577 lines
73 KiB
XML
577 lines
73 KiB
XML
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<mods:title id="2D364611919F59BF7F060846AE46FA50">From a pair to a dozen: the piscivorous species of Haplochromis (Cichlidae) from the Lake Edward system</mods:title>
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<mods:namePart id="31A15A2B8A6B2AB7B883DABCD5935230">Vranken, Nathan</mods:namePart>
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<mods:affiliation id="BA709820D666D4F3E101D1E42E75FF10">Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium. & Royal Museum for Central Africa, Biology department, Section Vertebrates, Leuvensesteenweg 13, 3080 Tervuren, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="0A6C68496CCEAB72F19C0D3EE62B563A" type="email">nathan.vranken@kuleuven.be</mods:nameIdentifier>
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<mods:namePart id="A98C2C0C085976C49C610002D2A1C0AF">Steenberge, Maarten Van</mods:namePart>
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<mods:affiliation id="C92772B73E3C81F0B63CB399DBDBE1A6">Royal Belgian Institute of Natural Sciences, Operational Directorate Taxonomy and Phylogeny, Vautierstraat 29, 1000 Brussels, Belgium. & Masaryk University, Department of Botany and Zoology, Kotlářská 2, 611 37 Brno, Czech Republic.</mods:affiliation>
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<mods:nameIdentifier id="1E949DDF26A99A2A8A7CAF4B7D1A1B92" type="email">mvansteenberge@naturalsciences.be</mods:nameIdentifier>
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<mods:namePart id="2E6EE75A80157E74FA566A54A11C4C57">Heylen, Annelies</mods:namePart>
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<mods:affiliation id="22DF4FC21EE90A55E81B9EC574913A04">KU Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="D4EFA3FEB895A1B56A3DBF994213AF73" type="email">annelies.heylen@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="4CE220228F9BF1F55295BC8667384BCF">Decru, Eva</mods:namePart>
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<mods:affiliation id="F58C252EAE79816C3D9513DBA098A1B8">KU Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium.</mods:affiliation>
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<mods:affiliation id="0578DCAECDDAAB2447DBF1B79BE25722">Royal Museum for Central Africa, Biology department, Section Vertebrates, Leuvensesteenweg 13, 3080 Tervuren, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="66CBD073108859DE5F81F9A8FBB05D11" type="email">jos.snoeks@africamuseum.be</mods:nameIdentifier>
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<mods:title id="1DD16AE39D402A127D07B77AE653656A">European Journal of Taxonomy</mods:title>
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<treatment id="038F87A66121DA72FDB5554024ADFF2B" ID-DOI="http://doi.org/10.5281/zenodo.6484183" ID-GBIF-Taxon="197197505" ID-Zenodo-Dep="6484183" LSID="urn:lsid:zoobank.org:act:18BC8C85-5A02-48F0-8D90-7B069122A09D" httpUri="http://treatment.plazi.org/id/038F87A66121DA72FDB5554024ADFF2B" lastPageId="42" lastPageNumber="43" pageId="38" pageNumber="39">
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<taxonomicName id="4C264D336121DA7EFDB555402543FE87" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[615,874,626,653]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="aquila" status="sp. nov.">
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<emphasis id="B952EAA26121DA7EFDB555402543FE87" bold="true" box="[615,874,626,653]" italics="true" pageId="38" pageNumber="39">Haplochromis aquila</emphasis>
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<taxonomicNameLabel id="A26157D96121DA7EFCA0554125E4FE87" box="[882,973,627,653]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C33C653B6121DA7EFD5A558F2585FEDD" box="[648,940,701,727]" pageId="38" pageNumber="39" type="description">
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<paragraph id="8B9936B06121DA7EFD5A558F2585FEDD" blockId="38.[369,1219,626,727]" box="[648,940,701,727]" pageId="38" pageNumber="39">
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<figureCitation id="131D2A356121DA7EFD5A558F24D9FEDD" box="[648,752,701,727]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="5.[190,234,1822,1848]" captionStartId-1="6.[189,231,1892,1918]" captionTargetBox-0="[193,1398,734,1726]" captionTargetBox-1="[208,1335,272,1830]" captionTargetId-0="figure-217@5.[189,1399,730,1726]" captionTargetId-1="figure-18@6.[208,1335,272,1830]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Fig. 1. The Lake Edward system, consisting of Lakes Edward and George that are connected through the Kazinga Channel. The catch localities of all examined specimens are indicated: Haplochromis sp. ‘latifrons’ (), H. mentatus Regan, 1925 (), H. sp. ‘rex’ (), H. sp. ‘simba’ (▼), H. sp. ‘glaucus’ (), H. sp. ‘aquila’ (), H. sp. ‘kimondo’ (●), H. sp. ‘falcatus’ (), H. sp. ‘curvidens’ (), H. sp. ‘pardus’ (), H. sp. ‘quasimodo’ (▲), and H. squamipinnis Regan, 1921 (█). The type localities of H. mentatus and H. squamipinnis are not shown as the exact locations are unknown." captionText-1="Fig. 2. a. Differences in total number of outer teeth and premaxillary dentition between the macro- and microdontic groups of piscivores from the Lake Edward system.b. Schematic presentation of the dominant male colour patterns in life, in the various groups studied. For H. sp. ‘latifrons’ the female and for H. sp. ‘curvidens’ the non-dominant colour patterns are shown, as those of dominant males are unknown." figureDoi-0="http://doi.org/10.5281/zenodo.6484155" figureDoi-1="http://doi.org/10.5281/zenodo.6484157" httpUri-0="https://zenodo.org/record/6484155/files/figure.png" httpUri-1="https://zenodo.org/record/6484157/files/figure.png" pageId="38" pageNumber="39">Figs 1–2</figureCitation>
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,
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<figureCitation id="131D2A356121DA7EFD2E558F2560FEDD" box="[764,841,701,727]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="39.[189,232,1963,1989]" captionStartId-1="40.[189,232,1892,1918]" captionStartId-2="42.[189,232,1963,1989]" captionTargetBox-0="[189,1398,1249,1831]" captionTargetBox-1="[189,1393,275,1861]" captionTargetBox-2="[189,1397,947,1862]" captionTargetId-0="figure-226@39.[189,1399,1249,1831]" captionTargetId-1="figure-19@40.[188,1393,265,1861]" captionTargetId-2="figure-180@42.[189,1400,947,1864]" captionTargetPageId-0="39" captionTargetPageId-1="40" captionTargetPageId-2="42" captionText-0="Fig. 20. Haplochromis aquila sp. nov., holotype, ♂, 113.6 mm SL (RMCA 2018.008.P.0355). Drawn by N. Vranken." captionText-1="Fig. 21. Haplochromis aquila sp. nov. a. Photograph of preserved holotype (RMCA 2018.008.P.0355; 113.6 mm SL). b. X-ray image of holotype. c–d. Photographs of freshly caught specimens. c. Dominant male, the holotype. d. Female (RMCA 2018.008.P.0352; 108.7 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." captionText-2="Fig. 22. Haplochromis aquila sp. nov. (RMCA 2018.008.P.0356; 122.9 mm SL). a. Dorsal view of the lower pharyngeal jaw. b. Lateral view of the lower pharyngeal jaw." figureDoi-0="http://doi.org/10.5281/zenodo.6484209" figureDoi-1="http://doi.org/10.5281/zenodo.6484211" figureDoi-2="http://doi.org/10.5281/zenodo.6484213" httpUri-0="https://zenodo.org/record/6484209/files/figure.png" httpUri-1="https://zenodo.org/record/6484211/files/figure.png" httpUri-2="https://zenodo.org/record/6484213/files/figure.png" pageId="38" pageNumber="39">20–22</figureCitation>
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;
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<tableCitation id="C6A4030B6121DA7EFC87558F2585FEDD" box="[853,940,701,727]" captionStart="Table 1" captionStartId="9.[182,252,548,576]" captionTargetPageId="9" captionText="Table 1. (continued on the next three pages). Measurements and counts of the piscivorous species of Haplochromis from the Lake Edward system." httpUri="http://table.plazi.org/id/DF596638610EDA51FF6455162158FE4A" pageId="38" pageNumber="39" tableUuid="DF596638610EDA51FF6455162158FE4A">Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C653B6121DA7FFF6F543324EBFDC4" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="diagnosis">
|
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<paragraph id="8B9936B06121DA7EFF6F543327E0FF11" blockId="38.[189,457,769,795]" box="[189,457,769,795]" pageId="38" pageNumber="39">
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<heading id="D0D181DC6121DA7EFF6F543327E0FF11" bold="true" box="[189,457,769,795]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
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<emphasis id="B952EAA26121DA7EFF6F543327E0FF11" bold="true" box="[189,457,769,795]" pageId="38" pageNumber="39">Differential diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9936B06121DA7EFF6F54032735FF9C" blockId="38.[189,1399,817,918]" pageId="38" pageNumber="39">Species with a piscivorous morphology; eye large [ED 30.0–31.5 (mean 30.6) % HL]; outer oral teeth few and large [UOT 25–37 (median 31)]; dominant males light grey with a black head and a bright red anal fin.</paragraph>
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<paragraph id="8B9936B06121DA7EFF6F54F22539F82E" blockId="38.[189,1398,959,1061]" pageId="38" pageNumber="39">
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Amongst piscivorous species from the Lake Edward system,
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<taxonomicName id="4C264D336121DA7EFC5454F225DFFFD3" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[902,1014,959,986]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="aquila" status="sp. nov.">
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<emphasis id="B952EAA26121DA7EFC5454F225B2FFD3" box="[902,923,960,985]" italics="true" pageId="38" pageNumber="39">H</emphasis>
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||
.
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<emphasis id="B952EAA26121DA7EFC78548D25DFFFD3" box="[938,1014,959,985]" italics="true" pageId="38" pageNumber="39">aquila</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A26157D96121DA7EFC2E54F2227FFFD0" box="[1020,1110,960,986]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
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differs from
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<taxonomicName id="4C264D336121DA7EFB3D54F2235FFFD3" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[1263,1398,959,986]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="latifrons" status="sp. nov.">
|
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<emphasis id="B952EAA26121DA7EFB3D54F2232DFFD3" box="[1263,1284,960,985]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
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<emphasis id="B952EAA26121DA7EFAC1548D235FFFD3" box="[1299,1398,959,985]" italics="true" pageId="38" pageNumber="39">latifrons</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26157D96121DA7EFF6F54D72732FFF5" box="[189,283,997,1023]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
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,
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<taxonomicName id="4C264D336121DA7EFEFB54D42790FFF5" authorityName="Regan" authorityYear="1925" box="[297,441,997,1023]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="mentatus">
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||
<emphasis id="B952EAA26121DA7EFEFB54D42717FFF5" box="[297,318,998,1023]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFE9C54D42790FFF5" box="[334,441,998,1023]" italics="true" pageId="38" pageNumber="39">mentatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C264D336121DA7EFE1854D4243DFFF5" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[458,532,997,1023]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="rex" status="sp. nov.">
|
||
<emphasis id="B952EAA26121DA7EFE1854D427F6FFF5" box="[458,479,998,1023]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFE3D54D4243DFFF5" box="[495,532,998,1023]" italics="true" pageId="38" pageNumber="39">rex</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96121DA7EFDCF54D72452FFF5" box="[541,635,997,1023]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C264D336121DA7EFD5A54D424DCFFF5" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[648,757,997,1023]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="simba" status="sp. nov.">
|
||
<emphasis id="B952EAA26121DA7EFD5A54D424B4FFF5" box="[648,669,998,1023]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFD7C54D724DCFFF5" box="[686,757,997,1023]" italics="true" pageId="38" pageNumber="39">simba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96121DA7EFD2C54D72575FFF5" box="[766,860,997,1023]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="4C264D336121DA7EFC4F54D42237FFF5" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[925,1054,997,1023]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="glaucus" status="sp. nov.">
|
||
<emphasis id="B952EAA26121DA7EFC4F54D4259BFFF5" box="[925,946,998,1023]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFC1054D72237FFF5" box="[962,1054,997,1023]" italics="true" pageId="38" pageNumber="39">glaucus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96121DA7EFBF554D722AAFFF5" box="[1063,1155,997,1023]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by a larger eye [ED 30.0–31.5 (30.6) vs 22.2–29.9 (24.4–28.3) % HL].
|
||
</paragraph>
|
||
<paragraph id="8B9936B06121DA7EFF6F537C24FEF8DD" blockId="38.[188,1400,1101,1240]" pageId="38" pageNumber="39">
|
||
It further differs from
|
||
<taxonomicName id="4C264D336121DA7EFE69537C246AF862" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[443,579,1102,1128]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="latifrons" status="sp. nov.">
|
||
<emphasis id="B952EAA26121DA7EFE69537C27F9F86D" box="[443,464,1102,1127]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFE0D537C246AF862" box="[479,579,1102,1128]" italics="true" pageId="38" pageNumber="39">latifrons</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96121DA7EFD95537C2488F862" box="[583,673,1102,1128]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C264D336121DA7EFD05537C254FF86D" authorityName="Regan" authorityYear="1925" box="[727,870,1102,1128]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="mentatus">
|
||
<emphasis id="B952EAA26121DA7EFD05537C24C5F86D" box="[727,748,1102,1127]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFD29537C254FF86D" box="[763,870,1102,1127]" italics="true" pageId="38" pageNumber="39">mentatus</emphasis>
|
||
</taxonomicName>
|
||
by a shorter caudal peduncle [CPL 14.6–15.4 (15.0) vs 15.7–18.0 (16.6–17.0) % SL]; further from
|
||
<taxonomicName id="4C264D336121DA7EFCF95346259AF887" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[811,947,1139,1165]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="latifrons" status="sp. nov.">
|
||
<emphasis id="B952EAA26121DA7EFCF953462569F887" box="[811,832,1140,1165]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFC9D5341259AF887" box="[847,947,1139,1165]" italics="true" pageId="38" pageNumber="39">latifrons</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96121DA7EFC685341223CF887" box="[954,1045,1139,1165]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the absence vs presence of a well-defined mid-lateral band; further from
|
||
<taxonomicName id="4C264D336121DA7EFD0F53AB2547F8B8" authorityName="Regan" authorityYear="1925" box="[733,878,1177,1203]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="mentatus">
|
||
<emphasis id="B952EAA26121DA7EFD0F53AB24DBF8B8" box="[733,754,1177,1202]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFCD153AB2547F8B8" box="[771,878,1177,1202]" italics="true" pageId="38" pageNumber="39">mentatus</emphasis>
|
||
</taxonomicName>
|
||
by dominant males uniformly light grey vs yellow-green with a red anterior part of flank.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06121DA7EFF6F523324B5F96C" blockId="38.[188,1399,1281,1382]" pageId="38" pageNumber="39">
|
||
It further differs from
|
||
<taxonomicName id="4C264D336121DA7EFE0052302436F911" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[466,543,1281,1307]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="rex" status="sp. nov.">
|
||
<emphasis id="B952EAA26121DA7EFE00523027CEF911" box="[466,487,1282,1307]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFE2852302436F911" box="[506,543,1282,1307]" italics="true" pageId="38" pageNumber="39">rex</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96121DA7EFDF9523324A0F911" box="[555,649,1281,1307]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by a shallower lacrimal [LaD 17.0–19.1 (18.3) vs 18.9–22.5 (20.8) % HL] and dominant males light grey with black operculum and snout vs cream-coloured with an orange operculum and a light blue snout.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06121DA7EFF6F52A22358F9F9" blockId="38.[189,1398,1423,1524]" pageId="38" pageNumber="39">
|
||
It further differs from
|
||
<taxonomicName id="4C264D336121DA7EFE6452A22408F9A3" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[438,545,1423,1450]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="simba" status="sp. nov.">
|
||
<emphasis id="B952EAA26121DA7EFE6452A227E2F9A3" box="[438,459,1424,1449]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFE0852BD2408F9A3" box="[474,545,1423,1449]" italics="true" pageId="38" pageNumber="39">simba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96121DA7EFDF652A22454F9A0" box="[548,637,1424,1450]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by a larger number of scales between first anal-fin spine and upper lateral line (ULL-A 12–15, rarely 11 vs 9–11), an absent vs strongly developed mental prominence, and dominant males light grey with a black head vs uniformly yellow with an orange anterior part of flank.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06121DA7EFF6F512C275CFA88" blockId="38.[187,1397,1565,1666]" pageId="38" pageNumber="39">
|
||
It further differs from
|
||
<taxonomicName id="4C264D336121DA7EFE6B512C2410FA3D" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[441,569,1565,1592]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="glaucus" status="sp. nov.">
|
||
<emphasis id="B952EAA26121DA7EFE6B512C27E7FA3D" box="[441,462,1566,1591]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFE0F512F2410FA3D" box="[477,569,1565,1591]" italics="true" pageId="38" pageNumber="39">glaucus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96121DA7EFDEC512C24BEFA32" box="[574,663,1566,1592]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by a broader head [HW 40.1–43.7 (42.0) vs 38.9–40.9 (39.7) % HL] and dominant males light grey with bright red anal and caudal fins vs light blue with crimson anal and caudal fins.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06121DA7EFF6F519E227BFB1A" blockId="38.[189,1399,1707,1808]" pageId="38" pageNumber="39">
|
||
It differs from
|
||
<taxonomicName id="4C264D336121DA7EFEA1519E2428FACF" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[371,513,1707,1734]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="kimondo" status="sp. nov.">
|
||
<emphasis id="B952EAA26121DA7EFEA1519E27A1FACF" box="[371,392,1708,1733]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFE4951992428FACF" box="[411,513,1707,1733]" italics="true" pageId="38" pageNumber="39">kimondo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96121DA7EFDD9519E2445FACC" box="[523,620,1708,1734]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C264D336121DA7EFDA9519E2529FACF" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[635,768,1707,1734]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="falcatus" status="sp. nov.">
|
||
<emphasis id="B952EAA26121DA7EFDA9519E24B9FACF" box="[635,656,1708,1733]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFD7051992529FACF" box="[674,768,1707,1733]" italics="true" pageId="38" pageNumber="39">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96121DA7EFCD9519E2542FACC" box="[779,875,1708,1734]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C264D336121DA7EFCA8519E223DFACF" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[890,1044,1707,1734]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="curvidens" status="sp. nov.">
|
||
<emphasis id="B952EAA26121DA7EFCA8519E25A6FACF" box="[890,911,1708,1733]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFC735199223DFACF" box="[929,1044,1707,1733]" italics="true" pageId="38" pageNumber="39">curvidens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96121DA7EFBCD519E2256FACC" box="[1055,1151,1708,1734]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C264D336121DA7EFB5D519E232EFACF" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[1167,1287,1707,1734]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="pardus" status="sp. nov.">
|
||
<emphasis id="B952EAA26121DA7EFB5D519E228DFACF" box="[1167,1188,1708,1733]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFB645199232EFACF" box="[1206,1287,1707,1733]" italics="true" pageId="38" pageNumber="39">pardus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96121DA7EFAC0519E235BFACC" box="[1298,1394,1708,1734]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C264D336121DA7EFF6F51E3274DFAE0" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[189,356,1744,1771]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="quasimodo" status="sp. nov.">
|
||
<emphasis id="B952EAA26121DA7EFF6F51E326FBFAE0" box="[189,210,1745,1770]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFF3051E2274DFAE0" box="[226,356,1744,1770]" italics="true" pageId="38" pageNumber="39">quasimodo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96121DA7EFEBF51E327E2FAE1" box="[365,459,1745,1771]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="4C264D336121DA7EFDD951E324E4FAE0" authorityName="Regan" authorityYear="1921" box="[523,717,1745,1771]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="squamipinnis">
|
||
<emphasis id="B952EAA26121DA7EFDD951E32436FAE0" box="[523,543,1745,1770]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFDFD51E324E4FAE0" box="[559,717,1745,1770]" italics="true" pageId="38" pageNumber="39">squamipinnis</emphasis>
|
||
</taxonomicName>
|
||
by the combination of large vs small outer oral teeth and smaller number of outer upper jaw teeth [UOT 25–37 (31) vs 39–79 (45–58)].
|
||
</paragraph>
|
||
<paragraph id="8B9936B06121DA7EFF6F50082466FBE3" blockId="38.[188,1399,1849,2025]" pageId="38" pageNumber="39">
|
||
It further differs from
|
||
<taxonomicName id="4C264D336121DA7EFE1750082466FB59" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[453,591,1849,1876]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="kimondo" status="sp. nov.">
|
||
<emphasis id="B952EAA26121DA7EFE17500827F3FB59" box="[453,474,1850,1875]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFE3B500B2466FB59" box="[489,591,1849,1875]" italics="true" pageId="38" pageNumber="39">kimondo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96121DA7EFD8A5008249FFB5E" box="[600,694,1850,1876]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C264D336121DA7EFD105008256DFB59" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[706,836,1849,1876]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="falcatus" status="sp. nov.">
|
||
<emphasis id="B952EAA26121DA7EFD10500824FEFB59" box="[706,727,1850,1875]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFD34500B256DFB59" box="[742,836,1849,1875]" italics="true" pageId="38" pageNumber="39">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96121DA7EFC9C50082580FB5E" box="[846,937,1850,1876]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C264D336121DA7EFC31500822A0FB59" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[995,1161,1849,1876]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="quasimodo" status="sp. nov.">
|
||
<emphasis id="B952EAA26121DA7EFC31500825D1FB59" box="[995,1016,1850,1875]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFBD5500B22A0FB59" box="[1031,1161,1849,1875]" italics="true" pageId="38" pageNumber="39">quasimodo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96121DA7EFB40500822C4FB5E" box="[1170,1261,1850,1876]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by absence vs mostly presence of a well-defined mid-lateral band and dominant males light grey with a black head vs grey dorsally and yellow ventrally, olive-green with an orange-red anterior part of flank, or light grey dorsally and blue-black ventrally; further from
|
||
<taxonomicName id="4C264D336121DA7EFD3B5098255AFBC9" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[745,883,1961,1988]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="kimondo" status="sp. nov.">
|
||
<emphasis id="B952EAA26121DA7EFD3B509824D7FBC9" box="[745,766,1962,1987]" italics="true" pageId="38" pageNumber="39">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26121DA7EFCDF509B255AFBC9" box="[781,883,1961,1987]" italics="true" pageId="38" pageNumber="39">kimondo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96121DA7EFCAB509825FAFBCE" box="[889,979,1962,1988]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by a narrower head [HW 40.1–43.7 (42.0) vs 42.9–48.0 (45.1) % HL].
|
||
</paragraph>
|
||
<paragraph id="8B9936B06120DA7FFF6F56382406FD63" blockId="39.[189,1399,265,362]" pageId="39" pageNumber="40">
|
||
It further differs from
|
||
<taxonomicName id="4C264D336120DA7FFE1756382475FD29" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[453,604,265,292]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="curvidens" status="sp. nov.">
|
||
<emphasis id="B952EAA26120DA7FFE17563827F3FD29" box="[453,474,266,291]" italics="true" pageId="39" pageNumber="40">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26120DA7FFE3B563B2475FD29" box="[489,604,265,291]" italics="true" pageId="39" pageNumber="40">curvidens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96120DA7FFDB7563824E9FD2E" box="[613,704,266,292]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C264D336120DA7FFD2856382546FD29" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[762,879,265,292]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="pardus" status="sp. nov.">
|
||
<emphasis id="B952EAA26120DA7FFD2856382526FD29" box="[762,783,266,291]" italics="true" pageId="39" pageNumber="40">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26120DA7FFCCC563B2546FD29" box="[798,879,265,291]" italics="true" pageId="39" pageNumber="40">pardus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96120DA7FFCAB563825FDFD2E" box="[889,980,266,292]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by a deeper cheek [ChD 26.8–30.8 (28.3) vs 20.8–24.9 (22.5–23.2) % HL]; further from
|
||
<taxonomicName id="4C264D336120DA7FFCE8561F2586FD4C" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[826,943,300,327]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="pardus" status="sp. nov.">
|
||
<emphasis id="B952EAA26120DA7FFCE8561F2567FD4C" box="[826,846,301,326]" italics="true" pageId="39" pageNumber="40">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26120DA7FFC8C561E2586FD4C" box="[862,943,300,326]" italics="true" pageId="39" pageNumber="40">pardus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96120DA7FFC6B561F223DFD4D" box="[953,1044,301,327]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by dominant males light grey vs speckled to uniformly black.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06120DA7FFF6F56A324EBFDC4" blockId="39.[189,1399,400,463]" pageId="39" pageNumber="40">
|
||
It further differs from
|
||
<taxonomicName id="4C264D336120DA7FFE1356A024AAFDA1" authorityName="Regan" authorityYear="1921" box="[449,643,401,427]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="squamipinnis">
|
||
<emphasis id="B952EAA26120DA7FFE1356A027FFFDA1" box="[449,470,402,427]" italics="true" pageId="39" pageNumber="40">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26120DA7FFE3756A024AAFDA1" box="[485,643,402,427]" italics="true" pageId="39" pageNumber="40">squamipinnis</emphasis>
|
||
</taxonomicName>
|
||
by larger eyes [ED 30.0–31.5 (30.6) vs 23.1–29.7 (26.6) % HL] and dominant males light grey vs slate blue.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C653B6120DA7FFF6F56C4237AFE35" box="[189,1363,502,575]" pageId="39" pageNumber="40" type="etymology">
|
||
<paragraph id="8B9936B06120DA7FFF6F56C4276DFE1A" blockId="39.[189,324,502,528]" box="[189,324,502,528]" pageId="39" pageNumber="40">
|
||
<heading id="D0D181DC6120DA7FFF6F56C4276DFE1A" bold="true" box="[189,324,502,528]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
|
||
<emphasis id="B952EAA26120DA7FFF6F56C4276DFE1A" bold="true" box="[189,324,502,528]" pageId="39" pageNumber="40">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B06120DA7FFF6F5516237AFE35" blockId="39.[189,1363,548,575]" box="[189,1363,548,575]" pageId="39" pageNumber="40">
|
||
Specific name from the Latin ‘
|
||
<taxonomicName id="4C264D336120DA7FFDF655162459FE34" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[548,624,548,574]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="aquila">
|
||
<emphasis id="B952EAA26120DA7FFDF655162459FE34" box="[548,624,548,574]" italics="true" pageId="39" pageNumber="40">aquila</emphasis>
|
||
</taxonomicName>
|
||
’ for ‘eagle’; referring to predatory morphology and large eyes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C653B6120DA7FFF6F555727CCF827" pageId="39" pageNumber="40" type="materials_examined">
|
||
<paragraph id="8B9936B06120DA7FFF6F55572783FE75" blockId="39.[189,426,613,639]" box="[189,426,613,639]" pageId="39" pageNumber="40">
|
||
<heading id="D0D181DC6120DA7FFF6F55572783FE75" bold="true" box="[189,426,613,639]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
|
||
<emphasis id="B952EAA26120DA7FFF6F55572783FE75" bold="true" box="[189,426,613,639]" pageId="39" pageNumber="40">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B06120DA7FFF3855A6253FFEFF" blockId="39.[189,1399,660,757]" pageId="39" pageNumber="40">
|
||
<materialsCitation id="3B4E3CED6120DA7FFF3855A6253FFEFF" ID-GBIF-Occurrence="3861203310" collectingDate="2018-01-20" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" latitude="-0.092" location="Kayanja" longLatPrecision="1" longitude="29.758417" pageId="39" pageNumber="40" specimenCode="RMCA 2018.008.P.0355" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="holotype">
|
||
<emphasis id="B952EAA26120DA7FFF3855A62775FEA4" bold="true" box="[234,348,660,686]" pageId="39" pageNumber="40">
|
||
<typeStatus id="549D88126120DA7FFF3855A62775FEA4" box="[234,348,660,686]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F33176206120DA7FFF6F55852713FEDB" box="[189,314,695,721]" name="Uganda" pageId="39" pageNumber="40">UGANDA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D20FD396120DA7FFE85558A2746FED8" box="[343,367,696,722]" pageId="39" pageNumber="40" type="male">♂</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B556120DA7FFEAC558527DFFED8" box="[382,502,695,722]" metricMagnitude="-1" metricUnit="m" metricValue="1.136" pageId="39" pageNumber="40" unit="mm" value="113.6">113.6 mm</quantity>
|
||
SL;
|
||
<collectingRegion id="49E2F8526120DA7FFDE1558524FCFEDB" box="[563,725,695,721]" pageId="39" pageNumber="40">Lake Edward</collectingRegion>
|
||
,
|
||
<location id="8EF9606B6120DA7FFD315585256CFEDB" LSID="urn:lsid:plazi:treatment:038F87A66121DA72FDB5554024ADFF2B:8EF9606B6120DA7FFD315585256CFEDB" box="[739,837,695,721]" country="Uganda" latitude="-0.092" longLatPrecision="1" longitude="29.758417" name="Kayanja" pageId="39" pageNumber="40" stateProvince="Lake Edward">Kayanja</location>
|
||
offshore;
|
||
<geoCoordinate id="EE1250776120DA7FFC1255842270FEDB" box="[960,1113,694,722]" degrees="0" direction="south" minutes="05" orientation="latitude" pageId="39" pageNumber="40" precision="1" seconds="31.2" value="-0.092">0°05′31.2″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1250776120DA7FFBBA5584233AFED8" box="[1128,1299,694,722]" degrees="29" direction="east" minutes="45" orientation="longitude" pageId="39" pageNumber="40" precision="1" seconds="30.3" value="29.758417">29°45′30.3″ E</geoCoordinate>
|
||
;
|
||
<date id="FF9810706120DA7FFAF3558526D2FEFE" pageId="39" pageNumber="40" value="2018-01-20">
|
||
<collectingDate id="EFDCE9986120DA7FFAF3558526D2FEFE" pageId="39" pageNumber="40" value="2018-01-20">20 Jan. 2018</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D353666120DA7FFED555E82785FEFE" box="[263,428,730,756]" pageId="39" pageNumber="40">HIPE3 exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB6120DA7FFE3C55E8253BFEFF" box="[494,786,730,757]" pageId="39" pageNumber="40">RMCA 2018.008.P.0355</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B9936B06120DA7FFF38542E27CCF827" blockId="39.[189,1399,796,1069]" pageId="39" pageNumber="40">
|
||
<materialsCitation id="3B4E3CED6120DA7FFF38542E22BDFF77" ID-GBIF-Occurrence="3861203390" collectingDate="2018-01-18" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" latitude="-0.16802779" location="Katwe" longLatPrecision="1" longitude="29.874277" pageId="39" pageNumber="40" specimenCode="RMCA 2018.008.P.0349, RMCA 2018.008.P.0350, RMCA 2018.008.P.0351" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<emphasis id="B952EAA26120DA7FFF38542E274EFF3C" bold="true" box="[234,359,796,822]" pageId="39" pageNumber="40">
|
||
<typeStatus id="549D88126120DA7FFF38542E274EFF3C" box="[234,359,796,822]" pageId="39" pageNumber="40" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F33176206120DA7FFF6F540D2713FF53" box="[189,314,831,857]" name="Uganda" pageId="39" pageNumber="40">UGANDA</collectingCountry>
|
||
–
|
||
<emphasis id="B952EAA26120DA7FFEB0540D243DFF53" bold="true" box="[354,532,831,857]" pageId="39" pageNumber="40">
|
||
<collectingRegion id="49E2F8526120DA7FFEB0540D243DFF53" box="[354,532,831,857]" pageId="39" pageNumber="40">Lake Edward</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D20FD396120DA7FFDEA540D2440FF50" box="[568,617,831,858]" pageId="39" pageNumber="40" type="female">1 ♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D20FD396120DA7FFDAE540D24ECFF50" box="[636,709,831,858]" pageId="39" pageNumber="40" type="male">2 ♂♂</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B556120DA7FFD05540D25BDFF50" box="[727,916,831,858]" metricMagnitude="-1" metricUnit="m" metricValue="1.008" metricValueMax="1.177" metricValueMin="0.839" pageId="39" pageNumber="40" unit="mm" value="100.8" valueMax="117.7" valueMin="83.9">83.9–117.7 mm</quantity>
|
||
SL; islands near
|
||
<location id="8EF9606B6120DA7FFBA6547222EDFF50" LSID="urn:lsid:plazi:treatment:038F87A66121DA72FDB5554024ADFF2B:8EF9606B6120DA7FFBA6547222EDFF50" box="[1140,1220,832,858]" country="Uganda" latitude="-0.16802779" longLatPrecision="1" longitude="29.874277" name="Katwe" pageId="39" pageNumber="40" stateProvince="Lake Edward">Katwe</location>
|
||
;
|
||
<geoCoordinate id="EE1250776120DA7FFB07540F2358FF53" box="[1237,1393,829,857]" degrees="0" direction="south" minutes="10" orientation="latitude" pageId="39" pageNumber="40" precision="1" seconds="04.9" value="-0.16802779">0°10′04.9″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1250776120DA7FFF6F54532741FF77" box="[189,360,865,893]" degrees="29" direction="east" minutes="52" orientation="longitude" pageId="39" pageNumber="40" precision="1" seconds="27.4" value="29.874277">29°52′27.4″ E</geoCoordinate>
|
||
;
|
||
<date id="FF9810706120DA7FFEA45450243BFF76" box="[374,530,866,893]" pageId="39" pageNumber="40" value="2018-01-18">
|
||
<collectingDate id="EFDCE9986120DA7FFEA45450243BFF76" box="[374,530,866,893]" pageId="39" pageNumber="40" value="2018-01-18">18 Jan. 2018</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D353666120DA7FFDF2545024EEFF76" box="[544,711,866,892]" pageId="39" pageNumber="40">HIPE3 exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB6120DA7FFCDF545022BDFF77" box="[781,1172,866,893]" pageId="39" pageNumber="40">RMCA 2018.008.P.0349 to 0351</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED6120DA7FFB6254502479FFC9" ID-GBIF-Occurrence="3861203331" collectingDate="2018-01-19" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" latitude="-0.16802779" location="Katwe" longLatPrecision="1" longitude="29.874277" pageId="39" pageNumber="40" specimenCode="RMCA 2018.008.P.0352, RMCA 2018.008.P.0353" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<specimenCount id="9D20FD396120DA7FFB62545022F7FF77" box="[1200,1246,866,893]" pageId="39" pageNumber="40" type="female">1 ♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D20FD396120DA7FFB3D54502337FF77" box="[1263,1310,866,893]" pageId="39" pageNumber="40" type="male">1 ♂</specimenCount>
|
||
, 108.7,
|
||
<quantity id="4CDE9B556120DA7FFF6F54B7271EFFAA" box="[189,311,901,928]" metricMagnitude="-1" metricUnit="m" metricValue="1.176" pageId="39" pageNumber="40" unit="mm" value="117.6">117.6 mm</quantity>
|
||
SL; islands near
|
||
<location id="8EF9606B6120DA7FFDC654B4244DFFAA" LSID="urn:lsid:plazi:treatment:038F87A66121DA72FDB5554024ADFF2B:8EF9606B6120DA7FFDC654B4244DFFAA" box="[532,612,902,928]" country="Uganda" latitude="-0.16802779" longLatPrecision="1" longitude="29.874277" name="Katwe" pageId="39" pageNumber="40" stateProvince="Lake Edward">Katwe</location>
|
||
;
|
||
<geoCoordinate id="EE1250776120DA7FFDA654B62526FFAA" box="[628,783,900,928]" degrees="0" direction="south" minutes="10" orientation="latitude" pageId="39" pageNumber="40" precision="1" seconds="04.9" value="-0.16802779">0°10′04.9″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1250776120DA7FFCF254B625E4FFAA" box="[800,973,900,928]" degrees="29" direction="east" minutes="52" orientation="longitude" pageId="39" pageNumber="40" precision="1" seconds="27.4" value="29.874277">29°52′27.4″ E</geoCoordinate>
|
||
;
|
||
<date id="FF9810706120DA7FFC0C54B42257FFAA" box="[990,1150,902,928]" pageId="39" pageNumber="40" value="2018-01-19">
|
||
<collectingDate id="EFDCE9986120DA7FFC0C54B42257FFAA" box="[990,1150,902,928]" pageId="39" pageNumber="40" value="2018-01-19">19 Jan. 2018</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D353666120DA7FFB5D54B42311FF95" box="[1167,1336,901,928]" pageId="39" pageNumber="40">HIPE3 exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB6120DA7FFF6F549B2479FFC9" box="[189,592,936,963]" pageId="39" pageNumber="40">RMCA 2018.008.P.0352 to 0353</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED6120DA7FFDA7549B227CFFED" ID-GBIF-Occurrence="3861203407" collectingDate="2018-01-19" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" latitude="-0.20966667" location="Kazinga Channel" longLatPrecision="1" longitude="29.883749" pageId="39" pageNumber="40" specimenCode="RMCA 2018.008.P.0354" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<specimenCount id="9D20FD396120DA7FFDA7549B2480FFC9" box="[629,681,937,963]" pageId="39" pageNumber="40" type="male">1 ♂</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B556120DA7FFD6E549A2503FFC9" box="[700,810,936,963]" metricMagnitude="-2" metricUnit="m" metricValue="8.559999999999999" pageId="39" pageNumber="40" unit="mm" value="85.6">85.6 mm</quantity>
|
||
SL; mouth of
|
||
<location id="8EF9606B6120DA7FFC3F549B22EBFFC9" LSID="urn:lsid:plazi:treatment:038F87A66121DA72FDB5554024ADFF2B:8EF9606B6120DA7FFC3F549B22EBFFC9" box="[1005,1218,937,963]" country="Uganda" latitude="-0.20966667" longLatPrecision="1" longitude="29.883749" name="Kazinga Channel" pageId="39" pageNumber="40" stateProvince="Lake Edward">Kazinga Channel</location>
|
||
;
|
||
<geoCoordinate id="EE1250776120DA7FFB0654952358FFC9" box="[1236,1393,935,963]" degrees="0" direction="south" minutes="12" orientation="latitude" pageId="39" pageNumber="40" precision="1" seconds="34.8" value="-0.20966667">0°12′34.8″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1250776120DA7FFF6F54F82745FFEC" box="[189,364,970,998]" degrees="29" direction="east" minutes="53" orientation="longitude" pageId="39" pageNumber="40" precision="1" seconds="01.5" value="29.883749">29°53′01.5″ E</geoCoordinate>
|
||
;
|
||
<date id="FF9810706120DA7FFEAC54FE240AFFEC" box="[382,547,972,998]" pageId="39" pageNumber="40" value="2018-01-19">
|
||
<collectingDate id="EFDCE9986120DA7FFEAC54FE240AFFEC" box="[382,547,972,998]" pageId="39" pageNumber="40" value="2018-01-19">19 Jan. 2018</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D353666120DA7FFDE754FE24C8FFEC" box="[565,737,972,998]" pageId="39" pageNumber="40">HIPE3 exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB6120DA7FFCFC54FE227CFFED" box="[814,1109,972,999]" pageId="39" pageNumber="40">RMCA 2018.008.P.0354</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED6120DA7FFBA854FE27C8F827" ID-GBIF-Occurrence="3861203314" collectingDate="2018-01-24" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" latitude="-0.40444446" location="Rwenshama" longLatPrecision="1" longitude="29.773556" pageId="39" pageNumber="40" specimenCode="RMCA 2018.008.P.0356" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<specimenCount id="9D20FD396120DA7FFBA854FE2286FFEC" box="[1146,1199,972,998]" pageId="39" pageNumber="40" type="male">1 ♂</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B556120DA7FFB1354FE2316FFEC" box="[1217,1343,972,998]" metricMagnitude="-1" metricUnit="m" metricValue="1.229" pageId="39" pageNumber="40" unit="mm" value="122.9">122.9 mm</quantity>
|
||
SL;
|
||
<geoCoordinate id="EE1250776120DA7FFF6F54DF2771F803" box="[189,344,1005,1033]" degrees="0" direction="south" minutes="24" orientation="latitude" pageId="39" pageNumber="40" precision="1" seconds="16.0" value="-0.40444446">0°24′16.0″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1250776120DA7FFEBB54DF243FF800" box="[361,534,1005,1034]" degrees="29" direction="east" minutes="46" orientation="longitude" pageId="39" pageNumber="40" precision="1" seconds="24.8" value="29.773556">29°46′24.8″ E</geoCoordinate>
|
||
;
|
||
<date id="FF9810706120DA7FFDF454DD24EEF803" box="[550,711,1007,1034]" pageId="39" pageNumber="40" value="2018-01-24">
|
||
<collectingDate id="EFDCE9986120DA7FFDF454DD24EEF803" box="[550,711,1007,1034]" pageId="39" pageNumber="40" value="2018-01-24">24 Jan. 2018</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D353666120DA7FFD0554DD25A9F803" box="[727,896,1007,1033]" pageId="39" pageNumber="40">HIPE3 exped.</collectorName>
|
||
leg.; bought at
|
||
<location id="8EF9606B6120DA7FFB9554DD22FFF803" LSID="urn:lsid:plazi:treatment:038F87A66121DA72FDB5554024ADFF2B:8EF9606B6120DA7FFB9554DD22FFF803" box="[1095,1238,1007,1033]" country="Uganda" latitude="-0.40444446" longLatPrecision="1" longitude="29.773556" name="Rwenshama" pageId="39" pageNumber="40" stateProvince="Lake Edward">Rwenshama</location>
|
||
landing site;
|
||
<specimenCode id="DB809ECB6120DA7FFF6F532027C8F827" box="[189,481,1042,1069]" pageId="39" pageNumber="40">RMCA 2018.008.P.0356</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5966386120DA7FFF6F50992759FBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484209" ID-Zenodo-Dep="6484209" httpUri="https://zenodo.org/record/6484209/files/figure.png" pageId="39" pageNumber="40" startId="39.[189,232,1963,1989]" targetBox="[189,1398,1249,1831]" targetPageId="39">
|
||
<paragraph id="8B9936B06120DA7FFF6F50992759FBE2" blockId="39.[189,1400,1963,2025]" pageId="39" pageNumber="40">
|
||
<emphasis id="B952EAA26120DA7FFF6F50992735FBCF" bold="true" box="[189,284,1963,1989]" pageId="39" pageNumber="40">Fig. 20.</emphasis>
|
||
<taxonomicName id="4C264D336120DA7FFEF75099240AFBCF" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[293,547,1963,1989]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="aquila" status="sp. nov.">
|
||
<emphasis id="B952EAA26120DA7FFEF75099240AFBCF" box="[293,547,1963,1989]" italics="true" pageId="39" pageNumber="40">Haplochromis aquila</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96120DA7FFDFC509E24A5FBCC" box="[558,652,1964,1990]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂, 113.6 mm SL (RMCA 2018.008.P.0355). Drawn by N. Vranken.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF596638612FDA70FF6F505624E5FBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484211" ID-Zenodo-Dep="6484211" httpUri="https://zenodo.org/record/6484211/files/figure.png" pageId="40" pageNumber="41" startId="40.[189,232,1892,1918]" targetBox="[189,1393,275,1861]" targetPageId="40">
|
||
<paragraph id="8B9936B0612FDA70FF6F505624E5FBE2" blockId="40.[188,1399,1892,2025]" pageId="40" pageNumber="41">
|
||
<emphasis id="B952EAA2612FDA70FF6F50562732FB75" bold="true" box="[189,283,1892,1919]" pageId="40" pageNumber="41">Fig. 21.</emphasis>
|
||
<taxonomicName id="4C264D33612FDA70FEF150572409FB75" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[291,544,1893,1919]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="aquila" status="sp. nov.">
|
||
<emphasis id="B952EAA2612FDA70FEF150572409FB75" box="[291,544,1893,1919]" italics="true" pageId="40" pageNumber="41">Haplochromis aquila</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D9612FDA70FDFB505724ACFB75" box="[553,645,1893,1919]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B952EAA2612FDA70FD5E505724B2FB75" bold="true" box="[652,667,1893,1919]" pageId="40" pageNumber="41">a</emphasis>
|
||
. Photograph of preserved holotype (RMCA 2018.008.P.0355; 113.6 mm SL).
|
||
<emphasis id="B952EAA2612FDA70FEA050BA27ABFBA8" bold="true" box="[370,386,1928,1954]" pageId="40" pageNumber="41">b</emphasis>
|
||
. X-ray image of holotype.
|
||
<emphasis id="B952EAA2612FDA70FD6C50BA24C3FBA8" bold="true" box="[702,746,1928,1954]" pageId="40" pageNumber="41">c–d</emphasis>
|
||
. Photographs of freshly caught specimens.
|
||
<emphasis id="B952EAA2612FDA70FB3750BA22DBFBA8" bold="true" box="[1253,1266,1928,1954]" pageId="40" pageNumber="41">c</emphasis>
|
||
. Dominant male, the holotype.
|
||
<emphasis id="B952EAA2612FDA70FE62509927E9FBCF" bold="true" box="[432,448,1963,1989]" pageId="40" pageNumber="41">d</emphasis>
|
||
. Female (RMCA 2018.008.P.0352; 108.7 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33C653B612EDA72FF6F563A22B7FE83" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="description">
|
||
<paragraph id="8B9936B0612EDA71FF6F563A2767FD28" blockId="41.[189,334,264,290]" box="[189,334,264,290]" pageId="41" pageNumber="42">
|
||
<heading id="D0D181DC612EDA71FF6F563A2767FD28" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
|
||
<emphasis id="B952EAA2612EDA71FF6F563A2767FD28" bold="true" box="[189,334,264,290]" pageId="41" pageNumber="42">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B0612EDA71FF6F56052456FED1" blockId="41.[188,1400,311,731]" pageId="41" pageNumber="42">
|
||
Based on
|
||
<specimenCount id="9D20FD39612EDA71FEE5560A27E3FD5B" box="[311,458,311,338]" pageId="41" pageNumber="42" type="generic">8 specimens</specimenCount>
|
||
(
|
||
<quantity id="4CDE9B55612EDA71FE0E560A24B2FD58" box="[476,667,312,338]" metricMagnitude="-1" metricUnit="m" metricValue="1.034" metricValueMax="1.229" metricValueMin="0.839" pageId="41" pageNumber="42" unit="mm" value="103.4" valueMax="122.9" valueMin="83.9">83.9–122.9 mm</quantity>
|
||
SL); body shallow (
|
||
<tableCitation id="C6A4030B612EDA71FC44560525D9FD58" box="[918,1008,311,338]" captionStart="Table 1" captionStartId="9.[182,252,548,576]" captionTargetPageId="9" captionText="Table 1. (continued on the next three pages). Measurements and counts of the piscivorous species of Haplochromis from the Lake Edward system." httpUri="http://table.plazi.org/id/DF596638610EDA51FF6455162158FE4A" pageId="41" pageNumber="42" tableUuid="DF596638610EDA51FF6455162158FE4A">Table 1</tableCitation>
|
||
) and oval (
|
||
<figureCitation id="131D2A35612EDA71FB52560522F2FD58" box="[1152,1243,311,338]" captionStart="Fig" captionStartId="39.[189,232,1963,1989]" captionTargetBox="[189,1398,1249,1831]" captionTargetId="figure-226@39.[189,1399,1249,1831]" captionTargetPageId="39" captionText="Fig. 20. Haplochromis aquila sp. nov., holotype, ♂, 113.6 mm SL (RMCA 2018.008.P.0355). Drawn by N. Vranken." figureDoi="http://doi.org/10.5281/zenodo.6484209" httpUri="https://zenodo.org/record/6484209/files/figure.png" pageId="41" pageNumber="42">Fig. 20</figureCitation>
|
||
). Head long, narrow, and with a straight to gently convex dorsal outline; interorbital area narrow; eye average in size in comparison to generalised
|
||
<taxonomicName id="4C264D33612EDA71FD9E56B224E3FD93" authorityName="Trewavas" authorityYear="1933" box="[588,714,383,409]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="elegans">
|
||
<emphasis id="B952EAA2612EDA71FD9E56B22448FD93" box="[588,609,384,409]" italics="true" pageId="41" pageNumber="42">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA2612EDA71FDA2564D24E3FD93" box="[624,714,383,409]" italics="true" pageId="41" pageNumber="42">elegans</emphasis>
|
||
</taxonomicName>
|
||
(but large for a piscivorous species); cheek deep; lacrimal average in depth. Snout average in length, rounded in dorsal view, narrow, relatively blunt, and slopes gently at 35–45°; premaxillary pedicel long and (strongly) prominent. Jaws isognathous to weakly prognathous, long, narrow, and rounded in dorsal view; gape large and slopes at 25–35°; maxilla extends to vertical through anterior margin of pupil. Lower jaw relatively stout and with straight ventral outline in lateral view, mental prominence absent or weakly developed, and lower jaw side nearly flat with an inclination of 25–35° to horizontal in anterior view. Upper jaw expanded anteriorly. Lips and oral mucosa relatively large. Neurocranium average in depth, ethmo-vomerine block decurved, preorbital region shallow (18–24% NL), orbital region average in depth (30–33% NL), and supraoccipital crest shallow and wedge-shaped (
|
||
<figureCitation id="131D2A35612EDA71FDD955F2245BFED1" box="[523,626,704,731]" captionStart="Fig" captionStartId="40.[189,232,1892,1918]" captionTargetBox="[189,1393,275,1861]" captionTargetId="figure-19@40.[188,1393,265,1861]" captionTargetPageId="40" captionText="Fig. 21. Haplochromis aquila sp. nov. a. Photograph of preserved holotype (RMCA 2018.008.P.0355; 113.6 mm SL). b. X-ray image of holotype. c–d. Photographs of freshly caught specimens. c. Dominant male, the holotype. d. Female (RMCA 2018.008.P.0352; 108.7 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484211" httpUri="https://zenodo.org/record/6484211/files/figure.png" pageId="41" pageNumber="42">Fig. 21b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9936B0612EDA71FF6F5430276AF830" blockId="41.[189,1401,770,1082]" pageId="41" pageNumber="42">
|
||
Outer oral teeth few, unicuspid, and very large. Necks stout, conical, and straight; crowns straight to weakly recurved and acutely pointed. Dental arcades with anterior half expanded laterally. Outer teeth widely and irregularly set with neck-distances of 1–4 neck-widths. No enlarged teeth posterior in upper jaw. Inner teeth small, recurved, a mixture of unicuspid and weakly tricuspid, weakly tricuspid rare in large specimens (>
|
||
<quantity id="4CDE9B55612EDA71FE1F54A32418FFA6" box="[461,561,913,940]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="41" pageNumber="42" unit="mm" value="100.0">100 mm</quantity>
|
||
SL), and acutely pointed in all specimens. Tooth bands very slender crescent-shaped with 1–2 rows of inner teeth, and narrow posteriorly until only outer row remains past ¾ length of tooth band in upper jaw, past ⅔ length of tooth band in lower jaw. Inner teeth closely and regularly set on 1 outer neck-width from outer row; implantation recumbent; size uniform throughout tooth band.
|
||
</paragraph>
|
||
<paragraph id="8B9936B0612EDA71FF6F53502317F900" blockId="41.[188,1400,1122,1291]" pageId="41" pageNumber="42">
|
||
Lower pharyngeal bone long, very narrow, slim, and shallow over whole length (
|
||
<figureCitation id="131D2A35612EDA71FB51535022F5F876" box="[1155,1244,1122,1148]" captionStart="Fig" captionStartId="42.[189,232,1963,1989]" captionTargetBox="[189,1397,947,1862]" captionTargetId="figure-180@42.[189,1400,947,1864]" captionTargetPageId="42" captionText="Fig. 22. Haplochromis aquila sp. nov. (RMCA 2018.008.P.0356; 122.9 mm SL). a. Dorsal view of the lower pharyngeal jaw. b. Lateral view of the lower pharyngeal jaw." figureDoi="http://doi.org/10.5281/zenodo.6484213" httpUri="https://zenodo.org/record/6484213/files/figure.png" pageId="41" pageNumber="42">Fig. 22</figureCitation>
|
||
). Pharyngeal teeth relatively large and slender; major cusps acutely pointed; cusp gaps concave; minor cusps and cusp protuberances very small. Teeth in two median longitudinal rows equal in size and form to lateral teeth,
|
||
<quantity id="4CDE9B55612EDA71FF6F53FF26D0F8ED" box="[189,249,1229,1255]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="41" pageNumber="42" unit="in" value="12.0">12 in</quantity>
|
||
each row. Posterior transverse row with 16–17 teeth, implanted erectly with a lateral inclination; major cusps nearly straight, bluntly pointed, and laterally compressed; minor cusps mostly absent.
|
||
</paragraph>
|
||
<paragraph id="8B9936B0612EDA71FF6F5200235EF946" blockId="41.[189,1399,1330,1357]" box="[189,1399,1330,1357]" pageId="41" pageNumber="42">Chest scales small; transition to larger flank scales gradual. Minute scales on proximal half of caudal fin.</paragraph>
|
||
<paragraph id="8B9936B0612EDA71FF6F524624F9F9F3" blockId="41.[189,1399,1396,1529]" pageId="41" pageNumber="42">Caudal fin emarginate; dorsal and anal fins reach to between verticals through caudal-fin base and two scales anterior to this vertical. Pectoral fin reaches to between genital opening and second anal-fin spine; pelvic fin reaches to genital opening in females, to between first and second anal-fin spines in males; first pelvic fin slightly elongated in all specimens.</paragraph>
|
||
<paragraph id="8B9936B0612EDA71FF6F51132594FA55" blockId="41.[189,1399,1569,1632]" pageId="41" pageNumber="42">Ceratobranchial gill rakers in outer row of first gill arch short, stout, and simple; posteriormost rakers anvil-shaped or bifid. Epibranchial gill rakers slender and simple.</paragraph>
|
||
<paragraph id="8B9936B0612EDA71FF6F51B5278AFAAB" blockId="41.[189,419,1671,1697]" box="[189,419,1671,1697]" pageId="41" pageNumber="42">
|
||
<heading id="D0D181DC612EDA71FF6F51B5278AFAAB" bold="true" box="[189,419,1671,1697]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
|
||
<emphasis id="B952EAA2612EDA71FF6F51B5278AFAAB" bold="true" box="[189,419,1671,1697]" pageId="41" pageNumber="42">Colouration in life</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B0612EDA71FF6F518424B0FB55" blockId="41.[188,1399,1718,1888]" pageId="41" pageNumber="42">
|
||
Dominant males: body metallic grey; dorsum greyish; belly speckled black; chest, cheek, snout, lower jaw, and lips black; eye with (dark) grey outer ring and golden inner ring (
|
||
<figureCitation id="131D2A35612EDA71FBC551E82253FAFE" box="[1047,1146,1754,1780]" captionStart="Fig" captionStartId="40.[189,232,1892,1918]" captionTargetBox="[189,1393,275,1861]" captionTargetId="figure-19@40.[188,1393,265,1861]" captionTargetPageId="40" captionText="Fig. 21. Haplochromis aquila sp. nov. a. Photograph of preserved holotype (RMCA 2018.008.P.0355; 113.6 mm SL). b. X-ray image of holotype. c–d. Photographs of freshly caught specimens. c. Dominant male, the holotype. d. Female (RMCA 2018.008.P.0352; 108.7 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484211" httpUri="https://zenodo.org/record/6484211/files/figure.png" pageId="41" pageNumber="42">Fig. 21c</figureCitation>
|
||
). Nostril, interorbital, and lacrimal stripes very faint. Pectoral fin hyaline, pelvic fin black; dorsal fin sooty and with black lappets; anal and caudal fins bright red and with dusky bases; anal fin with dusky posterior part and 1–2 large orange egg-spots with dusky rings.
|
||
</paragraph>
|
||
<paragraph id="8B9936B0612EDA72FF6F50B5242BFD4C" blockId="41.[187,1399,1927,2025]" lastBlockId="42.[189,1399,265,327]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
|
||
Females and juveniles: body and dorsal part of head silver with yellowish sheen; belly, chest, operculum, cheek, lower jaw, and lips white; snout dusky; eye with (dark) grey outer ring and silver inner ring (
|
||
<figureCitation id="131D2A35612EDA71FF1450FC2704FBE3" box="[198,301,1998,2025]" captionStart="Fig" captionStartId="40.[189,232,1892,1918]" captionTargetBox="[189,1393,275,1861]" captionTargetId="figure-19@40.[188,1393,265,1861]" captionTargetPageId="40" captionText="Fig. 21. Haplochromis aquila sp. nov. a. Photograph of preserved holotype (RMCA 2018.008.P.0355; 113.6 mm SL). b. X-ray image of holotype. c–d. Photographs of freshly caught specimens. c. Dominant male, the holotype. d. Female (RMCA 2018.008.P.0352; 108.7 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484211" httpUri="https://zenodo.org/record/6484211/files/figure.png" pageId="41" pageNumber="42">Fig. 21d</figureCitation>
|
||
). Nostril and interorbital stripes faint. Pectoral fin hyaline, pelvic fin white; dorsal and caudal fins dusky; dorsal fin with black lappets; caudal fin with a yellowish base and a faint red distal part; anal fin yellow with a red sheen.
|
||
</paragraph>
|
||
<paragraph id="8B9936B0612DDA72FF6F565C27F9FD8D" blockId="42.[189,464,365,392]" box="[189,464,365,392]" pageId="42" pageNumber="43">
|
||
<heading id="D0D181DC612DDA72FF6F565C27F9FD8D" bold="true" box="[189,464,365,392]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
|
||
<emphasis id="B952EAA2612DDA72FF6F565C27F9FD8D" bold="true" box="[189,464,365,392]" pageId="42" pageNumber="43">Preserved colouration</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B0612DDA72FF6F56AE22B7FE83" blockId="42.[189,1400,412,650]" pageId="42" pageNumber="43">
|
||
Body brown; dorsum dark-brown; ventral part of body and operculum yellowish; chest and lower jaw black in males; cheek yellowish in females, dark-brown in males; snout dusky (
|
||
<figureCitation id="131D2A35612DDA72FB7A568D2326FDD3" box="[1192,1295,447,473]" captionStart="Fig" captionStartId="40.[189,232,1892,1918]" captionTargetBox="[189,1393,275,1861]" captionTargetId="figure-19@40.[188,1393,265,1861]" captionTargetPageId="40" captionText="Fig. 21. Haplochromis aquila sp. nov. a. Photograph of preserved holotype (RMCA 2018.008.P.0355; 113.6 mm SL). b. X-ray image of holotype. c–d. Photographs of freshly caught specimens. c. Dominant male, the holotype. d. Female (RMCA 2018.008.P.0352; 108.7 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484211" httpUri="https://zenodo.org/record/6484211/files/figure.png" pageId="42" pageNumber="43">Fig. 21a</figureCitation>
|
||
). Nostril and interorbital stripes faint; lacrimal stripe narrow and well-defined; mental blotch present; vertical preopercular stripe well-defined in males. Pectoral fin hyaline; pelvic fin yellowish with blackish first rays in females, black in males; dorsal fin dusky and with black lappets and maculated posterior part; anal fin yellowish with a dusky overlay and, in males, with black lappets and 1–2 large egg spots; caudal fin with a dusky base, a hyaline distal part, and, in males, whole ventral half hyaline.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C653B612DDA72FF6F558224ADFF2B" pageId="42" pageNumber="43" type="distribution">
|
||
<paragraph id="8B9936B0612DDA72FF6F558227D8FEC1" blockId="42.[189,497,688,715]" box="[189,497,688,715]" pageId="42" pageNumber="43">
|
||
<heading id="D0D181DC612DDA72FF6F558227D8FEC1" bold="true" box="[189,497,688,715]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
|
||
<emphasis id="B952EAA2612DDA72FF6F558227D8FEC1" bold="true" box="[189,497,688,715]" pageId="42" pageNumber="43">Distribution and ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B0612DDA72FF6F55ED24ADFF2B" blockId="42.[187,1398,735,801]" pageId="42" pageNumber="43">Only known from Lake Edward, found in inshore areas over muddy substrates. Based on its morphology, most probably a piscivorous species.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |