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<document id="41DE6C358ADDFCD8671FC1B556506077" ID-CLB-Dataset="132136" ID-DOI="10.11646/phytotaxa.559.2.10" ID-GBIF-Dataset="a03e7f76-4232-4462-9519-5334c16d1291" ID-ISSN="1179-3163" ID-Zenodo-Dep="7021471" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1661409513770" checkinUser="plazi" docAuthor="He, Hai &amp; Zhang, Li-Bing" docDate="2022" docId="03D287C06A2E3F542BD94A27FEE5F867" docLanguage="en" docName="phytotaxa.559.2.10.pdf" docOrigin="Phytotaxa 559 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.559.2.10" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Polystichum gonggashanense H. S. Kung &amp; Li Bing Zhang ex H. He &amp; Li Bing Zhang 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="210" masterDocId="FFEBFFB86A2E3F572B514D16FFBCFFFB" masterDocTitle="Polystichum gonggashanense (Dryopteridaceae): a new fern from Sichuan, China" masterLastPageNumber="212" masterPageNumber="207" pageNumber="207" updateTime="1699372086959" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E7FA656E86A99008736626E42F3A677F">Polystichum gonggashanense (Dryopteridaceae): a new fern from Sichuan, China</mods:title>
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<mods:namePart id="305D27993D437CFF137B790CF1484DE2">He, Hai</mods:namePart>
<mods:nameIdentifier id="48E6997ED216B5F59843439102560D29" type="ORCID">0000-0001-5362-9177</mods:nameIdentifier>
<mods:affiliation id="45AF44B6DABC82175FFCEFBA265985A8">College of Life Sciences, Chongqing Normal University, Shapingba, Chongqing 401331, China &amp; hehai @ cqnu. edu. cn; https: // orcid. org / 0000 - 0001 - 5362 - 9177</mods:affiliation>
<mods:nameIdentifier id="D856EE0F8161626DBE61CC925FE3EA4C" type="email">hehai@cqnu.edu.cn</mods:nameIdentifier>
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<mods:namePart id="27E8270E4BDDF0DF7D46962F89B4A1CE">Zhang, Li-Bing</mods:namePart>
<mods:nameIdentifier id="33A8A0D46CAB7107EBC83661B51BEBF1" type="ORCID">0000-0002-4905-040X</mods:nameIdentifier>
<mods:affiliation id="26BCA4DA6C0FF73F7DADBA3FE4878350">Missouri Botanical Garden, 4344 Shaw Blvd., St. Louis, Missouri 63110, U. S. A. and Chengdu Institute of Biology, Chinese Academy &amp; Libing. Zhang @ mobot. org; https: // orcid. org / 0000 - 0002 - 4905 - 040 X</mods:affiliation>
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<mods:title id="040D951AB22B73D7AD3F9332D1E7DB4B">Phytotaxa</mods:title>
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<mods:date id="1DF357A4FB5C0081027831103E1832E3">2022</mods:date>
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<mods:number id="716C2B704AD0A9177998E906B71F56C3">2022-08-23</mods:number>
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<mods:number id="EF475594FBD2F36372478C20C053A60A">559</mods:number>
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<treatment id="03D287C06A2E3F542BD94A27FEE5F867" ID-DOI="http://doi.org/10.5281/zenodo.7037110" ID-GBIF-Taxon="199861358" ID-Zenodo-Dep="7037110" LSID="urn:lsid:plazi:treatment:03D287C06A2E3F542BD94A27FEE5F867" httpUri="http://treatment.plazi.org/id/03D287C06A2E3F542BD94A27FEE5F867" lastPageId="3" lastPageNumber="210" pageId="0" pageNumber="207">
<subSubSection id="C361655D6A2E3F572BD94A27FB1DF8BD" box="[136,1185,1841,1863]" pageId="0" pageNumber="207" type="nomenclature">
<paragraph id="8BC436D66A2E3F572BD94A27FB1DF8BD" blockId="0.[136,1452,1841,2079]" box="[136,1185,1841,1863]" pageId="0" pageNumber="207">
<taxonomicName id="4C7B4D556A2E3F572BD94A27FC79F8BC" ID-CoL="BDD7B" authority="H.S.Kung &amp; Li Bing Zhang ex H.He &amp; Li Bing Zhang" authorityName="H. S. Kung &amp; Li Bing Zhang ex H. He &amp; Li Bing Zhang" authorityYear="2022" box="[136,965,1841,1863]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="0" pageNumber="207" phylum="Tracheophyta" rank="species" species="gonggashanense" status="sp. nov.">
<emphasis id="B90FEAC46A2E3F572BD94A27FE12F8BC" bold="true" box="[136,430,1841,1863]" italics="true" pageId="0" pageNumber="207">Polystichum gonggashanense</emphasis>
H.S.Kung &amp; Li Bing Zhang ex H.He &amp; Li Bing Zhang
</taxonomicName>
,
<taxonomicNameLabel id="A23C57BF6A2E3F57289F4A24FBA4F8BD" box="[974,1048,1841,1863]" pageId="0" pageNumber="207" rank="species">
<emphasis id="B90FEAC46A2E3F57289F4A24FBAEF8BC" box="[974,1042,1842,1863]" italics="true" pageId="0" pageNumber="207">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="13402A536A2E3F572F744A27FBDBF8BD" box="[1061,1127,1841,1863]" captionStart="FIGURE 1" captionStartId="1.[136,229,2027,2049]" captionTargetBox="[188,1393,210,1971]" captionTargetId="figure-19@1.[151,1436,187,2000]" captionTargetPageId="1" captionText="FIGURE 1. An illustration of Polystichum gonggashanense.—A. Habit.—B. Pinna.—C. Petiole scales.—D. Rachis scales.—E. Microscales.—F. Indusium (based on Li-Bing Zhang 169; drawn by Jian Chen)." figureDoi="http://doi.org/10.5281/zenodo.7021473" httpUri="https://zenodo.org/record/7021473/files/figure.png" pageId="0" pageNumber="207">Figs. 1</figureCitation>
&amp;
<figureCitation id="13402A536A2E3F572FD74A27FB28F8BD" box="[1158,1172,1841,1862]" captionStart="FIGURE 2" captionStartId="2.[136,229,1946,1968]" captionTargetBox="[159,1428,205,1904]" captionTargetId="figure-29@2.[151,1436,190,1922]" captionTargetPageId="2" captionText="FIGURE 2. A color plate of Polystichum gonggashanense.—A. The highest peak of Mt.Gongga where the new species was discovered.— B. Habit.—C. Lower portion of petiole showing blackish brown scales.—D, E. Portions of adaxial pinnae showing scales.—F. Portion of abaxial pinna showing sori and scales." figureDoi="http://doi.org/10.5281/zenodo.7021475" httpUri="https://zenodo.org/record/7021475/files/figure.png" pageId="0" pageNumber="207">2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C361655D6A2E3F572BD94A43FC0BF875" pageId="0" pageNumber="207" type="materials_examined">
<paragraph id="8BC436D66A2E3F572BD94A43FC0BF875" blockId="0.[136,1452,1841,2079]" pageId="0" pageNumber="207">
<emphasis id="B90FEAC46A2E3F572BD94A43FF06F891" bold="true" box="[136,186,1877,1898]" pageId="0" pageNumber="207">Type</emphasis>
:—
<materialsCitation id="3B133C8B6A2E3F572B864A43FC0FF875" ID-GBIF-Occurrence="3894379301" collectingDate="2010-07-20" country="China" county="Luding County" elevation="2950" location="Hailuogou" municipality="Mt. Gongga" pageId="0" pageNumber="207" specimenCount="1" stateProvince="Sichuan" typeStatus="holotype">
<collectingCountry id="F36C76466A2E3F572B864A43FE94F891" box="[215,296,1877,1898]" name="China" pageId="0" pageNumber="207">CHINA</collectingCountry>
.
<collectingRegion id="49BFF8346A2E3F572A604A43FE3EF890" box="[305,386,1877,1899]" country="China" name="Sichuan" pageId="0" pageNumber="207">Sichuan</collectingRegion>
:
<collectingCounty id="62A54E5A6A2E3F572ADA4A43FDA3F891" box="[395,543,1877,1899]" pageId="0" pageNumber="207">Luding County</collectingCounty>
,
<collectingMunicipality id="6BA0ACAC6A2E3F5729794A43FD20F891" box="[552,668,1877,1898]" pageId="0" pageNumber="207">Mt. Gongga</collectingMunicipality>
(Minyar Gonkar),
<location id="8EA4600D6A2E3F5728024A43FC07F890" LSID="urn:lsid:plazi:treatment:03D287C06A2E3F542BD94A27FEE5F867:8EA4600D6A2E3F5728024A43FC07F890" box="[851,955,1877,1899]" country="China" county="Luding County" municipality="Mt. Gongga" name="Hailuogou" pageId="0" pageNumber="207" stateProvince="Sichuan">Hailuogou</location>
, #3
<superScript id="7C0E9B9E6A2E3F57288D4A42FC54F89B" attach="left" box="[988,1000,1876,1888]" fontSize="5" pageId="0" pageNumber="207">rd</superScript>
<location id="8EA4600D6A2E3F5728BF4A43FB9BF891" LSID="urn:lsid:plazi:treatment:03D287C06A2E3F542BD94A27FEE5F867:8EA4600D6A2E3F5728BF4A43FB9BF891" box="[1006,1063,1877,1898]" country="China" county="Luding County" municipality="Mt. Gongga" name="Camp" pageId="0" pageNumber="207" stateProvince="Sichuan">Camp</location>
to
<location id="8EA4600D6A2E3F572F174A43FB5DF890" LSID="urn:lsid:plazi:treatment:03D287C06A2E3F542BD94A27FEE5F867:8EA4600D6A2E3F572F174A43FB5DF890" box="[1094,1249,1877,1899]" country="China" county="Luding County" municipality="Mt. Gongga" name="Chengmendong" pageId="0" pageNumber="207" stateProvince="Sichuan">Chengmendong</location>
, elev. ca.
<quantity id="4C839B336A2E3F572E6F4A43FA36F891" box="[1342,1418,1877,1899]" metricMagnitude="3" metricUnit="m" metricValue="2.95" pageId="0" pageNumber="207" unit="m" value="2950.0">
<elevation id="0056D1E56A2E3F572E6F4A43FA36F891" box="[1342,1418,1877,1899]" metricMagnitude="3" metricUnit="m" metricValue="2.95" pageId="0" pageNumber="207" unit="m" value="2950.0">2950 m</elevation>
</quantity>
,
<date id="FFC510166A2E3F572EC24A43FEA1F875" pageId="0" pageNumber="207" value="2010-07-20">
<collectingDate id="EF81E9FE6A2E3F572EC24A43FEA1F875" pageId="0" pageNumber="207" value="2010-07-20">20 July 2010</collectingDate>
</date>
,
<emphasis id="B90FEAC46A2E3F572A764A6CFE1DF875" box="[295,417,1913,1935]" italics="true" pageId="0" pageNumber="207">Hai He 1190</emphasis>
(
<emphasis id="B90FEAC46A2E3F572AE14A6FFE4AF875" box="[432,502,1913,1934]" italics="true" pageId="0" pageNumber="207">HG037</emphasis>
) (
<typeStatus id="54C088746A2E3F57295A4A6FFDDDF874" box="[523,609,1913,1935]" pageId="0" pageNumber="207" type="holotype">holotype</typeStatus>
CDBI; isotypes CDBI, CTC, MO)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C361655D6A2E3F572BD94A8BFAFBF7E4" pageId="0" pageNumber="207" type="diagnosis">
<paragraph id="8BC436D66A2E3F572BD94A8BFAFBF7E4" blockId="0.[136,1452,1841,2079]" pageId="0" pageNumber="207">
<emphasis id="B90FEAC46A2E3F572BD94A8BFF57F848" bold="true" box="[136,235,1949,1971]" pageId="0" pageNumber="207">Diagnosis</emphasis>
:—The new species is most similar to
<taxonomicName id="4C7B4D556A2E3F5729364A8BFC90F848" box="[615,812,1949,1971]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="0" pageNumber="207" phylum="Tracheophyta" rank="species" species="sinense">
<emphasis id="B90FEAC46A2E3F5729364A8BFC90F848" box="[615,812,1949,1971]" italics="true" pageId="0" pageNumber="207">Polystichum sinense</emphasis>
</taxonomicName>
(Christ in
<bibRefCitation id="EFEA4B276A2E3F5728C94A8BFB1EF849" author="Baroni, E. &amp; Christ, H." box="[920,1186,1949,1971]" pageId="0" pageNumber="207" pagination="288 - 292" refId="ref2327" refString="Baroni, E. &amp; Christ, H. (1901) Filices plantaeque filicibus affines in Shan-si septentrionali, provinceae Imperii Sinensis, a Rev. Patre Josepho Giraldi collectae. Nuovo Giornale Botanico Italiano, n. ser. 10: 288 - 292." type="book chapter" year="1901">Baroni &amp; Christ 1901: 289</bibRefCitation>
)
<bibRefCitation id="EFEA4B276A2E3F572FE04A8BFAE7F849" author="Christ, H." box="[1201,1371,1949,1971]" pageId="0" pageNumber="207" pagination="1 - 69" refId="ref2446" refString="Christ, H. (1905) Les collections de fougeres de la Chine au Museum d'Histoire Naturelle de Paris. Bulletin de la Societe Botanique de France 52 (Memoires 1): 1 - 69. https: // doi. org / 10.1080 / 00378941.1905.10829209" type="journal article" year="1905">Christ (1905: 30)</bibRefCitation>
, but the former has an oblong-lanceolate lamina with the lower portion only slightly contracted toward the lamina base, petiole scales that are blackish brown, and rachis scales that are narrowly lanceolate. In contrast, the latter has an elliptic-lanceolate lamina with the lower portion substantially contracted toward the lamina base, petiole scales that are brown, and broad-lanceolate rachis scales.
</paragraph>
</subSubSection>
<caption id="DF04665E6A2F3F562BD94AFDFC3AF7DE" ID-DOI="http://doi.org/10.5281/zenodo.7021473" ID-Zenodo-Dep="7021473" httpUri="https://zenodo.org/record/7021473/files/figure.png" pageId="1" pageNumber="208" startId="1.[136,229,2027,2049]" targetBox="[188,1393,210,1971]" targetPageId="1">
<paragraph id="8BC436D66A2F3F562BD94AFDFC3AF7DE" blockId="1.[136,1452,2027,2085]" pageId="1" pageNumber="208">
<emphasis id="B90FEAC46A2F3F562BD94AFDFEB9F7FB" bold="true" box="[136,261,2027,2049]" pageId="1" pageNumber="208">FIGURE 1.</emphasis>
An illustration of
<taxonomicName id="4C7B4D556A2F3F562A834AFDFD4AF7FB" authorityName="H. S. Kung &amp; Li Bing Zhang ex H. He &amp; Li Bing Zhang" authorityYear="2022" box="[466,758,2027,2048]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="208" phylum="Tracheophyta" rank="species" species="gonggashanense">
<emphasis id="B90FEAC46A2F3F562A834AFDFD4AF7FB" box="[466,758,2027,2048]" italics="true" pageId="1" pageNumber="208">Polystichum gonggashanense</emphasis>
</taxonomicName>
.
<emphasis id="B90FEAC46A2F3F5629AD4AFAFCADF7FA" box="[764,785,2028,2049]" italics="true" pageId="1" pageNumber="208">—</emphasis>
A. Habit.—B. Pinna.—C. Petiole scales.—D. Rachis scales.—E. Microscales.—F. Indusium (based on
<emphasis id="B90FEAC46A2F3F562AA44506FD17F7DF" box="[501,683,2063,2085]" italics="true" pageId="1" pageNumber="208">Li-Bing Zhang 169</emphasis>
; drawn by Jian Chen).
</paragraph>
</caption>
<caption id="DF04665E6A2C3F552BD94A8CFE45F803" ID-DOI="http://doi.org/10.5281/zenodo.7021475" ID-Zenodo-Dep="7021475" httpUri="https://zenodo.org/record/7021475/files/figure.png" pageId="2" pageNumber="209" startId="2.[136,229,1946,1968]" targetBox="[159,1428,205,1904]" targetPageId="2">
<paragraph id="8BC436D66A2C3F552BD94A8CFE45F803" blockId="2.[136,1451,1946,2040]" pageId="2" pageNumber="209">
<emphasis id="B90FEAC46A2C3F552BD94A8CFF40F854" bold="true" box="[136,252,1946,1968]" pageId="2" pageNumber="209">FIGURE 2.</emphasis>
A color plate of
<taxonomicName id="4C7B4D556A2C3F552AC94A8CFD0FF854" authorityName="H. S. Kung &amp; Li Bing Zhang ex H. He &amp; Li Bing Zhang" authorityYear="2022" box="[408,691,1946,1967]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="209" phylum="Tracheophyta" rank="species" species="gonggashanense">
<emphasis id="B90FEAC46A2C3F552AC94A8CFD0FF854" box="[408,691,1946,1967]" italics="true" pageId="2" pageNumber="209">Polystichum gonggashanense</emphasis>
</taxonomicName>
.—A. The highest peak of Mt. Gongga where the new species was discovered.— B. Habit.—C. Lower portion of petiole showing blackish brown scales.—D, E. Portions of adaxial pinnae showing scales.—F. Portion of abaxial pinna showing sori and scales.
</paragraph>
</caption>
<subSubSection id="C361655D6A2D3F542BD94D88FAC8FD77" pageId="3" pageNumber="210" type="description">
<paragraph id="8BC436D66A2D3F542BD94D88FAC8FD77" blockId="3.[136,1452,158,1948]" pageId="3" pageNumber="210">
<emphasis id="B90FEAC46A2D3F542BD94D88FEAEFF43" bold="true" box="[136,274,158,184]" pageId="3" pageNumber="210">Description</emphasis>
:—Plants
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tall. Rhizomes erect; fronds cespitose. Petioles
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long,
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in diam., stramineous; large scales ovate or ovate-lanceolate often
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long,
<quantity id="4C839B336A2D3F5428CA4DD4FC4DFF26" box="[923,1009,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="210" unit="mm" value="8.5">8.5 mm</quantity>
wide, acute at apex, ciliate or subentire on margins, the proximal ones uniformly blackish brown or brown, thick chartaceous, distally transitioning gradually to bicolorous with paler-colored margins, then to concolorous light yellow-brown and membranous; small scales linear to lanceolate. Laminae oblong-lanceolate,
<quantity id="4C839B336A2D3F54293B4C38FD66FEB2" box="[618,730,302,329]" metricMagnitude="-1" metricUnit="m" metricValue="5.3" metricValueMax="5.7" metricValueMin="4.9" pageId="3" pageNumber="210" unit="cm" value="53.0" valueMax="57.0" valueMin="49.0">49–57 cm</quantity>
long,
<quantity id="4C839B336A2D3F5428754C38FC29FEB2" box="[804,917,302,329]" metricMagnitude="-1" metricUnit="m" metricValue="1.85" metricValueMax="2.1" metricValueMin="1.6" pageId="3" pageNumber="210" unit="cm" value="18.5" valueMax="21.0" valueMin="16.0">16–21 cm</quantity>
wide, elongated-acuminate or caudate at apex, lower portion sterile, bipinnate. Rachises ca.
<quantity id="4C839B336A2D3F5429264C44FD0AFE96" box="[631,694,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="210" unit="mm" value="2.0">2 mm</quantity>
in diam.; rachis scales narrow-lanceolate, brownish, long-acuminate at apex, subentire on margins, spreading or reflexed. Pinnae 22–31 pairs, spreading, alternate, short-petiolate, middle pinnae
<quantity id="4C839B336A2D3F542B8B4C8CFEEBFE4E" box="[218,343,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="3" pageNumber="210" unit="cm" value="2.0" valueMax="2.5" valueMin="1.5">1.5–2.5 cm</quantity>
apart,
<quantity id="4C839B336A2D3F542AF54C8CFDA6FE4E" box="[420,538,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="7.25" metricValueMax="10.0" metricValueMin="4.5" pageId="3" pageNumber="210" unit="cm" value="7.25" valueMax="10.0" valueMin="4.5">4.5–10 cm</quantity>
long,
<quantity id="4C839B336A2D3F5429314C8CFD61FE4E" box="[608,733,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="2.0999999999999996" metricValueMax="2.3" metricValueMin="1.9" pageId="3" pageNumber="210" unit="cm" value="2.0999999999999996" valueMax="2.3" valueMin="1.9">1.9–2.3 cm</quantity>
wide, lanceolate, apex acuminate or caudate, base inequilateral, 1-pinnate; basal pinnae slightly shorter, 4.5–7.0 cm long, with 7–11 pairs of free pinnules; pinna rachis with linearlanceolate small scales (these denser abaxially), ca.
<quantity id="4C839B336A2D3F5429814CF4FCAEFE06" box="[720,786,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="210" unit="mm" value="5.0">5 mm</quantity>
long. Pinnules 13–16 pairs, sessile, spreading, herbaceous, deltoid-ovate,
<quantity id="4C839B336A2D3F542A7A4F10FE19FDDA" box="[299,421,518,545]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="3" pageNumber="210" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
long,
<quantity id="4C839B336A2D3F542ABF4F10FDCAFDDA" box="[494,630,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="5.5" metricValueMin="4.1" pageId="3" pageNumber="210" unit="mm" value="4.8" valueMax="5.5" valueMin="4.1">4.1–5.5 mm</quantity>
wide, auriculae deltate, base cuneate and slightly adnate (only the distal third of the pinna), apex acute, margins aristate-dentate, with 1.5–3.5-mm-long linear microscales on both surfaces, basal acroscopic pair the longest and lobed; veins 5–7 pairs per pinnule, dichotomous, slightly visible. Sori light brown, one row on each side of midrib, close to midrib; indusia peltate, rotund, membranous, margins subentire.
</paragraph>
</subSubSection>
<subSubSection id="C361655D6A2D3F542BEC4F80FA9EFD4B" box="[189,1314,662,689]" pageId="3" pageNumber="210" type="etymology">
<paragraph id="8BC436D66A2D3F542BEC4F80FA9EFD4B" blockId="3.[136,1452,158,1948]" box="[189,1314,662,689]" pageId="3" pageNumber="210">
<emphasis id="B90FEAC46A2D3F542BEC4F80FE82FD4B" bold="true" box="[189,318,662,688]" pageId="3" pageNumber="210">Etymology</emphasis>
:—Derived from
<emphasis id="B90FEAC46A2D3F5429504F80FD36FD4B" box="[513,650,662,688]" italics="true" pageId="3" pageNumber="210">gonggashan</emphasis>
in Chinese pinyin, meaning Mt. Gongga (Minyar Gonkar).
</paragraph>
</subSubSection>
<subSubSection id="C361655D6A2D3F542BEC4FACFC8AFD2F" box="[189,822,698,725]" pageId="3" pageNumber="210" type="vernacular_names">
<paragraph id="8BC436D66A2D3F542BEC4FACFC8AFD2F" blockId="3.[136,1452,158,1948]" box="[189,822,698,725]" pageId="3" pageNumber="210">
<emphasis id="B90FEAC46A2D3F542BEC4FACFE37FD2F" bold="true" box="[189,395,698,724]" pageId="3" pageNumber="210">Vernacular name</emphasis>
:—
<emphasis id="B90FEAC46A2D3F542AFE4FADFD87FD29" bold="true" box="[431,571,699,722]" pageId="3" pageNumber="210">ūḝ山φƦ</emphasis>
(gong ga shan er jue).
</paragraph>
</subSubSection>
<subSubSection id="C361655D6A2D3F542BEC4FC8FB44FCE6" pageId="3" pageNumber="210" type="distribution">
<paragraph id="8BC436D66A2D3F542BEC4FC8FB44FCE6" blockId="3.[136,1452,158,1948]" pageId="3" pageNumber="210">
<emphasis id="B90FEAC46A2D3F542BEC4FC8FEF3FD03" bold="true" box="[189,335,734,760]" pageId="3" pageNumber="210">Distribution</emphasis>
:—It was found on the eastern, southeastern, and western slopes of Mt. Gongga in Jiulong, Luding, and Kangding counties,
<collectingRegion id="49BFF8346A2D3F542AC94E14FDE3FCE7" box="[408,607,770,796]" country="China" name="Sichuan" pageId="3" pageNumber="210">Sichuan Province</collectingRegion>
, Southwest
<collectingCountry id="F36C76466A2D3F5429B64E14FC91FCE7" box="[743,813,770,796]" name="China" pageId="3" pageNumber="210">China</collectingCountry>
, with elevational range of
<quantity id="4C839B336A2D3F542F074E14FB48FCE6" box="[1110,1268,770,797]" metricMagnitude="3" metricUnit="m" metricValue="2.85" metricValueMax="3.2" metricValueMin="2.5" pageId="3" pageNumber="210" unit="m" value="2850.0" valueMax="3200.0" valueMin="2500.0">2500–3200 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C361655D6A2D3F542BEC4E30FEC0FC0F" pageId="3" pageNumber="210" type="conservation">
<paragraph id="8BC436D66A2D3F542BEC4E30FEC0FC0F" blockId="3.[136,1452,158,1948]" pageId="3" pageNumber="210">
<emphasis id="B90FEAC46A2D3F542BEC4E30FE5BFCBB" bold="true" box="[189,487,806,832]" pageId="3" pageNumber="210">Conservation assessment</emphasis>
:—Based on thorough explorations by Kai-Yong Lang
<emphasis id="B90FEAC46A2D3F542F1B4E31FB3EFCBB" box="[1098,1154,806,832]" italics="true" pageId="3" pageNumber="210">et al.</emphasis>
in 1982, by L-BZ in 1989, 1990, and 1993, and by HH in 2010, only four very small populations each with a few individuals were found.All these populations are in high elevations of the same mountain. The expansion of the touristic facilities and global warming threaten the survival of
<taxonomicName id="4C7B4D556A2D3F542AF04E84FD4DFC57" authorityName="H. S. Kung &amp; Li Bing Zhang ex H. He &amp; Li Bing Zhang" authorityYear="2022" box="[417,753,914,940]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="gonggashanense">
<emphasis id="B90FEAC46A2D3F542AF04E84FD4DFC57" box="[417,753,914,940]" italics="true" pageId="3" pageNumber="210">Polystichum gonggashanense</emphasis>
</taxonomicName>
and the population in the first locality already disappeared. We suggest that the status of
<taxonomicName id="4C7B4D556A2D3F542A844EA0FC9FFC2B" authorityName="H. S. Kung &amp; Li Bing Zhang ex H. He &amp; Li Bing Zhang" authorityYear="2022" box="[469,803,950,976]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="gonggashanense">
<emphasis id="B90FEAC46A2D3F542A844EA0FC9FFC2B" box="[469,803,950,976]" italics="true" pageId="3" pageNumber="210">Polystichum gonggashanense</emphasis>
</taxonomicName>
be listed as CR (Critically Endangered) following IUCN criteria (
<bibRefCitation id="EFEA4B276A2D3F542BB74ECCFECCFC0F" author="IUCN Standards &amp; Petitions Committee" box="[230,368,986,1012]" pageId="3" pageNumber="210" refId="ref2494" refString="IUCN Standards and Petitions Committee (2019) Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Committee. Available from: http: // www. iucnredlist. org / documents / RedListGuidelines. pdf (Accessed 28 December 2021)." type="book" year="2019">IUCN 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C361655D6A2D3F542BEC4EE8FDB8FB53" pageId="3" pageNumber="210" type="materials_examined">
<paragraph id="8BC436D66A2D3F542BEC4EE8FDB8FB53" blockId="3.[136,1452,158,1948]" pageId="3" pageNumber="210">
<materialsCitation id="3B133C8B6A2D3F542BEC4EE8FDBCFB53" ID-GBIF-Occurrence="3894379302" collectingDate="1982-07" collectingDateMax="1990-10" collectingDateMin="1982-07" country="China" county="Luding County" elevation="2980" location="Hailuogou" municipality="Mt. Gongga" pageId="3" pageNumber="210" specimenCount="2" stateProvince="Sichuan" typeStatus="paratype">
<emphasis id="B90FEAC46A2D3F542BEC4EE8FD5EFBE3" bold="true" box="[189,738,1022,1048]" pageId="3" pageNumber="210">
Additional specimens examined (
<typeStatus id="54C088746A2D3F5429304EE8FD67FBE3" box="[609,731,1022,1048]" pageId="3" pageNumber="210" type="paratype">paratypes</typeStatus>
)
</emphasis>
:—
<collectingCountry id="F36C76466A2D3F5428554EE8FCDEFBE3" box="[772,866,1022,1048]" name="China" pageId="3" pageNumber="210">CHINA</collectingCountry>
.
<collectingRegion id="49BFF8346A2D3F5428244EE8FC68FBE3" box="[885,980,1022,1048]" country="China" name="Sichuan" pageId="3" pageNumber="210">Sichuan</collectingRegion>
:
<collectingCounty id="62A54E5A6A2D3F5428B64EE8FB20FBE3" box="[999,1180,1022,1048]" pageId="3" pageNumber="210">Luding County</collectingCounty>
,
<collectingMunicipality id="6BA0ACAC6A2D3F542FFE4EE9FAFCFBE3" box="[1199,1344,1022,1049]" pageId="3" pageNumber="210">Mt. Gongga</collectingMunicipality>
(Minyar Gonkar),
<location id="8EA4600D6A2D3F542BA24934FED0FBC7" LSID="urn:lsid:plazi:treatment:03D287C06A2E3F542BD94A27FEE5F867:8EA4600D6A2D3F542BA24934FED0FBC7" box="[243,364,1058,1084]" country="China" county="Luding County" municipality="Mt. Gongga" name="Hailuogou" pageId="3" pageNumber="210" stateProvince="Sichuan">Hailuogou</location>
, elev. ca.
<quantity id="4C839B336A2D3F542A894934FD93FBC6" box="[472,559,1058,1085]" metricMagnitude="3" metricUnit="m" metricValue="2.98" pageId="3" pageNumber="210" unit="m" value="2980.0">
<elevation id="0056D1E56A2D3F542A894934FD93FBC6" box="[472,559,1058,1085]" metricMagnitude="3" metricUnit="m" metricValue="2.98" pageId="3" pageNumber="210" unit="m" value="2980.0">2980 m</elevation>
</quantity>
,
<date id="FFC510166A2D3F54296B4934FD44FBC7" box="[570,760,1058,1084]" pageId="3" pageNumber="210" value="1989-10-29">
<collectingDate id="EF81E9FE6A2D3F54296B4934FD44FBC7" box="[570,760,1058,1084]" pageId="3" pageNumber="210" value="1989-10-29">29 October 1989</collectingDate>
</date>
,
<emphasis id="B90FEAC46A2D3F5428524935FC64FBC7" box="[771,984,1058,1084]" italics="true" pageId="3" pageNumber="210">
<location id="8EA4600D6A2D3F5428524935FC1BFBC7" LSID="urn:lsid:plazi:treatment:03D287C06A2E3F542BD94A27FEE5F867:8EA4600D6A2D3F5428524935FC1BFBC7" box="[771,935,1058,1084]" country="China" county="Luding County" municipality="Mt. Gongga" name="Li-Bing Zhang" pageId="3" pageNumber="210" stateProvince="Sichuan">Li-Bing Zhang</location>
169
</emphasis>
(KUN, PYU); ibid., 1990,
<emphasis id="B90FEAC46A2D3F542E594935FF0CFB9B" italics="true" pageId="3" pageNumber="210">
<location id="8EA4600D6A2D3F542E594935FA10FBC7" LSID="urn:lsid:plazi:treatment:03D287C06A2E3F542BD94A27FEE5F867:8EA4600D6A2D3F542E594935FA10FBC7" box="[1288,1452,1058,1084]" country="China" county="Luding County" municipality="Mt. Gongga" name="Li-Bing Zhang" pageId="3" pageNumber="210" stateProvince="Sichuan">Li-Bing Zhang</location>
911
</emphasis>
(CDBI);
<location id="8EA4600D6A2D3F542A4D4950FE75FB9B" LSID="urn:lsid:plazi:treatment:03D287C06A2E3F542BD94A27FEE5F867:8EA4600D6A2D3F542A4D4950FE75FB9B" box="[284,457,1094,1120]" country="China" county="Luding County" municipality="Mt. Gongga" name="Jiulong County" pageId="3" pageNumber="210" stateProvince="Sichuan">Jiulong County</location>
,
<location id="8EA4600D6A2D3F542A854950FD8CFB9B" LSID="urn:lsid:plazi:treatment:03D287C06A2E3F542BD94A27FEE5F867:8EA4600D6A2D3F542A854950FD8CFB9B" box="[468,560,1094,1120]" country="China" county="Luding County" municipality="Mt. Gongga" name="Hongba" pageId="3" pageNumber="210" stateProvince="Sichuan">Hongba</location>
, elev. ca.
<quantity id="4C839B336A2D3F5429CD4950FD48FB9A" box="[668,756,1094,1121]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="210" unit="m" value="2500.0">
<elevation id="0056D1E56A2D3F5429CD4950FD48FB9A" box="[668,756,1094,1121]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="210" unit="m" value="2500.0">2500 m</elevation>
</quantity>
, on forest margin,
<date id="FFC510166A2D3F5428974950FBDFFB9B" box="[966,1123,1094,1120]" pageId="3" pageNumber="210" value="1990-10">
<collectingDate id="EF81E9FE6A2D3F5428974950FBDFFB9B" box="[966,1123,1094,1120]" pageId="3" pageNumber="210" value="1990-10">October 1990</collectingDate>
</date>
,
<emphasis id="B90FEAC46A2D3F542F3F4951FAEDFB9B" box="[1134,1361,1094,1120]" italics="true" pageId="3" pageNumber="210">
<location id="8EA4600D6A2D3F542F3F4951FAAEFB9B" LSID="urn:lsid:plazi:treatment:03D287C06A2E3F542BD94A27FEE5F867:8EA4600D6A2D3F542F3F4951FAAEFB9B" box="[1134,1298,1094,1120]" country="China" county="Luding County" municipality="Mt. Gongga" name="Li-Bing Zhang" pageId="3" pageNumber="210" stateProvince="Sichuan">Li-Bing Zhang</location>
2786
</emphasis>
(CDBI, PYU);
<location id="8EA4600D6A2D3F542B89497CFE1DFB7F" LSID="urn:lsid:plazi:treatment:03D287C06A2E3F542BD94A27FEE5F867:8EA4600D6A2D3F542B89497CFE1DFB7F" box="[216,417,1130,1156]" country="China" county="Luding County" municipality="Mt. Gongga" name="Kangding County" pageId="3" pageNumber="210" stateProvince="Sichuan">Kangding County</location>
,
<location id="8EA4600D6A2D3F542AFD497CFE4CFB7F" LSID="urn:lsid:plazi:treatment:03D287C06A2E3F542BD94A27FEE5F867:8EA4600D6A2D3F542AFD497CFE4CFB7F" box="[428,496,1130,1156]" country="China" county="Luding County" municipality="Mt. Gongga" name="Liuba" pageId="3" pageNumber="210" stateProvince="Sichuan">Liuba</location>
, elev. ca.
<quantity id="4C839B336A2D3F54290D497CFD0FFB7E" box="[604,691,1130,1157]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="3" pageNumber="210" unit="m" value="3200.0">
<elevation id="0056D1E56A2D3F54290D497CFD0FFB7E" box="[604,691,1130,1157]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="3" pageNumber="210" unit="m" value="3200.0">3200 m</elevation>
</quantity>
,
<date id="FFC510166A2D3F5429EE497CFC93FB7F" box="[703,815,1130,1156]" pageId="3" pageNumber="210" value="1982-07">
<collectingDate id="EF81E9FE6A2D3F5429EE497CFC93FB7F" box="[703,815,1130,1156]" pageId="3" pageNumber="210" value="1982-07">July 1982</collectingDate>
</date>
,
<emphasis id="B90FEAC46A2D3F54286B497DFBEFFB7F" box="[826,1107,1130,1156]" italics="true" pageId="3" pageNumber="210">Kai-Yong Lang et al. 752</emphasis>
(KUN, PE); same town, 1990,
<emphasis id="B90FEAC46A2D3F542BD94999FEE6FB53" box="[136,346,1166,1192]" italics="true" pageId="3" pageNumber="210">Li-Bing Zhang s.n.</emphasis>
(CDBI, PYU)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C361655D6A2D3F542BEC49A4FEE5F867" pageId="3" pageNumber="210" type="discussion">
<paragraph id="8BC436D66A2D3F542BEC49A4FDDEFAEF" blockId="3.[136,1452,158,1948]" pageId="3" pageNumber="210">
<emphasis id="B90FEAC46A2D3F542BEC49A4FE38FB37" bold="true" box="[189,388,1202,1228]" pageId="3" pageNumber="210">Taxonomic notes</emphasis>
:—We examined all specimens available at CTC and it appears that all sporangia were dehisced. There were fewer than 32 spores per sporangium and most sporangia had only 10–20 spores upon examination. It is possible that the new species is apomictic.
</paragraph>
<paragraph id="8BC436D66A2D3F542BEC4809FB75F987" blockId="3.[136,1452,158,1948]" pageId="3" pageNumber="210">
In middle to high elevations of the Sino-Himalayan region species of
<taxonomicName id="4C7B4D556A2D3F5428884808FBDFFAC3" authorityName="Roth" authorityYear="1800" box="[985,1123,1310,1336]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="genus">
<emphasis id="B90FEAC46A2D3F5428884808FBDFFAC3" box="[985,1123,1310,1336]" italics="true" pageId="3" pageNumber="210">Polystichum</emphasis>
</taxonomicName>
are common, most of which belong to
<emphasis id="B90FEAC46A2D3F542BAD4855FEAEFAA7" box="[252,274,1347,1372]" italics="true" pageId="3" pageNumber="210">P.</emphasis>
sect.
<emphasis id="B90FEAC46A2D3F542A074854FE73FAA7" box="[342,463,1346,1372]" italics="true" pageId="3" pageNumber="210">Hypopeltis</emphasis>
(
<bibRefCitation id="EFEA4B276A2D3F542AB14854FD7FFAA7" author="Michaux, A." box="[480,707,1346,1372]" pageId="3" pageNumber="210" refId="ref2811" refString="Michaux, A. (1803) Flora Boreali-Abericana, sistens carcteres plantarum quas in America septentrionali collegit et detexit Andreas" type="book" year="1803">Michaux 1803: 266</bibRefCitation>
) T.
<bibRefCitation id="EFEA4B276A2D3F5429B64855FC09FAA7" author="Moore, T." box="[743,949,1346,1373]" pageId="3" pageNumber="210" refId="ref2868" refString="Moore, T. (1857 - 1862) Index Filicum. W. Pamplin, London, 396 pp." type="book" year="1857">Moore (1857: 84)</bibRefCitation>
taxonomically (
<bibRefCitation id="EFEA4B276A2D3F542F3E4854FA1CFAA7" author="Zhang, L. - B. &amp; Barrington, D. S." box="[1135,1440,1346,1372]" pageId="3" pageNumber="210" refId="ref3016" refString="Zhang, L. - B. &amp; Barrington, D. S. (2013) Polystichum Roth. In: Wu, Z. - Y., Raven, P. H. &amp; Hong, D. - Y. (Eds.) Flora of China, vols. 2 - 3. Science Press, Beijing &amp; Missouri Botanical Presss, St. Louis, pp. 629 - 713." type="book" year="2013">Zhang &amp; Barrington 2013</bibRefCitation>
). The taxonomy of this section is difficult partially because the species are over-described (
<bibRefCitation id="EFEA4B276A2D3F542F244870FA1CFA7B" author="Zhang, L. - B. &amp; Barrington, D. S." box="[1141,1440,1382,1408]" pageId="3" pageNumber="210" refId="ref3016" refString="Zhang, L. - B. &amp; Barrington, D. S. (2013) Polystichum Roth. In: Wu, Z. - Y., Raven, P. H. &amp; Hong, D. - Y. (Eds.) Flora of China, vols. 2 - 3. Science Press, Beijing &amp; Missouri Botanical Presss, St. Louis, pp. 629 - 713." type="book" year="2013">Zhang &amp; Barrington 2013</bibRefCitation>
). Between 1983 and 1993, seven “species” were described for
<taxonomicName id="4C7B4D556A2D3F542811489DFC13FA5F" box="[832,943,1419,1444]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="sinense">
<emphasis id="B90FEAC46A2D3F542811489DFC13FA5F" box="[832,943,1419,1444]" italics="true" pageId="3" pageNumber="210">P. sinense</emphasis>
</taxonomicName>
alone (Ching in
<bibRefCitation id="EFEA4B276A2D3F542F21489DFB67FA5F" author="Wu, C. - Y." box="[1136,1243,1418,1445]" pageId="3" pageNumber="210" refId="ref2916" refString="Wu, C. - Y. (1983) Flora Xizangica, vol. 1. Science Press, Beijing, 791 pp." type="book" year="1983">Wu 1983</bibRefCitation>
, Ching &amp; S.K.Wu in
<bibRefCitation id="EFEA4B276A2D3F542BF548B9FEB2FA33" author="Wu, C. - Y." box="[164,270,1454,1481]" pageId="3" pageNumber="210" refId="ref2916" refString="Wu, C. - Y. (1983) Flora Xizangica, vol. 1. Science Press, Beijing, 791 pp." type="book" year="1983">Wu 1983</bibRefCitation>
,
<bibRefCitation id="EFEA4B276A2D3F542A4948B9FE2EFA33" author="Yang, C. - Y." box="[280,402,1454,1481]" pageId="3" pageNumber="210" refId="ref2941" refString="Yang, C. - Y. (1992) Flora Xinjiangensis, vol. 1. Xinjiang Science &amp; Technology &amp; Hygiene Publishing House, Urumuqi, 337 pp." type="book" year="1992">Yang 1992</bibRefCitation>
) including
<taxonomicName id="4C7B4D556A2D3F54294348B9FBA7FA33" authority="Chang Y. Yang (1992: 302)" authorityName="Chang Y. Yang" authorityYear="1992" box="[530,1051,1454,1481]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="alatawshanicum">
<emphasis id="B90FEAC46A2D3F54294348B9FD5AFA33" box="[530,742,1454,1480]" italics="true" pageId="3" pageNumber="210">P. alatawshanicum</emphasis>
Chang Y.
<bibRefCitation id="EFEA4B276A2D3F54280348B9FBA7FA33" author="Yang, C. - Y." box="[850,1051,1454,1481]" pageId="3" pageNumber="210" refId="ref2941" refString="Yang, C. - Y. (1992) Flora Xinjiangensis, vol. 1. Xinjiang Science &amp; Technology &amp; Hygiene Publishing House, Urumuqi, 337 pp." type="book" year="1992">Yang (1992: 302)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C7B4D556A2D3F542F7748B9FAACFA33" authority="Ching" authorityName="Ching" box="[1062,1296,1454,1480]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="atuntzeense">
<emphasis id="B90FEAC46A2D3F542F7748B9FB79FA33" box="[1062,1221,1455,1480]" italics="true" pageId="3" pageNumber="210">P. atuntzeense</emphasis>
Ching
</taxonomicName>
in
<bibRefCitation id="EFEA4B276A2D3F542E6548B9FF01FA17" author="Wu, C. - Y." pageId="3" pageNumber="210" refId="ref2916" refString="Wu, C. - Y. (1983) Flora Xizangica, vol. 1. Science Press, Beijing, 791 pp." type="book" year="1983">Wu (1983: 231)</bibRefCitation>
,
<taxonomicName id="4C7B4D556A2D3F542B9D48C5FDD4FA17" authority="Ching &amp; S. K. Wu" authorityName="Ching &amp; S. K. Wu" box="[204,616,1490,1516]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="austrotibeticum">
<emphasis id="B90FEAC46A2D3F542B9D48C5FE25FA17" box="[204,409,1490,1516]" italics="true" pageId="3" pageNumber="210">P. austrotibeticum</emphasis>
Ching &amp; S.K.Wu
</taxonomicName>
in
<bibRefCitation id="EFEA4B276A2D3F5429C048C5FCF5FA17" author="Wu, C. - Y." box="[657,841,1490,1517]" pageId="3" pageNumber="210" refId="ref2916" refString="Wu, C. - Y. (1983) Flora Xizangica, vol. 1. Science Press, Beijing, 791 pp." type="book" year="1983">Wu (1983: 233)</bibRefCitation>
,
<taxonomicName id="4C7B4D556A2D3F54280948C5FB1BFA17" authority="Ching &amp; S. K. Wu" authorityName="Ching &amp; S. K. Wu" box="[856,1191,1490,1516]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="decorum">
<emphasis id="B90FEAC46A2D3F54280948C5FC64FA17" box="[856,984,1490,1516]" italics="true" pageId="3" pageNumber="210">P. decorum</emphasis>
Ching &amp; S.K.Wu
</taxonomicName>
in
<bibRefCitation id="EFEA4B276A2D3F542F8148C5FA35FA17" author="Wu, C. - Y." box="[1232,1417,1490,1517]" pageId="3" pageNumber="210" refId="ref2916" refString="Wu, C. - Y. (1983) Flora Xizangica, vol. 1. Science Press, Beijing, 791 pp." type="book" year="1983">Wu (1983: 229)</bibRefCitation>
,
<taxonomicName id="4C7B4D556A2D3F542EC648C5FE7BF9EB" authority="Ching &amp; S. K. Wu" authorityName="Ching &amp; S. K. Wu" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="ellipticum">
<emphasis id="B90FEAC46A2D3F542EC648C5FF44F9EB" italics="true" pageId="3" pageNumber="210">P. ellipticum</emphasis>
Ching &amp; S.K.Wu
</taxonomicName>
in
<bibRefCitation id="EFEA4B276A2D3F542AA148E1FD15F9EB" author="Wu, C. - Y." box="[496,681,1526,1553]" pageId="3" pageNumber="210" refId="ref2916" refString="Wu, C. - Y. (1983) Flora Xizangica, vol. 1. Science Press, Beijing, 791 pp." type="book" year="1983">Wu (1983: 233)</bibRefCitation>
,
<taxonomicName id="4C7B4D556A2D3F5429E648E1FC2DF9EB" authority="Ching" authorityName="Ching" box="[695,913,1526,1552]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="lhasaense">
<emphasis id="B90FEAC46A2D3F5429E648E1FCFFF9EB" box="[695,835,1526,1552]" italics="true" pageId="3" pageNumber="210">P. lhasaense</emphasis>
Ching
</taxonomicName>
in
<bibRefCitation id="EFEA4B276A2D3F5428EB48E1FBCFF9EB" author="Wu, C. - Y." box="[954,1139,1526,1553]" pageId="3" pageNumber="210" refId="ref2916" refString="Wu, C. - Y. (1983) Flora Xizangica, vol. 1. Science Press, Beijing, 791 pp." type="book" year="1983">Wu (1983: 230)</bibRefCitation>
, and
<taxonomicName id="4C7B4D556A2D3F542FE248E1FED2F9CF" authority="Chang Y. Yang (1992: 302)" authorityName="Chang Y. Yang" authorityYear="1992" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="parasinense">
<emphasis id="B90FEAC46A2D3F542FE248E1FAE4F9EB" box="[1203,1368,1527,1552]" italics="true" pageId="3" pageNumber="210">P. parasinense</emphasis>
Chang Y.
<bibRefCitation id="EFEA4B276A2D3F542BCE4B0DFED2F9CF" author="Yang, C. - Y." box="[159,366,1562,1589]" pageId="3" pageNumber="210" refId="ref2941" refString="Yang, C. - Y. (1992) Flora Xinjiangensis, vol. 1. Xinjiang Science &amp; Technology &amp; Hygiene Publishing House, Urumuqi, 337 pp." type="book" year="1992">Yang (1992: 302)</bibRefCitation>
</taxonomicName>
. Relevant molecular data for
<taxonomicName id="4C7B4D556A2D3F54299A4B0CFCE9F9CF" authorityName="Roth" authorityYear="1800" box="[715,853,1562,1588]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="genus">
<emphasis id="B90FEAC46A2D3F54299A4B0CFCE9F9CF" box="[715,853,1562,1588]" italics="true" pageId="3" pageNumber="210">Polystichum</emphasis>
</taxonomicName>
so far have not yet been based on extensive taxon sampling of
<emphasis id="B90FEAC46A2D3F542A424B29FE94F9A3" box="[275,296,1599,1624]" italics="true" pageId="3" pageNumber="210">P.</emphasis>
sect.
<emphasis id="B90FEAC46A2D3F542A364B28FE5CF9A3" box="[359,480,1598,1624]" italics="true" pageId="3" pageNumber="210">Hypopeltis</emphasis>
(
<bibRefCitation id="EFEA4B276A2D3F542ABF4B28FD2AF9A3" author="Liu, H. - M. &amp; Zhang, X. - C. &amp; Chen, Z. - D. &amp; Qiu, Y. - L." box="[494,662,1598,1625]" pageId="3" pageNumber="210" pagination="631 - 638" refId="ref2727" refString="Liu, H. - M., Zhang, X. - C., Chen, Z. - D. &amp; Qiu, Y. - L. (2007) Inclusion of the Eastern Asia endemic genus Sorolepidium in Polystichum (Dryopteridaceae): Evidence from the chloroplast rbcL gene and morphological characteristics. Chinese Science Bulletin 52: 631 - 638. https: // doi. org / 10.1007 / s 11434 - 007 - 0115 - 2" type="journal article" year="2007">
Liu
<emphasis id="B90FEAC46A2D3F54294C4B29FDF2F9A3" box="[541,590,1598,1624]" italics="true" pageId="3" pageNumber="210">et al</emphasis>
. 2007
</bibRefCitation>
,
<bibRefCitation id="EFEA4B276A2D3F5429F04B29FC1EF9A3" author="Le Pechon, T. &amp; He, H. &amp; Zhang, L. &amp; Zhou, X. - M. &amp; Gao, X. - F. &amp; Zhang, L. - B." box="[673,930,1598,1625]" pageId="3" pageNumber="210" pagination="55" refId="ref2548" refString="Le Pechon, T., He, H., Zhang, L., Zhou, X. - M., Gao, X. - F. &amp; Zhang, L. - B. (2016 a) Using a multilocus phylogeny to test morphology-based classifications of one of the largest fern genera Polystichum (Dryopteridaceae). BMC Evolutionary Biology 16: 55. https: // doi. org / 10.1186 / s 12862 - 016 - 0626 - z" type="journal article" year="2016">
Le Péchon
<emphasis id="B90FEAC46A2D3F54284C4B29FCF2F9A3" box="[797,846,1598,1624]" italics="true" pageId="3" pageNumber="210">et al</emphasis>
. 2016a
</bibRefCitation>
,
<bibRefCitation id="EFEA4B276A2D3F5428FC4B28FC44F9A3" author="Le Pechon, T. &amp; Zhang, L. &amp; Zhou, X. - M. &amp; He, H. &amp; Gao, X. - F. &amp; Zhang, L. - B." box="[941,1016,1598,1624]" pageId="3" pageNumber="210" pagination="324 - 336" refId="ref2635" refString="Le Pechon, T., Zhang, L., Zhou, X. - M., He, H., Gao, X. - F. &amp; Zhang, L. - B. (2016 b) A well-sampled phylogenetic analysis of the polystichoid ferns (Dryopteridaceae) suggests a complex biogeographical history involving both boreotropical migrations and recent transoceanic dispersals. Molecular Phylogenetics and Evolution 98: 324 - 336. https: // doi. org / 10.1016 / j. ympev. 2016.02.018" type="journal article" year="2016">2016b</bibRefCitation>
).
<taxonomicName id="4C7B4D556A2D3F542F5B4B28FAEAF9A3" authorityName="H. S. Kung &amp; Li Bing Zhang ex H. He &amp; Li Bing Zhang" authorityYear="2022" box="[1034,1366,1598,1624]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="gonggashanense">
<emphasis id="B90FEAC46A2D3F542F5B4B28FAEAF9A3" box="[1034,1366,1598,1624]" italics="true" pageId="3" pageNumber="210">Polystichum gonggashanense</emphasis>
</taxonomicName>
has not been described so far partially because it is a rare species endemic to a small area in Mt. Gongga.
</paragraph>
<paragraph id="8BC436D66A2D3F542BEC4B90FF57F8F7" blockId="3.[136,1452,158,1948]" pageId="3" pageNumber="210">
Notably,
<taxonomicName id="4C7B4D556A2D3F542A724B90FDA9F95B" box="[291,533,1670,1696]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="decorum">
<emphasis id="B90FEAC46A2D3F542A724B90FDA9F95B" box="[291,533,1670,1696]" italics="true" pageId="3" pageNumber="210">Polystichum decorum</emphasis>
</taxonomicName>
, thought as conspecific with
<taxonomicName id="4C7B4D556A2D3F5428094B91FB91F95B" authorityName="H. S. Kung &amp; Li Bing Zhang ex H. He &amp; Li Bing Zhang" authorityYear="2022" box="[856,1069,1670,1696]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="gonggashanense">
<emphasis id="B90FEAC46A2D3F5428094B91FB91F95B" box="[856,1069,1670,1696]" italics="true" pageId="3" pageNumber="210">P. gonggashanense</emphasis>
</taxonomicName>
by the reviewers of
<emphasis id="B90FEAC46A2D3F542E5C4B91FF41F93F" italics="true" pageId="3" pageNumber="210">Acta Botanica Yunnanica</emphasis>
as stated in the introduction, is a synonym of
<taxonomicName id="4C7B4D556A2D3F54285C4BBDFCC0F93F" box="[781,892,1707,1732]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="sinense">
<emphasis id="B90FEAC46A2D3F54285C4BBDFCC0F93F" box="[781,892,1707,1732]" italics="true" pageId="3" pageNumber="210">P. sinense</emphasis>
</taxonomicName>
. The distinction between
<taxonomicName id="4C7B4D556A2D3F542FF24BBDFAC7F93F" authorityName="H. S. Kung &amp; Li Bing Zhang ex H. He &amp; Li Bing Zhang" authorityYear="2022" box="[1187,1403,1706,1732]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="gonggashanense">
<emphasis id="B90FEAC46A2D3F542FF24BBDFAC7F93F" box="[1187,1403,1706,1732]" italics="true" pageId="3" pageNumber="210">P. gonggashanense</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7B4D556A2D3F542BD94BD9FF4AF913" box="[136,246,1743,1768]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="sinense">
<emphasis id="B90FEAC46A2D3F542BD94BD9FF4AF913" box="[136,246,1743,1768]" italics="true" pageId="3" pageNumber="210">P. sinense</emphasis>
</taxonomicName>
is not neglectable in the color of petiole scales, the lamina shape, and the shape of rachis scales (see above;
<tableCitation id="C6F9036D6A2D3F542BD94BE4FF66F8F7" box="[136,218,1778,1804]" captionStart="TABLE 1" captionStartId="4.[136,224,159,183]" captionTargetPageId="4" captionText="TABLE 1. Morphological comparison among Polystichum gongboense, P. gonggashanense, and P. haleakalense, and P. kangdingense." httpUri="http://table.plazi.org/id/DF04665E6A2A3F532BD94D89FE9EFF20" pageId="3" pageNumber="210" tableUuid="DF04665E6A2A3F532BD94D89FE9EFF20">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BC436D66A2D3F542BEC4A01FEE5F867" blockId="3.[136,1452,158,1948]" pageId="3" pageNumber="210">
In addition to
<taxonomicName id="4C7B4D556A2D3F542A064A00FD85F8CB" box="[343,569,1814,1840]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="sinense">
<emphasis id="B90FEAC46A2D3F542A064A00FD85F8CB" box="[343,569,1814,1840]" italics="true" pageId="3" pageNumber="210">Polystichum sinense</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7B4D556A2D3F5429174A01FCA0F8CB" authorityName="H. S. Kung &amp; Li Bing Zhang ex H. He &amp; Li Bing Zhang" authorityYear="2022" box="[582,796,1814,1840]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="gonggashanense">
<emphasis id="B90FEAC46A2D3F5429174A01FCA0F8CB" box="[582,796,1814,1840]" italics="true" pageId="3" pageNumber="210">P. gonggashanense</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="4C7B4D556A2D3F54288A4A01FA10F8CB" authority="Brackenridge (1854: 204)" authorityName="Brackenridge" authorityPageNumber="204" authorityYear="1854" box="[987,1452,1814,1841]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="haleakalense">
<emphasis id="B90FEAC46A2D3F54288A4A01FB3BF8CB" box="[987,1159,1814,1840]" italics="true" pageId="3" pageNumber="210">P. haleakalense</emphasis>
<bibRefCitation id="EFEA4B276A2D3F542FDC4A00FA10F8CB" author="Brackenridge, W. D." box="[1165,1452,1814,1841]" pageId="3" pageNumber="210" refId="ref2374" refString="Brackenridge, W. D. (1854) Botany. Cryptogamia. Filices, including Lycopodiaceae and Hydropterides. In: Wilkes, C. (Ed.) United States Exploring Expedition. During the years 1838, 1839, 1840, 1841, 1842. Under the command of Charles Wilkes, U. S. N., vol. 16. pp. 1 - 357." type="journal volume" year="1854">Brackenridge (1854: 204)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C7B4D556A2D3F542BE84A2DFBB3F8AE" authority="H. S. Kung &amp; Li Bing Zhang ex Li Bing Zhang (2012: 58)" authorityName="H. S. Kung &amp; Li Bing Zhang ex Li Bing Zhang" authorityPageNumber="58" authorityYear="2012" box="[185,1039,1850,1877]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="210" phylum="Tracheophyta" rank="species" species="kangdingense">
<emphasis id="B90FEAC46A2D3F542BE84A2DFECDF8AF" box="[185,369,1850,1876]" italics="true" pageId="3" pageNumber="210">P. kangdingense</emphasis>
H.S.Kung &amp; Li Bing Zhang ex Li Bing
<bibRefCitation id="EFEA4B276A2D3F5428164A2CFBB3F8AE" author="Zhang, L. - B." box="[839,1039,1850,1877]" pageId="3" pageNumber="210" pagination="57 - 60" refId="ref2972" refString="Zhang, L. - B. (2012) Taxonomic and nomenclatural notes on the fern genus Polystichum (Dryopteridaceae) from China. Phytotaxa 60: 57 - 60. https: // doi. org / 10.11646 / phytotaxa. 60.1.8" type="journal article" year="2012">Zhang (2012: 58)</bibRefCitation>
</taxonomicName>
in having bipinnate laminae, ovatelanceolate petiole scales, lanceolate rachis scales, and linear microscales. These four species are compared with one another in
<tableCitation id="C6F9036D6A2D3F542BAE4A94FEE5F867" box="[255,345,1922,1948]" captionStart="TABLE 1" captionStartId="4.[136,224,159,183]" captionTargetPageId="4" captionText="TABLE 1. Morphological comparison among Polystichum gongboense, P. gonggashanense, and P. haleakalense, and P. kangdingense." httpUri="http://table.plazi.org/id/DF04665E6A2A3F532BD94D89FE9EFF20" pageId="3" pageNumber="210" tableUuid="DF04665E6A2A3F532BD94D89FE9EFF20">Table 1.</tableCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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