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<mods:title id="CCB575D50C9916295396CEEAB161DBF0">Revision of the African cichlid fish genus Ctenochromis (Teleostei, Cichliformes), including a description of the new genus Shuja from Lake Tanganyika and the new species Ctenochromis scatebra from northern Tanzania</mods:title>
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<mods:namePart id="9703A6737CEA7ABEB2CCD43B167A54DA">Genner, Martin J.</mods:namePart>
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<mods:affiliation id="B0DC865FA921FD3FFFDB74A37D137164">School of Biological Sciences, University of Bristol, Life Sciences Building, 24 Tyndall Avenue, Bristol, BS 8 1 TQ, United Kingdom.</mods:affiliation>
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<mods:namePart id="1BAC42C69F34336CA750E8FE97677C97">Hsu, Ling-Lan</mods:namePart>
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<mods:affiliation id="DEC31A8CA66389DF3D57D9B7FD79142C">School of Biological Sciences, University of Bristol, Life Sciences Building, 24 Tyndall Avenue, Bristol, BS 8 1 TQ, United Kingdom.</mods:affiliation>
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<mods:affiliation id="B27B0D10E73D357A02B037E8EACE3495">School of Biological Sciences, University of Bristol, Life Sciences Building, 24 Tyndall Avenue, Bristol, BS 8 1 TQ, United Kingdom.</mods:affiliation>
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<mods:affiliation id="358EB30FEF6E5C6408CB1B52C336853C">Department of Biological and Marine Sciences, University of Hull, Hull, HU 5 7 RX, United Kingdom.</mods:affiliation>
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<mods:namePart id="7BD55E21B5B4D6F7543825B8129DDC6A">Saxon, Andrew D.</mods:namePart>
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<mods:affiliation id="272FEA3CA8718BB317ACB7A672652916">Tanzania Fisheries Research Institute (TAFIRI) P. O. Box 9750, Dar es Salaam, Tanzania.</mods:affiliation>
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<mods:affiliation id="AF92586B2AC5B91275F31FF1C1AD866B">Tanzania Fisheries Research Institute (TAFIRI) P. O. Box 9750, Dar es Salaam, Tanzania.</mods:affiliation>
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<mods:affiliation id="EA4F4A08248821D360BA401ABE432B74">School of Biological Sciences, Bangor University, Bangor, Gwynedd, LL 57 2 UW, United Kingdom.</mods:affiliation>
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<mods:title id="FC2EA90B2CC6D76BA6850CBDE069B0C3">European Journal of Taxonomy</mods:title>
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<mods:date id="674F302E5BABD4486629E945BAEEC54D">2022</mods:date>
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<taxonomicName id="4C397ACC6032EC2B5259FD4A664F3184" authority="Gunther, 1894" authorityName="Gunther" authorityYear="1894" box="[560,881,681,708]" class="Actinopterygii" family="Cichlidae" genus="Shuja" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="41" phylum="Chordata" rank="species" species="horei" status="gen. et comb. nov.">
<emphasis id="B94DDD5D6032EC2B5259FD4A67FE3183" bold="true" box="[560,704,681,708]" italics="true" pageId="18" pageNumber="41">Shuja horei</emphasis>
<bibRefCitation id="EFA87CBE6032EC2B52A1FD4A664F3184" author="Gunther A." box="[712,881,681,707]" pageId="18" pageNumber="41" pagination="628 - 632" refId="ref15026" refString="Gunther A. 1894. Descriptions of the reptiles and fishes collected by Mr. E. Coode-Hore on Lake Tanganyika. Proceedings of the Zoological Society of London 1893: 628 - 632." type="journal article" year="1894">Günther, 1894</bibRefCitation>
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<figureCitation id="13021DCA6032EC2B5374FD2F665C31A0" box="[797,866,716,743]" captionStart="Fig" captionStartId="18.[227,270,1856,1882]" captionTargetBox="[227,1391,1213,1815]" captionTargetId="figure-345@18.[227,1436,1213,1815]" captionTargetPageId="18" captionText="Fig. 5. Shuja horei gen. et comb. nov.a. Illustration from the original type specimen (Günther 1894).b. A freshly caught specimen collected in 2016 from the Malagarasi River, Ilagala (BMNH 2021.7.15.14). c. Radiographs of the type series (syntypes) from the Natural History Museum (BMNH 1889.1.30.13– 15). Note the prognathous jaw that distinguishes Shuja Genner, Ngatunga &amp; Turner gen. nov. from other genera within the Tropheini Poll, 1986. Radiographs from the Natural History Museum, London (Creative Commons Attribution License (CC BY 4.0)). Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6554365" httpUri="https://zenodo.org/record/6554365/files/figure.png" pageId="18" pageNumber="41">Fig. 5</figureCitation>
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<paragraph id="8B86014F6032EC2B508AFCEF67E03060" blockId="18.[227,796,780,873]" box="[227,734,780,807]" pageId="18" pageNumber="41">
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<treatmentCitation id="0A98275E6032EC2B508AFCEF67453061" author="Gunther A." box="[227,635,780,806]" page="630" pageId="18" pageNumber="41" year="1894">
<taxonomicName id="4C397ACC6032EC2B508AFCEF67453061" authority="Gunther, 1894: 630" authorityName="Gunther" authorityPageNumber="630" authorityYear="1894" box="[227,635,780,806]" class="Actinopterygii" family="Pomacentridae" genus="Chromis" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="41" phylum="Chordata" rank="species" species="horei">
<emphasis id="B94DDD5D6032EC2B508AFCEF64B23061" box="[227,396,780,806]" italics="true" pageId="18" pageNumber="41">Chromis horei</emphasis>
<bibRefCitation id="EFA87CBE6032EC2B51FDFCEF67453061" author="Gunther A." box="[404,635,780,806]" pageId="18" pageNumber="41" pagination="628 - 632" refId="ref15026" refString="Gunther A. 1894. Descriptions of the reptiles and fishes collected by Mr. E. Coode-Hore on Lake Tanganyika. Proceedings of the Zoological Society of London 1893: 628 - 632." type="journal article" year="1894">Günther, 1894: 630</bibRefCitation>
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, pl. 58a.
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<treatmentCitation id="0A98275E6032EC2B508AFCCE67BE300F" author="Boulenger G. A." box="[227,640,813,840]" page="122" pageId="18" pageNumber="41" year="1899">
<taxonomicName id="4C397ACC6032EC2B508AFCCE67BE300F" ID-CoL="56WCN" authority="Boulenger, 1899: 122" authorityName="Boulenger" authorityPageNumber="122" authorityYear="1899" box="[227,640,813,840]" class="Actinopterygii" family="Cichlidae" genus="Tilapia" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="41" phylum="Chordata" rank="species" species="horii">
<emphasis id="B94DDD5D6032EC2B508AFCCE64463000" box="[227,376,813,839]" italics="true" pageId="18" pageNumber="41">Tilapia horii</emphasis>
<bibRefCitation id="EFA87CBE6032EC2B51E9FCCE67BE300F" author="Boulenger G. A." box="[384,640,813,840]" pageId="18" pageNumber="41" pagination="98 - 144" refId="ref14171" refString="Boulenger G. A. 1899. A revision of the African and Syrian fishes of the family Cichlidae - Part II. 1. Proceedings of the Zoological Society of London 67: 98 - 144. https: // doi. org / 10.1111 / j. 1469 - 7998.1899. tb 06852. x" type="journal article" year="1899">Boulenger, 1899: 122</bibRefCitation>
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.
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<treatmentCitationGroup id="AB2926616032EC2B508AFCAC6622302E" box="[227,796,847,873]" pageId="18" pageNumber="41">
<treatmentCitation id="0A98275E6032EC2B508AFCAC6626302E" author="Borodin N. A." box="[227,792,847,873]" page="29" pageId="18" pageNumber="41" year="1936">
<taxonomicName id="4C397ACC6032EC2B508AFCAC6626302E" ID-CoL="7KTCW" authority="Borodin, 1936: 29" authorityName="Borodin" authorityPageNumber="29" authorityYear="1936" box="[227,792,847,873]" class="Actinopterygii" family="Cichlidae" genus="Tilapia" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="41" phylum="Chordata" rank="subSpecies" species="fasciata" subSpecies="tanganaicae">
<emphasis id="B94DDD5D6032EC2B508AFCAC6708302E" box="[227,566,847,873]" italics="true" pageId="18" pageNumber="41">Tilapia fasciata tanganaicae</emphasis>
<bibRefCitation id="EFA87CBE6032EC2B5257FCAC6626302E" author="Borodin N. A." box="[574,792,847,873]" pageId="18" pageNumber="41" pagination="1 - 34" refId="ref14139" refString="Borodin N. A. 1936. On a collection of freshwater fishes from Lakes Nyasa, Tanganyika and Viktoria in Central Africa. Zoologische Jahrbucher 68: 1 - 34." type="journal article" year="1936">Borodin, 1936: 29</bibRefCitation>
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.
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<treatmentCitationGroup id="AB2926616032EC2B508AFC6D60A230EF" box="[227,1436,910,937]" pageId="18" pageNumber="41">
<taxonomicName id="4C397ACC6032EC2B508AFC6D64F130EF" ID-CoL="3JLRR" baseAuthorityName="Takahashi" baseAuthorityYear="2003" box="[227,463,910,936]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="41" phylum="Chordata" rank="species" species="horei">
<emphasis id="B94DDD5D6032EC2B508AFC6D64F130EF" box="[227,463,910,936]" italics="true" pageId="18" pageNumber="41">Haplochromis horei</emphasis>
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–
<treatmentCitation id="0A98275E6032EC2B5185FC6C67E030EF" author="Trewavas E." box="[492,734,910,937]" page="242" pageId="18" pageNumber="41" year="1946">
<bibRefCitation id="EFA87CBE6032EC2B5185FC6C67E030EF" author="Trewavas E." box="[492,734,910,937]" pageId="18" pageNumber="41" pagination="240 - 246" refId="ref16992" refString="Trewavas E. 1946. The types of African cichlid fishes described by Borodin in 1931 and 1936, and of two species described by Boulenger in 1901. Proceedings of the Zoological Society of London 116: 240 - 246. https: // doi. org / 10.1111 / j. 1096 - 3642.1946. tb 00121. x" type="journal article" year="1946">Trewavas 1946: 242</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0A98275E6032EC2B5365FC6C617F30EF" author="Van Oijen M. J. P. &amp; Snoeks J. &amp; Skelton P. H. &amp; Marchal C. &amp; Teugels G. G." box="[780,1089,910,937]" page="131" pageId="18" pageNumber="41" year="1991">
<bibRefCitation id="EFA87CBE6032EC2B5365FC6C617F30EF" author="Van Oijen M. J. P. &amp; Snoeks J. &amp; Skelton P. H. &amp; Marchal C. &amp; Teugels G. G." box="[780,1089,910,937]" pageId="18" pageNumber="41" pagination="100 - 184" refId="ref17133" refString="Van Oijen M. J. P., Snoeks J., Skelton P. H., Marchal C. &amp; Teugels G. G. 1991. Haplochromis. In: Daget J., Gosse J. P., Teugels G. G. &amp; Thys Van Den Audenaerde D. F. E. (eds) Check List of the Freshwater Fishes of Africa = Catalogue des poissons d'eau douce d'Afrique. CLOFFA 4: 100 - 184. Musee royal de" type="journal article" year="1991">
van Oijen
<emphasis id="B94DDD5D6032EC2B53EFFC6C66FF30EF" box="[902,961,910,936]" italics="true" pageId="18" pageNumber="41">et al.</emphasis>
1991: 131
</bibRefCitation>
</treatmentCitation>
. —
<bibRefCitation id="EFA87CBE6032EC2B5419FC6D60A230EF" author="Wamuini Lunkayilakio S. &amp; Vreven E. J." box="[1136,1436,910,936]" pageId="18" pageNumber="41" pagination="279 - 287" refId="ref17435" refString="Wamuini Lunkayilakio S. &amp; Vreven E. J. 2010. ' Haplochromis ' snoeksi, a new species from the Inkisi River basin, Lower Congo (Perciformes: Cichlidae). Ichthyological Explorations of Freshwaters 21: 279 - 287." type="journal article" year="2010">Wamuini Lunkayilakio &amp;</bibRefCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B86014F6032EC2B5179FC5360A23769" blockId="18.[227,1436,910,1105]" pageId="18" pageNumber="41">
Vreven 2010: 280.
<taxonomicName id="4C397ACC6032EC2B508AFC32670830AC" ID-CoL="7KTCW" authorityName="Borodin" authorityYear="1936" box="[227,566,977,1003]" class="Actinopterygii" family="Cichlidae" genus="Tilapia" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="41" phylum="Chordata" rank="subSpecies" species="fasciata" subSpecies="tanganaicae">
<emphasis id="B94DDD5D6032EC2B508AFC32670830AC" box="[227,566,977,1003]" italics="true" pageId="18" pageNumber="41">Tilapia fasciata tanganaicae</emphasis>
</taxonomicName>
–
<bibRefCitation id="EFA87CBE6032EC2B523AFC32667830AC" author="Trewavas E." box="[595,838,977,1003]" pageId="18" pageNumber="41" pagination="240 - 246" refId="ref16992" refString="Trewavas E. 1946. The types of African cichlid fishes described by Borodin in 1931 and 1936, and of two species described by Boulenger in 1901. Proceedings of the Zoological Society of London 116: 240 - 246. https: // doi. org / 10.1111 / j. 1096 - 3642.1946. tb 00121. x" type="journal article" year="1946">Trewavas 1946: 242</bibRefCitation>
.
<taxonomicName id="4C397ACC6032EC2B508AFC1164F4374B" ID-CoL="56WCN" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[227,458,1010,1036]" class="Actinopterygii" family="Cichlidae" genus="Ctenochromis" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="41" phylum="Chordata" rank="species" species="horii">
<emphasis id="B94DDD5D6032EC2B508AFC1164F4374B" box="[227,458,1010,1036]" italics="true" pageId="18" pageNumber="41">Ctenochromis horii</emphasis>
</taxonomicName>
–
<bibRefCitation id="EFA87CBE6032EC2B518EFC116612374B" author="Greenwood P. H." box="[487,812,1010,1036]" pageId="18" pageNumber="41" pagination="265 - 322" refId="ref14894" refString="Greenwood P. H. 1979. Towards a phyletic classification of the ' genus' Haplochromis (Pisces, Cichlidae) and related taxa. Part I. Bulletin of the British Museum of Natural History (Zoology) 35: 265 - 322. https: // doi. org / 10.5962 / bhl. part. 20455" type="journal article" year="1979">Greenwood 1979: 289–290</bibRefCitation>
.
<taxonomicName id="4C397ACC6032EC2B508AFBF764F53769" ID-CoL="325KX" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[227,459,1044,1070]" class="Actinopterygii" family="Cichlidae" genus="Ctenochromis" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="41" phylum="Chordata" rank="species" species="horei">
<emphasis id="B94DDD5D6032EC2B508AFBF764F53769" box="[227,459,1044,1070]" italics="true" pageId="18" pageNumber="41">Ctenochromis horei</emphasis>
</taxonomicName>
–
<bibRefCitation id="EFA87CBE6032EC2B518EFBF7663F3769" author="Poll M. &amp; Gosse J. - P." box="[487,769,1043,1070]" pageId="18" pageNumber="41" pagination="1 - 324" refId="ref16011" refString="Poll M. &amp; Gosse J. - P. 1995. Genera des poissons d'eau douce de l'Afrique. Memoire de la Classe des Sciences. Academie royale de Belgique 9: 1 - 324." type="journal article" year="1995">Poll &amp; Gosse 1995: 229</bibRefCitation>
. —
<bibRefCitation id="EFA87CBE6032EC2B5347FBF7617F3769" author="De Vos L. D. G. &amp; Seegers L. &amp; Taverne L. &amp; Thys van den Audenaerde D. F. E." box="[814,1089,1044,1070]" pageId="18" pageNumber="41" refId="ref14561" refString="De Vos L. D. G., Seegers L., Taverne L. &amp; Thys van den Audenaerde D. F. E. 2001. L'ichthyofaune du bassin de la Malagarasi (systeme du Lac Tanganyika): une synthese de la connaissance actuelle. Annales" type="book" year="2001">
De Vos
<emphasis id="B94DDD5D6032EC2B53E1FBF766FC3769" box="[904,962,1044,1070]" italics="true" pageId="18" pageNumber="41">et al.</emphasis>
2001: 130
</bibRefCitation>
, 132. —
<bibRefCitation id="EFA87CBE6032EC2B54C1FBF760A23769" author="Kullander S. &amp; Roberts T." box="[1192,1436,1044,1070]" pageId="18" pageNumber="41" pagination="355 - 376" refId="ref15468" refString="Kullander S. &amp; Roberts T. 2011. Out of Lake Tanganyika: endemic lake fishes inhabit rapids of the Lukuga River. Ichthyological Explorations of Freshwaters 22: 355 - 376." type="journal article" year="2011">Kullander &amp; Roberts</bibRefCitation>
</paragraph>
<paragraph id="8B86014F6032EC2B5166FBD5665E3717" blockId="18.[227,1436,910,1105]" box="[271,864,1078,1105]" pageId="18" pageNumber="41">
2011: 362, 369. —
<bibRefCitation id="EFA87CBE6032EC2B519BFBD567E83717" author="Konings A." box="[498,726,1078,1105]" pageId="18" pageNumber="41" refId="ref15345" refString="Konings A. 2015. Tanganyika Cichlids in their Natural Habitat. 3 rd edition. Cichlid Press, El Paso, TX." type="book" year="2015">Konings 2015: 223</bibRefCitation>
;
<bibRefCitation id="EFA87CBE6032EC2B528BFBD566633717" author="Konings A." box="[738,861,1078,1104]" pageId="18" pageNumber="41" refId="ref15369" refString="Konings A. 2019. Tanganyika Cichlids in their Natural Habitat. 4 th edition. Cichlid Press, El Paso, TX." type="book" year="2019">2019: 237</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="DF4651C76032EC2B508AF8A3616F3B4C" ID-DOI="http://doi.org/10.5281/zenodo.6554365" ID-Zenodo-Dep="6554365" httpUri="https://zenodo.org/record/6554365/files/figure.png" pageId="18" pageNumber="41" startId="18.[227,270,1856,1882]" targetBox="[227,1391,1213,1815]" targetPageId="18">
<paragraph id="8B86014F6032EC2B508AF8A3616F3B4C" blockId="18.[223,1438,1856,2059]" pageId="18" pageNumber="41">
<emphasis id="B94DDD5D6032EC2B508AF8A36411341C" bold="true" box="[227,303,1856,1883]" pageId="18" pageNumber="41">Fig. 5.</emphasis>
<taxonomicName id="4C397ACC6032EC2B515CF8A26485341C" authorityName="Genner &amp; Hsu &amp; Collins &amp; Smith &amp; Saxon &amp; Shechonge &amp; Ngatunga &amp; Turner" authorityYear="2022" baseAuthorityName="Genner &amp; Hsu &amp; Collins &amp; Smith &amp; Saxon &amp; Shechonge &amp; Ngatunga &amp; Turner" baseAuthorityYear="1894" box="[309,443,1857,1883]" class="Actinopterygii" family="Cichlidae" genus="Shuja" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="41" phylum="Chordata" rank="species" species="horei" status="gen. et comb. nov.">
<emphasis id="B94DDD5D6032EC2B515CF8A26485341C" box="[309,443,1857,1883]" italics="true" pageId="18" pageNumber="41">Shuja horei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E60266032EC2B51A8F8A267AC341C" box="[449,658,1857,1883]" pageId="18" pageNumber="41" rank="species">gen. et comb. nov.</taxonomicNameLabel>
<emphasis id="B94DDD5D6032EC2B52FCF8A2679A341C" bold="true" box="[661,676,1857,1883]" pageId="18" pageNumber="41">a</emphasis>
. Illustration from the original type specimen (
<bibRefCitation id="EFA87CBE6032EC2B54DCF8A26065341C" author="Gunther A." box="[1205,1371,1857,1883]" pageId="18" pageNumber="41" pagination="628 - 632" refId="ref15026" refString="Gunther A. 1894. Descriptions of the reptiles and fishes collected by Mr. E. Coode-Hore on Lake Tanganyika. Proceedings of the Zoological Society of London 1893: 628 - 632." type="journal article" year="1894">Günther 1894</bibRefCitation>
).
<emphasis id="B94DDD5D6032EC2B5502F8A26045341C" bold="true" box="[1387,1403,1857,1883]" pageId="18" pageNumber="41">b</emphasis>
. A freshly caught specimen collected in 2016 from the Malagarasi River, Ilagala (BMNH 2021.7.15.14).
<emphasis id="B94DDD5D6032EC2B508DF86465D134E6" bold="true" box="[228,239,1927,1953]" pageId="18" pageNumber="41">c</emphasis>
. Radiographs of the type series (syntypes) from the Natural History Museum (BMNH 1889.1.30.13– 15). Note the prognathous jaw that distinguishes
<taxonomicName id="4C397ACC6032EC2B5355F84961E13482" authority="Genner, Ngatunga &amp; Turner" authorityName="Genner, Ngatunga &amp; Turner" authorityYear="2022" box="[828,1247,1962,1989]" class="Actinopterygii" family="Cichlidae" genus="Shuja" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="41" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B94DDD5D6032EC2B5355F84966413483" box="[828,895,1962,1988]" italics="true" pageId="18" pageNumber="41">Shuja</emphasis>
Genner, Ngatunga &amp; Turner
</taxonomicName>
<taxonomicNameLabel id="A27E60266032EC2B5483F84860673482" box="[1258,1369,1963,1989]" pageId="18" pageNumber="41" rank="genus">gen. nov.</taxonomicNameLabel>
from other genera within the
<taxonomicName id="4C397ACC6032EC2B5195F82D67CA34A0" authority="Poll, 1986" authorityName="Poll" authorityYear="1986" box="[508,756,1997,2024]" class="Actinopterygii" family="Cichlidae" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="40" phylum="Chordata" rank="tribe" tribe="Tropheini">
Tropheini
<bibRefCitation id="EFA87CBE6032EC2B521EF82D67CA34A0" author="Poll M." box="[631,756,1997,2024]" pageId="18" pageNumber="41" pagination="1 - 163" refId="ref15977" refString="Poll M. 1986. Classification des Cichlidae du lac Tanganika. Tribus, genres et especes. Memoires de l'Academie royale de Belgique, Classe des Sciences 45: 1 - 163." type="journal article" year="1986">Poll, 1986</bibRefCitation>
</taxonomicName>
. Radiographs from the Natural History Museum, London (Creative Commons Attribution License (CC BY 4.0)). Scale bars:
<quantity id="4CC1ACAA6032EC2B539EF81261733B4C" box="[1015,1101,2033,2059]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="41" unit="mm" value="10.0">10 mm</quantity>
.
</paragraph>
</caption>
<subSubSection id="C32352C46033EC2A508AFECD60483163" pageId="19" pageNumber="42" type="materials_examined">
<paragraph id="8B86014F6033EC2A508AFECD64ED320F" blockId="19.[227,467,302,328]" box="[227,467,302,328]" pageId="19" pageNumber="42">
<heading id="D0CEB6236033EC2A508AFECD64ED320F" bold="true" box="[227,467,302,328]" fontSize="11" level="3" pageId="19" pageNumber="42" reason="3">
<emphasis id="B94DDD5D6033EC2A508AFECD64ED320F" bold="true" box="[227,467,302,328]" pageId="19" pageNumber="42">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B86014F6033EC2A5179FEBF676832FA" blockId="19.[227,1436,348,445]" pageId="19" pageNumber="42">
<materialsCitation id="3B510B126033EC2A5179FEBF676D32FA" ID-GBIF-Occurrence="3770149309" collectionCode="BMNH" country="Tanzania" location="Lake Tanganyika" municipality="Lake Tanganyika" pageId="19" pageNumber="42" specimenCode="BMNH 1889.1.30.13, BMNH 1889.1.30.15" specimenCount="3" typeStatus="syntype">
<emphasis id="B94DDD5D6033EC2A5179FEBF64BF3231" bold="true" box="[272,385,348,374]" pageId="19" pageNumber="42">
<typeStatus id="5482BFED6033EC2A5179FEBF64BF3231" box="[272,385,348,374]" pageId="19" pageNumber="42" type="syntype">Syntypes</typeStatus>
</emphasis>
<collectingCountry id="F32E41DF6033EC2A508AFE9D644632DF" box="[227,376,382,408]" name="Tanzania" pageId="19" pageNumber="42">TANZANIA</collectingCountry>
•
<specimenCount id="9D3FCAC66033EC2A51F1FE9D670632DF" box="[408,568,382,408]" pageId="19" pageNumber="42">3 individuals</specimenCount>
;
<locationDeviation id="9A34186D6033EC2A5221FE9C678232DF" box="[584,700,382,409]" pageId="19" pageNumber="42">near Ujiji</locationDeviation>
(
<bibRefCitation id="EFA87CBE6033EC2A52A6FE9D66AB32DE" author="Boulenger G. A." box="[719,917,382,409]" pageId="19" pageNumber="42" refId="ref14228" refString="Boulenger G. A. 1915. Catalogue of the Fresh-Water Fishes of Africa in the British Museum (Natural History). Volume 3. Trustees of the British Museum of Natural History, London." type="book" year="1915">Boulenger 1915</bibRefCitation>
),
<collectingMunicipality id="6BE29B356033EC2A53C7FE9D61BE32DF" box="[942,1152,382,408]" pageId="19" pageNumber="42">Lake Tanganyika</collectingMunicipality>
;
<specimenCode id="DB9FA9346033EC2A54F9FE9C60B032DF" box="[1168,1422,382,409]" pageId="19" pageNumber="42">BMNH 1889.1.30.13</specimenCode>
–
<specimenCode id="DB9FA9346033EC2A508AFE4064E232FA" box="[227,476,418,445]" pageId="19" pageNumber="42">BMNH 1889.1.30.15</specimenCode>
(
<figureCitation id="13021DCA6033EC2A5185FE41677732FA" box="[492,585,418,445]" captionStart="Fig" captionStartId="18.[227,270,1856,1882]" captionTargetBox="[227,1391,1213,1815]" captionTargetId="figure-345@18.[227,1436,1213,1815]" captionTargetPageId="18" captionText="Fig. 5. Shuja horei gen. et comb. nov.a. Illustration from the original type specimen (Günther 1894).b. A freshly caught specimen collected in 2016 from the Malagarasi River, Ilagala (BMNH 2021.7.15.14). c. Radiographs of the type series (syntypes) from the Natural History Museum (BMNH 1889.1.30.13– 15). Note the prognathous jaw that distinguishes Shuja Genner, Ngatunga &amp; Turner gen. nov. from other genera within the Tropheini Poll, 1986. Radiographs from the Natural History Museum, London (Creative Commons Attribution License (CC BY 4.0)). Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6554365" httpUri="https://zenodo.org/record/6554365/files/figure.png" pageId="19" pageNumber="42">Fig. 5C</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B86014F6033EC2A5179FE07673432B9" blockId="19.[227,1398,484,548]" box="[272,522,484,510]" pageId="19" pageNumber="42">
<emphasis id="B94DDD5D6033EC2A5179FE07673432B9" bold="true" box="[272,522,484,510]" pageId="19" pageNumber="42">Additional material</emphasis>
</paragraph>
<paragraph id="8B86014F6033EC2A508AFDEA60483163" blockId="19.[227,1398,484,548]" box="[227,1398,521,548]" pageId="19" pageNumber="42">
<materialsCitation id="3B510B126033EC2A508AFDEA604D3163" ID-GBIF-Occurrence="3770149306" box="[227,1395,521,548]" collectingDate="2016-07-28" collectionCode="BMNH" country="Tanzania" location="Malagarasi River" municipality="Ilagala" pageId="19" pageNumber="42" specimenCode="BMNH 2021.7.15.14" specimenCount="1">
<collectingCountry id="F32E41DF6033EC2A508AFDEA64463164" box="[227,376,521,547]" name="Tanzania" pageId="19" pageNumber="42">TANZANIA</collectingCountry>
•
<specimenCount id="9D3FCAC66033EC2A51E6FDEA671E3164" box="[399,544,521,547]" pageId="19" pageNumber="42">1 individual</specimenCount>
;
<location id="8EE657946033EC2A5245FDEA67C63164" LSID="urn:lsid:plazi:treatment:0390B0596032EC2A5259FD4A679131D2:8EE657946033EC2A5245FDEA67C63164" box="[556,760,521,547]" country="Tanzania" municipality="Ilagala" name="Malagarasi River" pageId="19" pageNumber="42">Malagarasi River</location>
,
<collectingMunicipality id="6BE29B356033EC2A536DFDEA66693164" box="[772,855,521,547]" pageId="19" pageNumber="42">Ilagala</collectingMunicipality>
;
<date id="FF87278F6033EC2A530AFDEA66C93164" box="[867,1015,521,547]" pageId="19" pageNumber="42" value="2016-07-28">
<collectingDate id="EFC3DE676033EC2A530AFDEA66C93164" box="[867,1015,521,547]" pageId="19" pageNumber="42" value="2016-07-28">28 Jul. 2016</collectingDate>
</date>
;
<specimenCode id="DB9FA9346033EC2A546AFDEA61C23163" box="[1027,1276,521,548]" pageId="19" pageNumber="42">BMNH 2021.7.15.14</specimenCode>
(
<figureCitation id="13021DCA6033EC2A5565FDEA60573163" box="[1292,1385,521,548]" captionStart="Fig" captionStartId="18.[227,270,1856,1882]" captionTargetBox="[227,1391,1213,1815]" captionTargetId="figure-345@18.[227,1436,1213,1815]" captionTargetPageId="18" captionText="Fig. 5. Shuja horei gen. et comb. nov.a. Illustration from the original type specimen (Günther 1894).b. A freshly caught specimen collected in 2016 from the Malagarasi River, Ilagala (BMNH 2021.7.15.14). c. Radiographs of the type series (syntypes) from the Natural History Museum (BMNH 1889.1.30.13– 15). Note the prognathous jaw that distinguishes Shuja Genner, Ngatunga &amp; Turner gen. nov. from other genera within the Tropheini Poll, 1986. Radiographs from the Natural History Museum, London (Creative Commons Attribution License (CC BY 4.0)). Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6554365" httpUri="https://zenodo.org/record/6554365/files/figure.png" pageId="19" pageNumber="42">Fig. 5B</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32352C46033EC2A508AFDA8679131D2" box="[227,687,587,661]" pageId="19" pageNumber="42" type="distribution">
<paragraph id="8B86014F6033EC2A508AFDA864423122" blockId="19.[227,380,587,613]" box="[227,380,587,613]" pageId="19" pageNumber="42">
<heading id="D0CEB6236033EC2A508AFDA864423122" bold="true" box="[227,380,587,613]" fontSize="11" level="3" pageId="19" pageNumber="42" reason="3">
<emphasis id="B94DDD5D6033EC2A508AFDA864423122" bold="true" box="[227,380,587,613]" pageId="19" pageNumber="42">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B86014F6033EC2A508AFD98679131D2" blockId="19.[227,687,635,661]" box="[227,687,635,661]" pageId="19" pageNumber="42">Lake Tanganyika and proximate rivers.</paragraph>
</subSubSection>
</treatment>
</document>
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