, E—female); F, U. tenoris sp. nov.; G, U. pseudinsolita sp. nov.; H, P. apotropa Hendel; I, “Euxesta” insolita Hendel; J, Gen. sp. near insolita; K, Axiologina ferrumequinum Hendel. Abbreviations. Veins and cells: bm—basal medial cell; br—basal radial cell; c—costal cell; C—costal vein; cua—anterior cubital cell; CuA+CuP— anterior branch of cubital vein + posterior branch of cubital vein; dm—discal medial cell; dm-m—discal medial crossvein; h—humeral crossvein; M1—first branch of media; m1—first medial cell; M4—fourth branch of media; m4—fourth medial cell; ptstg—pterostigma; R1—anterior branch of radius; r1—first radial cell; r2+3—second + third radial cell; R2+3—second branch of radius; R 4+5 —third branch of radius; r 4+5 —fourth + fifth radial cell; r-m—radial-medial crossvein; Sc—subcostal vein. Wing pattern: DB = discal band; HB = humeral band; AAB = anterior apical band; SAB = subapical band; SCB = subcostal band. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023159" httpUri="https://zenodo.org/record/14023159/files/figure.png" pageId="98" pageNumber="99">Fig. 4 I</figureCitation>
), clearly differing by the at least partly hyaline or yellow wing base (in
<taxonomicName id="4C59E3BD1E02FF96FCF15096FC60FE35" box="[839,954,475,501]" class="Insecta" family="Ulidiidae" genus="Euxesta" kingdom="Animalia" order="Diptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="insolita">
<emphasis id="B92D442C1E02FF96FCF15096FC60FE35" box="[839,954,475,501]" italics="true" pageId="98" pageNumber="99">E. insolita</emphasis>
</taxonomicName>
completely black); as well as by the almost twice larger size (in
<taxonomicName id="4C59E3BD1E02FF96FECD50B2FE34FDD9" box="[379,494,511,537]" class="Insecta" family="Ulidiidae" genus="Euxesta" kingdom="Animalia" order="Diptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="insolita">
<emphasis id="B92D442C1E02FF96FECD50B2FE34FDD9" box="[379,494,511,537]" italics="true" pageId="98" pageNumber="99">E. insolita</emphasis>
</taxonomicName>
wing length = 3.0
<quantity id="4CA135DB1E02FF96FD7C50B3FCF9FDD9" box="[714,803,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="98" pageNumber="99" unit="mm" value="3.5">3.5 mm</quantity>
), and the epistome / clypeus sizes.
</paragraph>
</subSubSection>
<subSubSection id="C343CBB51E02FF91FF71536FFE4FFF71" lastPageId="101" lastPageNumber="102" pageId="98" pageNumber="99" type="description">
<paragraph id="8BE6983E1E02FF96FF71536FFB2BFBBD" blockId="98.[151,1437,150,2013]" pageId="98" pageNumber="99">
<emphasis id="B92D442C1E02FF96FF71536FFE8AFDFC" bold="true" box="[199,336,546,572]" pageId="98" pageNumber="99">Description</emphasis>
. Male. Head (
<figureCitation id="136284BB1E02FF96FE58536EFDA4FDFD" box="[494,638,546,573]" captionStart="FIGURE 55" captionStartId="99.[151,250,1375,1399]" captionTargetBox="[151,1434,181,1345]" captionTargetId="figure-20@99.[151,1435,181,1351]" captionTargetPageId="99" captionText="FIGURE 55. Ulivellia pseudinsolita sp. nov. holotype ♂. A, habitus, left and dorsal; B, same, dorsal; C, head, anterior; D, same, left; E, mesonotum, dorsal; F, wing, anterobasal part; G, labels. Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14023310" httpUri="https://zenodo.org/record/14023310/files/figure.png" pageId="98" pageNumber="99">Figs 55 CD</figureCitation>
) ratio (length: height: width) = 1: 1.4: 1.7, frons, parafacial, and gena yellowish brown, face and occiput black. Frons shiny brown, narrowed posteriorly (
<figureCitation id="136284BB1E02FF96FBF1530AFB6EFDA1" box="[1095,1204,582,609]" captionStart="FIGURE 55" captionStartId="99.[151,250,1375,1399]" captionTargetBox="[151,1434,181,1345]" captionTargetId="figure-20@99.[151,1435,181,1351]" captionTargetPageId="99" captionText="FIGURE 55. Ulivellia pseudinsolita sp. nov. holotype ♂. A, habitus, left and dorsal; B, same, dorsal; C, head, anterior; D, same, left; E, mesonotum, dorsal; F, wing, anterobasal part; G, labels. Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14023310" httpUri="https://zenodo.org/record/14023310/files/figure.png" pageId="98" pageNumber="99">Fig. 55 C</figureCitation>
), 1.2× as long (from the lunule to inner vertical seta) as wide at lunule and 1.5× longer than wide at vertex or 0.85× as long (from the lunule to anterior ocellus) as wide at lunule and 1.1× longer than wide at vertex, with metallic shining, brown or black ocellar triangle and vertical plates; orbits, vertex, gena and occiput silver-white microtrichose; parafacial narrow, 0.3× broader than postpedicel, sparsely white microtrichose. Frontal plates with 68 proclinate parafrontal setulae and 67 moderately strong, frontal setae and the frontal vitta with 3 interfrontal setae on each side, anteriormost crossing, longer than other setulae and half as long as outer vertical seta; 1 very long orbital seta 1.2× longer than ocellar seta, 0.8× longer than inner vertical seta and twice as long as outer vertical seta and 1 reclinate anterior orbital seta as long as longest frontal setae; postocellar seta long, 0.7× longer than inner vertical seta; outer vertical seta short, 0.4× longer than inner vertical seta and twice as long as postocular setae forming one row, allied by 1520 dorsal and 1825 ventral occipital setae. Lunule indistinguishable. Eye 1.6× higher than long. Face above transverse fold 0.6× higher than wide at fold, brown, white microtrichose; epistome entirely black or dark brown, sparsely white microtrichose, with metallic cyan sheen, strongly produced anteriorly. Clypeus shiny brown, sparsely white microtrichose, 0.3× higher than epistome. Gena brownish yellow, with long genal seta and 45 peristomal setae anterior of it, 0.40.5× longer than genal seta. Occiput black, except postgena partly brown, with cyan metallic sheen, greyish microtrichose; ventral part of occiput (postgena) with 57 setae almost as long as genal seta.
</paragraph>
<paragraph id="8BE6983E1E02FF96FF7155CAFE64FACD" blockId="98.[151,1437,150,2013]" pageId="98" pageNumber="99">Antenna brown; scape and pedicel with black setulae; postpedicel brown, whitish microtrichose, twice as long as wide, apically rounded; arista black to brown at base, almost bare. Mouthparts black, prementum black, with steel sheen. Palp brown, moderately wide crescent, apically rounded, with 912 long black setulae, of them medial setulae longer than others.</paragraph>
<paragraph id="8BE6983E1E02FF96FF71545AFABDFA25" blockId="98.[151,1437,150,2013]" pageId="98" pageNumber="99">
<emphasis id="B92D442C1E02FF96FF71545AFEFAFAF1" bold="true" box="[199,288,1303,1329]" pageId="98" pageNumber="99">Thorax</emphasis>
(
<figureCitation id="136284BB1E02FF96FE86545AFE40FAF1" box="[304,410,1302,1329]" captionStart="FIGURE 55" captionStartId="99.[151,250,1375,1399]" captionTargetBox="[151,1434,181,1345]" captionTargetId="figure-20@99.[151,1435,181,1351]" captionTargetPageId="99" captionText="FIGURE 55. Ulivellia pseudinsolita sp. nov. holotype ♂. A, habitus, left and dorsal; B, same, dorsal; C, head, anterior; D, same, left; E, mesonotum, dorsal; F, wing, anterobasal part; G, labels. Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14023310" httpUri="https://zenodo.org/record/14023310/files/figure.png" pageId="98" pageNumber="99">Fig. 55 E</figureCitation>
) brown to black, with bluish sheen and sparse white microtrichia not obscuring underlying cuticle. Mesonotal scutum 1.1×as long as wide; with brown microtrichia posterolateral to transverse suture, black setulose, with 1216 rows of setulae between rows of dorsocentral setulae; acrostichal prescutellar setae present; prescutellar area setulose behind anterior dorsocentral setae to acrostichal seta. Scutellum dorsally very slightly convex, black, almost matt, sparsely microtrichose, without setulae, laterally with bluish sheen. Subscutellum and mediotergite black, grey microtrichose. Other setae as described for the genus. All the setae and setulae black.
</paragraph>
<paragraph id="8BE6983E1E02FF96FF7154A3FAE2F81D" blockId="98.[151,1437,150,2013]" pageId="98" pageNumber="99">
<emphasis id="B92D442C1E02FF96FF7154A3FED2F9C8" bold="true" box="[199,264,1518,1544]" pageId="98" pageNumber="99">Wing</emphasis>
(
<figureCitation id="136284BB1E02FF96FEA154A2FEA1F9C9" box="[279,379,1519,1545]" captionStart="FIGURE 4" captionStartId="11.[151,250,1686,1710]" captionTargetBox="[247,1341,187,1657]" captionTargetId="figure-20@11.[241,1346,181,1662]" captionTargetPageId="11" captionText="FIGURE 4. Ulivellia, Polyteloptera and similar genera, wing. A, U. amnoni sp. nov.; B, U. inversa Speiser; C, U. laetitiae sp. nov.; DE, U. arcuata sp. nov. (D—male, E—female); F, U. tenoris sp. nov.; G, U. pseudinsolita sp. nov.; H, P. apotropa Hendel; I, “Euxesta” insolita Hendel; J, Gen. sp. near insolita; K, Axiologina ferrumequinum Hendel. Abbreviations. Veins and cells: bm—basal medial cell; br—basal radial cell; c—costal cell; C—costal vein; cua—anterior cubital cell; CuA+CuP— anterior branch of cubital vein + posterior branch of cubital vein; dm—discal medial cell; dm-m—discal medial crossvein; h—humeral crossvein; M1—first branch of media; m1—first medial cell; M4—fourth branch of media; m4—fourth medial cell; ptstg—pterostigma; R1—anterior branch of radius; r1—first radial cell; r2+3—second + third radial cell; R2+3—second branch of radius; R 4+5 —third branch of radius; r 4+5 —fourth + fifth radial cell; r-m—radial-medial crossvein; Sc—subcostal vein. Wing pattern: DB = discal band; HB = humeral band; AAB = anterior apical band; SAB = subapical band; SCB = subcostal band. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023159" httpUri="https://zenodo.org/record/14023159/files/figure.png" pageId="98" pageNumber="99">Figs 4 G</figureCitation>
,
<figureCitation id="136284BB1E02FF96FE3054A3FE21F9C9" box="[390,507,1518,1545]" captionStart="FIGURE 55" captionStartId="99.[151,250,1375,1399]" captionTargetBox="[151,1434,181,1345]" captionTargetId="figure-20@99.[151,1435,181,1351]" captionTargetPageId="99" captionText="FIGURE 55. Ulivellia pseudinsolita sp. nov. holotype ♂. A, habitus, left and dorsal; B, same, dorsal; C, head, anterior; D, same, left; E, mesonotum, dorsal; F, wing, anterobasal part; G, labels. Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14023310" httpUri="https://zenodo.org/record/14023310/files/figure.png" pageId="98" pageNumber="99">55 A, B, F</figureCitation>
)
<quantity id="4CA135DB1E02FF96FDBC54A3FDBAF9C9" box="[522,608,1518,1545]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="98" pageNumber="99" unit="mm" value="5.5">5.5 mm</quantity>
(
<specimenCount id="9D5F53B71E02FF96FDD954A2FD5FF9C9" box="[623,645,1519,1545]" count="1" pageId="98" pageNumber="99" type="male"></specimenCount>
) long, 3× longer than wide; basicostal cell hyaline or yellowish; costal cell 5× longer than wide, slightly widened apically, brown in basal half, with costa very slightly curved posteriorly, but forming neither lobe nor cleft before apex of vein Sc; pterostigma entirely brown, narrow triangular, 4.4× longer than wide, vein R
<subScript id="17DD9A7B1E02FF96FEE85720FEBCF9BB" attach="left" box="[350,358,1645,1659]" fontSize="6" pageId="98" pageNumber="99">1</subScript>
with 1517 setulae dorsal to Sc apex, terminating at level of crossvein dm-m; vein R
<subScript id="17DD9A7B1E02FF96FA9C5720FA99F9BB" attach="left" box="[1322,1347,1645,1659]" fontSize="6" pageId="98" pageNumber="99">2+3</subScript>
weakly arcuate, subparallel to costa, turning towards it before very apex. Crossvein r-m distal to center of cell dm. Cell r
<subScript id="17DD9A7B1E02FF96FF1757F8FF60F903" attach="left" box="[161,186,1717,1731]" fontSize="6" pageId="98" pageNumber="99">4+5</subScript>
5.8× longer than wide, widened distally of crossvein dm-m, narrowed apically. Cell m
<subScript id="17DD9A7B1E02FF96FB1157F8FB75F903" attach="left" box="[1191,1199,1717,1731]" fontSize="6" pageId="98" pageNumber="99">1</subScript>
narrowly triangular; the ultimate section of M
<subScript id="17DD9A7B1E02FF96FE0F5794FE1BF927" attach="left" box="[441,449,1753,1767]" fontSize="6" pageId="98" pageNumber="99">1</subScript>
3.7× longer than crossvein dm-m and 2.8× longer than penultimate section, markedly deepened posteriorly at middle. Vein CuA Z-shaped, forming moderately long posteroapical lobe of cell cua along vein CuP, 1.3× longer than its anterior shoulder. Wing pattern with subbasal crossband from basal half of costal cell through cell cua into alula, wide oblique brown discal crossband from pterostigma to base of cell m
<subScript id="17DD9A7B1E02FF96FB5D5608FB29F893" attach="left" box="[1259,1267,1861,1875]" fontSize="6" pageId="98" pageNumber="99">4</subScript>
except its very base and apex; narrow and oblique preapical crossband from cell r
<subScript id="17DD9A7B1E02FF96FC3A5624FC4EF8B7" attach="left" box="[908,916,1897,1911]" fontSize="6" pageId="98" pageNumber="99">1</subScript>
through crossvein dm-m merging with apical crossband in cell r
<subScript id="17DD9A7B1E02FF96FED356C0FEB7F85B" attach="left" box="[357,365,1933,1947]" fontSize="6" pageId="98" pageNumber="99">1</subScript>
reaching apex of cell m
<subScript id="17DD9A7B1E02FF96FDCB56C0FD5FF85B" attach="left" box="[637,645,1933,1947]" fontSize="6" pageId="98" pageNumber="99">1</subScript>
; hyaline interspace between discal and preapical crossband 1.31.6 as wide as discal crossband in cell dm (
<figureCitation id="136284BB1E02FF96FD9956D2FD56F879" box="[559,652,1951,1977]" captionStart="FIGURE 4" captionStartId="11.[151,250,1686,1710]" captionTargetBox="[247,1341,187,1657]" captionTargetId="figure-20@11.[241,1346,181,1662]" captionTargetPageId="11" captionText="FIGURE 4. Ulivellia, Polyteloptera and similar genera, wing. A, U. amnoni sp. nov.; B, U. inversa Speiser; C, U. laetitiae sp. nov.; DE, U. arcuata sp. nov. (D—male, E—female); F, U. tenoris sp. nov.; G, U. pseudinsolita sp. nov.; H, P. apotropa Hendel; I, “Euxesta” insolita Hendel; J, Gen. sp. near insolita; K, Axiologina ferrumequinum Hendel. Abbreviations. Veins and cells: bm—basal medial cell; br—basal radial cell; c—costal cell; C—costal vein; cua—anterior cubital cell; CuA+CuP— anterior branch of cubital vein + posterior branch of cubital vein; dm—discal medial cell; dm-m—discal medial crossvein; h—humeral crossvein; M1—first branch of media; m1—first medial cell; M4—fourth branch of media; m4—fourth medial cell; ptstg—pterostigma; R1—anterior branch of radius; r1—first radial cell; r2+3—second + third radial cell; R2+3—second branch of radius; R 4+5 —third branch of radius; r 4+5 —fourth + fifth radial cell; r-m—radial-medial crossvein; Sc—subcostal vein. Wing pattern: DB = discal band; HB = humeral band; AAB = anterior apical band; SAB = subapical band; SCB = subcostal band. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023159" httpUri="https://zenodo.org/record/14023159/files/figure.png" pageId="98" pageNumber="99">Fig. 4 G</figureCitation>
,
<figureCitation id="136284BB1E02FF96FD2C56D3FD09F879" box="[666,723,1950,1977]" captionStart="FIGURE 55" captionStartId="99.[151,250,1375,1399]" captionTargetBox="[151,1434,181,1345]" captionTargetId="figure-20@99.[151,1435,181,1351]" captionTargetPageId="99" captionText="FIGURE 55. Ulivellia pseudinsolita sp. nov. holotype ♂. A, habitus, left and dorsal; B, same, dorsal; C, head, anterior; D, same, left; E, mesonotum, dorsal; F, wing, anterobasal part; G, labels. Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14023310" httpUri="https://zenodo.org/record/14023310/files/figure.png" pageId="98" pageNumber="99">55 A</figureCitation>
). Cell cup and anal lobe pale grey, narrower than cell cua. Alula 3× longer than wide, grey in apical 0.50.6. Calypters white, with white cilia. Halter dark brown to black.
</paragraph>
<caption id="DF26C8B61E03FF97FF215412FBD0FA5A" ID-DOI="http://doi.org/10.5281/zenodo.14023310" ID-Zenodo-Dep="14023310" httpUri="https://zenodo.org/record/14023310/files/figure.png" pageId="99" pageNumber="100" startId="99.[151,250,1375,1399]" targetBox="[151,1434,181,1345]" targetPageId="99" targetType="figure">
<paragraph id="8BE6983E1E03FF97FF215412FBD0FA5A" blockId="99.[151,1436,1375,1435]" pageId="99" pageNumber="100">
<emphasis id="B92D442C1E03FF97FF215412FEF8FAB7" bold="true" box="[151,290,1375,1399]" pageId="99" pageNumber="100">FIGURE 55.</emphasis>
<taxonomicName id="4C59E3BD1E03FF97FE9C5412FDC8FAB7" authorityName="Kovac &amp; Kameneva &amp; Korneyev &amp; Araújo &amp; Savaris &amp; Smit &amp; Schneider &amp; Schreiber &amp; Korneyev" authorityYear="2024" box="[298,530,1375,1399]" class="Insecta" family="Ulidiidae" genus="Ulivellia" kingdom="Animalia" order="Diptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="pseudinsolita" status="sp. nov.">
<emphasis id="B92D442C1E03FF97FE9C5412FDC8FAB7" box="[298,530,1375,1399]" italics="true" pageId="99" pageNumber="100">Ulivellia pseudinsolita</emphasis>
</taxonomicName>
<emphasis id="B92D442C1E03FF97FDAD5412FDB7FAB7" bold="true" box="[539,621,1375,1399]" pageId="99" pageNumber="100">
<taxonomicNameLabel id="A21EF9571E03FF97FDAD5412FDB7FAB7" box="[539,621,1375,1399]" pageId="99" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype ♂.
<emphasis id="B92D442C1E03FF97FD4F5412FCD1FAB7" bold="true" box="[761,779,1375,1399]" pageId="99" pageNumber="100">A</emphasis>
, habitus, left and dorsal;
<emphasis id="B92D442C1E03FF97FBA05412FBFDFAB7" bold="true" box="[1046,1063,1375,1399]" pageId="99" pageNumber="100">B</emphasis>
, same, dorsal;
<emphasis id="B92D442C1E03FF97FB705412FB02FAB7" bold="true" box="[1222,1240,1375,1399]" pageId="99" pageNumber="100">C</emphasis>
, head, anterior;
<emphasis id="B92D442C1E03FF97FA325412FA4CFAB7" bold="true" box="[1412,1430,1375,1399]" pageId="99" pageNumber="100">D</emphasis>
, same, left;
<emphasis id="B92D442C1E03FF97FEB154CEFEC2FA5B" bold="true" box="[263,280,1411,1435]" pageId="99" pageNumber="100">E</emphasis>
, mesonotum, dorsal;
<emphasis id="B92D442C1E03FF97FE4554CEFDD8FA5B" bold="true" box="[499,514,1411,1435]" pageId="99" pageNumber="100">F</emphasis>
, wing, anterobasal part;
<emphasis id="B92D442C1E03FF97FD4B54CEFCCBFA5B" bold="true" box="[765,785,1411,1435]" pageId="99" pageNumber="100">G</emphasis>
, labels. Scale bar 2 mm.
</paragraph>
</caption>
<paragraph id="8BE6983E1E03FF97FF7154A0FD7EF957" blockId="99.[151,1437,1517,2012]" pageId="99" pageNumber="100">
<emphasis id="B92D442C1E03FF97FF7154A0FF25F9C7" bold="true" box="[199,255,1517,1543]" pageId="99" pageNumber="100">Legs</emphasis>
(
<figureCitation id="136284BB1E03FF97FEBB54A0FEACF9C7" box="[269,374,1517,1544]" captionStart="FIGURE 55" captionStartId="99.[151,250,1375,1399]" captionTargetBox="[151,1434,181,1345]" captionTargetId="figure-20@99.[151,1435,181,1351]" captionTargetPageId="99" captionText="FIGURE 55. Ulivellia pseudinsolita sp. nov. holotype ♂. A, habitus, left and dorsal; B, same, dorsal; C, head, anterior; D, same, left; E, mesonotum, dorsal; F, wing, anterobasal part; G, labels. Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14023310" httpUri="https://zenodo.org/record/14023310/files/figure.png" pageId="99" pageNumber="100">Fig. 55 A</figureCitation>
) with yellow or brownish yellow coxae and trochanters, femora and tibiae entirely black, except coxae and knees narrowly brown, with bluish sheen; tarsi entirely brownish yellow, black setose and setulose. Fore femur with 2 rows of posterodorsal and posterior setae slightly shorter than femur width, and 45 strong posteroventral setae in apical half. Mid femur anteriorly with row of short suberect setulae; mid tibia with one strong medioventral seta 2.1× longer than tibia width.
</paragraph>
<paragraph id="8BE6983E1E03FF97FF7157ECFC83F91F" blockId="99.[151,1437,1517,2012]" pageId="99" pageNumber="100">
<emphasis id="B92D442C1E03FF97FF7157ECFEE6F97B" bold="true" box="[199,316,1697,1723]" pageId="99" pageNumber="100">Abdomen</emphasis>
moderately elongate (
<figureCitation id="136284BB1E03FF97FD8057ECFD12F97C" box="[566,712,1697,1724]" captionStart="FIGURE 55" captionStartId="99.[151,250,1375,1399]" captionTargetBox="[151,1434,181,1345]" captionTargetId="figure-20@99.[151,1435,181,1351]" captionTargetPageId="99" captionText="FIGURE 55. Ulivellia pseudinsolita sp. nov. holotype ♂. A, habitus, left and dorsal; B, same, dorsal; C, head, anterior; D, same, left; E, mesonotum, dorsal; F, wing, anterobasal part; G, labels. Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14023310" httpUri="https://zenodo.org/record/14023310/files/figure.png" pageId="99" pageNumber="100">Figs 55 A, B</figureCitation>
,
<figureCitation id="136284BB1E03FF97FD6257ECFCD3F97B" box="[724,777,1697,1724]" captionStart="FIGURE 56" captionStartId="100.[151,250,1759,1783]" captionTargetBox="[158,1425,188,1728]" captionTargetId="figure-30@100.[151,1436,181,1735]" captionTargetPageId="100" captionText="FIGURE 56. Ulivellia pseudinsolita sp. nov. holotype ♂. A, postabdomen, posterior; B C, same, ventral; D, same, ventrolateral; E, surstyli and hypandrium, ventral; F, preabdomen, ventral. Abbreviations: cerc—cerci; ej apod—ejaculatory apodeme; epand—epandrium; ph g—phallic guide; ph—phallus; pregt—pregonite; p l sur—posterior lobe of surstylus; v l sur—ventral lobe of surstylus; st 2— second sternite; st 3—third sternite; st 4—fourth sternite; st 5—fifth sternite." figureDoi="http://doi.org/10.5281/zenodo.14023312" httpUri="https://zenodo.org/record/14023312/files/figure.png" pageId="99" pageNumber="100">56 F</figureCitation>
), black, sparsely whitish microtrichose, with cyan metallic sheen. Tergite 5 not microtrichose, somewhat shagreened, dull.
</paragraph>
<paragraph id="8BE6983E1E03FF97FF7157A4FD16F81B" blockId="99.[151,1437,1517,2012]" pageId="99" pageNumber="100">
Cerci wide oval, moderately developed, partly fused at bases elongate, sack-like; setulose (
<figureCitation id="136284BB1E03FF97FB4557A4FA4AF8C3" box="[1267,1424,1769,1796]" captionStart="FIGURE 56" captionStartId="100.[151,250,1759,1783]" captionTargetBox="[158,1425,188,1728]" captionTargetId="figure-30@100.[151,1436,181,1735]" captionTargetPageId="100" captionText="FIGURE 56. Ulivellia pseudinsolita sp. nov. holotype ♂. A, postabdomen, posterior; B C, same, ventral; D, same, ventrolateral; E, surstyli and hypandrium, ventral; F, preabdomen, ventral. Abbreviations: cerc—cerci; ej apod—ejaculatory apodeme; epand—epandrium; ph g—phallic guide; ph—phallus; pregt—pregonite; p l sur—posterior lobe of surstylus; v l sur—ventral lobe of surstylus; st 2— second sternite; st 3—third sternite; st 4—fourth sternite; st 5—fifth sternite." figureDoi="http://doi.org/10.5281/zenodo.14023312" httpUri="https://zenodo.org/record/14023312/files/figure.png" pageId="99" pageNumber="100">Figs 56 AD</figureCitation>
). Epandrium ovoid, dorsally short, moderately setulose; surstyli basally wide, outer surstylus flattened, posteroventrally directed, apically oval with single trichoid sensillum; inner surstylus with four very strong claw-like mesally directed prensisetae (
<figureCitation id="136284BB1E03FF97FE935618FE3EF8AF" box="[293,484,1877,1904]" captionStart="FIGURE 56" captionStartId="100.[151,250,1759,1783]" captionTargetBox="[158,1425,188,1728]" captionTargetId="figure-30@100.[151,1436,181,1735]" captionTargetPageId="100" captionText="FIGURE 56. Ulivellia pseudinsolita sp. nov. holotype ♂. A, postabdomen, posterior; B C, same, ventral; D, same, ventrolateral; E, surstyli and hypandrium, ventral; F, preabdomen, ventral. Abbreviations: cerc—cerci; ej apod—ejaculatory apodeme; epand—epandrium; ph g—phallic guide; ph—phallus; pregt—pregonite; p l sur—posterior lobe of surstylus; v l sur—ventral lobe of surstylus; st 2— second sternite; st 3—third sternite; st 4—fourth sternite; st 5—fifth sternite." figureDoi="http://doi.org/10.5281/zenodo.14023312" httpUri="https://zenodo.org/record/14023312/files/figure.png" pageId="99" pageNumber="100">Figs 56 A, C, D</figureCitation>
). Pregonites symmetrical, almost flat, with 5 moderately developed setulae ventro-mesally, conspicuously anterior to postgonites (
<figureCitation id="136284BB1E03FF97FD045634FC9DF853" box="[690,839,1913,1940]" captionStart="FIGURE 56" captionStartId="100.[151,250,1759,1783]" captionTargetBox="[158,1425,188,1728]" captionTargetId="figure-30@100.[151,1436,181,1735]" captionTargetPageId="100" captionText="FIGURE 56. Ulivellia pseudinsolita sp. nov. holotype ♂. A, postabdomen, posterior; B C, same, ventral; D, same, ventrolateral; E, surstyli and hypandrium, ventral; F, preabdomen, ventral. Abbreviations: cerc—cerci; ej apod—ejaculatory apodeme; epand—epandrium; ph g—phallic guide; ph—phallus; pregt—pregonite; p l sur—posterior lobe of surstylus; v l sur—ventral lobe of surstylus; st 2— second sternite; st 3—third sternite; st 4—fourth sternite; st 5—fifth sternite." figureDoi="http://doi.org/10.5281/zenodo.14023312" httpUri="https://zenodo.org/record/14023312/files/figure.png" pageId="99" pageNumber="100">Figs 56 C, E</figureCitation>
). Postgonites on laterobasal part of the phallic guide lobes, with 45 small trichoid sensilla (
<figureCitation id="136284BB1E03FF97FDEF56D0FD2AF877" box="[601,752,1949,1976]" captionStart="FIGURE 56" captionStartId="100.[151,250,1759,1783]" captionTargetBox="[158,1425,188,1728]" captionTargetId="figure-30@100.[151,1436,181,1735]" captionTargetPageId="100" captionText="FIGURE 56. Ulivellia pseudinsolita sp. nov. holotype ♂. A, postabdomen, posterior; B C, same, ventral; D, same, ventrolateral; E, surstyli and hypandrium, ventral; F, preabdomen, ventral. Abbreviations: cerc—cerci; ej apod—ejaculatory apodeme; epand—epandrium; ph g—phallic guide; ph—phallus; pregt—pregonite; p l sur—posterior lobe of surstylus; v l sur—ventral lobe of surstylus; st 2— second sternite; st 3—third sternite; st 4—fourth sternite; st 5—fifth sternite." figureDoi="http://doi.org/10.5281/zenodo.14023312" httpUri="https://zenodo.org/record/14023312/files/figure.png" pageId="99" pageNumber="100">Figs 56 C, E</figureCitation>
). Phallus moderately short and narrow, about 2.53× longer than epandrium high, thin and bare (
<figureCitation id="136284BB1E03FF97FD98568CFD1AF81B" box="[558,704,1985,2012]" captionStart="FIGURE 56" captionStartId="100.[151,250,1759,1783]" captionTargetBox="[158,1425,188,1728]" captionTargetId="figure-30@100.[151,1436,181,1735]" captionTargetPageId="100" captionText="FIGURE 56. Ulivellia pseudinsolita sp. nov. holotype ♂. A, postabdomen, posterior; B C, same, ventral; D, same, ventrolateral; E, surstyli and hypandrium, ventral; F, preabdomen, ventral. Abbreviations: cerc—cerci; ej apod—ejaculatory apodeme; epand—epandrium; ph g—phallic guide; ph—phallus; pregt—pregonite; p l sur—posterior lobe of surstylus; v l sur—ventral lobe of surstylus; st 2— second sternite; st 3—third sternite; st 4—fourth sternite; st 5—fifth sternite." figureDoi="http://doi.org/10.5281/zenodo.14023312" httpUri="https://zenodo.org/record/14023312/files/figure.png" pageId="99" pageNumber="100">Figs 56 AE</figureCitation>
).
</paragraph>
<caption id="DF26C8B61E04FF90FF215792FB40F8A3" ID-DOI="http://doi.org/10.5281/zenodo.14023312" ID-Zenodo-Dep="14023312" httpUri="https://zenodo.org/record/14023312/files/figure.png" pageId="100" pageNumber="101" startId="100.[151,250,1759,1783]" targetBox="[158,1425,188,1728]" targetPageId="100" targetType="figure">
<paragraph id="8BE6983E1E04FF90FF215792FB40F8A3" blockId="100.[151,1437,1759,1891]" pageId="100" pageNumber="101">
<emphasis id="B92D442C1E04FF90FF215792FEC5F937" bold="true" box="[151,287,1759,1783]" pageId="100" pageNumber="101">FIGURE 56.</emphasis>
<taxonomicName id="4C59E3BD1E04FF90FE955792FDD2F937" authorityName="Kovac &amp; Kameneva &amp; Korneyev &amp; Araújo &amp; Savaris &amp; Smit &amp; Schneider &amp; Schreiber &amp; Korneyev" authorityYear="2024" box="[291,520,1759,1783]" class="Insecta" family="Ulidiidae" genus="Ulivellia" kingdom="Animalia" order="Diptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="pseudinsolita" status="sp. nov.">
<emphasis id="B92D442C1E04FF90FE955792FDD2F937" box="[291,520,1759,1783]" italics="true" pageId="100" pageNumber="101">Ulivellia pseudinsolita</emphasis>
</taxonomicName>
<emphasis id="B92D442C1E04FF90FDBB5792FD86F937" bold="true" box="[525,604,1759,1783]" pageId="100" pageNumber="101">
<taxonomicNameLabel id="A21EF9571E04FF90FDBB5792FD86F937" box="[525,604,1759,1783]" pageId="100" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype ♂.
<emphasis id="B92D442C1E04FF90FD6B5792FD35F937" bold="true" box="[733,751,1759,1783]" pageId="100" pageNumber="101">A</emphasis>
, postabdomen, posterior;
<emphasis id="B92D442C1E04FF90FC455792FBC1F937" bold="true" box="[1011,1051,1759,1783]" pageId="100" pageNumber="101">B C</emphasis>
, same, ventral;
<emphasis id="B92D442C1E04FF90FB0E5792FB10F937" bold="true" box="[1208,1226,1759,1783]" pageId="100" pageNumber="101">D</emphasis>
, same, ventrolateral;
<emphasis id="B92D442C1E04FF90FF21564EFF72F8DB" bold="true" box="[151,168,1795,1819]" pageId="100" pageNumber="101">E</emphasis>
, surstyli and hypandrium, ventral;
<emphasis id="B92D442C1E04FF90FD93564EFDEEF8DB" bold="true" box="[549,564,1795,1819]" pageId="100" pageNumber="101">F</emphasis>
, preabdomen, ventral. Abbreviations: cerc—cerci; ej apod—ejaculatory apodeme; epand—epandrium; ph g—phallic guide; ph—phallus; pregt—pregonite; p l sur—posterior lobe of surstylus; v l sur—ventral lobe of surstylus; st 2— second sternite; st 3—third sternite; st 4—fourth sternite; st 5—fifth sternite.
</paragraph>
</caption>
<paragraph id="8BE6983E1E05FF91FF7151DAFE4FFF71" blockId="101.[151,1436,151,249]" box="[199,405,151,177]" pageId="101" pageNumber="102">
<emphasis id="B92D442C1E05FF91FF7151DAFEC4FF71" bold="true" box="[199,286,151,177]" pageId="101" pageNumber="102">Female</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="C343CBB51E05FF91FF7151F6FAEAFF39" pageId="101" pageNumber="102" type="etymology">
<paragraph id="8BE6983E1E05FF91FF7151F6FAEAFF39" blockId="101.[151,1436,151,249]" pageId="101" pageNumber="102">
<emphasis id="B92D442C1E05FF91FF7151F6FE94FF15" bold="true" box="[199,334,187,213]" pageId="101" pageNumber="102">Etymology.</emphasis>
The species name is an adjective and is composed of the Greek “pseud-” (“false”) and the species name
<taxonomicName id="4C59E3BD1E05FF91FF6D5192FEE4FF39" box="[219,318,223,249]" class="Insecta" family="Ulidiidae" genus="Euxesta" kingdom="Animalia" order="Diptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="insolita">“insolita</taxonomicName>
”, reflecting the superficial similarity of the wing pattern to
<taxonomicName id="4C59E3BD1E05FF91FC615192FAF6FF39" authority="Hendel, 1909" authorityName="Hendel" authorityYear="1909" box="[983,1324,223,249]" class="Insecta" family="Ulidiidae" genus="Euxesta" kingdom="Animalia" order="Diptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="insolita">
<emphasis id="B92D442C1E05FF91FC615192FB51FF39" box="[983,1163,223,249]" italics="true" pageId="101" pageNumber="102">Euxesta insolita</emphasis>
Hendel, 1909
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>