treatments-xml/data/03/B8/87/03B887E8FFB3A602FF14AAD560C7FDD9.xml

<document id="0C87897B8464BB19BE5CD553DBE62D72" ID-CLB-Dataset="20328" ID-DOI="10.11646/zootaxa.5105.4.1" ID-GBIF-Dataset="45708bdc-2d73-4b5d-91d9-75d9f31665a2" ID-ISSN="1175-5326" ID-Zenodo-Dep="6333771" ID-ZooBank="E81605D2-5B90-446A-B209-57EEFC8EA396" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646640338754" checkinUser="plazi" docAuthor="Huang, Hao &amp; Song, Kui" docDate="2022" docId="03B887E8FFB3A602FF14AAD560C7FDD9" docLanguage="en" docName="zootaxa.5105.4.1.pdf" docOrigin="Zootaxa 5105 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Colias thrasibulus subsp. angustomarginata Huang &amp; Song 2022, ssp. n." docType="treatment" docVersion="7" lastPageNumber="463" masterDocId="FF81FF90FFB8A60EFF83A826621BFFC0" masterDocTitle="A review of the genus Colias Fabricius, 1807 (Lepidoptera: Pieridae) from Qinghai with descriptions of a new subspecies of Colias thrasibulus Fruhstorfer, 1910" masterLastPageNumber="489" masterPageNumber="451" pageNumber="462" updateTime="1699317299447" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6F572056D5434E6882909E6358ED344B">A review of the genus Colias Fabricius, 1807 (Lepidoptera: Pieridae) from Qinghai with descriptions of a new subspecies of Colias thrasibulus Fruhstorfer, 1910</mods:title>
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<mods:namePart id="D141FE3972FDC06729063F1ED69127F9">Huang, Hao</mods:namePart>
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<mods:namePart id="934F0B204915140A0A12B6FC1F50CED5">Song, Kui</mods:namePart>
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<mods:affiliation id="34D9BD88708F4EC51D74C61D7B849B14">School of Economics and Management, Qinghai Nationalities University, Bayi Road No. 3, Xining, 810007, Qinghai, P. R. China. 76342870 @ qq. com; https: // orcid. org / 0000 - 0002 - 6928 - 7869</mods:affiliation>
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<taxonomicName id="4C114D7DFFB3A605FF14AAD56042FCCD" authority="Huang &amp; Song, 2022" authorityName="Huang &amp; Song" authorityYear="2022" box="[151,601,755,782]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="subSpecies" species="thrasibulus" status="subsp. nov." subSpecies="angustomarginata">
<emphasis id="B965EAECFFB3A605FF14AAD56042FCCD" bold="true" box="[151,601,755,782]" italics="true" pageId="11" pageNumber="462">Colias thrasibulus angustomarginata</emphasis>
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<taxonomicNameLabel id="A2565797FFB3A605FDE3AAD260B5FCCE" box="[608,686,756,782]" pageId="11" pageNumber="462" rank="subSpecies">ssp. n.</taxonomicNameLabel>
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<paragraph id="8BAE36FEFFB3A605FF14AB316065FCF2" blockId="11.[151,686,755,818]" box="[151,638,791,818]" pageId="11" pageNumber="462">
(
<figureCitation id="132A2A7BFFB3A605FF1CAB3162E6FCF2" box="[159,253,791,818]" captionStart="FIGURES 21–23" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[223,1364,181,1920]" captionTargetId="figure-21@20.[223,1364,181,1920]" captionTargetPageId="20" captionText="FIGURES 21–23. Habitus of Colias taxa ♂♂ &amp; ♀♀ at same scale (Upper side only). 21—Colias thrasibulus thrasibulus; 22—Colias thrasibulus fasciata; 23—Colias thrasibulus angustomarginata. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.6333789" httpUri="https://zenodo.org/record/6333789/files/figure.png" pageId="11" pageNumber="462">Figs. 23</figureCitation>
, 
<figureCitation id="132A2A7BFFB3A605FE8AAB31633CFCF1" box="[265,295,791,817]" captionStart="FIGURES 24–26" captionStartId="21.[151,264,1943,1967]" captionTargetBox="[223,1364,181,1920]" captionTargetId="figure-19@21.[223,1364,181,1920]" captionTargetPageId="21" captionText="FIGURES 24–26. Habitus of Colias taxa ♂♂ &amp; ♀♀ at same scale (Underside only). 24—Colias thrasibulus thrasibulus; 25—Colias thrasibulus fasciata; 26—Colias thrasibulus angustomarginata. Corresponding to figs. 21-23. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.6333791" httpUri="https://zenodo.org/record/6333791/files/figure.png" pageId="11" pageNumber="462">26</figureCitation>
, 
<figureCitation id="132A2A7BFFB3A605FEB0AB3E6360FCF2" box="[307,379,792,818]" captionStart-0="FIGURE 42" captionStart-1="FIGURE 43" captionStartId-0="24.[151,250,1943,1967]" captionStartId-1="25.[151,250,1943,1967]" captionTargetBox-0="[216,1372,181,1920]" captionTargetBox-1="[218,1370,181,1920]" captionTargetId-0="figure-21@24.[216,1372,181,1920]" captionTargetId-1="figure-19@25.[218,1370,181,1920]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="FIGURE 42. ♂ genitalia and 8th tergum (with superuncus) in lateral view at same scale. Scale bar = 1 mm." captionText-1="FIGURE 43. 8th tergum in dorsal view at same scale to show superuncus. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6333797" figureDoi-1="http://doi.org/10.5281/zenodo.6333799" httpUri-0="https://zenodo.org/record/6333797/files/figure.png" httpUri-1="https://zenodo.org/record/6333799/files/figure.png" pageId="11" pageNumber="462">42–43</figureCitation>
, 
<figureCitation id="132A2A7BFFB3A605FE04AB3163D4FCF2" box="[391,463,791,818]" captionStart-0="FIGURE 45" captionStart-1="FIGURE 46" captionStartId-0="27.[151,250,1943,1967]" captionStartId-1="28.[151,250,1943,1967]" captionTargetBox-0="[216,1372,181,1920]" captionTargetBox-1="[216,1372,181,1909]" captionTargetId-0="figure-19@27.[216,1372,181,1920]" captionTargetId-1="figure-21@28.[216,1372,181,1920]" captionTargetPageId-0="27" captionTargetPageId-1="28" captionText-0="FIGURE 45. ♂ genitalia in dorsal view at same scale. Scale bar = 1 mm." captionText-1="FIGURE 46. Ring in lateral view and scaphium in ventral view at same scale. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6333807" figureDoi-1="http://doi.org/10.5281/zenodo.6333809" httpUri-0="https://zenodo.org/record/6333807/files/figure.png" httpUri-1="https://zenodo.org/record/6333809/files/figure.png" pageId="11" pageNumber="462">45–46</figureCitation>
, 
<figureCitation id="132A2A7BFFB3A605FE58AB3163E3FCF2" box="[475,504,791,818]" captionStart="FIGURE 48" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[216,1372,181,1920]" captionTargetId="figure-21@30.[216,1372,181,1920]" captionTargetPageId="30" captionText="FIGURE 48. Phallus in lateral view and coecum-process in full-face view at same scale. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6333813" httpUri="https://zenodo.org/record/6333813/files/figure.png" pageId="11" pageNumber="462">48</figureCitation>
–50, 58, 67)
</paragraph>
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<emphasis id="B965EAECFFB3A605FF14AB46635CFCB9" bold="true" box="[151,327,863,890]" pageId="11" pageNumber="462">Type material.</emphasis>
<materialsCitation id="3B793CA3FFB3A605FED3AB4662FEFC02" ID-GBIF-Occurrence="3698883356" collectingDate="2021-07-28" collectorName="K. Song" country="China" county="Dulan County" elevation="4050" location="Tibetan Autonomous Prefecture" municipality="Haixi Mongol" pageId="11" pageNumber="462" specimenCount="1" specimenCount-male="1" stateProvince="Qinghai" typeStatus="holotype">
<emphasis id="B965EAECFFB3A605FED3AB4663A6FCBA" bold="true" box="[336,445,864,890]" pageId="11" pageNumber="462">
<typeStatus id="54AA885CFFB3A605FED3AB4663A6FCBA" box="[336,445,864,890]" pageId="11" pageNumber="462" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="132A2A7BFFB3A605FE4DAB79604BFCBA" box="[462,592,863,890]" captionStart="FIGURES 21–23" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[223,1364,181,1920]" captionTargetId="figure-21@20.[223,1364,181,1920]" captionTargetPageId="20" captionText="FIGURES 21–23. Habitus of Colias taxa ♂♂ &amp; ♀♀ at same scale (Upper side only). 21—Colias thrasibulus thrasibulus; 22—Colias thrasibulus fasciata; 23—Colias thrasibulus angustomarginata. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.6333789" httpUri="https://zenodo.org/record/6333789/files/figure.png" pageId="11" pageNumber="462">Fig. 23 - P</figureCitation>
<figureCitation id="132A2A7BFFB3A605FDD3AB466040FCBA" box="[592,603,864,890]" captionStart="FIGURES 1–3" captionStartId="13.[151,264,1943,1967]" captionTargetBox="[217,1370,181,1920]" captionTargetId="figure-19@13.[217,1370,181,1920]" captionTargetPageId="13" captionText="FIGURES 1–3. Habitus of Colias taxa ♂♂ &amp; ♀♀ at same scale (Upper half- upper side; lower half- underside). 1—Colias adelaidae adelaidae; 2—Colias arida cakana; 3—Colias wanda wanda. Serial number unique for each specimen. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.6333775" httpUri="https://zenodo.org/record/6333775/files/figure.png" pageId="11" pageNumber="462">1</figureCitation>
): 
<specimenCount id="9D17FD77FFB3A605FDF0AB466091FCBA" box="[627,650,864,890]" pageId="11" pageNumber="462" type="male">♂</specimenCount>
, 
<collectingCountry id="F306766EFFB3A605FD1BAB4660EEFCBA" box="[664,757,864,890]" name="China" pageId="11" pageNumber="462">CHINA</collectingCountry>
: 
<collectingRegion id="49D5F81CFFB3A605FC82AB7961D2FCB9" box="[769,969,863,889]" country="China" name="Qinghai" pageId="11" pageNumber="462">Qinghai province</collectingRegion>
, 
<collectingMunicipality id="6BCAAC84FFB3A605FC55AB79666DFCB9" box="[982,1142,863,889]" pageId="11" pageNumber="462">Haixi Mongol</collectingMunicipality>
and 
<location id="8ECE6025FFB3A605FB33AB796316FC5D" LSID="urn:lsid:plazi:treatment:03B887E8FFB3A602FF14AAD560C7FDD9:8ECE6025FFB3A605FB33AB796316FC5D" country="China" county="Dulan County" municipality="Haixi Mongol" name="Tibetan Autonomous Prefecture" pageId="11" pageNumber="462" stateProvince="Qinghai">Tibetan Autonomous Prefecture</location>
, 
<collectingCounty id="62CF4E72FFB3A605FE9FABA563A4FC5E" box="[284,447,899,926]" pageId="11" pageNumber="462">Dulan County</collectingCounty>
, ca 
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SW. of the county, 
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<elevation id="003CD1CDFFB3A605FCA8ABA261D9FC5E" box="[811,962,900,926]" metricMagnitude="3" metricUnit="m" metricValue="4.05" metricValueMax="4.1" metricValueMin="4.0" pageId="11" pageNumber="462" unit="m" value="4050.0" valueMax="4100.0" valueMin="4000.0">4000–4100m</elevation>
</quantity>
, 
<date id="FFAF103EFFB3A605FC53ABA26647FC5E" box="[976,1116,900,926]" pageId="11" pageNumber="462" value="2021-07-28">
<collectingDate id="EFEBE9D6FFB3A605FC53ABA26647FC5E" box="[976,1116,900,926]" pageId="11" pageNumber="462" value="2021-07-28">28.VII.2021</collectingDate>
</date>
, 
<collectorName id="26E45328FFB3A605FBE8ABA266D0FC5E" box="[1131,1227,900,926]" pageId="11" pageNumber="462">K. Song</collectorName>
leg., deposited in SNUC
</materialsCitation>
. 
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<emphasis id="B965EAECFFB3A605FF72AB8E6372FC02" bold="true" box="[241,361,936,962]" pageId="11" pageNumber="462">
<typeStatus id="54AA885CFFB3A605FF72AB8E6372FC02" box="[241,361,936,962]" pageId="11" pageNumber="462" type="paratype">Paratypes</typeStatus>
</emphasis>
: 
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, 
<specimenCount id="9D17FD77FFB3A605FE56AB8E600FFC02" box="[469,532,936,962]" pageId="11" pageNumber="462" type="female">3 ♀♀</specimenCount>
(
<specimenCount id="9D17FD77FFB3A605FDA6AB8E606EFC02" box="[549,629,936,962]" pageId="11" pageNumber="462" type="male">13 ♂♂</specimenCount>
, 
<specimenCount id="9D17FD77FFB3A605FD01AB8E60D9FC02" box="[642,706,936,962]" pageId="11" pageNumber="462" type="female">2 ♀♀</specimenCount>
in CSK, 
<specimenCount id="9D17FD77FFB3A605FCAFAB8E6175FC02" box="[812,878,936,962]" pageId="11" pageNumber="462" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="9D17FD77FFB3A605FCF8AB8E61BDFC02" box="[891,934,936,962]" pageId="11" pageNumber="462" type="female">1 ♀</specimenCount>
in 
<collectionCode id="ED00AE3BFFB3A605FC48AB8E6611FC02" box="[971,1034,936,962]" pageId="11" pageNumber="462">CHH</collectionCode>
), same locality as holotype, 14.VII. &amp; 
<date id="FFAF103EFFB3A605FF37ABEA636AFC26" box="[180,369,972,998]" pageId="11" pageNumber="462" value="2021-07-28" valueMax="2021-07-29" valueMin="2021-07-28">
<collectingDate id="EFEBE9D6FFB3A605FF37ABEA636AFC26" box="[180,369,972,998]" pageId="11" pageNumber="462" value="2021-07-28" valueMax="2021-07-29" valueMin="2021-07-28">28–29.VII. 2021</collectingDate>
</date>
, 
<collectorName id="26E45328FFB3A605FEFFABEA63C3FC26" box="[380,472,972,998]" pageId="11" pageNumber="462">K. Song</collectorName>
leg
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..
</paragraph>
<paragraph id="8BAE36FEFFB3A605FF44ABD660B9FB59" blockId="11.[151,1437,863,2042]" pageId="11" pageNumber="462">
<emphasis id="B965EAECFFB3A605FF44ABD66011FBC9" bold="true" box="[199,522,1007,1034]" pageId="11" pageNumber="462">
Male 
<typeStatus id="54AA885CFFB3A605FE8EABD6636FFBCA" box="[269,372,1008,1034]" pageId="11" pageNumber="462" type="holotype">holotype</typeStatus>
description.
</emphasis>
Length of forewing: 
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. Eye smooth. Antenna dark brown above and pale greyish brown below, with club mostly blackish brown, and with shaft obscurely segmented by pale scales. Body above black, clad with long yellowish setae, below densely clad with yellow scales. Frons and palpi clad with yellow scales, mixed with pale yellow and grayish setae. Legs clad with yellow scales, mixed with yellow setae. Both wings same shaped as in other two subspecies.
</paragraph>
<paragraph id="8BAE36FEFFB3A605FF44AC85635FFB22" blockId="11.[151,1437,863,2042]" pageId="11" pageNumber="462">Wing-pattern generally as in other two subspecies. Ciliae pale yellow on both wings both sides, mixed sparsely with pink ones.</paragraph>
<paragraph id="8BAE36FEFFB3A605FF44ACCD66F2FAC5" blockId="11.[151,1437,863,2042]" box="[199,1257,1259,1286]" pageId="11" pageNumber="462">
Male genitalia as in other two subspecies, not separable from most of the other 
<taxonomicName id="4C114D7DFFB3A605FBC3ACCD66F2FAC5" box="[1088,1257,1259,1285]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B965EAECFFB3A605FBC3ACCD6692FAC5" box="[1088,1161,1259,1285]" italics="true" pageId="11" pageNumber="462">Colias</emphasis>
species.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30B6575FFB3A605FF44AD2962E6FA79" pageId="11" pageNumber="462" type="description">
<paragraph id="8BAE36FEFFB3A605FF44AD2961A6FAB1" blockId="11.[151,1437,863,2042]" pageId="11" pageNumber="462">
<emphasis id="B965EAECFFB3A605FF44AD296031FAEA" bold="true" box="[199,554,1295,1322]" pageId="11" pageNumber="462">
Description of male 
<typeStatus id="54AA885CFFB3A605FE37AD366031FAEA" box="[436,554,1296,1322]" pageId="11" pageNumber="462" type="paratype">paratypes</typeStatus>
</emphasis>
. Length of forewing: 
<quantity id="4CE99B1BFFB3A605FC9FAD36618DFAE9" box="[796,918,1295,1322]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="11" pageNumber="462" unit="mm" value="24.5" valueMax="26.0" valueMin="23.0">23– 26mm</quantity>
, mostly 
<quantity id="4CE99B1BFFB3A605FC71AD366676FAEA" box="[1010,1133,1295,1322]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="11" pageNumber="462" unit="mm" value="24.5" valueMax="25.0" valueMin="24.0">24– 25mm</quantity>
. Ground color of wings upper side pale yellow, occasionally whitish (Specimen P2). Individual variations mainly found in marginal markings on forewing upper side and postdiscal fasciae on both wings upper side.
</paragraph>
<paragraph id="8BAE36FEFFB3A605FF44AD5D62E6FA79" blockId="11.[151,1437,863,2042]" pageId="11" pageNumber="462">
<emphasis id="B965EAECFFB3A605FF44AD5D6058FA56" bold="true" box="[199,579,1403,1430]" pageId="11" pageNumber="462">
Description of female 
<typeStatus id="54AA885CFFB3A605FE4EAD5A6058FA56" box="[461,579,1404,1430]" pageId="11" pageNumber="462" type="paratype">paratypes</typeStatus>
</emphasis>
. Length of forewing: 
<quantity id="4CE99B1BFFB3A605FCBAAD5A61AAFA55" box="[825,945,1403,1430]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="11" pageNumber="462" unit="mm" value="24.5" valueMax="26.0" valueMin="23.0">23- 26mm</quantity>
. Upper side ground color more whitish than in males.
</paragraph>
</subSubSection>
<subSubSection id="C30B6575FFB3A605FF44ADE2630CF9C1" pageId="11" pageNumber="462" type="etymology">
<paragraph id="8BAE36FEFFB3A605FF44ADE2630CF9C1" blockId="11.[151,1437,863,2042]" pageId="11" pageNumber="462">
<emphasis id="B965EAECFFB3A605FF44ADE26355FA1E" bold="true" box="[199,334,1476,1502]" pageId="11" pageNumber="462">Etymology.</emphasis>
The specific name refers to the narrower marginal markings on forewing upper side of the new subspecies.
</paragraph>
</subSubSection>
<subSubSection id="C30B6575FFB3A605FF44AE2D677EF919" pageId="11" pageNumber="462" type="diagnosis">
<paragraph id="8BAE36FEFFB3A605FF44AE2D677EF919" blockId="11.[151,1437,863,2042]" pageId="11" pageNumber="462">
<emphasis id="B965EAECFFB3A605FF44AE2D6321F9E5" bold="true" box="[199,314,1547,1573]" pageId="11" pageNumber="462">Diagnosis</emphasis>
. This new subspecies can be distinguished from 
<taxonomicName id="4C114D7DFFB3A605FCE6AE2A660DF9E5" authorityName="thrasibulus Fruhstorfer" authorityYear="1910" box="[869,1046,1547,1574]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="subSpecies" species="thrasibulus" subSpecies="thrasibulus">
ssp. 
<emphasis id="B965EAECFFB3A605FC1BAE2D660DF9E5" box="[920,1046,1547,1573]" italics="true" pageId="11" pageNumber="462">thrasibulus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C114D7DFFB3A605FBCDAE2A66C1F9E6" authorityName=": Kocman" authorityYear="2003" box="[1102,1242,1548,1574]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="subSpecies" species="thrasibulus" subSpecies="fasciata">
ssp. 
<emphasis id="B965EAECFFB3A605FB02AE2A66C1F9E6" box="[1153,1242,1548,1574]" italics="true" pageId="11" pageNumber="462">fasciata</emphasis>
</taxonomicName>
by the following combination of characters: 1) forewing upper side marginal markings, especially the one on vein 5, constantly narrower than in other subspecies; 2) size generally much larger than in 
<taxonomicName id="4C114D7DFFB3A605FC31AE726678F9AD" authorityName="thrasibulus Fruhstorfer" authorityYear="1910" box="[946,1123,1619,1646]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="subSpecies" species="thrasibulus" subSpecies="thrasibulus">
ssp. 
<emphasis id="B965EAECFFB3A605FC66AE756678F9AD" box="[997,1123,1619,1645]" italics="true" pageId="11" pageNumber="462">thrasibulus</emphasis>
</taxonomicName>
, slightly larger than in 
<taxonomicName id="4C114D7DFFB3A605FAF2AE7262EBF952" authorityName=": Kocman" authorityYear="2003" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="subSpecies" species="thrasibulus" subSpecies="fasciata">
ssp. 
<emphasis id="B965EAECFFB3A605FF14AE5E62EBF952" box="[151,240,1656,1682]" italics="true" pageId="11" pageNumber="462">fasciata</emphasis>
</taxonomicName>
; 3) forewing upper side postdiscal fasciae less developed than in other subspecies, duller and narrower, often more or less interrupted by veins; 4) hindwing upper side postdiscal fasciae usually obsolete (invariably well developed in 
<taxonomicName id="4C114D7DFFB3A605FEADAEE663A2F91A" authorityName=": Kocman" authorityYear="2003" box="[302,441,1728,1754]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="subSpecies" species="thrasibulus" subSpecies="fasciata">
ssp. 
<emphasis id="B965EAECFFB3A605FEE3AEE663A2F91A" box="[352,441,1728,1754]" italics="true" pageId="11" pageNumber="462">fasciata</emphasis>
</taxonomicName>
); 5) hindwing upper side discal area paler than in 
<taxonomicName id="4C114D7DFFB3A605FC68AEE6666DF91A" authorityName=": Kocman" authorityYear="2003" box="[1003,1142,1728,1754]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="subSpecies" species="thrasibulus" subSpecies="fasciata">
ssp. 
<emphasis id="B965EAECFFB3A605FB9EAEE6666DF91A" box="[1053,1142,1728,1754]" italics="true" pageId="11" pageNumber="462">fasciata</emphasis>
</taxonomicName>
, not so black-dusted.
</paragraph>
</subSubSection>
<subSubSection id="C30B6575FFB3A602FF44AEC260C7FDD9" lastPageId="12" lastPageNumber="463" pageId="11" pageNumber="462" type="discussion">
<paragraph id="8BAE36FEFFB3A602FF44AEC2605CFE49" blockId="11.[151,1437,863,2042]" lastBlockId="12.[151,1437,151,537]" lastPageId="12" lastPageNumber="463" pageId="11" pageNumber="462">
<emphasis id="B965EAECFFB3A605FF44AEC2632FF93E" bold="true" box="[199,308,1764,1790]" pageId="11" pageNumber="462">Remarks</emphasis>
. Only two subspecies of 
<taxonomicName id="4C114D7DFFB3A605FDCAAEC5610DF93D" authorityName="Fruhstorfer" authorityYear="1910" box="[585,790,1763,1789]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="thrasibulus">
<emphasis id="B965EAECFFB3A605FDCAAEC5610DF93D" box="[585,790,1763,1789]" italics="true" pageId="11" pageNumber="462">Colias thrasibulus</emphasis>
</taxonomicName>
were considered as valid in Grieshuber et al’s (2012) revision and 
<bibRefCitation id="EF804B0FFFB3A605FF7FAF2163C5F8E2" author="Grieshuber, J" box="[252,478,1799,1826]" pageId="11" pageNumber="462" refId="ref14730" refString="Grieshuber, J (2014) Guide to the butterflies of the Palearctic region. Pieridae Part II. Omnes Artes, Milano, 86 pp." type="book" year="2014">Grieshuber’s (2014)</bibRefCitation>
Guide book: 
<taxonomicName id="4C114D7DFFB3A605FDF4AF2E613CF8E1" authorityName="thrasibulus Fruhstorfer" authorityYear="1910" box="[631,807,1799,1826]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="subSpecies" species="thrasibulus" subSpecies="thrasibulus">
ssp. 
<emphasis id="B965EAECFFB3A605FD2AAF21613CF8E1" box="[681,807,1799,1825]" italics="true" pageId="11" pageNumber="462">thrasibulus</emphasis>
</taxonomicName>
known from Ladakh Range (Kardung La and Chonging Valley, N. or NW. of Kardung La- TL of 
<taxonomicName id="4C114D7DFFB3A605FDE1AF0D60FBF885" authorityName="Fruhstorfer" authorityYear="1910" box="[610,736,1835,1861]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="thrasibulus">
<emphasis id="B965EAECFFB3A605FDE1AF0D60FBF885" box="[610,736,1835,1861]" italics="true" pageId="11" pageNumber="462">thrasibulus</emphasis>
</taxonomicName>
), Zanskar Range (Taglang La - TL of 
<emphasis id="B965EAECFFB3A605FB0DAF0D6707F885" box="[1166,1308,1835,1861]" italics="true" pageId="11" pageNumber="462">zanskarensis</emphasis>
; Nimaling; vicinity of Morari Lake), E. Karakorum Range (Sasser La, Shimshin Jilga - Cretschmar, [1935]), W. 
<collectingRegion id="49D5F81CFFB3A605FA80AF696726F8A9" box="[1283,1341,1871,1897]" country="China" name="Xizang" pageId="11" pageNumber="462">Tibet</collectingRegion>
(Zhabu, Rebang, Ritu - 
<bibRefCitation id="EF804B0FFFB3A605FEC4AF5263A3F84E" author="Lee, C. - L." box="[327,440,1908,1934]" pageId="11" pageNumber="462" refId="ref15770" refString="Lee, C. - L. (1982) Lepidoptera: Rhopalocera. In: Huang, F. - S. (Ed.), Insects of Xizang, Vol. 2. Science Press, Beijing, pp. 127 - 154." type="book" year="1982">Lee, 1982</bibRefCitation>
&amp; 
<bibRefCitation id="EF804B0FFFB3A605FE5CAF526049F84E" author="Wu, C. - S." box="[479,594,1908,1934]" pageId="11" pageNumber="462" refId="ref16533" refString="Wu, C. - S. (2010) Fauna Sinica. Insecta Vol. 52. Lepidoptera Pieridae. Science Press, Beijing, 416 pp., 16 pls. [in Chinese]" type="book" year="2010">Wu, 2010</bibRefCitation>
; Yanhu, Gejie - 
<bibRefCitation id="EF804B0FFFB3A605FC88AF5561A8F84E" author="Verhulst, J." box="[779,947,1907,1934]" pageId="11" pageNumber="462" refId="ref16453" refString="Verhulst, J. (2001) Les Colias du Globe. Monograph of the genus Colias. Plates. Goecke &amp; Evers, Kelterns, 45 pp., 183 pls. [in French]" type="book" year="2001">Verhulst, 2001</bibRefCitation>
; W. of Paryang - 
<bibRefCitation id="EF804B0FFFB3A605FBFEAF556759F84E" author="Grieshuber, J" box="[1149,1346,1907,1934]" pageId="11" pageNumber="462" refId="ref14730" refString="Grieshuber, J (2014) Guide to the butterflies of the Palearctic region. Pieridae Part II. Omnes Artes, Milano, 86 pp." type="book" year="2014">Grieshuber, 2014</bibRefCitation>
) and C. 
<collectingRegion id="49D5F81CFFB3A605FF14AFB162CAF871" box="[151,209,1943,1969]" country="China" name="Xizang" pageId="11" pageNumber="462">Tibet</collectingRegion>
(S. of Namco Lake - 
<bibRefCitation id="EF804B0FFFB3A605FE49AFB16077F872" author="Hoshiai, A." box="[458,620,1943,1970]" pageId="11" pageNumber="462" pagination="8 - 10" refId="ref15351" refString="Hoshiai, A. (1996) Notes of Colias from China and surrounding area (1). In: Koiwaya, S. (Ed.), Studies of Chinese Butterflies. Vol. 3. S. Koiwaya, Tokyo, pp. 211 - 217, pls. 8 - 10. [in Japanese]" type="book chapter" year="1996">Hoshiai, 1996</bibRefCitation>
); and 
<taxonomicName id="4C114D7DFFB3A605FD32AFBE6126F872" authorityName=": Kocman" authorityYear="2003" box="[689,829,1944,1970]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="subSpecies" species="thrasibulus" subSpecies="fasciata">
ssp. 
<emphasis id="B965EAECFFB3A605FD67AFBE6126F872" box="[740,829,1944,1970]" italics="true" pageId="11" pageNumber="462">fasciata</emphasis>
</taxonomicName>
known only from Yushu, SE. 
<collectingRegion id="49D5F81CFFB3A605FB17AFB166E9F871" box="[1172,1266,1943,1969]" country="China" name="Qinghai" pageId="11" pageNumber="462">Qinghai</collectingRegion>
. 
<taxonomicName id="4C114D7DFFB3A605FB7EAFBE633EF815" authorityName="Hoshiai" authorityYear="1996" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="subSpecies" species="thrasibulus" subSpecies="zanskarensis">
<emphasis id="B965EAECFFB3A605FB7EAFBE633EF815" italics="true" pageId="11" pageNumber="462">C. thrasibulus zanskarensis</emphasis>
</taxonomicName>
was retained by 
<bibRefCitation id="EF804B0FFFB3A605FE64AF9D608CF816" author="Verhulst, J." box="[487,663,1979,2006]" pageId="11" pageNumber="462" refId="ref16453" refString="Verhulst, J. (2001) Les Colias du Globe. Monograph of the genus Colias. Plates. Goecke &amp; Evers, Kelterns, 45 pp., 183 pls. [in French]" type="book" year="2001">Verhulst (2001)</bibRefCitation>
as good subspecies for the populations from Zanskar Range and W. 
<collectingRegion id="49D5F81CFFB3A605FF14AFF962CFF839" box="[151,212,2015,2041]" country="China" name="Xizang" pageId="11" pageNumber="462">Tibet</collectingRegion>
, but was regarded by 
<bibRefCitation id="EF804B0FFFB3A605FE57AFF960F2F83A" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[468,745,2015,2042]" pageId="11" pageNumber="462" refId="ref15180" refString="Grieshuber, J., Worthy, R. &amp; Lamas, G. (2012) The genus Colias Fabricius, 1807. Jan Haugum's annotated catalogue of the Old World Colias. Tshikolovets Publications, Pardubice, 438 pp." type="book" year="2012">
Grieshuber 
<emphasis id="B965EAECFFB3A605FDD9AFC6608EF839" box="[602,661,2015,2041]" italics="true" pageId="11" pageNumber="462">et al.</emphasis>
(2012)
</bibRefCitation>
as synonym of 
<taxonomicName id="4C114D7DFFB3A605FC20AFC6664EF839" authorityName="thrasibulus Fruhstorfer" authorityYear="1910" box="[931,1109,2015,2042]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="subSpecies" species="thrasibulus" subSpecies="thrasibulus">
ssp. 
<emphasis id="B965EAECFFB3A605FC54AFF9664EF839" box="[983,1109,2015,2041]" italics="true" pageId="11" pageNumber="462">thrasibulus</emphasis>
</taxonomicName>
as “it is within the range of variation of nominotypical 
<taxonomicName id="4C114D7DFFB4A602FE4DA8B16074FF71" authorityName="Fruhstorfer" authorityYear="1910" box="[462,623,151,177]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="thrasibulus">
<emphasis id="B965EAECFFB4A602FE4DA8B16074FF71" box="[462,623,151,177]" italics="true" pageId="12" pageNumber="463">C. thrasibulus</emphasis>
</taxonomicName>
”. It is difficult to sort the population from S. of Namco, C. 
<collectingRegion id="49D5F81CFFB4A602FAA1A8B16744FF71" box="[1314,1375,151,177]" country="China" name="Xizang" pageId="12" pageNumber="463">Tibet</collectingRegion>
, as it is intermediate between 
<taxonomicName id="4C114D7DFFB4A602FE32A89D607FFF15" authorityName="thrasibulus Fruhstorfer" authorityYear="1910" box="[433,612,187,213]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subSpecies" species="thrasibulus" subSpecies="thrasibulus">
ssp. 
<emphasis id="B965EAECFFB4A602FE65A89D607FFF15" box="[486,612,187,213]" italics="true" pageId="12" pageNumber="463">thrasibulus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C114D7DFFB4A602FD22A89D6134FF15" authorityName=": Kocman" authorityYear="2003" box="[673,815,187,213]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subSpecies" species="thrasibulus" subSpecies="fasciata">
ssp. 
<emphasis id="B965EAECFFB4A602FD55A89D6134FF15" box="[726,815,187,213]" italics="true" pageId="12" pageNumber="463">fasciata</emphasis>
</taxonomicName>
in distribution and wing-characters. 
<taxonomicName id="4C114D7DFFB4A602FB59A89D6776FF15" authorityName=": Kocman" authorityYear="2003" box="[1242,1389,187,213]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subSpecies" species="thrasibulus" subSpecies="fasciata">
Ssp. 
<emphasis id="B965EAECFFB4A602FA97A89D6776FF15" box="[1300,1389,187,213]" italics="true" pageId="12" pageNumber="463">fasciata</emphasis>
</taxonomicName>
has only slight difference from 
<taxonomicName id="4C114D7DFFB4A602FE53A8F9609AFF39" authorityName="thrasibulus Fruhstorfer" authorityYear="1910" box="[464,641,223,249]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subSpecies" species="thrasibulus" subSpecies="thrasibulus">
ssp. 
<emphasis id="B965EAECFFB4A602FD80A8F9609AFF39" box="[515,641,223,249]" italics="true" pageId="12" pageNumber="463">thrasibulus</emphasis>
</taxonomicName>
in the darker discal area on hindwing upper side, however, the newly collected specimens of 
<taxonomicName id="4C114D7DFFB4A602FE1CA9256030FEDD" authorityName=": Kocman" authorityYear="2003" box="[415,555,259,285]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subSpecies" species="thrasibulus" subSpecies="fasciata">
ssp. 
<emphasis id="B965EAECFFB4A602FE51A9256030FEDD" box="[466,555,259,285]" italics="true" pageId="12" pageNumber="463">fasciata</emphasis>
</taxonomicName>
from Yushu confirm that this slight difference is rather constant. It is possible that 
<taxonomicName id="4C114D7DFFB4A602FF4BA9016348FE81" authorityName=": Kocman" authorityYear="2003" box="[200,339,295,321]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subSpecies" species="thrasibulus" subSpecies="fasciata">
ssp. 
<emphasis id="B965EAECFFB4A602FF79A9016348FE81" box="[250,339,295,321]" italics="true" pageId="12" pageNumber="463">fasciata</emphasis>
</taxonomicName>
is only the end of a cline of nominotypical 
<taxonomicName id="4C114D7DFFB4A602FCB8A90161C2FE81" authorityName="Fruhstorfer" authorityYear="1910" box="[827,985,295,321]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="thrasibulus">
<emphasis id="B965EAECFFB4A602FCB8A90161C2FE81" box="[827,985,295,321]" italics="true" pageId="12" pageNumber="463">C. thrasibulus</emphasis>
</taxonomicName>
. On the other hand, the new subspecies, 
<emphasis id="B965EAECFFB4A602FF14A96D6378FEA4" box="[151,355,331,356]" italics="true" pageId="12" pageNumber="463">angustomarginata</emphasis>
appears to produce a entirely different phenotype with large size and obsolete black markings and suffusion on the wings upper side.
</paragraph>
<paragraph id="8BAE36FEFFB4A602FF44A9B560C7FDD9" blockId="12.[151,1437,151,537]" pageId="12" pageNumber="463">
The zoogeographical pattern of 
<taxonomicName id="4C114D7DFFB4A602FDB9A9B56110FE6D" authorityName="Fruhstorfer" authorityYear="1910" box="[570,779,403,429]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="thrasibulus">
<emphasis id="B965EAECFFB4A602FDB9A9B56110FE6D" box="[570,779,403,429]" italics="true" pageId="12" pageNumber="463">Colias thrasibulus</emphasis>
</taxonomicName>
recalls that of 
<taxonomicName id="4C114D7DFFB4A602FC3DA9B5675CFE6D" authority="(Esper, 1783)" baseAuthorityName="Esper" baseAuthorityYear="1783" box="[958,1351,403,429]" class="Insecta" family="Nymphalidae" genus="Hipparchia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="autonoe">
<emphasis id="B965EAECFFB4A602FC3DA9B566B9FE6C" box="[958,1186,403,429]" italics="true" pageId="12" pageNumber="463">Hipparchia autonoe</emphasis>
(Esper, 1783)
</taxonomicName>
(
<taxonomicName id="4C114D7DFFB4A602FADAA9B562E1FE11" class="Insecta" family="Nymphalidae" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="family">Nymphalidae</taxonomicName>
: 
<taxonomicName id="4C114D7DFFB4A602FE85A991637BFE12" authorityName="Boisduval" authorityYear="1833" box="[262,352,439,466]" class="Insecta" family="Nymphalidae" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="tribe" tribe="Satyrini">Satyrini</taxonomicName>
) in Tibetan plateau: 
<taxonomicName id="4C114D7DFFB4A602FDC8A99160D2FE10" baseAuthorityName="Esper" baseAuthorityYear="1783" box="[587,713,439,464]" class="Insecta" family="Nymphalidae" genus="Hipparchia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="autonoe">
<emphasis id="B965EAECFFB4A602FDC8A99160D2FE10" box="[587,713,439,464]" italics="true" pageId="12" pageNumber="463">H. autonoe</emphasis>
</taxonomicName>
changes a little from W. 
<collectingRegion id="49D5F81CFFB4A602FC6BA9916639FE11" box="[1000,1058,439,465]" country="China" name="Xizang" pageId="12" pageNumber="463">Tibet</collectingRegion>
(
<taxonomicName id="4C114D7DFFB4A602FBB0A991678FFE11" authority="Markhasiov, 2021" authorityName="Markhasiov" authorityYear="2021" box="[1075,1428,439,465]" pageId="12" pageNumber="463" rank="subSpecies" subSpecies="mabcha">
ssp. 
<emphasis id="B965EAECFFB4A602FBE4A99166DBFE11" box="[1127,1216,439,465]" italics="true" pageId="12" pageNumber="463">mabcha</emphasis>
Markhasiov, 2021
</taxonomicName>
) to Yushu, SE. 
<collectingRegion id="49D5F81CFFB4A602FEB5A9FD638BFE35" box="[310,400,475,501]" country="China" name="Qinghai" pageId="12" pageNumber="463">Qinghai</collectingRegion>
(
<taxonomicName id="4C114D7DFFB4A602FE22A9FD60ADFE35" authority="Lang, 2019" authorityName="Lang" authorityYear="2019" box="[417,694,475,501]" pageId="12" pageNumber="463" rank="subSpecies" subSpecies="songkui">
ssp. 
<emphasis id="B965EAECFFB4A602FE50A9FD6031FE35" box="[467,554,475,501]" italics="true" pageId="12" pageNumber="463">songkui</emphasis>
Lang, 2019
</taxonomicName>
), but turns to be much brighter at Dulan-Balong areas, C. 
<collectingRegion id="49D5F81CFFB4A602FAC1A9FD6787FE35" box="[1346,1436,475,501]" country="China" name="Qinghai" pageId="12" pageNumber="463">Qinghai</collectingRegion>
(
<taxonomicName id="4C114D7DFFB4A602FF23A9D960CAFDD9" authority="Sbordoni, Bozano &amp; Cesaroni, 2018" authorityName="Sbordoni, Bozano &amp; Cesaroni" authorityYear="2018" box="[160,721,511,537]" pageId="12" pageNumber="463" rank="subSpecies" subSpecies="arcellae">
ssp. 
<emphasis id="B965EAECFFB4A602FF51A9D96336FDD9" box="[210,301,511,537]" italics="true" pageId="12" pageNumber="463">arcellae</emphasis>
Sbordoni, Bozano &amp; Cesaroni, 2018
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>