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treatments-xml/data/03/95/6A/03956A03A37E7F68FF67FD3E8A85DC00.xml
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<document id="10DCB777C5EC8C4509B4537C65C466D5" ID-CLB-Dataset="32021" ID-DOI="10.11646/zootaxa.4323.3.1" ID-GBIF-Dataset="f1be4566-adeb-4995-83c7-2aebab3c7458" ID-ISSN="1175-5326" ID-Zenodo-Dep="919606" ID-ZooBank="3E58F99B-2A30-437A-95Ae-D68A1D3Ecc45" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1506057765384" checkinUser="plazi" docAuthor="Schmidt, Joachim" docDate="2017" docId="03956A03A37E7F68FF67FD3E8A85DC00" docLanguage="en" docName="zootaxa.4323.3.1.pdf" docOrigin="Zootaxa 4323 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichotichnus (Bottchrus) panchhaseensis Schmidt 2017, sp. n." docType="treatment" docVersion="7" lastPageNumber="322" masterDocId="FFAC127BA36D7F7DFFF0FFAE8B5CD930" masterDocTitle="Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species" masterLastPageNumber="358" masterPageNumber="301" pageNumber="320" updateTime="1698468866275" updateUser="ExternalLinkService">
<mods:mods id="20B0B1AAA711EC17B38C2291275963CC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3DC949332ED421AF8BC120CA0EF635FB">
<mods:title id="624EB627E203162E848BE6D8DC80F712">Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species</mods:title>
</mods:titleInfo>
<mods:name id="0C873028A4AFBA8508A1BCF89EBD4EEF" type="personal">
<mods:role id="51EC92E09C2D7D61B0ADD03054818B7F">
<mods:roleTerm id="FED32478D29C0030EDDD07980FC5D38B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="09BE1EFF29447C552D93E1547A069D40">Schmidt, Joachim</mods:namePart>
</mods:name>
<mods:typeOfResource id="CD71F511CD4192DCAAA04A3463952E64">text</mods:typeOfResource>
<mods:relatedItem id="F14C49589047725998881618D9A9F166" type="host">
<mods:titleInfo id="A38C37933D7657B795A19A8C196AD7E1">
<mods:title id="CD7ADD4C409070295932F03F9851D827">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="359E0772BC050B7D62CE8B46446BD0EA">
<mods:date id="188D89647894DF8708842D4A1186C333">2017</mods:date>
<mods:detail id="9CD70946EF9BD65FA1C2B01AE2466021" type="pubDate">
<mods:number id="AB37FC970F3982AAB4ECCA39DE9C5D23">2017-09-22</mods:number>
</mods:detail>
<mods:detail id="200C27563DD8DE34FDFDBA695335DEBD" type="volume">
<mods:number id="FFA92F4F90DF687BDCD1BA4047C68EF2">4323</mods:number>
</mods:detail>
<mods:detail id="5327BC492081140839D83350145DA1ED" type="issue">
<mods:number id="97A22D1FE102B2AD384EF571600D208E">3</mods:number>
</mods:detail>
<mods:extent id="3620211F6FD9B9FC1D4B182A1D9F4DD8" unit="page">
<mods:start id="4E8BF426D1874AA4AB34C4594D2DE183">301</mods:start>
<mods:end id="3C73D34D331C5E74D16BDE8683BE1F27">358</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A08E3F2F60A8E233F84A6D907C5186B1">journal article</mods:classification>
<mods:identifier id="661FA813C82249A061B3A4943FF3F82F" type="CLB-Dataset">32021</mods:identifier>
<mods:identifier id="2FBC4F7F8E86DD2D0EDBAA42C4BBAA1A" type="DOI">10.11646/zootaxa.4323.3.1</mods:identifier>
<mods:identifier id="D327C20973219A5A5C0D93043DE23EA5" type="GBIF-Dataset">f1be4566-adeb-4995-83c7-2aebab3c7458</mods:identifier>
<mods:identifier id="10290B5EFB6000C70B8186A4B94399C4" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7A9A3DDEA7948F50C89FB55810F3D6EA" type="Zenodo-Dep">919606</mods:identifier>
<mods:identifier id="A4190649BF6E05D545329363A9CCE2B1" type="ZooBank">3E58F99B-2A30-437A-95Ae-D68A1D3Ecc45</mods:identifier>
</mods:mods>
<treatment id="03956A03A37E7F68FF67FD3E8A85DC00" ID-DOI="http://doi.org/10.5281/zenodo.6015751" ID-GBIF-Taxon="134839413" ID-Zenodo-Dep="6015751" LSID="urn:lsid:plazi:treatment:03956A03A37E7F68FF67FD3E8A85DC00" httpUri="http://treatment.plazi.org/id/03956A03A37E7F68FF67FD3E8A85DC00" lastPageId="21" lastPageNumber="322" pageId="19" pageNumber="320">
<subSubSection id="C326889EA37E7F6EFF67FD3E8AF8DBFB" pageId="19" pageNumber="320" type="nomenclature">
<paragraph id="8B83DB15A37E7F6EFF67FD3E89BEDB9A" blockId="19.[151,738,656,715]" box="[151,738,656,682]" pageId="19" pageNumber="320">
<heading id="D0CB6C79A37E7F6EFF67FD3E89BEDB9A" bold="true" box="[151,738,656,682]" fontSize="11" level="1" pageId="19" pageNumber="320" reason="1">
<emphasis id="B9480707A37E7F6EFF67FD3E89BEDB9A" bold="true" box="[151,738,656,682]" pageId="19" pageNumber="320">
<taxonomicName id="4C3CA096A37E7F6EFF67FD3E89C4DB9A" ID-CoL="58LHN" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[151,664,656,682]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="panchhaseensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A37E7F6EFF67FD3E8A1DDB9A" bold="true" box="[151,321,656,682]" italics="true" pageId="19" pageNumber="320">Trichotichnus</emphasis>
(
<emphasis id="B9480707A37E7F6EFEA2FD3E8A95DB9A" bold="true" box="[338,457,656,682]" italics="true" pageId="19" pageNumber="320">Bottchrus</emphasis>
)
<emphasis id="B9480707A37E7F6EFE2AFD3E89C4DB9A" bold="true" box="[474,664,656,682]" italics="true" pageId="19" pageNumber="320">panchhaseensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA37E7F6EFD6FFD3E89BEDB9A" box="[671,738,656,682]" pageId="19" pageNumber="320" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B83DB15A37E7F6EFF67FD1C8AF8DBFB" blockId="19.[151,738,656,715]" box="[151,420,690,715]" pageId="19" pageNumber="320">
(
<figureCitation id="1307C790A37E7F6EFF6FFD1C8BA9DBFB" box="[159,245,690,715]" captionStart="FIGURES 38 – 41" captionStartId="15.[151,264,1942,1965]" captionTargetBox="[273,1314,196,1918]" captionTargetId="figure@15.[271,1315,193,1921]" captionTargetPageId="15" captionText="FIGURES 38 – 41. Himalayan species of Trichotichnus subgenus Bottchrus, photographs of habitus. 38, T. schawalleri sp. n., holotype; 39, T. obliquebasalis sp. n., paratype, male; 40, T. siklesensis sp. n., paratype, male; 41, T. panchhaseensis sp. n., paratype, male." httpUri="https://zenodo.org/record/919620/files/figure.png" pageId="19" pageNumber="320">Figs 41</figureCitation>
,
<figureCitation id="1307C790A37E7F6EFEF1FD1C8A16DBFB" box="[257,330,690,715]" captionStart="FIGURES 53 – 59" captionStartId="20.[151,264,1678,1701]" captionTargetBox="[195,1394,776,1638]" captionTargetId="figure@20.[176,1411,769,1657]" captionTargetPageId="20" captionText="FIGURES 53 – 59. Trichotichnus (Bottchrus) panchhaseensis sp. n. (53 – 55, paratypes; 56 – 59, holotype). 53, left metepisternum; 54, apex of last (VII) abdominal sternite, male; 55, same, female; 56 – 57, median lobe of aedeagus; 58, terminal lamella; 59, apical capitulum. 53 – 55, ventral view; 56, lateral view; 57 – 58, dorsal view; 59, caudal view. Scale bars = 0.25 mm (A: Figs 53 – 57; B: Figs 58 – 59)." httpUri="https://zenodo.org/record/919632/files/figure.png" pageId="19" pageNumber="320">53–59</figureCitation>
,
<figureCitation id="1307C790A37E7F6EFEA5FD1C8ADDDBFB" box="[341,385,690,715]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="19" pageNumber="320">151</figureCitation>
,
<figureCitation id="1307C790A37E7F6EFE7DFD1D8AC7DBFA" box="[397,411,691,714]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="19" pageNumber="320">
<emphasis id="B9480707A37E7F6EFE7DFD1D8AC7DBFA" box="[397,411,691,714]" italics="true" pageId="19" pageNumber="320">7</emphasis>
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C326889EA37E7F6EFF67FD54890DDA93" pageId="19" pageNumber="320" type="materials_examined">
<paragraph id="8B83DB15A37E7F6EFF67FD548A14DA07" blockId="19.[151,1437,762,2011]" pageId="19" pageNumber="320">
<materialsCitation id="3B54D148A37E7F6EFF67FD548A14DA07" ID-GBIF-Occurrence="1655986858" collectingDate="97-05-18" collectorName="W Pokhara &amp; Schmidt" country="Nepal" elevation="2150" location="Annapurna" municipality="Mt. Panchhase" pageId="19" pageNumber="320" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B9480707A37E7F6EFF67FD548A18DA23" bold="true" box="[151,324,762,787]" pageId="19" pageNumber="320">
<typeStatus id="548765B7A37E7F6EFF67FD548B8EDA23" box="[151,210,762,787]" pageId="19" pageNumber="320">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="548765B7A37E7F6EFEAAFD548A9BDA23" box="[346,455,762,787]" pageId="19" pageNumber="320" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D3A109CA37E7F6EFE28FD558AB3DA22" box="[472,495,763,786]" pageId="19" pageNumber="320" type="male">♂</specimenCount>
, “
<collectingCountry id="F32B9B85A37E7F6EFDFBFD55893ADA23" box="[523,614,763,787]" name="Nepal" pageId="19" pageNumber="320">NEPAL</collectingCountry>
<location id="8EE38DCEA37E7F6EFD84FD5589A8DA23" LSID="urn:lsid:plazi:treatment:03956A03A37E7F68FF67FD3E8A85DC00:8EE38DCEA37E7F6EFD84FD5589A8DA23" box="[628,756,763,787]" country="Nepal" municipality="Mt. Panchhase" name="Annapurna" pageId="19" pageNumber="320">Annapurna</location>
,
<collectingMunicipality id="6BE7416FA37E7F6EFCF5FD5588E8DA23" box="[773,948,762,787]" pageId="19" pageNumber="320">Mt. Panchhase</collectingMunicipality>
,
<elevation id="00113C26A37E7F6EFC36FD558F36DA22" box="[966,1130,763,787]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="19" pageNumber="320" unit="m" value="2150.0" valueMax="2300.0" valueMin="2000.0">
<quantity id="4CC476F0A37E7F6EFC36FD558F36DA22" box="[966,1130,763,787]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="19" pageNumber="320" unit="m" value="2150.0" valueMax="2300.0" valueMin="2000.0">2000–2300 m</quantity>
</elevation>
,
<collectorName id="26C9BEC3A37E7F6EFB8DFD558E59DA23" box="[1149,1285,762,787]" pageId="19" pageNumber="320">W Pokhara</collectorName>
,
<collectingDate id="EFC6043DA37E7F6EFAE6FD548E30DA23" box="[1302,1388,762,787]" pageId="19" pageNumber="320" value="97-05-18">
<date id="FF82FDD5A37E7F6EFAE6FD548E30DA23" box="[1302,1388,762,787]" pageId="19" pageNumber="320" value="97-05-18">18.5.97</date>
</collectingDate>
, lg.
<collectorName id="26C9BEC3A37E7F6EFF67FCB08BABDA07" box="[151,247,798,823]" pageId="19" pageNumber="320">Schmidt</collectorName>
” (cJS).
</materialsCitation>
</paragraph>
<paragraph id="8B83DB15A37E7F6EFF37FCED890DDA93" blockId="19.[151,1437,762,2011]" pageId="19" pageNumber="320">
<materialsCitation id="3B54D148A37E7F6EFF37FCED8BE7DA4E" ID-GBIF-Occurrence="1655986861" collectingDate="97-05-18" collectionCode="ZIN" collectorName="W Pokhara &amp; Schmidt" country="Nepal" elevation="2150" location="Pokhara Mt" municipality="Mt. Panchhase" pageId="19" pageNumber="320" specimenCount="34" specimenCount-female="16" specimenCount-male="18" typeStatus="paratype">
<typeStatus id="548765B7A37E7F6EFF37FCED8A65DA6B" box="[199,313,835,859]" pageId="19" pageNumber="320" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D3A109CA37E7F6EFEB3FCED8A21DA6A" box="[323,381,835,859]" pageId="19" pageNumber="320" type="male">17 ♂</specimenCount>
,
<specimenCount id="9D3A109CA37E7F6EFE7AFCEC8A9FDA6A" box="[394,451,834,859]" pageId="19" pageNumber="320" type="female">15 ♀</specimenCount>
, same data as holotype (cJS,
<collectionCode id="ED2D43D0A37E7F6EFCE0FCED881FDA6B" box="[784,835,835,859]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="19" pageNumber="320">ZIN</collectionCode>
);
<specimenCount id="9D3A109CA37E7F6EFCA7FCED88D8DA6A" box="[855,900,835,859]" pageId="19" pageNumber="320" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D3A109CA37E7F6EFC60FCED88E7DA6A" box="[912,955,835,859]" pageId="19" pageNumber="320" type="female">1 ♀</specimenCount>
, “
<collectingCountry id="F32B9B85A37E7F6EFC23FCED8F72DA6B" box="[979,1070,835,859]" name="Nepal" pageId="19" pageNumber="320">NEPAL</collectingCountry>
HIMALAYA
<quantity id="4CC476F0A37E7F6EFB20FCED8E4BDA6B" box="[1232,1303,834,859]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="320" unit="km" value="20.0">20 km</quantity>
W
<location id="8EE38DCEA37E7F6EFAB0FCEC8BE7DA4E" LSID="urn:lsid:plazi:treatment:03956A03A37E7F68FF67FD3E8A85DC00:8EE38DCEA37E7F6EFAB0FCEC8BE7DA4E" country="Nepal" name="Pokhara Mt" pageId="19" pageNumber="320">Pokhara Mt</location>
</materialsCitation>
.
<materialsCitation id="3B54D148A37E7F6EFF35FCC8890DDA93" ID-GBIF-Occurrence="1655986855" collectingDate="1997-05-21" collectingDateMax="2015-06-18" collectingDateMin="1997-05-21" collectionCode="SMTD" collectorName="Pokhara &amp; Kande &amp; E. Kucera" country="Nepal" elevation="2400" location="Panchhase" pageId="19" pageNumber="320" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="8EE38DCEA37E7F6EFF35FCC88A1CDA4F" LSID="urn:lsid:plazi:treatment:03956A03A37E7F68FF67FD3E8A85DC00:8EE38DCEA37E7F6EFF35FCC88A1CDA4F" box="[197,320,870,895]" country="Nepal" name="Panchhase" pageId="19" pageNumber="320">Panchhase</location>
”, “NW slope,
<elevation id="00113C26A37E7F6EFE10FCC9896BDA4E" box="[480,567,871,895]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="19" pageNumber="320" unit="m" value="2400.0">
<quantity id="4CC476F0A37E7F6EFE10FCC9896BDA4E" box="[480,567,871,895]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="19" pageNumber="320" unit="m" value="2400.0">2400 m</quantity>
</elevation>
,
<collectingDate id="EFC6043DA37E7F6EFDB3FCC989E4DA4F" box="[579,696,871,895]" pageId="19" pageNumber="320" value="1997-05-21">
<date id="FF82FDD5A37E7F6EFDB3FCC989E4DA4F" box="[579,696,871,895]" pageId="19" pageNumber="320" value="1997-05-21">21.V.1997</date>
</collectingDate>
, lg. Jäger” (
<collectionCode id="ED2D43D0A37E7F6EFCCDFCC988CDDA4F" box="[829,913,871,895]" pageId="19" pageNumber="320">SMTD</collectionCode>
);
<specimenCount id="9D3A109CA37E7F6EFC54FCC9888CDA4E" box="[932,976,871,895]" pageId="19" pageNumber="320" type="male">1 ♂</specimenCount>
, “
<collectingCountry id="F32B9B85A37E7F6EFC15FCC98F1CDA4F" box="[997,1088,871,895]" name="Nepal" pageId="19" pageNumber="320">NEPAL</collectingCountry>
central,
<collectorName id="26C9BEC3A37E7F6EFB50FCC88FA0DA4F" box="[1184,1276,870,895]" pageId="19" pageNumber="320">Pokhara</collectorName>
–
<collectorName id="26C9BEC3A37E7F6EFAE8FCC88E3DDA4F" box="[1304,1377,870,895]" pageId="19" pageNumber="320">Kande</collectorName>
env.,
<collectingDate id="EFC6043DA37E7F6EFF67FC248A69DA93" box="[151,309,906,931]" pageId="19" pageNumber="320" value="2015-06-04" valueMax="2015-06-18" valueMin="2015-06-04">
<date id="FF82FDD5A37E7F6EFF67FC248A69DA93" box="[151,309,906,931]" pageId="19" pageNumber="320" value="2015-06-04" valueMax="2015-06-18" valueMin="2015-06-04">4–18.VI.2015</date>
</collectingDate>
, lgt.
<collectorName id="26C9BEC3A37E7F6EFE9CFC258A83DA93" box="[364,479,907,931]" pageId="19" pageNumber="320">E. Kučera</collectorName>
” (cKUČ).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C326889EA37E7F68FF37FC008848D8E0" lastPageId="21" lastPageNumber="322" pageId="19" pageNumber="320" type="description">
<paragraph id="8B83DB15A37E7F6EFF37FC008F0CDAF6" blockId="19.[151,1437,762,2011]" box="[199,1104,942,967]" pageId="19" pageNumber="320">
<emphasis id="B9480707A37E7F6EFF37FC008A0DDAF7" bold="true" box="[199,337,942,967]" pageId="19" pageNumber="320">Description</emphasis>
. Habitus as in
<figureCitation id="1307C790A37E7F6EFE08FC008917DAF7" box="[504,587,942,967]" captionStart="FIGURES 38 – 41" captionStartId="15.[151,264,1942,1965]" captionTargetBox="[273,1314,196,1918]" captionTargetId="figure@15.[271,1315,193,1921]" captionTargetPageId="15" captionText="FIGURES 38 – 41. Himalayan species of Trichotichnus subgenus Bottchrus, photographs of habitus. 38, T. schawalleri sp. n., holotype; 39, T. obliquebasalis sp. n., paratype, male; 40, T. siklesensis sp. n., paratype, male; 41, T. panchhaseensis sp. n., paratype, male." httpUri="https://zenodo.org/record/919620/files/figure.png" pageId="19" pageNumber="320">Fig. 41</figureCitation>
. Body length
<quantity id="4CC476F0A37E7F6EFD17FC008833DAF6" box="[743,879,942,967]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="6.6" metricValueMin="5.6" pageId="19" pageNumber="320" unit="mm" value="6.1" valueMax="6.6" valueMin="5.6">5.6–6.6 mm</quantity>
, width
<quantity id="4CC476F0A37E7F6EFC32FC018F10DAF6" box="[962,1100,943,967]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.8" metricValueMin="2.4" pageId="19" pageNumber="320" unit="mm" value="2.5999999999999996" valueMax="2.8" valueMin="2.4">2.4–2.8 mm</quantity>
.
</paragraph>
<paragraph id="8B83DB15A37E7F6EFF37FC7C8E61DD03" blockId="19.[151,1437,762,2011]" pageId="19" pageNumber="320">Colour: Body piceous black, rather shiny on dorsum, with distinct blue tinge on elytra and pronotum; labrum, in some specimens also base of mandibles, very narrow margins of pronotum and apical portion of elytral margins slightly paler; palpi, antennae and legs brownish yellow, femora and metatrochanteres distinctly infuscate.</paragraph>
<paragraph id="8B83DB15A37E7F6EFF37FB908A57DF42" blockId="19.[151,1437,762,2011]" pageId="19" pageNumber="320">
Head: Comparatively large, with wide neck (in males, HWmax/PWmax = 0.68–0.71, HWmin/PWmax = 0.53– 0.56; in females, these indices 0.71–0.73 and 0.56–0.59, respectively). Eyes moderately or markedly convex (in males, HWmax/HWmin = 1.25–1.28, and in females, 1.24–1.26), in lateral view wide oval, almost round, occasionally slightly elongate. Tempora about one third to half as long as eye, slightly convex, somewhat abruptly sloped to neck. Genae noticeably wider than width of antennomere 1, with few short setae. Lateral margin of head between antenna and eye, like in all the preceding species, slightly protruded laterally, forming an obtuse angle near eyes, distinctly bordered. Area between supraorbital furrow and upper margin of eye approximately as wide as or slightly shorter than width of antennomere 2 basally. Supraorbital setigerous pore located just behind level of posterior margin of eye, separated from supraorbital furrow approximately by distance slightly more than width of antennomere 3 apically. Labrum slightly concave anteriorly. Clypeus almost straight, only slightly emarginate along anterior margin, slightly convex basally, flatly sloped to apex, with a moderately large setigerous pore at each apical angle. Frons and vertex convex.
<collectingRegion id="49F815F7A37E7F6EFDA2FA6589EDDCD2" box="[594,689,1483,1506]" country="Myanmar" name="Chin" pageId="19" pageNumber="320">Mentum</collectingRegion>
and submentum completely fused, submentum with one long seta on each side, usually also with a short seta laterally of long seta. Left mandible not truncate, blunted at apex. Dorsal surface smooth, highly obliterate transverse meshes visible only laterally behind eyes. Antennae comparatively short, with antennomeres 4–8 slightly longer than wide, in male extended, in female not extended to basal edge of pronotum.
</paragraph>
<paragraph id="8B83DB15A37E7F69FF37F9D188B1D9E5" blockId="19.[151,1437,762,2011]" lastBlockId="20.[151,1437,151,717]" lastPageId="20" lastPageNumber="321" pageId="19" pageNumber="320">Pronotum: Moderately transverse (PWmax/PL = 1.30–1.37), widest at the end of anterior third, slightly or moderately narrowed posteriad (PWmax/PWmin = 1.21–1.30), with one lateral seta inserted slightly before the widest point. Sides rounded anteriorly, almost straight in basal third, and slightly sinuate in very short distance just before basal angles. Apical margin slightly emarginate, bordered almost throughout, border shortly interrupted at middle. Apical angles very slightly protruded anteriad, narrowly rounded at tip. Basal margin nearly straight, vaguely bordered laterally, slightly longer than apical margin and slightly shorter than base of elytra between humeral angles. Basal angles very slightly obtuse, subdenticulate. Pronotal disc moderately convex, strongly sloped to apical angles and moderately so to basal ones, not depressed basally, but slightly flattened at basal angles. Lateral gutter very narrow, indistinctly widened in apical half. Median line fine, not deepened, ended markedly before apical and basal margins. Basal foveae distinct, small, oval, very shallow. Surface sparsely, somewhat coarsely and irregularly punctate in basal and latero-basal areas before lateral seta, with few very fine punctures at apical margin. Pronotal disc with very obliterate microsculpture along sides.</paragraph>
<paragraph id="8B83DB15A3797F69FF37FF71885CDB0D" blockId="20.[151,1437,151,717]" pageId="20" pageNumber="321">Elytra: In lateral and caudal view convex, in dorsal view oval, comparatively elongate (in males, EL/EW = 1.41–1.47; EL/PL = 2.38–2.45; EW/PWmax = 1.24–1.27; in females, these indices 1.41–1.45, 2.43–2.48, 1.25– 1.29, respectively), widest slightly behind middle, with somewhat acute apex. Humeri prominent, angularly rounded at tip, without denticle. Subapical sinuation very shallow. Sutural angle rectangular, not blunted at tip. Basal edge nearly straight, slightly curved just at humerus, forming a sharp, obtuse angle with lateral margin. Striae slightly impressed along entire length. Intervals slightly convex throughout, weakly narrowed posteriad. Parascutellar setigerous pore large. Parascutellar striole short, at most as long as distance from parascutellar pore to suture, with apex free. Interval 3 with a small discal setigerous pore at stria 2 behind middle. Marginal umbilicate series widely interrupted at middle, with 6–8 pores in anterior group and 8–9 pores in posterior group. Microsculpture not evident even at high magnification.</paragraph>
<paragraph id="8B83DB15A3797F69FF37FDE9890FDB50" blockId="20.[151,1437,151,717]" box="[199,595,583,608]" pageId="20" pageNumber="321">Hindwings: Reduced to tiny scales.</paragraph>
<paragraph id="8B83DB15A3797F69FF37FDC2885DDBFD" blockId="20.[151,1437,151,717]" pageId="20" pageNumber="321">
Ventral surface: Prosternum and metasternum finely pubescent. Metepisternum (
<figureCitation id="1307C790A3797F69FBA1FDC28FFFDBB5" box="[1105,1187,620,645]" captionStart="FIGURES 53 – 59" captionStartId="20.[151,264,1678,1701]" captionTargetBox="[195,1394,776,1638]" captionTargetId="figure@20.[176,1411,769,1657]" captionTargetPageId="20" captionText="FIGURES 53 – 59. Trichotichnus (Bottchrus) panchhaseensis sp. n. (53 – 55, paratypes; 56 – 59, holotype). 53, left metepisternum; 54, apex of last (VII) abdominal sternite, male; 55, same, female; 56 – 57, median lobe of aedeagus; 58, terminal lamella; 59, apical capitulum. 53 – 55, ventral view; 56, lateral view; 57 – 58, dorsal view; 59, caudal view. Scale bars = 0.25 mm (A: Figs 53 – 57; B: Figs 58 – 59)." httpUri="https://zenodo.org/record/919632/files/figure.png" pageId="20" pageNumber="321">Fig. 53</figureCitation>
) markedly wider than long, moderately narrowed posteriad. Apex of last visible (VII) abdominal sternite in male widely rounded (
<figureCitation id="1307C790A3797F69FA9FFD218BEADBFD" captionStart="FIGURES 53 – 59" captionStartId="20.[151,264,1678,1701]" captionTargetBox="[195,1394,776,1638]" captionTargetId="figure@20.[176,1411,769,1657]" captionTargetPageId="20" captionText="FIGURES 53 – 59. Trichotichnus (Bottchrus) panchhaseensis sp. n. (53 – 55, paratypes; 56 – 59, holotype). 53, left metepisternum; 54, apex of last (VII) abdominal sternite, male; 55, same, female; 56 – 57, median lobe of aedeagus; 58, terminal lamella; 59, apical capitulum. 53 – 55, ventral view; 56, lateral view; 57 – 58, dorsal view; 59, caudal view. Scale bars = 0.25 mm (A: Figs 53 – 57; B: Figs 58 – 59)." httpUri="https://zenodo.org/record/919632/files/figure.png" pageId="20" pageNumber="321">Fig. 54</figureCitation>
), in female narrowly or angularly rounded (
<figureCitation id="1307C790A3797F69FD50FD1A89A9DBFD" box="[672,757,692,717]" captionStart="FIGURES 53 – 59" captionStartId="20.[151,264,1678,1701]" captionTargetBox="[195,1394,776,1638]" captionTargetId="figure@20.[176,1411,769,1657]" captionTargetPageId="20" captionText="FIGURES 53 – 59. Trichotichnus (Bottchrus) panchhaseensis sp. n. (53 – 55, paratypes; 56 – 59, holotype). 53, left metepisternum; 54, apex of last (VII) abdominal sternite, male; 55, same, female; 56 – 57, median lobe of aedeagus; 58, terminal lamella; 59, apical capitulum. 53 – 55, ventral view; 56, lateral view; 57 – 58, dorsal view; 59, caudal view. Scale bars = 0.25 mm (A: Figs 53 – 57; B: Figs 58 – 59)." httpUri="https://zenodo.org/record/919632/files/figure.png" pageId="20" pageNumber="321">Fig. 55</figureCitation>
).
</paragraph>
<caption id="DF438B9DA3797F69FF67F9208A81DE30" httpUri="https://zenodo.org/record/919632/files/figure.png" pageId="20" pageNumber="321" targetBox="[195,1394,776,1638]" targetPageId="20">
<paragraph id="8B83DB15A3797F69FF67F9208A81DE30" blockId="20.[151,1436,1678,1792]" pageId="20" pageNumber="321">
<emphasis id="B9480707A3797F69FF67F9208A03DF94" bold="true" box="[151,351,1678,1701]" pageId="20" pageNumber="321">FIGURES 53–59.</emphasis>
<taxonomicName id="4C3CA096A3797F69FE81F920886FDF94" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[369,819,1678,1700]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="panchhaseensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3797F69FE81F9208AA0DF94" box="[369,508,1678,1700]" italics="true" pageId="20" pageNumber="321">Trichotichnus</emphasis>
(
<emphasis id="B9480707A3797F69FDE7F9208926DF94" box="[535,634,1678,1700]" italics="true" pageId="20" pageNumber="321">Bottchrus</emphasis>
)
<emphasis id="B9480707A3797F69FD64F920886FDF94" box="[660,819,1678,1700]" italics="true" pageId="20" pageNumber="321">panchhaseensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3797F69FCB5F92188D6DF94" box="[837,906,1679,1700]" pageId="20" pageNumber="321" rank="species">
<emphasis id="B9480707A3797F69FCB5F92188D6DF94" bold="true" box="[837,906,1679,1700]" pageId="20" pageNumber="321">sp. n.</emphasis>
</taxonomicNameLabel>
(53–55, paratypes; 56–59, holotype). 53, left metepisternum; 54, apex of last (VII) abdominal sternite, male; 55, same, female; 56–57, median lobe of aedeagus; 58, terminal lamella; 59, apical capitulum. 53–55, ventral view; 56, lateral view; 57–58, dorsal view; 59, caudal view. Scale bars = 0.25 mm (A: Figs 53–57; B: Figs 58–59).
</paragraph>
</caption>
<paragraph id="8B83DB15A3797F69FF37F8868BA2DEE1" blockId="20.[151,1437,1831,2037]" pageId="20" pageNumber="321">Legs: Protibia very finely longitudinally sulcate in basal third, occasionally sulcus extremely short. Profemur with five to seven setae on anterio-ventral margin. Tarsi short, metatarsus noticeably shorter than HWmin, with metatarsomeres (particularly 2–4) markedly widened posteriad, metatarsomere 1 approximately as long as metatarsomeres 2+3. Tarsomere 5 with two pairs of ventro-lateral setae. In male, pro- and mesotarsi moderately widened.</paragraph>
<paragraph id="8B83DB15A3797F68FF37F8728952D9E5" blockId="20.[151,1437,1831,2037]" lastBlockId="21.[151,1437,151,1328]" lastPageId="21" lastPageNumber="322" pageId="20" pageNumber="321">Female genitalia: Laterotergite (hemisternite) symmetrical, with two thick setae apically. Basal stylomere with one preapical spine on external margin. Apical stylomere moderately curved, with a peg-like spine at both ventral and dorsal edges of outer margin.</paragraph>
<paragraph id="8B83DB15A3787F68FF37FF718848D8E0" blockId="21.[151,1437,151,1328]" pageId="21" pageNumber="322">
Male genitalia: Median lobe of aedeagus (
<figureCitation id="1307C790A3787F68FD6FFF71887CD9C8" box="[671,800,223,248]" captionStart="FIGURES 53 – 59" captionStartId="20.[151,264,1678,1701]" captionTargetBox="[195,1394,776,1638]" captionTargetId="figure@20.[176,1411,769,1657]" captionTargetPageId="20" captionText="FIGURES 53 – 59. Trichotichnus (Bottchrus) panchhaseensis sp. n. (53 – 55, paratypes; 56 – 59, holotype). 53, left metepisternum; 54, apex of last (VII) abdominal sternite, male; 55, same, female; 56 – 57, median lobe of aedeagus; 58, terminal lamella; 59, apical capitulum. 53 – 55, ventral view; 56, lateral view; 57 – 58, dorsal view; 59, caudal view. Scale bars = 0.25 mm (A: Figs 53 – 57; B: Figs 58 – 59)." httpUri="https://zenodo.org/record/919632/files/figure.png" pageId="21" pageNumber="322">Figs 56–59</figureCitation>
) in lateral view arcuate, C-shaped, strongly bent ventrad just after basal bulb and not bent ventrad in its apical third, with slightly convex ventral margin in its apical half; in dorsal view slightly curved to right, somewhat evenly narrowed to apex in distal half. Terminal lamella (
<figureCitation id="1307C790A3787F68FAB1FE898EC8D870" box="[1345,1428,295,320]" captionStart="FIGURES 53 – 59" captionStartId="20.[151,264,1678,1701]" captionTargetBox="[195,1394,776,1638]" captionTargetId="figure@20.[176,1411,769,1657]" captionTargetPageId="20" captionText="FIGURES 53 – 59. Trichotichnus (Bottchrus) panchhaseensis sp. n. (53 – 55, paratypes; 56 – 59, holotype). 53, left metepisternum; 54, apex of last (VII) abdominal sternite, male; 55, same, female; 56 – 57, median lobe of aedeagus; 58, terminal lamella; 59, apical capitulum. 53 – 55, ventral view; 56, lateral view; 57 – 58, dorsal view; 59, caudal view. Scale bars = 0.25 mm (A: Figs 53 – 57; B: Figs 58 – 59)." httpUri="https://zenodo.org/record/919632/files/figure.png" pageId="21" pageNumber="322">Fig. 58</figureCitation>
) short, narrowed to apex, slightly sinuate at sides just before capitulum. Apical capitulum in lateral view (
<figureCitation id="1307C790A3787F68FAB1FEE28EC8D855" box="[1345,1428,332,357]" captionStart="FIGURES 53 – 59" captionStartId="20.[151,264,1678,1701]" captionTargetBox="[195,1394,776,1638]" captionTargetId="figure@20.[176,1411,769,1657]" captionTargetPageId="20" captionText="FIGURES 53 – 59. Trichotichnus (Bottchrus) panchhaseensis sp. n. (53 – 55, paratypes; 56 – 59, holotype). 53, left metepisternum; 54, apex of last (VII) abdominal sternite, male; 55, same, female; 56 – 57, median lobe of aedeagus; 58, terminal lamella; 59, apical capitulum. 53 – 55, ventral view; 56, lateral view; 57 – 58, dorsal view; 59, caudal view. Scale bars = 0.25 mm (A: Figs 53 – 57; B: Figs 58 – 59)." httpUri="https://zenodo.org/record/919632/files/figure.png" pageId="21" pageNumber="322">Fig. 56</figureCitation>
) more markedly prominent dorsally than ventrally, strongly oblique, with ventral flange situated much more proximally than dorsal flange, in caudal view (
<figureCitation id="1307C790A3787F68FD44FE3A8854D89D" box="[692,776,404,429]" captionStart="FIGURES 53 – 59" captionStartId="20.[151,264,1678,1701]" captionTargetBox="[195,1394,776,1638]" captionTargetId="figure@20.[176,1411,769,1657]" captionTargetPageId="20" captionText="FIGURES 53 – 59. Trichotichnus (Bottchrus) panchhaseensis sp. n. (53 – 55, paratypes; 56 – 59, holotype). 53, left metepisternum; 54, apex of last (VII) abdominal sternite, male; 55, same, female; 56 – 57, median lobe of aedeagus; 58, terminal lamella; 59, apical capitulum. 53 – 55, ventral view; 56, lateral view; 57 – 58, dorsal view; 59, caudal view. Scale bars = 0.25 mm (A: Figs 53 – 57; B: Figs 58 – 59)." httpUri="https://zenodo.org/record/919632/files/figure.png" pageId="21" pageNumber="322">Fig. 59</figureCitation>
) somewhat elongate oval. Internal sac with characteristic pattern of longitudinal spiny folds, without larger spines.
</paragraph>
</subSubSection>
<subSubSection id="C326889EA3787F68FF37FE7288F9DDF5" pageId="21" pageNumber="322" type="etymology">
<paragraph id="8B83DB15A3787F68FF37FE728AB6DB27" blockId="21.[151,1437,151,1328]" pageId="21" pageNumber="322">
<emphasis id="B9480707A3787F68FF37FE728A1AD8C5" bold="true" box="[199,326,476,501]" pageId="21" pageNumber="322">Etymology</emphasis>
. The specific name refers to the Mt. Panchhase on the southern slope of the Annapurna Himal where the new species occurs.
</paragraph>
<paragraph id="8B83DB15A3787F68FF37FD8A88F9DDF5" blockId="21.[151,1437,151,1328]" pageId="21" pageNumber="322">
<emphasis id="B9480707A3787F68FF37FD8A8A06DB0D" bold="true" box="[199,346,548,573]" pageId="21" pageNumber="322">Comparison</emphasis>
. In size, general appearance and short metepisterna,
<taxonomicName id="4C3CA096A3787F68FC5EFD8B8FF0DB0D" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[942,1196,548,573]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="panchhaseensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3787F68FC5EFD8B889EDB0C" box="[942,962,549,572]" italics="true" pageId="21" pageNumber="322">T.</emphasis>
(
<emphasis id="B9480707A3787F68FC24FD8B88B9DB0C" box="[980,997,549,572]" italics="true" pageId="21" pageNumber="322">B</emphasis>
.)
<emphasis id="B9480707A3787F68FC0CFD8A8FF0DB0D" box="[1020,1196,548,573]" italics="true" pageId="21" pageNumber="322">panchhaseensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3787F68FB44FD8A8FA8DB0D" box="[1204,1268,548,573]" pageId="21" pageNumber="322" rank="species">
<emphasis id="B9480707A3787F68FB44FD8A8FA8DB0D" bold="true" box="[1204,1268,548,573]" pageId="21" pageNumber="322">sp. n.</emphasis>
</taxonomicNameLabel>
is similar to
<taxonomicName id="4C3CA096A3787F68FA77FD8B8A18DB50" authorityName="Kirschenhofer" authorityYear="1992" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="holzschuhi" subGenus="Bottchrus">
<emphasis id="B9480707A3787F68FA77FD8B8EC1DB0C" box="[1415,1437,549,572]" italics="true" pageId="21" pageNumber="322">T.</emphasis>
(
<emphasis id="B9480707A3787F68FF50FDE78BEDDB50" box="[160,177,585,608]" italics="true" pageId="21" pageNumber="322">B</emphasis>
.)
<emphasis id="B9480707A3787F68FF3CFDE98A18DB50" box="[204,324,583,608]" italics="true" pageId="21" pageNumber="322">holzschuhi</emphasis>
</taxonomicName>
and the other species described above, but may be discriminated among them in the following combination of the distinctive characters: body comparatively more elongate, with relatively long pronotum and elytra; pronotal base straight nearly throughout, not oblique laterally; protibia very finely longitudinally sulcate in basal third; median lobe of aedeagus arcuate but not bent ventrad in its apical portion, with convex ventral margin in apical half and with very short terminal lamella (
<figureCitation id="1307C790A3787F68FD04FD798827DBC0" box="[756,891,727,752]" captionStart="FIGURES 53 – 59" captionStartId="20.[151,264,1678,1701]" captionTargetBox="[195,1394,776,1638]" captionTargetId="figure@20.[176,1411,769,1657]" captionTargetPageId="20" captionText="FIGURES 53 – 59. Trichotichnus (Bottchrus) panchhaseensis sp. n. (53 – 55, paratypes; 56 – 59, holotype). 53, left metepisternum; 54, apex of last (VII) abdominal sternite, male; 55, same, female; 56 – 57, median lobe of aedeagus; 58, terminal lamella; 59, apical capitulum. 53 – 55, ventral view; 56, lateral view; 57 – 58, dorsal view; 59, caudal view. Scale bars = 0.25 mm (A: Figs 53 – 57; B: Figs 58 – 59)." httpUri="https://zenodo.org/record/919632/files/figure.png" pageId="21" pageNumber="322">Figs 56–58</figureCitation>
); its apical capitulum more strongly prominent dorsally than ventrally, with ventral flange situated much more proximally than dorsal flange, in caudal view (
<figureCitation id="1307C790A3787F68FA9FFD528BE8DA08" captionStart="FIGURES 53 – 59" captionStartId="20.[151,264,1678,1701]" captionTargetBox="[195,1394,776,1638]" captionTargetId="figure@20.[176,1411,769,1657]" captionTargetPageId="20" captionText="FIGURES 53 – 59. Trichotichnus (Bottchrus) panchhaseensis sp. n. (53 – 55, paratypes; 56 – 59, holotype). 53, left metepisternum; 54, apex of last (VII) abdominal sternite, male; 55, same, female; 56 – 57, median lobe of aedeagus; 58, terminal lamella; 59, apical capitulum. 53 – 55, ventral view; 56, lateral view; 57 – 58, dorsal view; 59, caudal view. Scale bars = 0.25 mm (A: Figs 53 – 57; B: Figs 58 – 59)." httpUri="https://zenodo.org/record/919632/files/figure.png" pageId="21" pageNumber="322">Fig. 59</figureCitation>
) somewhat elongate oval; internal sac without spines. The most distinctive characters also include mentum completely fused with submentum, pronotal lateral gutters slightly widened in apical half, microsculpture on pronotal and elytral discs not developed, humeral denticle absent, last visible (VII) abdominal sternite in female narrowly or angularly rounded at apex (
<figureCitation id="1307C790A3787F68FD54FC228858DA95" box="[676,772,908,933]" captionStart="FIGURES 53 – 59" captionStartId="20.[151,264,1678,1701]" captionTargetBox="[195,1394,776,1638]" captionTargetId="figure@20.[176,1411,769,1657]" captionTargetPageId="20" captionText="FIGURES 53 – 59. Trichotichnus (Bottchrus) panchhaseensis sp. n. (53 – 55, paratypes; 56 – 59, holotype). 53, left metepisternum; 54, apex of last (VII) abdominal sternite, male; 55, same, female; 56 – 57, median lobe of aedeagus; 58, terminal lamella; 59, apical capitulum. 53 – 55, ventral view; 56, lateral view; 57 – 58, dorsal view; 59, caudal view. Scale bars = 0.25 mm (A: Figs 53 – 57; B: Figs 58 – 59)." httpUri="https://zenodo.org/record/919632/files/figure.png" pageId="21" pageNumber="322">Fig. 55</figureCitation>
), and femora distinctly infuscate.
<taxonomicName id="4C3CA096A3787F68FB33FC228A1BDAF8" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="panchhaseensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3787F68FB33FC228E00DA95" box="[1219,1372,908,933]" italics="true" pageId="21" pageNumber="322">Trichotichnus</emphasis>
(
<emphasis id="B9480707A3787F68FA8AFC238ED7DA94" box="[1402,1419,909,932]" italics="true" pageId="21" pageNumber="322">B</emphasis>
.)
<emphasis id="B9480707A3787F68FF67FC018A1BDAF8" box="[151,327,943,968]" italics="true" pageId="21" pageNumber="322">panchhaseensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3787F68FEBFFC018ACCDAF8" box="[335,400,943,968]" pageId="21" pageNumber="322" rank="species">
<emphasis id="B9480707A3787F68FEBFFC018ACCDAF8" bold="true" box="[335,400,943,968]" pageId="21" pageNumber="322">sp. n.</emphasis>
</taxonomicNameLabel>
distinctly differs from the sympatric
<taxonomicName id="4C3CA096A3787F68FCCCFC1F88B2DAF8" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[828,1006,943,968]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="parvulus" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3787F68FCCCFC1F880DDAF8" box="[828,849,945,968]" italics="true" pageId="21" pageNumber="322">T.</emphasis>
(
<emphasis id="B9480707A3787F68FC92FC1F882FDAF8" box="[866,883,945,968]" italics="true" pageId="21" pageNumber="322">B</emphasis>
.)
<emphasis id="B9480707A3787F68FC7CFC0188B2DAF8" box="[908,1006,943,968]" italics="true" pageId="21" pageNumber="322">parvulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3787F68FC07FC018F64DAF8" box="[1015,1080,943,968]" pageId="21" pageNumber="322" rank="species">
<emphasis id="B9480707A3787F68FC07FC018F64DAF8" bold="true" box="[1015,1080,943,968]" pageId="21" pageNumber="322">sp. n.</emphasis>
</taxonomicNameLabel>
in having greater size (
<quantity id="4CC476F0A3787F68FAB6FC018B9FDADC" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="6.6" metricValueMin="5.6" pageId="21" pageNumber="322" unit="mm" value="6.1" valueMax="6.6" valueMin="5.6">5.6–6.6 mm</quantity>
versus
<quantity id="4CC476F0A3787F68FEE7FC7A8AFDDADC" box="[279,417,980,1005]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.4" metricValueMin="4.8" pageId="21" pageNumber="322" unit="mm" value="5.1" valueMax="5.4" valueMin="4.8">4.8–5.4 mm</quantity>
), mentum and submentum completely fused [in
<taxonomicName id="4C3CA096A3787F68FC49FC7B8F3BDADD" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[953,1127,980,1005]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="parvulus" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3787F68FC49FC7B8892DADC" box="[953,974,981,1004]" italics="true" pageId="21" pageNumber="322">T.</emphasis>
(
<emphasis id="B9480707A3787F68FC2DFC7B88B2DADC" box="[989,1006,981,1004]" italics="true" pageId="21" pageNumber="322">B</emphasis>
.)
<emphasis id="B9480707A3787F68FBF5FC7A8F3BDADD" box="[1029,1127,980,1005]" italics="true" pageId="21" pageNumber="322">parvulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3787F68FB9DFC7A8FF0DADD" box="[1133,1196,980,1005]" pageId="21" pageNumber="322" rank="species">
<emphasis id="B9480707A3787F68FB9DFC7A8FF0DADD" bold="true" box="[1133,1196,980,1005]" pageId="21" pageNumber="322">sp. n.</emphasis>
</taxonomicNameLabel>
, separated medially], protibia finely sulcate, elytral sutural angle less strongly produced posteriad, almost rectangular at apex, last visible (VII) abdominal sternite in female narrowly or angularly rounded at apex [in
<taxonomicName id="4C3CA096A3787F68FBD9FBB38FF7DD05" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[1065,1195,1052,1077]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="parvulus" status="sp. nov.">
<emphasis id="B9480707A3787F68FBD9FBB38FF7DD05" box="[1065,1195,1052,1077]" italics="true" pageId="21" pageNumber="322">T. parvulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3787F68FB46FBB28FA5DD05" box="[1206,1273,1052,1077]" pageId="21" pageNumber="322" rank="species">
<emphasis id="B9480707A3787F68FB46FBB28FA5DD05" bold="true" box="[1206,1273,1052,1077]" pageId="21" pageNumber="322">sp. n.</emphasis>
</taxonomicNameLabel>
, more widely rounded (
<figureCitation id="1307C790A3787F68FEF2FB918A0BDD68" box="[258,343,1087,1112]" captionStart="FIGURES 22 – 31" captionStartId="10.[151,264,1454,1477]" captionTargetBox="[159,1414,724,1410]" captionTargetId="figure@10.[151,1436,707,1434]" captionTargetPageId="10" captionText="FIGURES 22 – 31. Trichotichnus (Bottchrus) parvulus sp. n. (22 – 24, 28, paratypes from Mt. Panchhase; 25 – 27, holotype; 29 – 31, paratype from Krapa Danda mountain range). 22, mentum and submentum; 23, apex of last (VII) abdominal sternite, male; 24, same, female; 25, 27, 29, 31, median lobe of aedeagus; 26, 30, terminal lamella and apical capitulum of aedeagus; 28, left metepisternum. 22, 24, 28, ventral view; 25, 29, lateral view; 27, 31, dorsal view; 26, 30, caudal view. Scale bars = 0.25 mm (A: Figs 22 – 25, 27 – 29, 31; B: Figs 26 – 30)." httpUri="https://zenodo.org/record/919614/files/figure.png" pageId="21" pageNumber="322">Fig. 24</figureCitation>
)], and the penis structure as described above [for comparison, the median lobe of
<taxonomicName id="4C3CA096A3787F68FB1CFBEF8EC0DD68" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[1260,1436,1087,1112]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="parvulus" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3787F68FB1CFBEF8E5CDD68" box="[1260,1280,1089,1112]" italics="true" pageId="21" pageNumber="322">T.</emphasis>
(
<emphasis id="B9480707A3787F68FAE1FBEF8E7EDD68" box="[1297,1314,1089,1112]" italics="true" pageId="21" pageNumber="322">B</emphasis>
.)
<emphasis id="B9480707A3787F68FACAFB918EC0DD68" box="[1338,1436,1087,1112]" italics="true" pageId="21" pageNumber="322">parvulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3787F68FF67FBCA8B84DD4D" box="[151,216,1124,1149]" pageId="21" pageNumber="322" rank="species">
<emphasis id="B9480707A3787F68FF67FBCA8B84DD4D" bold="true" box="[151,216,1124,1149]" pageId="21" pageNumber="322">sp. n.</emphasis>
</taxonomicNameLabel>
with longer terminal lamella, its apical capitulum in lateral view oblique, with acute dorsal flange located less distally than rounded ventral flange (
<figureCitation id="1307C790A3787F68FD99FB2989AEDD90" box="[617,754,1159,1184]" captionStart="FIGURES 22 – 31" captionStartId="10.[151,264,1454,1477]" captionTargetBox="[159,1414,724,1410]" captionTargetId="figure@10.[151,1436,707,1434]" captionTargetPageId="10" captionText="FIGURES 22 – 31. Trichotichnus (Bottchrus) parvulus sp. n. (22 – 24, 28, paratypes from Mt. Panchhase; 25 – 27, holotype; 29 – 31, paratype from Krapa Danda mountain range). 22, mentum and submentum; 23, apex of last (VII) abdominal sternite, male; 24, same, female; 25, 27, 29, 31, median lobe of aedeagus; 26, 30, terminal lamella and apical capitulum of aedeagus; 28, left metepisternum. 22, 24, 28, ventral view; 25, 29, lateral view; 27, 31, dorsal view; 26, 30, caudal view. Scale bars = 0.25 mm (A: Figs 22 – 25, 27 – 29, 31; B: Figs 26 – 30)." httpUri="https://zenodo.org/record/919614/files/figure.png" pageId="21" pageNumber="322">Figs. 25, 29</figureCitation>
), in caudal view apical capitulum somewhat triangular (
<figureCitation id="1307C790A3787F68FA9AFB298BBCDDF4" captionStart="FIGURES 22 – 31" captionStartId="10.[151,264,1454,1477]" captionTargetBox="[159,1414,724,1410]" captionTargetId="figure@10.[151,1436,707,1434]" captionTargetPageId="10" captionText="FIGURES 22 – 31. Trichotichnus (Bottchrus) parvulus sp. n. (22 – 24, 28, paratypes from Mt. Panchhase; 25 – 27, holotype; 29 – 31, paratype from Krapa Danda mountain range). 22, mentum and submentum; 23, apex of last (VII) abdominal sternite, male; 24, same, female; 25, 27, 29, 31, median lobe of aedeagus; 26, 30, terminal lamella and apical capitulum of aedeagus; 28, left metepisternum. 22, 24, 28, ventral view; 25, 29, lateral view; 27, 31, dorsal view; 26, 30, caudal view. Scale bars = 0.25 mm (A: Figs 22 – 25, 27 – 29, 31; B: Figs 26 – 30)." httpUri="https://zenodo.org/record/919614/files/figure.png" pageId="21" pageNumber="322">Figs 26, 30</figureCitation>
), and internal sac with a small group of small spines medially].
</paragraph>
</subSubSection>
<subSubSection id="C326889EA3787F68FF37FB618A85DC00" pageId="21" pageNumber="322" type="distribution">
<paragraph id="8B83DB15A3787F68FF37FB618A85DC00" blockId="21.[151,1437,151,1328]" pageId="21" pageNumber="322">
<emphasis id="B9480707A3787F68FF37FB618A05DDD8" bold="true" box="[199,345,1231,1256]" pageId="21" pageNumber="322">Distribution</emphasis>
(
<figureCitation id="1307C790A3787F68FE9BFB618A93DDD8" box="[363,463,1231,1256]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="21" pageNumber="322">Fig. 151</figureCitation>
,
<figureCitation id="1307C790A3787F68FE2DFB7E8AB7DDD7" box="[477,491,1232,1255]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="21" pageNumber="322">
<emphasis id="B9480707A3787F68FE2DFB7E8AB7DDD7" box="[477,491,1232,1255]" italics="true" pageId="21" pageNumber="322">7</emphasis>
</figureCitation>
). Slopes of Mt. Panchhase approximately
<quantity id="4CC476F0A3787F68FC29FB7E8F7FDDD8" box="[985,1059,1231,1256]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="322" unit="km" value="20.0">20 km</quantity>
west of Pokhara city in western
<collectingRegion id="49F815F7A3787F68FF67FB5A8BB6DC3D" box="[151,234,1268,1293]" country="Sri Lanka" name="Central" pageId="21" pageNumber="322">Central</collectingRegion>
<collectingCountry id="F32B9B85A3787F68FF04FB5A8A66DC3D" box="[244,314,1268,1293]" name="Nepal" pageId="21" pageNumber="322">Nepal</collectingCountry>
, Kaski District, at altitudes of
<quantity id="4CC476F0A3787F68FD59FB5A8817DC3C" box="[681,843,1268,1292]" metricMagnitude="3" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="21" pageNumber="322" unit="m" value="2200.0" valueMax="2400.0" valueMin="2000.0">2000–2400 m</quantity>
. The species occurs there sympatricly with
<taxonomicName id="4C3CA096A3787F68FAA1FB5B8BA5DC00" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="parvulus" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3787F68FAA1FB5B8E3ADC3C" box="[1361,1382,1269,1292]" italics="true" pageId="21" pageNumber="322">T.</emphasis>
(
<emphasis id="B9480707A3787F68FA8AFB5B8ED7DC3C" box="[1402,1419,1269,1292]" italics="true" pageId="21" pageNumber="322">B</emphasis>
.)
<emphasis id="B9480707A3787F68FF67FAB98BA5DC00" box="[151,249,1303,1328]" italics="true" pageId="21" pageNumber="322">parvulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3787F68FEF0FAB98A63DC00" box="[256,319,1303,1328]" pageId="21" pageNumber="322" rank="species">
<emphasis id="B9480707A3787F68FEF0FAB98A63DC00" bold="true" box="[256,319,1303,1328]" pageId="21" pageNumber="322">sp. n.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="1307C790A3787F68FEBEFAB98AF3DC00" box="[334,431,1303,1328]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="21" pageNumber="322">Fig. 151</figureCitation>
,
<emphasis id="B9480707A3787F68FE4AFAB68A94DC00" box="[442,456,1304,1328]" italics="true" pageId="21" pageNumber="322">3</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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