treatments-xml/data/88/04/6E/88046EA8E25D52A7E7DD35FA11578D08.xml
2024-06-21 12:42:55 +02:00

404 lines
42 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.123.1448" ID-GBIF-Dataset="f1dbd205-6f2f-4feb-867e-d52041258d12" ID-PMC="PMC3175121" ID-Pensoft-Pub="1313-2970-123-1" ID-PubMed="21998529" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-123-1" ModsDocOrigin="ZooKeys 123" ModsDocTitle="Australian Assassins, Part I: A review of the Assassin Spiders (Araneae, Archaeidae) of mid-eastern Australia" checkinTime="1451249962699" checkinUser="pensoft" docAuthor="Rix, Michael G. &amp; Harvey, Mark S." docDate="2011" docId="88046EA8E25D52A7E7DD35FA11578D08" docLanguage="en" docName="ZooKeys 123: 1-100" docOrigin="ZooKeys 123" docSource="http://dx.doi.org/10.3897/zookeys.123.1448" docTitle="Austrarchaea Forster &amp; Platnick 1984" docType="treatment" docVersion="4" lastPageNumber="10" masterDocId="A428FFEAFD3A7353FFACFFBCFFFA6D47" masterDocTitle="Australian Assassins, Part I: A review of the Assassin Spiders (Araneae, Archaeidae) of mid-eastern Australia" masterLastPageNumber="100" masterPageNumber="1" pageNumber="7" updateTime="1668152002706" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Australian Assassins, Part I: A review of the Assassin Spiders (Araneae, Archaeidae) of mid-eastern Australia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rix, Michael G.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Harvey, Mark S.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>123</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>100</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.123.1448</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.123.1448</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-123-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152030486" LSID="urn:lsid:plazi:treatment:88046EA8E25D52A7E7DD35FA11578D08" httpUri="http://treatment.plazi.org/id/88046EA8E25D52A7E7DD35FA11578D08" lastPageId="11" lastPageNumber="10" pageId="6" pageNumber="7">
<subSubSection pageId="6" pageNumber="7" type="nomenclature">
<paragraph pageId="6" pageNumber="7">
Genus
<taxonomicName LSID="http://species-id.net/wiki/Austrarchaea" authority="Forster &amp; Platnick, 1984" authorityName="Forster &amp; Platnick" authorityYear="1984" class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Austrarchaea Forster &amp; Platnick, 1984</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="reference_group">
<paragraph pageId="6" pageNumber="7">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Austrarchaea</taxonomicName>
Forster &amp; Platnick, 1984: 21;
<bibRefCitation author="Platnick, NI" journalOrPublisher="Bulletin of the British Arachnological Society" pageId="39" pageNumber="40" pagination="259 - 261" title="On Western Australian Austrarchaea (Araneae, Archaeidae)." volume="8" year="1991 b">Platnick 1991b</bibRefCitation>
: 259;
<bibRefCitation author="Main, BY" journalOrPublisher="Western Australian Naturalist" pageId="39" pageNumber="40" pagination="151 - 154" title="Additional records of the Gondwanan spider Austrarchaea from southwestern Australia." volume="20" year="1995">Main 1995</bibRefCitation>
: 151;
<bibRefCitation author="Harvey, MS" journalOrPublisher="Records of the Western Australian Museum" pageId="39" pageNumber="40" pagination="35 - 37" title="A new species of Austrarchaea (Araneae: Archaeidae) from Western Australia." volume="21" year="2002 a">Harvey 2002a</bibRefCitation>
: 35.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="type species">
<paragraph pageId="6" pageNumber="7">Type species.</paragraph>
<paragraph pageId="6" pageNumber="7">
<taxonomicName family="Archaeidae" genus="Archaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaea nodosa" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nodosa">Archaea nodosa</taxonomicName>
Forster, 1956, by original designation.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="diagnosis">
<paragraph pageId="6" pageNumber="7">Diagnosis.</paragraph>
<paragraph pageId="6" pageNumber="7">
Species of
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Austrarchaea</taxonomicName>
can be distinguished from all other extant
<taxonomicName family="Archaeidae" lsidName="" pageId="6" pageNumber="7" rank="family">Archaeidae</taxonomicName>
(i.e. Malagasy and African species of
<taxonomicName class="Arachnida" family="Archaeidae" genus="Eriauchenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eriauchenius" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Eriauchenius</taxonomicName>
and
<taxonomicName class="Arachnida" family="Archaeidae" genus="Afrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afrarchaea" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Afrarchaea</taxonomicName>
) by the presence of numerous, clustered spermathecae in females (Figs 5D, 10G, 14G) and by the presence of a long, wiry embolus on the pedipalp of males (Figs 10E, 15E, 27E) (
<bibRefCitation author="Forster, RR" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="39" pageNumber="40" pagination="1 - 106" title="A review of the archaeid spiders and their relatives, with notes on the limits of the superfamily Palpimanoidea (Arachnida, Araneae)." volume="178" year="1984">Forster and Platnick 1984</bibRefCitation>
,
<bibRefCitation author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="39" pageNumber="40" pagination="255 - 296" title="A revision of the assassin spiders of the Eriauchenius gracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="doi:10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood 2008</bibRefCitation>
). The remarkable, elevated shape of the carapace (Figs 4A-C, 10A-B) and the very long chelicerae (Figs 4B, 4D) will also immediately separate this genus from all other Australian spiders.
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="description">
<paragraph pageId="6" pageNumber="7">Description.</paragraph>
<paragraph pageId="6" pageNumber="7">Small, haplogyne, araneomorph spiders; total length 2.5 to 5.0.</paragraph>
<paragraph pageId="6" pageNumber="7">Colouration: Body colouration cryptic and relatively uniform across species, usually with only subtle intraspecific variation in abdominal patterning; carapace, sternum and chelicerae tan brown to dark reddish-brown, interspersed with darker regions of granulate cuticle (Fig. 5), covered in highly reflective setae; legs tan-brown to darker reddish-brown, with pattern of darker annulations on distal segments; abdomen mottled with beige and variable hues of grey-brown (Figs 5E-G), with darker sclerites, scutes and sclerotic spots (Figs 5A-B); paler beige markings due to reflective, subcuticular guanine crystals (Fig. 5B); antero-lateral face of abdomen always with large, humeral patch of reflective guanine crystals (Figs 5A, 5E-G).</paragraph>
<paragraph lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
Cephalothorax: Carapace greatly elevated anteriorly (CH/CL ratio usually 2.0-2.4; Fig. 6), with raised, highly modified pars cephalica forming
<normalizedToken originalValue="neck">'neck'</normalizedToken>
and bulbous
<normalizedToken originalValue="head">'head'</normalizedToken>
(see
<bibRefCitation author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="39" pageNumber="40" pagination="255 - 296" title="A revision of the assassin spiders of the Eriauchenius gracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="doi:10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood 2008</bibRefCitation>
) (Figs 4A-C);
<normalizedToken originalValue="neck">'neck'</normalizedToken>
with concomitantly long diastema (see
<bibRefCitation author="Schuett, K" journalOrPublisher="Humboldt-Universitaet zu Berlin, Berlin" pageId="39" pageNumber="40" title="The limits and phylogeny of the Araneoidea (Arachnida, Araneae)." year="2002">
<normalizedToken originalValue="Schütt">Schuett</normalizedToken>
2002
</bibRefCitation>
) between cheliceral bases and anterior margin of carapace, fused along entire length with sclerotised cuticle (Fig. 4C); cheliceral bases emanating from broad, fully-enclosed cheliceral foramen situated at front of
<normalizedToken originalValue="head">'head'</normalizedToken>
(Figs 4A-B); posterior
<normalizedToken originalValue="head">'head'</normalizedToken>
region usually also bearing two pairs of rudimentary protrusions or
<normalizedToken originalValue="horns">'horns'</normalizedToken>
, each typically terminating in a short, thickened seta (Fig. 4A). Carapace with densely granulate cuticular microstructure (Fig. 4G), covered in larger setose tubercles arranged in clusters or distinct rows (Figs 4C, 4E); each tubercle bearing single densely plumose or ciliate seta; setose tubercles largest on
<normalizedToken originalValue="neck">'neck'</normalizedToken>
and pars thoracica (Figs 4C, 4E). Eight eyes present on anterior margin of
<normalizedToken originalValue="head">'head'</normalizedToken>
, in four widely separated diads (Figs 4A-B); AME largest, widely separated, directed antero-laterally on rounded ocular bulge (Fig. 4B); PME situated closely posterior to AME, directed obliquely on postero-lateral side of ocular bulge; lateral eyes contiguous, with shared raised bases, directed ventro-laterally on widest lateral margin of
<normalizedToken originalValue="head">'head'</normalizedToken>
<pageBreakToken pageId="7" pageNumber="8" start="start">(</pageBreakToken>
Figs 4A-B). Sternum longer than wide, covered in setose tubercles; lateral margins separated from dorsal pleural sclerite extending between coxae I-IV. Labium subtriangular, not fused to sternum, directed antero-ventrally at oblique angle to sternum; labrum with pair of divergent projections on anterior surface. Maxillae large (Fig. 4C), straddling labium and labrum, converging distally; serrula a single row of teeth. Chelicerae very long, spear-like, distally divergent (Figs 4B, 4D, 4F), usually with proximal bulging projection (Fig. 4B); both sexes with oval, ectal stridulatory file adjacent to pedipalps (Fig. 4F); males usually also with brush (Figs 4F, 12C, 19C, 22C), short comb (Figs 14C, 18C) or dense tuft (Figs 16C, 17C) of accessory setae on anterior face of paturon. Chelicerae armed with three rows of peg teeth; anterior (prolateral) row with two peg teeth near tip of fang; posterior (retrolateral) row with single peg tooth near tip of fang; median (prolateral) row with more than 15 peg teeth extending along inner prolateral margin of paturon to near base of fang; median row with approximately nine porrect, comb-like peg teeth adjacent to fang, several larger, flattened, spiniform peg teeth near tip of fang, and additional progressively shorter, spiniform peg teeth along inner paturon (Fig. 4D); cheliceral retromargin also with four or five true teeth and prominent cheliceral gland mound.
</paragraph>
<paragraph pageId="7" pageNumber="8">
Legs and female pedipalp: Legs (longest to shortest) 1
<normalizedToken originalValue="42">-4-2-</normalizedToken>
3, covered with short plumose setae; spines absent; patella I long, greater than one-third length of femur I. Trichobothria present on tibiae and metatarsi of legs; tibiae I-IV each with two trichobothria; metatarsi I-IV each with single trichobothrium; bothrial bases with strongly ridged hood. Tarsi shorter than metatarsi, with capsulate tarsal organ and three claws; tarsi, metatarsi and distal tibiae of legs I-II usually with ventral and pro-ventral rows of moveable, spatulate setae. Female pedipalp with long, porrect trochanter and small tarsal claw; tibia with two dorsal trichobothria.
</paragraph>
<paragraph pageId="7" pageNumber="8">Abdomen: Abdomen arched anteriorly, rounded-subtriangular in lateral view, usually with four to six large hump-like tubercles on dorsal surface (Figs 5A, 5E-G); cuticle covered with short plumose setae and numerous sclerotic spots (Figs 5A-B). Epigastric region with sclerotised (setose) book lung covers and dorsal and ventral plates surrounding pedicel (Fig. 5C) (plates fused in males); dorsal pedicel plate with transverse ridges; females with median genital plate and sclerotised lateral sigillae (Figs 5C-D); males with broad dorsal scute fused anteriorly to epigastric sclerites, with or without additional paired sclerites associated with hump-like tubercles (Fig. 5A). Six spinnerets, surrounded by thickened cuticle; ALS largest, PMS smallest; colulus absent. Posterior pair of divided tracheal spiracles situated anterior to spinnerets; males also with transverse row of epiandrous gland spigots situated closely anterior to epigastric furrow.</paragraph>
<paragraph lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Genitalia: Female genitalia haplogyne, with sclerotised, strongly arched genital plate anterior to epigastric furrow (Figs 5C-D); internally with gonopore leading to large, spherical membranous bursa (Fig. 17G; see also
<bibRefCitation author="Forster, RR" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="39" pageNumber="40" pagination="1 - 106" title="A review of the archaeid spiders and their relatives, with notes on the limits of the superfamily Palpimanoidea (Arachnida, Araneae)." volume="178" year="1984">Forster and Platnick 1984</bibRefCitation>
, fig. 57) overlying two separate, radiating clusters of sclerotised anterior spermathecae (Figs 5D, 10G, 14G, 19G). Male pedipalp with complex, expandable pyriform bulb (Figs 10E, 19E, 23E, 24E), consisting of smooth tegulum, proximal
<normalizedToken originalValue="subtegulum">'subtegulum'</normalizedToken>
and associated tegular groove with basal haematodocha (Figs 10E, 23E, 27E) (similar to
<taxonomicName family="Mecysmaucheniidae" lsidName="" pageId="7" pageNumber="8" rank="family">Mecysmaucheniidae</taxonomicName>
and potentially analogous to the subtegular division of Entelegynae); distal tegulum with excavate, rimmed cavity surrounding massive, inflatable haematodochal complex
<pageBreakToken pageId="8" pageNumber="9" start="start">incorporating</pageBreakToken>
distal embolus, basal embolic sclerite and multiple tegular sclerites (Figs 26D, 27E) (see below); distal haematodochal complex with balloon-like proximal portion (anchored by distal rim of tegulum) and sinuous, tapering embolic portion (anchored by flexible, hinged retro-ventral conductor) (Figs 26D, 27E). Unexpanded pedipalp with folded, wiry embolus abutting conductor (Fig. 17E); tegular sclerites embedded pro-distally (Fig. 20E); pedipalpal expansion and haematodochal inflation (e.g. see Figs 14E, 23E, 26D, 27E) resulting in significant conformational changes to shape of conductor, length and orientation of embolus, and relative position of tegular sclerites.
</paragraph>
<paragraph pageId="8" pageNumber="9">
As noted by
<bibRefCitation author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="39" pageNumber="40" pagination="255 - 296" title="A revision of the assassin spiders of the Eriauchenius gracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="doi:10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood (2008)</bibRefCitation>
, the homology of the tegular sclerites among archaeid genera remains unclear, and this is especially true for
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Austrarchaea</taxonomicName>
relative to Malagasy and African taxa. For the purposes of this revision, and for an easy comparison among species of
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Austrarchaea</taxonomicName>
from mid-eastern Australia, the moveable tegular sclerites of the pedipalp are here numbered (1-3), relative to their pro-distal position within the unexpanded tegular cavity (e.g. see Figs 11F, 17F). Tegular sclerite 1 (TS 1) is a porrect, variably spiniform (Fig. 25F), rod-like (Fig. 20E) or filiform (Fig. 10F) process (breakable in some specimens; Fig. 21F) that originates near the prolateral base of the conductor, adjacent to the embedded base of the proximal embolic sclerite; during pedipalpal expansion this sclerite usually remains distally directed, positioned adjacent to the embolic haematodocha (Figs 26D, 27E). Tegular sclerite 2 (TS 2) is a distinctive, pointed, usually spur-like process, angled obliquely towards the conductor (Figs 11F, 25F), which is closely associated with the adjacent tegular sclerite 2a (TS 2a); in the unexpanded state, the sinuous, filiform TS 2a is usually obscured and
<normalizedToken originalValue="locked">'locked'</normalizedToken>
within a folded groove along the margin of TS 2 (see
<bibRefCitation author="Forster, RR" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="39" pageNumber="40" pagination="1 - 106" title="A review of the archaeid spiders and their relatives, with notes on the limits of the superfamily Palpimanoidea (Arachnida, Araneae)." volume="178" year="1984">Forster and Platnick 1984</bibRefCitation>
, figs 60, 62). Tegular sclerite 3 (TS 3) is the most disto-dorsally positioned of the tegular sclerites, with a broader, more plate-like morphology relative to TS 1-2, usually visible as a distally pointed or rod-like projection beyond the retro-distal rim of the tegulum (Figs 14E-F, 17E-F, 20D-E).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="distribution">
<paragraph pageId="8" pageNumber="9">Distribution.</paragraph>
<paragraph pageId="8" pageNumber="9">
Assassin spiders occur in mesic habitats throughout south-eastern, south-western and north-eastern mainland Australia (Fig. 2), usually in montane rainforests (Figs 30C, 38C, 41C) and wet eucalypt forests (Figs 39C, 42C, 45C), but occasionally in temperate heathlands or lowland rainforests (Fig. 40C). In south-eastern Australia they occur on Kangaroo Island (South Australia) and along the Great Dividing Range, from Grampians National Park in south-western Victoria north to Kroombit Tops National Park in south-eastern Queensland. In south-western Western Australia they occur from the Leeuwin-Naturaliste National Park east to Cape Le Grand National Park, with outlying populations in the Porongurup and Stirling Range National Parks. In north-eastern Queensland archaeids occur along the Great Dividing Range, from Eungella National Park near Mackay north to the Mount Finnigan Uplands, near Cooktown. Although this distribution is markedly concordant with the distribution of closed and tall open forests in
<normalizedToken originalValue="Australias">Australia's</normalizedToken>
east and extreme south-west (see Specht 1981), assassin spiders appear to be notably absent from Tasmania, from the Australian Alps and from the 'St Lawrence
<normalizedToken originalValue="Gap">Gap'</normalizedToken>
(Webb and Tracey 1981) (Fig. 2), as evidenced by the lack of museum specimens and despite targeted searches by the senior author.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="composition">
<paragraph pageId="9" pageNumber="10">
<pageBreakToken pageId="9" pageNumber="10" start="start">Composition</pageBreakToken>
.
</paragraph>
<paragraph pageId="9" pageNumber="10">
Five described species -
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea daviesae" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="daviesae">Austrarchaea daviesae</taxonomicName>
Forster &amp; Platnick, 1984,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea hickmani" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="hickmani">Austrarchaea hickmani</taxonomicName>
(Butler, 1929),
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea mainae" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mainae">Austrarchaea mainae</taxonomicName>
Platnick, 1991b,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea nodosa" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="nodosa">Austrarchaea nodosa</taxonomicName>
(Forster, 1956) and
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea robinsi" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="robinsi">Austrarchaea robinsi</taxonomicName>
Harvey, 2002a - and the 17 new species from mid-eastern Australia:
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea alani" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="alani">Austrarchaea alani</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea aleenae" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aleenae">Austrarchaea aleenae</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea binfordae" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="binfordae">Austrarchaea binfordae</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea christopheri" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="christopheri">Austrarchaea christopheri</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea clyneae" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="clyneae">Austrarchaea clyneae</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea cunninghami" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="cunninghami">Austrarchaea cunninghami</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea dianneae" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="dianneae">Austrarchaea dianneae</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea harmsi" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="harmsi">Austrarchaea harmsi</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea helenae" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="helenae">Austrarchaea helenae</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea judyae" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="judyae">Austrarchaea judyae</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea mascordi" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mascordi">Austrarchaea mascordi</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea mcguiganae" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mcguiganae">Austrarchaea mcguiganae</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea milledgei" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="milledgei">Austrarchaea milledgei</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea monteithi" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="monteithi">Austrarchaea monteithi</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea platnickorum" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="platnickorum">Austrarchaea platnickorum</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea raveni" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="raveni">Austrarchaea raveni</taxonomicName>
sp. n. and
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea smithae" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="smithae">Austrarchaea smithae</taxonomicName>
sp. n.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="remarks">
<paragraph pageId="9" pageNumber="10">Remarks.</paragraph>
<paragraph pageId="9" pageNumber="10">
At least three clades of
<taxonomicName family="Archaeidae" lsidName="" pageId="9" pageNumber="10" rank="family">Archaeidae</taxonomicName>
can be recognised in Australia (Fig. 3B; see also
<bibRefCitation author="Wood, HM" editor="Żabka, M" journalOrPublisher="Akademia Podlaska, Siedlce" pageId="39" pageNumber="40" title="Archaeid and mecysmaucheniid spiders and their relatives (Araneae: Archaeidae, Mecysmaucheniidae): phylogeny, biogeography and evolution of the carapace morphology." volumeTitle="18 th International Congress of Arachnology, Book of Abstracts, July 2010." year="2010">Wood et al. 2010</bibRefCitation>
): a mid-eastern Australian clade, distributed from southern New South Wales to south-eastern Queensland (including the enigmatic, basal species
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea monteithi" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="monteithi">Austrarchaea monteithi</taxonomicName>
sp. n.); a north-eastern Queensland clade, endemic to tropical Queensland; and a southern Australian clade, known from Victoria, South Australia and south-western Western Australia. For the purposes of this revision, mid-eastern Australian species are diagnosed relative only to other related species from mid-eastern Australia (i.e.
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea nodosa" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="nodosa">Austrarchaea nodosa</taxonomicName>
and its closest relatives; Fig. 3B), all of which possess five or six dorsal hump-like tubercles on the abdomen (Figs 5F-G) and have a carapace height to carapace length (CH/CL) ratio ≥ 2.00.
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea daviesae" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="daviesae">Austrarchaea daviesae</taxonomicName>
and related species from north-eastern Queensland have only two pairs of hump-like tubercles on the abdomen (Fig. 5E), and
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea hickmani" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="hickmani">Austrarchaea hickmani</taxonomicName>
,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea robinsi" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="robinsi">Austrarchaea robinsi</taxonomicName>
and
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea mainae" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mainae">Austrarchaea mainae</taxonomicName>
from southern Australia have a carapace height to carapace length (CH/CL) ratio significantly less than 2.00 (M. Rix, pers. obs.).
</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" type="key to the species of austrarchaea known from mid-eastern australia (males required)">
<paragraph pageId="9" pageNumber="10">Key to the species of Austrarchaea known from mid-eastern Australia (males required)</paragraph>
<paragraph lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
<table lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
<tr pageId="9" pageNumber="10">
<td colspan="1" pageId="9" pageNumber="10" rowspan="1">Fig. 5F</td>
<td colspan="1" pageId="9" pageNumber="10" rowspan="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea monteithi" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="monteithi">Austrarchaea monteithi</taxonomicName>
</td>
</tr>
<tr pageId="9" pageNumber="10">
<td colspan="1" pageId="9" pageNumber="10" rowspan="1">Fig. 5G</td>
</tr>
<tr pageId="9" pageNumber="10">
<td colspan="1" pageId="9" pageNumber="10" rowspan="1">Figs 16C17C23C</td>
</tr>
<tr pageId="9" pageNumber="10">
<td colspan="1" pageId="9" pageNumber="10" rowspan="1">Figs 12C19C22CFigs 14C18C</td>
</tr>
<tr pageId="9" pageNumber="10">
<td colspan="1" pageId="9" pageNumber="10" rowspan="1">Fig. 16C</td>
<td colspan="1" pageId="9" pageNumber="10" rowspan="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea harmsi" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="harmsi">Austrarchaea harmsi</taxonomicName>
</td>
</tr>
<tr pageId="9" pageNumber="10">
<td colspan="1" pageId="9" pageNumber="10" rowspan="1">Figs 17C23C</td>
</tr>
<tr pageId="9" pageNumber="10">
<td colspan="1" pageId="9" pageNumber="10" rowspan="1">Figs 17D-FFig. 17F</td>
<td colspan="1" pageId="9" pageNumber="10" rowspan="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea aleenae" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aleenae">Austrarchaea aleenae</taxonomicName>
</td>
</tr>
<tr pageId="9" pageNumber="10">
<td colspan="1" pageId="9" pageNumber="10" rowspan="1">Fig. 23EFig. 23E</td>
<td colspan="1" pageId="9" pageNumber="10" rowspan="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea milledgei" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="milledgei">Austrarchaea milledgei</taxonomicName>
</td>
</tr>
<tr pageId="9" pageNumber="10">
<td colspan="1" pageId="9" pageNumber="10" rowspan="1">Figs 20C-E</td>
<td colspan="1" pageId="9" pageNumber="10" rowspan="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea christopheri" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="christopheri">Austrarchaea christopheri</taxonomicName>
</td>
</tr>
<tr pageId="10" pageNumber="11">
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">
<pageBreakToken pageId="10" pageNumber="11" start="start">Figs</pageBreakToken>
18F25FFigs 10F22F
</td>
</tr>
<tr pageId="10" pageNumber="11">
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">Figs 8A, 8HFigs 8A, 8C-D, 8H9D</td>
</tr>
<tr pageId="10" pageNumber="11">
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">Figs 8F-G, 8I9C, 9F-I</td>
</tr>
<tr pageId="10" pageNumber="11">
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">Figs 14C18C</td>
</tr>
<tr pageId="10" pageNumber="11">
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">Figs 11C15C</td>
</tr>
<tr pageId="10" pageNumber="11">
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">Figs 14D-FFig. 14E</td>
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea raveni" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="raveni">Austrarchaea raveni</taxonomicName>
</td>
</tr>
<tr pageId="10" pageNumber="11">
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">Figs 18D-EFigs 18D-F</td>
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea alani" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="alani">Austrarchaea alani</taxonomicName>
</td>
</tr>
<tr pageId="10" pageNumber="11">
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">Figs 22E-F</td>
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea binfordae" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="binfordae">Austrarchaea binfordae</taxonomicName>
</td>
</tr>
<tr pageId="10" pageNumber="11">
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">Figs 11F15F</td>
</tr>
<tr pageId="10" pageNumber="11">
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">Figs 15D-FFig. 8C</td>
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea judyae" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="judyae">Austrarchaea judyae</taxonomicName>
</td>
</tr>
<tr pageId="10" pageNumber="11">
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">Figs 11D-EFig. 8H</td>
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea dianneae" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dianneae">Austrarchaea dianneae</taxonomicName>
</td>
</tr>
<tr pageId="10" pageNumber="11">
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">Figs 10F24F</td>
</tr>
<tr pageId="10" pageNumber="11">
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">Figs 12F13E21F25F</td>
</tr>
<tr pageId="10" pageNumber="11">
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">Figs 10D-E</td>
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea nodosa" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nodosa">Austrarchaea nodosa</taxonomicName>
</td>
</tr>
<tr pageId="10" pageNumber="11">
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">Figs 24D-E</td>
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea mascordi" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mascordi">Austrarchaea mascordi</taxonomicName>
</td>
</tr>
<tr pageId="10" pageNumber="11">
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">Fig. 27E</td>
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea mcguiganae" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mcguiganae">Austrarchaea mcguiganae</taxonomicName>
</td>
</tr>
<tr pageId="10" pageNumber="11">
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">Figs 12F13E21F25F26D</td>
</tr>
<tr pageId="10" pageNumber="11">
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">Fig. 9GFig. 9G</td>
<td colspan="1" pageId="10" pageNumber="11" rowspan="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea smithae" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="smithae">Austrarchaea smithae</taxonomicName>
</td>
</tr>
<tr pageId="11" pageNumber="12">
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">
<pageBreakToken pageId="11" pageNumber="12" start="start">Figs</pageBreakToken>
8F-G9C, 9IFigs 8F-G9C, 9I
</td>
</tr>
<tr pageId="11" pageNumber="12">
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">Fig. 8F</td>
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea clyneae" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="clyneae">Austrarchaea clyneae</taxonomicName>
</td>
</tr>
<tr pageId="11" pageNumber="12">
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">Figs 8G9C, 9I</td>
</tr>
<tr pageId="11" pageNumber="12">
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">Figs 21F26D</td>
</tr>
<tr pageId="11" pageNumber="12">
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">Fig. 12F</td>
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea cunninghami" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cunninghami">Austrarchaea cunninghami</taxonomicName>
</td>
</tr>
<tr pageId="11" pageNumber="12">
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">Fig. 26D</td>
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea helenae" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="helenae">Austrarchaea helenae</taxonomicName>
</td>
</tr>
<tr pageId="11" pageNumber="12">
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">Fig. 21F</td>
<td colspan="1" pageId="11" pageNumber="12" rowspan="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea platnickorum" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="platnickorum">Austrarchaea platnickorum</taxonomicName>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>