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<document id="5D1FDF568B8D3A81C5AAD93282B35893" ID-CLB-Dataset="20446" ID-DOI="10.11646/zootaxa.5104.2.5" ID-GBIF-Dataset="dc85c0df-79ee-4aab-b21d-2df3a61d8ac0" ID-ISSN="1175-5326" ID-Zenodo-Dep="6280839" ID-ZooBank="DE993A29-98B4-4DA0-85CF-9B8D4868A454" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645780355554" checkinUser="plazi" docAuthor="Sonar, Mohan A., Pawar, Ravi V. &amp; Wayal, Dyaneshwar V." docDate="2022" docId="03D287EDEC7BB133478FFF5775A6FBF8" docLanguage="en" docName="zootaxa.5104.2.5.pdf" docOrigin="Zootaxa 5104 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thalamoporella badvei Sonar &amp; Pawar &amp; Wayal 2022, n. sp." docType="treatment" docVersion="8" lastPageNumber="256" masterDocId="FFEBFF95EC7EB1364718FF8C7716FF80" masterDocTitle="Fossil Thalamoporellidae (Bryozoa) from Paleogene-Neogene sediments of western Kachchh, Gujarat, India" masterLastPageNumber="274" masterPageNumber="251" pageNumber="256" updateTime="1699316508393" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3C77B0723999531EDF4E2AE84F9E3F78">Fossil Thalamoporellidae (Bryozoa) from Paleogene-Neogene sediments of western Kachchh, Gujarat, India</mods:title>
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<mods:namePart id="53EFA4DED15ACC4614B271F8950D8271">Sonar, Mohan A.</mods:namePart>
<mods:affiliation id="E96A725C87260B210C6572410431A717">Department of Geology, Government Institute of Science, Aurangabad, Maharashtra, India.</mods:affiliation>
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<mods:namePart id="EF1A6E4708B55F37AE8B7E18760E81DF">Pawar, Ravi V.</mods:namePart>
<mods:affiliation id="E50B61D28DAD7A52747192773E14D2F3">Department of Geology, Government Institute of Science, Aurangabad, Maharashtra, India.</mods:affiliation>
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<mods:namePart id="D6DFD6FFE13DDC5888E64933CA793FF5">Wayal, Dyaneshwar V.</mods:namePart>
<mods:affiliation id="BAEE335E87C47CE1B19A2E57867FAFFF">Department of Geology, Government Institute of Science, Aurangabad, Maharashtra, India.</mods:affiliation>
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<mods:date id="D8AF31EE0FF5EA2EFC96289C9560EC33">2022</mods:date>
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<subSubSection id="C3616570EC7BB133478FFF5776DEFF74" box="[151,456,219,244]" pageId="5" pageNumber="256" type="nomenclature">
<paragraph id="8BC436FBEC7BB133478FFF5776DEFF74" blockId="5.[151,456,219,281]" box="[151,456,219,244]" pageId="5" pageNumber="256">
<heading id="D08C8197EC7BB133478FFF5776DEFF74" bold="true" box="[151,456,219,244]" fontSize="10" level="2" pageId="5" pageNumber="256" reason="2">
<emphasis id="B90FEAE9EC7BB133478FFF5776DEFF74" bold="true" box="[151,456,219,244]" pageId="5" pageNumber="256">
<taxonomicName id="4C7B4D78EC7BB133478FFF57769FFF73" authority="Sonar &amp; Pawar &amp; Wayal, 2022" authorityName="Sonar &amp; Pawar &amp; Wayal" authorityYear="2022" box="[151,393,219,243]" class="Gymnolaemata" family="Thalamoporellidae" genus="Thalamoporella" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="256" phylum="Bryozoa" rank="species" species="badvei" status="sp. nov.">Thalamoporella badvei</taxonomicName>
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<emphasis id="B90FEAE9EC7BB1334697FF5076B5FF74" box="[399,419,220,244]" inLineHeading="true" pageId="5" pageNumber="256" reason="1">n.</emphasis>
sp.
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</emphasis>
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</paragraph>
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<subSubSection id="C3616570EC7BB133478FFF72765AFE98" box="[151,332,254,281]" pageId="5" pageNumber="256" type="description">
<paragraph id="8BC436FBEC7BB133478FFF72765AFE98" blockId="5.[151,456,219,281]" box="[151,332,254,281]" pageId="5" pageNumber="256">
(
<figureCitation id="13402A7EEC7BB13347B8FF7277F2FE99" box="[160,228,254,281]" captionStart="FIGURE 5" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[151,1436,1244,1920]" captionTargetId="figure-77@4.[151,1436,1244,1920]" captionTargetPageId="4" captionText="FIGURE 5. Thalamoporella badvei n. sp. (Holotype GIS/B 0111). A. General view of colony showing autozooids. B. Detail of an anomalous or damaged zooid. C. Enlargement of autozooids, orifices and an avicularium. Scale bars: A, 500 µm; B, 100 µm; C, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.6280845" httpUri="https://zenodo.org/record/6280845/files/figure.png" pageId="5" pageNumber="256">Fig. 5</figureCitation>
,
<tableCitation id="C6F90340EC7BB13347F7FF727655FE98" box="[239,323,254,280]" captionStart="TABLE 2" captionStartId="5.[151,239,1191,1215]" captionTargetPageId="5" captionText="TABLE 2. Measurements (in mm) of Thalamoporella badvei n. sp." pageId="5" pageNumber="256">Table 2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3616570EC7BB133478FFEC9760FFE28" pageId="5" pageNumber="256" type="materials_examined">
<paragraph id="8BC436FBEC7BB133478FFEC9760FFE28" blockId="5.[151,1437,325,1144]" pageId="5" pageNumber="256">
<emphasis id="B90FEAE9EC7BB133478FFEC97693FEDF" bold="true" box="[151,389,325,351]" pageId="5" pageNumber="256">Material examined.</emphasis>
<materialsCitation id="3B133CA6EC7BB1334689FECA758EFEE0" ID-GBIF-Occurrence="3501212307" box="[401,664,326,352]" collectionCode="GIS/B 0111" pageId="5" pageNumber="256" specimenCount="1" typeStatus="holotype">
<typeStatus id="54C08859EC7BB1334689FECA76E8FEE0" box="[401,510,326,352]" pageId="5" pageNumber="256" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED6AAE3EEC7BB1334514FECA758EFEE0" box="[524,664,326,352]" pageId="5" pageNumber="256">GIS/B 0111</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B133CA6EC7BB13345B0FECA7603FE28" ID-GBIF-Occurrence="3501212313" collectingDate="2011-01-14" county="Cliff" elevation="40" latitude="23.395277" location="Kharinadi Formation" longLatPrecision="21" longitude="68.76917" municipality="Laiyari" pageId="5" pageNumber="256" specimenCount="1" stateProvince="Delhi" typeStatus="paratype">
<typeStatus id="54C08859EC7BB13345B0FECA740FFEE0" box="[680,793,326,352]" pageId="5" pageNumber="256" type="paratype">Paratypes</typeStatus>
: GIS/B 01120130.
<collectingCounty id="62A54E77EC7BB1334313FECA7329FEE0" box="[1035,1087,326,352]" pageId="5" pageNumber="256">Cliff</collectingCounty>
section of
<collectingMunicipality id="6BA0AC81EC7BB13343D1FECA720BFEE0" box="[1225,1309,326,352]" pageId="5" pageNumber="256">Laiyari</collectingMunicipality>
,
<location id="8EA46020EC7BB1334234FECA7619FE04" LSID="urn:lsid:plazi:treatment:03D287EDEC7BB133478FFF5775A6FBF8:8EA46020EC7BB1334234FECA7619FE04" county="Cliff" latitude="23.395277" longLatPrecision="21" longitude="68.76917" municipality="Laiyari" name="Kharinadi Formation" pageId="5" pageNumber="256" stateProvince="Delhi">Kharinadi Formation</location>
,
<location id="8EA46020EC7BB1334601FEE676D7FE04" LSID="urn:lsid:plazi:treatment:03D287EDEC7BB133478FFF5775A6FBF8:8EA46020EC7BB1334601FEE676D7FE04" box="[281,449,362,388]" county="Cliff" latitude="23.395277" longLatPrecision="21" longitude="68.76917" municipality="Laiyari" name="lower Miocene" pageId="5" pageNumber="256" stateProvince="Delhi">lower Miocene</location>
(Aquitanian),
<geoCoordinate id="EE4F503CEC7BB133457BFEE675E3FE04" box="[611,757,362,388]" degrees="23" direction="north" minutes="23" orientation="latitude" pageId="5" pageNumber="256" precision="15" seconds="43" value="23.395277">23°2343” N</geoCoordinate>
,
<geoCoordinate id="EE4F503CEC7BB13345E7FEE6749BFE04" box="[767,909,362,388]" degrees="68" direction="east" minutes="46" orientation="longitude" pageId="5" pageNumber="256" precision="15" seconds="09" value="68.76917">68°4609” E</geoCoordinate>
,
<elevation id="0056D1C8EC7BB1334480FEE67329FE04" box="[920,1087,362,388]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="256" unit="m" value="40.0">
elevation
<quantity id="4C839B1EEC7BB133431DFEE67329FE04" box="[1029,1087,362,388]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="256" unit="m" value="40.0">40 m</quantity>
</elevation>
,
<date id="FFC5103BEC7BB1334352FEE67215FE04" box="[1098,1283,362,388]" pageId="5" pageNumber="256" value="2011-01-14">
<collectingDate id="EF81E9D3EC7BB1334352FEE67215FE04" box="[1098,1283,362,388]" pageId="5" pageNumber="256" value="2011-01-14">14 January 2011</collectingDate>
</date>
, DST project,
<location id="8EA46020EC7BB133478FFE0277DAFE28" LSID="urn:lsid:plazi:treatment:03D287EDEC7BB133478FFF5775A6FBF8:8EA46020EC7BB133478FFE0277DAFE28" box="[151,204,398,424]" county="Cliff" latitude="23.395277" longLatPrecision="21" longitude="68.76917" municipality="Laiyari" name="New" pageId="5" pageNumber="256" stateProvince="Delhi">New</location>
<collectingRegion id="49BFF819EC7BB13347CBFE027603FE28" box="[211,277,398,424]" country="India" name="Delhi" pageId="5" pageNumber="256">Delhi</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3616570EC7BB13347DFFE3E762AFE70" pageId="5" pageNumber="256" type="etymology">
<paragraph id="8BC436FBEC7BB13347DFFE3E762AFE70" blockId="5.[151,1437,325,1144]" pageId="5" pageNumber="256">
<emphasis id="B90FEAE9EC7BB13347DFFE3E7658FE4C" bold="true" box="[199,334,434,460]" pageId="5" pageNumber="256">Etymology.</emphasis>
The species is named after the late Dr. R. M. Badve, in acknowledgement of his contribution to palaeontology.
</paragraph>
</subSubSection>
<subSubSection id="C3616570EC7BB13347DFFE7575F1FD00" pageId="5" pageNumber="256" type="diagnosis">
<paragraph id="8BC436FBEC7BB13347DFFE7575F1FD00" blockId="5.[151,1437,325,1144]" pageId="5" pageNumber="256">
<emphasis id="B90FEAE9EC7BB13347DFFE757654FD93" bold="true" box="[199,322,505,531]" pageId="5" pageNumber="256">Diagnosis.</emphasis>
Colony encrusting. Autozooids mostly elongate-rectangular, arranged in quincunx, each bordered by raised, thick autozooidal boundaries. Orifice almost pyriform shaped. Adoral tubercles wanting. Opesiules subtriangular, equal in size. Cryptocyst smooth, minutely perforated. Avicularia small, triangular, placed proximal to the autozooid with distally directed acute rostrum.
</paragraph>
</subSubSection>
<subSubSection id="C3616570EC7BB13347DFFD057577FC20" pageId="5" pageNumber="256" type="description">
<paragraph id="8BC436FBEC7BB13347DFFD057577FC20" blockId="5.[151,1437,325,1144]" pageId="5" pageNumber="256">
<emphasis id="B90FEAE9EC7BB13347DFFD05764FFD23" bold="true" box="[199,345,649,675]" pageId="5" pageNumber="256">Description.</emphasis>
Colony encrusting, unilaminar, arranged in longitudinally series of alternating rows. Autozooids long, with equal, parallel walls, surrounded by distinct, thick, raised autozooidal boundaries, fused with those of adjacent autozooids (
<figureCitation id="13402A7EEC7BB1334698FD5E76CFFD6D" box="[384,473,722,749]" captionStart="FIGURE 5" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[151,1436,1244,1920]" captionTargetId="figure-77@4.[151,1436,1244,1920]" captionTargetPageId="4" captionText="FIGURE 5. Thalamoporella badvei n. sp. (Holotype GIS/B 0111). A. General view of colony showing autozooids. B. Detail of an anomalous or damaged zooid. C. Enlargement of autozooids, orifices and an avicularium. Scale bars: A, 500 µm; B, 100 µm; C, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.6280845" httpUri="https://zenodo.org/record/6280845/files/figure.png" pageId="5" pageNumber="256">Fig. 5A</figureCitation>
). Some autozooids exhibit anomalous growth (
<figureCitation id="13402A7EEC7BB13344C4FD5E7325FD6D" box="[988,1075,722,749]" captionStart="FIGURE 5" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[151,1436,1244,1920]" captionTargetId="figure-77@4.[151,1436,1244,1920]" captionTargetPageId="4" captionText="FIGURE 5. Thalamoporella badvei n. sp. (Holotype GIS/B 0111). A. General view of colony showing autozooids. B. Detail of an anomalous or damaged zooid. C. Enlargement of autozooids, orifices and an avicularium. Scale bars: A, 500 µm; B, 100 µm; C, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.6280845" httpUri="https://zenodo.org/record/6280845/files/figure.png" pageId="5" pageNumber="256">Fig. 5B</figureCitation>
). Orifices generally subpyriform and longer than wide, with raised margin, the distal border more arcuate and the proximal border slightly concave. Adoral areas narrow, without tubercles. Two equal subtriangular opesiules adjacent to lateral walls of autozooids away from the proximal lip of the orifice. Cryptocyst an extensive shelf, shallow, smooth, with minute perforation. Avicularia rare, small, triangular, distal to an autozooid at bifurcation of rows; rostrum acute with blunt tip, directed distally (
<figureCitation id="13402A7EEC7BB13347E1FC0A7644FC21" box="[249,338,902,929]" captionStart="FIGURE 5" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[151,1436,1244,1920]" captionTargetId="figure-77@4.[151,1436,1244,1920]" captionTargetPageId="4" captionText="FIGURE 5. Thalamoporella badvei n. sp. (Holotype GIS/B 0111). A. General view of colony showing autozooids. B. Detail of an anomalous or damaged zooid. C. Enlargement of autozooids, orifices and an avicularium. Scale bars: A, 500 µm; B, 100 µm; C, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.6280845" httpUri="https://zenodo.org/record/6280845/files/figure.png" pageId="5" pageNumber="256">Fig. 5C</figureCitation>
). Ovicells not observed.
</paragraph>
</subSubSection>
<subSubSection id="C3616570EC7BB13347DFFC2675A6FBF8" pageId="5" pageNumber="256" type="discussion">
<paragraph id="8BC436FBEC7BB13347DFFC2675A6FBF8" blockId="5.[151,1437,325,1144]" pageId="5" pageNumber="256">
<emphasis id="B90FEAE9EC7BB13347DFFC26762DFC44" bold="true" box="[199,315,938,964]" pageId="5" pageNumber="256">Remarks.</emphasis>
<taxonomicName id="4C7B4D78EC7BB133465AFC267462FC44" authority="Guha &amp; Gopikrishna, 2004" authorityName="Guha &amp; Gopikrishna" authorityYear="2004" box="[322,884,938,964]" class="Gymnolaemata" family="Thalamoporellidae" genus="Thalamoporella" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="256" phylum="Bryozoa" rank="species" species="voigti">
<emphasis id="B90FEAE9EC7BB133465AFC26752AFC44" box="[322,572,938,964]" italics="true" pageId="5" pageNumber="256">Thalamoporella voigti</emphasis>
<bibRefCitation id="EFEA4B0AEC7BB133455BFC267462FC44" author="Guha, A. K. &amp; Gopikrishna, K." box="[579,884,938,964]" pageId="5" pageNumber="256" pagination="1 - 51" refId="ref8888" refString="Guha, A. K. &amp; Gopikrishna, K. (2004) Fossil Thalamoporella (Bryozoa) from the Tertiary sequences of Western Kachchh, Gujarat, India. Irene McCulloch Monograph Series, 7, 1 - 51. https: // doi. org / 10.1127 / njgpm / 2004 / 2004 / 681" type="journal article" year="2004">Guha &amp; Gopikrishna, 2004</bibRefCitation>
</taxonomicName>
superficially resembles the present species in the shape of its avicularia. However,
<taxonomicName id="4C7B4D78EC7BB133451EFC437576FC68" authorityName="Guha &amp; Gopikrishna" authorityYear="2004" box="[518,608,975,1000]" class="Gymnolaemata" family="Thalamoporellidae" genus="Thalamoporella" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="256" phylum="Bryozoa" rank="species" species="voigti">
<emphasis id="B90FEAE9EC7BB133451EFC437576FC68" box="[518,608,975,1000]" italics="true" pageId="5" pageNumber="256">T. voigti</emphasis>
</taxonomicName>
has raised prominent adoral tubercles, more-vasiform autozooids bordered by very thick, raised finely beaded autozooidal boundaries and a coarsely perforated cryptocyst.
<taxonomicName id="4C7B4D78EC7BB13343F1FC7E7544FBB0" authority="Soule, Soule &amp; Chaney, 1999" authorityName="Soule, Soule &amp; Chaney" authorityYear="1999" class="Gymnolaemata" family="Thalamoporellidae" genus="Thalamoporella" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="256" phylum="Bryozoa" rank="species" species="spinosa">
<emphasis id="B90FEAE9EC7BB13343F1FC7E77FBFBB0" italics="true" pageId="5" pageNumber="256">Thalamoporella spinosa</emphasis>
<bibRefCitation id="EFEA4B0AEC7BB13347E0FB9A7544FBB0" author="Soule, D. F. &amp; Soule, J. D. &amp; Chaney, H. W." box="[248,594,1046,1072]" pageId="5" pageNumber="256" pagination="1 - 57" refId="ref10350" refString="Soule, D. F., Soule, J. D. &amp; Chaney, H. W. (1999) New species of Thalamoporella (Bryozoa) with acute or subacute avicularium mandibles and review of known species worldwide. Irene McCulloch Foundation Monograph Series, 4, 1 - 57." type="journal article" year="1999">Soule, Soule &amp; Chaney, 1999</bibRefCitation>
</taxonomicName>
shows some resemblance in the shape of orifices and thick autozooidal borders but differs from the present species in having large adoral tubercles, unequal opesiules, short autozooids and avicularia that differ in position, shape and size.
</paragraph>
</subSubSection>
</treatment>
</document>