201 lines
26 KiB
XML
201 lines
26 KiB
XML
<document id="A4050A742AAAD86275754E7020418BD3" ID-ISSN="11755334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629589942712" checkinUser="felipe" docAuthor="Vaz-De-Mello, Fernando Z." docDate="2008" docId="03D287CEFFA5AF08D1DBDC23FE1A1001" docLanguage="en" docName="zt01955p075.pdf" docOrigin="Zootaxa 1955" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Bradypodidium Vaz-De-Mello 2008, new genus" docType="treatment" docVersion="3" lastPageNumber="19" masterDocId="FFEBFFB6FFB4AF1AD14CDF74FFA61358" masterDocTitle="Synopsis of the new subtribe Scatimina (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini), with descriptions of twelve new genera and review of Genieridium, new genus" masterLastPageNumber="75" masterPageNumber="1" pageNumber="18" updateTime="1699081972826" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="A56A523E01CEA1BE11D421870F0D05C6">
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<mods:title id="0FE431CA36944262A5C1A1CA7ED9ED52">Synopsis of the new subtribe Scatimina (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini), with descriptions of twelve new genera and review of Genieridium, new genus</mods:title>
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<mods:name id="06A246FEA379A57F4C2C7CE02FCD542C" type="personal">
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<mods:roleTerm id="3BFDE6FCFD51CE7F62C884315CB0351F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A5417A5CDF87ED68FE8BE41BDE8B84DA">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:typeOfResource id="D4DAD7B63FAA2CA942EF31A0475676C3">text</mods:typeOfResource>
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<mods:titleInfo id="FE819A6A39D8D163795710B8993C6215">
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<mods:title id="93B79F52F1CEC5811DAFA8BCFD6BB091">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="BB3332C6A7CEE4C0660DE9C846BEA6AE">
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<mods:date id="43DD250BA7FF79677D9143C0E01CC2C3">2008</mods:date>
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<mods:detail id="427810B16E18AC7D78D23EA3E1A6109A" type="pubDate">
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<mods:number id="D025AEABEC5FFEE7A99ADE97A877286D">2008-12-05</mods:number>
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<mods:detail id="3F914E4601277DB37AE5B9C5F8599127" type="volume">
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<mods:number id="48CEF1510303E0A50BCDC246B71A53F5">1955</mods:number>
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<mods:start id="639AADAF0500799EEA0C2F5EA779971A">1</mods:start>
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<mods:end id="FD472C7DF1C8FB96B28B1BBFFCAA57C1">75</mods:end>
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<mods:classification id="F1AF8EB6D2EB1729BB6F7F7C7B5ED6B0">journal article</mods:classification>
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<mods:identifier id="943D2FD58BE229A0B2C9A0A5C28A8865" type="ISSN">11755334</mods:identifier>
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</mods:mods>
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<treatment id="03D287CEFFA5AF08D1DBDC23FE1A1001" LSID="urn:lsid:plazi:treatment:03D287CEFFA5AF08D1DBDC23FE1A1001" httpUri="http://treatment.plazi.org/id/03D287CEFFA5AF08D1DBDC23FE1A1001" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
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<subSubSection id="C3616553FFA5AF0BD1DBDC23FE591029" box="[151,511,855,881]" pageId="17" pageNumber="18" type="nomenclature">
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<paragraph id="8BC436D8FFA5AF0BD1DBDC23FE591029" blockId="17.[151,511,855,881]" box="[151,511,855,881]" pageId="17" pageNumber="18">
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<heading id="D08C81B4FFA5AF0BD1DBDC23FE591029" bold="true" box="[151,511,855,881]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
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<emphasis id="B90FEACAFFA5AF0BD1DBDC23FE591029" bold="true" box="[151,511,855,881]" pageId="17" pageNumber="18">
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2.
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<taxonomicName id="4C7B4D5BFFA5AF0BD1F8DC23FEC91029" ID-CoL="3CQT" authority="Vaz-De-Mello, 2008" authorityName="Vaz-De-Mello" authorityYear="2008" box="[180,367,855,881]" class="Insecta" family="Scarabaeidae" genus="Bradypodidium" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90FEACAFFA5AF0BD1F8DC23FEC91029" bold="true" box="[180,367,855,881]" italics="true" pageId="17" pageNumber="18">Bradypodidium</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A23C57B1FFA5AF0BD032DC23FE591029" box="[382,511,855,881]" pageId="17" pageNumber="18" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FFA5AF0BD1DBDCD6FA91176C" pageId="17" pageNumber="18" type="diagnosis">
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<paragraph id="8BC436D8FFA5AF0BD1DBDCD6FA91176C" blockId="17.[151,1436,930,2036]" pageId="17" pageNumber="18">
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<emphasis id="B90FEACAFFA5AF0BD1DBDCD6FEBF10E4" bold="true" box="[151,281,930,956]" pageId="17" pageNumber="18">Diagnosis:</emphasis>
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Small oval to elongate species (
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<figureCitation id="13402A5DFFA5AF0BD3E4DCD6FCA410E4" box="[680,770,930,956]" captionStart="FIGURES 15–22" captionStartId="56.[151,269,1962,1986]" captionTargetBox="[207,1374,197,1940]" captionTargetId="figure-57@56.[207,1374,197,1940]" captionTargetPageId="56" captionText="FIGURES 15–22. Dorsal views. 15–16: Besourenga: 15. B. sp.; 16. B. horacioi; 17. Bradypodidium sp.; 18. Degallieridium lilliputanum; 19. Eutrichillum sp.; 20. Feeridium woodruffi; 21–22: Genieridium: 21. G. bordoni; 22. G. cryptops." pageId="17" pageNumber="18">Fig. 17</figureCitation>
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), hairs very fine, clypeus with none, two or four teeth (
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<figureCitation id="13402A5DFFA5AF0BD1D3DCBEFE8810BC" box="[159,302,970,996]" captionStart="FIGURES 39–46" captionStartId="59.[151,269,1969,1993]" captionTargetBox="[211,1382,194,1944]" captionTargetId="figure-50@59.[211,1382,194,1948]" captionTargetPageId="59" captionText="FIGURES 39–46. 39–40: Besourenga minutus: 39. head; 40. parameres; 41. B. amarillai, head; 42. B. horacioi, ♂ protarsus; 43–44: Bradypodidium bradyporum: 43. head; 44. parameres; 45–46: B. adisi; 45. head; 46. ♂ protarsus." pageId="17" pageNumber="18">Figs. 43, 45</figureCitation>
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), elytral striae deeper and wider posteriorly, protibial teeth concentrated in the apical one-half of tibia; males with proclaws and last tarsomere strongly modified (
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<figureCitation id="13402A5DFFA5AF0BD287DC86FB8F1754" box="[971,1065,1010,1036]" captionStart="FIGURES 39–46" captionStartId="59.[151,269,1969,1993]" captionTargetBox="[211,1382,194,1944]" captionTargetId="figure-50@59.[211,1382,194,1948]" captionTargetPageId="59" captionText="FIGURES 39–46. 39–40: Besourenga minutus: 39. head; 40. parameres; 41. B. amarillai, head; 42. B. horacioi, ♂ protarsus; 43–44: Bradypodidium bradyporum: 43. head; 44. parameres; 45–46: B. adisi; 45. head; 46. ♂ protarsus." pageId="17" pageNumber="18">Fig. 46</figureCitation>
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); lacking apicomesal tooth in protibia; parameres elongated, flat and wider at apex, always with mesal apices superposed (
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<figureCitation id="13402A5DFFA5AF0BD59EDB6EFA8D176C" box="[1234,1323,1050,1076]" captionStart="FIGURES 39–46" captionStartId="59.[151,269,1969,1993]" captionTargetBox="[211,1382,194,1944]" captionTargetId="figure-50@59.[211,1382,194,1948]" captionTargetPageId="59" captionText="FIGURES 39–46. 39–40: Besourenga minutus: 39. head; 40. parameres; 41. B. amarillai, head; 42. B. horacioi, ♂ protarsus; 43–44: Bradypodidium bradyporum: 43. head; 44. parameres; 45–46: B. adisi; 45. head; 46. ♂ protarsus." pageId="17" pageNumber="18">Fig. 44</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FFA5AF0BD18ADB36FAC214AC" pageId="17" pageNumber="18" type="description">
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<paragraph id="8BC436D8FFA5AF0BD18ADB36FB84145C" blockId="17.[151,1436,930,2036]" pageId="17" pageNumber="18">
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<emphasis id="B90FEACAFFA5AF0BD18ADB36FEC61704" bold="true" box="[198,352,1090,1116]" pageId="17" pageNumber="18">Description:</emphasis>
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Size
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<quantity id="4C839B3DFFA5AF0BD0EEDB36FD951704" box="[418,563,1090,1116]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.7" metricValueMin="2.3" pageId="17" pageNumber="18" unit="mm" value="3.0" valueMax="3.7" valueMin="2.3">2.3–3.7 mm</quantity>
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. Body oval, with elytra clearly wider than prothorax (
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<figureCitation id="13402A5DFFA5AF0BD5E5DB36FAA61704" box="[1193,1280,1090,1116]" captionStart="FIGURES 15–22" captionStartId="56.[151,269,1962,1986]" captionTargetBox="[207,1374,197,1940]" captionTargetId="figure-57@56.[207,1374,197,1940]" captionTargetPageId="56" captionText="FIGURES 15–22. Dorsal views. 15–16: Besourenga: 15. B. sp.; 16. B. horacioi; 17. Bradypodidium sp.; 18. Degallieridium lilliputanum; 19. Eutrichillum sp.; 20. Feeridium woodruffi; 21–22: Genieridium: 21. G. bordoni; 22. G. cryptops." pageId="17" pageNumber="18">Fig. 17</figureCitation>
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); color tan to black, rarely with traces of metallic sheen. Clypeus with none, two, or four teeth (interspecific and sexually dimorphic in at least some species related to a wide variety of general head forms: transverse, rounded, or triangularly elongated). Clypeal border lateral to teeth straight to feebly rounded, continuous with genal border. Clypeofrontal and clypeogenal sutures indistinct, head dorsally flat except for anteromedian part of clypeus. Eyes, in dorsal view oval; interocular width 8 to 10 times eye width (
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<figureCitation id="13402A5DFFA5AF0BD28FDA7EFBF5167C" box="[963,1107,1290,1316]" captionStart="FIGURES 39–46" captionStartId="59.[151,269,1969,1993]" captionTargetBox="[211,1382,194,1944]" captionTargetId="figure-50@59.[211,1382,194,1948]" captionTargetPageId="59" captionText="FIGURES 39–46. 39–40: Besourenga minutus: 39. head; 40. parameres; 41. B. amarillai, head; 42. B. horacioi, ♂ protarsus; 43–44: Bradypodidium bradyporum: 43. head; 44. parameres; 45–46: B. adisi; 45. head; 46. ♂ protarsus." pageId="17" pageNumber="18">Figs. 43, 45</figureCitation>
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). Pronotum lacking anterior or posterior beads, lateral callosity weakly defined. Disc centrally with setose punctures. Hypomera laterally separated from pronotal disc by strong carina and bearing a strong posterior longitudinal carina parallel to the pronotal-hypomeral delimitation. Mesoepimera with anterior transverse carina. Metasternum with setose punctures on the disc. Elytral interstriae almost flat, each with two well-separated rows of setose punctures; striae wider and deeper on the posterior declivity. Pseudoepipleura gradually narrowed from anterior third to end of elytron, with setiferous punctures all along. Protibiae with three teeth, basal one frequently reduced; teeth concentrated in apical two-fifths of tibia; lateral tibial margin denticulate between base and basal tooth; ventral scale-like setae of tibial teeth absent. Mesotibiae and metatibiae elongate, much longer than five times their apical width. Metatarsus with first tarsomere flattened and oval, longer than second. Parameres bladelike, tips slightly overlapped, apically wider than at mid-length (
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<figureCitation id="13402A5DFFA5AF0BD2E8D9EEFC5915EC" box="[932,1023,1690,1716]" captionStart="FIGURES 39–46" captionStartId="59.[151,269,1969,1993]" captionTargetBox="[211,1382,194,1944]" captionTargetId="figure-50@59.[211,1382,194,1948]" captionTargetPageId="59" captionText="FIGURES 39–46. 39–40: Besourenga minutus: 39. head; 40. parameres; 41. B. amarillai, head; 42. B. horacioi, ♂ protarsus; 43–44: Bradypodidium bradyporum: 43. head; 44. parameres; 45–46: B. adisi; 45. head; 46. ♂ protarsus." pageId="17" pageNumber="18">Fig. 44</figureCitation>
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), about as long as half phallobase length or shorter. Coxites present, large and symmetrical. Spermatheca C-shaped, gradually narrowed posteriorly, basally globular and strongly modified in conical point at duct insertion.
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</paragraph>
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<paragraph id="8BC436D8FFA5AF0BD18AD866FAC214AC" blockId="17.[151,1436,930,2036]" pageId="17" pageNumber="18">
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<emphasis id="B90FEACAFFA5AF0BD18AD866FE621474" bold="true" box="[198,452,1810,1836]" pageId="17" pageNumber="18">Sexual dimorphism:</emphasis>
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Apical tarsomere of protarsi of male excavated dorsally to receive proclaws; claws strongly angulate at mid-length (
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<figureCitation id="13402A5DFFA5AF0BD365D84EFD20140C" box="[553,646,1850,1876]" captionStart="FIGURES 39–46" captionStartId="59.[151,269,1969,1993]" captionTargetBox="[211,1382,194,1944]" captionTargetId="figure-50@59.[211,1382,194,1948]" captionTargetPageId="59" captionText="FIGURES 39–46. 39–40: Besourenga minutus: 39. head; 40. parameres; 41. B. amarillai, head; 42. B. horacioi, ♂ protarsus; 43–44: Bradypodidium bradyporum: 43. head; 44. parameres; 45–46: B. adisi; 45. head; 46. ♂ protarsus." pageId="17" pageNumber="18">Fig. 46</figureCitation>
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). Metasternal disc somewhat concave in males, and very flat in females. Protibial teeth much wider and robust in females; males frequently with very reduced basal tooth. In species with variable clypeal shape, males often with more elongated clypeus when teeth are present, and reduced or absent lateral clypeal teeth; while females almost always have transverse or rounded head with four well developed clypeal teeth. Pygidium much more transverse and vertical in females than in males.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FFA6AF08D18ADFE3FD1A1382" pageId="18" pageNumber="19" type="type_taxon">
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<paragraph id="8BC436D8FFA6AF08D18ADFE3FD1A1382" blockId="18.[151,1437,151,338]" pageId="18" pageNumber="19">
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<emphasis id="B90FEACAFFA6AF08D18ADFE3FECB13E9" bold="true" box="[198,365,151,177]" pageId="18" pageNumber="19">
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<typeStatus id="54C0887AFFA6AF08D18ADFE3FEA213E9" box="[198,260,151,177]" pageId="18" pageNumber="19">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C7B4D5BFFA6AF08D03ADFECFCCE13EA" ID-CoL="58936" authority="Boucomont, 1928" authorityName=": Boucomont" authorityYear="1928" box="[374,872,152,178]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bradyporum">
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<emphasis id="B90FEACAFFA6AF08D03ADFECFD2D13E9" box="[374,651,152,177]" italics="true" pageId="18" pageNumber="19">Trichillum bradyporum</emphasis>
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<bibRefCitation id="EFEA4B29FFA6AF08D3D8DFECFCCE13EA" author="Boucomont, A." box="[660,872,152,178]" pageId="18" pageNumber="19" pagination="202 - 207" refId="ref47542" refString="Boucomont, A. (1928) Coprophages d'Amerique du Sud nouveaux ou peu connus. Bulletin de la Societe Entomologique de France, 1928, 186 - 194, 202 - 207." type="journal article" year="1928">Boucomont, 1928</bibRefCitation>
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</taxonomicName>
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(present designation) =
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<taxonomicName id="4C7B4D5BFFA6AF08D5C2DFECFE751382" ID-CoL="5X3QX" authority="(Boucomont, 1928)" authorityName="Vaz-De-Mello" authorityYear="2008" baseAuthorityName="Boucomont" baseAuthorityYear="1928" class="Insecta" family="Scarabaeidae" genus="Bradypodidium" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bradyporum" status="comb. nov.">
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<emphasis id="B90FEACAFFA6AF08D5C2DFECFF451381" italics="true" pageId="18" pageNumber="19">Bradypodidium bradyporum</emphasis>
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(
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<bibRefCitation id="EFEA4B29FFA6AF08D1BEDFB4FE6C1382" author="Boucomont, A." box="[242,458,192,218]" pageId="18" pageNumber="19" pagination="202 - 207" refId="ref47542" refString="Boucomont, A. (1928) Coprophages d'Amerique du Sud nouveaux ou peu connus. Bulletin de la Societe Entomologique de France, 1928, 186 - 194, 202 - 207." type="journal article" year="1928">Boucomont, 1928</bibRefCitation>
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||
)
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||
</taxonomicName>
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,
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<emphasis id="B90FEACAFFA6AF08D093DFCBFD131381" bold="true" box="[479,693,191,217]" pageId="18" pageNumber="19">
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||
<taxonomicNameLabel id="A23C57B1FFA6AF08D093DFCBFD131381" box="[479,693,191,217]" pageId="18" pageNumber="19" rank="species">new combination</taxonomicNameLabel>
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</emphasis>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FFA6AF08D18ADF93FD24120A" pageId="18" pageNumber="19" type="etymology">
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<paragraph id="8BC436D8FFA6AF08D18ADF93FD24120A" blockId="18.[151,1437,151,338]" pageId="18" pageNumber="19">
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<emphasis id="B90FEACAFFA6AF08D18ADF93FEF11259" bold="true" box="[198,343,231,257]" pageId="18" pageNumber="19">Etymology:</emphasis>
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Combination of
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<taxonomicName id="4C7B4D5BFFA6AF08D36EDF9CFD341259" authorityName="Linnaeus" authorityYear="1758" box="[546,658,232,257]" class="Mammalia" family="Bradypodidae" genus="Bradypus" kingdom="Animalia" order="Pilosa" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
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<emphasis id="B90FEACAFFA6AF08D36EDF9CFD341259" box="[546,658,232,257]" italics="true" pageId="18" pageNumber="19">Bradypus</emphasis>
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</taxonomicName>
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(generic name of the most common sloths [
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<taxonomicName id="4C7B4D5BFFA6AF08D5D7DF9CFA87125A" authorityName="Linnaeus" authorityYear="1758" box="[1179,1313,232,258]" class="Mammalia" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="class">Mammalia</taxonomicName>
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: Edentata:
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<taxonomicName id="4C7B4D5BFFA6AF08D1DBDE64FEE41272" authorityName="Gray" authorityYear="1821" box="[151,322,272,298]" class="Mammalia" family="Bradypodidae" kingdom="Animalia" order="Pilosa" pageId="18" pageNumber="19" phylum="Chordata" rank="family">Bradypodidae</taxonomicName>
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], in reference to the phoretic relation these scarabs have with them) and
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<taxonomicName id="4C7B4D5BFFA6AF08D5EBDE64FA971271" authorityName="Harold" authorityYear="1868" box="[1191,1329,272,297]" class="Insecta" family="Scarabaeidae" genus="Pedaridium" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEACAFFA6AF08D5EBDE64FA971271" box="[1191,1329,272,297]" italics="true" pageId="18" pageNumber="19">Pedaridium</emphasis>
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</taxonomicName>
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, a genus name.
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<taxonomicName id="4C7B4D5BFFA6AF08D1A9DE4CFE3D1209" authorityName="Vaz-De-Mello" authorityYear="2008" box="[229,411,312,337]" class="Insecta" family="Scarabaeidae" genus="Bradypodidium" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEACAFFA6AF08D1A9DE4CFE3D1209" box="[229,411,312,337]" italics="true" pageId="18" pageNumber="19">Bradypodidium</emphasis>
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</taxonomicName>
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is neuter in gender.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FFA6AF08D18ADEF3FF401292" pageId="18" pageNumber="19" type="distribution">
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<paragraph id="8BC436D8FFA6AF08D18ADEF3FF401292" blockId="18.[151,1436,391,857]" pageId="18" pageNumber="19">
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||
<emphasis id="B90FEACAFFA6AF08D18ADEF3FECF12F9" bold="true" box="[198,361,391,417]" pageId="18" pageNumber="19">Distribution:</emphasis>
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Widely co-distributed with sloths in the Neotropical region from
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<collectingCountry id="F36C7648FFA6AF08D52ADEFCFB4312FA" box="[1126,1253,392,418]" name="Costa Rica" pageId="18" pageNumber="19">Costa Rica</collectingCountry>
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to southeastern
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<collectingCountry id="F36C7648FFA6AF08D1DBDEC4FF441292" box="[151,226,432,458]" name="Brazil" pageId="18" pageNumber="19">Brazil</collectingCountry>
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.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3616553FFA6AF08D18ADEA3FE3F106A" pageId="18" pageNumber="19" type="discussion">
|
||
<paragraph id="8BC436D8FFA6AF08D18ADEA3FE2C11E2" blockId="18.[151,1436,391,857]" pageId="18" pageNumber="19">
|
||
<emphasis id="B90FEACAFFA6AF08D18ADEA3FEE712A9" bold="true" box="[198,321,471,497]" pageId="18" pageNumber="19">Remarks:</emphasis>
|
||
Collection records suggest an obligatory relationship with sloths.
|
||
<taxonomicName id="4C7B4D5BFFA6AF08D525DEACFAB912A9" authorityName="Vaz-De-Mello" authorityYear="2008" box="[1129,1311,472,497]" class="Insecta" family="Scarabaeidae" genus="Bradypodidium" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEACAFFA6AF08D525DEACFAB912A9" box="[1129,1311,472,497]" italics="true" pageId="18" pageNumber="19">Bradypodidium</emphasis>
|
||
</taxonomicName>
|
||
is closely related to
|
||
<taxonomicName id="4C7B4D5BFFA6AF08D043DD74FE0E1141" authorityName="Vaz-De-Mello" authorityYear="2008" box="[271,424,512,537]" class="Insecta" family="Scarabaeidae" genus="Trichillidium" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEACAFFA6AF08D043DD74FE0E1141" box="[271,424,512,537]" italics="true" pageId="18" pageNumber="19">Trichillidium</emphasis>
|
||
</taxonomicName>
|
||
, sharing with the latter protibial teeth concentrated in the apical half of tibia, slender dorsal setae, among many other characters. Monophyly of
|
||
<taxonomicName id="4C7B4D5BFFA6AF08D205DD5CFC591119" authorityName="Vaz-De-Mello" authorityYear="2008" box="[841,1023,552,577]" class="Insecta" family="Scarabaeidae" genus="Bradypodidium" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEACAFFA6AF08D205DD5CFC591119" box="[841,1023,552,577]" italics="true" pageId="18" pageNumber="19">Bradypodidium</emphasis>
|
||
</taxonomicName>
|
||
is supported by the relatively slen- der male tibia in relation to female, laminate overlapped parameres, relatively elongated body, transverse pygidium in females, and the association with sloths. See under
|
||
<taxonomicName id="4C7B4D5BFFA6AF08D2D8DD0CFB8611C9" authorityName="Vaz-De-Mello" authorityYear="2008" box="[916,1056,632,657]" class="Insecta" family="Scarabaeidae" genus="Onoreidium" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEACAFFA6AF08D2D8DD0CFB8611C9" box="[916,1056,632,657]" italics="true" pageId="18" pageNumber="19">Onoreidium</emphasis>
|
||
</taxonomicName>
|
||
for more remarks on
|
||
<emphasis id="B90FEACAFFA6AF08D469DD0CFE2511E1" italics="true" pageId="18" pageNumber="19">Trichillidium-Bradypodidium</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC436D8FFA6AF08D18ADDB3FE3F106A" blockId="18.[151,1436,391,857]" pageId="18" pageNumber="19">
|
||
<emphasis id="B90FEACAFFA6AF08D18ADDB3FEC911B9" bold="true" box="[198,367,711,737]" pageId="18" pageNumber="19">Composition:</emphasis>
|
||
Besides the
|
||
<typeStatus id="54C0887AFFA6AF08D34FDDBCFD9311BA" box="[515,565,712,738]" pageId="18" pageNumber="19">type</typeStatus>
|
||
species, the genus
|
||
<taxonomicName id="4C7B4D5BFFA6AF08D258DDBCFC6C11B9" authorityName="Vaz-De-Mello" authorityYear="2008" box="[788,970,712,737]" class="Insecta" family="Scarabaeidae" genus="Bradypodidium" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEACAFFA6AF08D258DDBCFC6C11B9" box="[788,970,712,737]" italics="true" pageId="18" pageNumber="19">Bradypodidium</emphasis>
|
||
</taxonomicName>
|
||
includes
|
||
<taxonomicName id="4C7B4D5BFFA6AF08D576DDBCFAF811BA" authority="(Ratcliffe, 1980)" baseAuthorityName="Ratcliffe" baseAuthorityYear="1980" box="[1082,1374,712,738]" class="Insecta" family="Scarabaeidae" genus="Bradypodidium" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="adisi" status="comb. nov.">
|
||
<emphasis id="B90FEACAFFA6AF08D576DDBCFB3511B9" box="[1082,1171,712,737]" italics="true" pageId="18" pageNumber="19">B. adisi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEA4B29FFA6AF08D5EDDDBCFAF011BA" author="Ratcliffe, B. C." box="[1185,1366,712,738]" pageId="18" pageNumber="19" pagination="337 - 350" refId="ref50629" refString="Ratcliffe, B. C. (1980) New species of Coprini (Coleoptera: Scarabaeidae: Scarabaeinae) taken from the pellage of three toed sloths (Bradypus tridactylus L.) (Edentata: Bradypodidae) in Central Amazonia with a brief commentary on scarab-sloth relationships. The Coleopterists Bulletin, 34 (4), 337 - 350." type="journal article" year="1980">Ratcliffe, 1980</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B90FEACAFFA6AF08D425DDB3FE951051" bold="true" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="A23C57B1FFA6AF08D425DDB3FE951051" pageId="18" pageNumber="19" rank="species">new combination</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C7B4D5BFFA6AF08D00FDD84FCE21052" authority="(Ferreira & Galileo, 1993)" authorityName="Vaz-De-Mello" authorityYear="2008" baseAuthorityName="Ferreira & Galileo" baseAuthorityYear="2008" box="[323,836,752,778]" class="Insecta" family="Scarabaeidae" genus="Bradypodidium" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="venezuelense" status="comb. nov.">
|
||
<emphasis id="B90FEACAFFA6AF08D00FDD84FE591051" box="[323,511,752,777]" italics="true" pageId="18" pageNumber="19">B. venezuelense</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEA4B29FFA6AF08D343DD84FC9D1052" author="Ferreira, A. M. R. M. & Galileo, M. H. M." box="[527,827,752,778]" pageId="18" pageNumber="19" pagination="3 - 70" refId="ref48083" refString="Ferreira, A. M. R. M. & Galileo, M. H. M. (1993) Revisao taxonomica do genero Pedaridium Harold, 1868 (Coleoptera, Scarabaeidae, Scarabaeinae, Coprini). Iheringia, Serie Zoologia, 74, 3 - 70." type="journal article" year="1993">Ferreira & Galileo, 1993</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B90FEACAFFA6AF08D21EDD9BFB8F1051" bold="true" box="[850,1065,751,777]" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="A23C57B1FFA6AF08D21EDD9BFB8F1051" box="[850,1065,751,777]" pageId="18" pageNumber="19" rank="species">new combination</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and at least three new species presently under study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3616553FFA6AF08D18ADC4BFE1A1001" box="[198,444,831,857]" pageId="18" pageNumber="19" type="materials_examined">
|
||
<paragraph id="8BC436D8FFA6AF08D18ADC4BFE1A1001" blockId="18.[151,1436,391,857]" box="[198,444,831,857]" pageId="18" pageNumber="19">
|
||
<emphasis id="B90FEACAFFA6AF08D18ADC4BFE1A1001" bold="true" box="[198,444,831,857]" pageId="18" pageNumber="19">Material examined:</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |