treatments-xml/data/03/90/E8/0390E827603CFF9D95D1F8A247F4F99C.xml

<document id="C1699C3BBCDA1E3FCCBEDA5CE7C59B15" ID-CLB-Dataset="22094" ID-DOI="10.5852/ejt.2015.162" ID-GBIF-Dataset="e7c55171-03d5-4714-a0c2-b7e70ab80fce" ID-ISSN="2118-9773" ID-Zenodo-Dep="3829511" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1504785881774" checkinUser="jeremy" docAuthor="Huber, Bernhard A., Nuñeza, Olga M. &amp; Ung, Charles Leh Moi" docDate="2015" docId="0390E827603CFF9D95D1F8A247F4F99C" docLanguage="en" docName="ejt-2015-162.pdf.imf" docOrigin="European Journal of Taxonomy 162" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Aetana Huber 2005" docType="treatment" docVersion="5" lastPageNumber="9" masterDocId="FFA9905F603FFF959719FF814549FFA4" masterDocTitle="Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae)" masterLastPageNumber="78" masterPageNumber="1" pageNumber="4" updateTime="1698468106960" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8A8CCD81F4219A02F051970BDDDCE256">Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana (Araneae, Pholcidae)</mods:title>
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<mods:namePart id="75365B2A17B4BC524938C735D2999A7A">Huber, Bernhard A.</mods:namePart>
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<mods:affiliation id="E51FCB3DDC84A69CFD2057C0C1C8D55D">Alexander Koenig Research Museum of Zoology, Adenauerallee 160, 53113 Bonn, Germany. &amp; urn: lsid: zoobank. org: author: 33607 F 65 - 19 BF- 4 DC 9 - 94 FD- 4 BB 88 CED 455 F</mods:affiliation>
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<mods:namePart id="1F5BE591B9A568A1F23A262FD7793A70">Nuñeza, Olga M.</mods:namePart>
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<mods:affiliation id="A61571C7AC385B68F59238495B8D9C03">Department of Biological Sciences, Mindanao State University - Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines. &amp; Email: olgamnuneza @ yahoo. com &amp; urn: lsid: zoobank. org: author: 6 D 14 FCCC-F 148 - 4293 - 9 F 23 - 0 EB 5 A 4119305</mods:affiliation>
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<mods:namePart id="F56D18FB32633CF38192C6272210D8AF">Ung, Charles Leh Moi</mods:namePart>
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<mods:affiliation id="111848F508E96338809B9055312CFB27">Sarawak Museum, Jalan Tun Abang Haji Openg, 93566 Kuching, Sarawak, Malaysia. &amp; Email: charllmu @ sarawak. gov. my &amp; urn: lsid: zoobank. org: author: 8738 F 336 - D 1 CC- 4 AD 3 - 97 E 4 - 82 BC 1729 A 123</mods:affiliation>
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<paragraph id="8B865931603CFF9695D1F8A246FEF89A" blockId="3.[712,951,1827,1854]" box="[712,951,1827,1854]" pageId="3" pageNumber="4">
<heading id="D0CEEE5D603CFF9695D1F8A246FEF89A" box="[712,951,1827,1854]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="4" reason="2">
<taxonomicName id="4C3922B2603CFF9695D1F8A246FEF89A" ID-CoL="PTX" ID-ENA="1966698" authority="Huber, 2005" authorityName="Huber" authorityYear="2005" box="[712,951,1827,1854]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B94D8523603CFF9695D1F8A24657F899" bold="true" box="[712,798,1827,1853]" italics="true" pageId="3" pageNumber="4">Aetana</emphasis>
Huber, 2005
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<treatmentCitation id="0A987F20603CFF9697FAF8E44770F8DB" author="Huber B. A." box="[227,569,1892,1919]" page="73" pageId="3" pageNumber="4" year="2005">
<taxonomicName id="4C3922B2603CFF9697FAF8E44770F8DB" ID-CoL="PTX" authority="Huber, 2005 a: 72 - 73" authorityName="Huber" authorityPageNumber="72 - 73" authorityYear="2005" box="[227,569,1892,1919]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B94D8523603CFF9697FAF8E4447FF8DB" box="[227,310,1893,1919]" italics="true" pageId="3" pageNumber="4">Aetana</emphasis>
<bibRefCitation id="EFA824C0603CFF969624F8E54770F8DB" author="Huber B. A." box="[317,569,1892,1919]" pageId="3" pageNumber="4" pagination="61 - 172" refId="ref27087" refString="Huber B. A. 2005 a. Revision of the genus Spermophora Hentz in Southeast Asia and on the Pacific Islands, with descriptions of three new genera (Araneae: Pholcidae). Zoologische Mededelingen 79 - 2 (4): 61 - 172." type="journal article" year="2005">Huber, 2005a: 72–73</bibRefCitation>
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.
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<typeStatus id="5482E793603CFF96955AF8E44734F8DB" box="[579,637,1893,1919]" pageId="3" pageNumber="4">Type</typeStatus>
species: 
<taxonomicName id="4C3922B2603CFF9695F1F8E4414BF8DB" ID-CoL="65BCT" authority="Huber, 2005" authorityName="Huber" authorityYear="2005" box="[744,1026,1892,1919]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="omayan">
<emphasis id="B94D8523603CFF9695F1F8E4462FF8DA" box="[744,870,1893,1919]" italics="true" pageId="3" pageNumber="4">A. omayan</emphasis>
Huber, 2005
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.
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<emphasis id="B94D8523603BFF9197FAFEB14415FEEE" bold="true" box="[227,348,304,330]" pageId="4" pageNumber="5">Diagnosis</emphasis>
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Even though the cladistic analysis identifies only two synapomorphies for 
<taxonomicName id="4C3922B2603BFF919378FEE141FDFEDE" authorityName="Huber" authorityYear="2005" box="[1121,1204,352,378]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B94D8523603BFF919378FEE141FDFEDE" box="[1121,1204,352,378]" italics="true" pageId="4" pageNumber="5">Aetana</emphasis>
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, the genus is fairly easily distinguished from the putatively closest relatives by the following characters: retrolateral trichobothrium on leg 1 very proximal (at &lt;5% of tibia length) and presence of curved hairs on tibiae and / or metatarsi (both in contrast to 
<taxonomicName id="4C3922B2603BFF91958EFE48467EFE47" authorityName="Hentz" authorityYear="1841" box="[663,823,457,483]" class="Arachnida" family="Pholcidae" genus="Spermophora" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B94D8523603BFF91958EFE48467EFE47" box="[663,823,457,483]" italics="true" pageId="4" pageNumber="5">Spermophora</emphasis>
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, 
<taxonomicName id="4C3922B2603BFF91945DFE484108FE40" authority="Huber, 2005" authorityName="Huber" authorityYear="2005" box="[836,1089,457,484]" class="Arachnida" family="Pholcidae" genus="Khorata" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B94D8523603BFF91945DFE4846EFFE47" box="[836,934,457,483]" italics="true" pageId="4" pageNumber="5">Khorata</emphasis>
Huber, 2005
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, 
<taxonomicName id="4C3922B2603BFF919352FE4841E4FE47" authorityName="Huber" authorityYear="2005" box="[1099,1197,457,483]" class="Arachnida" family="Pholcidae" genus="Savarna" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B94D8523603BFF919352FE4841E4FE47" box="[1099,1197,457,483]" italics="true" pageId="4" pageNumber="5">Savarna</emphasis>
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, and an undescribed genus from 
<collectingCountry id="F32E19A1603BFF91966BFE6D4493FDA2" box="[370,474,492,518]" name="Malaysia" pageId="4" pageNumber="5">Sarawak</collectingCountry>
, below called ‘Gen.n. Borneo’); sternum not dark, retrolateral trichobothrium on male palpal tibia in very distal position, and presence of epiandrous spigots (all in contrast to 
<taxonomicName id="4C3922B2603BFF91922AFD8E40DCFD8D" authorityName="Huber" authorityYear="2005" box="[1331,1429,527,553]" class="Arachnida" family="Pholcidae" genus="Khorata" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B94D8523603BFF91922AFD8E40DCFD8D" box="[1331,1429,527,553]" italics="true" pageId="4" pageNumber="5">Khorata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3922B2603BFF9197FAFDB2440CFDE9" authorityName="Huber" authorityYear="2005" box="[227,325,563,589]" class="Arachnida" family="Pholcidae" genus="Savarna" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B94D8523603BFF9197FAFDB2440CFDE9" box="[227,325,563,589]" italics="true" pageId="4" pageNumber="5">Savarna</emphasis>
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, and ‘Gen.n. Borneo’); male legs without spines and male palpal coxa unmodified (both in contrast to ‘Gen.n. Borneo’); ALS with only two spigots, epigynal plate without external pair of pockets, and female genitalia without unpaired posterior pocket (all in contrast to 
<taxonomicName id="4C3922B2603BFF919335FDF84185FD37" authorityName="Hentz" authorityYear="1841" box="[1068,1228,633,659]" class="Arachnida" family="Pholcidae" genus="Spermophora" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B94D8523603BFF919335FDF84185FD37" box="[1068,1228,633,659]" italics="true" pageId="4" pageNumber="5">Spermophora</emphasis>
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). Most characters previously thought to be diagnostic (
<bibRefCitation id="EFA824C0603BFF9195BAFD1D460BFD13" author="Huber B. A." box="[675,834,668,695]" pageId="4" pageNumber="5" pagination="61 - 172" refId="ref27087" refString="Huber B. A. 2005 a. Revision of the genus Spermophora Hentz in Southeast Asia and on the Pacific Islands, with descriptions of three new genera (Araneae: Pholcidae). Zoologische Mededelingen 79 - 2 (4): 61 - 172." type="journal article" year="2005">Huber 2005a</bibRefCitation>
) are rendered invalid due to the newly described species.
</paragraph>
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<paragraph id="8B865931603BFF9197FAFC81443DFCBE" blockId="4.[227,372,768,794]" box="[227,372,768,794]" pageId="4" pageNumber="5">
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<emphasis id="B94D8523603BFF9197FAFC81443DFCBE" bold="true" box="[227,372,768,794]" pageId="4" pageNumber="5">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B865931603BFF919609FCAE4419FCED" blockId="4.[227,1437,815,983]" box="[272,336,815,841]" pageId="4" pageNumber="5">
<emphasis id="B94D8523603BFF919609FCAE4419FCED" bold="true" box="[272,336,815,841]" pageId="4" pageNumber="5">Male</emphasis>
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<paragraph id="8B865931603BFF9197FAFCD246FFFC73" blockId="4.[227,1437,815,983]" pageId="4" pageNumber="5">
MEASUREMENTS. Total body length ~2.5–4.5 (smallest species in 
<taxonomicName id="4C3922B2603BFF9194FCFCD24126FCC8" authorityName="Huber" authorityYear="2005" box="[997,1135,850,877]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kinabalu">
<emphasis id="B94D8523603BFF9194FCFCD24126FCC8" box="[997,1135,850,877]" italics="true" pageId="4" pageNumber="5">A. kinabalu</emphasis>
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group; largest species in 
<taxonomicName id="4C3922B2603BFF9197FAFCF74428FC2B" authorityName="Huber" authorityYear="2005" box="[227,353,886,912]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="omayan">
<emphasis id="B94D8523603BFF9197FAFCF74428FC2B" box="[227,353,886,912]" italics="true" pageId="4" pageNumber="5">A. omayan</emphasis>
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group); carapace width 0.9–1.8; leg 1 length ~27–44; tibia 1 length ~6.0–11.0; tibia 2/tibia 4 length 0.92–1.08; tibia 1 L/d ~55–95 (the largest species, 
<taxonomicName id="4C3922B2603BFF9194BCFC18416AFC17" authorityName="Huber" authorityYear="2005" box="[933,1059,921,947]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="omayan">
<emphasis id="B94D8523603BFF9194BCFC18416AFC17" box="[933,1059,921,947]" italics="true" pageId="4" pageNumber="5">A. omayan</emphasis>
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, has the relatively thickest legs; the smallest species, 
<taxonomicName id="4C3922B2603BFF9196C0FC3D477BFC72" authorityName="Huber" authorityYear="2015" box="[473,562,956,982]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="gaya">
<emphasis id="B94D8523603BFF9196C0FC3D477BFC72" box="[473,562,956,982]" italics="true" pageId="4" pageNumber="5">A. gaya</emphasis>
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, has the relatively thinnest legs).
</paragraph>
<paragraph id="8B865931603BFF9197FAFC7A4474FB06" blockId="4.[227,1436,1019,1186]" pageId="4" pageNumber="5">
COLOR. In life mostly ochre-gray with brown and black marks (e.g., 
<figureCitation id="130245B4603BFF9194E5FC7A41CBFBB2" box="[1020,1154,1019,1046]" captionStart="Figs 51–56" captionStartId="21.[227,279,1966,1992]" captionTargetBox="[227,1437,665,1925]" captionTargetId="figure@21.[227,1437,665,1925]" captionTargetPageId="21" captionText="Figs 51–56. Live specimens. Aetana kiukoki group. 51–53. A. kiukoki Huber, sp. nov., ♂ from Baganihan (51), ♂ and ♀ from Santo Domingo (52–53), Mindanao. 54. A. paragua Huber, sp. nov., ♀ from Dinagat Island, Mindanao. 55–56. A. loboc Huber, sp. nov., ♂ and ♀ with eggsac from near Loboc, Bohol." figureDoi="http://doi.org/10.5281/zenodo.3829539" httpUri="https://zenodo.org/record/3829539/files/figure.png" pageId="4" pageNumber="5">Figs 51–56</figureCitation>
, 
<figureCitation id="130245B4603BFF919397FC7A41BEFBB2" box="[1166,1271,1019,1046]" captionStart="Figs 105–110" captionStartId="36.[227,278,1966,1992]" captionTargetBox="[228,1438,741,1921]" captionTargetId="figure@36.[228,1438,741,1921]" captionTargetPageId="36" captionText="Figs 105–110. Live specimens. Aetana kinabalu group. — 105–106. A. lambir Huber, sp. nov., ♂ and ♀ with eggsac from Lambir, Sarawak. — 107–108. A. poring Huber, sp. nov., ♂♂ from Mt. Kinabalu, Sabah. — 109– 110. A. indah Huber, sp. nov., adult and penultimate instar, ♂♂ from Crocker Range, Sabah." figureDoi="http://doi.org/10.5281/zenodo.3829557" httpUri="https://zenodo.org/record/3829557/files/figure.png" pageId="4" pageNumber="5">105–110</figureCitation>
), only 
<taxonomicName id="4C3922B2603BFF91925FFC7D4467FB9C" authority="Huber" authorityName="Huber" authorityYear="2015" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="libjo" status="sp. nov.">
<emphasis id="B94D8523603BFF91925FFC7D40D5FBB1" box="[1350,1436,1019,1046]" italics="true" pageId="4" pageNumber="5">A. libjo</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E3858603BFF919622FB9E44DFFB9D" box="[315,406,1055,1081]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C3922B2603BFF9196C8FB9E478DFB9C" authority="Huber" authorityName="Huber" authorityYear="2015" box="[465,708,1054,1081]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="baganihan" status="sp. nov.">
<emphasis id="B94D8523603BFF9196C8FB9E4739FB9C" box="[465,624,1054,1081]" italics="true" pageId="4" pageNumber="5">A. baganihan</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E3858603BFF9195C9FB9E4665FB9D" box="[720,812,1055,1081]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
with light brown to orange prosoma and palps (
<figureCitation id="130245B4603BFF919271FB9F446BFBF8" captionStart="Figs 6–12" captionStartId="9.[225,277,1966,1992]" captionTargetBox="[231,1441,413,1916]" captionTargetId="figure@9.[231,1441,413,1916]" captionTargetPageId="9" captionText="Figs 6–12. Live specimens. Aetana ocampoi group. 6–7. A. ocampoi Huber, sp. nov., ♂ from Mt. Isarog, Luzon. 8–9. A. libjo Huber, sp. nov., ♂ and ♀ from Dinagat Island, Mindanao. 10–12. A. baganihan Huber, sp. nov., ♂♂ from Baganighan, Mindanao." figureDoi="http://doi.org/10.5281/zenodo.3829521" httpUri="https://zenodo.org/record/3829521/files/figure.png" pageId="4" pageNumber="5">Figs 8–12</figureCitation>
); sternum never dark; legs usually with indistinct darker rings on femora (subdistally) and tibiae (proximally and subdistally); darker rings missing in 
<taxonomicName id="4C3922B2603BFF919440FBE4414DFBDB" authority="Huber" authorityName="Huber" authorityYear="2015" box="[857,1028,1125,1151]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="libjo" status="sp. nov.">
<emphasis id="B94D8523603BFF919440FBE446F8FBDB" box="[857,945,1125,1151]" italics="true" pageId="4" pageNumber="5">A. libjo</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E3858603BFF919309FBE44125FBDB" box="[1040,1132,1125,1151]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C3922B2603BFF9193BFFBE440D1FBDB" authority="Huber" authorityName="Huber" authorityYear="2015" box="[1190,1432,1125,1151]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="baganihan" status="sp. nov.">
<emphasis id="B94D8523603BFF9193BFFBE4400DFBDB" box="[1190,1348,1125,1151]" italics="true" pageId="4" pageNumber="5">A. baganihan</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E3858603BFF9197FAFB094474FB06" box="[227,317,1160,1186]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8B865931603BFF9197FAFB494765FA5F" blockId="4.[227,1437,1223,1531]" pageId="4" pageNumber="5">
BODY. Carapace either with shallow median furrow restricted to frontal part (
<figureCitation id="130245B4603BFF919341FB464196FB45" box="[1112,1247,1223,1249]" captionStart="Figs 62–69" captionStartId="23.[227,279,1896,1922]" captionTargetBox="[227,1437,421,1844]" captionTargetId="figure@23.[227,1437,421,1844]" captionTargetPageId="23" captionText="Figs 62–69. Aetana kiukoki Huber, sp. nov. 62–63. Male prosoma, oblique frontal and frontal views. 64. Female prosoma, frontal view. 65. Male clypeus and chelicerae, oblique frontal view. 66. Male clypeus modification, frontal view. 67. Epigynum, ventral view. 68. Male ALS and PMS. 69. Comb-hair on female tarsus 4. cl = clypeus; lca = lateral cheliceral apophysis. Scale lines: 62–64 = 400 µm; 65, 67 = 200 µm; 66 = 100 µm; 68–69 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3829543" httpUri="https://zenodo.org/record/3829543/files/figure.png" pageId="4" pageNumber="5">Figs 63–64</figureCitation>
, 
<figureCitation id="130245B4603BFF9193F0FB46405FFB45" box="[1257,1302,1223,1249]" captionStart="Figs 195–203. — 195–197" captionStartId="56.[227,279,1824,1850]" captionTargetBox="[242,1421,302,1783]" captionTargetId="figure@56.[242,1421,302,1783]" captionTargetPageId="56" captionText="Figs 195–203. — 195–197. Aetana abadae Huber, sp. nov. 195. Epigynum, ventral view. 196. Detail of preceding, showing opening of membranous pocket (arrow on Fig. 195). 197. Female ALS. — 198–203. A. omayan Huber, 2005. 198. Male prosoma, frontal view. 199. Left palp, retrolateral view. 200–201. Right procursus, prolatero-dorsal and prolateral views. 202. Detail of preceding. 203. Male palpal tarsal organ. b = genital bulb; e = embolus; f = femur; p = procursus; rp = retrolatero-ventral process; ti = tibia; tr = trochanter; vl = ventral lamina. Scale lines: 195, 199 = 300 µm; 196, 202 = 40 µm; 197, 203 = 10 µm; 198 = 500 µm; 200 = 200 µm; 201 = 100 µm" figureDoi="http://doi.org/10.5281/zenodo.3829584" httpUri="https://zenodo.org/record/3829584/files/figure.png" pageId="4" pageNumber="5">198</figureCitation>
) or without furrow (
<figureCitation id="130245B4603BFF91965CFB6B44E6FAA0" box="[325,431,1258,1284]" captionStart="Figs 118–125" captionStartId="39.[227,279,1896,1922]" captionTargetBox="[227,1437,374,1852]" captionTargetId="figure@39.[227,1437,374,1852]" captionTargetPageId="39" captionText="Figs 118–125. Aetana lambir Huber, sp. nov. 118–119. Male and female prosomata, frontal views. 120. Male palpal tarsal organ. 121. Left procursus, prolateral view. 122. Detail of preceding. 123. Right procursus (and femur apophysis), retrolateral view. 124. Female ALS. 125. Epigynum. f = femur; vl = ventral lamina. Scale lines: 118–119, 125 = 200 µm; 120 = 8 µm; 121 = 50 µm; 122 = 20 µm; 123 = 60 µm; 124 = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3829564" httpUri="https://zenodo.org/record/3829564/files/figure.png" pageId="4" pageNumber="5">Figs 118</figureCitation>
, 
<figureCitation id="130245B4603BFF9196A7FB6B44A7FAA0" box="[446,494,1258,1284]" captionStart="Figs 130–136" captionStartId="43.[225,277,1930,1956]" captionTargetBox="[227,1437,336,1884]" captionTargetId="figure@43.[227,1437,336,1884]" captionTargetPageId="43" captionText="Figs 130–136. Aetana gaya Huber, sp. nov. 130–131. Male prosoma, frontal and oblique frontal views. 132. Female prosoma, frontal view. 133. Detail of male metatarsus 1. 134. Male palpal tarsal organ. 135. Male carapace and ocular area. 136. Right procursus, retrolateral view. rp = retrolatero-ventral process; vl = ventral lamina. Scale lines: 130–132, 135 = 200 µm; 133 = 80 µm; 134 = 8 µm; 136 = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3829568" httpUri="https://zenodo.org/record/3829568/files/figure.png" pageId="4" pageNumber="5">130</figureCitation>
); ocular area raised, eye triads on short stalks directed toward lateral, in some species with median process (
<figureCitation id="130245B4603BFF91955CFA8C47EBFA8C" box="[581,674,1293,1320]" captionStart="Figs 75–79" captionStartId="28.[227,279,1966,1992]" captionTargetBox="[230,1434,356,1921]" captionTargetId="figure@28.[227,1437,353,1922]" captionTargetPageId="28" captionText="Figs 75–79. Aetana loboc Huber, sp. nov. 75–76. Left male palp, prolateral and retrolateral views. 77. Male prosoma, frontal view. 78–79. Cleared female genitalia, ventral and dorsal views. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829547" httpUri="https://zenodo.org/record/3829547/files/figure.png" pageId="4" pageNumber="5">Figs 77</figureCitation>
, 
<figureCitation id="130245B4603BFF9195B6FA8C4787FA8C" box="[687,718,1293,1320]" captionStart="Figs 96–98" captionStartId="33.[227,279,2001,2027]" captionTargetBox="[229,1429,577,1939]" captionTargetId="figure@33.[228,1438,564,1944]" captionTargetPageId="33" captionText="Figs 96–98. Aetana pasambai Huber, sp. nov. 96. Male prosoma, oblique frontal view. 97–98. Left male palp, prolateral and retrolateral views. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829553" httpUri="https://zenodo.org/record/3829553/files/figure.png" pageId="4" pageNumber="5">96</figureCitation>
) or with pair of processes arising from near ALE (
<figureCitation id="130245B4603BFF919235FA8C40D1FA8C" box="[1324,1432,1293,1320]" captionStart="Figs 219–223" captionStartId="60.[227,279,1932,1958]" captionTargetBox="[227,1437,378,1887]" captionTargetId="figure@60.[227,1437,378,1889]" captionTargetPageId="60" captionText="Figs 219–223. Aetana manansalai Huber, sp. nov. 219–220. Left male palp, prolateral and retrolateral views. 221. Male prosoma, oblique frontal view. 222–223. Cleared female genitalia, ventral and dorsal views. Arrow points at membranous pocket. rp = retrolatero-ventral process; vl = ventral lamina. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829590" httpUri="https://zenodo.org/record/3829590/files/figure.png" pageId="4" pageNumber="5">Figs 221</figureCitation>
, 
<figureCitation id="130245B4603BFF9197FAFAB0445DFAE8" box="[227,276,1329,1356]" captionStart="Figs 224–228" captionStartId="63.[227,279,1966,1992]" captionTargetBox="[229,1436,444,1924]" captionTargetId="figure@63.[227,1437,444,1924]" captionTargetPageId="63" captionText="Figs 224–228. Aetana lozadae Huber, sp. nov. 224–225. Left male palp, prolateral and retrolateral views. 226. Male prosoma, oblique frontal view. 227–228. Cleared female genitalia, ventral and dorsal views. rp = retrolatero-ventral process; vl = ventral lamina. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829592" httpUri="https://zenodo.org/record/3829592/files/figure.png" pageId="4" pageNumber="5">226</figureCitation>
). AME absent. Clypeus high, either unmodified, with small paired processes (
<figureCitation id="130245B4603BFF9193B3FAB04059FAEF" box="[1194,1296,1329,1355]" captionStart="Figs 189–194. — 189–193" captionStartId="55.[227,279,1894,1920]" captionTargetBox="[227,1436,450,1840]" captionTargetId="figure@55.[227,1437,450,1847]" captionTargetPageId="55" captionText="Figs 189–194. — 189–193. Aetana abadae Huber, sp. nov. 189–190. Left male palp, prolateral and retrolateral views. 191. Male prosoma, oblique frontal view. 192–193. Cleared female genitalia, ventral and dorsal views. — 194. A. omayan Huber, 2005, cleared female genitalia, dorsal view. Arrows point at membranous pockets. ip = internal sclerotized pocket; rp = retrolatero-ventral process; vl = ventral lamina. Scale lines: 0.5 mm (female genitalia at same scale)." figureDoi="http://doi.org/10.5281/zenodo.3829582" httpUri="https://zenodo.org/record/3829582/files/figure.png" pageId="4" pageNumber="5">Fig. 191</figureCitation>
), with large median process (
<figureCitation id="130245B4603BFF9196B3FAD5474EFACB" box="[426,519,1364,1391]" captionStart="Figs 57–61" captionStartId="22.[227,279,1966,1992]" captionTargetBox="[227,1432,469,1920]" captionTargetId="figure@22.[227,1437,469,1922]" captionTargetPageId="22" captionText="Figs 57–61. Aetana kiukoki Huber, sp. nov. 57–58. Left male palp, prolateral and retrolateral views. 59. Male prosoma, oblique frontal view. 60–61. Cleared female genitalia, ventral and dorsal views. b = genital bulb; e = embolus; p = procursus. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829541" httpUri="https://zenodo.org/record/3829541/files/figure.png" pageId="4" pageNumber="5">Figs 59</figureCitation>
, 
<figureCitation id="130245B4603BFF91950DFAD5477CFACA" box="[532,565,1364,1390]" captionStart="Figs 75–79" captionStartId="28.[227,279,1966,1992]" captionTargetBox="[230,1434,356,1921]" captionTargetId="figure@28.[227,1437,353,1922]" captionTargetPageId="28" captionText="Figs 75–79. Aetana loboc Huber, sp. nov. 75–76. Left male palp, prolateral and retrolateral views. 77. Male prosoma, frontal view. 78–79. Cleared female genitalia, ventral and dorsal views. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829547" httpUri="https://zenodo.org/record/3829547/files/figure.png" pageId="4" pageNumber="5">77</figureCitation>
), or with indistinct lateral ridges (
<figureCitation id="130245B4603BFF9194D0FAD5417CFACA" box="[969,1077,1364,1390]" captionStart="Figs 219–223" captionStartId="60.[227,279,1932,1958]" captionTargetBox="[227,1437,378,1887]" captionTargetId="figure@60.[227,1437,378,1889]" captionTargetPageId="60" captionText="Figs 219–223. Aetana manansalai Huber, sp. nov. 219–220. Left male palp, prolateral and retrolateral views. 221. Male prosoma, oblique frontal view. 222–223. Cleared female genitalia, ventral and dorsal views. Arrow points at membranous pocket. rp = retrolatero-ventral process; vl = ventral lamina. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829590" httpUri="https://zenodo.org/record/3829590/files/figure.png" pageId="4" pageNumber="5">Figs 221</figureCitation>
, 
<figureCitation id="130245B4603BFF91935BFAD5413AFACB" box="[1090,1139,1364,1391]" captionStart="Figs 224–228" captionStartId="63.[227,279,1966,1992]" captionTargetBox="[229,1436,444,1924]" captionTargetId="figure@63.[227,1437,444,1924]" captionTargetPageId="63" captionText="Figs 224–228. Aetana lozadae Huber, sp. nov. 224–225. Left male palp, prolateral and retrolateral views. 226. Male prosoma, oblique frontal view. 227–228. Cleared female genitalia, ventral and dorsal views. rp = retrolatero-ventral process; vl = ventral lamina. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829592" httpUri="https://zenodo.org/record/3829592/files/figure.png" pageId="4" pageNumber="5">226</figureCitation>
). Abdomen from slightly longer than high (
<figureCitation id="130245B4603BFF9196AAFAF64770FA36" box="[435,569,1399,1426]" captionStart="Figs 51–56" captionStartId="21.[227,279,1966,1992]" captionTargetBox="[227,1437,665,1925]" captionTargetId="figure@21.[227,1437,665,1925]" captionTargetPageId="21" captionText="Figs 51–56. Live specimens. Aetana kiukoki group. 51–53. A. kiukoki Huber, sp. nov., ♂ from Baganihan (51), ♂ and ♀ from Santo Domingo (52–53), Mindanao. 54. A. paragua Huber, sp. nov., ♀ from Dinagat Island, Mindanao. 55–56. A. loboc Huber, sp. nov., ♂ and ♀ with eggsac from near Loboc, Bohol." figureDoi="http://doi.org/10.5281/zenodo.3829539" httpUri="https://zenodo.org/record/3829539/files/figure.png" pageId="4" pageNumber="5">Figs 51–56</figureCitation>
) to almost cylindrical (
<figureCitation id="130245B4603BFF919453FAF646DFFA35" box="[842,918,1399,1425]" captionStart="Figs 6–12" captionStartId="9.[225,277,1966,1992]" captionTargetBox="[231,1441,413,1916]" captionTargetId="figure@9.[231,1441,413,1916]" captionTargetPageId="9" captionText="Figs 6–12. Live specimens. Aetana ocampoi group. 6–7. A. ocampoi Huber, sp. nov., ♂ from Mt. Isarog, Luzon. 8–9. A. libjo Huber, sp. nov., ♂ and ♀ from Dinagat Island, Mindanao. 10–12. A. baganihan Huber, sp. nov., ♂♂ from Baganighan, Mindanao." figureDoi="http://doi.org/10.5281/zenodo.3829521" httpUri="https://zenodo.org/record/3829521/files/figure.png" pageId="4" pageNumber="5">Figs 8</figureCitation>
, 
<figureCitation id="130245B4603BFF9194BAFAF6469BFA35" box="[931,978,1399,1425]" captionStart="Figs 99–104" captionStartId="35.[227,279,1930,1956]" captionTargetBox="[227,1437,731,1888]" captionTargetId="figure@35.[227,1437,731,1888]" captionTargetPageId="35" captionText="Figs 99–104. Live specimens. Aetana kinabalu group. — 99–103. A. kinabalu Huber, 2005. ♂ and ♀ with eggsac from Gunung Mulu, Sarawak (99, 100); ♀ with eggsac and ♂ from Crocker Range, Sabah (101, 102); and ♂ from Mt. Kinabalu, Sabah (103). — 104. A. gaya Huber, sp. nov., ♂ from Gaya Island, Sabah." figureDoi="http://doi.org/10.5281/zenodo.3829555" httpUri="https://zenodo.org/record/3829555/files/figure.png" pageId="4" pageNumber="5">102</figureCitation>
), pointed at spinnerets. Male gonopore with four epiandrous spigots in all species examined with SEM (e.g., 
<figureCitation id="130245B4603BFF919315FA1B412CFA10" box="[1036,1125,1434,1460]" captionStart="Figs 30–37" captionStartId="15.[227,279,1909,1935]" captionTargetBox="[227,1437,310,1867]" captionTargetId="figure@15.[227,1437,310,1867]" captionTargetPageId="15" captionText="Figs 30–37. Aetana libjo Huber, sp. nov. 30. Processes of right genital bulb, retrolateral view. 31. Detail of preceding. 32. Comb-hairs on female tarsus 4. 33. Processes of left genital bulb, prolateral view. 34. Detail of preceding. 35. Male palpal tarsal organ. 36. Female ALS. 37. Male gonopore. Scale lines: 30, 33 = 100 µm; 31, 37 = 40 µm; 32, 34 = 20 µm; 35–36 = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3829533" httpUri="https://zenodo.org/record/3829533/files/figure.png" pageId="4" pageNumber="5">Figs 37</figureCitation>
, 
<figureCitation id="130245B4603BFF919368FA1B41D6FA10" box="[1137,1183,1434,1460]" captionStart="Figs 137–144" captionStartId="44.[227,279,1930,1956]" captionTargetBox="[227,1437,404,1882]" captionTargetId="figure@44.[227,1437,404,1882]" captionTargetPageId="44" captionText="Figs 137–144. Aetana gaya Huber, sp. nov. 137. Left procursus, prolateral view. 138–140. Details of preceding. 141. Male spinnerets. 142. Male ALS. 143. Male gonopore. 144. Epigynum. b = genital bulb; f = femur; p = procursus. Scale lines: 137 = 100 µm; 138, 141 = 60 µm; 139–140 = 20 µm; 142 = 10 µm; 143 = 30 µm; 144 = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3829570" httpUri="https://zenodo.org/record/3829570/files/figure.png" pageId="4" pageNumber="5">143</figureCitation>
, 
<figureCitation id="130245B4603BFF9193B2FA1B4192FA11" box="[1195,1243,1434,1461]" captionStart="Figs 155–162" captionStartId="50.[227,279,1862,1888]" captionTargetBox="[227,1437,302,1837]" captionTargetId="figure@50.[227,1437,302,1837]" captionTargetPageId="50" captionText="Figs 155–162. Aetana indah Huber, sp. nov. 155–156. Left palp, prolatero-distal and retrolaterodistal views. 157. Left procursus, retrolatero-distal view. 158. Detail of left procursus (and trochanter apophysis), prolateral view. 159. Left embolus (arrow points at sperm duct opening). 160–161. Left male cheliceral apophyses, oblique frontal and frontal views. 162. Male gonopore. b = genital bulb; e = embolus; f = femur; p = procursus; ti = tibia; tr = trochanter. Scale lines: 155–156 = 200 µm; 157 = 100 µm; 158–160 = 60 µm; 161 = 80 µm; 162 = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.3829576" httpUri="https://zenodo.org/record/3829576/files/figure.png" pageId="4" pageNumber="5">162</figureCitation>
); each ALS with large widened spigot and pointed spigot, without further cylindrically shaped spigots (
<figureCitation id="130245B4603BFF9193EBFA3C4019FA73" box="[1266,1360,1469,1495]" captionStart="Figs 62–69" captionStartId="23.[227,279,1896,1922]" captionTargetBox="[227,1437,421,1844]" captionTargetId="figure@23.[227,1437,421,1844]" captionTargetPageId="23" captionText="Figs 62–69. Aetana kiukoki Huber, sp. nov. 62–63. Male prosoma, oblique frontal and frontal views. 64. Female prosoma, frontal view. 65. Male clypeus and chelicerae, oblique frontal view. 66. Male clypeus modification, frontal view. 67. Epigynum, ventral view. 68. Male ALS and PMS. 69. Comb-hair on female tarsus 4. cl = clypeus; lca = lateral cheliceral apophysis. Scale lines: 62–64 = 400 µm; 65, 67 = 200 µm; 66 = 100 µm; 68–69 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3829543" httpUri="https://zenodo.org/record/3829543/files/figure.png" pageId="4" pageNumber="5">Figs 68</figureCitation>
, 
<figureCitation id="130245B4603BFF919247FA3F40C6FA7C" box="[1374,1423,1470,1496]" captionStart="Figs 204–212" captionStartId="57.[227,279,1897,1923]" captionTargetBox="[227,1437,514,1857]" captionTargetId="figure@57.[227,1437,514,1857]" captionTargetPageId="57" captionText="Figs 204–212. A. omayan Huber, 2005. 204. Left procursus tip (ventral lamina), retrolateral view. 205. Right procursus tip (pointed sclerite and ventral lamina), prolateral view. 206. Male gonopore. 207. Detail of male tarsus 1. 208–209. Comb-hairs on male tarsus 4. 210–211. Male and female ALS. 212. Epigynum. Scale lines: 204 = 50 µm; 205 = 40 µm; 206, 208 = 30 µm; 207 = 60 µm; 209 = 8 µm; 210–211 = 20 µm; 212 = 400 µm." figureDoi="http://doi.org/10.5281/zenodo.3829586" httpUri="https://zenodo.org/record/3829586/files/figure.png" pageId="4" pageNumber="5">210</figureCitation>
); PMS with two spigots each.
</paragraph>
</subSubSection>
<subSubSection id="C3230ABA603BFF9197FAF9A14672F907" pageId="4" pageNumber="5" type="discussion">
<paragraph id="8B865931603BFF9197FAF9A14672F907" blockId="4.[227,1436,1568,1699]" pageId="4" pageNumber="5">
CHELICERAE. Very variable, distal apophyses ranging from short processes in frontal position (
<figureCitation id="130245B4603BFF919222F9A140DEF99F" box="[1339,1431,1568,1595]" captionStart="Figs 13–15" captionStartId="10.[227,279,1968,1994]" captionTargetBox="[234,1437,704,1917]" captionTargetId="figure@10.[227,1437,703,1921]" captionTargetPageId="10" captionText="Figs 13–15. Aetana ocampoi Huber, sp. nov. 13–14. Left male palp, prolateral and retrolateral views (asterisk: retrolateral process of bulb). 15. Male chelicerae, frontal view. b = genital bulb; p = procursus. Scale lines: 13–14 = 0.5 mm; 15 = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3829525" httpUri="https://zenodo.org/record/3829525/files/figure.png" pageId="4" pageNumber="5">Figs 15</figureCitation>
, 
<figureCitation id="130245B4603BFF9197FAF9C2444BF9F9" box="[227,258,1603,1629]" captionStart="Figs 38–44. — 38–40" captionStartId="16.[227,279,1906,1932]" captionTargetBox="[227,1432,599,1858]" captionTargetId="figure@16.[227,1437,593,1864]" captionTargetPageId="16" captionText="Figs 38–44. — 38–40. Aetana libjo Huber, sp. nov. 38–39. Left genital bulb in prolateral and retrolateral views. 40. Male chelicerae, frontal view. — 41–44. Aetana baganihan Huber, sp. nov. 41–42. Left genital bulb in prolateral and retrolateral views. 43–44. Cleared female genitalia, ventral and dorsal views (asterisk: median membranous structure). Scale lines: 38–39, 41–44 = 0.5 mm; 40 = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3829535" httpUri="https://zenodo.org/record/3829535/files/figure.png" pageId="4" pageNumber="5">40</figureCitation>
) to long processes in lateral position (
<figureCitation id="130245B4603BFF9195A2F9C2466DF9F9" box="[699,804,1603,1629]" captionStart="Figs 189–194. — 189–193" captionStartId="55.[227,279,1894,1920]" captionTargetBox="[227,1436,450,1840]" captionTargetId="figure@55.[227,1437,450,1847]" captionTargetPageId="55" captionText="Figs 189–194. — 189–193. Aetana abadae Huber, sp. nov. 189–190. Left male palp, prolateral and retrolateral views. 191. Male prosoma, oblique frontal view. 192–193. Cleared female genitalia, ventral and dorsal views. — 194. A. omayan Huber, 2005, cleared female genitalia, dorsal view. Arrows point at membranous pockets. ip = internal sclerotized pocket; rp = retrolatero-ventral process; vl = ventral lamina. Scale lines: 0.5 mm (female genitalia at same scale)." figureDoi="http://doi.org/10.5281/zenodo.3829582" httpUri="https://zenodo.org/record/3829582/files/figure.png" pageId="4" pageNumber="5">Figs 191</figureCitation>
, 
<figureCitation id="130245B4603BFF919429F9C24629F9FA" box="[816,864,1603,1630]" captionStart="Figs 224–228" captionStartId="63.[227,279,1966,1992]" captionTargetBox="[229,1436,444,1924]" captionTargetId="figure@63.[227,1437,444,1924]" captionTargetPageId="63" captionText="Figs 224–228. Aetana lozadae Huber, sp. nov. 224–225. Left male palp, prolateral and retrolateral views. 226. Male prosoma, oblique frontal view. 227–228. Cleared female genitalia, ventral and dorsal views. rp = retrolatero-ventral process; vl = ventral lamina. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829592" httpUri="https://zenodo.org/record/3829592/files/figure.png" pageId="4" pageNumber="5">226</figureCitation>
), absent in 
<taxonomicName id="4C3922B2603BFF9194FAF9C54111F9F9" authorityName="Huber" authorityYear="2015" box="[995,1112,1603,1630]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kiukoki">
<emphasis id="B94D8523603BFF9194FAF9C54111F9F9" box="[995,1112,1603,1630]" italics="true" pageId="4" pageNumber="5">A. kiukoki</emphasis>
</taxonomicName>
group; proximal apophyses usually present, large in 
<taxonomicName id="4C3922B2603BFF919519F9E6473CF924" authorityName="Huber" authorityYear="2015" box="[512,629,1638,1665]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kiukoki">
<emphasis id="B94D8523603BFF919519F9E6473CF924" box="[512,629,1638,1665]" italics="true" pageId="4" pageNumber="5">A. kiukoki</emphasis>
</taxonomicName>
group (
<figureCitation id="130245B4603BFF9195D6F9E7466FF925" box="[719,806,1638,1665]" captionStart="Figs 62–69" captionStartId="23.[227,279,1896,1922]" captionTargetBox="[227,1437,421,1844]" captionTargetId="figure@23.[227,1437,421,1844]" captionTargetPageId="23" captionText="Figs 62–69. Aetana kiukoki Huber, sp. nov. 62–63. Male prosoma, oblique frontal and frontal views. 64. Female prosoma, frontal view. 65. Male clypeus and chelicerae, oblique frontal view. 66. Male clypeus modification, frontal view. 67. Epigynum, ventral view. 68. Male ALS and PMS. 69. Comb-hair on female tarsus 4. cl = clypeus; lca = lateral cheliceral apophysis. Scale lines: 62–64 = 400 µm; 65, 67 = 200 µm; 66 = 100 µm; 68–69 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3829543" httpUri="https://zenodo.org/record/3829543/files/figure.png" pageId="4" pageNumber="5">Fig. 66</figureCitation>
), absent in 
<taxonomicName id="4C3922B2603BFF9194B3F9E64166F924" authorityName="Huber" authorityYear="2015" box="[938,1071,1639,1665]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ocampoi">
<emphasis id="B94D8523603BFF9194B3F9E64166F924" box="[938,1071,1639,1665]" italics="true" pageId="4" pageNumber="5">A. ocampoi</emphasis>
</taxonomicName>
group (
<figureCitation id="130245B4603BFF919390F9E741AAF925" box="[1161,1251,1638,1665]" captionStart="Figs 13–15" captionStartId="10.[227,279,1968,1994]" captionTargetBox="[234,1437,704,1917]" captionTargetId="figure@10.[227,1437,703,1921]" captionTargetPageId="10" captionText="Figs 13–15. Aetana ocampoi Huber, sp. nov. 13–14. Left male palp, prolateral and retrolateral views (asterisk: retrolateral process of bulb). 15. Male chelicerae, frontal view. b = genital bulb; p = procursus. Scale lines: 13–14 = 0.5 mm; 15 = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3829525" httpUri="https://zenodo.org/record/3829525/files/figure.png" pageId="4" pageNumber="5">Figs 15</figureCitation>
, 
<figureCitation id="130245B4603BFF9193F6F9E74058F924" box="[1263,1297,1638,1664]" captionStart="Figs 38–44. — 38–40" captionStartId="16.[227,279,1906,1932]" captionTargetBox="[227,1432,599,1858]" captionTargetId="figure@16.[227,1437,593,1864]" captionTargetPageId="16" captionText="Figs 38–44. — 38–40. Aetana libjo Huber, sp. nov. 38–39. Left genital bulb in prolateral and retrolateral views. 40. Male chelicerae, frontal view. — 41–44. Aetana baganihan Huber, sp. nov. 41–42. Left genital bulb in prolateral and retrolateral views. 43–44. Cleared female genitalia, ventral and dorsal views (asterisk: median membranous structure). Scale lines: 38–39, 41–44 = 0.5 mm; 40 = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3829535" httpUri="https://zenodo.org/record/3829535/files/figure.png" pageId="4" pageNumber="5">40</figureCitation>
); chelicerae without modified hairs; without stridulatory ridges.
</paragraph>
</subSubSection>
<subSubSection id="C3230ABA603BFF9097FAF9484143FD00" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8B865931603BFF9097FAF94840D5FE12" blockId="4.[227,1437,1737,2045]" lastBlockId="5.[227,1437,305,438]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
PALPS. Coxa unmodified; trochanter usually with one retrolateral to ventral process, sometimes provided with scales or teeth (
<figureCitation id="130245B4603BFF9196C5F96D470DF8A3" box="[476,580,1772,1799]" captionStart="Figs 155–162" captionStartId="50.[227,279,1862,1888]" captionTargetBox="[227,1437,302,1837]" captionTargetId="figure@50.[227,1437,302,1837]" captionTargetPageId="50" captionText="Figs 155–162. Aetana indah Huber, sp. nov. 155–156. Left palp, prolatero-distal and retrolaterodistal views. 157. Left procursus, retrolatero-distal view. 158. Detail of left procursus (and trochanter apophysis), prolateral view. 159. Left embolus (arrow points at sperm duct opening). 160–161. Left male cheliceral apophyses, oblique frontal and frontal views. 162. Male gonopore. b = genital bulb; e = embolus; f = femur; p = procursus; ti = tibia; tr = trochanter. Scale lines: 155–156 = 200 µm; 157 = 100 µm; 158–160 = 60 µm; 161 = 80 µm; 162 = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.3829576" httpUri="https://zenodo.org/record/3829576/files/figure.png" pageId="4" pageNumber="5">Fig. 158</figureCitation>
), in 
<taxonomicName id="4C3922B2603BFF919565F96D464AF8A2" ID-CoL="65BD8" authorityName="Huber" authorityYear="2015" box="[636,771,1772,1798]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ocampoi">
<emphasis id="B94D8523603BFF919565F96D464AF8A2" box="[636,771,1772,1798]" italics="true" pageId="4" pageNumber="5">A. ocampoi</emphasis>
</taxonomicName>
group fused to femur (
<figureCitation id="130245B4603BFF919302F96D413EF8A2" box="[1051,1143,1772,1798]" captionStart="Figs 13–15" captionStartId="10.[227,279,1968,1994]" captionTargetBox="[234,1437,704,1917]" captionTargetId="figure@10.[227,1437,703,1921]" captionTargetPageId="10" captionText="Figs 13–15. Aetana ocampoi Huber, sp. nov. 13–14. Left male palp, prolateral and retrolateral views (asterisk: retrolateral process of bulb). 15. Male chelicerae, frontal view. b = genital bulb; p = procursus. Scale lines: 13–14 = 0.5 mm; 15 = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3829525" httpUri="https://zenodo.org/record/3829525/files/figure.png" pageId="4" pageNumber="5">Figs 14</figureCitation>
, 
<figureCitation id="130245B4603BFF91939FF96D41EEF8A2" box="[1158,1191,1772,1798]" captionStart="Figs 26–29" captionStartId="14.[227,279,1932,1958]" captionTargetBox="[227,1433,346,1885]" captionTargetId="figure@14.[227,1437,346,1890]" captionTargetPageId="14" captionText="Figs 26–29. Aetana libjo Huber, sp. nov. 26–27. Left male palp, prolateral and retrolateral views (asterisk: retrolateral process of bulb). 28–29. Cleared female genitalia, ventral and dorsal views (asterisk: median membranous structure). b = genital bulb; p = procursus; rt = retrolateral trichobothrium. Scale lines: 26–27 = 0.5 mm; 28–29 = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3829531" httpUri="https://zenodo.org/record/3829531/files/figure.png" pageId="4" pageNumber="5">27</figureCitation>
); trochanter in some species with additional prolateral apophysis (
<figureCitation id="130245B4603BFF9195F3F88E4606F88D" box="[746,847,1807,1833]" captionStart="Figs 189–194. — 189–193" captionStartId="55.[227,279,1894,1920]" captionTargetBox="[227,1436,450,1840]" captionTargetId="figure@55.[227,1437,450,1847]" captionTargetPageId="55" captionText="Figs 189–194. — 189–193. Aetana abadae Huber, sp. nov. 189–190. Left male palp, prolateral and retrolateral views. 191. Male prosoma, oblique frontal view. 192–193. Cleared female genitalia, ventral and dorsal views. — 194. A. omayan Huber, 2005, cleared female genitalia, dorsal view. Arrows point at membranous pockets. ip = internal sclerotized pocket; rp = retrolatero-ventral process; vl = ventral lamina. Scale lines: 0.5 mm (female genitalia at same scale)." figureDoi="http://doi.org/10.5281/zenodo.3829582" httpUri="https://zenodo.org/record/3829582/files/figure.png" pageId="4" pageNumber="5">Fig. 189</figureCitation>
); femur rarely simple (
<figureCitation id="130245B4603BFF91934FF88E41E2F88D" box="[1110,1195,1807,1833]" captionStart="Figs 26–29" captionStartId="14.[227,279,1932,1958]" captionTargetBox="[227,1433,346,1885]" captionTargetId="figure@14.[227,1437,346,1890]" captionTargetPageId="14" captionText="Figs 26–29. Aetana libjo Huber, sp. nov. 26–27. Left male palp, prolateral and retrolateral views (asterisk: retrolateral process of bulb). 28–29. Cleared female genitalia, ventral and dorsal views (asterisk: median membranous structure). b = genital bulb; p = procursus; rt = retrolateral trichobothrium. Scale lines: 26–27 = 0.5 mm; 28–29 = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3829531" httpUri="https://zenodo.org/record/3829531/files/figure.png" pageId="4" pageNumber="5">Fig. 27</figureCitation>
), usually with one or more processes, in 
<taxonomicName id="4C3922B2603BFF9196A2F8B24709F8E8" authorityName="Huber" authorityYear="2005" box="[443,576,1842,1869]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kinabalu">
<emphasis id="B94D8523603BFF9196A2F8B24709F8E8" box="[443,576,1842,1869]" italics="true" pageId="4" pageNumber="5">A. kinabalu</emphasis>
</taxonomicName>
group with complex set of up to four processes (e.g., 
<figureCitation id="130245B4603BFF919382F8B34075F8E9" box="[1179,1340,1842,1869]" captionStart="Figs 150–154" captionStartId="49.[227,279,1934,1960]" captionTargetBox="[227,1433,540,1890]" captionTargetId="figure@49.[227,1437,540,1893]" captionTargetPageId="49" captionText="Figs 150–154. Aetana indah Huber, sp. nov. 150–151. Left male palp, prolateral and retrolateral views (arrow points at prolateral apophysis near hinge). 152. Male chelicerae, frontal view. 153–154. Cleared female genitalia, ventral and dorsal views (arrow points at membranous pocket). ip = internal sclerotized pocket; rp = retrolatero-ventral process; vl = ventral lamina. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829574" httpUri="https://zenodo.org/record/3829574/files/figure.png" pageId="4" pageNumber="5">Figs 150–151</figureCitation>
); patella either triangular in lateral view or ventral side longer than usual (e.g., 
<figureCitation id="130245B4603BFF919302F8D4413DF8D4" box="[1051,1140,1877,1904]" captionStart="Figs 13–15" captionStartId="10.[227,279,1968,1994]" captionTargetBox="[234,1437,704,1917]" captionTargetId="figure@10.[227,1437,703,1921]" captionTargetPageId="10" captionText="Figs 13–15. Aetana ocampoi Huber, sp. nov. 13–14. Left male palp, prolateral and retrolateral views (asterisk: retrolateral process of bulb). 15. Male chelicerae, frontal view. b = genital bulb; p = procursus. Scale lines: 13–14 = 0.5 mm; 15 = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3829525" httpUri="https://zenodo.org/record/3829525/files/figure.png" pageId="4" pageNumber="5">Figs 14</figureCitation>
, 
<figureCitation id="130245B4603BFF91939BF8D441EAF8D4" box="[1154,1187,1877,1904]" captionStart="Figs 57–61" captionStartId="22.[227,279,1966,1992]" captionTargetBox="[227,1432,469,1920]" captionTargetId="figure@22.[227,1437,469,1922]" captionTargetPageId="22" captionText="Figs 57–61. Aetana kiukoki Huber, sp. nov. 57–58. Left male palp, prolateral and retrolateral views. 59. Male prosoma, oblique frontal view. 60–61. Cleared female genitalia, ventral and dorsal views. b = genital bulb; e = embolus; p = procursus. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829541" httpUri="https://zenodo.org/record/3829541/files/figure.png" pageId="4" pageNumber="5">58</figureCitation>
); tibia either of usual shape (e.g., 
<figureCitation id="130245B4603BFF919668F8F84482F837" box="[369,459,1913,1939]" captionStart="Figs 13–15" captionStartId="10.[227,279,1968,1994]" captionTargetBox="[234,1437,704,1917]" captionTargetId="figure@10.[227,1437,703,1921]" captionTargetPageId="10" captionText="Figs 13–15. Aetana ocampoi Huber, sp. nov. 13–14. Left male palp, prolateral and retrolateral views (asterisk: retrolateral process of bulb). 15. Male chelicerae, frontal view. b = genital bulb; p = procursus. Scale lines: 13–14 = 0.5 mm; 15 = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3829525" httpUri="https://zenodo.org/record/3829525/files/figure.png" pageId="4" pageNumber="5">Figs 14</figureCitation>
, 
<figureCitation id="130245B4603BFF9196C0F8F8474FF837" box="[473,518,1913,1939]" captionStart="Figs 126–129" captionStartId="42.[227,279,1934,1960]" captionTargetBox="[227,1435,586,1879]" captionTargetId="figure@42.[227,1437,586,1886]" captionTargetPageId="42" captionText="Figs 126–129. Aetana gaya Huber, sp. nov. 126–127. Left male palp, prolateral and retrolateral views. 128–129. Cleared female genitalia, ventral and dorsal views (arrow points at membranous pocket). b = genital bulb; e = embolus; rp = retrolatero-ventral process; vl = ventral lamina. Scale lines: 126–127 = 0.5 mm; 128–129 = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3829566" httpUri="https://zenodo.org/record/3829566/files/figure.png" pageId="4" pageNumber="5">127</figureCitation>
) or rather small and slender (
<taxonomicName id="4C3922B2603BFF91947FF8F84695F837" authorityName="Huber" authorityYear="2015" box="[870,988,1913,1939]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kiukoki">
<emphasis id="B94D8523603BFF91947FF8F84695F837" box="[870,988,1913,1939]" italics="true" pageId="4" pageNumber="5">A. kiukoki</emphasis>
</taxonomicName>
group; e.g., 
<figureCitation id="130245B4603BFF91936AF8F84183F837" box="[1139,1226,1912,1939]" captionStart="Figs 57–61" captionStartId="22.[227,279,1966,1992]" captionTargetBox="[227,1432,469,1920]" captionTargetId="figure@22.[227,1437,469,1922]" captionTargetPageId="22" captionText="Figs 57–61. Aetana kiukoki Huber, sp. nov. 57–58. Left male palp, prolateral and retrolateral views. 59. Male prosoma, oblique frontal view. 60–61. Cleared female genitalia, ventral and dorsal views. b = genital bulb; e = embolus; p = procursus. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829541" httpUri="https://zenodo.org/record/3829541/files/figure.png" pageId="4" pageNumber="5">Fig. 58</figureCitation>
), with retrolateral trichobothrium in very distal position (close to tibia-tarsus joint; 
<figureCitation id="130245B4603BFF9194DDF81D4152F812" box="[964,1051,1948,1974]" captionStart="Figs 26–29" captionStartId="14.[227,279,1932,1958]" captionTargetBox="[227,1433,346,1885]" captionTargetId="figure@14.[227,1437,346,1890]" captionTargetPageId="14" captionText="Figs 26–29. Aetana libjo Huber, sp. nov. 26–27. Left male palp, prolateral and retrolateral views (asterisk: retrolateral process of bulb). 28–29. Cleared female genitalia, ventral and dorsal views (asterisk: median membranous structure). b = genital bulb; p = procursus; rt = retrolateral trichobothrium. Scale lines: 26–27 = 0.5 mm; 28–29 = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3829531" httpUri="https://zenodo.org/record/3829531/files/figure.png" pageId="4" pageNumber="5">Figs 27</figureCitation>
, 
<figureCitation id="130245B4603BFF91933CF81D411DF812" box="[1061,1108,1948,1974]" captionStart="Figs 113–117" captionStartId="38.[227,279,1930,1956]" captionTargetBox="[231,1434,479,1882]" captionTargetId="figure@38.[228,1438,476,1888]" captionTargetPageId="38" captionText="Figs 113–117. Aetana lambir Huber, sp. nov. 113–114. Left male palp, prolateral and retrolateral views. 115. Male chelicerae, frontal view. 116–117. Cleared female genitalia, ventral and dorsal views (arrow points at membranous pocket). b = genital bulb; e = embolus; rp = retrolatero-ventral process; rt = retrolateral trichobothrium; vl = ventral lamina. Scale lines: 113–114 = 0.5 mm; 115–117 = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3829562" httpUri="https://zenodo.org/record/3829562/files/figure.png" pageId="4" pageNumber="5">114</figureCitation>
); palpal tarsus small, usually with capsular tarsal organ (
<figureCitation id="130245B4603BFF919530F83E47DCF87D" box="[553,661,1983,2009]" captionStart="Figs 130–136" captionStartId="43.[225,277,1930,1956]" captionTargetBox="[227,1437,336,1884]" captionTargetId="figure@43.[227,1437,336,1884]" captionTargetPageId="43" captionText="Figs 130–136. Aetana gaya Huber, sp. nov. 130–131. Male prosoma, frontal and oblique frontal views. 132. Female prosoma, frontal view. 133. Detail of male metatarsus 1. 134. Male palpal tarsal organ. 135. Male carapace and ocular area. 136. Right procursus, retrolateral view. rp = retrolatero-ventral process; vl = ventral lamina. Scale lines: 130–132, 135 = 200 µm; 133 = 80 µm; 134 = 8 µm; 136 = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3829568" httpUri="https://zenodo.org/record/3829568/files/figure.png" pageId="4" pageNumber="5">Figs 134</figureCitation>
, 
<figureCitation id="130245B4603BFF9195BDF83E479AF87D" box="[676,723,1983,2009]" captionStart="Figs 195–203. — 195–197" captionStartId="56.[227,279,1824,1850]" captionTargetBox="[242,1421,302,1783]" captionTargetId="figure@56.[242,1421,302,1783]" captionTargetPageId="56" captionText="Figs 195–203. — 195–197. Aetana abadae Huber, sp. nov. 195. Epigynum, ventral view. 196. Detail of preceding, showing opening of membranous pocket (arrow on Fig. 195). 197. Female ALS. — 198–203. A. omayan Huber, 2005. 198. Male prosoma, frontal view. 199. Left palp, retrolateral view. 200–201. Right procursus, prolatero-dorsal and prolateral views. 202. Detail of preceding. 203. Male palpal tarsal organ. b = genital bulb; e = embolus; f = femur; p = procursus; rp = retrolatero-ventral process; ti = tibia; tr = trochanter; vl = ventral lamina. Scale lines: 195, 199 = 300 µm; 196, 202 = 40 µm; 197, 203 = 10 µm; 198 = 500 µm; 200 = 200 µm; 201 = 100 µm" figureDoi="http://doi.org/10.5281/zenodo.3829584" httpUri="https://zenodo.org/record/3829584/files/figure.png" pageId="4" pageNumber="5">203</figureCitation>
), exposed in 
<taxonomicName id="4C3922B2603BFF91946CF83E416AF87D" authority="Huber" authorityName="Huber" authorityYear="2015" box="[885,1059,1983,2009]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="libjo" status="sp. nov.">
<emphasis id="B94D8523603BFF91946CF83E4684F87D" box="[885,973,1983,2009]" italics="true" pageId="4" pageNumber="5">A. libjo</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E3858603BFF919328F84141C4F87E" box="[1073,1165,1984,2010]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="130245B4603BFF9193B9F83E41BEF87E" box="[1184,1271,1983,2010]" captionStart="Figs 30–37" captionStartId="15.[227,279,1909,1935]" captionTargetBox="[227,1437,310,1867]" captionTargetId="figure@15.[227,1437,310,1867]" captionTargetPageId="15" captionText="Figs 30–37. Aetana libjo Huber, sp. nov. 30. Processes of right genital bulb, retrolateral view. 31. Detail of preceding. 32. Comb-hairs on female tarsus 4. 33. Processes of left genital bulb, prolateral view. 34. Detail of preceding. 35. Male palpal tarsal organ. 36. Female ALS. 37. Male gonopore. Scale lines: 30, 33 = 100 µm; 31, 37 = 40 µm; 32, 34 = 20 µm; 35–36 = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3829533" httpUri="https://zenodo.org/record/3829533/files/figure.png" pageId="4" pageNumber="5">Fig. 35</figureCitation>
) (not clear in 
<taxonomicName id="4C3922B2603BFF9197FAF86244F1F858" authority="Huber" authorityName="Huber" authorityYear="2015" box="[227,440,2018,2045]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ocampoi" status="sp. nov.">
<emphasis id="B94D8523603BFF9197FAF862442FF858" box="[227,358,2019,2045]" italics="true" pageId="4" pageNumber="5">A. ocampoi</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E3858603BFF9196DAF8624755F859" box="[451,540,2019,2045]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C3922B2603BFF91954AF8624609F858" authority="Huber" authorityName="Huber" authorityYear="2015" box="[595,832,2018,2045]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="baganihan" status="sp. nov.">
<emphasis id="B94D8523603BFF91954AF86247A7F858" box="[595,750,2018,2045]" italics="true" pageId="4" pageNumber="5">A. baganihan</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E3858603BFF919452F86246E1F859" box="[843,936,2019,2045]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
); procursus usually complex with proximal and distal parts connected by membranous hinge; procursus in 
<taxonomicName id="4C3922B2603AFF9094A4FEB34178FEEF" authorityName="Huber" authorityYear="2015" box="[957,1073,305,332]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kiukoki">
<emphasis id="B94D8523603AFF9094A4FEB34178FEEF" box="[957,1073,305,332]" italics="true" pageId="5" pageNumber="6">A. kiukoki</emphasis>
</taxonomicName>
group unusually long (
<figureCitation id="130245B4603AFF909224FEB040D1FEE8" box="[1341,1432,305,332]" captionStart="Figs 57–61" captionStartId="22.[227,279,1966,1992]" captionTargetBox="[227,1432,469,1920]" captionTargetId="figure@22.[227,1437,469,1922]" captionTargetPageId="22" captionText="Figs 57–61. Aetana kiukoki Huber, sp. nov. 57–58. Left male palp, prolateral and retrolateral views. 59. Male prosoma, oblique frontal view. 60–61. Cleared female genitalia, ventral and dorsal views. b = genital bulb; e = embolus; p = procursus. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829541" httpUri="https://zenodo.org/record/3829541/files/figure.png" pageId="5" pageNumber="6">Figs 58</figureCitation>
, 
<figureCitation id="130245B4603AFF9097FAFED5444AFECB" box="[227,259,340,367]" captionStart="Figs 75–79" captionStartId="28.[227,279,1966,1992]" captionTargetBox="[230,1434,356,1921]" captionTargetId="figure@28.[227,1437,353,1922]" captionTargetPageId="28" captionText="Figs 75–79. Aetana loboc Huber, sp. nov. 75–76. Left male palp, prolateral and retrolateral views. 77. Male prosoma, frontal view. 78–79. Cleared female genitalia, ventral and dorsal views. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829547" httpUri="https://zenodo.org/record/3829547/files/figure.png" pageId="5" pageNumber="6">76</figureCitation>
), in 
<taxonomicName id="4C3922B2603AFF909628FED444FAFECA" authorityName="Huber" authorityYear="2015" box="[305,435,341,367]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ocampoi">
<emphasis id="B94D8523603AFF909628FED444FAFECA" box="[305,435,341,367]" italics="true" pageId="5" pageNumber="6">A. ocampoi</emphasis>
</taxonomicName>
group reduced to simple semi-transparent process (
<figureCitation id="130245B4603AFF9094E3FED5411BFECA" box="[1018,1106,340,366]" captionStart="Figs 13–15" captionStartId="10.[227,279,1968,1994]" captionTargetBox="[234,1437,704,1917]" captionTargetId="figure@10.[227,1437,703,1921]" captionTargetPageId="10" captionText="Figs 13–15. Aetana ocampoi Huber, sp. nov. 13–14. Left male palp, prolateral and retrolateral views (asterisk: retrolateral process of bulb). 15. Male chelicerae, frontal view. b = genital bulb; p = procursus. Scale lines: 13–14 = 0.5 mm; 15 = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3829525" httpUri="https://zenodo.org/record/3829525/files/figure.png" pageId="5" pageNumber="6">Figs 14</figureCitation>
, 
<figureCitation id="130245B4603AFF909345FED54134FECA" box="[1116,1149,340,366]" captionStart="Figs 26–29" captionStartId="14.[227,279,1932,1958]" captionTargetBox="[227,1433,346,1885]" captionTargetId="figure@14.[227,1437,346,1890]" captionTargetPageId="14" captionText="Figs 26–29. Aetana libjo Huber, sp. nov. 26–27. Left male palp, prolateral and retrolateral views (asterisk: retrolateral process of bulb). 28–29. Cleared female genitalia, ventral and dorsal views (asterisk: median membranous structure). b = genital bulb; p = procursus; rt = retrolateral trichobothrium. Scale lines: 26–27 = 0.5 mm; 28–29 = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3829531" httpUri="https://zenodo.org/record/3829531/files/figure.png" pageId="5" pageNumber="6">27</figureCitation>
); bulb either with weakly sclerotized embolus as only process (
<figureCitation id="130245B4603AFF909591FEF647A7FE36" box="[648,750,375,402]" captionStart="Figs 113–117" captionStartId="38.[227,279,1930,1956]" captionTargetBox="[231,1434,479,1882]" captionTargetId="figure@38.[228,1438,476,1888]" captionTargetPageId="38" captionText="Figs 113–117. Aetana lambir Huber, sp. nov. 113–114. Left male palp, prolateral and retrolateral views. 115. Male chelicerae, frontal view. 116–117. Cleared female genitalia, ventral and dorsal views (arrow points at membranous pocket). b = genital bulb; e = embolus; rp = retrolatero-ventral process; rt = retrolateral trichobothrium; vl = ventral lamina. Scale lines: 113–114 = 0.5 mm; 115–117 = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3829562" httpUri="https://zenodo.org/record/3829562/files/figure.png" pageId="5" pageNumber="6">Figs 114</figureCitation>
, 
<figureCitation id="130245B4603AFF9095EEFEF6466FFE36" box="[759,806,375,402]" captionStart="Figs 155–162" captionStartId="50.[227,279,1862,1888]" captionTargetBox="[227,1437,302,1837]" captionTargetId="figure@50.[227,1437,302,1837]" captionTargetPageId="50" captionText="Figs 155–162. Aetana indah Huber, sp. nov. 155–156. Left palp, prolatero-distal and retrolaterodistal views. 157. Left procursus, retrolatero-distal view. 158. Detail of left procursus (and trochanter apophysis), prolateral view. 159. Left embolus (arrow points at sperm duct opening). 160–161. Left male cheliceral apophyses, oblique frontal and frontal views. 162. Male gonopore. b = genital bulb; e = embolus; f = femur; p = procursus; ti = tibia; tr = trochanter. Scale lines: 155–156 = 200 µm; 157 = 100 µm; 158–160 = 60 µm; 161 = 80 µm; 162 = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.3829576" httpUri="https://zenodo.org/record/3829576/files/figure.png" pageId="5" pageNumber="6">159</figureCitation>
), in 
<taxonomicName id="4C3922B2603AFF90944CFEF9468EFE36" authorityName="Huber" authorityYear="2015" box="[853,967,376,402]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kiukoki">
<emphasis id="B94D8523603AFF90944CFEF9468EFE36" box="[853,967,376,402]" italics="true" pageId="5" pageNumber="6">A. kiukoki</emphasis>
</taxonomicName>
group with additional short membranous process (
<figureCitation id="130245B4603AFF909649FE1A44F8FE12" box="[336,433,411,438]" captionStart="Figs 57–61" captionStartId="22.[227,279,1966,1992]" captionTargetBox="[227,1432,469,1920]" captionTargetId="figure@22.[227,1437,469,1922]" captionTargetPageId="22" captionText="Figs 57–61. Aetana kiukoki Huber, sp. nov. 57–58. Left male palp, prolateral and retrolateral views. 59. Male prosoma, oblique frontal view. 60–61. Cleared female genitalia, ventral and dorsal views. b = genital bulb; e = embolus; p = procursus. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829541" httpUri="https://zenodo.org/record/3829541/files/figure.png" pageId="5" pageNumber="6">Figs 58</figureCitation>
, 
<figureCitation id="130245B4603AFF9096DAFE1A44AAFE12" box="[451,483,411,438]" captionStart="Figs 75–79" captionStartId="28.[227,279,1966,1992]" captionTargetBox="[230,1434,356,1921]" captionTargetId="figure@28.[227,1437,353,1922]" captionTargetPageId="28" captionText="Figs 75–79. Aetana loboc Huber, sp. nov. 75–76. Left male palp, prolateral and retrolateral views. 77. Male prosoma, frontal view. 78–79. Cleared female genitalia, ventral and dorsal views. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829547" httpUri="https://zenodo.org/record/3829547/files/figure.png" pageId="5" pageNumber="6">75</figureCitation>
), in 
<taxonomicName id="4C3922B2603AFF90953AFE1A47E7FE11" authorityName="Huber" authorityYear="2015" box="[547,686,411,437]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ocampoi">
<emphasis id="B94D8523603AFF90953AFE1A47E7FE11" box="[547,686,411,437]" italics="true" pageId="5" pageNumber="6">A. ocampoi</emphasis>
</taxonomicName>
group with additional novel processes (
<figureCitation id="130245B4603AFF9093BDFE1A4079FE11" box="[1188,1328,411,437]" captionStart="Figs 13–15" captionStartId="10.[227,279,1968,1994]" captionTargetBox="[234,1437,704,1917]" captionTargetId="figure@10.[227,1437,703,1921]" captionTargetPageId="10" captionText="Figs 13–15. Aetana ocampoi Huber, sp. nov. 13–14. Left male palp, prolateral and retrolateral views (asterisk: retrolateral process of bulb). 15. Male chelicerae, frontal view. b = genital bulb; p = procursus. Scale lines: 13–14 = 0.5 mm; 15 = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3829525" httpUri="https://zenodo.org/record/3829525/files/figure.png" pageId="5" pageNumber="6">Figs 13–14</figureCitation>
, 
<figureCitation id="130245B4603AFF90925BFE1A40D9FE12" box="[1346,1424,411,438]" captionStart="Figs 26–29" captionStartId="14.[227,279,1932,1958]" captionTargetBox="[227,1433,346,1885]" captionTargetId="figure@14.[227,1437,346,1890]" captionTargetPageId="14" captionText="Figs 26–29. Aetana libjo Huber, sp. nov. 26–27. Left male palp, prolateral and retrolateral views (asterisk: retrolateral process of bulb). 28–29. Cleared female genitalia, ventral and dorsal views (asterisk: median membranous structure). b = genital bulb; p = procursus; rt = retrolateral trichobothrium. Scale lines: 26–27 = 0.5 mm; 28–29 = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3829531" httpUri="https://zenodo.org/record/3829531/files/figure.png" pageId="5" pageNumber="6">26–27</figureCitation>
).
</paragraph>
<paragraph id="8B865931603AFF9097FAFE5B4143FD00" blockId="5.[227,1437,474,676]" pageId="5" pageNumber="6">
LEGS. Without spines; with curved hairs on tibiae and/or metatarsi (
<figureCitation id="130245B4603AFF909310FE5B4138FE50" box="[1033,1137,474,500]" captionStart="Figs 130–136" captionStartId="43.[225,277,1930,1956]" captionTargetBox="[227,1437,336,1884]" captionTargetId="figure@43.[227,1437,336,1884]" captionTargetPageId="43" captionText="Figs 130–136. Aetana gaya Huber, sp. nov. 130–131. Male prosoma, frontal and oblique frontal views. 132. Female prosoma, frontal view. 133. Detail of male metatarsus 1. 134. Male palpal tarsal organ. 135. Male carapace and ocular area. 136. Right procursus, retrolateral view. rp = retrolatero-ventral process; vl = ventral lamina. Scale lines: 130–132, 135 = 200 µm; 133 = 80 µm; 134 = 8 µm; 136 = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3829568" httpUri="https://zenodo.org/record/3829568/files/figure.png" pageId="5" pageNumber="6">Fig. 133</figureCitation>
); usually without or with few vertical hairs, only in 
<taxonomicName id="4C3922B2603AFF90950CFE7F47F7FDB3" authority="Huber" authorityName="Huber" authorityYear="2015" box="[533,702,509,536]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="libjo" status="sp. nov.">
<emphasis id="B94D8523603AFF90950CFE7F4725FDB3" box="[533,620,509,536]" italics="true" pageId="5" pageNumber="6">A. libjo</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E3858603AFF9095D0FE7F466DFDBC" box="[713,804,510,536]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C3922B2603AFF909445FE7F4102FDB3" authority="Huber" authorityName="Huber" authorityYear="2015" box="[860,1099,509,536]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="baganihan" status="sp. nov.">
<emphasis id="B94D8523603AFF909445FE7F46B0FDB3" box="[860,1017,509,536]" italics="true" pageId="5" pageNumber="6">A. baganihan</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E3858603AFF90934EFE7F41F8FDBC" box="[1111,1201,510,536]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
with one dense row retrolatero-dorsally on each tibia; retrolateral trichobothrium on tibia 1 very proximal (at 2–4% of tibia length), prolateral trichobothrium absent on tibia 1, present on other tibiae. Tarsus 1 usually with ~25–30 pseudosegments, fairly distinct distally; tarsus 4 with single row of ventral comb-hairs of a modified 
<taxonomicName id="4C3922B2603AFF9097FAFD0B4403FD00" authorityName="Thorell" authorityYear="1898" box="[227,330,650,676]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B94D8523603AFF9097FAFD0B4403FD00" box="[227,330,650,676]" italics="true" pageId="5" pageNumber="6">Belisana</emphasis>
</taxonomicName>
- 
<typeStatus id="5482E793603AFF909648FD0B44CCFD00" box="[337,389,650,676]" pageId="5" pageNumber="6">type</typeStatus>
(
<emphasis id="B94D8523603AFF90968EFD0B44FDFD00" box="[407,436,650,676]" italics="true" pageId="5" pageNumber="6">cf.</emphasis>
<bibRefCitation id="EFA824C0603AFF9096A2FD0B464AFD00" author="Huber B. A. &amp; Fleckenstein N." box="[443,771,650,676]" pageId="5" pageNumber="6" pagination="232 - 240" refId="ref27255" refString="Huber B. A. &amp; Fleckenstein N. 2008. Comb-hairs on the fourth tarsi in pholcid spiders (Araneae, Pholcidae). Journal of Arachnology 36: 232 - 240. http: // dx. doi. org / 10.1636 / CSh 07 - 71.1" type="journal article" year="2008">Huber &amp; Fleckenstein 2008</bibRefCitation>
; 
<figureCitation id="130245B4603AFF909416FD0B4620FD00" box="[783,873,650,676]" captionStart="Figs 30–37" captionStartId="15.[227,279,1909,1935]" captionTargetBox="[227,1437,310,1867]" captionTargetId="figure@15.[227,1437,310,1867]" captionTargetPageId="15" captionText="Figs 30–37. Aetana libjo Huber, sp. nov. 30. Processes of right genital bulb, retrolateral view. 31. Detail of preceding. 32. Comb-hairs on female tarsus 4. 33. Processes of left genital bulb, prolateral view. 34. Detail of preceding. 35. Male palpal tarsal organ. 36. Female ALS. 37. Male gonopore. Scale lines: 30, 33 = 100 µm; 31, 37 = 40 µm; 32, 34 = 20 µm; 35–36 = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3829533" httpUri="https://zenodo.org/record/3829533/files/figure.png" pageId="5" pageNumber="6">Figs 32</figureCitation>
, 
<figureCitation id="130245B4603AFF90946CFD0B46DCFD00" box="[885,917,650,676]" captionStart="Figs 62–69" captionStartId="23.[227,279,1896,1922]" captionTargetBox="[227,1437,421,1844]" captionTargetId="figure@23.[227,1437,421,1844]" captionTargetPageId="23" captionText="Figs 62–69. Aetana kiukoki Huber, sp. nov. 62–63. Male prosoma, oblique frontal and frontal views. 64. Female prosoma, frontal view. 65. Male clypeus and chelicerae, oblique frontal view. 66. Male clypeus modification, frontal view. 67. Epigynum, ventral view. 68. Male ALS and PMS. 69. Comb-hair on female tarsus 4. cl = clypeus; lca = lateral cheliceral apophysis. Scale lines: 62–64 = 400 µm; 65, 67 = 200 µm; 66 = 100 µm; 68–69 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3829543" httpUri="https://zenodo.org/record/3829543/files/figure.png" pageId="5" pageNumber="6">69</figureCitation>
, 
<figureCitation id="130245B4603AFF9094B8FD0B4688FD00" box="[929,961,650,676]" captionStart="Figs 80–86" captionStartId="29.[227,279,1932,1958]" captionTargetBox="[227,1437,463,1888]" captionTargetId="figure@29.[227,1437,463,1888]" captionTargetPageId="29" captionText="Figs 80–86. Aetana loboc Huber, sp. nov. 80. Female prosoma, frontal view. 81. Female ALS. 82. Comb-hair on female tarsus 4. 83. Detail of female tarsus 1. 84. Detail of female tarsus 4. 85. Epigynum. 86. Detail of preceding. Scale lines: 80, 85 = 200 µm; 81 = 10 µm; 82 = 8 µm; 83–84 = 40 µm; 86 = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3829549" httpUri="https://zenodo.org/record/3829549/files/figure.png" pageId="5" pageNumber="6">82</figureCitation>
, 
<figureCitation id="130245B4603AFF9094D4FD0B46B7FD00" box="[973,1022,650,676]" captionStart="Figs 204–212" captionStartId="57.[227,279,1897,1923]" captionTargetBox="[227,1437,514,1857]" captionTargetId="figure@57.[227,1437,514,1857]" captionTargetPageId="57" captionText="Figs 204–212. A. omayan Huber, 2005. 204. Left procursus tip (ventral lamina), retrolateral view. 205. Right procursus tip (pointed sclerite and ventral lamina), prolateral view. 206. Male gonopore. 207. Detail of male tarsus 1. 208–209. Comb-hairs on male tarsus 4. 210–211. Male and female ALS. 212. Epigynum. Scale lines: 204 = 50 µm; 205 = 40 µm; 206, 208 = 30 µm; 207 = 60 µm; 209 = 8 µm; 210–211 = 20 µm; 212 = 400 µm." figureDoi="http://doi.org/10.5281/zenodo.3829586" httpUri="https://zenodo.org/record/3829586/files/figure.png" pageId="5" pageNumber="6">209</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3230ABA603AFF909609FD4D4422FD42" box="[272,363,716,742]" pageId="5" pageNumber="6" type="description">
<paragraph id="8B865931603AFF909609FD4D4422FD42" blockId="5.[227,1437,716,1164]" box="[272,363,716,742]" pageId="5" pageNumber="6">
<emphasis id="B94D8523603AFF909609FD4D4422FD42" bold="true" box="[272,363,716,742]" pageId="5" pageNumber="6">Female</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3230ABA603AFF9097FAFD6F4620FB28" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8B865931603AFF9097FAFD6F4620FB28" blockId="5.[227,1437,716,1164]" pageId="5" pageNumber="6">
Similar to male but eye triads on lower humps (
<figureCitation id="130245B4603AFF90942AFD6F46DBFCAC" box="[819,914,750,776]" captionStart="Figs 62–69" captionStartId="23.[227,279,1896,1922]" captionTargetBox="[227,1437,421,1844]" captionTargetId="figure@23.[227,1437,421,1844]" captionTargetPageId="23" captionText="Figs 62–69. Aetana kiukoki Huber, sp. nov. 62–63. Male prosoma, oblique frontal and frontal views. 64. Female prosoma, frontal view. 65. Male clypeus and chelicerae, oblique frontal view. 66. Male clypeus modification, frontal view. 67. Epigynum, ventral view. 68. Male ALS and PMS. 69. Comb-hair on female tarsus 4. cl = clypeus; lca = lateral cheliceral apophysis. Scale lines: 62–64 = 400 µm; 65, 67 = 200 µm; 66 = 100 µm; 68–69 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3829543" httpUri="https://zenodo.org/record/3829543/files/figure.png" pageId="5" pageNumber="6">Figs 64</figureCitation>
, 
<figureCitation id="130245B4603AFF9094BBFD6E468BFCAD" box="[930,962,751,777]" captionStart="Figs 80–86" captionStartId="29.[227,279,1932,1958]" captionTargetBox="[227,1437,463,1888]" captionTargetId="figure@29.[227,1437,463,1888]" captionTargetPageId="29" captionText="Figs 80–86. Aetana loboc Huber, sp. nov. 80. Female prosoma, frontal view. 81. Female ALS. 82. Comb-hair on female tarsus 4. 83. Detail of female tarsus 1. 84. Detail of female tarsus 4. 85. Epigynum. 86. Detail of preceding. Scale lines: 80, 85 = 200 µm; 81 = 10 µm; 82 = 8 µm; 83–84 = 40 µm; 86 = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3829549" httpUri="https://zenodo.org/record/3829549/files/figure.png" pageId="5" pageNumber="6">80</figureCitation>
, 
<figureCitation id="130245B4603AFF9094CBFD6E46B7FCAD" box="[978,1022,751,777]" captionStart="Figs 118–125" captionStartId="39.[227,279,1896,1922]" captionTargetBox="[227,1437,374,1852]" captionTargetId="figure@39.[227,1437,374,1852]" captionTargetPageId="39" captionText="Figs 118–125. Aetana lambir Huber, sp. nov. 118–119. Male and female prosomata, frontal views. 120. Male palpal tarsal organ. 121. Left procursus, prolateral view. 122. Detail of preceding. 123. Right procursus (and femur apophysis), retrolateral view. 124. Female ALS. 125. Epigynum. f = femur; vl = ventral lamina. Scale lines: 118–119, 125 = 200 µm; 120 = 8 µm; 121 = 50 µm; 122 = 20 µm; 123 = 60 µm; 124 = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3829564" httpUri="https://zenodo.org/record/3829564/files/figure.png" pageId="5" pageNumber="6">119</figureCitation>
) and closer together than in male (sexual dimorphism low in 
<taxonomicName id="4C3922B2603AFF90952BFC9347F2FC88" ID-CoL="65BD8" authorityName="Huber" authorityYear="2015" box="[562,699,786,812]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ocampoi">
<emphasis id="B94D8523603AFF90952BFC9347F2FC88" box="[562,699,786,812]" italics="true" pageId="5" pageNumber="6">A. ocampoi</emphasis>
</taxonomicName>
group; in other groups, eye triads in females often less than half as wide apart as in males); clypeus and chelicerae unmodified; legs slightly shorter than in males (tibia 1 ~4.0–9.0). Females in 
<taxonomicName id="4C3922B2603AFF90955FFCD8478DFCD6" authorityName="Huber" authorityYear="2005" box="[582,708,857,883]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="omayan">
<emphasis id="B94D8523603AFF90955FFCD8478DFCD6" box="[582,708,857,883]" italics="true" pageId="5" pageNumber="6">A. omayan</emphasis>
</taxonomicName>
group with stridulatory apparatus between prosoma (elongate median plate on carapace posteriorly; 
<figureCitation id="130245B4603AFF909583FCFA4672FC32" box="[666,827,891,918]" captionStart="Figs 178–188" captionStartId="53.[227,279,1860,1886]" captionTargetBox="[228,1438,336,1814]" captionTargetId="figure@53.[228,1438,336,1814]" captionTargetPageId="53" captionText="Figs 178–188. Live specimens. Aetana omayan group. — 178–179. A. abadae Huber, sp. nov., ♂ and ♀ with eggsac from Twin Lakes, Negros. — 180–181. A. omayan Huber, 2005, ♂ and ♀ with eggsac from Baguio, Luzon. — 182–183. A. manansalai Huber, sp. nov., ♂ and ♀ from Mt. Banahaw, Luzon. — 184–185. A. lozadae Huber, sp. nov., ♂♂ from Mt. Isarog, Luzon. — 186–188. Female prosomata, showing stridulatory plates (arrows), in A. abadae Huber, sp. nov. (186), A. omayan Huber, 2005 (187) and A. manansalai Huber, sp. nov. (188)." figureDoi="http://doi.org/10.5281/zenodo.3829580" httpUri="https://zenodo.org/record/3829580/files/figure.png" pageId="5" pageNumber="6">Figs 186–188</figureCitation>
) against abdomen (indistinct hairless area frontally). Epigynum weakly to heavily sclerotized, sometimes with scape of variable length and shape (
<figureCitation id="130245B4603AFF909222FC1F40D1FC1D" box="[1339,1432,926,953]" captionStart="Figs 26–29" captionStartId="14.[227,279,1932,1958]" captionTargetBox="[227,1433,346,1885]" captionTargetId="figure@14.[227,1437,346,1890]" captionTargetPageId="14" captionText="Figs 26–29. Aetana libjo Huber, sp. nov. 26–27. Left male palp, prolateral and retrolateral views (asterisk: retrolateral process of bulb). 28–29. Cleared female genitalia, ventral and dorsal views (asterisk: median membranous structure). b = genital bulb; p = procursus; rt = retrolateral trichobothrium. Scale lines: 26–27 = 0.5 mm; 28–29 = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3829531" httpUri="https://zenodo.org/record/3829531/files/figure.png" pageId="5" pageNumber="6">Figs 28</figureCitation>
, 
<figureCitation id="130245B4603AFF9097FAFC43444DFC79" box="[227,260,962,989]" captionStart="Figs 57–61" captionStartId="22.[227,279,1966,1992]" captionTargetBox="[227,1432,469,1920]" captionTargetId="figure@22.[227,1437,469,1922]" captionTargetPageId="22" captionText="Figs 57–61. Aetana kiukoki Huber, sp. nov. 57–58. Left male palp, prolateral and retrolateral views. 59. Male prosoma, oblique frontal view. 60–61. Cleared female genitalia, ventral and dorsal views. b = genital bulb; e = embolus; p = procursus. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829541" httpUri="https://zenodo.org/record/3829541/files/figure.png" pageId="5" pageNumber="6">60</figureCitation>
), never with external pair of pockets in anterior epigynal plate but in 
<taxonomicName id="4C3922B2603AFF90934DFC434197FC78" authorityName="Huber" authorityYear="2005" box="[1108,1246,962,988]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kinabalu">
<emphasis id="B94D8523603AFF90934DFC434197FC78" box="[1108,1246,962,988]" italics="true" pageId="5" pageNumber="6">A. kinabalu</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3922B2603AFF909205FC4340D5FC78" authorityName="Huber" authorityYear="2005" box="[1308,1436,962,988]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="omayan">
<emphasis id="B94D8523603AFF909205FC4340D5FC78" box="[1308,1436,962,988]" italics="true" pageId="5" pageNumber="6">A. omayan</emphasis>
</taxonomicName>
groups sometimes with internal sclerotized pockets originating from ventral wall of uterus externus (
<figureCitation id="130245B4603AFF9097F2FB89441CFB86" box="[235,341,1032,1058]" captionStart="Figs 111–112" captionStartId="37.[227,279,1997,2023]" captionTargetBox="[239,1430,1527,1927]" captionTargetId="figure@37.[227,1437,1519,1939]" captionTargetPageId="37" captionText="Figs 111–112. Aetana kinabalu Huber, 2005. Cleared female genitalia, ventral and dorsal views (arrow in Fig. 112 points at membranous pocket). ip = internal sclerotized pocket. Scale line: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829560" httpUri="https://zenodo.org/record/3829560/files/figure.png" pageId="5" pageNumber="6">Figs 111</figureCitation>
, 
<figureCitation id="130245B4603AFF90967AFB894487FB86" box="[355,462,1032,1058]" captionStart="Figs 189–194. — 189–193" captionStartId="55.[227,279,1894,1920]" captionTargetBox="[227,1436,450,1840]" captionTargetId="figure@55.[227,1437,450,1847]" captionTargetPageId="55" captionText="Figs 189–194. — 189–193. Aetana abadae Huber, sp. nov. 189–190. Left male palp, prolateral and retrolateral views. 191. Male prosoma, oblique frontal view. 192–193. Cleared female genitalia, ventral and dorsal views. — 194. A. omayan Huber, 2005, cleared female genitalia, dorsal view. Arrows point at membranous pockets. ip = internal sclerotized pocket; rp = retrolatero-ventral process; vl = ventral lamina. Scale lines: 0.5 mm (female genitalia at same scale)." figureDoi="http://doi.org/10.5281/zenodo.3829582" httpUri="https://zenodo.org/record/3829582/files/figure.png" pageId="5" pageNumber="6">192–194</figureCitation>
), and in 
<taxonomicName id="4C3922B2603AFF909520FB8847F0FB86" authorityName="Huber" authorityYear="2005" box="[569,697,1033,1059]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="omayan">
<emphasis id="B94D8523603AFF909520FB8847F0FB86" box="[569,697,1033,1059]" italics="true" pageId="5" pageNumber="6">A. omayan</emphasis>
</taxonomicName>
group with pair of membranous external pockets in posterior epigynal area (
<figureCitation id="130245B4603AFF909696FBAA477DFBE1" box="[399,564,1067,1093]" captionStart="Figs 189–194. — 189–193" captionStartId="55.[227,279,1894,1920]" captionTargetBox="[227,1436,450,1840]" captionTargetId="figure@55.[227,1437,450,1847]" captionTargetPageId="55" captionText="Figs 189–194. — 189–193. Aetana abadae Huber, sp. nov. 189–190. Left male palp, prolateral and retrolateral views. 191. Male prosoma, oblique frontal view. 192–193. Cleared female genitalia, ventral and dorsal views. — 194. A. omayan Huber, 2005, cleared female genitalia, dorsal view. Arrows point at membranous pockets. ip = internal sclerotized pocket; rp = retrolatero-ventral process; vl = ventral lamina. Scale lines: 0.5 mm (female genitalia at same scale)." figureDoi="http://doi.org/10.5281/zenodo.3829582" httpUri="https://zenodo.org/record/3829582/files/figure.png" pageId="5" pageNumber="6">Figs 192, 194</figureCitation>
, 
<figureCitation id="130245B4603AFF909526FBAA4739FBE1" box="[575,624,1067,1093]" captionStart="Figs 219–223" captionStartId="60.[227,279,1932,1958]" captionTargetBox="[227,1437,378,1887]" captionTargetId="figure@60.[227,1437,378,1889]" captionTargetPageId="60" captionText="Figs 219–223. Aetana manansalai Huber, sp. nov. 219–220. Left male palp, prolateral and retrolateral views. 221. Male prosoma, oblique frontal view. 222–223. Cleared female genitalia, ventral and dorsal views. Arrow points at membranous pocket. rp = retrolatero-ventral process; vl = ventral lamina. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829590" httpUri="https://zenodo.org/record/3829590/files/figure.png" pageId="5" pageNumber="6">222</figureCitation>
). Internal genitalia with pair of pore plates, in 
<taxonomicName id="4C3922B2603AFF909397FBAD407EFBE1" authority="Huber" authorityName="Huber" authorityYear="2015" box="[1166,1335,1067,1094]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="libjo" status="sp. nov.">
<emphasis id="B94D8523603AFF909397FBAD41ACFBE1" box="[1166,1253,1067,1094]" italics="true" pageId="5" pageNumber="6">A. libjo</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E3858603AFF90925BFBAD40D4FBE2" box="[1346,1437,1068,1094]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C3922B2603AFF90960DFBCE474AFBCC" authority="Huber" authorityName="Huber" authorityYear="2015" box="[276,515,1102,1129]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="baganihan" status="sp. nov.">
<emphasis id="B94D8523603AFF90960DFBCE44F8FBCD" box="[276,433,1103,1129]" italics="true" pageId="5" pageNumber="6">A. baganihan</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E3858603AFF909517FBCE472EFBCD" box="[526,615,1103,1129]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
with unique median membranous structure (
<figureCitation id="130245B4603AFF909376FBCF4180FBCD" box="[1135,1225,1102,1129]" captionStart="Figs 26–29" captionStartId="14.[227,279,1932,1958]" captionTargetBox="[227,1433,346,1885]" captionTargetId="figure@14.[227,1437,346,1890]" captionTargetPageId="14" captionText="Figs 26–29. Aetana libjo Huber, sp. nov. 26–27. Left male palp, prolateral and retrolateral views (asterisk: retrolateral process of bulb). 28–29. Cleared female genitalia, ventral and dorsal views (asterisk: median membranous structure). b = genital bulb; p = procursus; rt = retrolateral trichobothrium. Scale lines: 26–27 = 0.5 mm; 28–29 = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3829531" httpUri="https://zenodo.org/record/3829531/files/figure.png" pageId="5" pageNumber="6">Figs 29</figureCitation>
, 
<figureCitation id="130245B4603AFF9093CCFBCE41BFFBCD" box="[1237,1270,1103,1129]" captionStart="Figs 38–44. — 38–40" captionStartId="16.[227,279,1906,1932]" captionTargetBox="[227,1432,599,1858]" captionTargetId="figure@16.[227,1437,593,1864]" captionTargetPageId="16" captionText="Figs 38–44. — 38–40. Aetana libjo Huber, sp. nov. 38–39. Left genital bulb in prolateral and retrolateral views. 40. Male chelicerae, frontal view. — 41–44. Aetana baganihan Huber, sp. nov. 41–42. Left genital bulb in prolateral and retrolateral views. 43–44. Cleared female genitalia, ventral and dorsal views (asterisk: median membranous structure). Scale lines: 38–39, 41–44 = 0.5 mm; 40 = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3829535" httpUri="https://zenodo.org/record/3829535/files/figure.png" pageId="5" pageNumber="6">44</figureCitation>
); in 
<taxonomicName id="4C3922B2603AFF909231FBCE40D4FBCD" authorityName="Huber" authorityYear="2015" box="[1320,1437,1103,1129]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kiukoki">
<emphasis id="B94D8523603AFF909231FBCE40D4FBCD" box="[1320,1437,1103,1129]" italics="true" pageId="5" pageNumber="6">A. kiukoki</emphasis>
</taxonomicName>
group with distinctive serrated ridges (
<figureCitation id="130245B4603AFF9095BEFBF3464AFB28" box="[679,771,1138,1164]" captionStart="Figs 57–61" captionStartId="22.[227,279,1966,1992]" captionTargetBox="[227,1432,469,1920]" captionTargetId="figure@22.[227,1437,469,1922]" captionTargetPageId="22" captionText="Figs 57–61. Aetana kiukoki Huber, sp. nov. 57–58. Left male palp, prolateral and retrolateral views. 59. Male prosoma, oblique frontal view. 60–61. Cleared female genitalia, ventral and dorsal views. b = genital bulb; e = embolus; p = procursus. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829541" httpUri="https://zenodo.org/record/3829541/files/figure.png" pageId="5" pageNumber="6">Figs 61</figureCitation>
, 
<figureCitation id="130245B4603AFF909416FBF34666FB28" box="[783,815,1138,1164]" captionStart="Figs 70–74" captionStartId="26.[227,279,1967,1993]" captionTargetBox="[227,1437,565,1925]" captionTargetId="figure@26.[227,1437,565,1925]" captionTargetPageId="26" captionText="Figs 70–74. Aetana paragua Huber, sp. nov. 70–71. Left male palp, prolateral and retrolateral views. 72. Male prosoma, frontal view. 73–74. Cleared female genitalia, ventral and dorsal views (arrows point at serrated ridges). rt = retrolateral trichobothrium. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829545" httpUri="https://zenodo.org/record/3829545/files/figure.png" pageId="5" pageNumber="6">73</figureCitation>
, 
<figureCitation id="130245B4603AFF909422FBF34615FB28" box="[827,860,1138,1164]" captionStart="Figs 75–79" captionStartId="28.[227,279,1966,1992]" captionTargetBox="[230,1434,356,1921]" captionTargetId="figure@28.[227,1437,353,1922]" captionTargetPageId="28" captionText="Figs 75–79. Aetana loboc Huber, sp. nov. 75–76. Left male palp, prolateral and retrolateral views. 77. Male prosoma, frontal view. 78–79. Cleared female genitalia, ventral and dorsal views. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829547" httpUri="https://zenodo.org/record/3829547/files/figure.png" pageId="5" pageNumber="6">79</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF4609B9603AFF9097F8F8514723F7AA" ID-DOI="http://doi.org/10.5281/zenodo.3829515" ID-Zenodo-Dep="3829515" httpUri="https://zenodo.org/record/3829515/files/figure.png" pageId="5" pageNumber="6" startId="5.[225,268,2000,2026]" targetBox="[225,1436,1257,1958]" targetPageId="5">
<paragraph id="8B865931603AFF9097F8F8514723F7AA" blockId="5.[225,1436,2000,2062]" pageId="5" pageNumber="6">
<emphasis id="B94D8523603AFF9097F8F8514478F84E" bold="true" box="[225,305,2000,2026]" pageId="5" pageNumber="6">Fig. 2.</emphasis>
Known distribution of 
<taxonomicName id="4C3922B2603AFF909552F85047D7F84F" authorityName="Huber" authorityYear="2005" box="[587,670,2001,2027]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B94D8523603AFF909552F85047D7F84F" box="[587,670,2001,2027]" italics="true" pageId="5" pageNumber="6">Aetana</emphasis>
</taxonomicName>
. Shown are also three undescribed species that are known from poorly preserved specimens only.
</paragraph>
</caption>
<subSubSection id="C3230ABA6039FF9297FAFEB0406DFE58" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="discussion">
<paragraph id="8B8659316039FF9397FAFEB04718FEEE" blockId="6.[227,593,304,331]" box="[227,593,304,331]" pageId="6" pageNumber="7">
<heading id="D0CEEE5D6039FF9397FAFEB04718FEEE" bold="true" box="[227,593,304,331]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B94D85236039FF9397FAFEB04718FEEE" bold="true" box="[227,593,304,331]" pageId="6" pageNumber="7">Monophyly and relationships</emphasis>
</heading>
</paragraph>
<paragraph id="8B8659316039FF9397FAFEDE46D4FDE8" blockId="6.[227,1436,351,589]" pageId="6" pageNumber="7">
Morphologically, the monophyly of 
<taxonomicName id="4C3922B26039FF939585FEE147A6FEDE" authorityName="Huber" authorityYear="2005" box="[668,751,352,378]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B94D85236039FF939585FEE147A6FEDE" box="[668,751,352,378]" italics="true" pageId="6" pageNumber="7">Aetana</emphasis>
</taxonomicName>
appears weakly supported. Only two characters support this node (
<figureCitation id="130245B46039FF93967DFE0344E5FE39" box="[356,428,386,413]" captionStart="Fig" captionStartId="2.[227,270,1966,1992]" captionTargetBox="[227,1416,466,1905]" captionTargetId="figure@2.[227,1436,461,1923]" captionTargetPageId="2" captionText="Fig. 1. Single most parsimonious cladogram resulting from analyses of the matrix in Appendix 1. For characters and character states see Appendix 3. Only unambiguous character changes are shown. See Cladistic analysis section for further details." figureDoi="http://doi.org/10.5281/zenodo.3829513" httpUri="https://zenodo.org/record/3829513/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
), both of them without homoplasy within the taxa included in the matrix but with considerable homoplasy among more distant 
<taxonomicName id="4C3922B26039FF9395E5FE274639FE64" authorityName="C.L.Koch" authorityYear="1850" box="[764,880,422,448]" class="Arachnida" family="Pholcidae" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subFamily" subFamily="Pholcinae">Pholcinae</taxonomicName>
relatives: (1) the very proximal position of the retrolateral trichobothrium on the leg tibiae (char. 30); and (2) the presence of curved hairs on leg tibiae and/or metatarsi (char. 31). However, preliminary analyses of molecular data (including many more potential close relatives) (A. Valdez-Mondragón, D. Dimitrov, B.A. Huber, unpubl. data) consistently support the monophyly of 
<taxonomicName id="4C3922B26039FF939503FDB24724FDE9" authorityName="Huber" authorityYear="2005" box="[538,621,563,589]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B94D85236039FF939503FDB24724FDE9" box="[538,621,563,589]" italics="true" pageId="6" pageNumber="7">Aetana</emphasis>
</taxonomicName>
with high support values.
</paragraph>
<paragraph id="8B8659316039FF9397FAFDF54104FC9A" blockId="6.[227,1437,628,831]" pageId="6" pageNumber="7">
The sister group of 
<taxonomicName id="4C3922B26039FF9396D0FDF54755FD2A" authorityName="Huber" authorityYear="2005" box="[457,540,628,654]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B94D85236039FF9396D0FDF54755FD2A" box="[457,540,628,654]" italics="true" pageId="6" pageNumber="7">Aetana</emphasis>
</taxonomicName>
appears much better supported by morphology. Four characters suggest that three genera together (
<taxonomicName id="4C3922B26039FF9396E9FD16471BFD15" authorityName="Huber" authorityYear="2005" box="[496,594,663,689]" class="Arachnida" family="Pholcidae" genus="Khorata" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B94D85236039FF9396E9FD16471BFD15" box="[496,594,663,689]" italics="true" pageId="6" pageNumber="7">Khorata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3922B26039FF93957BFD16478DFD15" authorityName="Huber" authorityYear="2005" box="[610,708,663,689]" class="Arachnida" family="Pholcidae" genus="Savarna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B94D85236039FF93957BFD16478DFD15" box="[610,708,663,689]" italics="true" pageId="6" pageNumber="7">Savarna</emphasis>
</taxonomicName>
, and ‘Gen.n. Borneo’) are sister to 
<taxonomicName id="4C3922B26039FF939377FD164188FD15" authorityName="Huber" authorityYear="2005" box="[1134,1217,663,689]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B94D85236039FF939377FD164188FD15" box="[1134,1217,663,689]" italics="true" pageId="6" pageNumber="7">Aetana</emphasis>
</taxonomicName>
. This, however, is in conflict with our preliminary molecular data that suggest a closer relationship between 
<taxonomicName id="4C3922B26039FF93920FFD3A4020FD71" authorityName="Huber" authorityYear="2005" box="[1302,1385,699,725]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B94D85236039FF93920FFD3A4020FD71" box="[1302,1385,699,725]" italics="true" pageId="6" pageNumber="7">Aetana</emphasis>
</taxonomicName>
and East Asian 
<taxonomicName id="4C3922B26039FF939675FD5C4745FD53" authorityName="Hentz" authorityYear="1841" box="[364,524,733,759]" class="Arachnida" family="Pholcidae" genus="Spermophora" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B94D85236039FF939675FD5C4745FD53" box="[364,524,733,759]" italics="true" pageId="6" pageNumber="7">Spermophora</emphasis>
</taxonomicName>
(incl. 
<taxonomicName id="4C3922B26039FF939545FD5F4797FD53" authorityName="Simon" authorityYear="1892" box="[604,734,733,760]" class="Arachnida" family="Pholcidae" genus="Spermophora" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="estebani">
<emphasis id="B94D85236039FF939545FD5F4797FD53" box="[604,734,733,760]" italics="true" pageId="6" pageNumber="7">S. estebani</emphasis>
</taxonomicName>
) than between 
<taxonomicName id="4C3922B26039FF939480FD5F46A5FD5C" authorityName="Huber" authorityYear="2005" box="[921,1004,734,760]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B94D85236039FF939480FD5F46A5FD5C" box="[921,1004,734,760]" italics="true" pageId="6" pageNumber="7">Aetana</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3922B26039FF939333FD5C41C5FD53" authorityName="Huber" authorityYear="2005" box="[1066,1164,733,759]" class="Arachnida" family="Pholcidae" genus="Khorata" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B94D85236039FF939333FD5C41C5FD53" box="[1066,1164,733,759]" italics="true" pageId="6" pageNumber="7">Khorata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3922B26039FF939384FD5F41B6FD5C" authorityName="Huber" authorityYear="2005" box="[1181,1279,734,760]" class="Arachnida" family="Pholcidae" genus="Savarna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B94D85236039FF939384FD5F41B6FD5C" box="[1181,1279,734,760]" italics="true" pageId="6" pageNumber="7">Savarna</emphasis>
</taxonomicName>
, and ‘Gen.n. Borneo’. Such a close relationship with East Asian 
<taxonomicName id="4C3922B26039FF939458FC8046A8FCBF" authorityName="Hentz" authorityYear="1841" box="[833,993,769,795]" class="Arachnida" family="Pholcidae" genus="Spermophora" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B94D85236039FF939458FC8046A8FCBF" box="[833,993,769,795]" italics="true" pageId="6" pageNumber="7">Spermophora</emphasis>
</taxonomicName>
was actually proposed in the original description (
<bibRefCitation id="EFA824C06039FF93966DFCA5475BFC9B" author="Huber B. A." box="[372,530,804,831]" pageId="6" pageNumber="7" pagination="61 - 172" refId="ref27087" refString="Huber B. A. 2005 a. Revision of the genus Spermophora Hentz in Southeast Asia and on the Pacific Islands, with descriptions of three new genera (Araneae: Pholcidae). Zoologische Mededelingen 79 - 2 (4): 61 - 172." type="journal article" year="2005">Huber 2005a</bibRefCitation>
), at that time without a formal cladistic analysis.
</paragraph>
<caption id="DF4609B96039FF9397FAF85346E6F7B4" ID-DOI="http://doi.org/10.5281/zenodo.3829517" ID-Zenodo-Dep="3829517" httpUri="https://zenodo.org/record/3829517/files/figure.png" pageId="6" pageNumber="7" startId="6.[227,279,2002,2028]" targetBox="[227,1436,883,1950]" targetPageId="6">
<paragraph id="8B8659316039FF9397FAF85340D5F848" blockId="6.[227,1436,2002,2029]" box="[227,1436,2002,2029]" pageId="6" pageNumber="7">
<emphasis id="B94D85236039FF9397FAF853441BF848" bold="true" box="[227,338,2002,2028]" pageId="6" pageNumber="7">Figs 3–4.</emphasis>
Known distributions of the 
<taxonomicName id="4C3922B26039FF939582F8524613F848" authorityName="Huber" authorityYear="2015" box="[667,858,2003,2029]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ocampoi">
<emphasis id="B94D85236039FF939582F8524613F848" box="[667,858,2003,2029]" italics="true" pageId="6" pageNumber="7">Aetana ocampoi</emphasis>
</taxonomicName>
(3) and 
<taxonomicName id="4C3922B26039FF9394A4F852417BF848" authorityName="Huber" authorityYear="2015" box="[957,1074,2002,2029]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kiukoki">
<emphasis id="B94D85236039FF9394A4F852417BF848" box="[957,1074,2002,2029]" italics="true" pageId="6" pageNumber="7">A. kiukoki</emphasis>
</taxonomicName>
(4) groups. The question mark
</paragraph>
<paragraph id="8B8659316039FF9397FAF87446E6F7B4" blockId="6.[227,317,2037,2063]" box="[227,943,2037,2064]" lastBlockId="6.[324,943,2037,2064]" pageId="6" pageNumber="7">
denotes a female specimen assigned tentatively to 
<taxonomicName id="4C3922B26039FF93942AF87746E0F7AB" authorityName="Huber" authorityYear="2015" box="[819,937,2037,2064]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kiukoki">
<emphasis id="B94D85236039FF93942AF87746E0F7AB" box="[819,937,2037,2064]" italics="true" pageId="6" pageNumber="7">A. kiukoki</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B8659316038FF9297FAFEB0406DFE58" blockId="7.[227,1437,305,508]" pageId="7" pageNumber="8">
Within 
<taxonomicName id="4C3922B26038FF929623FEB344C4FEE8" authorityName="Huber" authorityYear="2005" box="[314,397,306,332]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B94D85236038FF929623FEB344C4FEE8" box="[314,397,306,332]" italics="true" pageId="7" pageNumber="8">Aetana</emphasis>
</taxonomicName>
, four species groups receive strong support both from the morphological analysis herein (
<figureCitation id="130245B46038FF9297F2FED54478FECA" box="[235,305,340,366]" captionStart="Fig" captionStartId="2.[227,270,1966,1992]" captionTargetBox="[227,1416,466,1905]" captionTargetId="figure@2.[227,1436,461,1923]" captionTargetPageId="2" captionText="Fig. 1. Single most parsimonious cladogram resulting from analyses of the matrix in Appendix 1. For characters and character states see Appendix 3. Only unambiguous character changes are shown. See Cladistic analysis section for further details." figureDoi="http://doi.org/10.5281/zenodo.3829513" httpUri="https://zenodo.org/record/3829513/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
) and from our preliminary molecular data. Conflict exists regarding relationships among these groups. The present analysis supports a sister group relationship between the 
<taxonomicName id="4C3922B26038FF9293B9FEF9402FFE36" authorityName="Huber" authorityYear="2005" box="[1184,1382,376,402]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kinabalu">
<emphasis id="B94D85236038FF9293B9FEF9402FFE36" box="[1184,1382,376,402]" italics="true" pageId="7" pageNumber="8">Aetana kinabalu</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3922B26038FF9297FAFE1A4428FE11" authorityName="Huber" authorityYear="2005" box="[227,353,411,437]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="omayan">
<emphasis id="B94D85236038FF9297FAFE1A4428FE11" box="[227,353,411,437]" italics="true" pageId="7" pageNumber="8">A. omayan</emphasis>
</taxonomicName>
groups, and at least one of the three characters supporting this node is a unique feature (the long retrolateral membranous process on the procursus; char. 21). Our preliminary molecular data do not strongly support any sister group relationships among the four species groups within 
<taxonomicName id="4C3922B26038FF9293D3FE634054FE58" authorityName="Huber" authorityYear="2005" box="[1226,1309,482,508]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B94D85236038FF9293D3FE634054FE58" box="[1226,1309,482,508]" italics="true" pageId="7" pageNumber="8">Aetana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3230ABA6038FF9D97FAFDA240D4FE35" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="biology_ecology">
<paragraph id="8B8659316038FF9297FAFDA244EDFD98" blockId="7.[227,420,546,573]" box="[227,420,546,573]" pageId="7" pageNumber="8">
<heading id="D0CEEE5D6038FF9297FAFDA244EDFD98" bold="true" box="[227,420,546,573]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B94D85236038FF9297FAFDA244EDFD98" bold="true" box="[227,420,546,573]" pageId="7" pageNumber="8">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="8B8659316038FF9D97FAFDD340D4FE35" blockId="7.[227,1437,593,655]" lastBlockId="8.[227,1437,305,402]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Most species were collected in forests, from well protected spaces close to the ground, under rocks and logs, in small holes and cavities. Few species occur higher in the vegetation, even in places directly reached by the sun, most notably the closely related 
<taxonomicName id="4C3922B26037FF9D944CFEB34149FEEF" authority="Huber" authorityName="Huber" authorityYear="2015" box="[853,1024,305,332]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="libjo" status="sp. nov.">
<emphasis id="B94D85236037FF9D944CFEB346E4FEEF" box="[853,941,305,332]" italics="true" pageId="8" pageNumber="9">A. libjo</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E38586037FF9D9314FEB34120FEE8" box="[1037,1129,306,332]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C3922B26037FF9D93BCFEB340D1FEEF" authority="Huber" authorityName="Huber" authorityYear="2015" box="[1189,1432,305,332]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="baganihan" status="sp. nov.">
<emphasis id="B94D85236037FF9D93BCFEB3400DFEEF" box="[1189,1348,305,332]" italics="true" pageId="8" pageNumber="9">A. baganihan</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E38586037FF9D97FAFED44474FECB" box="[227,317,341,367]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
and the putatively close relatives 
<taxonomicName id="4C3922B26037FF9D95D5FED4461DFECA" authorityName="Huber" authorityYear="2005" box="[716,852,340,367]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="kinabalu">
<emphasis id="B94D85236037FF9D95D5FED4461DFECA" box="[716,852,340,367]" italics="true" pageId="8" pageNumber="9">A. kinabalu</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3922B26037FF9D9495FED44104FECA" authority="Huber" authorityName="Huber" authorityYear="2015" box="[908,1101,340,367]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lambir" status="sp. nov.">
<emphasis id="B94D85236037FF9D9495FED446B2FECA" box="[908,1019,340,367]" italics="true" pageId="8" pageNumber="9">A. lambir</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E38586037FF9D9340FED441FDFECB" box="[1113,1204,341,367]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
An exception is the 
<typeStatus id="5482E7936037FF9D97FAFEF9445FFE36" box="[227,278,376,402]" pageId="8" pageNumber="9">type</typeStatus>
species 
<taxonomicName id="4C3922B26037FF9D9661FEF944BCFE35" authorityName="Huber" authorityYear="2005" box="[376,501,376,402]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="omayan">
<emphasis id="B94D85236037FF9D9661FEF944BCFE35" box="[376,501,376,402]" italics="true" pageId="8" pageNumber="9">A. omayan</emphasis>
</taxonomicName>
, which was mainly collected in a cave but also among rocks in a treeless ravine.
</paragraph>
</subSubSection>
<caption id="DF4609B96038FF9297FAF8754028F7AB" ID-DOI="http://doi.org/10.5281/zenodo.3829519" ID-Zenodo-Dep="3829519" box="[227,1377,2036,2063]" httpUri="https://zenodo.org/record/3829519/files/figure.png" pageId="7" pageNumber="8" startId="7.[227,270,2036,2062]" targetBox="[227,1436,710,1988]" targetPageId="7">
<paragraph id="8B8659316038FF9297FAF8754028F7AB" blockId="7.[227,1377,2036,2063]" box="[227,1377,2036,2063]" pageId="7" pageNumber="8">
<emphasis id="B94D85236038FF9297FAF8754479F7AA" bold="true" box="[227,304,2036,2062]" pageId="7" pageNumber="8">Fig. 5.</emphasis>
Known distributions of the 
<taxonomicName id="4C3922B26038FF929563F87446E5F7AA" authority="(Borneo)" baseAuthorityName="Borneo" box="[634,940,2036,2063]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kinabalu">
<emphasis id="B94D85236038FF929563F8744675F7AA" box="[634,828,2036,2063]" italics="true" pageId="7" pageNumber="8">Aetana kinabalu</emphasis>
(Borneo)
</taxonomicName>
and 
<taxonomicName id="4C3922B26038FF9294FCF874404BF7AB" authority="(Philippines)" authorityName="Huber" authorityYear="2005" baseAuthorityName="Philippines" box="[997,1282,2036,2063]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="omayan">
<emphasis id="B94D85236038FF9294FCF874412AF7AA" box="[997,1123,2037,2063]" italics="true" pageId="7" pageNumber="8">A. omayan</emphasis>
(Philippines)
</taxonomicName>
groups.
</paragraph>
</caption>
<subSubSection id="C3230ABA6037FF9D97FAFE384103FBBE" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8B8659316037FF9D97FAFE38400AFDBD" blockId="8.[227,1437,441,538]" pageId="8" pageNumber="9">
All species seem to build simple domed sheet webs (like most other pholcids studied), but the sheets are unusually strongly domed in 
<taxonomicName id="4C3922B26037FF9D9561FE5C4661FE52" ID-CoL="65BCQ" authority="Huber" authorityName="Huber" authorityYear="2015" box="[632,808,476,503]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="libjo" status="sp. nov.">
<emphasis id="B94D85236037FF9D9561FE5C479BFE52" box="[632,722,476,503]" italics="true" pageId="8" pageNumber="9">A. libjo</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E38586037FF9D9421FE5C46DEFE53" box="[824,919,477,503]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C3922B26037FF9D94CEFE5C4186FE52" authority="Huber" authorityName="Huber" authorityYear="2015" box="[983,1231,476,503]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="baganihan" status="sp. nov.">
<emphasis id="B94D85236037FF9D94CEFE5C4130FE52" box="[983,1145,476,503]" italics="true" pageId="8" pageNumber="9">A. baganihan</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E38586037FF9D93C6FE5C4009FE53" box="[1247,1344,477,503]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, and in 
<taxonomicName id="4C3922B26037FF9D97FAFD814423FDBD" box="[227,362,511,538]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="kinabalu">
<emphasis id="B94D85236037FF9D97FAFD814423FDBD" box="[227,362,511,538]" italics="true" pageId="8" pageNumber="9">A. kinabalu</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3922B26037FF9D96BAFD81472DFDBD" authority="Huber" authorityName="Huber" authorityYear="2015" box="[419,612,511,538]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lambir" status="sp. nov.">
<emphasis id="B94D85236037FF9D96BAFD81475BFDBD" box="[419,530,511,538]" italics="true" pageId="8" pageNumber="9">A. lambir</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E38586037FF9D9569FD814782FDBE" box="[624,715,512,538]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
a second sheet occurs a few cm under the main sheet.
</paragraph>
<paragraph id="8B8659316037FF9D97FAFDC046BFFD61" blockId="8.[227,1436,577,709]" pageId="8" pageNumber="9">
In six cases, two species were found to share a locality, sometimes in different microhabitats (e.g., at Baganihan: 
<taxonomicName id="4C3922B26037FF9D9676FDE5477DFDDA" authority="Huber" authorityName="Huber" authorityYear="2015" box="[367,564,612,638]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="kiukoki" status="sp. nov.">
<emphasis id="B94D85236037FF9D9676FDE544AAFDDA" box="[367,483,612,638]" italics="true" pageId="8" pageNumber="9">A. kiukoki</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E38586037FF9D9526FDE447D1FDDB" box="[575,664,613,639]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
near the ground; 
<taxonomicName id="4C3922B26037FF9D947BFDE54118FDDA" authority="Huber" authorityName="Huber" authorityYear="2015" box="[866,1105,612,638]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="baganihan" status="sp. nov.">
<emphasis id="B94D85236037FF9D947BFDE546B6FDDA" box="[866,1023,612,638]" italics="true" pageId="8" pageNumber="9">A. baganihan</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E38586037FF9D9342FDE441FCFDDB" box="[1115,1205,613,639]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
among vegetation), sometimes in what seemed to be identical microhabitats (e.g., at Mt. Banahaw: 
<taxonomicName id="4C3922B26037FF9D9380FD0940D1FD05" authority="Huber" authorityName="Huber" authorityYear="2015" box="[1177,1432,647,674]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="manansalai" status="sp. nov.">
<emphasis id="B94D85236037FF9D9380FD09400DFD05" box="[1177,1348,647,674]" italics="true" pageId="8" pageNumber="9">A. manansalai</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E38586037FF9D97FAFD2A4474FD61" box="[227,317,683,709]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C3922B26037FF9D966FFD2A471DFD60" authority="Huber" authorityName="Huber" authorityYear="2015" box="[374,596,682,709]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="banahaw" status="sp. nov.">
<emphasis id="B94D85236037FF9D966FFD2A474AFD60" box="[374,515,682,709]" italics="true" pageId="8" pageNumber="9">A. banahaw</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E38586037FF9D9579FD2A47F4FD61" box="[608,701,683,709]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, both close to the ground).
</paragraph>
<paragraph id="8B8659316037FF9D97FAFD6D4621FCE8" blockId="8.[227,1436,748,845]" pageId="8" pageNumber="9">
In most species, males and females were often found together sharing a web. This was never observed in 
<taxonomicName id="4C3922B26037FF9D9618FC9144E3FC8D" authority="Huber" authorityName="Huber" authorityYear="2015" box="[257,426,783,810]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="libjo" status="sp. nov.">
<emphasis id="B94D85236037FF9D9618FC914411FC8D" box="[257,344,783,810]" italics="true" pageId="8" pageNumber="9">A. libjo</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E38586037FF9D96AFFC914758FC8E" box="[438,529,784,810]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C3922B26037FF9D9550FC914673FC8D" authority="Huber" authorityName="Huber" authorityYear="2015" box="[585,826,783,810]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="baganihan" status="sp. nov.">
<emphasis id="B94D85236037FF9D9550FC9147AEFC8D" box="[585,743,783,810]" italics="true" pageId="8" pageNumber="9">A. baganihan</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E38586037FF9D945FFC9146EAFC8E" box="[838,931,784,810]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, where males and females were sometimes observed very close to each other but in separate webs.
</paragraph>
<paragraph id="8B8659316037FF9D97FAFCF54103FBBE" blockId="8.[227,1437,884,1051]" pageId="8" pageNumber="9">
When disturbed, 
<taxonomicName id="4C3922B26037FF9D96B2FCF544B7FC2A" authorityName="Huber" authorityYear="2005" box="[427,510,884,910]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94D85236037FF9D96B2FCF544B7FC2A" box="[427,510,884,910]" italics="true" pageId="8" pageNumber="9">Aetana</emphasis>
</taxonomicName>
spiders tend to run toward the periphery of the web rather than to vibrate in the ‘typical’ pholcid way. Some do then vibrate vigorously for a very short time before becoming motionless and pressing their body against the substrate; others stop vibrating and start to gently move the abdomen in circles (
<taxonomicName id="4C3922B26037FF9D967DFC5F475AFC53" authority="Huber" authorityName="Huber" authorityYear="2015" box="[356,531,989,1016]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="libjo" status="sp. nov.">
<emphasis id="B94D85236037FF9D967DFC5F44F7FC53" box="[356,446,989,1016]" italics="true" pageId="8" pageNumber="9">A. libjo</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E38586037FF9D953BFC5F4736FC5C" box="[546,639,990,1016]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C3922B26037FF9D95A7FC5F46FDFC53" authority="Huber" authorityName="Huber" authorityYear="2015" box="[702,948,989,1016]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="baganihan" status="sp. nov.">
<emphasis id="B94D85236037FF9D95A7FC5F4617FC53" box="[702,862,989,1016]" italics="true" pageId="8" pageNumber="9">A. baganihan</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E38586037FF9D94DAFC5F416CFC5C" box="[963,1061,990,1016]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
). The closely related 
<taxonomicName id="4C3922B26037FF9D9233FC5F4467FBBE" authority="Huber" authorityName="Huber" authorityYear="2015" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="poring" status="sp. nov.">
<emphasis id="B94D85236037FF9D9233FC5F40D5FC53" box="[1322,1436,990,1016]" italics="true" pageId="8" pageNumber="9">A. poring</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E38586037FF9D9623FB8044DDFBBF" box="[314,404,1025,1051]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C3922B26037FF9D96D4FB8047CAFBBE" authority="Huber" authorityName="Huber" authorityYear="2015" box="[461,643,1024,1051]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="indah" status="sp. nov.">
<emphasis id="B94D85236037FF9D96D4FB804779FBBF" box="[461,560,1025,1051]" italics="true" pageId="8" pageNumber="9">A. indah</emphasis>
Huber
</taxonomicName>
, 
<taxonomicNameLabel id="A27E38586037FF9D9597FB8047A0FBBF" box="[654,745,1025,1051]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
barely reacted to disturbance.
</paragraph>
</subSubSection>
<subSubSection id="C3230ABA6037FF9D97FAFBC346D4FB50" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8B8659316037FF9D97FAFBC344CAFBF8" blockId="8.[227,387,1090,1116]" box="[227,387,1090,1116]" pageId="8" pageNumber="9">
<heading id="D0CEEE5D6037FF9D97FAFBC344CAFBF8" bold="true" box="[227,387,1090,1116]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B94D85236037FF9D97FAFBC344CAFBF8" bold="true" box="[227,387,1090,1116]" pageId="8" pageNumber="9">Composition</emphasis>
</heading>
</paragraph>
<paragraph id="8B8659316037FF9D97FAFBF146D4FB50" blockId="8.[227,1436,1136,1269]" pageId="8" pageNumber="9">
The genus now includes 18 described species. The high species turnover in the 
<collectingCountry id="F32E19A16037FF9D93D9FBF1400DFB2E" box="[1216,1348,1136,1162]" name="Philippines" pageId="8" pageNumber="9">Philippines</collectingCountry>
and in northern Borneo, together with the wide distribution of one species group (
<taxonomicName id="4C3922B26037FF9D935DFB154189FB0A" authorityName="Huber" authorityYear="2005" box="[1092,1216,1172,1198]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="omayan">
<emphasis id="B94D85236037FF9D935DFB154189FB0A" box="[1092,1216,1172,1198]" italics="true" pageId="8" pageNumber="9">A. omayan</emphasis>
</taxonomicName>
group; 
<collectingCountry id="F32E19A16037FF9D9201FB1540D5FB0A" box="[1304,1436,1172,1198]" name="Philippines" pageId="8" pageNumber="9">Philippines</collectingCountry>
to 
<collectingCountry id="F32E19A16037FF9D961BFB3644C1FB76" box="[258,392,1207,1234]" name="Fiji" pageId="8" pageNumber="9">Fiji Islands</collectingCountry>
) and the huge sampling gaps in eastern 
<collectingCountry id="F32E19A16037FF9D947CFB364693FB75" box="[869,986,1207,1233]" name="Indonesia" pageId="8" pageNumber="9">Indonesia</collectingCountry>
, New 
<collectingCountry id="F32E19A16037FF9D933EFB364136FB75" box="[1063,1151,1207,1233]" name="Guinea-Bissau" pageId="8" pageNumber="9">Guinea</collectingCountry>
, and east to 
<collectingCountry id="F32E19A16037FF9D920AFB364075FB75" box="[1299,1340,1207,1233]" name="Fiji" pageId="8" pageNumber="9">Fiji</collectingCountry>
suggest that at least several dozen further species are likely to exist.
</paragraph>
</subSubSection>
<subSubSection id="C3230ABA6037FF9D97FAFA9A47F4F99C" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8B8659316037FF9D97FAFA9A4435FA91" blockId="8.[227,380,1307,1333]" box="[227,380,1307,1333]" pageId="8" pageNumber="9">
<heading id="D0CEEE5D6037FF9D97FAFA9A4435FA91" bold="true" box="[227,380,1307,1333]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B94D85236037FF9D97FAFA9A4435FA91" bold="true" box="[227,380,1307,1333]" pageId="8" pageNumber="9">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B8659316037FF9D97FAFACB47F4F99C" blockId="8.[227,1436,1354,1592]" pageId="8" pageNumber="9">
Ranging from northern Borneo and the 
<collectingCountry id="F32E19A16037FF9D95A3FACB4677FAC0" box="[698,830,1354,1380]" name="Philippines" pageId="8" pageNumber="9">Philippines</collectingCountry>
to 
<collectingCountry id="F32E19A16037FF9D947FFACB46C6FAC0" box="[870,911,1354,1380]" name="Fiji" pageId="8" pageNumber="9">Fiji</collectingCountry>
(
<figureCitation id="130245B46037FF9D94B9FACB46A0FAC0" box="[928,1001,1354,1380]" captionStart="Fig" captionStartId="5.[225,268,2000,2026]" captionTargetBox="[225,1436,1257,1958]" captionTargetId="figure@5.[227,1436,1257,1957]" captionTargetPageId="5" captionText="Fig. 2. Known distribution of Aetana. Shown are also three undescribed species that are known from poorly preserved specimens only." figureDoi="http://doi.org/10.5281/zenodo.3829515" httpUri="https://zenodo.org/record/3829515/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
). Most gaps and missing records are probably due to lack of adequate sampling, but the absence of records from two areas might reflect real absence. First, our intensive collecting in western 
<collectingCountry id="F32E19A16037FF9D9461FA114694FA0E" box="[888,989,1424,1450]" name="Malaysia" pageId="8" pageNumber="9">Sarawak</collectingCountry>
(at seven localities ranging from the Pueh foothills in the west to Niah in the east) did not result in a single specimen of 
<taxonomicName id="4C3922B26037FF9D93A8FA35404DFA6A" authorityName="Huber" authorityYear="2005" box="[1201,1284,1460,1486]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94D85236037FF9D93A8FA35404DFA6A" box="[1201,1284,1460,1486]" italics="true" pageId="8" pageNumber="9">Aetana</emphasis>
</taxonomicName>
, while many specimens were collected at all eight localities in eastern 
<collectingCountry id="F32E19A16037FF9D948CFA5646B3FA55" box="[917,1018,1495,1521]" name="Malaysia" pageId="8" pageNumber="9">Sarawak</collectingCountry>
(east of Niah) and 
<collectingRegion id="49FD97D36037FF9D93FFFA564078FA55" box="[1254,1329,1495,1521]" country="Malaysia" name="Sabah" pageId="8" pageNumber="9">Sabah</collectingRegion>
. Second, northern 
<collectingCountry id="F32E19A16037FF9D9654FA7B44F0F9B0" box="[333,441,1530,1556]" name="Australia" pageId="8" pageNumber="9">Australia</collectingCountry>
is relatively well sampled, but a revision of all the material available in collections (Huber 2001) did not reveal any 
<taxonomicName id="4C3922B26037FF9D957AF99F47FFF99C" authorityName="Huber" authorityYear="2005" box="[611,694,1566,1592]" class="Arachnida" family="Pholcidae" genus="Aetana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94D85236037FF9D957AF99F47FFF99C" box="[611,694,1566,1592]" italics="true" pageId="8" pageNumber="9">Aetana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>