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<mods:title id="DF3AFF1812A3848C29985E86CE9FF3C9">Phylogeny and revision of Schraderiellus Rider, 1998 with the description of five new species (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini)</mods:title>
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<treatment id="038A87D6FFB6FFD9FF7FF98544DEFE68" ID-DOI="http://doi.org/10.5281/zenodo.4889943" ID-GBIF-Taxon="182955702" ID-Zenodo-Dep="4889943" LSID="urn:lsid:plazi:treatment:038A87D6FFB6FFD9FF7FF98544DEFE68" httpUri="http://treatment.plazi.org/id/038A87D6FFB6FFD9FF7FF98544DEFE68" lastPageId="9" lastPageNumber="56" pageId="8" pageNumber="55">
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<emphasis id="B957EAD2FFB6FFD8FF7FF98543A0F98A" bold="true" box="[151,738,1558,1584]" pageId="8" pageNumber="55">
Key to the species of
<taxonomicName id="4C234D43FFB6FFD8FE74F98543A0F98A" authority="Rider, 1998" authorityName="Rider" authorityYear="1998" box="[412,738,1558,1584]" class="Insecta" family="Pentatomidae" genus="Schraderiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B957EAD2FFB6FFD8FE74F985430EF98A" bold="true" box="[412,588,1558,1584]" italics="true" pageId="8" pageNumber="55">Schraderiellus</emphasis>
<bibRefCitation id="EFB24B31FFB6FFD8FDBBF98543A0F98A" author="Rider, D. A." box="[595,738,1558,1584]" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="8" pageNumber="55" pagination="504 - 510" part="100" refId="ref12493" refString="Rider, D. A. (1998) Nomenclatural changes in the Pentatomoidea (Hemiptera - Heteroptera: Pentatomidae, Tessaratomidae). III. Generic level changes. Proceedings of the Entomological Society of Washington, 100 (3), 504 - 510." title="Nomenclatural changes in the Pentatomoidea (Hemiptera - Heteroptera: Pentatomidae, Tessaratomidae). III. Generic level changes" type="journal article" year="1998">Rider, 1998</bibRefCitation>
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1 Connexivum, apexof hemelytra, radial vein, baseandapexof scutellumwithlargeyellowspots ........................ .............................................
<emphasis id="B957EAD2FFB6FFD8FCEAF9E945FBF92A" bold="true" box="[770,1209,1658,1680]" pageId="8" pageNumber="55">
<taxonomicName id="4C234D43FFB6FFD8FCEAF9E9452CF92A" authority="Roell &amp; Campos" authorityName="Roell &amp; Campos" box="[770,1134,1658,1680]" class="Insecta" family="Pentatomidae" genus="Schraderiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="luridomaculatus">
<emphasis id="B957EAD2FFB6FFD8FCEAF9E942FCF92A" bold="true" box="[770,958,1658,1680]" italics="true" pageId="8" pageNumber="55">S. luridomaculatus</emphasis>
Roell &amp; Campos
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, sp. nv.
</emphasis>
(
<figureCitation id="13182A45FFB6FFD8FB2EF9E94473F935" box="[1222,1329,1658,1679]" captionStart="FIGURE 3" captionStartId="4.[151,250,1955,1977]" captionTargetBox="[174,1428,196,1930]" captionTargetId="figure-86@4.[150,1436,193,1933]" captionTargetPageId="4" captionText="FIGURE 3. Species of Schraderiellus, dorsal and ventral habitus. A, B, S. hughesae; C, D, S. cinctus; E, F, S. luridomaculatus sp. nv.; G, H, S. rufilineatus sp. nv.; I, J, S. luteipedis sp. nv.; K, L, S. brevicolis sp. nv.; M, N, S. falcatus sp. nv.. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983228" httpUri="https://zenodo.org/record/4983228/files/figure.png" pageId="8" pageNumber="55">Figs. 3EF</figureCitation>
,
<figureCitation id="13182A45FFB6FFD8FAD3F9E9442CF935" box="[1339,1390,1658,1679]" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[314,1259,195,1859]" captionTargetId="figure-110@6.[314,1260,193,1868]" captionTargetPageId="6" captionText="FIGURE 5. Pygophores in posterior, dorsal and lateral views. AC, S. hughesae; DF, S. cinctus; GI, S. luridomaculatus sp. nv.; JL, S. rufilineatus sp. nv.; MO, S. luteipedis sp. nv.; PR, S. brevicolis sp. nv.; SU, S. falcatus sp. nv.. Abbreviations: dr, dorsal rim; il, inferior layer; p1, dorsal projection of the posterolateral angles; p2, basal projection of the posterolateral angles; pa, posterolateral angles; par, parameres; vr, ventral rim; X, segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983234" httpUri="https://zenodo.org/record/4983234/files/figure.png" pageId="8" pageNumber="55">5GI</figureCitation>
,
<figureCitation id="13182A45FFB6FFD8FA90F9E944D7F935" box="[1400,1429,1658,1679]" captionStart="FIGURE 8" captionStartId="9.[151,250,1935,1957]" captionTargetBox="[180,1393,531,1897]" captionTargetId="figure-449@9.[180,1394,525,1898]" captionTargetPageId="9" captionText="FIGURE 8. Female genital plates, posterior view. A, S. hughesae; B, S. cinctus; C, S. luridomaculatus sp. nv.; D, S. rufilineatus sp. nv.; E, S. luteipedis sp. nv.. Abbreviations. gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; laVIII, laterotergites VIII; laIX, laterotergites IX; X, segment X. Scale bars: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983240" httpUri="https://zenodo.org/record/4983240/files/figure.png" pageId="8" pageNumber="55">8C</figureCitation>
)
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<keyLead id="30D295F5FFB6FFD8FF7FF90444D9F973" pageId="8" pageNumber="55">- Connexivum immaculate or with small spots, or marked with a lateral yellowish to reddish band; hemelytra and scutellum occasionallywithsmallyellowspots ..................................................................... 2</keyLead>
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2 Tibiaeandtarsientirelyyellowishtolightbrown, contrastingwithdarkfemora (
<figureCitation id="13182A45FFB6FFD8FC0CF9414506F95D" box="[996,1092,1746,1767]" captionStart="FIGURE 3" captionStartId="4.[151,250,1955,1977]" captionTargetBox="[174,1428,196,1930]" captionTargetId="figure-86@4.[150,1436,193,1933]" captionTargetPageId="4" captionText="FIGURE 3. Species of Schraderiellus, dorsal and ventral habitus. A, B, S. hughesae; C, D, S. cinctus; E, F, S. luridomaculatus sp. nv.; G, H, S. rufilineatus sp. nv.; I, J, S. luteipedis sp. nv.; K, L, S. brevicolis sp. nv.; M, N, S. falcatus sp. nv.. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983228" httpUri="https://zenodo.org/record/4983228/files/figure.png" pageId="8" pageNumber="55">Fig. 3J, N</figureCitation>
) .......................... 3
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<paragraph id="8B9C36C0FFB6FFD8FF7FF97C44D7F8BE" blockId="8.[151,1436,1628,2031]" box="[151,1429,1775,1796]" pageId="8" pageNumber="55">
<keyLead id="30D295F5FFB6FFD8FF7FF97C44D7F8BE" box="[151,1429,1775,1796]" pageId="8" pageNumber="55">- Tibiaedarkbrown, concolorwithfemur, ortibiaelightbrownfromhalftotheapex; tarsivariable .................... 4</keyLead>
</paragraph>
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<paragraph id="8B9C36C0FFB6FFD8FF7FF89F44DEF885" blockId="8.[151,1436,1628,2031]" pageId="8" pageNumber="55">
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3 Connexivumnotpunctured; anterolateralmarginsof pronotum yellowish; parameres spatulated, slightlyfacingposteriorly... ........................................
<emphasis id="B957EAD2FFB6FFD8FD2FF8B94578F8FA" bold="true" box="[711,1082,1834,1856]" pageId="8" pageNumber="55">
<taxonomicName id="4C234D43FFB6FFD8FD2FF8B942ADF8FA" authority="Roell &amp; Campos" authorityName="Roell &amp; Campos" box="[711,1007,1834,1856]" class="Insecta" family="Pentatomidae" genus="Schraderiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="luteipedis">
<emphasis id="B957EAD2FFB6FFD8FD2FF8B9427CF8FA" bold="true" box="[711,830,1834,1856]" italics="true" pageId="8" pageNumber="55">S. luteipedis</emphasis>
Roell &amp; Campos
</taxonomicName>
, sp. nv.
</emphasis>
(
<figureCitation id="13182A45FFB6FFD8FBAFF8B945EAF885" box="[1095,1192,1834,1855]" captionStart="FIGURE 3" captionStartId="4.[151,250,1955,1977]" captionTargetBox="[174,1428,196,1930]" captionTargetId="figure-86@4.[150,1436,193,1933]" captionTargetPageId="4" captionText="FIGURE 3. Species of Schraderiellus, dorsal and ventral habitus. A, B, S. hughesae; C, D, S. cinctus; E, F, S. luridomaculatus sp. nv.; G, H, S. rufilineatus sp. nv.; I, J, S. luteipedis sp. nv.; K, L, S. brevicolis sp. nv.; M, N, S. falcatus sp. nv.. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983228" httpUri="https://zenodo.org/record/4983228/files/figure.png" pageId="8" pageNumber="55">Figs. 3I, J</figureCitation>
,
<figureCitation id="13182A45FFB6FFD8FB5BF8B945B6F885" box="[1203,1268,1834,1855]" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[314,1259,195,1859]" captionTargetId="figure-110@6.[314,1260,193,1868]" captionTargetPageId="6" captionText="FIGURE 5. Pygophores in posterior, dorsal and lateral views. AC, S. hughesae; DF, S. cinctus; GI, S. luridomaculatus sp. nv.; JL, S. rufilineatus sp. nv.; MO, S. luteipedis sp. nv.; PR, S. brevicolis sp. nv.; SU, S. falcatus sp. nv.. Abbreviations: dr, dorsal rim; il, inferior layer; p1, dorsal projection of the posterolateral angles; p2, basal projection of the posterolateral angles; pa, posterolateral angles; par, parameres; vr, ventral rim; X, segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983234" httpUri="https://zenodo.org/record/4983234/files/figure.png" pageId="8" pageNumber="55">5MO</figureCitation>
,
<figureCitation id="13182A45FFB6FFD8FB15F8B9445EF885" box="[1277,1308,1834,1855]" captionStart="FIGURE 6" captionStartId="7.[151,250,1967,1989]" captionTargetBox="[315,1253,194,1929]" captionTargetId="figure-68@7.[315,1260,193,1943]" captionTargetPageId="7" captionText="FIGURE 6. Parameres; dorsal, posterior, inner lateral and external lateral respectively. A, S. hughesae; B, S. cinctus; C, S. rufilineatus sp. nv.; D, S. luteipedis sp. nv.; E, S. brevicolis sp. nv.; F, S. falcatus sp. nv.. Scale bar: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4983236" httpUri="https://zenodo.org/record/4983236/files/figure.png" pageId="8" pageNumber="55">6D</figureCitation>
,
<figureCitation id="13182A45FFB6FFD8FACEF8B94404F885" box="[1318,1350,1834,1855]" captionStart="FIGURE 7" captionStartId="8.[151,250,854,876]" captionTargetBox="[159,1400,198,833]" captionTargetId="figure-898@8.[151,1436,193,833]" captionTargetPageId="8" captionText="FIGURE 7. Male internal genitalia (phallus), lateral view. A, S. hughesae; B, S. cinctus; C, S. rufilineatus sp. nv.; D, S. luteipedis sp. nv.; E, S. brevicolis sp. nv.; F, S. falcatus sp. nv.. Scale bars = 0.5mm" figureDoi="http://doi.org/10.5281/zenodo.4983238" httpUri="https://zenodo.org/record/4983238/files/figure.png" pageId="8" pageNumber="55">7D</figureCitation>
,
<figureCitation id="13182A45FFB6FFD8FAB8F8B9442EF885" box="[1360,1388,1834,1855]" captionStart="FIGURE 8" captionStartId="9.[151,250,1935,1957]" captionTargetBox="[180,1393,531,1897]" captionTargetId="figure-449@9.[180,1394,525,1898]" captionTargetPageId="9" captionText="FIGURE 8. Female genital plates, posterior view. A, S. hughesae; B, S. cinctus; C, S. luridomaculatus sp. nv.; D, S. rufilineatus sp. nv.; E, S. luteipedis sp. nv.. Abbreviations. gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; laVIII, laterotergites VIII; laIX, laterotergites IX; X, segment X. Scale bars: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983240" httpUri="https://zenodo.org/record/4983240/files/figure.png" pageId="8" pageNumber="55">8E</figureCitation>
,
<figureCitation id="13182A45FFB6FFD8FA9EF8B944D6F885" box="[1398,1428,1834,1855]" captionStart="FIGURE 9" captionStartId="10.[151,250,1802,1824]" captionTargetBox="[151,1357,233,1748]" captionTargetId="figure-88@10.[150,1436,193,1781]" captionTargetPageId="10" captionText="FIGURE 9. Female internal genitalia, posterior view. A, S. hughesae; B, S. cinctus; C, S. rufilineatus sp. nv.; D, S. luteipedis sp. nv.. Abbreviations: cs, capsula seminalis; gcIX, gonocoxites IX; gpIX, gonapophyses IX; id, inner ducts; laIX, laterotergites IX; md, medium ducts; od, outer ducts; pi, pars intermedialis; rs, ring sclerites; vi, vaginal intima. Scale bars: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4983242" httpUri="https://zenodo.org/record/4983242/files/figure.png" pageId="8" pageNumber="55">9D</figureCitation>
)
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- Connexivum slightly punctured; anterolateral margins of pronotum concolor with disc; parameres cylindrical, strongly facing posteriorly .......................................
<emphasis id="B957EAD2FFB6FFD8FCF5F8F745C2F8C0" bold="true" box="[797,1152,1892,1914]" pageId="8" pageNumber="55">
<taxonomicName id="4C234D43FFB6FFD8FCF5F8F74577F8C0" authority="Roell &amp; Campos" authorityName="Roell &amp; Campos" box="[797,1077,1892,1914]" class="Insecta" family="Pentatomidae" genus="Schraderiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="falcatus">
<emphasis id="B957EAD2FFB6FFD8FCF5F8F742C7F8C0" bold="true" box="[797,901,1892,1914]" italics="true" pageId="8" pageNumber="55">S. falcatus</emphasis>
Roell &amp; Campos
</taxonomicName>
, sp. nv.
</emphasis>
(
<figureCitation id="13182A45FFB6FFD8FB66F8F74446F8C3" box="[1166,1284,1892,1913]" captionStart="FIGURE 3" captionStartId="4.[151,250,1955,1977]" captionTargetBox="[174,1428,196,1930]" captionTargetId="figure-86@4.[150,1436,193,1933]" captionTargetPageId="4" captionText="FIGURE 3. Species of Schraderiellus, dorsal and ventral habitus. A, B, S. hughesae; C, D, S. cinctus; E, F, S. luridomaculatus sp. nv.; G, H, S. rufilineatus sp. nv.; I, J, S. luteipedis sp. nv.; K, L, S. brevicolis sp. nv.; M, N, S. falcatus sp. nv.. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983228" httpUri="https://zenodo.org/record/4983228/files/figure.png" pageId="8" pageNumber="55">Figs. 3MN</figureCitation>
,
<figureCitation id="13182A45FFB6FFD8FAE6F8F74405F8C3" box="[1294,1351,1892,1913]" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[314,1259,195,1859]" captionTargetId="figure-110@6.[314,1260,193,1868]" captionTargetPageId="6" captionText="FIGURE 5. Pygophores in posterior, dorsal and lateral views. AC, S. hughesae; DF, S. cinctus; GI, S. luridomaculatus sp. nv.; JL, S. rufilineatus sp. nv.; MO, S. luteipedis sp. nv.; PR, S. brevicolis sp. nv.; SU, S. falcatus sp. nv.. Abbreviations: dr, dorsal rim; il, inferior layer; p1, dorsal projection of the posterolateral angles; p2, basal projection of the posterolateral angles; pa, posterolateral angles; par, parameres; vr, ventral rim; X, segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983234" httpUri="https://zenodo.org/record/4983234/files/figure.png" pageId="8" pageNumber="55">5SU</figureCitation>
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<figureCitation id="13182A45FFB6FFD8FAB9F8F74428F8C3" box="[1361,1386,1892,1913]" captionStart="FIGURE 6" captionStartId="7.[151,250,1967,1989]" captionTargetBox="[315,1253,194,1929]" captionTargetId="figure-68@7.[315,1260,193,1943]" captionTargetPageId="7" captionText="FIGURE 6. Parameres; dorsal, posterior, inner lateral and external lateral respectively. A, S. hughesae; B, S. cinctus; C, S. rufilineatus sp. nv.; D, S. luteipedis sp. nv.; E, S. brevicolis sp. nv.; F, S. falcatus sp. nv.. Scale bar: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4983236" httpUri="https://zenodo.org/record/4983236/files/figure.png" pageId="8" pageNumber="55">6F</figureCitation>
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<figureCitation id="13182A45FFB6FFD8FA9CF8F744CDF8C3" box="[1396,1423,1892,1913]" captionStart="FIGURE 7" captionStartId="8.[151,250,854,876]" captionTargetBox="[159,1400,198,833]" captionTargetId="figure-898@8.[151,1436,193,833]" captionTargetPageId="8" captionText="FIGURE 7. Male internal genitalia (phallus), lateral view. A, S. hughesae; B, S. cinctus; C, S. rufilineatus sp. nv.; D, S. luteipedis sp. nv.; E, S. brevicolis sp. nv.; F, S. falcatus sp. nv.. Scale bars = 0.5mm" figureDoi="http://doi.org/10.5281/zenodo.4983238" httpUri="https://zenodo.org/record/4983238/files/figure.png" pageId="8" pageNumber="55">7F</figureCitation>
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<paragraph id="8B9C36C0FFB6FFD8FF7FF81144D9F80E" blockId="8.[151,1436,1628,2031]" pageId="8" pageNumber="55">
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4 Posterolateral angles of pygophore each with a projection directed dorsally (
<figureCitation id="13182A45FFB6FFD8FC39F8114579F82D" box="[977,1083,1922,1943]" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[314,1259,195,1859]" captionTargetId="figure-110@6.[314,1260,193,1868]" captionTargetPageId="6" captionText="FIGURE 5. Pygophores in posterior, dorsal and lateral views. AC, S. hughesae; DF, S. cinctus; GI, S. luridomaculatus sp. nv.; JL, S. rufilineatus sp. nv.; MO, S. luteipedis sp. nv.; PR, S. brevicolis sp. nv.; SU, S. falcatus sp. nv.. Abbreviations: dr, dorsal rim; il, inferior layer; p1, dorsal projection of the posterolateral angles; p2, basal projection of the posterolateral angles; pa, posterolateral angles; par, parameres; vr, ventral rim; X, segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983234" httpUri="https://zenodo.org/record/4983234/files/figure.png" pageId="8" pageNumber="55">Fig. 5C, p</figureCitation>
<figureCitation id="13182A45FFB6FFD8FBD2F8114506F82D" box="[1082,1092,1922,1943]" captionStart="FIGURE 1" captionStartId="2.[151,250,1767,1789]" captionTargetBox="[151,1435,196,1742]" captionTargetId="figure-62@2.[150,1436,193,1746]" captionTargetPageId="2" captionText="FIGURE 1. Strict consensus of 12 equally parsimonious tree calculated under implied weighting (k=9.344). (Black circles = nonhomoplasious transformations; white circles = homoplasious transformations. Numbers in red above the branches indicate the relative Bremer support values." figureDoi="http://doi.org/10.5281/zenodo.4983224" httpUri="https://zenodo.org/record/4983224/files/figure.png" pageId="8" pageNumber="55">1</figureCitation>
); posterior margins of gonocoxites VIIIovergonocoxitesIXconvex (
<figureCitation id="13182A45FFB6FFD8FDC4F80C43D6F80E" box="[556,660,1951,1972]" captionStart="FIGURE 8" captionStartId="9.[151,250,1935,1957]" captionTargetBox="[180,1393,531,1897]" captionTargetId="figure-449@9.[180,1394,525,1898]" captionTargetPageId="9" captionText="FIGURE 8. Female genital plates, posterior view. A, S. hughesae; B, S. cinctus; C, S. luridomaculatus sp. nv.; D, S. rufilineatus sp. nv.; E, S. luteipedis sp. nv.. Abbreviations. gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; laVIII, laterotergites VIII; laIX, laterotergites IX; X, segment X. Scale bars: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983240" httpUri="https://zenodo.org/record/4983240/files/figure.png" pageId="8" pageNumber="55">Fig. 8A, D</figureCitation>
) .............................................................. 5
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<paragraph id="8B9C36C0FFB6FFD8FF7FF82F44D9F855" blockId="8.[151,1436,1628,2031]" pageId="8" pageNumber="55">
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- Posterolateral angles of pygophore straight, without dorsal projections (
<figureCitation id="13182A45FFB6FFD8FC72F82F42A5F86B" box="[922,999,1980,2001]" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[314,1259,195,1859]" captionTargetId="figure-110@6.[314,1260,193,1868]" captionTargetPageId="6" captionText="FIGURE 5. Pygophores in posterior, dorsal and lateral views. AC, S. hughesae; DF, S. cinctus; GI, S. luridomaculatus sp. nv.; JL, S. rufilineatus sp. nv.; MO, S. luteipedis sp. nv.; PR, S. brevicolis sp. nv.; SU, S. falcatus sp. nv.. Abbreviations: dr, dorsal rim; il, inferior layer; p1, dorsal projection of the posterolateral angles; p2, basal projection of the posterolateral angles; pa, posterolateral angles; par, parameres; vr, ventral rim; X, segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983234" httpUri="https://zenodo.org/record/4983234/files/figure.png" pageId="8" pageNumber="55">Fig. 5R</figureCitation>
); posterior margins of gonocoxites VIII over gonocoxites IXrectilinear (
<figureCitation id="13182A45FFB6FFD8FE00F849432BF855" box="[488,617,2010,2031]" captionStart="FIGURE 8" captionStartId="9.[151,250,1935,1957]" captionTargetBox="[180,1393,531,1897]" captionTargetId="figure-449@9.[180,1394,525,1898]" captionTargetPageId="9" captionText="FIGURE 8. Female genital plates, posterior view. A, S. hughesae; B, S. cinctus; C, S. luridomaculatus sp. nv.; D, S. rufilineatus sp. nv.; E, S. luteipedis sp. nv.. Abbreviations. gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; laVIII, laterotergites VIII; laIX, laterotergites IX; X, segment X. Scale bars: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983240" httpUri="https://zenodo.org/record/4983240/files/figure.png" pageId="8" pageNumber="55">Fig. 8BC, E</figureCitation>
) ....................... .. .................. ...... ................ 6
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5 Dorsal surface of body dark brown with small yellow spots adjacent to the cicatrices on pronotum, on scutellum and hemelytra; anterolateral margin of pronotum and hemelytra dark brown or yellowish; margin of connexivum dark brown to black; parameres cylindrical, apexslightly flattened...........
<taxonomicName id="4C234D43FFB7FFD9FCF1FF41456FFF52" authority="(Ruckes, 1959)" baseAuthorityName="Ruckes" baseAuthorityYear="1959" box="[793,1069,210,232]" class="Insecta" family="Pentatomidae" genus="Schraderiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="hughesae">
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<emphasis id="B957EAD2FFB7FFD9FCF1FF4142D3FF52" bold="true" box="[793,913,210,232]" italics="true" pageId="9" pageNumber="56">S. hughesae</emphasis>
(
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(
<figureCitation id="13182A45FFB7FFD9FBD2FF4145EEFF5D" box="[1082,1196,210,231]" captionStart="FIGURE 3" captionStartId="4.[151,250,1955,1977]" captionTargetBox="[174,1428,196,1930]" captionTargetId="figure-86@4.[150,1436,193,1933]" captionTargetPageId="4" captionText="FIGURE 3. Species of Schraderiellus, dorsal and ventral habitus. A, B, S. hughesae; C, D, S. cinctus; E, F, S. luridomaculatus sp. nv.; G, H, S. rufilineatus sp. nv.; I, J, S. luteipedis sp. nv.; K, L, S. brevicolis sp. nv.; M, N, S. falcatus sp. nv.. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983228" httpUri="https://zenodo.org/record/4983228/files/figure.png" pageId="9" pageNumber="56">Figs. 3AB</figureCitation>
,
<figureCitation id="13182A45FFB7FFD9FB5DFF4145B2FF5D" box="[1205,1264,210,231]" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[314,1259,195,1859]" captionTargetId="figure-110@6.[314,1260,193,1868]" captionTargetPageId="6" captionText="FIGURE 5. Pygophores in posterior, dorsal and lateral views. AC, S. hughesae; DF, S. cinctus; GI, S. luridomaculatus sp. nv.; JL, S. rufilineatus sp. nv.; MO, S. luteipedis sp. nv.; PR, S. brevicolis sp. nv.; SU, S. falcatus sp. nv.. Abbreviations: dr, dorsal rim; il, inferior layer; p1, dorsal projection of the posterolateral angles; p2, basal projection of the posterolateral angles; pa, posterolateral angles; par, parameres; vr, ventral rim; X, segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983234" httpUri="https://zenodo.org/record/4983234/files/figure.png" pageId="9" pageNumber="56">5AC</figureCitation>
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<figureCitation id="13182A45FFB7FFD9FB13FF414458FF5D" box="[1275,1306,210,231]" captionStart="FIGURE 6" captionStartId="7.[151,250,1967,1989]" captionTargetBox="[315,1253,194,1929]" captionTargetId="figure-68@7.[315,1260,193,1943]" captionTargetPageId="7" captionText="FIGURE 6. Parameres; dorsal, posterior, inner lateral and external lateral respectively. A, S. hughesae; B, S. cinctus; C, S. rufilineatus sp. nv.; D, S. luteipedis sp. nv.; E, S. brevicolis sp. nv.; F, S. falcatus sp. nv.. Scale bar: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4983236" httpUri="https://zenodo.org/record/4983236/files/figure.png" pageId="9" pageNumber="56">6A</figureCitation>
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<figureCitation id="13182A45FFB7FFD9FACCFF414401FF5D" box="[1316,1347,210,231]" captionStart="FIGURE 7" captionStartId="8.[151,250,854,876]" captionTargetBox="[159,1400,198,833]" captionTargetId="figure-898@8.[151,1436,193,833]" captionTargetPageId="8" captionText="FIGURE 7. Male internal genitalia (phallus), lateral view. A, S. hughesae; B, S. cinctus; C, S. rufilineatus sp. nv.; D, S. luteipedis sp. nv.; E, S. brevicolis sp. nv.; F, S. falcatus sp. nv.. Scale bars = 0.5mm" figureDoi="http://doi.org/10.5281/zenodo.4983238" httpUri="https://zenodo.org/record/4983238/files/figure.png" pageId="9" pageNumber="56">7A</figureCitation>
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<figureCitation id="13182A45FFB7FFD9FAA5FF41442EFF5D" box="[1357,1388,210,231]" captionStart="FIGURE 8" captionStartId="9.[151,250,1935,1957]" captionTargetBox="[180,1393,531,1897]" captionTargetId="figure-449@9.[180,1394,525,1898]" captionTargetPageId="9" captionText="FIGURE 8. Female genital plates, posterior view. A, S. hughesae; B, S. cinctus; C, S. luridomaculatus sp. nv.; D, S. rufilineatus sp. nv.; E, S. luteipedis sp. nv.. Abbreviations. gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; laVIII, laterotergites VIII; laIX, laterotergites IX; X, segment X. Scale bars: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983240" httpUri="https://zenodo.org/record/4983240/files/figure.png" pageId="9" pageNumber="56">8A</figureCitation>
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<figureCitation id="13182A45FFB7FFD9FA9EFF4144D7FF5D" box="[1398,1429,210,231]" captionStart="FIGURE 9" captionStartId="10.[151,250,1802,1824]" captionTargetBox="[151,1357,233,1748]" captionTargetId="figure-88@10.[150,1436,193,1781]" captionTargetPageId="10" captionText="FIGURE 9. Female internal genitalia, posterior view. A, S. hughesae; B, S. cinctus; C, S. rufilineatus sp. nv.; D, S. luteipedis sp. nv.. Abbreviations: cs, capsula seminalis; gcIX, gonocoxites IX; gpIX, gonapophyses IX; id, inner ducts; laIX, laterotergites IX; md, medium ducts; od, outer ducts; pi, pars intermedialis; rs, ring sclerites; vi, vaginal intima. Scale bars: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4983242" httpUri="https://zenodo.org/record/4983242/files/figure.png" pageId="9" pageNumber="56">9A</figureCitation>
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- Dorsal surface of body without spots or with a conspicuous spot at apex of each radial vein; band lining the body reddish, includingconnexivum; parameresspatular, basecylindricalandapex expanded, flattened ............................. ......................................
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<taxonomicName id="4C234D43FFB7FFD9FD58FEB942AFFEFA" authority="Roell &amp; Campos" authorityName="Roell &amp; Campos" box="[688,1005,298,320]" class="Insecta" family="Pentatomidae" genus="Schraderiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="rufilineatus">
<emphasis id="B957EAD2FFB7FFD9FD58FEB9427FFEFA" bold="true" box="[688,829,298,320]" italics="true" pageId="9" pageNumber="56">S. rufilineatus</emphasis>
Roell &amp; Campos
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, sp. nv.
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(
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6 Anterolateral margin of pronotum and of hemelytra, and lateral margin of connexivum orange or yellowish, including connexivum; posterolateral angles of pygophore long and divergent.................................................... .................................................
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<emphasis id="B957EAD2FFB7FFD9FCDCFE1142D6FE22" bold="true" box="[820,916,386,408]" italics="true" pageId="9" pageNumber="56">S. cinctus</emphasis>
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(
<figureCitation id="13182A45FFB7FFD9FBD6FE1145EDFE2D" box="[1086,1199,386,407]" captionStart="FIGURE 3" captionStartId="4.[151,250,1955,1977]" captionTargetBox="[174,1428,196,1930]" captionTargetId="figure-86@4.[150,1436,193,1933]" captionTargetPageId="4" captionText="FIGURE 3. Species of Schraderiellus, dorsal and ventral habitus. A, B, S. hughesae; C, D, S. cinctus; E, F, S. luridomaculatus sp. nv.; G, H, S. rufilineatus sp. nv.; I, J, S. luteipedis sp. nv.; K, L, S. brevicolis sp. nv.; M, N, S. falcatus sp. nv.. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983228" httpUri="https://zenodo.org/record/4983228/files/figure.png" pageId="9" pageNumber="56">Figs. 3CD</figureCitation>
,
<figureCitation id="13182A45FFB7FFD9FB51FE1145B5FE2D" box="[1209,1271,386,407]" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[314,1259,195,1859]" captionTargetId="figure-110@6.[314,1260,193,1868]" captionTargetPageId="6" captionText="FIGURE 5. Pygophores in posterior, dorsal and lateral views. AC, S. hughesae; DF, S. cinctus; GI, S. luridomaculatus sp. nv.; JL, S. rufilineatus sp. nv.; MO, S. luteipedis sp. nv.; PR, S. brevicolis sp. nv.; SU, S. falcatus sp. nv.. Abbreviations: dr, dorsal rim; il, inferior layer; p1, dorsal projection of the posterolateral angles; p2, basal projection of the posterolateral angles; pa, posterolateral angles; par, parameres; vr, ventral rim; X, segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983234" httpUri="https://zenodo.org/record/4983234/files/figure.png" pageId="9" pageNumber="56">5 DF</figureCitation>
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<figureCitation id="13182A45FFB7FFD9FAE8FE11445CFE2D" box="[1280,1310,386,407]" captionStart="FIGURE 6" captionStartId="7.[151,250,1967,1989]" captionTargetBox="[315,1253,194,1929]" captionTargetId="figure-68@7.[315,1260,193,1943]" captionTargetPageId="7" captionText="FIGURE 6. Parameres; dorsal, posterior, inner lateral and external lateral respectively. A, S. hughesae; B, S. cinctus; C, S. rufilineatus sp. nv.; D, S. luteipedis sp. nv.; E, S. brevicolis sp. nv.; F, S. falcatus sp. nv.. Scale bar: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4983236" httpUri="https://zenodo.org/record/4983236/files/figure.png" pageId="9" pageNumber="56">6B</figureCitation>
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<figureCitation id="13182A45FFB7FFD9FAC0FE114407FE2D" box="[1320,1349,386,407]" captionStart="FIGURE 7" captionStartId="8.[151,250,854,876]" captionTargetBox="[159,1400,198,833]" captionTargetId="figure-898@8.[151,1436,193,833]" captionTargetPageId="8" captionText="FIGURE 7. Male internal genitalia (phallus), lateral view. A, S. hughesae; B, S. cinctus; C, S. rufilineatus sp. nv.; D, S. luteipedis sp. nv.; E, S. brevicolis sp. nv.; F, S. falcatus sp. nv.. Scale bars = 0.5mm" figureDoi="http://doi.org/10.5281/zenodo.4983238" httpUri="https://zenodo.org/record/4983238/files/figure.png" pageId="9" pageNumber="56">7B</figureCitation>
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<figureCitation id="13182A45FFB7FFD9FAB8FE11442CFE2D" box="[1360,1390,386,407]" captionStart="FIGURE 8" captionStartId="9.[151,250,1935,1957]" captionTargetBox="[180,1393,531,1897]" captionTargetId="figure-449@9.[180,1394,525,1898]" captionTargetPageId="9" captionText="FIGURE 8. Female genital plates, posterior view. A, S. hughesae; B, S. cinctus; C, S. luridomaculatus sp. nv.; D, S. rufilineatus sp. nv.; E, S. luteipedis sp. nv.. Abbreviations. gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; laVIII, laterotergites VIII; laIX, laterotergites IX; X, segment X. Scale bars: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983240" httpUri="https://zenodo.org/record/4983240/files/figure.png" pageId="9" pageNumber="56">8B</figureCitation>
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<figureCitation id="13182A45FFB7FFD9FA90FE1144D7FE2D" box="[1400,1429,386,407]" captionStart="FIGURE 9" captionStartId="10.[151,250,1802,1824]" captionTargetBox="[151,1357,233,1748]" captionTargetId="figure-88@10.[150,1436,193,1781]" captionTargetPageId="10" captionText="FIGURE 9. Female internal genitalia, posterior view. A, S. hughesae; B, S. cinctus; C, S. rufilineatus sp. nv.; D, S. luteipedis sp. nv.. Abbreviations: cs, capsula seminalis; gcIX, gonocoxites IX; gpIX, gonapophyses IX; id, inner ducts; laIX, laterotergites IX; md, medium ducts; od, outer ducts; pi, pars intermedialis; rs, ring sclerites; vi, vaginal intima. Scale bars: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4983242" httpUri="https://zenodo.org/record/4983242/files/figure.png" pageId="9" pageNumber="56">9B</figureCitation>
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<paragraph id="8B9C36C0FFB7FFD9FF7FFE0C44DEFE68" blockId="9.[151,1436,151,466]" pageId="9" pageNumber="56">
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- Anterolateral margin of pronotum and of hemelytra black or yellowish, margin of connexivum black; posterolateral angles of pygophoreshortandparallel .......................
<emphasis id="B957EAD2FFB7FFD9FCFEFE2F45C8FE68" bold="true" box="[790,1162,444,466]" pageId="9" pageNumber="56">
<taxonomicName id="4C234D43FFB7FFD9FCFEFE2F457CFE68" authority="Roell &amp; Campos" authorityName="Roell &amp; Campos" box="[790,1086,444,466]" class="Insecta" family="Pentatomidae" genus="Schraderiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="brevicolis">
<emphasis id="B957EAD2FFB7FFD9FCFEFE2F42CDFE68" bold="true" box="[790,911,444,466]" italics="true" pageId="9" pageNumber="56">S. brevicolis</emphasis>
Roell &amp; Campos
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, sp. nv.
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(
<figureCitation id="13182A45FFB7FFD9FB7FFE2E444AFE68" box="[1175,1288,445,466]" captionStart="FIGURE 3" captionStartId="4.[151,250,1955,1977]" captionTargetBox="[174,1428,196,1930]" captionTargetId="figure-86@4.[150,1436,193,1933]" captionTargetPageId="4" captionText="FIGURE 3. Species of Schraderiellus, dorsal and ventral habitus. A, B, S. hughesae; C, D, S. cinctus; E, F, S. luridomaculatus sp. nv.; G, H, S. rufilineatus sp. nv.; I, J, S. luteipedis sp. nv.; K, L, S. brevicolis sp. nv.; M, N, S. falcatus sp. nv.. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983228" httpUri="https://zenodo.org/record/4983228/files/figure.png" pageId="9" pageNumber="56">Figs. 3KL</figureCitation>
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<figureCitation id="13182A45FFB7FFD9FABBFE2E442DFE68" box="[1363,1391,445,466]" captionStart="FIGURE 6" captionStartId="7.[151,250,1967,1989]" captionTargetBox="[315,1253,194,1929]" captionTargetId="figure-68@7.[315,1260,193,1943]" captionTargetPageId="7" captionText="FIGURE 6. Parameres; dorsal, posterior, inner lateral and external lateral respectively. A, S. hughesae; B, S. cinctus; C, S. rufilineatus sp. nv.; D, S. luteipedis sp. nv.; E, S. brevicolis sp. nv.; F, S. falcatus sp. nv.. Scale bar: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4983236" httpUri="https://zenodo.org/record/4983236/files/figure.png" pageId="9" pageNumber="56">6E</figureCitation>
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<figureCitation id="13182A45FFB7FFD9FA91FE2E44D7FE68" box="[1401,1429,445,466]" captionStart="FIGURE 7" captionStartId="8.[151,250,854,876]" captionTargetBox="[159,1400,198,833]" captionTargetId="figure-898@8.[151,1436,193,833]" captionTargetPageId="8" captionText="FIGURE 7. Male internal genitalia (phallus), lateral view. A, S. hughesae; B, S. cinctus; C, S. rufilineatus sp. nv.; D, S. luteipedis sp. nv.; E, S. brevicolis sp. nv.; F, S. falcatus sp. nv.. Scale bars = 0.5mm" figureDoi="http://doi.org/10.5281/zenodo.4983238" httpUri="https://zenodo.org/record/4983238/files/figure.png" pageId="9" pageNumber="56">7E</figureCitation>
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