606 lines
70 KiB
XML
606 lines
70 KiB
XML
<document id="4F08E8275A70E33A2087DEF37FF8524F" ID-DOI="10.11646/zootaxa.3641.4.8" ID-GBIF-Dataset="f472225b-2aaf-4e42-ad50-f464340a15f6" ID-ISSN="1175-5326" ID-Zenodo-Dep="222413" ID-ZooBank="1D35547B-9E6C-4A18-832A-E4C8B5A31B40" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460596226550" checkinUser="plazi" docAuthor="Lee, B. Y., Ng, N. K. & Ng, L." docDate="2013" docId="03F5333FFFD9FF0A6CF047B3FF6BFAB2" docLanguage="en" docName="zt03641p432.pdf" docOrigin="Zootaxa 3641 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clistocoeloma balansae A. Milne-Edwards 1873" docType="treatment" docVersion="7" lastPageNumber="428" masterDocId="FFCC4B47FFD8FF026C67451EFFBDFB1E" masterDocTitle="On the identity of Clistocoeloma balansae A. Milne-Edwards, 1873, and C. tectum (Rathbun, 1914), with description of a new species from the West Pacific (Crustacea: Decapoda: Sesarmidae)" masterLastPageNumber="432" masterPageNumber="420" pageNumber="421" updateTime="1698326858605" updateUser="plazi">
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<mods:title id="1C44276588DE505D890D29E6F9C2E78F">On the identity of Clistocoeloma balansae A. Milne-Edwards, 1873, and C. tectum (Rathbun, 1914), with description of a new species from the West Pacific (Crustacea: Decapoda: Sesarmidae)</mods:title>
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<mods:namePart id="0D5536CB945F010CA1990B3C3093553B">Lee, B. Y.</mods:namePart>
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<mods:namePart id="62563A5DD7DF67B1B307268CA5AED969">Ng, N. K.</mods:namePart>
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<mods:namePart id="B2B0323DEDA4F7E8690B774889B2DAAB">Ng, L.</mods:namePart>
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<mods:typeOfResource id="56E86C9FB6713E2055B3BC3DD9AA61BA">text</mods:typeOfResource>
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<mods:title id="D40754D4B0249EF72F8C0F7F82CD8649">Zootaxa</mods:title>
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<mods:part id="4447630FD6E441ED77D1DAA2C4689B92">
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<mods:date id="4BEAE4DA6BEB41670D93AE0EFAC88096">2013</mods:date>
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<mods:number id="F052D3E14EB24C315066DC0A22093CCA">3641</mods:number>
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<mods:classification id="AF559AFC632273CDE9C4F3625CD865A5">journal article</mods:classification>
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<mods:identifier id="8F66E1B0BE8341001EF9C1711A80BEFD" type="DOI">10.11646/zootaxa.3641.4.8</mods:identifier>
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<treatment id="03F5333FFFD9FF0A6CF047B3FF6BFAB2" ID-DOI="http://doi.org/10.5281/zenodo.6147166" ID-GBIF-Taxon="119566892" ID-Zenodo-Dep="6147166" LSID="urn:lsid:plazi:treatment:03F5333FFFD9FF0A6CF047B3FF6BFAB2" httpUri="http://treatment.plazi.org/id/03F5333FFFD9FF0A6CF047B3FF6BFAB2" lastPageId="8" lastPageNumber="428" pageId="1" pageNumber="421">
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<subSubSection id="C346D1A2FFD9FF036CF047B3FEF8F852" pageId="1" pageNumber="421" type="nomenclature">
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<paragraph id="8BE38229FFD9FF036CF047B3FD51F9D9" blockId="1.[151,748,685,744]" box="[151,748,685,711]" pageId="1" pageNumber="421">
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<heading id="D0AB3545FFD9FF036CF047B3FD51F9D9" bold="true" box="[151,748,685,711]" fontSize="11" level="1" pageId="1" pageNumber="421" reason="1">
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<taxonomicName id="4C5CF9AAFFD9FF036CF047B3FD51F9D9" ID-CoL="W7TZ" authority="A. Milne-Edwards, 1873" authorityName="A. Milne-Edwards" authorityYear="1873" box="[151,748,685,711]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="421" phylum="Arthropoda" rank="species" species="balansae">
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<emphasis id="B9285E3BFFD9FF036CF047B3FD51F9D9" bold="true" box="[151,748,685,711]" pageId="1" pageNumber="421">
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<emphasis id="B9285E3BFFD9FF036CF047B3FE08F9D9" bold="true" box="[151,437,685,711]" italics="true" pageId="1" pageNumber="421">Clistocoeloma balansae</emphasis>
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A. Milne-Edwards, 1873
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BE38229FFD9FF036CF047CEFEAFF9F6" blockId="1.[151,748,685,744]" box="[151,274,720,744]" pageId="1" pageNumber="421">
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(
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<figureCitation id="13679EACFFD9FF036CF847CEFEB4F9F6" box="[159,265,720,744]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="2.[151,250,1910,1933]" captionStartId-1="3.[151,250,1726,1749]" captionStartId-2="4.[151,250,1704,1727]" captionStartId-3="5.[151,250,1982,2005]" captionStartId-4="6.[151,250,1124,1147]" captionStartId-5="7.[151,250,1880,1903]" captionTargetBox-0="[188,1398,301,1888]" captionTargetBox-1="[188,1398,193,1703]" captionTargetBox-2="[188,1398,176,1685]" captionTargetBox-3="[302,1285,373,1961]" captionTargetBox-4="[172,1368,241,1053]" captionTargetBox-5="[216,1398,1033,1780]" captionTargetId-0="figure@2.[188,1398,301,1889]" captionTargetId-1="figure@3.[188,1398,193,1705]" captionTargetId-2="figure@4.[188,1398,175,1687]" captionTargetId-3="figure@5.[302,1285,373,1961]" captionTargetId-4="figure@6.[151,1436,176,1121]" captionTargetId-5="figure@7.[188,1398,951,1858]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionText-0="FIGURE 1. Clistocoeloma balansae A. Milne-Edwards, 1873, lectotype male (20.7 × 19.7 mm) (MNHN B 10912), New Caledonia. A, dorsal overall view; B, ventral view of abdomen and thoracic sternum; C, frontal view of carapace. (Photographs by T. Naruse)" captionText-1="FIGURE 2. Sesarma (Sesarm a) tectum Rathbun, 1914, holotype female (19.5 × 18.6 mm) (USNM 45766), Philippines. A, dorsal overall view; B, ventral view of abdomen and thoracic sternum; C, frontal view of carapace; D, posterior view of carapace." captionText-2="FIGURE 3. Clistocoeloma balansae A. Milne-Edwards, 1873. A – C, male (15.9 × 15.5 mm) (ZRC 2012.702), Philippines; D, female (17.2 × 16.9 mm) (ZRC 2012.702), Philippines. A, dorsal overall view; B, ventral view of abdomen and thoracic sternum; C, D posterior view of carapace." captionText-3="FIGURE 4. Clistocoeloma balansae A. Milne-Edwards, 1873, male (15.9 × 15.5 mm) (ZRC 2012.702), Philippines. A, outer view of left chela; B, inner view of left chela; C, dorsal marginal view of left chela." captionText-4="FIGURE 5. Clistocoeloma balansae A. Milne-Edwards, 1873, male (15.9 × 15.5 mm) (ZRC 2012.702), Philippines. A, male abdomen; B, G 1 dorsal view; C, G 1 lateral view; D, G 1 ventral view. G 1 denuded." captionText-5="FIGURE 6. Clistocoeloma balansae A. Milne-Edwards, 1873 (after A. Milne-Edwards 1873: pl. 17 fig. 1). A, dorsal overall view; B, frontal view of carapace; C, male abdomen; D, outer view of left chela." httpUri-0="https://zenodo.org/record/222414/files/figure.png" httpUri-1="https://zenodo.org/record/222415/files/figure.png" httpUri-2="https://zenodo.org/record/222416/files/figure.png" httpUri-3="https://zenodo.org/record/222417/files/figure.png" httpUri-4="https://zenodo.org/record/222418/files/figure.png" httpUri-5="https://zenodo.org/record/222419/files/figure.png" pageId="1" pageNumber="421">Figs. 1–6</figureCitation>
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)
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</paragraph>
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<paragraph id="8BE38229FFD9FF036CF04606FEF8F852" blockId="1.[151,1437,792,1121]" pageId="1" pageNumber="421">
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<taxonomicName id="4C5CF9AAFFD9FF036CF04606FD7FF830" authority="A. Milne-Edwards, 1873: 311" authorityName="A. Milne-Edwards" authorityPageNumber="311" authorityYear="1873" box="[151,706,792,814]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="421" phylum="Arthropoda" rank="species" species="balansae">
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<emphasis id="B9285E3BFFD9FF036CF04606FE36F830" box="[151,395,792,814]" italics="true" pageId="1" pageNumber="421">Clistocoeloma balansae</emphasis>
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A. Milne-Edwards, 1873: 311
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</taxonomicName>
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, 312, 331, pl. 17: fig. 1, 1a–c (
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<typeStatus id="54E73C8BFFD9FF036F9E4607FB9BF830" box="[1017,1062,793,814]" pageId="1" pageNumber="421">type</typeStatus>
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locality Nouvelle Calédonie [=
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<collectingCountry id="F34BC2B9FFD9FF03690A4607FE88F852" name="New Caledonia" pageId="1" pageNumber="421">New Caledonia</collectingCountry>
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]).
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</paragraph>
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</subSubSection>
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<subSubSection id="C346D1A2FFD9FF036CF0464BFC62FF7E" pageId="1" pageNumber="421" type="description">
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<paragraph id="8BE38229FFD9FF036CF0464BFE52F8D9" blockId="1.[151,1437,792,1121]" pageId="1" pageNumber="421">
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<taxonomicName id="4C5CF9AAFFD9FF036CF0464BFE36F875" box="[151,395,853,875]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="421" phylum="Arthropoda" rank="species" species="balansae">
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<emphasis id="B9285E3BFFD9FF036CF0464BFE36F875" box="[151,395,853,875]" italics="true" pageId="1" pageNumber="421">Clistocoeloma balansae</emphasis>
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</taxonomicName>
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—Kingsley 1880: 219;
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<collectingCountry id="F34BC2B9FFD9FF036E1D4648FD25F875" box="[634,664,854,875]" name="South Georgia and the South Sandwich Islands" pageId="1" pageNumber="421">De</collectingCountry>
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<collectingCountry id="F34BC2B9FFD9FF036EF94648FD71F875" box="[670,716,854,875]" name="Isle of Man" pageId="1" pageNumber="421">Man</collectingCountry>
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1888: 195–196;
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<collectingCountry id="F34BC2B9FFD9FF036F1D4648FC2AF875" box="[890,919,854,875]" name="South Georgia and the South Sandwich Islands" pageId="1" pageNumber="421">De</collectingCountry>
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<collectingCountry id="F34BC2B9FFD9FF036FF94648FC71F875" box="[926,972,854,875]" name="Isle of Man" pageId="1" pageNumber="421">Man</collectingCountry>
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1896: 339, 340, 342, 343; Alcock 1900: 429; Tesch 1917: 222–225, 231 (list), 260 (key); Shen 1933: 52, 54 (key); Chopra & Das 1937: 433, fig. 15d (list); Edmondson 1951: 240 (list); Hsueh & Huang 1996: 63 (list); Rahayu & Takeda 2000: 38 (list); Davie 2002: 221; Komai
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<emphasis id="B9285E3BFFD9FF036940468AFAE8F8B7" box="[1319,1365,915,937]" italics="true" pageId="1" pageNumber="421">et al</emphasis>
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. 2004: 41; Ng
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<emphasis id="B9285E3BFFD9FF036D7446ADFEFBF8D9" box="[275,326,945,968]" italics="true" pageId="1" pageNumber="421">et al.</emphasis>
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2008: 220 (list).
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</paragraph>
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<paragraph id="8BE38229FFD9FF036CF046CFFCB2F8F8" blockId="1.[151,1437,792,1121]" box="[151,783,976,998]" pageId="1" pageNumber="421">
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<taxonomicName id="4C5CF9AAFFD9FF036CF046CFFDD6F8F8" authority="Rathbun, 1914: 78" authorityName="Rathbun" authorityPageNumber="78" authorityYear="1914" box="[151,619,976,998]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="421" phylum="Arthropoda" rank="species" species="tectum" subGenus="Sesarma">
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<emphasis id="B9285E3BFFD9FF036CF046CFFF53F8F8" box="[151,238,977,998]" italics="true" pageId="1" pageNumber="421">Sesarma</emphasis>
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(
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<emphasis id="B9285E3BFFD9FF036C9A46CFFEE9F8F8" box="[253,340,977,998]" italics="true" pageId="1" pageNumber="421">Sesarma</emphasis>
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)
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<emphasis id="B9285E3BFFD9FF036D0446CFFE1AF8F8" box="[355,423,977,998]" italics="true" pageId="1" pageNumber="421">tectum</emphasis>
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Rathbun, 1914: 78
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</taxonomicName>
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, 79 [synonymy]
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</paragraph>
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<paragraph id="8BE38229FFD9FF036CF046F1FC43FF1A" blockId="1.[151,1437,792,1121]" box="[151,1022,1006,1029]" pageId="1" pageNumber="421">
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<taxonomicName id="4C5CF9AAFFD9FF036CF046F1FE1AFF1B" box="[151,423,1007,1029]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="421" phylum="Arthropoda" rank="species" species="tectum" subGenus="Sesarma">
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<emphasis id="B9285E3BFFD9FF036CF046F1FF53FF1A" box="[151,238,1007,1028]" italics="true" pageId="1" pageNumber="421">Sesarma</emphasis>
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(
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<emphasis id="B9285E3BFFD9FF036C9A46F1FEE9FF1A" box="[253,340,1007,1028]" italics="true" pageId="1" pageNumber="421">Sesarma</emphasis>
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)
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<emphasis id="B9285E3BFFD9FF036D0446EEFE1AFF1B" box="[355,423,1008,1029]" italics="true" pageId="1" pageNumber="421">tectum</emphasis>
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</taxonomicName>
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—Estampador 1937: 537 (list); Estampador 1959: 94 (list).
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</paragraph>
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<paragraph id="8BE38229FFD9FF036CF04113FC62FF7E" blockId="1.[151,1437,792,1121]" pageId="1" pageNumber="421">
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<taxonomicName id="4C5CF9AAFFD9FF036CF04113FECEFF3D" box="[151,371,1037,1059]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="421" phylum="Arthropoda" rank="species" species="tectum">
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<emphasis id="B9285E3BFFD9FF036CF04113FECEFF3D" box="[151,371,1037,1059]" italics="true" pageId="1" pageNumber="421">Clistocoeloma tectum</emphasis>
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</taxonomicName>
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—Tesch 1917: 222–227, 231 (list), 260 (key), pls. 17: fig. 3, 3a–c; Shen, 1933: 53, 54 (key); Chopra & Das 1937: 433; Ward 1941: 2 (list); Edmondson 1951: 240; Hsueh & Huang 1996: 63 (list); Niem 1996: 332 (list); Rahayu & Takeda 2000: 35, 38; Komai
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<emphasis id="B9285E3BFFD9FF036E6F4152FD8EFF7F" box="[520,563,1099,1121]" italics="true" pageId="1" pageNumber="421">et al</emphasis>
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. 2004: 41 (list); Ng
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<emphasis id="B9285E3BFFD9FF036F644152FC92FF7F" box="[771,815,1099,1121]" italics="true" pageId="1" pageNumber="421">et al</emphasis>
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. 2008: 220 (list).
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</paragraph>
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</subSubSection>
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<subSubSection id="C346D1A2FFD9FF036CF04193FABFFDFB" pageId="1" pageNumber="421" type="materials_examined">
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<paragraph id="8BE38229FFD9FF036CF04193FABFFDFB" blockId="1.[151,1436,1165,2017]" pageId="1" pageNumber="421">
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<emphasis id="B9285E3BFFD9FF036CF04193FE39FFB8" bold="true" box="[151,388,1165,1190]" pageId="1" pageNumber="421">Material examined:</emphasis>
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<specimenCount id="9D5A49A0FFD9FF036DEB4193FE64FFBB" box="[396,473,1165,1189]" pageId="1" pageNumber="421" type="male">1 male</specimenCount>
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(20.7 ×
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<quantity id="4CA42FCCFFD9FF036E5A4193FD19FFBB" box="[573,676,1165,1189]" metricMagnitude="-2" metricUnit="m" metricValue="1.97" pageId="1" pageNumber="421" unit="mm" value="19.7">19.7 mm</quantity>
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) (MNHN B10912) [
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<typeStatus id="54E73C8BFFD9FF036FEA4193FC44FFBB" box="[909,1017,1165,1189]" pageId="1" pageNumber="421" type="lectotype">lectotype</typeStatus>
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, herein designated],
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<collectingCountry id="F34BC2B9FFD9FF0368824190FA25FFBB" box="[1253,1432,1165,1189]" name="New Caledonia" pageId="1" pageNumber="421">New Caledonia</collectingCountry>
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, coll. B. Balansa, no collection date.—
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<specimenCount id="9D5A49A0FFD9FF036E0E41AFFD71FFD7" box="[617,716,1201,1225]" pageId="1" pageNumber="421" type="female">1 female</specimenCount>
|
||
(18.9 ×
|
||
<quantity id="4CA42FCCFFD9FF036F2641AFFC10FFD7" box="[833,941,1201,1225]" metricMagnitude="-2" metricUnit="m" metricValue="1.73" pageId="1" pageNumber="421" unit="mm" value="17.3">17.3 mm</quantity>
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) (MNHN B10912) [
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||
<typeStatus id="54E73C8BFFD9FF0368C041AFFAF9FFD7" box="[1191,1348,1201,1225]" pageId="1" pageNumber="421" type="paralectotype">paralectotype</typeStatus>
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, herein designated],
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<collectingCountry id="F34BC2B9FFD9FF036D4B41C8FE5CFFF3" box="[300,481,1237,1261]" name="New Caledonia" pageId="1" pageNumber="421">New Caledonia</collectingCountry>
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||
, coll. Balansa, no collection date.—
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<specimenCount id="9D5A49A0FFD9FF036FF141CBFC4AFFF3" box="[918,1015,1237,1261]" pageId="1" pageNumber="421" type="female">1 female</specimenCount>
|
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(19.5 ×
|
||
<quantity id="4CA42FCCFFD9FF03680041CBFB6FFFF3" box="[1127,1234,1237,1261]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" pageId="1" pageNumber="421" unit="mm" value="18.6">18.6 mm</quantity>
|
||
) (USNM 45766) [
|
||
<typeStatus id="54E73C8BFFD9FF036CFA41E7FEBFFE0F" box="[157,258,1273,1297]" pageId="1" pageNumber="421" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C5CF9AAFFD9FF036D5F41E4FC8BFE0F" authority="Rathbun, 1914" authorityName="Rathbun" authorityYear="1914" box="[312,822,1273,1297]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="421" phylum="Arthropoda" rank="species" species="tectum" subGenus="Sesarma">
|
||
<emphasis id="B9285E3BFFD9FF036D5F41E4FE25FE0F" box="[312,408,1274,1297]" italics="true" pageId="1" pageNumber="421">Sesarma</emphasis>
|
||
(
|
||
<emphasis id="B9285E3BFFD9FF036DD641E4FDACFE0F" box="[433,529,1274,1297]" italics="true" pageId="1" pageNumber="421">Sesarma</emphasis>
|
||
)
|
||
<emphasis id="B9285E3BFFD9FF036E4D41E4FDC8FE0F" box="[554,629,1274,1297]" italics="true" pageId="1" pageNumber="421">tectum</emphasis>
|
||
Rathbun, 1914
|
||
</taxonomicName>
|
||
], off northern Luzon, Palaui I., Port
|
||
<collectingCountry id="F34BC2B9FFD9FF03696141E7FA25FE0F" box="[1286,1432,1273,1297]" name="Saint Vincent and the Grenadines" pageId="1" pageNumber="421">San Vicente</collectingCountry>
|
||
,
|
||
<collectingCountry id="F34BC2B9FFD9FF036CF04003FEA5FE2B" box="[151,280,1309,1333]" name="Philippines" pageId="1" pageNumber="421">Philippines</collectingCountry>
|
||
, coll.
|
||
<date id="FFE2A4E9FFD9FF036D3B4003FD89FE2B" box="[348,564,1309,1333]" pageId="1" pageNumber="421" value="1908-11-14">14 November 1908</date>
|
||
.—Additional material:
|
||
<specimenCount id="9D5A49A0FFD9FF036F234003FC32FE2B" box="[836,911,1309,1333]" pageId="1" pageNumber="421" type="male">1 male</specimenCount>
|
||
(11.8 ×
|
||
<quantity id="4CA42FCCFFD9FF036F974003FBE5FE2B" box="[1008,1112,1309,1333]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="1" pageNumber="421" unit="mm" value="11.2">11.2 mm</quantity>
|
||
),
|
||
<specimenCount id="9D5A49A0FFD9FF03680C4003FB70FE2B" box="[1131,1229,1309,1333]" pageId="1" pageNumber="421" type="female">1 female</specimenCount>
|
||
(16.5 ×
|
||
<quantity id="4CA42FCCFFD9FF0369494003FA29FE2B" box="[1326,1428,1309,1333]" metricMagnitude="-2" metricUnit="m" metricValue="1.53" pageId="1" pageNumber="421" unit="mm" value="15.3">15.3 mm</quantity>
|
||
) (ZRC 2009.0898), Kawasan, Matutinao, Cebu,
|
||
<collectingCountry id="F34BC2B9FFD9FF036ECF405FFC94FE47" box="[680,809,1345,1369]" name="Philippines" pageId="1" pageNumber="421">Philippines</collectingCountry>
|
||
, coll. H. C. Liu
|
||
<emphasis id="B9285E3BFFD9FF036FBD405CFBAFFE47" box="[986,1042,1345,1369]" italics="true" pageId="1" pageNumber="421">et al.</emphasis>
|
||
,
|
||
<date id="FFE2A4E9FFD9FF036878405FFB66FE47" box="[1055,1243,1345,1370]" pageId="1" pageNumber="421" value="2001-01-12">12 January 2001</date>
|
||
.—
|
||
<specimenCount id="9D5A49A0FFD9FF036898405FFAF9FE47" box="[1279,1348,1345,1369]" pageId="1" pageNumber="421" type="male">1 male</specimenCount>
|
||
(23.3 ×
|
||
<quantity id="4CA42FCCFFD9FF036CF0407BFF43FE63" box="[151,254,1381,1405]" metricMagnitude="-2" metricUnit="m" metricValue="2.26" pageId="1" pageNumber="421" unit="mm" value="22.6">22.6 mm</quantity>
|
||
) (ZRC 2009.0900), Gilatilan, south of Matutinao, Cebu,
|
||
<collectingCountry id="F34BC2B9FFD9FF036FEC407BFBB1FE63" box="[907,1036,1381,1405]" name="Philippines" pageId="1" pageNumber="421">Philippines</collectingCountry>
|
||
, coll. H. C. Liu
|
||
<emphasis id="B9285E3BFFD9FF0368AE4079FABEFE60" box="[1225,1283,1382,1406]" italics="true" pageId="1" pageNumber="421">et al.</emphasis>
|
||
,
|
||
<date id="FFE2A4E9FFD9FF036975407BFF6EFEBF" pageId="1" pageNumber="421" value="2001-02-12">12 February 2001</date>
|
||
.—
|
||
<specimenCount id="9D5A49A0FFD9FF036C914097FE80FEBF" box="[246,317,1417,1441]" pageId="1" pageNumber="421" type="male">1 male</specimenCount>
|
||
(15.9 ×
|
||
<quantity id="4CA42FCCFFD9FF036DFC4097FDBEFEBF" box="[411,515,1417,1441]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="1" pageNumber="421" unit="mm" value="15.5">15.5 mm</quantity>
|
||
),
|
||
<specimenCount id="9D5A49A0FFD9FF036E714097FD3FFEBF" box="[534,642,1417,1441]" pageId="1" pageNumber="421" type="female">2 females</specimenCount>
|
||
(18.7 ×
|
||
<quantity id="4CA42FCCFFD9FF036E874097FCFBFEBF" box="[736,838,1417,1441]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="1" pageNumber="421" unit="mm" value="17.8">17.8 mm</quantity>
|
||
, 17.2 ×
|
||
<quantity id="4CA42FCCFFD9FF036FC74097FBB9FEBF" box="[928,1028,1417,1441]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" pageId="1" pageNumber="421" unit="mm" value="16.9">16.9 mm</quantity>
|
||
) (ZRC 2012.702), PANGLAO 2004 Expedition, station M45, adjacent to Nipa palms, freshwater spring on bank, Abatan River, Bohol I.,
|
||
<collectingCountry id="F34BC2B9FFD9FF03697940B3FA21FEDB" box="[1310,1436,1453,1477]" name="Philippines" pageId="1" pageNumber="421">Philippines</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE68E4EEFFD9FF036CF840CFFEBAFEF7" box="[159,263,1489,1513]" direction="north" orientation="latitude" pageId="1" pageNumber="421" precision="92" value="9.731667">9º43.9’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE68E4EEFFD9FF036D7C40CFFE20FEF7" box="[283,413,1489,1513]" direction="east" orientation="longitude" pageId="1" pageNumber="421" precision="92" value="123.89">123º53.4’E</geoCoordinate>
|
||
), coll. PANGLAO 2004 Expedition,
|
||
<date id="FFE2A4E9FFD9FF036F0540CFFBB1FEF7" box="[866,1036,1489,1513]" pageId="1" pageNumber="421" value="2004-06-29">29 June 2004</date>
|
||
.—
|
||
<specimenCount id="9D5A49A0FFD9FF03684840CFFBC0FEF7" box="[1071,1149,1489,1513]" pageId="1" pageNumber="421" type="male">1 male</specimenCount>
|
||
(7.4 ×
|
||
<quantity id="4CA42FCCFFD9FF03688140CFFAF8FEF7" box="[1254,1349,1489,1513]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="1" pageNumber="421" unit="mm" value="7.1">7.1 mm</quantity>
|
||
) (ZRC 2012.703), PANGLAO 2004 Expedition, station M57, fringe mangrove, Sungcolan inlet, Panglao I.,
|
||
<collectingCountry id="F34BC2B9FFD9FF03697940EBFA21FD13" box="[1310,1436,1525,1549]" name="Philippines" pageId="1" pageNumber="421">Philippines</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE68E4EEFFD9FF036CF84307FEBAFD2F" box="[159,263,1561,1585]" direction="north" orientation="latitude" pageId="1" pageNumber="421" precision="92" value="9.633333">9º38.0’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE68E4EEFFD9FF036D724307FE2AFD2F" box="[277,407,1561,1585]" direction="east" orientation="longitude" pageId="1" pageNumber="421" precision="92" value="123.831665">123º49.9’E</geoCoordinate>
|
||
), coll. PANGLAO 2004 Expedition,
|
||
<date id="FFE2A4E9FFD9FF036F594307FC7AFD2F" box="[830,967,1561,1585]" pageId="1" pageNumber="421" value="2004-07-04">4 July 2004</date>
|
||
.—
|
||
<specimenCount id="9D5A49A0FFD9FF036F8D4307FB82FD2F" box="[1002,1087,1561,1585]" pageId="1" pageNumber="421" type="male">3 males</specimenCount>
|
||
(15.5 ×
|
||
<quantity id="4CA42FCCFFD9FF0368C34307FAB0FD2F" box="[1188,1293,1561,1585]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="1" pageNumber="421" unit="mm" value="14.7">14.7 mm</quantity>
|
||
, 12.3 ×
|
||
<quantity id="4CA42FCCFFD9FF03690B4307FF7BFD4B" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="1" pageNumber="421" unit="mm" value="11.7">11.7 mm</quantity>
|
||
, 9.0 ×
|
||
<quantity id="4CA42FCCFFD9FF036D7D4323FECAFD4B" box="[282,375,1597,1621]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="421" unit="mm" value="8.5">8.5 mm</quantity>
|
||
),
|
||
<specimenCount id="9D5A49A0FFD9FF036DEB4323FE46FD4B" box="[396,507,1597,1621]" pageId="1" pageNumber="421" type="female">2 females</specimenCount>
|
||
(21.7 ×
|
||
<quantity id="4CA42FCCFFD9FF036E044323FD70FD4B" box="[611,717,1597,1621]" metricMagnitude="-2" metricUnit="m" metricValue="2.12" pageId="1" pageNumber="421" unit="mm" value="21.2">21.2 mm</quantity>
|
||
, 14.0 ×
|
||
<quantity id="4CA42FCCFFD9FF036F484323FC25FD4B" box="[815,920,1597,1621]" metricMagnitude="-2" metricUnit="m" metricValue="2.12" pageId="1" pageNumber="421" unit="mm" value="21.2">21.2 mm</quantity>
|
||
) (ZRC 2012.704), Kawasan Fall, Matutinau, Cebu,
|
||
<collectingCountry id="F34BC2B9FFD9FF036C84437FFED9FD67" box="[227,356,1633,1657]" name="Philippines" pageId="1" pageNumber="421">Philippines</collectingCountry>
|
||
, coll. P. K. L. Ng
|
||
<emphasis id="B9285E3BFFD9FF036E5B437CFDC3FD67" box="[572,638,1633,1657]" italics="true" pageId="1" pageNumber="421">et al.,</emphasis>
|
||
<date id="FFE2A4E9FFD9FF036EE0437FFC9FFD67" box="[647,802,1633,1657]" pageId="1" pageNumber="421" value="2003-07-30">30 July 2003</date>
|
||
.—
|
||
<specimenCount id="9D5A49A0FFD9FF036F22437FFC19FD67" box="[837,932,1633,1657]" pageId="1" pageNumber="421" type="female">1 female</specimenCount>
|
||
(17.5 ×
|
||
<quantity id="4CA42FCCFFD9FF03686B437FFBCEFD67" box="[1036,1139,1633,1657]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="1" pageNumber="421" unit="mm" value="16.5">16.5 mm</quantity>
|
||
) (ZRC 2012.700), station VM4, intertidal, between Rose point and Nasouli River,
|
||
<collectingCountry id="F34BC2B9FFD9FF036F534398FC2DFD83" box="[820,912,1670,1693]" name="Vanuatu" pageId="1" pageNumber="421">Vanuatu</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE68E4EEFFD9FF036FC4439BFBA8FD83" box="[931,1045,1669,1693]" direction="south" orientation="latitude" pageId="1" pageNumber="421" precision="92" value="-15.581667">15º34.9’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE68E4EEFFD9FF036842439BFB1AFD83" box="[1061,1191,1669,1693]" direction="east" orientation="longitude" pageId="1" pageNumber="421" precision="92" value="167.03">167º01.8’E</geoCoordinate>
|
||
), coll. SANTO 2006 Expedition,
|
||
<date id="FFE2A4E9FFD9FF036D4643B7FDBCFDDF" box="[289,513,1705,1729]" pageId="1" pageNumber="421" value="2006-09-11">11 September 2006</date>
|
||
.—
|
||
<specimenCount id="9D5A49A0FFD9FF036E4343B7FDD6FDDF" box="[548,619,1705,1729]" pageId="1" pageNumber="421" type="male">1 male</specimenCount>
|
||
(11.2 ×
|
||
<quantity id="4CA42FCCFFD9FF036EAA43B7FC8EFDDF" box="[717,819,1705,1729]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="421" unit="mm" value="10.5">10.5 mm</quantity>
|
||
) (ZRC 2012.701), station VM15, estuarine mangrove, Wambu River,
|
||
<collectingCountry id="F34BC2B9FFD9FF036D5843D0FE27FDFB" box="[319,410,1742,1765]" name="Vanuatu" pageId="1" pageNumber="421">Vanuatu</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE68E4EEFFD9FF036DCC43D3FDA1FDFB" box="[427,540,1741,1765]" direction="south" orientation="latitude" pageId="1" pageNumber="421" precision="92" value="-15.558333">15º33.5’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE68E4EEFFD9FF036E4F43D3FD17FDFB" box="[552,682,1741,1765]" direction="east" orientation="longitude" pageId="1" pageNumber="421" precision="92" value="167.14">167º08.4’E</geoCoordinate>
|
||
), coll. SANTO 2006 Expedition,
|
||
<date id="FFE2A4E9FFD9FF03684643D3FB43FDFB" box="[1057,1278,1741,1765]" pageId="1" pageNumber="421" value="2006-09-17">17 September 2006</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346D1A2FFD9FF046CA043EFFBA9FD74" lastPageId="6" lastPageNumber="426" pageId="1" pageNumber="421" type="diagnosis">
|
||
<paragraph id="8BE38229FFD9FF036CA043EFFA21FCFF" blockId="1.[151,1436,1165,2017]" pageId="1" pageNumber="421">
|
||
<emphasis id="B9285E3BFFD9FF036CA043EFFEFCFC14" bold="true" box="[199,321,1777,1802]" pageId="1" pageNumber="421">Diagnosis.</emphasis>
|
||
Carapace almost square, covered with dense coat of setae, among which stands small tufts of setae resembling tubercles; denuded carapace with uneven surface, with pronounced tubercles (
|
||
<figureCitation id="13679EACFFD9FF0368DE420BFAB5FC33" box="[1209,1288,1813,1837]" captionStart="FIGURE 1" captionStartId="2.[151,250,1910,1933]" captionTargetBox="[188,1398,301,1888]" captionTargetId="figure@2.[188,1398,301,1889]" captionTargetPageId="2" captionText="FIGURE 1. Clistocoeloma balansae A. Milne-Edwards, 1873, lectotype male (20.7 × 19.7 mm) (MNHN B 10912), New Caledonia. A, dorsal overall view; B, ventral view of abdomen and thoracic sternum; C, frontal view of carapace. (Photographs by T. Naruse)" httpUri="https://zenodo.org/record/222414/files/figure.png" pageId="1" pageNumber="421">Figs. 1</figureCitation>
|
||
A, 2A, 3A). Postfrontal lobes distinct, divided by deep longitudinal grooves, distal lobes separated into 2 distinct tubercles by shallow groove (
|
||
<figureCitation id="13679EACFFD9FF036D324243FE1CFC6B" box="[341,417,1885,1909]" captionStart="FIGURE 1" captionStartId="2.[151,250,1910,1933]" captionTargetBox="[188,1398,301,1888]" captionTargetId="figure@2.[188,1398,301,1889]" captionTargetPageId="2" captionText="FIGURE 1. Clistocoeloma balansae A. Milne-Edwards, 1873, lectotype male (20.7 × 19.7 mm) (MNHN B 10912), New Caledonia. A, dorsal overall view; B, ventral view of abdomen and thoracic sternum; C, frontal view of carapace. (Photographs by T. Naruse)" httpUri="https://zenodo.org/record/222414/files/figure.png" pageId="1" pageNumber="421">Figs. 1</figureCitation>
|
||
A, 2A, 3A). Inner orbital angle present, pronounced (
|
||
<figureCitation id="13679EACFFD9FF0368614243FBEFFC6B" box="[1030,1106,1885,1909]" captionStart="FIGURE 1" captionStartId="2.[151,250,1910,1933]" captionTargetBox="[188,1398,301,1888]" captionTargetId="figure@2.[188,1398,301,1889]" captionTargetPageId="2" captionText="FIGURE 1. Clistocoeloma balansae A. Milne-Edwards, 1873, lectotype male (20.7 × 19.7 mm) (MNHN B 10912), New Caledonia. A, dorsal overall view; B, ventral view of abdomen and thoracic sternum; C, frontal view of carapace. (Photographs by T. Naruse)" httpUri="https://zenodo.org/record/222414/files/figure.png" pageId="1" pageNumber="421">Figs. 1</figureCitation>
|
||
C, 2C). Anterolateral margin with 3 triangular teeth, inclusive of external orbital angle. Cheliped with dorsal surface of carpus with dense coat of setae; covered with tubercles when denuded; single longitudinal pectinated ridge present on dorsal surface of chela, with about 30 fine closely spaced teeth; dorsal surface of movable dactylus of chela with 14–20 tubercles (
|
||
<figureCitation id="13679EACFFD9FF03692E42D7FA33FCFF" box="[1353,1422,1993,2017]" captionStart="FIGURE 4" captionStartId="5.[151,250,1982,2005]" captionTargetBox="[302,1285,373,1961]" captionTargetId="figure@5.[302,1285,373,1961]" captionTargetPageId="5" captionText="FIGURE 4. Clistocoeloma balansae A. Milne-Edwards, 1873, male (15.9 × 15.5 mm) (ZRC 2012.702), Philippines. A, outer view of left chela; B, inner view of left chela; C, dorsal marginal view of left chela." httpUri="https://zenodo.org/record/222417/files/figure.png" pageId="1" pageNumber="421">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE38229FFDAFF006CF04586FADCFBE9" blockId="2.[151,1436,151,247]" pageId="2" pageNumber="422">
|
||
Inner surface of chela granular, with single vertical row of approximately 9 granules in adult males (
|
||
<figureCitation id="13679EACFFDAFF0069544589FAC6FBB1" box="[1331,1403,151,175]" captionStart="FIGURE 4" captionStartId="5.[151,250,1982,2005]" captionTargetBox="[302,1285,373,1961]" captionTargetId="figure@5.[302,1285,373,1961]" captionTargetPageId="5" captionText="FIGURE 4. Clistocoeloma balansae A. Milne-Edwards, 1873, male (15.9 × 15.5 mm) (ZRC 2012.702), Philippines. A, outer view of left chela; B, inner view of left chela; C, dorsal marginal view of left chela." httpUri="https://zenodo.org/record/222417/files/figure.png" pageId="2" pageNumber="422">Fig. 4</figureCitation>
|
||
B). Tubercle present behind pectinated ridge on inner surface of male chela (
|
||
<figureCitation id="13679EACFFDAFF006FAC45A2FBA7FBCA" box="[971,1050,188,212]" captionStart="FIGURE 4" captionStartId="5.[151,250,1982,2005]" captionTargetBox="[302,1285,373,1961]" captionTargetId="figure@5.[302,1285,373,1961]" captionTargetPageId="5" captionText="FIGURE 4. Clistocoeloma balansae A. Milne-Edwards, 1873, male (15.9 × 15.5 mm) (ZRC 2012.702), Philippines. A, outer view of left chela; B, inner view of left chela; C, dorsal marginal view of left chela." httpUri="https://zenodo.org/record/222417/files/figure.png" pageId="2" pageNumber="422">Figs. 4</figureCitation>
|
||
A, 4C). Tip of G1 with dense tufts of setae, single row of setae on exterior margin of G1; wide chitinous tip visible when denuded (
|
||
<figureCitation id="13679EACFFDAFF0068A945C1FAA7FBE9" box="[1230,1306,223,247]" captionStart="FIGURE 5" captionStartId="6.[151,250,1124,1147]" captionTargetBox="[172,1368,241,1053]" captionTargetId="figure@6.[151,1436,176,1121]" captionTargetPageId="6" captionText="FIGURE 5. Clistocoeloma balansae A. Milne-Edwards, 1873, male (15.9 × 15.5 mm) (ZRC 2012.702), Philippines. A, male abdomen; B, G 1 dorsal view; C, G 1 lateral view; D, G 1 ventral view. G 1 denuded." httpUri="https://zenodo.org/record/222418/files/figure.png" pageId="2" pageNumber="422">Figs. 5</figureCitation>
|
||
B–D).
|
||
</paragraph>
|
||
<caption id="DF23D2A1FFDAFF006CF04268FE9CFCD4" httpUri="https://zenodo.org/record/222414/files/figure.png" pageId="2" pageNumber="422" targetBox="[188,1398,301,1888]" targetPageId="2">
|
||
<paragraph id="8BE38229FFDAFF006CF04268FE9CFCD4" blockId="2.[151,1436,1910,1994]" pageId="2" pageNumber="422">
|
||
<emphasis id="B9285E3BFFDAFF006CF04268FEACFC95" bold="true" box="[151,273,1910,1933]" pageId="2" pageNumber="422">FIGURE 1</emphasis>
|
||
.
|
||
<taxonomicName id="4C5CF9AAFFDAFF006D464268FC94FC92" authority="A. Milne-Edwards, 1873" authorityName="A. Milne-Edwards" authorityYear="1873" box="[289,809,1910,1932]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="422" phylum="Arthropoda" rank="species" species="balansae">
|
||
<emphasis id="B9285E3BFFDAFF006D464268FDA4FC92" box="[289,537,1910,1932]" italics="true" pageId="2" pageNumber="422">Clistocoeloma balansae</emphasis>
|
||
A. Milne-Edwards, 1873
|
||
</taxonomicName>
|
||
, lectotype male (20.7 × 19.7 mm) (MNHN B10912), New Caledonia. A, dorsal overall view; B, ventral view of abdomen and thoracic sternum; C, frontal view of carapace. (Photographs by T. Naruse)
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF23D2A1FFDBFF016CF043A0FF4AFC0F" httpUri="https://zenodo.org/record/222415/files/figure.png" pageId="3" pageNumber="423" targetBox="[188,1398,193,1703]" targetPageId="3">
|
||
<paragraph id="8BE38229FFDBFF016CF043A0FF4AFC0F" blockId="3.[151,1436,1726,1809]" pageId="3" pageNumber="423">
|
||
<emphasis id="B9285E3BFFDBFF016CF043A0FEB2FDCD" bold="true" box="[151,271,1726,1749]" pageId="3" pageNumber="423">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="4C5CF9AAFFDBFF016D7943A1FEC8FDCA" box="[286,373,1727,1748]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="423" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9285E3BFFDBFF016D7943A1FEC8FDCA" box="[286,373,1727,1748]" italics="true" pageId="3" pageNumber="423">Sesarma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B9285E3BFFDBFF016DE143A1FE6CFDCA" box="[390,465,1727,1748]" italics="true" pageId="3" pageNumber="423">Sesarm</emphasis>
|
||
a)
|
||
<taxonomicName id="4C5CF9AAFFDBFF016D8A43A1FD6EFDCA" authority="Rathbun, 1914" authorityName="Rathbun" authorityYear="1914" box="[493,723,1726,1748]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="423" phylum="Arthropoda" rank="species" species="tectum">
|
||
<emphasis id="B9285E3BFFDBFF016D8A43A1FD8CFDCA" box="[493,561,1727,1748]" italics="true" pageId="3" pageNumber="423">tectum</emphasis>
|
||
Rathbun, 1914
|
||
</taxonomicName>
|
||
, holotype female (19.5 × 18.6 mm) (USNM 45766), Philippines. A, dorsal overall view; B, ventral view of abdomen and thoracic sternum; C, frontal view of carapace; D, posterior view of carapace.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE38229FFDBFF016CA04223FC69FCF8" blockId="3.[151,1436,1853,2022]" pageId="3" pageNumber="423">
|
||
<emphasis id="B9285E3BFFDBFF016CA04223FEC9FC48" bold="true" box="[199,372,1853,1878]" pageId="3" pageNumber="423">Redescription.</emphasis>
|
||
Carapace slightly wider than long, almost square, covered with dense coat of setae, among which stands small tufts of setae resembling tubercles. Denuded carapace with uneven surface; pronounced tubercles on carapace defined by deep grooves. Postfrontal lobes distinct, divided by deep longitudinal grooves, median pair broad, prominent, distal lobes split into 2 distinct tubercles by shallow groove. Regions relatively well defined by tubercles on carapace. Inner orbital angle present, pronounced.
|
||
</paragraph>
|
||
<caption id="DF23D2A1FFDCFF066CF043B6FD82FDE5" httpUri="https://zenodo.org/record/222416/files/figure.png" pageId="4" pageNumber="424" targetBox="[188,1398,176,1685]" targetPageId="4">
|
||
<paragraph id="8BE38229FFDCFF066CF043B6FD82FDE5" blockId="4.[151,1436,1704,1787]" pageId="4" pageNumber="424">
|
||
<emphasis id="B9285E3BFFDCFF066CF043B6FEB0FDA3" bold="true" box="[151,269,1704,1727]" pageId="4" pageNumber="424">FIGURE 3</emphasis>
|
||
.
|
||
<taxonomicName id="4C5CF9AAFFDCFF066D7D43B6FCA8FDA0" authority="A. Milne-Edwards, 1873" authorityName="A. Milne-Edwards" authorityYear="1873" box="[282,789,1704,1726]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="424" phylum="Arthropoda" rank="species" species="balansae">
|
||
<emphasis id="B9285E3BFFDCFF066D7D43B6FDB2FDA0" box="[282,527,1704,1726]" italics="true" pageId="4" pageNumber="424">Clistocoeloma balansae</emphasis>
|
||
A. Milne-Edwards, 1873
|
||
</taxonomicName>
|
||
. A–C, male (15.9 × 15.5 mm) (ZRC 2012.702), Philippines; D, female (17.2 × 16.9 mm) (ZRC 2012.702), Philippines. A, dorsal overall view; B, ventral view of abdomen and thoracic sternum; C, D posterior view of carapace.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE38229FFDCFF066CA04236FAD4FC7A" blockId="4.[151,1436,1832,2036]" pageId="4" pageNumber="424">Anterolateral margin with 3 triangular teeth, inclusive of external orbital angle: third tooth largest, roundest; posterolateral margin convex. Entire margin of carapace fringed with dense, short setae, which obscures teeth.</paragraph>
|
||
<paragraph id="8BE38229FFDCFF076CA0426EFC03FA21" blockId="4.[151,1436,1832,2036]" lastBlockId="5.[151,1436,151,319]" lastPageId="5" lastPageNumber="425" pageId="4" pageNumber="424">Chelipeds equal or subequal in size; large, robust in males; small, slender in females; chela approximately 2.1 times dactylus length. Merus covered with dense coat of setae, smooth when denuded. Dorsal surface of carpus covered with small tufts of setae resembling tubercles; revealing tubercles when denuded. Outer surface of palm smooth when denuded. Dorsal surface of movable dactylus of chela with 14–20 tubercles, evenly spaced, proximal tubercles small, increasing in size medially, decreasing in size distally. Tubercles on movable dactylus of chela present in female but smaller, less distinct. Single longitudinal pectinated ridge on dorsal surface of chela, with about 30 fine teeth. Inner surface of chela granular, with single vertical row of approximately 9 granules in adult males. Granules on inner surface of palm not obvious or absent in juveniles, females. Tubercle present behind pectinated ridge on inner surface of male chela; less distinct on females.</paragraph>
|
||
<caption id="DF23D2A1FFDDFF076CF042A0FC59FCED" httpUri="https://zenodo.org/record/222417/files/figure.png" pageId="5" pageNumber="425" targetBox="[302,1285,373,1961]" targetPageId="5">
|
||
<paragraph id="8BE38229FFDDFF076CF042A0FC59FCED" blockId="5.[151,1436,1982,2035]" pageId="5" pageNumber="425">
|
||
<emphasis id="B9285E3BFFDDFF076CF042A0FEB1FCCD" bold="true" box="[151,268,1982,2005]" pageId="5" pageNumber="425">FIGURE 4</emphasis>
|
||
.
|
||
<taxonomicName id="4C5CF9AAFFDDFF076D7D42A0FCA8FCCA" authority="A. Milne-Edwards, 1873" authorityName="A. Milne-Edwards" authorityYear="1873" box="[282,789,1982,2004]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="425" phylum="Arthropoda" rank="species" species="balansae">
|
||
<emphasis id="B9285E3BFFDDFF076D7D42A0FDB2FCCA" box="[282,527,1982,2004]" italics="true" pageId="5" pageNumber="425">Clistocoeloma balansae</emphasis>
|
||
A. Milne-Edwards, 1873
|
||
</taxonomicName>
|
||
, male (15.9 × 15.5 mm) (ZRC 2012.702), Philippines. A, outer view of left chela; B, inner view of left chela; C, dorsal marginal view of left chela.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF23D2A1FFDEFF046CF0417AFC60FF87" httpUri="https://zenodo.org/record/222418/files/figure.png" pageId="6" pageNumber="426" targetBox="[172,1368,241,1053]" targetPageId="6">
|
||
<paragraph id="8BE38229FFDEFF046CF0417AFC60FF87" blockId="6.[151,1436,1124,1178]" pageId="6" pageNumber="426">
|
||
<emphasis id="B9285E3BFFDEFF046CF0417AFEAEFF64" bold="true" box="[151,275,1124,1147]" pageId="6" pageNumber="426">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C5CF9AAFFDEFF046D7C417BFCABFF64" authority="A. Milne-Edwards, 1873" authorityName="A. Milne-Edwards" authorityYear="1873" box="[283,790,1125,1147]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="426" phylum="Arthropoda" rank="species" species="balansae">
|
||
<emphasis id="B9285E3BFFDEFF046D7C417BFDB2FF65" box="[283,527,1125,1147]" italics="true" pageId="6" pageNumber="426">Clistocoeloma balansae</emphasis>
|
||
A. Milne-Edwards, 1873
|
||
</taxonomicName>
|
||
, male (15.9 × 15.5 mm) (ZRC 2012.702), Philippines. A, male abdomen; B, G1 dorsal view; C, G1 lateral view; D, G1 ventral view. G1 denuded.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE38229FFDEFF046CA041D8FBF8FE54" blockId="6.[151,1437,1222,2002]" pageId="6" pageNumber="426">Walking legs covered with dense coat of setae, with small tufts of setae resembling tubercles, smooth when denuded; longer setae on dorsal, ventral margins of entire leg except dactylus; meri with deep groove subdistally. Third walking leg: merus approximately 2 times longer than wide; combined length of carpus, propodus slightly shorter than merus (c. 0.8 times); propodus approximately 2 times longer than wide.</paragraph>
|
||
<paragraph id="8BE38229FFDEFF046CA04048FAF6FEC4" blockId="6.[151,1437,1222,2002]" pageId="6" pageNumber="426">
|
||
Thoracic sternum setose, covered with short setae. Abdomen of male narrow, widest at somite 3; long setae fringing margins.
|
||
<collectingCountry id="F34BC2B9FFDEFF046D024064FE22FE8C" box="[357,415,1402,1426]" name="Malta" pageId="6" pageNumber="426">Male</collectingCountry>
|
||
abdomen with telson approximately 1.2 times longer than wide. Female abdomen wide, rounded; telson half embedded in somite 6.
|
||
<collectingCountry id="F34BC2B9FFDEFF046EE54080FD06FEA8" box="[642,699,1438,1462]" name="Malta" pageId="6" pageNumber="426">Male</collectingCountry>
|
||
G1 stout, but relatively slender; subdistally wide. Tip of male G1 with dense tufts of setae, single row of setae on exterior margin; wide chitinous tip revealed when denuded.
|
||
</paragraph>
|
||
<paragraph id="8BE38229FFDEFF046CA040F8FBA9FD74" blockId="6.[151,1437,1222,2002]" pageId="6" pageNumber="426">
|
||
<emphasis id="B9285E3BFFDEFF046CA040F8FE33FEE1" bold="true" box="[199,398,1510,1535]" pageId="6" pageNumber="426">Ecological notes.</emphasis>
|
||
This species is found in mangroves. The dense coat of setae on the carapace and legs of the crab is similar in colour as the mud that the crab inhabits, suggesting a possible form of camouflage. According to the collector, this species is very rare in its
|
||
<typeStatus id="54E73C8BFFDEFF046E104331FD1AFD59" box="[631,679,1583,1607]" pageId="6" pageNumber="426">type</typeStatus>
|
||
locality [“
|
||
<emphasis id="B9285E3BFFDEFF046F794331FD2BFD74" italics="true" pageId="6" pageNumber="426">
|
||
Cette espèce est fort rare à la
|
||
<collectingCountry id="F34BC2B9FFDEFF04680A4330FAEDFD58" box="[1133,1360,1582,1606]" name="New Caledonia" pageId="6" pageNumber="426">Nouvelle-Calédonie</collectingCountry>
|
||
, où M. Balansa l’a trouvée au milieu des Palétuviers
|
||
</emphasis>
|
||
”] (A. Milne-Edwards 1873: 312).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346D1A2FFDEFF046CA04368FB3AFD90" box="[199,1159,1654,1679]" pageId="6" pageNumber="426" type="distribution">
|
||
<paragraph id="8BE38229FFDEFF046CA04368FB3AFD90" blockId="6.[151,1437,1222,2002]" box="[199,1159,1654,1679]" pageId="6" pageNumber="426">
|
||
<emphasis id="B9285E3BFFDEFF046CA04368FEDDFD91" bold="true" box="[199,352,1654,1679]" pageId="6" pageNumber="426">Distribution.</emphasis>
|
||
<collectingCountry id="F34BC2B9FFDEFF046D004369FDA5FD90" box="[359,536,1654,1678]" name="New Caledonia" pageId="6" pageNumber="426">New Caledonia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F34BC2B9FFDEFF046E454369FDC3FD90" box="[546,638,1655,1678]" name="Vanuatu" pageId="6" pageNumber="426">Vanuatu</collectingCountry>
|
||
and,
|
||
<collectingCountry id="F34BC2B9FFDEFF046EDB4368FC87FD90" box="[700,826,1654,1678]" name="Philippines" pageId="6" pageNumber="426">Philippines</collectingCountry>
|
||
(Panglao and Palaui Islands).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346D1A2FFDEFF046CA04385FA21FCCC" pageId="6" pageNumber="426" type="discussion">
|
||
<paragraph id="8BE38229FFDEFF046CA04385FC05FC5C" blockId="6.[151,1437,1222,2002]" pageId="6" pageNumber="426">
|
||
<emphasis id="B9285E3BFFDEFF046CA04385FE86FDAD" bold="true" box="[199,315,1691,1715]" pageId="6" pageNumber="426">Remarks.</emphasis>
|
||
Alphonse Milne-Edwards (1873: 311–312) made clear in the description that he had at least one male and one female specimen. All extant specimens are
|
||
<typeStatus id="54E73C8BFFDEFF046F2643A1FC1EFDC9" box="[833,931,1727,1751]" pageId="6" pageNumber="426" type="syntype">syntypes</typeStatus>
|
||
because no
|
||
<typeStatus id="54E73C8BFFDEFF04685C43A0FB20FDC8" box="[1083,1181,1726,1750]" pageId="6" pageNumber="426" type="holotype">holotype</typeStatus>
|
||
was indicated. In the MNHN, there is one dried male and one dried female specimen with the former corresponding in size to the specimen measured by A. Milne-Edwards (1873: 312). The dried male specimen measuring 20.7 ×
|
||
<quantity id="4CA42FCCFFDEFF0469534218FA20FC00" box="[1332,1437,1798,1822]" metricMagnitude="-2" metricUnit="m" metricValue="1.97" pageId="6" pageNumber="426" unit="mm" value="19.7">19.7 mm</quantity>
|
||
(MNHN B10912a) is herein designated as the
|
||
<typeStatus id="54E73C8BFFDEFF046EF84234FCB8FC5C" box="[671,773,1834,1858]" pageId="6" pageNumber="426" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C5CF9AAFFDEFF046F4C4235FC0CFC5C" box="[811,945,1834,1858]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="426" phylum="Arthropoda" rank="species" species="balansae">
|
||
<emphasis id="B9285E3BFFDEFF046F4C4235FC83FC5C" box="[811,830,1835,1858]" italics="true" pageId="6" pageNumber="426">C</emphasis>
|
||
.
|
||
<emphasis id="B9285E3BFFDEFF046F2B4234FC0CFC5C" box="[844,945,1834,1858]" italics="true" pageId="6" pageNumber="426">balansae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE38229FFDEFF046CA04251FA21FCCC" blockId="6.[151,1437,1222,2002]" pageId="6" pageNumber="426">
|
||
<taxonomicName id="4C5CF9AAFFDEFF046CA04251FD1CFC78" authority="Rathbun, 1914" authorityName="Rathbun" authorityYear="1914" box="[199,673,1870,1895]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="426" phylum="Arthropoda" rank="species" species="tectum" subGenus="Sesarma">
|
||
<emphasis id="B9285E3BFFDEFF046CA04251FE95FC78" box="[199,296,1871,1894]" italics="true" pageId="6" pageNumber="426">Sesarma</emphasis>
|
||
(
|
||
<emphasis id="B9285E3BFFDEFF046D5F4251FE25FC78" box="[312,408,1871,1894]" italics="true" pageId="6" pageNumber="426">Sesarma</emphasis>
|
||
)
|
||
<emphasis id="B9285E3BFFDEFF046DCF424EFE4EFC79" box="[424,499,1872,1895]" italics="true" pageId="6" pageNumber="426">tectum</emphasis>
|
||
Rathbun, 1914
|
||
</taxonomicName>
|
||
, was described on the basis of a large female specimen (19.5 ×
|
||
<quantity id="4CA42FCCFFDEFF04690C4250FF7BFC94" metricMagnitude="-2" metricUnit="m" metricValue="1.86" pageId="6" pageNumber="426" unit="mm" value="18.6">18.6 mm</quantity>
|
||
; USNM 45766) from Palaui I., Luzon,
|
||
<collectingCountry id="F34BC2B9FFDEFF046EE2426CFCBAFC94" box="[645,775,1906,1930]" name="Philippines" pageId="6" pageNumber="426">Philippines</collectingCountry>
|
||
. While it was described at length, no figure was provided. The species has never been figured. Tesch (1917) identified one male and three females from Talaud I. (= Talaut Archipelago, off northern Sulawesi) and one female from Nias I. (western Sumatra, Indian Ocean) as
|
||
<taxonomicName id="4C5CF9AAFFDEFF04696B42A5FAD1FCCC" box="[1292,1388,1979,2002]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="426" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9285E3BFFDEFF04696B42A5FAD1FCCC" box="[1292,1388,1979,2002]" italics="true" pageId="6" pageNumber="426">Sesarma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B9285E3BFFDEFF04691A42A5FA2EFCCC" box="[1405,1427,1979,2002]" italics="true" pageId="6" pageNumber="426">S.</emphasis>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346D1A2FFDFFF0A6CF04587FF6BFAB2" lastPageId="8" lastPageNumber="428" pageId="7" pageNumber="427" type="description">
|
||
<paragraph id="8BE38229FFDFFF056CF04587FB7CF861" blockId="7.[151,1436,151,896]" pageId="7" pageNumber="427">
|
||
<taxonomicName id="4C5CF9AAFFDFFF056CF04587FF5FFBAE" box="[151,226,153,176]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="427" phylum="Arthropoda" rank="species" species="tectum">
|
||
<emphasis id="B9285E3BFFDFFF056CF04587FF5FFBAE" box="[151,226,153,176]" italics="true" pageId="7" pageNumber="427">tectum</emphasis>
|
||
</taxonomicName>
|
||
although he did not examine Rathbun’s
|
||
<typeStatus id="54E73C8BFFDFFF056ECF4586FD61FBAE" box="[680,732,152,176]" pageId="7" pageNumber="427">type</typeStatus>
|
||
. Tesch also transferred the species to
|
||
<taxonomicName id="4C5CF9AAFFDFFF0568E54586FA99FBAE" box="[1154,1316,152,176]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="427" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9285E3BFFDFFF0568E54586FA99FBAE" box="[1154,1316,152,176]" italics="true" pageId="7" pageNumber="427">Clistocoeloma</emphasis>
|
||
</taxonomicName>
|
||
. Although he was confident in his identification, Tesch (1917: 223) commented “
|
||
<taxonomicName id="4C5CF9AAFFDFFF056FCB45A2FBA2FBCA" box="[940,1055,188,212]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="427" phylum="Arthropoda" rank="species" species="tectum">
|
||
<emphasis id="B9285E3BFFDFFF056FCB45A2FC7BFBCA" box="[940,966,188,212]" italics="true" pageId="7" pageNumber="427">Cl</emphasis>
|
||
.
|
||
<emphasis id="B9285E3BFFDFFF056FB345A3FBA2FBCA" box="[980,1055,189,212]" italics="true" pageId="7" pageNumber="427">tectum</emphasis>
|
||
</taxonomicName>
|
||
bears a very great resemblance to
|
||
<taxonomicName id="4C5CF9AAFFDFFF056CF045FEFDBAFBE9" authority="A. Milne-Edwards" authorityName="A. Milne-Edwards" box="[151,519,223,248]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="427" phylum="Arthropoda" rank="species" species="balansae">
|
||
<emphasis id="B9285E3BFFDFFF056CF045FEFF0FFBE6" box="[151,178,224,248]" italics="true" pageId="7" pageNumber="427">Cl</emphasis>
|
||
.
|
||
<emphasis id="B9285E3BFFDFFF056CA545FEFE9AFBE6" box="[194,295,224,248]" italics="true" pageId="7" pageNumber="427">balansae</emphasis>
|
||
A. Milne-Edwards
|
||
</taxonomicName>
|
||
, on account of the carapace being quadrate, and the distance between the outer orbital angles only slightly or not at all exceeding the length of the carapace, and the external postfrontal lobes being subdivided into two small tubercles. It differs by the course of the upper orbital border that is not waved, by the tubercles on the carapace, the shape of the abdomen of the 3 and probably by some other characters”. Tesch must have based his observations on the different male abdominal shapes on his single male specimen and the figure in A. Milne-Edwards (1873) since Rathbun’s (1914)
|
||
<typeStatus id="54E73C8BFFDFFF056F5C448BFCD6FAB3" box="[827,875,405,429]" pageId="7" pageNumber="427">type</typeStatus>
|
||
was a female. While the form of the supraorbital margin is somewhat variable, the male abdomen as figured by A. Milne-Edwards (1873: pl 17, fig. 1b) is unusual, being evenly triangular, unlike that of typical
|
||
<taxonomicName id="4C5CF9AAFFDFFF056ED144C2FCE5FAEA" box="[694,856,476,500]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="427" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9285E3BFFDFFF056ED144C2FCE5FAEA" box="[694,856,476,500]" italics="true" pageId="7" pageNumber="427">Clistocoeloma</emphasis>
|
||
</taxonomicName>
|
||
(and most sesarmids) that is more quadrate with somite 6 relatively broader (e.g., in the present
|
||
<taxonomicName id="4C5CF9AAFFDFFF056EC4471EFC95F906" box="[675,808,512,536]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="427" phylum="Arthropoda" rank="species" species="balansae">
|
||
<emphasis id="B9285E3BFFDFFF056EC4471EFC95F906" box="[675,808,512,536]" italics="true" pageId="7" pageNumber="427">C. balansae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="13679EACFFDFFF056F5144E1FC3CF909" box="[822,897,511,535]" captionStart="FIGURE 3" captionStartId="4.[151,250,1704,1727]" captionTargetBox="[188,1398,176,1685]" captionTargetId="figure@4.[188,1398,175,1687]" captionTargetPageId="4" captionText="FIGURE 3. Clistocoeloma balansae A. Milne-Edwards, 1873. A – C, male (15.9 × 15.5 mm) (ZRC 2012.702), Philippines; D, female (17.2 × 16.9 mm) (ZRC 2012.702), Philippines. A, dorsal overall view; B, ventral view of abdomen and thoracic sternum; C, D posterior view of carapace." httpUri="https://zenodo.org/record/222416/files/figure.png" pageId="7" pageNumber="427">Figs. 3</figureCitation>
|
||
B, 6A). The telson of the dried
|
||
<typeStatus id="54E73C8BFFDFFF0568BC44E1FAFFF909" box="[1243,1346,511,535]" pageId="7" pageNumber="427" type="lectotype">lectotype</typeStatus>
|
||
male of
|
||
<taxonomicName id="4C5CF9AAFFDFFF056CF0473BFEA2F922" box="[151,287,548,572]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="427" phylum="Arthropoda" rank="species" species="balansae">
|
||
<emphasis id="B9285E3BFFDFFF056CF0473BFF17F922" box="[151,170,549,572]" italics="true" pageId="7" pageNumber="427">C</emphasis>
|
||
.
|
||
<emphasis id="B9285E3BFFDFFF056CDD473AFEA2F922" box="[186,287,548,572]" italics="true" pageId="7" pageNumber="427">balansae</emphasis>
|
||
</taxonomicName>
|
||
is damaged by glue, but the shape of the other somites (
|
||
<figureCitation id="13679EACFFDFFF056FC9473AFC52F922" box="[942,1007,548,572]" captionStart="FIGURE 1" captionStartId="2.[151,250,1910,1933]" captionTargetBox="[188,1398,301,1888]" captionTargetId="figure@2.[188,1398,301,1889]" captionTargetPageId="2" captionText="FIGURE 1. Clistocoeloma balansae A. Milne-Edwards, 1873, lectotype male (20.7 × 19.7 mm) (MNHN B 10912), New Caledonia. A, dorsal overall view; B, ventral view of abdomen and thoracic sternum; C, frontal view of carapace. (Photographs by T. Naruse)" httpUri="https://zenodo.org/record/222414/files/figure.png" pageId="7" pageNumber="427">Fig. 1</figureCitation>
|
||
B) is typical for many
|
||
<taxonomicName id="4C5CF9AAFFDFFF05689D473AFA21F922" box="[1274,1436,548,572]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="427" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9285E3BFFDFFF05689D473AFA21F922" box="[1274,1436,548,572]" italics="true" pageId="7" pageNumber="427">Clistocoeloma</emphasis>
|
||
</taxonomicName>
|
||
species and does not match the drawing in A. Milne-Edwards (1873: pl. 17 fig. 1b). The abdomen of “
|
||
<taxonomicName id="4C5CF9AAFFDFFF05696D4756FA32F97E" box="[1290,1423,584,608]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="427" phylum="Arthropoda" rank="species" species="balansae">
|
||
<emphasis id="B9285E3BFFDFFF05696D4756FAA0F941" box="[1290,1309,584,607]" italics="true" pageId="7" pageNumber="427">C</emphasis>
|
||
.
|
||
<emphasis id="B9285E3BFFDFFF05694D4756FA32F97E" box="[1322,1423,584,608]" italics="true" pageId="7" pageNumber="427">balansae</emphasis>
|
||
</taxonomicName>
|
||
” figured by A. Milne-Edwards almost certainly belongs to a species of the family
|
||
<taxonomicName id="4C5CF9AAFFDFFF05684B4772FB20F99A" box="[1068,1181,620,644]" class="Malacostraca" family="Varunidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="427" phylum="Arthropoda" rank="family">Varunidae</taxonomicName>
|
||
and was probably just mislabeled; this seems especially likely since two varunids were depicted on the same plate, viz.
|
||
<taxonomicName id="4C5CF9AAFFDFFF056892478EFDFDF9D2" authority="A. Milne-Edwards, 1873" authorityName="A. Milne-Edwards" authorityYear="1873" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="427" phylum="Arthropoda" rank="species" species="barbatus">
|
||
<emphasis id="B9285E3BFFDFFF056892478EFF46F9D2" italics="true" pageId="7" pageNumber="427">Ptychognathus barbatus</emphasis>
|
||
(A. Milne-Edwards, 1873)
|
||
</taxonomicName>
|
||
(as
|
||
<taxonomicName id="4C5CF9AAFFDFFF056E1B47AAFC26F9D2" box="[636,923,692,716]" class="Malacostraca" family="Varunidae" genus="Gnathograpsus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="427" phylum="Arthropoda" rank="species" species="barbatus">
|
||
<emphasis id="B9285E3BFFDFFF056E1B47AAFC26F9D2" box="[636,923,692,716]" italics="true" pageId="7" pageNumber="427">Gnathograpsus barbatus</emphasis>
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName id="4C5CF9AAFFDFFF056F8E47AAFAB2F9D2" box="[1001,1295,692,716]" class="Malacostraca" family="Varunidae" genus="Pseudograpsus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="427" phylum="Arthropoda" rank="species" species="elongatus">
|
||
<emphasis id="B9285E3BFFDFFF056F8E47AAFAB2F9D2" box="[1001,1295,692,716]" italics="true" pageId="7" pageNumber="427">Pseudograpsus elongatus</emphasis>
|
||
</taxonomicName>
|
||
(A. Milne- Edwards, 1873) (as
|
||
<taxonomicName id="4C5CF9AAFFDFFF056D1047C7FD2CF9EE" box="[375,657,728,752]" class="Malacostraca" family="Varunidae" genus="Heterograpsus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="427" phylum="Arthropoda" rank="species" species="elongatus">
|
||
<emphasis id="B9285E3BFFDFFF056D1047C7FD2CF9EE" box="[375,657,728,752]" italics="true" pageId="7" pageNumber="427">Heterograpsus elongatus</emphasis>
|
||
</taxonomicName>
|
||
). Other than the shape of the male abdomen, we can discern no other major differences between the specimens described by Tesch (1917) and the present specimens of
|
||
<taxonomicName id="4C5CF9AAFFDFFF05697447E3FA21F80A" box="[1299,1436,764,788]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="427" phylum="Arthropoda" rank="species" species="balansae">
|
||
<emphasis id="B9285E3BFFDFFF05697447E3FA9BF80A" box="[1299,1318,765,788]" italics="true" pageId="7" pageNumber="427">C</emphasis>
|
||
.
|
||
<emphasis id="B9285E3BFFDFFF05695047E2FA21F80A" box="[1335,1436,764,788]" italics="true" pageId="7" pageNumber="427">balansae</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F34BC2B9FFDFFF056CB2463FFE3AF829" box="[213,391,799,824]" name="New Caledonia" pageId="7" pageNumber="427">New Caledonia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F34BC2B9FFDFFF056DF5463FFE50F826" box="[402,493,801,824]" name="Vanuatu" pageId="7" pageNumber="427">Vanuatu</collectingCountry>
|
||
and the
|
||
<collectingCountry id="F34BC2B9FFDFFF056E284601FD70F829" box="[591,717,799,823]" name="Philippines" pageId="7" pageNumber="427">Philippines</collectingCountry>
|
||
and we provisionally treat his records as this species. The male specimen from Talaud I. is likely to be real
|
||
<taxonomicName id="4C5CF9AAFFDFFF056EED465BFCACF842" box="[650,785,836,860]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="427" phylum="Arthropoda" rank="species" species="balansae">
|
||
<emphasis id="B9285E3BFFDFFF056EED465BFD20F842" box="[650,669,837,860]" italics="true" pageId="7" pageNumber="427">C</emphasis>
|
||
.
|
||
<emphasis id="B9285E3BFFDFFF056ECB465AFCACF842" box="[684,785,836,860]" italics="true" pageId="7" pageNumber="427">balansae</emphasis>
|
||
</taxonomicName>
|
||
as these islands are just south of the
|
||
<collectingCountry id="F34BC2B9FFDFFF0568DC465AFA80F842" box="[1211,1341,836,860]" name="Philippines" pageId="7" pageNumber="427">Philippines</collectingCountry>
|
||
. Tesch’s (1917) specimens from Nias will need to be re-examined as this locality is in the Indian Ocean.
|
||
</paragraph>
|
||
<caption id="DF23D2A1FFDFFF056CF04246FC7BFC92" httpUri="https://zenodo.org/record/222419/files/figure.png" pageId="7" pageNumber="427" targetBox="[216,1398,1033,1780]" targetPageId="7">
|
||
<paragraph id="8BE38229FFDFFF056CF04246FC7BFC92" blockId="7.[151,1436,1880,1932]" pageId="7" pageNumber="427">
|
||
<emphasis id="B9285E3BFFDFFF056CF04246FEA9FC73" bold="true" box="[151,276,1880,1903]" pageId="7" pageNumber="427">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C5CF9AAFFDFFF056D7C4246FCA8FC70" authority="A. Milne-Edwards, 1873" authorityName="A. Milne-Edwards" authorityYear="1873" box="[283,789,1880,1902]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="427" phylum="Arthropoda" rank="species" species="balansae">
|
||
<emphasis id="B9285E3BFFDFFF056D7C4246FDADFC70" box="[283,528,1880,1902]" italics="true" pageId="7" pageNumber="427">Clistocoeloma balansae</emphasis>
|
||
A. Milne-Edwards, 1873
|
||
</taxonomicName>
|
||
(after A. Milne-Edwards 1873: pl. 17 fig. 1). A, dorsal overall view; B, frontal view of carapace; C, male abdomen; D, outer view of left chela.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE38229FFD0FF0A6CA04589FF6BFAB2" blockId="8.[151,1436,151,428]" pageId="8" pageNumber="428">
|
||
The
|
||
<typeStatus id="54E73C8BFFD0FF0A6C9E4589FEE6FBB1" box="[249,347,151,175]" pageId="8" pageNumber="428" type="holotype">holotype</typeStatus>
|
||
of
|
||
<emphasis id="B9285E3BFFD0FF0A6DE64586FE32FBB1" box="[385,399,152,175]" italics="true" pageId="8" pageNumber="428">S</emphasis>
|
||
. (
|
||
<emphasis id="B9285E3BFFD0FF0A6DC14586FE09FBB1" box="[422,436,152,175]" italics="true" pageId="8" pageNumber="428">S</emphasis>
|
||
.)
|
||
<taxonomicName id="4C5CF9AAFFD0FF0A6DAC4587FDABFBAE" box="[459,534,153,176]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="428" phylum="Arthropoda" rank="species" species="tectum">
|
||
<emphasis id="B9285E3BFFD0FF0A6DAC4587FDABFBAE" box="[459,534,153,176]" italics="true" pageId="8" pageNumber="428">tectum</emphasis>
|
||
</taxonomicName>
|
||
is one of the largest
|
||
<taxonomicName id="4C5CF9AAFFD0FF0A6E994586FC1DFBAE" box="[766,928,152,176]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="428" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9285E3BFFD0FF0A6E994586FC1DFBAE" box="[766,928,152,176]" italics="true" pageId="8" pageNumber="428">Clistocoeloma</emphasis>
|
||
</taxonomicName>
|
||
specimens known and can be separated from
|
||
<taxonomicName id="4C5CF9AAFFD0FF0A6CF045A3FEA0FBCA" box="[151,285,188,212]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="428" phylum="Arthropoda" rank="species" species="balansae">
|
||
<emphasis id="B9285E3BFFD0FF0A6CF045A3FF17FBCA" box="[151,170,189,212]" italics="true" pageId="8" pageNumber="428">C</emphasis>
|
||
.
|
||
<emphasis id="B9285E3BFFD0FF0A6CDF45A2FEA0FBCA" box="[184,285,188,212]" italics="true" pageId="8" pageNumber="428">balansae</emphasis>
|
||
</taxonomicName>
|
||
only by the form of the posterior carapace margin. The margin is slightly sinuous to almost straight in the typical
|
||
<taxonomicName id="4C5CF9AAFFD0FF0A6D7D45FEFE1EFBE6" box="[282,419,224,248]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="428" phylum="Arthropoda" rank="species" species="balansae">
|
||
<emphasis id="B9285E3BFFD0FF0A6D7D45FEFE90FBE9" box="[282,301,224,247]" italics="true" pageId="8" pageNumber="428">C</emphasis>
|
||
.
|
||
<emphasis id="B9285E3BFFD0FF0A6D5945FEFE1EFBE6" box="[318,419,224,248]" italics="true" pageId="8" pageNumber="428">balansae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13679EACFFD0FF0A6DD245C1FDBBFBE9" box="[437,518,223,247]" captionStart="FIGURE 3" captionStartId="4.[151,250,1704,1727]" captionTargetBox="[188,1398,176,1685]" captionTargetId="figure@4.[188,1398,175,1687]" captionTargetPageId="4" captionText="FIGURE 3. Clistocoeloma balansae A. Milne-Edwards, 1873. A – C, male (15.9 × 15.5 mm) (ZRC 2012.702), Philippines; D, female (17.2 × 16.9 mm) (ZRC 2012.702), Philippines. A, dorsal overall view; B, ventral view of abdomen and thoracic sternum; C, D posterior view of carapace." httpUri="https://zenodo.org/record/222416/files/figure.png" pageId="8" pageNumber="428">Figs. 3</figureCitation>
|
||
C, D) whereas, the posterior carapace margin is deeply concave medially in
|
||
<taxonomicName id="4C5CF9AAFFD0FF0A69E545FEFF5FFA02" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="428" phylum="Arthropoda" rank="species" species="tectum">
|
||
<emphasis id="B9285E3BFFD0FF0A69E545FEFF5FFA02" italics="true" pageId="8" pageNumber="428">C. tectum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13679EACFFD0FF0A6C93441AFE81FA02" box="[244,316,260,284]" captionStart="FIGURE 2" captionStartId="3.[151,250,1726,1749]" captionTargetBox="[188,1398,193,1703]" captionTargetId="figure@3.[188,1398,193,1705]" captionTargetPageId="3" captionText="FIGURE 2. Sesarma (Sesarm a) tectum Rathbun, 1914, holotype female (19.5 × 18.6 mm) (USNM 45766), Philippines. A, dorsal overall view; B, ventral view of abdomen and thoracic sternum; C, frontal view of carapace; D, posterior view of carapace." httpUri="https://zenodo.org/record/222415/files/figure.png" pageId="8" pageNumber="428">Fig. 2</figureCitation>
|
||
D). A careful examination of the posterior carapace margin of the
|
||
<typeStatus id="54E73C8BFFD0FF0A682F441AFB17FA02" box="[1096,1194,260,284]" pageId="8" pageNumber="428" type="holotype">holotype</typeStatus>
|
||
female of
|
||
<taxonomicName id="4C5CF9AAFFD0FF0A694A441BFA21FA02" box="[1325,1436,261,284]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="428" phylum="Arthropoda" rank="species" species="tectum">
|
||
<emphasis id="B9285E3BFFD0FF0A694A441BFAFDFA02" box="[1325,1344,261,284]" italics="true" pageId="8" pageNumber="428">C</emphasis>
|
||
.
|
||
<emphasis id="B9285E3BFFD0FF0A6936441BFA21FA02" box="[1361,1436,261,284]" italics="true" pageId="8" pageNumber="428">tectum</emphasis>
|
||
</taxonomicName>
|
||
shows that the margin is slightly asymmetrical, indicating that an injury had occurred previously and the present form is the result of uneven growth after healing. Other than this feature, it agrees with
|
||
<taxonomicName id="4C5CF9AAFFD0FF0A68EA4453FAA8FA7A" box="[1165,1301,332,356]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="428" phylum="Arthropoda" rank="species" species="balansae">
|
||
<emphasis id="B9285E3BFFD0FF0A68EA4453FB1DFA7A" box="[1165,1184,333,356]" italics="true" pageId="8" pageNumber="428">C</emphasis>
|
||
.
|
||
<emphasis id="B9285E3BFFD0FF0A68D74452FAA8FA7A" box="[1200,1301,332,356]" italics="true" pageId="8" pageNumber="428">balansae</emphasis>
|
||
</taxonomicName>
|
||
in all other diagnostic characters.
|
||
<collectingCountry id="F34BC2B9FFD0FF0A6DF6446EFE0DFA96" box="[401,432,368,392]" name="American Samoa" pageId="8" pageNumber="428">As</collectingCountry>
|
||
such, we synonymise
|
||
<emphasis id="B9285E3BFFD0FF0A6ECA446EFD06FA99" box="[685,699,368,391]" italics="true" pageId="8" pageNumber="428">S</emphasis>
|
||
. (
|
||
<emphasis id="B9285E3BFFD0FF0A6EB4446EFD5CFA99" box="[723,737,368,391]" italics="true" pageId="8" pageNumber="428">S</emphasis>
|
||
.)
|
||
<taxonomicName id="4C5CF9AAFFD0FF0A6E9E446FFC4EFA99" authority="Rathbun, 1914" authorityName="Rathbun" authorityYear="1914" box="[761,1011,367,392]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="428" phylum="Arthropoda" rank="species" species="tectum">
|
||
<emphasis id="B9285E3BFFD0FF0A6E9E446FFCF9FA96" box="[761,836,369,392]" italics="true" pageId="8" pageNumber="428">tectum</emphasis>
|
||
Rathbun, 1914
|
||
</taxonomicName>
|
||
, with
|
||
<taxonomicName id="4C5CF9AAFFD0FF0A6850446EFF6FFAB2" authority="A. Milne-Edwards, 1873" authorityName="A. Milne-Edwards" authorityYear="1873" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="428" phylum="Arthropoda" rank="species" species="balansae">
|
||
<emphasis id="B9285E3BFFD0FF0A6850446EFBF7FA99" box="[1079,1098,368,391]" italics="true" pageId="8" pageNumber="428">C</emphasis>
|
||
.
|
||
<emphasis id="B9285E3BFFD0FF0A683F446EFB00FA96" box="[1112,1213,368,392]" italics="true" pageId="8" pageNumber="428">balansae</emphasis>
|
||
A. Milne-Edwards, 1873
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |