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<document id="C4101C048C7F8491279CFD77C163AE51" ID-CLB-Dataset="30161" ID-DOI="10.11646/zootaxa.4415.1.3" ID-GBIF-Dataset="7d19d012-8cc3-4355-88d9-e90526d4dcef" ID-ISSN="1175-5326" ID-Zenodo-Dep="1241791" ID-ZooBank="EB339387-497F-47E1-918E-0EC5060F2D90" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1525597777960" checkinUser="plazi" docAuthor="Malipatil, M. B. &amp; Scudder, G. G. E." docDate="2018" docId="03F08D2BFF9F0E73DBA5FC2FFA74F80A" docLanguage="en" docName="zootaxa.4415.1.3.pdf.imf" docOrigin="Zootaxa 4415 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraindopamphantus bruneiensis Malipatil &amp; Scudder 2018, sp. nov." docType="treatment" docVersion="7" masterDocId="FFC9F553FF9D0E78DB32FFA1FFE8FFDB" masterDocTitle="ParaindOPaMPHantus bruneiensis gen. nov. et sp. nov., as the first representative of the subfamily Pamphantinae from South East Asia (Hemiptera: Heteroptera: Geocoridae: Pamphantinae: Indopamphantini)" updateTime="1698540943884" updateUser="ExternalLinkService">
<mods:mods id="D8F4387670E0C67D3BBA6F19D82A23A3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AA8357E278750BC28776084457EE17E6">
<mods:title id="9820BD654B3C79570A5C4D9336C6F3A7">ParaindOPaMPHantus bruneiensis gen. nov. et sp. nov., as the first representative of the subfamily Pamphantinae from South East Asia (Hemiptera: Heteroptera: Geocoridae: Pamphantinae: Indopamphantini)</mods:title>
</mods:titleInfo>
<mods:name id="158CCCEBB6942FD8FAA33461686EA730" type="personal">
<mods:role id="59EAEE7A6ADD5C620E885E54565D8802">
<mods:roleTerm id="503CB696A96E1D790B779CD30DDF00C4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A9D51CF5126A0EE7D806B5A79136E3BC">Malipatil, M. B.</mods:namePart>
</mods:name>
<mods:name id="D660CCDA7725FBC6251B885B82196029" type="personal">
<mods:role id="B05BE4BBB3112A2C41BA62020D469307">
<mods:roleTerm id="E04ECD14127BF020D44F837C97E36C1B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5407288EBB7EC5D703709C7630D6284C">Scudder, G. G. E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="AA9C41CEE2F6D5670F72439C316CF2A0">text</mods:typeOfResource>
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<mods:title id="7C5A9FA897AA5848920A4C5245C63AA4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C1F363A6FB852616DD5D4FEC9A40328D">
<mods:date id="E6F5E6140F05D3F0A4B92A1994ACC2C6">2018</mods:date>
<mods:detail id="0E4272273AB1097C3DABE3D1F551C99D" type="pubDate">
<mods:number id="5709B3FB875953A60F76441E983227F8">2018-04-27</mods:number>
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<mods:detail id="9C9C8FBC970034F2575DA26B0ED77508" type="volume">
<mods:number id="EB3D6B33858B0450A79F03E637D5F7BE">4415</mods:number>
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<mods:number id="7D8C6BDFE9ED46DE24D61DCB79AA30B5">1</mods:number>
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<mods:classification id="8D41D58ECD3E294E614EB050127607DC">journal volume</mods:classification>
<mods:identifier id="DA9109608F4991C67FB009F03BC35278" type="CLB-Dataset">30161</mods:identifier>
<mods:identifier id="33DDEFE32172701F6339C884C21697E4" type="DOI">10.11646/zootaxa.4415.1.3</mods:identifier>
<mods:identifier id="A247DDAEF273F18E9D16DDDA6B7E78D0" type="GBIF-Dataset">7d19d012-8cc3-4355-88d9-e90526d4dcef</mods:identifier>
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<treatment id="03F08D2BFF9F0E73DBA5FC2FFA74F80A" ID-DOI="http://doi.org/10.5281/zenodo.5986813" ID-GBIF-Taxon="143811186" ID-Zenodo-Dep="5986813" LSID="urn:lsid:plazi:treatment:03F08D2BFF9F0E73DBA5FC2FFA74F80A" httpUri="http://treatment.plazi.org/id/03F08D2BFF9F0E73DBA5FC2FFA74F80A" lastPageId="11" pageId="2">
<subSubSection id="C3436FB6FF9F0E7ADBA5FC2FFEC8FC11" pageId="2" type="nomenclature">
<paragraph id="8BE63C3DFF9F0E7ADBA5FC2FFD7DFC73" blockId="2.[151,661,910,970]" box="[151,661,910,936]" pageId="2">
<heading id="D0AE8B51FF9F0E7ADBA5FC2FFD7DFC73" bold="true" box="[151,661,910,936]" fontSize="11" level="1" pageId="2" reason="1">
<emphasis id="B92DE02FFF9F0E7ADBA5FC2FFD7DFC73" bold="true" box="[151,661,910,936]" pageId="2">
<taxonomicName id="4C5947BEFF9F0E7ADBA5FC2FFDD8FC73" ID-CoL="4CX8F" authority="Malipatil &amp; Scudder, 2018" authorityName="Malipatil &amp; Scudder" authorityYear="2018" box="[151,560,910,936]" class="Insecta" family="Geocoridae" genus="Paraindopamphantus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" phylum="Arthropoda" rank="species" species="bruneiensis" status="sp. nov.">
<emphasis id="B92DE02FFF9F0E7ADBA5FC2FFDD8FC73" bold="true" box="[151,560,910,936]" italics="true" pageId="2">Paraindopamphantus bruneiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21E5D54FF9F0E7AD90AFC2FFD7DFC73" box="[568,661,910,936]" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE63C3DFF9F0E7ADBA5FC10FEC8FC11" blockId="2.[151,661,910,970]" box="[151,288,945,970]" pageId="2">
(
<figureCitation id="136220B8FF9F0E7ADBADFC10FEFFFC11" box="[159,279,945,970]" captionStart-0="FIGURES 12" captionStart-1="FIGURES 34" captionStart-2="FIGURES 56" captionStart-3="FIGURES 78" captionStart-4="FIGURES 910" captionStart-5="FIGURES 1112" captionStart-6="FIGURES 1314" captionStart-7="FIGURES 1517" captionStartId-0="3.[151,264,1975,1998]" captionStartId-1="4.[151,264,1965,1988]" captionStartId-2="5.[151,264,1965,1988]" captionStartId-3="6.[151,264,1964,1987]" captionStartId-4="7.[151,264,1801,1824]" captionStartId-5="8.[151,264,1573,1596]" captionStartId-6="9.[151,264,1668,1691]" captionStartId-7="10.[151,264,1935,1958]" captionTargetBox-0="[151,1436,360,1921]" captionTargetBox-1="[371,1220,193,1946]" captionTargetBox-2="[367,1217,184,1940]" captionTargetBox-3="[151,1435,212,1922]" captionTargetBox-4="[151,1436,190,1780]" captionTargetBox-5="[151,1436,298,1552]" captionTargetBox-6="[151,1436,304,1648]" captionTargetBox-7="[186,1406,190,1918]" captionTargetId-0="figure@3.[151,1436,360,1922]" captionTargetId-1="figure@4.[371,1220,190,1946]" captionTargetId-2="figure@5.[367,1217,184,1940]" captionTargetId-3="figure@6.[151,1436,212,1923]" captionTargetId-4="figure@7.[151,1436,190,1780]" captionTargetId-5="figure@8.[151,1436,298,1552]" captionTargetId-6="figure@9.[151,1436,304,1648]" captionTargetId-7="figure@10.[186,1407,190,1918]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionText-0="FIGURES 12. Paraindopamphantus bruneiensis, holotype female:1, habitus, dorsal view; 2, habitus, lateral view." captionText-1="FIGURES 34. Paraindopamphantus bruneiensis, holotype female: 3, head and pronotum, dorsal view; 4, head and part of pronotum, lateral view. gs, gular swelling." captionText-2="FIGURES 56. Paraindopamphantus bruneiensis, holotype female: 5, head, ventral view; 6, gular swelling, enlarged. gs, gular swelling." captionText-3="FIGURES 78. Paraindopamphantus bruneiensis, holotype female: 7, meso- and metathoracic area, lateral view; 8, posterior part of thorax and abdomen, dorsal view.sgsp, scent gland spout; ssp, scutellar spine." captionText-4="FIGURES 910. Paraindopamphantus bruneiensis, holotype female: 9, part of thorax and abdomen, ventral view; 10, abdominal dorsum. sgs, scent gland scar; sgsp, scent gland spout; IIVII, abdominal tergites." captionText-5="FIGURES 1112. Paraindopamphantus bruneiensis, holotype female: 11, abdominal terga; 12, abdomen, lateral view. sgs, scent gland scar; sp, spiracle; IIVII, abdominal tergites." captionText-6="FIGURES 1314. Paraindopamphantus bruneiensis, holotype female, abdominal venter. tr, trichobothria; IIIV, abdominal sternites." captionText-7="FIGURES 1517. Paraindopamphantus bruneiensis, holotype female: 15, spermatheca; 16,17, ovipositor. Abbreviations: ga, gonangulum; 1gpo, 2gpo, first and second gonapophysis; 1gx, 2gx, first and second gonocoxa; 1r, 2r, first and second ramus; sb, spermathecal bulb; spd, spermathecal duct." httpUri-0="https://zenodo.org/record/1241793/files/figure.png" httpUri-1="https://zenodo.org/record/1241795/files/figure.png" httpUri-2="https://zenodo.org/record/1241797/files/figure.png" httpUri-3="https://zenodo.org/record/1241799/files/figure.png" httpUri-4="https://zenodo.org/record/1241801/files/figure.png" httpUri-5="https://zenodo.org/record/1241803/files/figure.png" httpUri-6="https://zenodo.org/record/1241805/files/figure.png" httpUri-7="https://zenodo.org/record/1241807/files/figure.png" pageId="2">Figs. 117</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3436FB6FF9F0E7ADBA5FC58FB02FBEE" pageId="2" type="materials_examined">
<paragraph id="8BE63C3DFF9F0E7ADBA5FC58FB02FBEE" blockId="2.[151,1437,1017,2025]" pageId="2">
<materialsCitation id="3B313660FF9F0E7ADBA5FC58FB02FBEE" ID-GBIF-Occurrence="1841139505" collectingDate="1982-08-20" collectingDateMax="1982-09-10" collectingDateMin="1982-08-20" collectionCode="NHM" collectorName="Lamunin, N. E &amp; Stork, BM" country="Brunei" location="Type" pageId="2" specimenCount="2" typeStatus="holotype">
<emphasis id="B92DE02FFF9F0E7ADBA5FC58FE88FBC9" bold="true" box="[151,352,1017,1042]" pageId="2">
<typeStatus id="54E2829FFF9F0E7ADBA5FC58FF3AFBC9" box="[151,210,1017,1042]" pageId="2">
<location id="8E866AE6FF9F0E7ADBA5FC58FF3AFBC9" LSID="urn:lsid:plazi:treatment:03F08D2BFF9F0E73DBA5FC2FFA74F80A:8E866AE6FF9F0E7ADBA5FC58FF3AFBC9" box="[151,210,1017,1042]" country="Brunei" name="Type" pageId="2">Type</location>
</typeStatus>
specimens:
</emphasis>
<typeStatus id="54E2829FFF9F0E7ADA5FFC58FE3DFBC9" box="[365,469,1017,1042]" pageId="2" type="holotype">Holotype</typeStatus>
female,
<collectingCountry id="F34E7CADFF9F0E7AD973FC5BFD44FBCA" box="[577,684,1018,1041]" name="Brunei" pageId="2">BRUNEI</collectingCountry>
, Bukit Sulang, nr
<collectorName id="26AC59EBFF9F0E7AD8BFFC58FBDDFBCA" box="[909,1077,1017,1042]" pageId="2">Lamunin, N.E</collectorName>
.
<collectorName id="26AC59EBFF9F0E7ADF77FC58FB29FBCA" box="[1093,1217,1017,1042]" pageId="2">Stork, BM</collectorName>
1982-388 fogging
<date id="FFE71AFDFF9F0E7ADBA5FBBCFE80FBED" box="[151,360,1053,1078]" pageId="2" value="1982-08-20" valueMax="1982-09-10" valueMin="1982-08-20">
<collectingDate id="EFA3E315FF9F0E7ADBA5FBBCFE80FBED" box="[151,360,1053,1078]" pageId="2" value="1982-08-20" valueMax="1982-09-10" valueMin="1982-08-20">20.viii10.ix.1982</collectingDate>
</date>
; insecticide fogging Tree 16. Dipterocarp.
<taxonomicName id="4C5947BEFF9F0E7AD86AFBBCFB9CFBED" authority="Foxw." authorityName="Foxw." box="[856,1140,1053,1078]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Shorea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="2" phylum="Tracheophyta" rank="species" species="johorensis">
<emphasis id="B92DE02FFF9F0E7AD86AFBBCFBC9FBED" box="[856,1057,1053,1078]" italics="true" pageId="2">Shorea johorensis</emphasis>
Foxw.
</taxonomicName>
, in
<collectionCode id="ED48A4F8FF9F0E7ADF93FBBFFB0EFBEE" box="[1185,1254,1054,1077]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="2">NHM</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3436FB6FF9F0E73DBF5FBE0FB54F96A" lastPageId="11" pageId="2" type="description">
<paragraph id="8BE63C3DFF9F0E7ADBF5FBE0FA89FACA" blockId="2.[151,1437,1017,2025]" pageId="2">
<emphasis id="B92DE02FFF9F0E7ADBF5FBE0FEB1FB81" bold="true" box="[199,345,1089,1114]" pageId="2">Description.</emphasis>
Body ground colour black, shiny to sub-shiny (
<figureCitation id="136220B8FF9F0E7AD8B0FBE0FBE3FB81" box="[898,1035,1089,1114]" captionStart="FIGURES 12" captionStartId="3.[151,264,1975,1998]" captionTargetBox="[151,1436,360,1921]" captionTargetId="figure@3.[151,1436,360,1922]" captionTargetPageId="3" captionText="FIGURES 12. Paraindopamphantus bruneiensis, holotype female:1, habitus, dorsal view; 2, habitus, lateral view." httpUri="https://zenodo.org/record/1241793/files/figure.png" pageId="2">Figs. 1 &amp; 2</figureCitation>
). Head above postero-laterally and behind eyes with pale markings. Eyes reddish-fuscous. Antennae with distal 2 segments fuscous with basal areas lighter as for 2nd and part of 1
<taxonomicName id="4C5947BEFF9F0E7ADAD8FB29FDB2FB7E" box="[490,602,1160,1189]" pageId="2" rank="subSpecies" subSpecies="segment">st segment</taxonomicName>
(
<figureCitation id="136220B8FF9F0E7AD959FB2DFD5AFB7E" box="[619,690,1164,1189]" captionStart="FIGURES 34" captionStartId="4.[151,264,1965,1988]" captionTargetBox="[371,1220,193,1946]" captionTargetId="figure@4.[371,1220,190,1946]" captionTargetPageId="4" captionText="FIGURES 34. Paraindopamphantus bruneiensis, holotype female: 3, head and pronotum, dorsal view; 4, head and part of pronotum, lateral view. gs, gular swelling." httpUri="https://zenodo.org/record/1241795/files/figure.png" pageId="2">Fig. 4</figureCitation>
). Pronotal posterior lobe with a median elongate light patch as in
<figureCitation id="136220B8FF9F0E7ADBA5FB11FF37FB13" box="[151,223,1200,1225]" captionStart="FIGURES 34" captionStartId="4.[151,264,1965,1988]" captionTargetBox="[371,1220,193,1946]" captionTargetId="figure@4.[371,1220,190,1946]" captionTargetPageId="4" captionText="FIGURES 34. Paraindopamphantus bruneiensis, holotype female: 3, head and pronotum, dorsal view; 4, head and part of pronotum, lateral view. gs, gular swelling." httpUri="https://zenodo.org/record/1241795/files/figure.png" pageId="2">Fig. 3</figureCitation>
. Scutellum uniformly black. Hemelytra as in
<figureCitation id="136220B8FF9F0E7AD9CCFB11FCAEFB13" box="[766,838,1200,1225]" captionStart="FIGURES 78" captionStartId="6.[151,264,1964,1987]" captionTargetBox="[151,1435,212,1922]" captionTargetId="figure@6.[151,1436,212,1923]" captionTargetPageId="6" captionText="FIGURES 78. Paraindopamphantus bruneiensis, holotype female: 7, meso- and metathoracic area, lateral view; 8, posterior part of thorax and abdomen, dorsal view.sgsp, scent gland spout; ssp, scutellar spine." httpUri="https://zenodo.org/record/1241799/files/figure.png" pageId="2">Fig. 8</figureCitation>
, corium with 3 broadly spaced light bands across; membrane with narrow dirty white margin and with a dirty white narrow band across near its base (
<figureCitation id="136220B8FF9F0E7ADE3BFB75FAA7FB36" box="[1289,1359,1236,1261]" captionStart="FIGURES 78" captionStartId="6.[151,264,1964,1987]" captionTargetBox="[151,1435,212,1922]" captionTargetId="figure@6.[151,1436,212,1923]" captionTargetPageId="6" captionText="FIGURES 78. Paraindopamphantus bruneiensis, holotype female: 7, meso- and metathoracic area, lateral view; 8, posterior part of thorax and abdomen, dorsal view.sgsp, scent gland spout; ssp, scutellar spine." httpUri="https://zenodo.org/record/1241799/files/figure.png" pageId="2">Fig. 8</figureCitation>
). Legs with coxae fuscous, femora mostly fuscous, basal area of mid and hind femora pale, tibiae fuscous, tarsi pale.
</paragraph>
<paragraph id="8BE63C3DFF9F0E7ADBF5FABDFB79FA7B" blockId="2.[151,1437,1017,2025]" pageId="2">Body and appendages clothed with numerous elongate, upright, golden setae. Head above and pronotum covered with yellow setae. Head ventrally with a few sparse setae on either side of swollen gular areas. Antennal segments covered with short dense setae. Legs, particularly femora and tibiae with long yellow shiny setae, tibia with shorter setae; thorax ventrally without setae, but abdomen ventrally with bristly setae.</paragraph>
<paragraph id="8BE63C3DFF9F0E7ADBF5FA0CFDDDFA1E" blockId="2.[151,1437,1017,2025]" box="[199,565,1452,1477]" pageId="2">
<emphasis id="B92DE02FFF9F0E7ADBF5FA0CFE81FA1F" box="[199,361,1453,1476]" italics="true" pageId="2">Measurements</emphasis>
:
<typeStatus id="54E2829FFF9F0E7ADA4AFA0DFE32FA1E" box="[376,474,1452,1477]" pageId="2" type="holotype">holotype</typeStatus>
female.
</paragraph>
<paragraph id="8BE63C3DFF9F0E7ADBF5FA71FD28F9D6" blockId="2.[151,1437,1017,2025]" pageId="2">
<emphasis id="B92DE02FFF9F0E7ADBF5FA71FEE9FA32" box="[199,257,1488,1513]" italics="true" pageId="2">Body</emphasis>
: Very elongate, linear (
<figureCitation id="136220B8FF9F0E7AD93AFA71FD63FA33" box="[520,651,1488,1513]" captionStart="FIGURES 12" captionStartId="3.[151,264,1975,1998]" captionTargetBox="[151,1436,360,1921]" captionTargetId="figure@3.[151,1436,360,1922]" captionTargetPageId="3" captionText="FIGURES 12. Paraindopamphantus bruneiensis, holotype female:1, habitus, dorsal view; 2, habitus, lateral view." httpUri="https://zenodo.org/record/1241793/files/figure.png" pageId="2">Figs. 1 &amp; 2</figureCitation>
), length 7.90, maximum width across abdomen 1.38, minimum width across wings on second abdominal segment 0.69.
</paragraph>
<paragraph id="8BE63C3DFF9F0E7ADBF5F9B9FEBFF8EE" blockId="2.[151,1437,1017,2025]" pageId="2">
<emphasis id="B92DE02FFF9F0E7ADBF5F9B9FEECF9EA" box="[199,260,1560,1585]" italics="true" pageId="2">Head</emphasis>
: Above smooth, without punctures but with one pair of faintly depressed short lines in anterior half well above juga (
<figureCitation id="136220B8FF9F0E7ADA11F99DFE8FF98E" box="[291,359,1596,1621]" captionStart="FIGURES 34" captionStartId="4.[151,264,1965,1988]" captionTargetBox="[371,1220,193,1946]" captionTargetId="figure@4.[371,1220,190,1946]" captionTargetPageId="4" captionText="FIGURES 34. Paraindopamphantus bruneiensis, holotype female: 3, head and pronotum, dorsal view; 4, head and part of pronotum, lateral view. gs, gular swelling." httpUri="https://zenodo.org/record/1241795/files/figure.png" pageId="2">Fig. 3</figureCitation>
), with tylus rounded and as long as juga, eyes large, occupying most of lateral surface of head but ending well before rounded base of head; length head 0.96; width across eyes 1.44; interocular space 0.87; interocellar space 0.64; eye-ocellar space 0.05; eye length 0.69; eye width 0.29. Labium slightly exceeding fore coxae, 2nd segment reaching base of head; length of segments: I 0.50; II 0.52; III 0.34; IV 0.43. Length of labrum 0.50. Antennae with 1
<taxonomicName id="4C5947BEFF9F0E7ADAA6F96FFDECF931" box="[404,516,1742,1770]" pageId="2" rank="subSpecies" subSpecies="segment">st segment</taxonomicName>
very short, slightly exceeding apex of tylus, 3rd and 4th much thicker than basal two segments, 3rd segment club shaped and densely covered with erect bristles; length of segments: I 0.23; II 0.78; III 0.66; IV 0.73.
</paragraph>
<paragraph id="8BE63C3DFF9F0E7BDBF5F8E1FBB4FEC6" blockId="2.[151,1437,1017,2025]" lastBlockId="3.[151,1437,151,285]" lastPageId="3" pageId="2">
<emphasis id="B92DE02FFF9F0E7ADBF5F8E1FEF0F882" box="[199,280,1856,1881]" italics="true" pageId="2">Thorax</emphasis>
: Pronotum (
<figureCitation id="136220B8FF9F0E7ADA9DF8E1FE1FF882" box="[431,503,1856,1881]" captionStart="FIGURES 34" captionStartId="4.[151,264,1965,1988]" captionTargetBox="[371,1220,193,1946]" captionTargetId="figure@4.[371,1220,190,1946]" captionTargetPageId="4" captionText="FIGURES 34. Paraindopamphantus bruneiensis, holotype female: 3, head and pronotum, dorsal view; 4, head and part of pronotum, lateral view. gs, gular swelling." httpUri="https://zenodo.org/record/1241795/files/figure.png" pageId="2">Fig. 3</figureCitation>
) with posterior margin evenly and shallowly concave in middle, anterior lobe smooth, shiny, except broad anterior collar that is dull and non-shiny, posterior lobe smooth, without punctures; median length 2.07 (anterior lobe 0.80, median stalk 0.40, posterior lobe 0.87); width posterior margin 1.17; width anterior margin 0.87. Thoracic scent gland evaporative area as in
<figureCitation id="136220B8FF9F0E7AD89BF80DFC1BF81F" box="[937,1011,1964,1989]" captionStart="FIGURES 78" captionStartId="6.[151,264,1964,1987]" captionTargetBox="[151,1435,212,1922]" captionTargetId="figure@6.[151,1436,212,1923]" captionTargetPageId="6" captionText="FIGURES 78. Paraindopamphantus bruneiensis, holotype female: 7, meso- and metathoracic area, lateral view; 8, posterior part of thorax and abdomen, dorsal view.sgsp, scent gland spout; ssp, scutellar spine." httpUri="https://zenodo.org/record/1241799/files/figure.png" pageId="2">Fig. 7</figureCitation>
. Scutellum length 0.59; width 0.50. Hemelytra generally narrowed with lateral margins concave and highly constricted behind level of claval commissure (
<figureCitation id="136220B8FF9E0E7BDA01FF36FE93FF6B" box="[307,379,151,176]" captionStart="FIGURES 78" captionStartId="6.[151,264,1964,1987]" captionTargetBox="[151,1435,212,1922]" captionTargetId="figure@6.[151,1436,212,1923]" captionTargetPageId="6" captionText="FIGURES 78. Paraindopamphantus bruneiensis, holotype female: 7, meso- and metathoracic area, lateral view; 8, posterior part of thorax and abdomen, dorsal view.sgsp, scent gland spout; ssp, scutellar spine." httpUri="https://zenodo.org/record/1241799/files/figure.png" pageId="3">Fig. 8</figureCitation>
) and behind at about base of abdomen and second abdominal segment (
<figureCitation id="136220B8FF9E0E7BDFD5FF36FAD8FF6B" box="[1255,1328,151,176]" captionStart="FIGURES 78" captionStartId="6.[151,264,1964,1987]" captionTargetBox="[151,1435,212,1922]" captionTargetId="figure@6.[151,1436,212,1923]" captionTargetPageId="6" captionText="FIGURES 78. Paraindopamphantus bruneiensis, holotype female: 7, meso- and metathoracic area, lateral view; 8, posterior part of thorax and abdomen, dorsal view.sgsp, scent gland spout; ssp, scutellar spine." httpUri="https://zenodo.org/record/1241799/files/figure.png" pageId="3">Fig. 8</figureCitation>
); corium elongately tapering to reach abdominal tergum 3, apical corial margin almost straight, with costal margin gradually slightly dilated, widest at about mid length; length of hemelytra 4.60; length of corium 2.30; claval commissure 0.46; membrane shiny, with 3 faint longitudinal veins (
<figureCitation id="136220B8FF9E0E7BD9CCFEA5FCA8FEC7" box="[766,832,260,285]" captionStart="FIGURES 78" captionStartId="6.[151,264,1964,1987]" captionTargetBox="[151,1435,212,1922]" captionTargetId="figure@6.[151,1436,212,1923]" captionTargetPageId="6" captionText="FIGURES 78. Paraindopamphantus bruneiensis, holotype female: 7, meso- and metathoracic area, lateral view; 8, posterior part of thorax and abdomen, dorsal view.sgsp, scent gland spout; ssp, scutellar spine." httpUri="https://zenodo.org/record/1241799/files/figure.png" pageId="3">Fig. 8</figureCitation>
); width membrane 1.15.
</paragraph>
<caption id="DF266CB5FF9E0E7BDBA5F816FAD9F816" box="[151,1329,1975,1998]" httpUri="https://zenodo.org/record/1241793/files/figure.png" pageId="3" startId="3.[151,264,1975,1998]" targetBox="[151,1436,360,1921]" targetPageId="3">
<paragraph id="8BE63C3DFF9E0E7BDBA5F816FAD9F816" blockId="3.[151,1329,1975,1998]" box="[151,1329,1975,1998]" pageId="3">
<emphasis id="B92DE02FFF9E0E7BDBA5F816FED2F817" bold="true" box="[151,314,1975,1998]" pageId="3">FIGURES 12.</emphasis>
<emphasis id="B92DE02FFF9E0E7BDA72F816FD75F816" box="[320,669,1975,1997]" italics="true" pageId="3">
<taxonomicName id="4C5947BEFF9E0E7BDA72F816FD71F816" authorityName="Malipatil &amp; Scudder" authorityYear="2018" box="[320,665,1975,1997]" class="Insecta" family="Geocoridae" genus="Paraindopamphantus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" phylum="Arthropoda" rank="species" species="bruneiensis">Paraindopamphantus bruneiensis</taxonomicName>
,
</emphasis>
holotype female: 1, habitus, dorsal view; 2, habitus, lateral view.
</paragraph>
</caption>
<caption id="DF266CB5FF990E7CDBA5F80CFDAAF838" httpUri="https://zenodo.org/record/1241795/files/figure.png" pageId="4" startId="4.[151,264,1965,1988]" targetBox="[371,1220,193,1946]" targetPageId="4">
<paragraph id="8BE63C3DFF990E7CDBA5F80CFDAAF838" blockId="4.[151,1436,1965,2019]" pageId="4">
<emphasis id="B92DE02FFF990E7CDBA5F80CFED4F818" bold="true" box="[151,316,1965,1988]" pageId="4">FIGURES 34.</emphasis>
<emphasis id="B92DE02FFF990E7CDA76F80FFD4AF81F" box="[324,674,1966,1988]" italics="true" pageId="4">
<taxonomicName id="4C5947BEFF990E7CDA76F80FFD76F81F" authorityName="Malipatil &amp; Scudder" authorityYear="2018" box="[324,670,1966,1988]" class="Insecta" family="Geocoridae" genus="Paraindopamphantus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" phylum="Arthropoda" rank="species" species="bruneiensis">Paraindopamphantus bruneiensis</taxonomicName>
,
</emphasis>
holotype female: 3, head and pronotum, dorsal view; 4, head and part of pronotum, lateral view. gs, gular swelling.
</paragraph>
</caption>
<caption id="DF266CB5FF980E7DDBA5F80CFEC6F838" httpUri="https://zenodo.org/record/1241797/files/figure.png" pageId="5" startId="5.[151,264,1965,1988]" targetBox="[367,1217,184,1940]" targetPageId="5">
<paragraph id="8BE63C3DFF980E7DDBA5F80CFEC6F838" blockId="5.[151,1435,1965,2019]" pageId="5">
<emphasis id="B92DE02FFF980E7DDBA5F80CFED5F818" bold="true" box="[151,317,1965,1988]" pageId="5">FIGURES 56.</emphasis>
<emphasis id="B92DE02FFF980E7DDA74F80FFD4EF81F" box="[326,678,1966,1988]" italics="true" pageId="5">
<taxonomicName id="4C5947BEFF980E7DDA74F80FFD49F81F" authorityName="Malipatil &amp; Scudder" authorityYear="2018" box="[326,673,1966,1988]" class="Insecta" family="Geocoridae" genus="Paraindopamphantus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" phylum="Arthropoda" rank="species" species="bruneiensis">Paraindopamphantus bruneiensis</taxonomicName>
,
</emphasis>
holotype female: 5, head, ventral view; 6, gular swelling, enlarged. gs, gular swelling.
</paragraph>
</caption>
<caption id="DF266CB5FF9B0E7EDBA5F80DFC10F83A" httpUri="https://zenodo.org/record/1241799/files/figure.png" pageId="6" startId="6.[151,264,1964,1987]" targetBox="[151,1435,212,1922]" targetPageId="6">
<paragraph id="8BE63C3DFF9B0E7EDBA5F80DFC10F83A" blockId="6.[151,1436,1964,2017]" pageId="6">
<emphasis id="B92DE02FFF9B0E7EDBA5F80DFED3F819" bold="true" box="[151,315,1964,1987]" pageId="6">FIGURES 78.</emphasis>
<emphasis id="B92DE02FFF9B0E7EDA73F80DFD77F819" box="[321,671,1964,1986]" italics="true" pageId="6">
<taxonomicName id="4C5947BEFF9B0E7EDA73F80DFD73F819" authorityName="Malipatil &amp; Scudder" authorityYear="2018" box="[321,667,1964,1986]" class="Insecta" family="Geocoridae" genus="Paraindopamphantus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" phylum="Arthropoda" rank="species" species="bruneiensis">Paraindopamphantus bruneiensis</taxonomicName>
,
</emphasis>
holotype female: 7, meso- and metathoracic area, lateral view; 8, posterior part of thorax and abdomen, dorsal view. sgsp, scent gland spout; ssp, scutellar spine.
</paragraph>
</caption>
<caption id="DF266CB5FF9A0E7FDBA5F8A8FBABF8E5" httpUri="https://zenodo.org/record/1241801/files/figure.png" pageId="7" startId="7.[151,264,1801,1824]" targetBox="[151,1436,190,1780]" targetPageId="7">
<paragraph id="8BE63C3DFF9A0E7FDBA5F8A8FBABF8E5" blockId="7.[151,1435,1801,1854]" pageId="7">
<emphasis id="B92DE02FFF9A0E7FDBA5F8A8FEA5F8C5" bold="true" box="[151,333,1801,1824]" pageId="7">FIGURES 910.</emphasis>
<emphasis id="B92DE02FFF9A0E7FDA6BF8A8FD53F8C4" box="[345,699,1801,1823]" italics="true" pageId="7">
<taxonomicName id="4C5947BEFF9A0E7FDA6BF8A8FD5FF8C4" authorityName="Malipatil &amp; Scudder" authorityYear="2018" box="[345,695,1801,1823]" class="Insecta" family="Geocoridae" genus="Paraindopamphantus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" phylum="Arthropoda" rank="species" species="bruneiensis">Paraindopamphantus bruneiensis</taxonomicName>
,
</emphasis>
holotype female: 9, part of thorax and abdomen, ventral view; 10, abdominal dorsum. sgs, scent gland scar; sgsp, scent gland spout; IIVII, abdominal tergites.
</paragraph>
</caption>
<paragraph id="8BE63C3DFF9A0E7FDBF5F8C2FF2BF804" blockId="7.[151,1437,1891,2015]" pageId="7">
<emphasis id="B92DE02FFF9A0E7FDBF5F8C2FED9F8A7" box="[199,305,1891,1916]" italics="true" pageId="7">Abdomen</emphasis>
: Gradually widened from anterior margin of segment III to V, then gradually narrowed posteriorly (
<figureCitation id="136220B8FF9A0E7FDBADF825FF1AF846" box="[159,242,1924,1949]" captionStart="FIGURES 910" captionStartId="7.[151,264,1801,1824]" captionTargetBox="[151,1436,190,1780]" captionTargetId="figure@7.[151,1436,190,1780]" captionTargetPageId="7" captionText="FIGURES 910. Paraindopamphantus bruneiensis, holotype female: 9, part of thorax and abdomen, ventral view; 10, abdominal dorsum. sgs, scent gland scar; sgsp, scent gland spout; IIVII, abdominal tergites." httpUri="https://zenodo.org/record/1241801/files/figure.png" pageId="7">Fig. 10</figureCitation>
). Tergum uniformly sclerotized (
<figureCitation id="136220B8FF9A0E7FD951F825FD5EF846" box="[611,694,1924,1949]" captionStart="FIGURES 910" captionStartId="7.[151,264,1801,1824]" captionTargetBox="[151,1436,190,1780]" captionTargetId="figure@7.[151,1436,190,1780]" captionTargetPageId="7" captionText="FIGURES 910. Paraindopamphantus bruneiensis, holotype female: 9, part of thorax and abdomen, ventral view; 10, abdominal dorsum. sgs, scent gland scar; sgsp, scent gland spout; IIVII, abdominal tergites." httpUri="https://zenodo.org/record/1241801/files/figure.png" pageId="7">Fig. 10</figureCitation>
); abdominal sternum V broad and complete, sternum VI in female only slightly narrowed mesally by ovipositor but complete anteriorly (
<figureCitation id="136220B8FF9A0E7FD8F4F804FBB9F865" box="[966,1105,1957,1982]" captionStart-0="FIGURES 1112" captionStart-1="FIGURES 1314" captionStartId-0="8.[151,264,1573,1596]" captionStartId-1="9.[151,264,1668,1691]" captionTargetBox-0="[151,1436,298,1552]" captionTargetBox-1="[151,1436,304,1648]" captionTargetId-0="figure@8.[151,1436,298,1552]" captionTargetId-1="figure@9.[151,1436,304,1648]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 1112. Paraindopamphantus bruneiensis, holotype female: 11, abdominal terga; 12, abdomen, lateral view. sgs, scent gland scar; sp, spiracle; IIVII, abdominal tergites." captionText-1="FIGURES 1314. Paraindopamphantus bruneiensis, holotype female, abdominal venter. tr, trichobothria; IIIV, abdominal sternites." httpUri-0="https://zenodo.org/record/1241803/files/figure.png" httpUri-1="https://zenodo.org/record/1241805/files/figure.png" pageId="7">Figs. 1214</figureCitation>
), uniformly sclerotized (
<figureCitation id="136220B8FF9A0E7FDE5DF804FF5EF804" captionStart="FIGURES 1314" captionStartId="9.[151,264,1668,1691]" captionTargetBox="[151,1436,304,1648]" captionTargetId="figure@9.[151,1436,304,1648]" captionTargetPageId="9" captionText="FIGURES 1314. Paraindopamphantus bruneiensis, holotype female, abdominal venter. tr, trichobothria; IIIV, abdominal sternites." httpUri="https://zenodo.org/record/1241805/files/figure.png" pageId="7">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="8BE63C3DFF950E70DBF5FF36FDAFFF23" blockId="8.[151,1436,151,248]" pageId="8">
<emphasis id="B92DE02FFF950E70DBF5FF36FE6FFF6B" box="[199,391,151,176]" italics="true" pageId="8">Female genitalia</emphasis>
: As in generic description (
<figureCitation id="136220B8FF950E70D9F4FF36FCBAFF6B" box="[710,850,151,176]" captionStart="FIGURES 1517" captionStartId="10.[151,264,1935,1958]" captionTargetBox="[186,1406,190,1918]" captionTargetId="figure@10.[186,1407,190,1918]" captionTargetPageId="10" captionText="FIGURES 1517. Paraindopamphantus bruneiensis, holotype female: 15, spermatheca; 16,17, ovipositor. Abbreviations: ga, gonangulum; 1gpo, 2gpo, first and second gonapophysis; 1gx, 2gx, first and second gonocoxa; 1r, 2r, first and second ramus; sb, spermathecal bulb; spd, spermathecal duct." httpUri="https://zenodo.org/record/1241807/files/figure.png" pageId="8">Figs. 1517</figureCitation>
). Spermatheca with apical bulb heavily sclerotized, spherical, lacking flanges, duct near base of bulb moderately long, saccoid, then narrowing to a 2-3 turned tube, uniformly lightly sclerotized (
<figureCitation id="136220B8FF950E70DAD4FF7EFDD2FF23" box="[486,570,223,248]" captionStart="FIGURES 1517" captionStartId="10.[151,264,1935,1958]" captionTargetBox="[186,1406,190,1918]" captionTargetId="figure@10.[186,1407,190,1918]" captionTargetPageId="10" captionText="FIGURES 1517. Paraindopamphantus bruneiensis, holotype female: 15, spermatheca; 16,17, ovipositor. Abbreviations: ga, gonangulum; 1gpo, 2gpo, first and second gonapophysis; 1gx, 2gx, first and second gonocoxa; 1r, 2r, first and second ramus; sb, spermathecal bulb; spd, spermathecal duct." httpUri="https://zenodo.org/record/1241807/files/figure.png" pageId="8">Fig. 15</figureCitation>
).
</paragraph>
<caption id="DF266CB5FF950E70DBA5F984FD3DF981" httpUri="https://zenodo.org/record/1241803/files/figure.png" pageId="8" startId="8.[151,264,1573,1596]" targetBox="[151,1436,298,1552]" targetPageId="8">
<paragraph id="8BE63C3DFF950E70DBA5F984FD3DF981" blockId="8.[151,1435,1573,1626]" pageId="8">
<emphasis id="B92DE02FFF950E70DBA5F984FEBDF9E0" bold="true" box="[151,341,1573,1596]" pageId="8">FIGURES 1112.</emphasis>
<emphasis id="B92DE02FFF950E70DA6FF984FD55F9E0" box="[349,701,1573,1595]" italics="true" pageId="8">
<taxonomicName id="4C5947BEFF950E70DA6FF984FD51F9E0" authorityName="Malipatil &amp; Scudder" authorityYear="2018" box="[349,697,1573,1595]" class="Insecta" family="Geocoridae" genus="Paraindopamphantus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" phylum="Arthropoda" rank="species" species="bruneiensis">Paraindopamphantus bruneiensis</taxonomicName>
,
</emphasis>
holotype female: 11, abdominal terga; 12, abdomen, lateral view. sgs, scent gland scar; sp, spiracle; IIVII, abdominal tergites.
</paragraph>
</caption>
<paragraph id="8BE63C3DFF950E70DBF5F926FE2DF944" blockId="8.[151,1437,1671,2020]" box="[199,453,1671,1696]" pageId="8">
<emphasis id="B92DE02FFF950E70DBF5F926FEC8F97B" bold="true" box="[199,288,1671,1696]" pageId="8">Nymph</emphasis>
(
<figureCitation id="136220B8FF950E70DA1DF926FE50F944" box="[303,440,1671,1696]" captionStart-0="FIGURES 1819" captionStart-1="FIGURES 2021" captionStartId-0="11.[151,264,1446,1469]" captionStartId-1="12.[151,264,1991,2014]" captionTargetBox-0="[191,1396,180,1435]" captionTargetBox-1="[167,1425,190,1974]" captionTargetId-0="figure@11.[191,1396,180,1436]" captionTargetId-1="figure@12.[167,1425,190,1975]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURES 1819. Paraindopamphantus bruneiensis, fifth instar nymph: 18, dorsal view; 19, lateral view; ms, mesonotal swelling; psp, pronotal spines; tas, thoracic-abdominal swelling; wp, wing pad." captionText-1="FIGURES 2021. Paraindopamphantus bruneiensis,fifth instar nymph:20, head, dorsal view; 21, right antenna." httpUri-0="https://zenodo.org/record/1241809/files/figure.png" httpUri-1="https://zenodo.org/record/1241811/files/figure.png" pageId="8">Figs. 1821</figureCitation>
).
</paragraph>
<paragraph id="8BE63C3DFF950E70DBF5F90AFAA0F93C" blockId="8.[151,1437,1671,2020]" pageId="8">
<materialsCitation id="3B313660FF950E70DBF5F90AFAA0F93C" ID-GBIF-Occurrence="1841139506" collectingDate="1982-08-20" collectingDateMax="1982-09-10" collectingDateMin="1982-08-20" collectionCode="NHM" collectorName="Lamunin, N. E &amp; Stork, BM" country="Brunei" location="Specimen" pageId="8" specimenCount="1">
<emphasis id="B92DE02FFF950E70DBF5F90AFE55F91F" bold="true" box="[199,445,1707,1732]" pageId="8">
<location id="8E866AE6FF950E70DBF5F90AFED2F91F" LSID="urn:lsid:plazi:treatment:03F08D2BFF9F0E73DBA5FC2FFA74F80A:8E866AE6FF950E70DBF5F90AFED2F91F" box="[199,314,1707,1732]" country="Brunei" name="Specimen" pageId="8">Specimen</location>
examined:
</emphasis>
One fifth instar nymph,
<collectingCountry id="F34E7CADFF950E70D9E0F90DFCD5F918" box="[722,829,1708,1731]" name="Brunei" pageId="8">BRUNEI</collectingCountry>
, Bukit Sulang, nr
<collectorName id="26AC59EBFF950E70DF35F90AFB42F918" box="[1031,1194,1707,1732]" pageId="8">Lamunin, N.E</collectorName>
.
<collectorName id="26AC59EBFF950E70DF86F90AFAC2F918" box="[1204,1322,1707,1732]" pageId="8">Stork, BM</collectorName>
1982-388 fogging
<date id="FFE71AFDFF950E70DBC7F96EFE2EF933" box="[245,454,1743,1768]" pageId="8" value="1982-08-20" valueMax="1982-09-10" valueMin="1982-08-20">
<collectingDate id="EFA3E315FF950E70DBC7F96EFE2EF933" box="[245,454,1743,1768]" pageId="8" value="1982-08-20" valueMax="1982-09-10" valueMin="1982-08-20">20.viii10.ix.1982</collectingDate>
</date>
; insecticide fogging Tree 16. Dipterocarp.
<taxonomicName id="4C5947BEFF950E70D884F96EFB3AF93C" authority="Foxw." authorityName="Foxw." box="[950,1234,1743,1768]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Shorea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="8" phylum="Tracheophyta" rank="species" species="johorensis">
<emphasis id="B92DE02FFF950E70D884F96EFB97F933" box="[950,1151,1743,1768]" italics="true" pageId="8">Shorea johorensis</emphasis>
Foxw.
</taxonomicName>
, in
<collectionCode id="ED48A4F8FF950E70DFCDF971FAACF93C" box="[1279,1348,1744,1767]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="8">NHM</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE63C3DFF950E70DBF5F952FC37F8F4" blockId="8.[151,1437,1671,2020]" pageId="8">
<emphasis id="B92DE02FFF950E70DBF5F952FEB1F8D7" bold="true" box="[199,345,1779,1804]" pageId="8">Description.</emphasis>
Body ground colour reddish to straw coloured, sub shiny as in
<figureCitation id="136220B8FF950E70DF2AF952FB5CF8D0" box="[1048,1204,1779,1804]" captionStart="FIGURES 1819" captionStartId="11.[151,264,1446,1469]" captionTargetBox="[191,1396,180,1435]" captionTargetId="figure@11.[191,1396,180,1436]" captionTargetPageId="11" captionText="FIGURES 1819. Paraindopamphantus bruneiensis, fifth instar nymph: 18, dorsal view; 19, lateral view; ms, mesonotal swelling; psp, pronotal spines; tas, thoracic-abdominal swelling; wp, wing pad." httpUri="https://zenodo.org/record/1241809/files/figure.png" pageId="8">Figs. 18 &amp; 19</figureCitation>
. Head and pronotum with sparse setae, antennae and legs with denser and longer setae (
<figureCitation id="136220B8FF950E70D84DF8B6FC3AF8F4" box="[895,978,1815,1840]" captionStart="FIGURES 1819" captionStartId="11.[151,264,1446,1469]" captionTargetBox="[191,1396,180,1435]" captionTargetId="figure@11.[191,1396,180,1436]" captionTargetPageId="11" captionText="FIGURES 1819. Paraindopamphantus bruneiensis, fifth instar nymph: 18, dorsal view; 19, lateral view; ms, mesonotal swelling; psp, pronotal spines; tas, thoracic-abdominal swelling; wp, wing pad." httpUri="https://zenodo.org/record/1241809/files/figure.png" pageId="8">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="8BE63C3DFF950E70DBF5F89AFF27F8AC" blockId="8.[151,1437,1671,2020]" pageId="8">
<emphasis id="B92DE02FFF950E70DBF5F89AFEE9F88F" box="[199,257,1851,1876]" italics="true" pageId="8">Body</emphasis>
: Very elongate, strongly myrmecomorphic (
<figureCitation id="136220B8FF950E70D9DDF89AFC63F888" box="[751,907,1851,1876]" captionStart="FIGURES 1819" captionStartId="11.[151,264,1446,1469]" captionTargetBox="[191,1396,180,1435]" captionTargetId="figure@11.[191,1396,180,1436]" captionTargetPageId="11" captionText="FIGURES 1819. Paraindopamphantus bruneiensis, fifth instar nymph: 18, dorsal view; 19, lateral view; ms, mesonotal swelling; psp, pronotal spines; tas, thoracic-abdominal swelling; wp, wing pad." httpUri="https://zenodo.org/record/1241809/files/figure.png" pageId="8">Figs. 18 &amp; 19</figureCitation>
), length 5.52, maximum width across abdomen 1.33.
</paragraph>
<paragraph id="8BE63C3DFF950E71DBF5F822FD90FF25" blockId="8.[151,1437,1671,2020]" lastBlockId="9.[151,1436,151,254]" lastPageId="9" pageId="8">
<emphasis id="B92DE02FFF950E70DBF5F822FEECF847" box="[199,260,1923,1948]" italics="true" pageId="8">Head</emphasis>
: Above smooth, without punctures (e.g.,
<figureCitation id="136220B8FF950E70D9EAF822FCC4F840" box="[728,812,1923,1948]" captionStart="FIGURES 2021" captionStartId="12.[151,264,1991,2014]" captionTargetBox="[167,1425,190,1974]" captionTargetId="figure@12.[167,1425,190,1975]" captionTargetPageId="12" captionText="FIGURES 2021. Paraindopamphantus bruneiensis,fifth instar nymph:20, head, dorsal view; 21, right antenna." httpUri="https://zenodo.org/record/1241811/files/figure.png" pageId="8">Fig. 20</figureCitation>
), covered with sparse setae, with eyes large, occupying most of lateral surface of head but ending well before rounded base of head, not nearing pronotum; length head 1.01; width across eyes 1.28; interocular space 0.69; eye length 0.57; eye width 0.27. Labium slightly exceeding fore coxae, all segments slender; length of segments: I 0.27; II 0.39; III 0.27; IV 0.39. Labrum as long as 1
<taxonomicName id="4C5947BEFF940E71DE7EFF36FA74FF68" box="[1356,1436,151,179]" pageId="9" rank="subSpecies" subSpecies="labial">st labial</taxonomicName>
segment, length 0.27. Antennae with 1
<taxonomicName id="4C5947BEFF940E71D963FF1CFD29FF01" box="[593,705,189,218]" pageId="9" rank="subSpecies" subSpecies="segment">st segment</taxonomicName>
very short (
<figureCitation id="136220B8FF940E71D87CFF60FC4AFF01" box="[846,930,193,218]" captionStart="FIGURES 2021" captionStartId="12.[151,264,1991,2014]" captionTargetBox="[167,1425,190,1974]" captionTargetId="figure@12.[167,1425,190,1975]" captionTargetPageId="12" captionText="FIGURES 2021. Paraindopamphantus bruneiensis,fifth instar nymph:20, head, dorsal view; 21, right antenna." httpUri="https://zenodo.org/record/1241811/files/figure.png" pageId="9">Fig. 21</figureCitation>
), slightly exceeding apex of tylus; length of segments: I 0.36; II 0.62; III 0.52; IV 0.62.
</paragraph>
<caption id="DF266CB5FF940E71DBA5F925FF1BF961" httpUri="https://zenodo.org/record/1241805/files/figure.png" pageId="9" startId="9.[151,264,1668,1691]" targetBox="[151,1436,304,1648]" targetPageId="9">
<paragraph id="8BE63C3DFF940E71DBA5F925FF1BF961" blockId="9.[151,1436,1668,1722]" pageId="9">
<emphasis id="B92DE02FFF940E71DBA5F925FEBEF941" bold="true" box="[151,342,1668,1691]" pageId="9">FIGURES 1314.</emphasis>
<emphasis id="B92DE02FFF940E71DA6CF924FD56F940" box="[350,702,1669,1691]" italics="true" pageId="9">
<taxonomicName id="4C5947BEFF940E71DA6CF924FD52F940" authorityName="Malipatil &amp; Scudder" authorityYear="2018" box="[350,698,1669,1691]" class="Insecta" family="Geocoridae" genus="Paraindopamphantus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" phylum="Arthropoda" rank="species" species="bruneiensis">Paraindopamphantus bruneiensis</taxonomicName>
,
</emphasis>
holotype female, abdominal venter. tr, trichobothria; IIIV, abdominal sternites.
</paragraph>
</caption>
<paragraph id="8BE63C3DFF940E71DBF5F947FCC1F854" blockId="9.[151,1437,1766,1935]" pageId="9">
<emphasis id="B92DE02FFF940E71DBF5F947FEF0F924" box="[199,280,1766,1791]" italics="true" pageId="9">Thorax</emphasis>
: Pronotum with narrow lateral carina (
<figureCitation id="136220B8FF940E71D9E0F947FC9AF924" box="[722,882,1766,1791]" captionStart="FIGURES 1819" captionStartId="11.[151,264,1446,1469]" captionTargetBox="[191,1396,180,1435]" captionTargetId="figure@11.[191,1396,180,1436]" captionTargetPageId="11" captionText="FIGURES 1819. Paraindopamphantus bruneiensis, fifth instar nymph: 18, dorsal view; 19, lateral view; ms, mesonotal swelling; psp, pronotal spines; tas, thoracic-abdominal swelling; wp, wing pad." httpUri="https://zenodo.org/record/1241809/files/figure.png" pageId="9">Figs. 18 &amp; 19</figureCitation>
), posterior margin with a pair of large up-curved spine-like processes, anterior lobe broader and larger than posterior lobe, smooth, shiny, collar dull and non-shiny (
<figureCitation id="136220B8FF940E71DBADF88FFF1FF89C" box="[159,247,1838,1863]" captionStart="FIGURES 1819" captionStartId="11.[151,264,1446,1469]" captionTargetBox="[191,1396,180,1435]" captionTargetId="figure@11.[191,1396,180,1436]" captionTargetPageId="11" captionText="FIGURES 1819. Paraindopamphantus bruneiensis, fifth instar nymph: 18, dorsal view; 19, lateral view; ms, mesonotal swelling; psp, pronotal spines; tas, thoracic-abdominal swelling; wp, wing pad." httpUri="https://zenodo.org/record/1241809/files/figure.png" pageId="9">Fig. 19</figureCitation>
); below with a narrow median groove for receipt of labium; median length 1.15 (anterior lobe 0.50; posterior lobe 0.35); width at posterior margin 0.64; width at anterior margin 0.92. Wing pads with bases each with a blunt dorsal swelling (
<figureCitation id="136220B8FF940E71DA97F8D7FE1FF854" box="[421,503,1910,1935]" captionStart="FIGURES 1819" captionStartId="11.[151,264,1446,1469]" captionTargetBox="[191,1396,180,1435]" captionTargetId="figure@11.[191,1396,180,1436]" captionTargetPageId="11" captionText="FIGURES 1819. Paraindopamphantus bruneiensis, fifth instar nymph: 18, dorsal view; 19, lateral view; ms, mesonotal swelling; psp, pronotal spines; tas, thoracic-abdominal swelling; wp, wing pad." httpUri="https://zenodo.org/record/1241809/files/figure.png" pageId="9">Fig. 19</figureCitation>
), length of wing pads 2.66.
</paragraph>
<caption id="DF266CB5FF970E72DBA5F82EFD87F838" httpUri="https://zenodo.org/record/1241807/files/figure.png" pageId="10" startId="10.[151,264,1935,1958]" targetBox="[186,1406,190,1918]" targetPageId="10">
<paragraph id="8BE63C3DFF970E72DBA5F82EFD87F838" blockId="10.[151,1436,1935,2019]" pageId="10">
<emphasis id="B92DE02FFF970E72DBA5F82EFEBCF87E" bold="true" box="[151,340,1935,1958]" pageId="10">FIGURES 1517.</emphasis>
<emphasis id="B92DE02FFF970E72DA69F831FD51F87D" box="[347,697,1936,1958]" italics="true" pageId="10">
<taxonomicName id="4C5947BEFF970E72DA69F831FD5DF87D" authorityName="Malipatil &amp; Scudder" authorityYear="2018" box="[347,693,1936,1958]" class="Insecta" family="Geocoridae" genus="Paraindopamphantus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" phylum="Arthropoda" rank="species" species="bruneiensis">Paraindopamphantus bruneiensis</taxonomicName>
,
</emphasis>
holotype female: 15, spermatheca; 16,17, ovipositor. Abbreviations: ga, gonangulum; 1gpo, 2gpo, first and second gonapophysis; 1gx, 2gx, first and second gonocoxa; 1r, 2r, first and second ramus; sb, spermathecal bulb; spd, spermathecal duct.
</paragraph>
</caption>
<caption id="DF266CB5FF960E73DBA5FA07FC51FA07" httpUri="https://zenodo.org/record/1241809/files/figure.png" pageId="11" startId="11.[151,264,1446,1469]" targetBox="[191,1396,180,1435]" targetPageId="11">
<paragraph id="8BE63C3DFF960E73DBA5FA07FC51FA07" blockId="11.[151,1436,1446,1500]" pageId="11">
<emphasis id="B92DE02FFF960E73DBA5FA07FEBFFA67" bold="true" box="[151,343,1446,1469]" pageId="11">FIGURES 1819.</emphasis>
<emphasis id="B92DE02FFF960E73DA53FA06FD2AFA66" box="[353,706,1447,1469]" italics="true" pageId="11">
<taxonomicName id="4C5947BEFF960E73DA53FA06FD56FA66" authorityName="Malipatil &amp; Scudder" authorityYear="2018" box="[353,702,1447,1469]" class="Insecta" family="Geocoridae" genus="Paraindopamphantus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" phylum="Arthropoda" rank="species" species="bruneiensis">Paraindopamphantus bruneiensis</taxonomicName>
,
</emphasis>
fifth instar nymph: 18, dorsal view; 19, lateral view; ms, mesonotal swelling; psp, pronotal spines; tas, thoracic-abdominal swelling; wp, wing pad.
</paragraph>
</caption>
<paragraph id="8BE63C3DFF960E73DBF5F9A9FB54F96A" blockId="11.[151,1436,1544,2001]" pageId="11">
<emphasis id="B92DE02FFF960E73DBF5F9A9FED9F9FA" box="[199,305,1544,1569]" italics="true" pageId="11">Abdomen</emphasis>
: Gradually widened from anterior margin of segment III to V, then gradually narrowed posteriorly (
<figureCitation id="136220B8FF960E73DBADF98DFED4F99E" box="[159,316,1580,1605]" captionStart="FIGURES 1819" captionStartId="11.[151,264,1446,1469]" captionTargetBox="[191,1396,180,1435]" captionTargetId="figure@11.[191,1396,180,1436]" captionTargetPageId="11" captionText="FIGURES 1819. Paraindopamphantus bruneiensis, fifth instar nymph: 18, dorsal view; 19, lateral view; ms, mesonotal swelling; psp, pronotal spines; tas, thoracic-abdominal swelling; wp, wing pad." httpUri="https://zenodo.org/record/1241809/files/figure.png" pageId="11">Figs. 18 &amp; 19</figureCitation>
); very narrow, tubular and with a distinct swollen node around junction of thorax and abdomen as in
<figureCitation id="136220B8FF960E73DBA5F9F1FF03F9B2" box="[151,235,1616,1641]" captionStart="FIGURES 1819" captionStartId="11.[151,264,1446,1469]" captionTargetBox="[191,1396,180,1435]" captionTargetId="figure@11.[191,1396,180,1436]" captionTargetPageId="11" captionText="FIGURES 1819. Paraindopamphantus bruneiensis, fifth instar nymph: 18, dorsal view; 19, lateral view; ms, mesonotal swelling; psp, pronotal spines; tas, thoracic-abdominal swelling; wp, wing pad." httpUri="https://zenodo.org/record/1241809/files/figure.png" pageId="11">Fig. 19</figureCitation>
. Sutures between all abdominal terga distinct, those between terga III-IV only slightly and those between IV-V and V-VI strongly curved caudad from margin to meson and with distinct scent gland scars which are subequal (the latter fractionally wider) in width, scar between III-IV indistinct (
<figureCitation id="136220B8FF960E73DF23F939FB47F96A" box="[1041,1199,1688,1713]" captionStart="FIGURES 1819" captionStartId="11.[151,264,1446,1469]" captionTargetBox="[191,1396,180,1435]" captionTargetId="figure@11.[191,1396,180,1436]" captionTargetPageId="11" captionText="FIGURES 1819. Paraindopamphantus bruneiensis, fifth instar nymph: 18, dorsal view; 19, lateral view; ms, mesonotal swelling; psp, pronotal spines; tas, thoracic-abdominal swelling; wp, wing pad." httpUri="https://zenodo.org/record/1241809/files/figure.png" pageId="11">Figs. 18 &amp; 19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3436FB6FF960E73DBF5F91CFD21F90E" box="[199,713,1725,1750]" pageId="11" type="distribution">
<paragraph id="8BE63C3DFF960E73DBF5F91CFD21F90E" blockId="11.[151,1436,1544,2001]" box="[199,713,1725,1750]" pageId="11">
<emphasis id="B92DE02FFF960E73DBF5F91CFE88F90D" bold="true" box="[199,352,1725,1750]" pageId="11">Distribution.</emphasis>
<collectingCountry id="F34E7CADFF960E73DA55F91CFE5BF90D" box="[359,435,1725,1750]" name="Brunei" pageId="11">Brunei</collectingCountry>
on the island of Borneo.
</paragraph>
</subSubSection>
<subSubSection id="C3436FB6FF960E73DBF5F941FA74F80A" pageId="11" type="etymology">
<paragraph id="8BE63C3DFF960E73DBF5F941FED7F8C5" blockId="11.[151,1436,1544,2001]" pageId="11">
<emphasis id="B92DE02FFF960E73DBF5F941FEA6F922" bold="true" box="[199,334,1760,1785]" pageId="11">Etymology.</emphasis>
The new species name proposed here,
<emphasis id="B92DE02FFF960E73D837F943FC41F922" box="[773,937,1760,1785]" italics="true" pageId="11">
<taxonomicName id="4C5947BEFF960E73D837F943FC4DF922" authorityName="Malipatil &amp; Scudder" authorityYear="2018" box="[773,933,1760,1785]" class="Insecta" family="Geocoridae" genus="Paraindopamphantus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" phylum="Arthropoda" rank="species" species="bruneiensis">P. bruneiensis</taxonomicName>
,
</emphasis>
alludes to the country where the species has been collected.
</paragraph>
<paragraph id="8BE63C3DFF960E73DBF5F889FA74F80A" blockId="11.[151,1436,1544,2001]" pageId="11">
<emphasis id="B92DE02FFF960E73DBF5F889FEF9F89A" bold="true" box="[199,273,1832,1857]" pageId="11">Notes.</emphasis>
Both the holotype female and the fifth instar nymph were collected from the canopy of
<taxonomicName id="4C5947BEFF960E73DE7FF889FEB2F8BE" authority="Foxw." authorityName="Foxw." class="Magnoliopsida" family="Dipterocarpaceae" genus="Shorea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="11" phylum="Tracheophyta" rank="species" species="johorensis">
<emphasis id="B92DE02FFF960E73DE7FF889FEE3F8BD" italics="true" pageId="11">Shorea johorensis</emphasis>
Foxw.
</taxonomicName>
trees.
<emphasis id="B92DE02FFF960E73DA98F8ECFD96F8BD" box="[426,638,1869,1894]" italics="true" pageId="11">
<taxonomicName id="4C5947BEFF960E73DA98F8ECFD92F8BD" box="[426,634,1869,1894]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Shorea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="11" phylum="Tracheophyta" rank="species" species="johorensis">Shorea johorensis</taxonomicName>
,
</emphasis>
commonly known “Light Red Meranti”, belongs to the plant family
<taxonomicName id="4C5947BEFF960E73DBA5F8D1FEB7F852" box="[151,351,1904,1929]" class="Magnoliopsida" family="Dipterocarpaceae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Malvales" pageId="11" phylum="Tracheophyta" rank="family">Dipterocarpaceae</taxonomicName>
. It is a large emergent tree growing typically to
<quantity id="4CA191D8FF960E73D8ABF8D1FC3DF853" box="[921,981,1904,1929]" metricMagnitude="1" metricUnit="m" metricValue="6.5" pageId="11" unit="m" value="65.0">65 m</quantity>
tall. It produces a high-grade light red meranti timber and for this reason it is a priority plantation tree in Indonesia. It is found in Indonesia (Kalimantan, Sumatera); Malaysia (Peninsular Malaysia, Sabah, Sarawak), and is a critically endangered species (
<bibRefCitation id="EFC841CCFF960E73DFC8F819FA78F80A" box="[1274,1424,1976,2001]" pageId="11" refId="ref3782" refString="Ashton, P. 1998. Shorea johorensis. The IUCN Red List of Threatened Species 1998: e. T 33119 A 9758343. https: // doi. org / 10.2305 / IUCN. UK. 1998. RLTS. T 33119 A 9758343. en" type="journal article">Ashton 1998</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>