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<document id="15EE3D3BFE6C333665369A13231D6A8E" ID-CLB-Dataset="46662" ID-DOI="10.5281/zenodo.203293" ID-GBIF-Dataset="c01126cf-9cac-4a2a-b574-d6ea42e427a8" ID-ISSN="1175-5326" ID-Zenodo-Dep="203293" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460369354800" checkinUser="plazi" docAuthor="López-García, Margarita M., Méndez-Rojas, Diana M. &amp; Navarrete-Heredia, José Luís" docDate="2011" docId="53688790FFDDFFF821FD793B0273C165" docLanguage="en" docName="zt02916p064.pdf" docOrigin="Zootaxa 2916" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Megarthrus andinus López-García, Méndez-Rojas &amp; Navarrete-Heredia, 2011, sp. nov." docType="treatment" docVersion="9" lastPageNumber="64" masterDocId="AF51FFE8FFDDFFFA216A7D5F004AC36C" masterDocTitle="First record and a new species of Megarthrus Curtis (Coleoptera, Staphylinidae, Proteininae) from the Colombian Central Andes" masterLastPageNumber="64" masterPageNumber="62" pageNumber="62" updateTime="1698295557277" updateUser="ExternalLinkService">
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<mods:title id="269641B4DF55203380842EABE06AF2FE">First record and a new species of Megarthrus Curtis (Coleoptera, Staphylinidae, Proteininae) from the Colombian Central Andes</mods:title>
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<mods:namePart id="AC9533F4BE190AD0690CB0ED62CD129A">López-García, Margarita M.</mods:namePart>
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<mods:namePart id="096E1606014EAB1486E6E78F36163FD7">Méndez-Rojas, Diana M.</mods:namePart>
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<mods:namePart id="B358A0602B594ABC18D8364E86649B7D">Navarrete-Heredia, José Luís</mods:namePart>
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<mods:date id="FC45058C059CE57AA7B3CAF9AFD9EC44">2011</mods:date>
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<treatment id="53688790FFDDFFF821FD793B0273C165" ID-DOI="http://doi.org/10.5281/zenodo.6184642" ID-GBIF-Taxon="119411455" ID-Zenodo-Dep="6184642" LSID="urn:lsid:plazi:treatment:53688790FFDDFFF821FD793B0273C165" httpUri="http://treatment.plazi.org/id/53688790FFDDFFF821FD793B0273C165" lastPageId="2" lastPageNumber="64" pageId="0" pageNumber="62">
<subSubSection id="93DB650DFFDDFFFA21FD793B019FC710" box="[151,469,1124,1148]" pageId="0" pageNumber="62" type="nomenclature">
<paragraph id="DB7E3686FFDDFFFA21FD793B019FC710" blockId="0.[151,469,1124,1148]" box="[151,469,1124,1148]" pageId="0" pageNumber="62">
<heading id="803681EAFFDDFFFA21FD793B019FC710" bold="true" box="[151,469,1124,1148]" fontSize="10" level="2" pageId="0" pageNumber="62" reason="2">
<emphasis id="E9B5EA94FFDDFFFA21FD793B019FC710" bold="true" box="[151,469,1124,1148]" pageId="0" pageNumber="62">
<taxonomicName id="1CC14D05FFDDFFFA21FD793B0133C710" ID-CoL="6R7VH" box="[151,377,1124,1148]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="62" phylum="Arthropoda" rank="species" species="andinus" status="sp. nov.">
<emphasis id="E9B5EA94FFDDFFFA21FD793B0133C710" bold="true" box="[151,377,1124,1148]" italics="true" pageId="0" pageNumber="62">Megarthrus andinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F28657EFFFDDFFFA2015793B019FC710" box="[383,469,1124,1148]" pageId="0" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="93DB650DFFDDFFFA21FD79F6012CC644" pageId="0" pageNumber="62" type="materials_examined">
<paragraph id="DB7E3686FFDDFFFA21FD79F6012CC644" blockId="0.[151,1436,1193,1979]" pageId="0" pageNumber="62">
<emphasis id="E9B5EA94FFDDFFFA21FD79F6014FC7AD" bold="true" box="[151,261,1193,1217]" pageId="0" pageNumber="62">
<typeStatus id="047A8824FFDDFFFA21FD79F6014BC7AD" box="[151,257,1193,1217]" pageId="0" pageNumber="62" type="holotype">Holotype</typeStatus>
.
</emphasis>
3.
<collectingCountry id="A3D67616FFDDFFFA204779F601F4C7AC" box="[301,446,1193,1216]" name="Colombia" pageId="0" pageNumber="62">COLOMBIA</collectingCountry>
, Quindio Dept., Calarca, Planadas,
<geoCoordinate id="BEF55041FFDDFFFA225279F60382C7AC" box="[824,968,1193,1216]" direction="north" orientation="latitude" pageId="0" pageNumber="62" precision="1" value="4.437028">4°2613.3” N</geoCoordinate>
<geoCoordinate id="BEF55041FFDDFFFA22A579F60414C7AC" box="[975,1118,1193,1216]" direction="west" orientation="longitude" pageId="0" pageNumber="62" precision="15" value="-75.624725">75°3729” W</geoCoordinate>
,
<quantity id="1C399B63FFDDFFFA250279F604F0C7AC" box="[1128,1210,1193,1216]" metricMagnitude="3" metricUnit="m" metricValue="3.011" pageId="0" pageNumber="62" unit="m" value="3011.0">3011 m</quantity>
,
<date id="AF7F1046FFDDFFFA25AF79F60567C7AC" box="[1221,1325,1193,1216]" pageId="0" pageNumber="62" value="2009-05-14">14.v.2009</date>
(Méndez- Rojas D. &amp; López-García M.) ex leaf litter in cloud forest,
<collectionCode id="BDD0AE43FFDDFFFA224D79930326C78F" box="[807,876,1228,1251]" pageId="0" pageNumber="62">CIUQ</collectionCode>
.
<emphasis id="E9B5EA94FFDDFFFA221279930453C788" bold="true" box="[888,1049,1228,1252]" pageId="0" pageNumber="62">
<typeStatus id="047A8824FFDDFFFA2212799303A0C788" box="[888,1002,1228,1252]" pageId="0" pageNumber="62" type="paratype">Paratypes</typeStatus>
(5).
</emphasis>
Same data as
<typeStatus id="047A8824FFDDFFFA25D179930552C78F" box="[1211,1304,1228,1251]" pageId="0" pageNumber="62" type="holotype">holotype</typeStatus>
(1 3: ICN- MHN); same data as
<typeStatus id="047A8824FFDDFFFA201279B00192C66A" box="[376,472,1263,1286]" pageId="0" pageNumber="62" type="holotype">holotype</typeStatus>
, except for:
<date id="AF7F1046FFDDFFFA233179B002F5C66A" box="[603,703,1263,1286]" pageId="0" pageNumber="62" value="2011-01-27">27.i.2011</date>
(1 3, 1 Ƥ:
<collectionCode id="BDD0AE43FFDDFFFA225279B00334C66A" box="[824,894,1263,1286]" pageId="0" pageNumber="62">CIUQ</collectionCode>
); same data as
<typeStatus id="047A8824FFDDFFFA257779B00437C66A" box="[1053,1149,1263,1286]" pageId="0" pageNumber="62" type="holotype">holotype</typeStatus>
, except for:
<date id="AF7F1046FFDDFFFA246A79B00517C66A" box="[1280,1373,1263,1286]" pageId="0" pageNumber="62" value="2011-02-03">3.ii.2011</date>
(1 3: IAvH, 1 Ƥ:
<collectionCode id="BDD0AE43FFDDFFFA207F784E0111C644" box="[277,347,1297,1320]" pageId="0" pageNumber="62">CIUQ</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="93DB650DFFDDFFFB219F786B009FC48B" lastPageId="1" lastPageNumber="63" pageId="0" pageNumber="62" type="description">
<paragraph id="DB7E3686FFDDFFFA219F786B04C5C577" blockId="0.[151,1436,1193,1979]" pageId="0" pageNumber="62">
<emphasis id="E9B5EA94FFDDFFFA219F786B0133C620" bold="true" box="[245,377,1332,1356]" pageId="0" pageNumber="62">Description</emphasis>
. Habitus as in
<figureCitation id="43FA2A03FFDDFFFA234F786B022AC627" box="[549,608,1332,1355]" captionStart="FIGURES 1 9" captionStartId="1.[151,264,1673,1696]" captionTargetBox="[211,1370,260,1552]" captionTargetId="figure@1.[151,1436,194,1649]" captionTargetPageId="1" captionText="FIGURES 1 9. Megarthrus andinus sp. nov. Habitus (scale bar = 1 mm) (1); antenna (2); medial process of abdominal sternites II and III, in lateral view (anterior to left) (3); male sternite VIII, dissected (4); ventral (5) and lateral (6) aedeagus; female sternite VIII (7); ventral (8) and lateral (9) female genital segment (scale bars = 0.2 mm)." httpUri="https://zenodo.org/record/203294/files/figure.png" pageId="0" pageNumber="62">Fig.1</figureCitation>
. Combined length of pronotum and elytra=
<quantity id="1C399B63FFDDFFFA2521786B0484C627" box="[1099,1230,1332,1355]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="0" pageNumber="62" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4-1.6 mm</quantity>
; maximal pronotal width= 1
<quantity id="1C399B63FFDDFFFA206E78080113C602" box="[260,345,1367,1390]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="0" pageNumber="62" unit="mm" value="1.1">1.1 mm</quantity>
. Body dark brown with pronotum, antennae, legs, mentum and maxillary palpi somewhat paler; scape, pedicel and antennomeres 1, 2 and 11 paler than antennomeres 3 10. Dorsal pubescence fairly uniform, denser on pronotum than on elytron and abdomen; frontal setae directed forward, median area of frons less dense than on vertex; pronotum, elytra and abdomen with pubescence oriented posteriorly, elytral setae curved and recumbent, becoming denser near humeral area and with interspaces as wide as puncture diameters, abdominal pubescence becoming denser on lateral margins of tergites. Metasternal punctation deep with its median line almost impunctate.
</paragraph>
<paragraph id="DB7E3686FFDDFFFA21AD7B7801AFC5CA" blockId="0.[151,1436,1193,1979]" pageId="0" pageNumber="62">
Anterior margin of head non-truncate, nearly rounded. Temple rounded to subquadrate in dorsal view. Submentum surface weakly convex. Antenna filiform, 2.6 2.8 times longer than pronotum; with scape and pedicel sub-cylindrical and not flattened, segment 4 symmetrical and narrower than the remaining antennomeres which increase gradually in width, from segment 5 (
<figureCitation id="43FA2A03FFDDFFFA20FD7BD00193C5CA" box="[407,473,1679,1702]" captionStart="FIGURES 1 9" captionStartId="1.[151,264,1673,1696]" captionTargetBox="[211,1370,260,1552]" captionTargetId="figure@1.[151,1436,194,1649]" captionTargetPageId="1" captionText="FIGURES 1 9. Megarthrus andinus sp. nov. Habitus (scale bar = 1 mm) (1); antenna (2); medial process of abdominal sternites II and III, in lateral view (anterior to left) (3); male sternite VIII, dissected (4); ventral (5) and lateral (6) aedeagus; female sternite VIII (7); ventral (8) and lateral (9) female genital segment (scale bars = 0.2 mm)." httpUri="https://zenodo.org/record/203294/files/figure.png" pageId="0" pageNumber="62">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="DB7E3686FFDDFFFA21AD7BED018DC4D7" blockId="0.[151,1436,1193,1979]" pageId="0" pageNumber="62">
Pronotum 1.1 times longer than head; median portion convex with medial groove impunctate and flat. Pronotal disc shallowly depressed along basal and apical portion of lateral edge which is more deeply depressed medially. Hypomeron smooth with its inner edge wider in its median portion. Prosternal medial ridge absent; anterior prosternal margin bordered with a regular row of five longitudinal ridges. Protrochanter lacking transverse ridge. Scutellum with anterior border rounded and posterior border slightly acute, margins raised and with pubescence oriented posteriorly. Elytra 2.1 times longer than pronotum (
<figureCitation id="43FA2A03FFDDFFFA20B87A00025BC41A" box="[466,529,1887,1910]" captionStart="FIGURES 1 9" captionStartId="1.[151,264,1673,1696]" captionTargetBox="[211,1370,260,1552]" captionTargetId="figure@1.[151,1436,194,1649]" captionTargetPageId="1" captionText="FIGURES 1 9. Megarthrus andinus sp. nov. Habitus (scale bar = 1 mm) (1); antenna (2); medial process of abdominal sternites II and III, in lateral view (anterior to left) (3); male sternite VIII, dissected (4); ventral (5) and lateral (6) aedeagus; female sternite VIII (7); ventral (8) and lateral (9) female genital segment (scale bars = 0.2 mm)." httpUri="https://zenodo.org/record/203294/files/figure.png" pageId="0" pageNumber="62">Fig. 1</figureCitation>
), weakly narrowed at base; humeral callus raised; disc with conspicuous swellings; anterior margin convex and inner apical angle obtuse. Abdominal sternites II and III with medial processes as in
<figureCitation id="43FA2A03FFDDFFFA243C7ADD05DDC4F5" box="[1366,1431,1922,1945]" captionStart="FIGURES 1 9" captionStartId="1.[151,264,1673,1696]" captionTargetBox="[211,1370,260,1552]" captionTargetId="figure@1.[151,1436,194,1649]" captionTargetPageId="1" captionText="FIGURES 1 9. Megarthrus andinus sp. nov. Habitus (scale bar = 1 mm) (1); antenna (2); medial process of abdominal sternites II and III, in lateral view (anterior to left) (3); male sternite VIII, dissected (4); ventral (5) and lateral (6) aedeagus; female sternite VIII (7); ventral (8) and lateral (9) female genital segment (scale bars = 0.2 mm)." httpUri="https://zenodo.org/record/203294/files/figure.png" pageId="0" pageNumber="62">Fig. 3</figureCitation>
.
<collectingCountry id="A3D67616FFDDFFFA21FD7AFB0086C4D7" box="[151,204,1956,1979]" name="India" pageId="0" pageNumber="62">Hind</collectingCountry>
wings fully developed.
</paragraph>
<caption id="8FBE660EFFDCFFFB21FD7BD6045DC5B1" httpUri="https://zenodo.org/record/203294/files/figure.png" pageId="1" pageNumber="63" targetBox="[211,1370,260,1552]" targetPageId="1">
<paragraph id="DB7E3686FFDCFFFB21FD7BD6045DC5B1" blockId="1.[151,1436,1673,1758]" pageId="1" pageNumber="63">
<emphasis id="E9B5EA94FFDCFFFB21FD7BD60170C5F3" bold="true" box="[151,314,1673,1696]" pageId="1" pageNumber="63">FIGURES 19.</emphasis>
<taxonomicName id="1CC14D05FFDCFFFB20557BD50247C5CC" box="[319,525,1674,1696]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="andinus" status="sp. nov.">
<emphasis id="E9B5EA94FFDCFFFB20557BD50247C5CC" box="[319,525,1674,1696]" italics="true" pageId="1" pageNumber="63">Megarthrus andinus</emphasis>
</taxonomicName>
<emphasis id="E9B5EA94FFDCFFFB23797BD50229C5F3" bold="true" box="[531,611,1674,1695]" pageId="1" pageNumber="63">
<taxonomicNameLabel id="F28657EFFFDCFFFB23797BD50229C5F3" box="[531,611,1674,1695]" pageId="1" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Habitus (scale bar=1 mm) (1); antenna (2); medial process of abdominal sternites II and III, in lateral view (anterior to left) (3); male sternite VIII, dissected (4); ventral (5) and lateral (6) aedeagus; female sternite VIII (7); ventral (8) and lateral (9) female genital segment (scale bars= 0.2 mm).
</paragraph>
</caption>
<paragraph id="DB7E3686FFDCFFFB21AD7A7C009FC48B" blockId="1.[151,1436,1827,2023]" pageId="1" pageNumber="63">
<emphasis id="E9B5EA94FFDCFFFB21AD7A7C014DC456" box="[199,263,1827,1850]" italics="true" pageId="1" pageNumber="63">Male:</emphasis>
Frontoclypeal area not modified. Protarsamere 1 lacking adhesive setae. Mesofemur about as long as metafemur. Mesotibia shorter than metatibia. Metatarsomere 1 somewhat shorter than combined length of metatarsomeres 2 4. Peg-like setae arranged in a single row on mesotrochanter, mesotibia and metatibia. Sternite VIII as in
<figureCitation id="43FA2A03FFDCFFFB21D87AD400B9C4CE" box="[178,243,1931,1954]" captionStart="FIGURES 1 9" captionStartId="1.[151,264,1673,1696]" captionTargetBox="[211,1370,260,1552]" captionTargetId="figure@1.[151,1436,194,1649]" captionTargetPageId="1" captionText="FIGURES 1 9. Megarthrus andinus sp. nov. Habitus (scale bar = 1 mm) (1); antenna (2); medial process of abdominal sternites II and III, in lateral view (anterior to left) (3); male sternite VIII, dissected (4); ventral (5) and lateral (6) aedeagus; female sternite VIII (7); ventral (8) and lateral (9) female genital segment (scale bars = 0.2 mm)." httpUri="https://zenodo.org/record/203294/files/figure.png" pageId="1" pageNumber="63">Fig. 4</figureCitation>
. Sternite IX lacking subbasal protuberance. Aedeagus as in
<figureCitation id="43FA2A03FFDCFFFB221E7AD40399C4CE" box="[884,979,1931,1954]" captionStart="FIGURES 1 9" captionStartId="1.[151,264,1673,1696]" captionTargetBox="[211,1370,260,1552]" captionTargetId="figure@1.[151,1436,194,1649]" captionTargetPageId="1" captionText="FIGURES 1 9. Megarthrus andinus sp. nov. Habitus (scale bar = 1 mm) (1); antenna (2); medial process of abdominal sternites II and III, in lateral view (anterior to left) (3); male sternite VIII, dissected (4); ventral (5) and lateral (6) aedeagus; female sternite VIII (7); ventral (8) and lateral (9) female genital segment (scale bars = 0.2 mm)." httpUri="https://zenodo.org/record/203294/files/figure.png" pageId="1" pageNumber="63">Figs 5, 6</figureCitation>
.
<emphasis id="E9B5EA94FFDCFFFB22B47AD40464C4CE" box="[990,1070,1931,1954]" italics="true" pageId="1" pageNumber="63">Female</emphasis>
: Abdominal tergite VIII lacking a medioapical projection. Sternite VIII as in
<figureCitation id="43FA2A03FFDCFFFB23357AF102D5C4A9" box="[607,671,1966,1989]" captionStart="FIGURES 1 9" captionStartId="1.[151,264,1673,1696]" captionTargetBox="[211,1370,260,1552]" captionTargetId="figure@1.[151,1436,194,1649]" captionTargetPageId="1" captionText="FIGURES 1 9. Megarthrus andinus sp. nov. Habitus (scale bar = 1 mm) (1); antenna (2); medial process of abdominal sternites II and III, in lateral view (anterior to left) (3); male sternite VIII, dissected (4); ventral (5) and lateral (6) aedeagus; female sternite VIII (7); ventral (8) and lateral (9) female genital segment (scale bars = 0.2 mm)." httpUri="https://zenodo.org/record/203294/files/figure.png" pageId="1" pageNumber="63">Fig. 7</figureCitation>
. Genital segment as in
<figureCitation id="43FA2A03FFDCFFFB22FC7AF103BFC4A9" box="[918,1013,1966,1989]" captionStart="FIGURES 1 9" captionStartId="1.[151,264,1673,1696]" captionTargetBox="[211,1370,260,1552]" captionTargetId="figure@1.[151,1436,194,1649]" captionTargetPageId="1" captionText="FIGURES 1 9. Megarthrus andinus sp. nov. Habitus (scale bar = 1 mm) (1); antenna (2); medial process of abdominal sternites II and III, in lateral view (anterior to left) (3); male sternite VIII, dissected (4); ventral (5) and lateral (6) aedeagus; female sternite VIII (7); ventral (8) and lateral (9) female genital segment (scale bars = 0.2 mm)." httpUri="https://zenodo.org/record/203294/files/figure.png" pageId="1" pageNumber="63">Figs 8 9</figureCitation>
; gonocoxal plate lacking a mediodorsal ridge.
</paragraph>
</subSubSection>
<subSubSection id="93DB650DFFDFFFF821AD7DC8045BC3C3" box="[199,1041,151,175]" pageId="2" pageNumber="64" type="etymology">
<paragraph id="DB7E3686FFDFFFF821AD7DC8045BC3C3" blockId="2.[151,1437,151,521]" box="[199,1041,151,175]" pageId="2" pageNumber="64">
<emphasis id="E9B5EA94FFDFFFF821AD7DC8010DC3C3" bold="true" box="[199,327,151,175]" pageId="2" pageNumber="64">Etymology.</emphasis>
The specific epithet is an adjective referring to the Andean region.
</paragraph>
</subSubSection>
<subSubSection id="93DB650DFFDFFFF821AD7DE504B5C255" pageId="2" pageNumber="64" type="distribution">
<paragraph id="DB7E3686FFDFFFF821AD7DE504B5C255" blockId="2.[151,1437,151,521]" pageId="2" pageNumber="64">
<emphasis id="E9B5EA94FFDFFFF821AD7DE50113C3BE" bold="true" box="[199,345,186,210]" pageId="2" pageNumber="64">Distribution.</emphasis>
<taxonomicName id="1CC14D05FFDFFFF820357DE5019CC3BD" box="[351,470,186,209]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="64" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="E9B5EA94FFDFFFF820357DE5019CC3BD" box="[351,470,186,209]" italics="true" pageId="2" pageNumber="64">M. andinus</emphasis>
</taxonomicName>
is known only from its
<typeStatus id="047A8824FFDFFFF823BE7DE50348C3BD" box="[724,770,186,209]" pageId="2" pageNumber="64">type</typeStatus>
locality, which is a patch of high Andean forest on the Central Cordillera of the Colombian Andes. The species appears to be restricted to a very specific and small area, because after sampling different forests by sifting leaf litter it was exclusively collected at this locality. This pattern is in accord with the narrowly endemic distributions recorded for other Neotropical
<taxonomicName id="1CC14D05FFDFFFF822377C7D0391C255" box="[861,987,290,313]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="E9B5EA94FFDFFFF822377C7D0391C255" box="[861,987,290,313]" italics="true" pageId="2" pageNumber="64">Megarthrus</emphasis>
</taxonomicName>
species (
<bibRefCitation id="BF504B77FFDFFFF825577C7D04B9C255" author="Cuccodoro" box="[1085,1267,290,313]" pageId="2" pageNumber="64" refString="Cuccodoro, G. (2011) Revision of the Neotropical types of Megarthrus Curtis, 1829 and description of two new species from Costa Rica and Peru (Coleoptera, Staphylinidae, Proteininae). Revue Suisse de Zoologie, 118 (1), 107 147." type="journal article" year="2011">Cuccodoro 2011</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="93DB650DFFDFFFF821AD7C1B0273C165" pageId="2" pageNumber="64" type="discussion">
<paragraph id="DB7E3686FFDFFFF821AD7C1B0273C165" blockId="2.[151,1437,151,521]" pageId="2" pageNumber="64">
<emphasis id="E9B5EA94FFDFFFF821AD7C1B017FC230" bold="true" box="[199,309,324,348]" pageId="2" pageNumber="64">Remarks.</emphasis>
From Neotropical
<taxonomicName id="1CC14D05FFDFFFF823687C1A0235C230" box="[514,639,325,348]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="E9B5EA94FFDFFFF823687C1A0235C230" box="[514,639,325,348]" italics="true" pageId="2" pageNumber="64">Megarthrus</emphasis>
</taxonomicName>
species,
<taxonomicName id="1CC14D05FFDFFFF8238B7C1A03B7C230" authority="Bierig 1940" authorityName="Bierig" authorityYear="1940" box="[737,1021,325,348]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="64" phylum="Arthropoda" rank="species" species="mammiger">
<emphasis id="E9B5EA94FFDFFFF8238B7C1A033DC230" box="[737,887,325,348]" italics="true" pageId="2" pageNumber="64">M. mammiger</emphasis>
<bibRefCitation id="BF504B77FFDFFFF822147C1A03B7C230" author="Bierig" box="[894,1021,325,348]" pageId="2" pageNumber="64" refString="Bierig, A. (1940) Proteinini (Col. Staph.) Costaricenses. Revista de Entomologia, 11, 373 380." type="journal article" year="1940">Bierig 1940</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="A3D67616FFDFFFF825547C1A04F9C230" box="[1086,1203,325,348]" name="Costa Rica" pageId="2" pageNumber="64">Costa Rica</collectingCountry>
is the most similar to the Colombian new species, but the latter differs in the following characters: body smaller (3
<quantity id="1C399B63FFDFFFF825C27C3704B7C213" box="[1192,1277,360,383]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="64" unit="mm" value="3.4">3.4 mm</quantity>
instead of
<quantity id="1C399B63FFDFFFF824117C37008FC2CD" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="64" unit="mm" value="3.8">3.8 mm</quantity>
); antennae longer, antennomeres 1, 2 and 11 paler, whereas in
<taxonomicName id="1CC14D05FFDFFFF822037CD503B5C2CD" box="[873,1023,394,417]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="64" phylum="Arthropoda" rank="species" species="mammiger">
<emphasis id="E9B5EA94FFDFFFF822037CD503B5C2CD" box="[873,1023,394,417]" italics="true" pageId="2" pageNumber="64">M. mammiger</emphasis>
</taxonomicName>
antennomeres 10 11 are paler; elytral and pronotal setae oriented posteriorly (fairly straight in
<taxonomicName id="1CC14D05FFDFFFF823937CF203C5C2A8" box="[761,911,429,452]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="64" phylum="Arthropoda" rank="species" species="mammiger">
<emphasis id="E9B5EA94FFDFFFF823937CF203C5C2A8" box="[761,911,429,452]" italics="true" pageId="2" pageNumber="64">M. mammiger</emphasis>
</taxonomicName>
); male sternite VIII with apical margin rounded, bilobed, very wide and with medial emargination deep and U-shaped; aedeagus smaller and broader with parameres slightly shorter and with apex rounded.
</paragraph>
</subSubSection>
</treatment>
</document>