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<document id="5FD1858428E5976FDEDC676E778BC00E" ID-CLB-Dataset="22402" ID-DOI="10.5852/ejt.2015.128" ID-GBIF-Dataset="44b592de-b878-417f-80e1-a651f83183ad" ID-ISSN="2118-9773" ID-Zenodo-Dep="3785153" ID-ZooBank="FBAF2DED-5FD6-4F22-8A3A-E94733A92D73" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588612901601" checkinUser="carolina" docAuthor="Constant, Jérôme" docDate="2015" docId="03EFE724FFA0332F2748950B19DA8BF1" docLanguage="en" docName="ejt-2015-128.pdf.imf" docOrigin="European Journal of Taxonomy 128" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pyrops whiteheadi" docType="treatment" docVersion="6" lastPageNumber="21" masterDocId="FFD69F5CFFAC333B253C915F1A378828" masterDocTitle="Review of the effusus group of the Lanternfly genus Pyrops Spinola, 1839, with one new species and notes on trophobiosis (Hemiptera: Fulgoromorpha: Fulgoridae)" masterLastPageNumber="23" masterPageNumber="1" pageNumber="13" updateTime="1698838393947" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="B8DFABEFB5221F456999BD739132A81F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D7213E1AE0F1737F7A664905D293B9B0">
<mods:title id="586C3CFC299CCEC1C28EF1A7CC77F0D5">Review of the effusus group of the Lanternfly genus Pyrops Spinola, 1839, with one new species and notes on trophobiosis (Hemiptera: Fulgoromorpha: Fulgoridae)</mods:title>
</mods:titleInfo>
<mods:name id="D7F3735D43294C946E6289AD3C8C2A2E" type="personal">
<mods:role id="858F72722E2EE66E3358328B8A17BE1F">
<mods:roleTerm id="22287EE168B6B8322A9DA7EA6AFB3B40">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3F23C015BC9C4F4B1985CE9AB03CD357">Constant, Jérôme</mods:namePart>
</mods:name>
<mods:typeOfResource id="BFF1ABEB511149BBBEABD0775021744A">text</mods:typeOfResource>
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<mods:titleInfo id="21299A84E4D12D245884505F33DD332F">
<mods:title id="B600C7746FEFDCBF7BCAA6D2037E0151">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="108CD9D89877148AE636EE8FF7B4424D">
<mods:date id="BE519FCC0332AAEB97BBEEB622E4A88F">2015</mods:date>
<mods:detail id="9FC312D7942BF9AFC7A815EFBB171C94" type="pubDate">
<mods:number id="5874090BD5C990AA978907AEB524F12C">2015-07-08</mods:number>
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<mods:detail id="673A5C1685707459BD7AF90054D8B0CB" type="issue">
<mods:number id="D88E928B307D2A34A3E51C5521ADFDB2">128</mods:number>
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<mods:classification id="A4F779731CC974A4FA9F3E1AA1769D00">journal article</mods:classification>
<mods:identifier id="A56C113CF022580D2404EDC7479B766A" type="CLB-Dataset">22402</mods:identifier>
<mods:identifier id="698DF957D0260A629FD7BCCD0B7CFE51" type="DOI">10.5852/ejt.2015.128</mods:identifier>
<mods:identifier id="5F52C199E646239947D03815952D6856" type="GBIF-Dataset">44b592de-b878-417f-80e1-a651f83183ad</mods:identifier>
<mods:identifier id="8E41FA9BBF6B45BEACE0BBF5AB1F187B" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="1E70EED2DFADC10AEEF1B33CA74E5B1E" type="Zenodo-Dep">3785153</mods:identifier>
<mods:identifier id="0E0416DBDE58E9B41E9D3961E31DC11C" type="ZooBank">FBAF2DED-5FD6-4F22-8A3A-E94733A92D73</mods:identifier>
</mods:mods>
<treatment id="03EFE724FFA0332F2748950B19DA8BF1" ID-DOI="http://doi.org/10.5281/zenodo.3794753" ID-GBIF-Taxon="163887574" ID-Zenodo-Dep="3794753" LSID="urn:lsid:plazi:treatment:03EFE724FFA0332F2748950B19DA8BF1" httpUri="http://treatment.plazi.org/id/03EFE724FFA0332F2748950B19DA8BF1" lastPageId="20" lastPageNumber="21" pageId="12" pageNumber="13">
<subSubSection id="C35C05B9FFA033372748950B1E3C8C46" box="[628,1035,1107,1134]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BF95632FFA033372748950B1E3C8C46" blockId="12.[628,1035,1107,1170]" box="[628,1035,1107,1134]" pageId="12" pageNumber="13">
<heading id="D0B1E15EFFA033372748950B1E3C8C46" box="[628,1035,1107,1134]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
<taxonomicName id="4C462DB1FFA033372748950B1E3C8C46" ID-CoL="4QVBT" authority="(Distant, 1889)" baseAuthorityName="Distant" baseAuthorityYear="1889" box="[628,1035,1107,1134]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFA033372748950B19658C46" bold="true" box="[628,850,1107,1134]" italics="true" pageId="12" pageNumber="13">Pyrops whiteheadi</emphasis>
(
<bibRefCitation id="EFD72BC3FFA03337265E950B1E348C46" author="Distant" box="[866,1027,1108,1134]" firstAuthor="Distant" pageId="12" pageNumber="13" pagination="419 - 422" refId="ref9724" refString="Distant W. L. 1889. XLIX. Descriptions of new species of Rhynchota collected on or near the Kina Balu Mountain, North Borneo. Annals and Magazine of Natural History (Series 6) 3: 419 - 422." type="journal article" year="1889">Distant, 1889</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35C05B9FFA0333727AA952819DF8CB9" box="[662,1000,1143,1170]" pageId="12" pageNumber="13" type="description">
<paragraph id="8BF95632FFA0333727AA952819DF8CB9" blockId="12.[628,1035,1107,1170]" box="[662,1000,1143,1170]" pageId="12" pageNumber="13">
<figureCitation id="137D4AB7FFA0333727AA952818EA8CBA" box="[662,733,1143,1170]" captionStart="Fig" captionStartId="13.[227,270,1968,1994]" captionTargetBox="[228,1429,803,1928]" captionTargetId="figure@13.[227,1436,801,1928]" captionTargetPageId="13" captionText="Fig. 5. Pyrops whiteheadi ♂, Sabah, Danum Valley (RMNH). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (D–E not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785163" httpUri="https://zenodo.org/record/3785163/files/figure.png" pageId="12" pageNumber="13">Figs 5</figureCitation>
A–E, 6, 9A–C, 12A–E
</paragraph>
</subSubSection>
<subSubSection id="C35C05B9FFA0333725DF95E718E88F37" pageId="12" pageNumber="13" type="reference_group">
<paragraph id="8BF95632FFA0333725DF95E71BD38CDE" blockId="12.[227,1436,1208,1270]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="AB56711CFFA0333725DF95E71BD38CDE" pageId="12" pageNumber="13">
<treatmentCitation id="0AE77023FFA0333725DF95E718A88CFA" author="Distant W. L." box="[227,671,1208,1234]" page="422" pageId="12" pageNumber="13" year="1889">
<taxonomicName id="4C462DB1FFA0333725DF95E718A88CFA" ID-CoL="6JQHY" authority="Distant 1889: 422" authorityName="Distant" authorityPageNumber="422" authorityYear="1889" box="[227,671,1208,1234]" class="Insecta" family="Fulgoridae" genus="Fulgora" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFA0333725DF95E71BFD8CFA" box="[227,458,1208,1234]" italics="true" pageId="12" pageNumber="13">Fulgora whiteheadi</emphasis>
<bibRefCitation id="EFD72BC3FFA0333724F395E718A88CFA" author="Distant" box="[463,671,1208,1234]" firstAuthor="Distant" pageId="12" pageNumber="13" pagination="419 - 422" refId="ref9724" refString="Distant W. L. 1889. XLIX. Descriptions of new species of Rhynchota collected on or near the Kina Balu Mountain, North Borneo. Annals and Magazine of Natural History (Series 6) 3: 419 - 422." type="journal article" year="1889">Distant 1889: 422</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54FDE890FFA03337279295E618DF8CFB" box="[686,744,1209,1235]" pageId="12" pageNumber="13">Type</typeStatus>
in NHM) [described and compared to
<taxonomicName id="4C462DB1FFA03337219A95E61F068CFA" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1840" box="[1190,1329,1208,1234]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="delesserti">
<emphasis id="B9328A20FFA03337219A95E61F068CFA" box="[1190,1329,1208,1234]" italics="true" pageId="12" pageNumber="13">P. delesserti</emphasis>
</taxonomicName>
(Guérin- Méneville, 1840)].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF95632FFA0333725DF94421B698D89" blockId="12.[227,1437,1308,1652]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="AB56711CFFA0333725DF94421B698D89" pageId="12" pageNumber="13">
<taxonomicName id="4C462DB1FFA0333725DF94421BFB8D1F" ID-CoL="6JQHY" authorityName="Distant" authorityYear="1889" box="[227,460,1309,1335]" class="Insecta" family="Fulgoridae" genus="Fulgora" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFA0333725DF94421BFB8D1F" box="[227,460,1309,1335]" italics="true" pageId="12" pageNumber="13">Fulgora whiteheadi</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AE77023FFA0333724D0944218F78D10" author="Distant W. L." box="[492,704,1309,1336]" page="518" pageId="12" pageNumber="13" year="1891">
<bibRefCitation id="EFD72BC3FFA0333724D0944218F78D10" author="Distant" box="[492,704,1309,1336]" firstAuthor="Distant" pageId="12" pageNumber="13" pagination="517 - 519" refId="ref9767" refString="Distant W. L. 1891. XXIII. Descriptions of four new species of the genus Fulgora. Transactions of the Entomological Society of London 1891: 517 - 519." type="journal article" year="1891">Distant 1891: 518</bibRefCitation>
</treatmentCitation>
[compared to
<taxonomicName id="4C462DB1FFA033372652944119EA8D1F" ID-CoL="4QV9M" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[878,989,1309,1335]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFA033372652944119EA8D1F" box="[878,989,1309,1335]" italics="true" pageId="12" pageNumber="13">P. effusus</emphasis>
</taxonomicName>
]. —
<treatmentCitation id="0AE77023FFA03337212094421F3C8D1F" author="Lallemand V." box="[1052,1291,1309,1335]" page="88" pageId="12" pageNumber="13" year="1963">
<bibRefCitation id="EFD72BC3FFA03337212094421F3C8D1F" author="Lallemand" box="[1052,1291,1309,1335]" firstAuthor="Lallemand" pageId="12" pageNumber="13" pagination="1 - 99" refId="ref10154" refString="Lallemand V. 1963. Revision des Fulgoridae (Homoptera). Deuxieme partie. Faunes asiatique et australienne. Memoires de l'Institut royal des Sciences naturelles de Belgique (Series 2) 75: 1 - 99." type="journal article" year="1963">Lallemand 1963: 88</bibRefCitation>
</treatmentCitation>
[considered as senior synonym of
<taxonomicName id="4C462DB1FFA033372731941E184B8D72" ID-CoL="4QV9M" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[525,636,1344,1370]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFA033372731941E184B8D72" box="[525,636,1344,1370]" italics="true" pageId="12" pageNumber="13">P. effusus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C462DB1FFA03337278B941E19BF8D72" authorityName="Baker" authorityYear="1925" box="[695,904,1344,1370]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="viridicastaneus">
<emphasis id="B9328A20FFA03337278B941E19BF8D72" box="[695,904,1344,1370]" italics="true" pageId="12" pageNumber="13">P. viridicastaneus</emphasis>
</taxonomicName>
(erroneous); described, keyed; interpretation of the species mixing
<taxonomicName id="4C462DB1FFA033372733943B18498D55" ID-CoL="4QV9M" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[527,638,1379,1405]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFA033372733943B18498D55" box="[527,638,1379,1405]" italics="true" pageId="12" pageNumber="13">P. effusus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C462DB1FFA033372786943B196E8D55" ID-CoL="6JQHY" authorityName="Distant" authorityYear="1889" box="[698,857,1379,1405]" class="Insecta" family="Fulgoridae" genus="Fulgora" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFA033372786943B196E8D55" box="[698,857,1379,1405]" italics="true" pageId="12" pageNumber="13">P. whiteheadi</emphasis>
</taxonomicName>
]. —
<treatmentCitation id="0AE77023FFA0333726A5943C1E8C8D55" author="Sato M. &amp; Nagai S." box="[921,1211,1379,1405]" page="308" pageId="12" pageNumber="13" year="1994">
<bibRefCitation id="EFD72BC3FFA0333726A5943C1E8C8D55" author="Sato &amp; Nagai" box="[921,1211,1379,1405]" firstAuthor="Sato" pageId="12" pageNumber="13" pagination="307 - 314" refId="ref10496" refString="Sato M. &amp; Nagai S. 1994. Taxonomic notes on the Southeast Asian species of the genus Fulgora (Homoptera: Fulgoridae). Transactions of the Shikoku Entomological Society 20 (3 - 4): 307 - 314." type="journal article" year="1994">Satô &amp; Nagai 1994: 308</bibRefCitation>
</treatmentCitation>
[compared with
<taxonomicName id="4C462DB1FFA0333720BB943B1B7A8D88" ID-CoL="9F54Z" baseAuthorityName="Sato &amp; Nagai" baseAuthorityYear="1994" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="gunjii">
<emphasis id="B9328A20FFA0333720BB943B1B7A8D88" italics="true" pageId="12" pageNumber="13">P. gunjii</emphasis>
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF95632FFA0333725DF94F519438DCF" blockId="12.[227,1437,1308,1652]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="AB56711CFFA0333725DF94F519438DCF" pageId="12" pageNumber="13">
<taxonomicName id="4C462DB1FFA0333725DF94F51BB88DEC" ID-CoL="9XR5P" authorityName="Distant" authorityYear="1891" box="[227,399,1450,1476]" class="Insecta" family="Fulgoridae" genus="Fulgora" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="effusa">
<emphasis id="B9328A20FFA0333725DF94F51BB88DEC" box="[227,399,1450,1476]" italics="true" pageId="12" pageNumber="13">Fulgora effusa</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AE77023FFA03337249694F5185D8DED" author="Baker C. F." box="[426,618,1450,1477]" page="359" pageId="12" pageNumber="13" year="1925">
<bibRefCitation id="EFD72BC3FFA03337249694F5185D8DED" author="Baker" box="[426,618,1450,1477]" firstAuthor="Baker" pageId="12" pageNumber="13" pagination="343 - 364" refId="ref9357" refString="Baker C. F. 1925. Remarks on certain Indo-Malayan Fulgora, with special reference to Philippines species. Philippines Journal of Science 28: 343 - 364." type="journal article" year="1925">Baker 1925: 359</bibRefCitation>
</treatmentCitation>
[comparison between
<taxonomicName id="4C462DB1FFA03337264D94F419EA8DEC" ID-CoL="4QV9M" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[881,989,1450,1476]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFA03337264D94F419EA8DEC" box="[881,989,1450,1476]" italics="true" pageId="12" pageNumber="13">P. effusus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C462DB1FFA03337212894F41E878DEC" ID-CoL="6JQHY" authorityName="Distant" authorityYear="1889" box="[1044,1200,1450,1476]" class="Insecta" family="Fulgoridae" genus="Fulgora" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFA03337212894F41E878DEC" box="[1044,1200,1450,1476]" italics="true" pageId="12" pageNumber="13">P. whiteheadi</emphasis>
</taxonomicName>
based on specimens of
<taxonomicName id="4C462DB1FFA03337241494911BF18DCF" ID-CoL="6JQHY" authorityName="Distant" authorityYear="1889" box="[296,454,1485,1511]" class="Insecta" family="Fulgoridae" genus="Fulgora" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFA03337241494911BF18DCF" box="[296,454,1485,1511]" italics="true" pageId="12" pageNumber="13">P. whiteheadi</emphasis>
</taxonomicName>
erroneously identified as
<taxonomicName id="4C462DB1FFA0333727C9949119548DCF" ID-CoL="4QV9M" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[757,867,1485,1511]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFA0333727C9949119548DCF" box="[757,867,1485,1511]" italics="true" pageId="12" pageNumber="13">P. effusus</emphasis>
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF95632FFA0333725DF94AE1E4E8E23" blockId="12.[227,1437,1308,1652]" box="[227,1145,1520,1547]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="AB56711CFFA0333725DF94AE1E4E8E23" box="[227,1145,1520,1547]" pageId="12" pageNumber="13">
<taxonomicName id="4C462DB1FFA0333725DF94AE1BDD8E22" box="[227,490,1520,1546]" class="Insecta" family="Fulgoridae" genus="Laternaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFA0333725DF94AE1BDD8E22" box="[227,490,1520,1546]" italics="true" pageId="12" pageNumber="13">Laternaria whiteheadi</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AE77023FFA03337273B94AF18928E22" box="[519,677,1520,1546]" pageId="12" pageNumber="13">
<bibRefCitation id="EFD72BC3FFA03337273B94AF18928E22" author="Metcalf" box="[519,677,1520,1546]" firstAuthor="Metcalf" pageId="12" pageNumber="13" refId="ref10291" refString="Metcalf Z. P. 1947. General Catalogue of the Homoptera. Fascicle IV Fulgoroidea. Part 9 Fulgoridae. North Carolina State College, Raleigh (U. S. A.)." type="book" year="1947">Metcalf 1947</bibRefCitation>
</treatmentCitation>
[transferred to
<taxonomicName id="4C462DB1FFA03337266694AE19EE8E22" authorityName="Linnaeus" authorityYear="1764" box="[858,985,1521,1546]" class="Insecta" family="Fulgoridae" genus="Laternaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9328A20FFA03337266694AE19EE8E22" box="[858,985,1521,1546]" italics="true" pageId="12" pageNumber="13">Laternaria</emphasis>
</taxonomicName>
; catalogued].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF95632FFA0333725DF974B18B78E5C" blockId="12.[227,1437,1308,1652]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="AB56711CFFA0333725DF974B18B78E5C" pageId="12" pageNumber="13">
<taxonomicName id="4C462DB1FFA0333725DF974B1BF68E05" ID-CoL="4QVBT" baseAuthorityName="Distant" baseAuthorityYear="1889" box="[227,449,1555,1581]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFA0333725DF974B1BF68E05" box="[227,449,1555,1581]" italics="true" pageId="12" pageNumber="13">Pyrops whiteheadi</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AE77023FFA0333724D9974C192A8E05" author="Nagai S. &amp; Porion T." box="[485,797,1555,1581]" page="26" pageId="12" pageNumber="13" year="1996">
<bibRefCitation id="EFD72BC3FFA0333724D9974C192A8E05" author="Nagai &amp; Porion" box="[485,797,1555,1581]" firstAuthor="Nagai" pageId="12" pageNumber="13" refId="ref10327" refString="Nagai S. &amp; Porion T. 1996. Fulgoridae 2: Catalogue illustre des faunes asiatique et australienne. Sciences Nat, Compiegne." type="book" year="1996">Nagai &amp; Porion 1996: 26</bibRefCitation>
</treatmentCitation>
[transferred to
<taxonomicName id="4C462DB1FFA0333726E2974B1E078E05" ID-CoL="765X" authorityName="Spinola" authorityYear="1839" box="[990,1072,1556,1581]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9328A20FFA0333726E2974B1E078E05" box="[990,1072,1556,1581]" italics="true" pageId="12" pageNumber="13">Pyrops</emphasis>
</taxonomicName>
; listed], pl. 11, fig. 160 [
<typeStatus id="54FDE890FFA03337205B974B1FAB8E06" box="[1383,1436,1556,1582]" pageId="12" pageNumber="13">type</typeStatus>
illustrated]. —
<treatmentCitation id="0AE77023FFA033372480976918588E79" author="Liang A. - P." box="[444,623,1590,1617]" page="45" pageId="12" pageNumber="13" year="1998">
<bibRefCitation id="EFD72BC3FFA033372480976918588E79" author="Liang" box="[444,623,1590,1617]" firstAuthor="Liang" pageId="12" pageNumber="13" pagination="41 - 47" refId="ref10253" refString="Liang A. - P. 1998. Nomenclatorial notes on the Oriental lantern fly genus Pyrops Spinola (Hemiptera: Fulgoroidea: Fulgoridae). Acta Zootaxonomica Sinica 23 (1): 41 - 47." type="journal article" year="1998">Liang 1998: 45</bibRefCitation>
</treatmentCitation>
[listed]. —
<treatmentCitation id="0AE77023FFA0333727C297691E2E8E78" author="Nagai S. &amp; Porion T." box="[766,1049,1590,1617]" page="6" pageId="12" pageNumber="13" year="2004">
<bibRefCitation id="EFD72BC3FFA0333727C297691E2E8E78" author="Nagai &amp; Porion" box="[766,1049,1590,1617]" firstAuthor="Nagai" pageId="12" pageNumber="13" refId="ref10378" refString="Nagai S. &amp; Porion T. 2004. Fulgoridae 2, supplement 2: Nouveaux Fulgoridae d'Asie du Sud-Est. Hillside Books, Canterbury." type="book" year="2004">Nagai &amp; Porion 2004: 6</bibRefCitation>
</treatmentCitation>
[comparison with
<taxonomicName id="4C462DB1FFA0333721CB97681F528E79" ID-CoL="4QV9M" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[1271,1381,1591,1617]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFA0333721CB97681F528E79" box="[1271,1381,1591,1617]" italics="true" pageId="12" pageNumber="13">P. effusus</emphasis>
</taxonomicName>
], pl. 1, fig. 6 [illustration of habitus].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF95632FFA0333725DF97C318E88EF1" blockId="12.[227,1436,1691,1823]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="AB56711CFFA0333725DF97C318E88EF1" pageId="12" pageNumber="13">
non
<taxonomicName id="4C462DB1FFA03337242B97C418368E9D" ID-CoL="6JQHY" authorityName="Distant" authorityYear="1889" box="[279,513,1691,1717]" class="Insecta" family="Fulgoridae" genus="Fulgora" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFA03337242B97C418368E9D" box="[279,513,1691,1717]" italics="true" pageId="12" pageNumber="13">Fulgora whiteheadi</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AE77023FFA03337271C97C418D28E9D" box="[544,741,1691,1717]" pageId="12" pageNumber="13">
<bibRefCitation id="EFD72BC3FFA03337271C97C418D28E9D" author="Lallemand" box="[544,741,1691,1717]" firstAuthor="Lallemand" pageId="12" pageNumber="13" pagination="1 - 99" refId="ref10154" refString="Lallemand V. 1963. Revision des Fulgoridae (Homoptera). Deuxieme partie. Faunes asiatique et australienne. Memoires de l'Institut royal des Sciences naturelles de Belgique (Series 2) 75: 1 - 99." type="journal article" year="1963">Lallemand 1963</bibRefCitation>
</treatmentCitation>
: pl. X, figs 11–13 [illustration of the genitalia of
<taxonomicName id="4C462DB1FFA03337201197C31FAB8E9D" ID-CoL="4QV9M" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[1325,1436,1691,1717]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFA03337201197C31FAB8E9D" box="[1325,1436,1691,1717]" italics="true" pageId="12" pageNumber="13">P. effusus</emphasis>
</taxonomicName>
erroneously identified as
<taxonomicName id="4C462DB1FFA03337270C97E018F98EF0" ID-CoL="6JQHY" authorityName="Distant" authorityYear="1889" box="[560,718,1726,1752]" class="Insecta" family="Fulgoridae" genus="Fulgora" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFA03337270C97E018F98EF0" box="[560,718,1726,1752]" italics="true" pageId="12" pageNumber="13">P. whiteheadi</emphasis>
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF95632FFA0333725DF97BD18E88F37" blockId="12.[227,1436,1691,1823]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="AB56711CFFA0333725DF97BD18E88F37" pageId="12" pageNumber="13">
non
<taxonomicName id="4C462DB1FFA03337242B97BD1BC68ED4" ID-CoL="4QVBT" baseAuthorityName="Distant" baseAuthorityYear="1889" box="[279,497,1762,1788]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFA03337242B97BD1BC68ED4" box="[279,497,1762,1788]" italics="true" pageId="12" pageNumber="13">Pyrops whiteheadi</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AE77023FFA03337273397BE19268ED3" box="[527,785,1761,1788]" pageId="12" pageNumber="13">
<bibRefCitation id="EFD72BC3FFA03337273397BE19268ED3" author="Nagai &amp; Porion" box="[527,785,1761,1788]" firstAuthor="Nagai" pageId="12" pageNumber="13" refId="ref10327" refString="Nagai S. &amp; Porion T. 1996. Fulgoridae 2: Catalogue illustre des faunes asiatique et australienne. Sciences Nat, Compiegne." type="book" year="1996">Nagai &amp; Porion 1996</bibRefCitation>
</treatmentCitation>
: pl. 17, fig. 213 [illustration of a specimen of
<taxonomicName id="4C462DB1FFA03337201197BD1FAB8ED4" ID-CoL="4QV9M" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[1325,1436,1762,1788]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFA03337201197BD1FAB8ED4" box="[1325,1436,1762,1788]" italics="true" pageId="12" pageNumber="13">P. effusus</emphasis>
</taxonomicName>
erroneously identified as
<taxonomicName id="4C462DB1FFA03337270C965918F98F37" ID-CoL="4QVBT" baseAuthorityName="Distant" baseAuthorityYear="1889" box="[560,718,1797,1823]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFA03337270C965918F98F37" box="[560,718,1797,1823]" italics="true" pageId="12" pageNumber="13">P. whiteheadi</emphasis>
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35C05B9FFA0333625DF9619190B8947" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8BF95632FFA0333725DF96191B6B8F48" blockId="12.[227,348,1862,1888]" box="[227,348,1862,1888]" pageId="12" pageNumber="13">
<heading id="D0B1E15EFFA0333725DF96191B6B8F48" bold="true" box="[227,348,1862,1888]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B9328A20FFA0333725DF96191B6B8F48" bold="true" box="[227,348,1862,1888]" pageId="12" pageNumber="13">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BF95632FFA0333725DF962A1FAA8FFD" blockId="12.[227,1437,1909,2006]" pageId="12" pageNumber="13">
In addition to the characters defining the
<taxonomicName id="4C462DB1FFA0333727E2962A19188FA7" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[734,815,1909,1935]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFA0333727E2962A19188FA7" box="[734,815,1909,1935]" italics="true" pageId="12" pageNumber="13">effusus</emphasis>
</taxonomicName>
group, the species can be recognized by: (1) head entirely blue (often faded to greenish in collection specimens) (
<figureCitation id="137D4AB7FFA03337268296C71E378F9B" box="[958,1024,1944,1971]" captionStart="Fig" captionStartId="13.[227,270,1968,1994]" captionTargetBox="[228,1429,803,1928]" captionTargetId="figure@13.[227,1436,801,1928]" captionTargetPageId="13" captionText="Fig. 5. Pyrops whiteheadi ♂, Sabah, Danum Valley (RMNH). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (D–E not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785163" httpUri="https://zenodo.org/record/3785163/files/figure.png" pageId="12" pageNumber="13">Fig. 5</figureCitation>
D–E); (2) pronotum yellow-brown, mesonotum blue-brown (
<figureCitation id="137D4AB7FFA03337273296E418598FFE" box="[526,622,1979,2006]" captionStart="Fig" captionStartId="13.[227,270,1968,1994]" captionTargetBox="[228,1429,803,1928]" captionTargetId="figure@13.[227,1436,801,1928]" captionTargetPageId="13" captionText="Fig. 5. Pyrops whiteheadi ♂, Sabah, Danum Valley (RMNH). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (D–E not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785163" httpUri="https://zenodo.org/record/3785163/files/figure.png" pageId="12" pageNumber="13">Fig. 5A</figureCitation>
); (3) tegmina green with small yellow spots also beyond nodal line;
</paragraph>
<paragraph id="8BF95632FFA1333625DF906D190B8947" blockId="13.[227,1436,305,367]" pageId="13" pageNumber="14">
spots rounded and not coalescent (
<figureCitation id="137D4AB7FFA133362742906E18E98964" box="[638,734,305,332]" captionStart="Fig" captionStartId="13.[227,270,1968,1994]" captionTargetBox="[228,1429,803,1928]" captionTargetId="figure@13.[227,1436,801,1928]" captionTargetPageId="13" captionText="Fig. 5. Pyrops whiteheadi ♂, Sabah, Danum Valley (RMNH). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (D–E not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785163" httpUri="https://zenodo.org/record/3785163/files/figure.png" pageId="13" pageNumber="14">Fig. 5A</figureCitation>
); (4) tegmina without large pale yellow patch at nodal line and with apical margin slightly infuscate (
<figureCitation id="137D4AB7FFA1333627EC900B19188947" box="[720,815,340,367]" captionStart="Fig" captionStartId="13.[227,270,1968,1994]" captionTargetBox="[228,1429,803,1928]" captionTargetId="figure@13.[227,1436,801,1928]" captionTargetPageId="13" captionText="Fig. 5. Pyrops whiteheadi ♂, Sabah, Danum Valley (RMNH). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Head and thorax, lateral view. E. Frons, normal view. Scale bar = 30 mm (D–E not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3785163" httpUri="https://zenodo.org/record/3785163/files/figure.png" pageId="13" pageNumber="14">Fig. 5A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35C05B9FFA1333525DF90C91FB389F0" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BF95632FFA1333625DF90C91BE78998" blockId="13.[227,464,406,432]" box="[227,464,406,432]" pageId="13" pageNumber="14">
<heading id="D0B1E15EFFA1333625DF90C91BE78998" bold="true" box="[227,464,406,432]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B9328A20FFA1333625DF90C91BE78998" bold="true" box="[227,464,406,432]" pageId="13" pageNumber="14">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BF95632FFA13336242C909A1B8889F6" blockId="13.[227,1436,452,584]" box="[272,447,452,479]" pageId="13" pageNumber="14">
<emphasis id="B9328A20FFA13336242C909A1B8889F6" bold="true" box="[272,447,452,479]" pageId="13" pageNumber="14">
<typeStatus id="54FDE890FFA13336242C909A1B7A89F7" box="[272,333,453,479]" pageId="13" pageNumber="14">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="8BF95632FFA1333625DF90B71BC78A60" blockId="13.[227,1436,452,584]" pageId="13" pageNumber="14">
<collectingCountry id="F35116A2FFA1333625DF90B71BB78A2A" box="[227,384,488,514]" name="Malaysia" pageId="13" pageNumber="14">MALAYSIA</collectingCountry>
:
<typeStatus id="54FDE890FFA1333624B790B718328A2A" box="[395,517,488,514]" pageId="13" pageNumber="14" type="lectotype">Lectotype</typeStatus>
:
<specimenCount id="9D409DBBFFA13336272D90B7181E8A2A" box="[529,553,488,514]" pageId="13" pageNumber="14" type="male">♂</specimenCount>
, here designated to provide a reference standard for the species (examined on photographs): East Malaysia, [Kina Balu (Whitehead)] [
<emphasis id="B9328A20FFA13336268493541E098A0D" box="[952,1086,523,549]" italics="true" pageId="13" pageNumber="14">Whiteheadi</emphasis>
Dist.] [Distant Coll. 1911— 383.] [Type] (
<collectionCode id="ED57CEF7FFA1333624B993711BD38A60" box="[389,484,558,584]" pageId="13" pageNumber="14">BMNH</collectionCode>
).
</paragraph>
<paragraph id="8BF95632FFA1333625DF93301E2B8A85" blockId="13.[227,1436,623,685]" pageId="13" pageNumber="14">
<collectingCountry id="F35116A2FFA1333625DF93301BB78AA1" box="[227,384,623,649]" name="Malaysia" pageId="13" pageNumber="14">MALAYSIA</collectingCountry>
:
<typeStatus id="54FDE890FFA1333624B3933018088AA1" box="[399,575,623,649]" pageId="13" pageNumber="14" type="paralectotype">Paralectotypes</typeStatus>
:
<specimenCount id="9D409DBBFFA133362772933018A18AA2" box="[590,662,623,650]" pageId="13" pageNumber="14" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D409DBBFFA13336279B933018D98AA2" box="[679,750,623,650]" pageId="13" pageNumber="14" type="female">2 ♀♀</specimenCount>
(examined on photographs): East Malaysia, [Kina Balu (Whitehead)] [
<emphasis id="B9328A20FFA1333624AC93CC18218A85" box="[400,534,659,685]" italics="true" pageId="13" pageNumber="14">Whiteheadi</emphasis>
Dist.] [Distant Coll. 1911—383.] (
<collectionCode id="ED57CEF7FFA13336268D93CC1E278A85" box="[945,1040,659,685]" pageId="13" pageNumber="14">BMNH</collectionCode>
).
</paragraph>
<caption id="DF3906BAFFA1333625DF96EF18688039" ID-DOI="http://doi.org/10.5281/zenodo.3785163" ID-Zenodo-Dep="3785163" httpUri="https://zenodo.org/record/3785163/files/figure.png" pageId="13" pageNumber="14" startId="13.[227,270,1968,1994]" targetBox="[228,1429,803,1928]" targetPageId="13">
<paragraph id="8BF95632FFA1333625DF96EF1FAA8FC5" blockId="13.[227,1437,1968,2030]" pageId="13" pageNumber="14">
<emphasis id="B9328A20FFA1333625DF96EF1B038FE2" bold="true" box="[227,308,1968,1994]" pageId="13" pageNumber="14">Fig. 5.</emphasis>
<taxonomicName id="4C462DB1FFA13336240396EE18298FE2" baseAuthorityName="Distant" baseAuthorityYear="1889" box="[319,542,1968,1994]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFA13336240396EE18298FE2" box="[319,542,1968,1994]" italics="true" pageId="13" pageNumber="14">Pyrops whiteheadi</emphasis>
</taxonomicName>
♂, Sabah, Danum Valley (RMNH).
<emphasis id="B9328A20FFA1333626DD96EF19C18FE2" bold="true" box="[993,1014,1968,1994]" pageId="13" pageNumber="14">A</emphasis>
. Habitus, dorsal view.
<emphasis id="B9328A20FFA13336202F96EF1F108FE2" bold="true" box="[1299,1319,1968,1994]" pageId="13" pageNumber="14">B</emphasis>
. Habitus, ventral view.
<emphasis id="B9328A20FFA1333624B8968C1BAE8FC5" bold="true" box="[388,409,2003,2029]" pageId="13" pageNumber="14">C</emphasis>
. Habitus, lateral view.
<emphasis id="B9328A20FFA133362793968B18F38FC6" bold="true" box="[687,708,2004,2030]" pageId="13" pageNumber="14">D</emphasis>
. Head and thorax, lateral view.
<emphasis id="B9328A20FFA13336217F968B1E608FC6" bold="true" box="[1091,1111,2004,2030]" pageId="13" pageNumber="14">E</emphasis>
. Frons, normal view. Scale
</paragraph>
<paragraph id="8BF95632FFA1333625DF96A918688039" blockId="13.[227,607,2038,2065]" box="[227,607,2038,2065]" pageId="13" pageNumber="14">
<pageTitle id="CBD98E55FFA1333625DF96A918688039" box="[227,607,2038,2065]" pageId="13" pageNumber="14">bar = 30 mm (D–E not to scale).</pageTitle>
</paragraph>
</caption>
<paragraph id="8BF95632FFA23335242C906F18308962" blockId="14.[227,1437,304,472]" box="[272,519,304,330]" pageId="14" pageNumber="15">
<emphasis id="B9328A20FFA23335242C906F18308962" bold="true" box="[272,519,304,330]" pageId="14" pageNumber="15">Additional material</emphasis>
</paragraph>
<paragraph id="8BF95632FFA2333525DF900B1FB389F0" blockId="14.[227,1437,304,472]" pageId="14" pageNumber="15">
<collectingCountry id="F35116A2FFA2333525DF900B1BB78946" box="[227,384,340,366]" name="Malaysia" pageId="14" pageNumber="15">MALAYSIA</collectingCountry>
:
<specimenCount id="9D409DBBFFA2333524B6900B1B808946" box="[394,439,340,366]" pageId="14" pageNumber="15" type="male">1 ♂</specimenCount>
,
<collectingRegion id="498298D0FFA2333524FF900B18398946" box="[451,526,340,366]" country="Malaysia" name="Sabah" pageId="14" pageNumber="15">Sabah</collectingRegion>
, Kinabalu National Park, Poring Hot Spring,
<date id="FFF870F2FFA233352123900B1E8E8946" box="[1055,1209,340,366]" pageId="14" pageNumber="15" value="1999-04-21">21 Apr. 1999</date>
, T. Trilar (
<collectionCode id="ED57CEF7FFA23335200E900B1FB88946" box="[1330,1423,340,366]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="14" pageNumber="15" type="Museum">RBINS</collectionCode>
);
<specimenCount id="9D409DBBFFA2333525DF90281B2689BA" box="[227,273,375,402]" pageId="14" pageNumber="15" type="female">1 ♀</specimenCount>
,
<collectingRegion id="498298D0FFA23335241E90281B5A89B9" box="[290,365,375,401]" country="Malaysia" name="Sabah" pageId="14" pageNumber="15">Sabah</collectingRegion>
,
<quantity id="4CBEFBD7FFA23335244790281BF089B9" box="[379,455,375,401]" metricMagnitude="4" metricUnit="m" metricValue="2.3" pageId="14" pageNumber="15" unit="km" value="23.0">23 km</quantity>
W of Sandakan, Sepilok tree tower,
<geoCoordinate id="EE7230F5FFA2333526BB902819EA89BA" box="[903,989,375,402]" degrees="5" direction="north" minutes="49" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="5.8166666">5°49’N</geoCoordinate>
<geoCoordinate id="EE7230F5FFA2333526D490281E6D89B9" box="[1000,1114,375,401]" degrees="118" direction="east" minutes="06" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="118.1">118°06’E</geoCoordinate>
,
<date id="FFF870F2FFA23335215490281F3789B9" box="[1128,1280,375,402]" pageId="14" pageNumber="15" value="1987-11-01">1 Nov. 1987</date>
,
<quantity id="4CBEFBD7FFA23335203290281F4D89BA" box="[1294,1402,375,402]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="14" pageNumber="15" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0–100 m</quantity>
, J. Huisman &amp; R. de Jong (
<collectionCode id="ED57CEF7FFA2333524C390C51869899C" box="[511,606,410,436]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="15" type="Museum">RMNH</collectionCode>
);
<specimenCount id="9D409DBBFFA23335274D90C518A8899D" box="[625,671,410,437]" pageId="14" pageNumber="15" type="male">1 ♂</specimenCount>
,
<collectingRegion id="498298D0FFA23335279090C518C0899C" box="[684,759,410,436]" country="Malaysia" name="Sabah" pageId="14" pageNumber="15">Sabah</collectingRegion>
, Danum Valley,
<quantity id="4CBEFBD7FFA23335268B90C51E36899C" box="[951,1025,410,436]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="15" unit="km" value="70.0">70 km</quantity>
W of Lahad Datu, rainforest along narrow stream, at light, next to bridge Nature Trail,
<quantity id="4CBEFBD7FFA23335267F90E219B789F0" box="[835,896,445,472]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="15" unit="m" value="50.0">50 m</quantity>
,
<date id="FFF870F2FFA2333526B190E11E2989F0" box="[909,1054,446,472]" pageId="14" pageNumber="15" value="1989-12-01">1 Dec. 1989</date>
, M.J. &amp; J.P. Duffels (
<collectionCode id="ED57CEF7FFA23335202590E11F4F89F0" box="[1305,1400,446,472]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="15" type="Museum">RMNH</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="DF3906BAFFA2333525DF96A919318039" ID-DOI="http://doi.org/10.5281/zenodo.3785165" ID-Zenodo-Dep="3785165" box="[227,774,2038,2065]" httpUri="https://zenodo.org/record/3785165/files/figure.png" pageId="14" pageNumber="15" startId="14.[227,270,2038,2064]" targetBox="[227,1435,518,1988]" targetPageId="14">
<paragraph id="8BF95632FFA2333525DF96A919318039" blockId="14.[227,774,2038,2065]" box="[227,774,2038,2065]" pageId="14" pageNumber="15">
<emphasis id="B9328A20FFA2333525DF96A91B078038" bold="true" box="[227,304,2038,2064]" pageId="14" pageNumber="15">Fig. 6.</emphasis>
<taxonomicName id="4C462DB1FFA23335240496A81BD58038" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[312,482,2038,2064]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFA23335240496A81BD58038" box="[312,482,2038,2064]" italics="true" pageId="14" pageNumber="15">Pyrops effusus</emphasis>
</taxonomicName>
group, distribution map.
</paragraph>
</caption>
<caption id="DF3906BAFFA3333425DF96EF18C18038" ID-DOI="http://doi.org/10.5281/zenodo.3785167" ID-Zenodo-Dep="3785167" httpUri="https://zenodo.org/record/3785167/files/figure.png" pageId="15" pageNumber="16" startId="15.[227,279,1968,1994]" targetBox="[227,1436,344,1927]" targetPageId="15">
<paragraph id="8BF95632FFA3333425DF96EF1FAA8FC5" blockId="15.[227,1437,1968,2030]" pageId="15" pageNumber="16">
<emphasis id="B9328A20FFA3333425DF96EF1B668FE2" bold="true" box="[227,337,1968,1994]" pageId="15" pageNumber="16">Figs 7–9.</emphasis>
<taxonomicName id="4C462DB1FFA33334246A96EE1BC88FE2" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[342,511,1968,1994]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFA33334246A96EE1BC88FE2" box="[342,511,1968,1994]" italics="true" pageId="15" pageNumber="16">Pyrops effusus</emphasis>
</taxonomicName>
group, ♂ genitalia.
<emphasis id="B9328A20FFA3333427D996EF18C38FE2" bold="true" box="[741,756,1968,1994]" pageId="15" pageNumber="16">7</emphasis>
.
<taxonomicName id="4C462DB1FFA33334263D96EE199D8FE2" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[769,938,1968,1994]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFA33334263D96EE199D8FE2" box="[769,938,1968,1994]" italics="true" pageId="15" pageNumber="16">Pyrops effusus</emphasis>
</taxonomicName>
.
<emphasis id="B9328A20FFA33334268496EF19F08FE2" bold="true" box="[952,967,1968,1994]" pageId="15" pageNumber="16">8</emphasis>
.
<taxonomicName id="4C462DB1FFA3333426E896EE1E588FE2" baseAuthorityName="Sato &amp; Nagai" baseAuthorityYear="1994" box="[980,1135,1969,1994]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="gunjii">
<emphasis id="B9328A20FFA3333426E896EE1E588FE2" box="[980,1135,1969,1994]" italics="true" pageId="15" pageNumber="16">Pyrops gunjii</emphasis>
</taxonomicName>
.
<emphasis id="B9328A20FFA33334214196EF1EBB8FE2" bold="true" box="[1149,1164,1968,1994]" pageId="15" pageNumber="16">9</emphasis>
.
<taxonomicName id="4C462DB1FFA3333421A596EE1F458FE2" baseAuthorityName="Distant" baseAuthorityYear="1889" box="[1177,1394,1968,1994]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFA3333421A596EE1F458FE2" box="[1177,1394,1968,1994]" italics="true" pageId="15" pageNumber="16">Pyrops whiteheadi</emphasis>
</taxonomicName>
. —
<emphasis id="B9328A20FFA3333425DF968C1ACF8FC5" bold="true" box="[227,248,2003,2029]" pageId="15" pageNumber="16">A</emphasis>
. Lateral view.
<emphasis id="B9328A20FFA33334249B968C1B8C8FC5" bold="true" box="[423,443,2003,2029]" pageId="15" pageNumber="16">B</emphasis>
. Dorsal view.
<emphasis id="B9328A20FFA333342759968C184D8FC5" bold="true" box="[613,634,2003,2029]" pageId="15" pageNumber="16">C</emphasis>
. Posteroventral view.
<emphasis id="B9328A20FFA3333426BC968B19968FC6" box="[896,929,2004,2030]" italics="true" pageId="15" pageNumber="16">An</emphasis>
= anal tube,
<emphasis id="B9328A20FFA333342106968B1E788FC6" box="[1082,1103,2004,2030]" italics="true" pageId="15" pageNumber="16">G</emphasis>
= gonostylus,
<emphasis id="B9328A20FFA3333421C1968B1F2B8FC5" box="[1277,1308,2004,2029]" italics="true" pageId="15" pageNumber="16">Py</emphasis>
= pygofer.
</paragraph>
<paragraph id="8BF95632FFA3333425DF96A918C18038" blockId="15.[227,758,2038,2065]" box="[227,758,2038,2065]" pageId="15" pageNumber="16">
<pageTitle id="CBD98E55FFA3333425DF96A918C18038" box="[227,758,2038,2065]" pageId="15" pageNumber="16">Scale bar = 1 mm. Photographs by J. Brecko.</pageTitle>
</paragraph>
</caption>
<subSubSection id="C35C05B9FFBC332B242C906E1B45899C" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8BF95632FFBC332B242C906E1B7C8963" blockId="16.[227,1437,305,436]" box="[272,331,305,331]" pageId="16" pageNumber="17">
<emphasis id="B9328A20FFBC332B242C906E1B7C8963" bold="true" box="[272,331,305,331]" pageId="16" pageNumber="17">Note</emphasis>
</paragraph>
<paragraph id="8BF95632FFBC332B25DF900B1B45899C" blockId="16.[227,1437,305,436]" pageId="16" pageNumber="17">
One specimen labelled:
<collectingRegion id="498298D0FFBC332B24C0900B18518946" box="[508,614,340,366]" country="Malaysia" name="Selangor" pageId="16" pageNumber="17">Selangor</collectingRegion>
, Banting, 2 Oct. [19]89, Ismail &amp; Ruslan in the collections of BMNH, has been examined on photograph. The location is in West
<collectingCountry id="F35116A2FFBC332B26AA90281E3589B9" box="[918,1026,375,401]" name="Malaysia" pageId="16" pageNumber="17">Malaysia</collectingCountry>
and the specimen is assumed to be mislabelled.
</paragraph>
</subSubSection>
<caption id="DF3906BAFFBC332B25DF96D11E31803A" ID-DOI="http://doi.org/10.5281/zenodo.3785169" ID-Zenodo-Dep="3785169" httpUri="https://zenodo.org/record/3785169/files/figure.png" pageId="16" pageNumber="17" startId="16.[227,270,1934,1960]" targetBox="[227,1436,495,1889]" targetPageId="16">
<paragraph id="8BF95632FFBC332B25DF96D11FAB8FC6" blockId="16.[227,1436,1934,2031]" pageId="16" pageNumber="17">
<emphasis id="B9328A20FFBC332B25DF96D11B4A8F80" bold="true" box="[227,381,1934,1961]" pageId="16" pageNumber="17">Fig. 10. — A</emphasis>
.
<taxonomicName id="4C462DB1FFBC332B24B096D018028F80" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[396,565,1934,1960]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFBC332B24B096D018028F80" box="[396,565,1934,1960]" italics="true" pageId="16" pageNumber="17">Pyrops effusus</emphasis>
</taxonomicName>
, in natura, Borneo, Kalimantan, Wehea Protected Forest, 25 Sep. 2010, K. Norby. —
<emphasis id="B9328A20FFBC332B245F96ED1B408FE4" bold="true" box="[355,375,1970,1996]" pageId="16" pageNumber="17">B</emphasis>
.
<taxonomicName id="4C462DB1FFBC332B24B496ED18108FE3" baseAuthorityName="Sato &amp; Nagai" baseAuthorityYear="1994" box="[392,551,1970,1995]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="gunjii">
<emphasis id="B9328A20FFBC332B24B496ED18108FE3" box="[392,551,1970,1995]" italics="true" pageId="16" pageNumber="17">Pyrops gunjii</emphasis>
</taxonomicName>
, in natura, Borneo, East Kalimantan, 0°32’4.70”S 117°27’46.73”E, 3 Jul. 2011, M. Hauskrecht. —
<emphasis id="B9328A20FFBC332B2732968B18738FC6" bold="true" box="[526,580,2004,2030]" pageId="16" pageNumber="17">C–F</emphasis>
.
<taxonomicName id="4C462DB1FFBC332B276F968B19548FC6" authorityName="Baker" authorityYear="1925" box="[595,867,2004,2030]" class="Insecta" family="Fulgoridae" genus="Fulgora" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="viridicastanea">
<emphasis id="B9328A20FFBC332B276F968B19548FC6" box="[595,867,2004,2030]" italics="true" pageId="16" pageNumber="17">Fulgora viridicastanea</emphasis>
</taxonomicName>
, holotype (Photographs A. Kennedy).
<emphasis id="B9328A20FFBC332B2016968B1F088FC6" bold="true" box="[1322,1343,2004,2030]" pageId="16" pageNumber="17">C</emphasis>
. Dorsal
</paragraph>
<paragraph id="8BF95632FFBC332B25DF96A71E31803A" blockId="16.[227,1030,2040,2066]" box="[227,1030,2040,2066]" pageId="16" pageNumber="17">
view.
<emphasis id="B9328A20FFBC332B241596A71B09803A" bold="true" box="[297,318,2040,2066]" pageId="16" pageNumber="17">D</emphasis>
. Left lateral view.
<emphasis id="B9328A20FFBC332B272796A71818803A" bold="true" box="[539,559,2040,2066]" pageId="16" pageNumber="17">E</emphasis>
. Head, normal view of frons.
<emphasis id="B9328A20FFBC332B26B396A71996803A" bold="true" box="[911,929,2040,2066]" pageId="16" pageNumber="17">F</emphasis>
. Labels.
</paragraph>
</caption>
<subSubSection id="C35C05B9FFBD3329242C906F1BCE8C30" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
<paragraph id="8BF95632FFBD332A242C906F186C8963" blockId="17.[227,1437,304,402]" box="[272,603,304,331]" pageId="17" pageNumber="18">
<emphasis id="B9328A20FFBD332A242C906F186C8963" bold="true" box="[272,603,304,331]" pageId="17" pageNumber="18">Examined on photographs</emphasis>
</paragraph>
<paragraph id="8BF95632FFBD332A25DF900B1FAA89B9" blockId="17.[227,1437,304,402]" pageId="17" pageNumber="18">
EAST
<collectingCountry id="F35116A2FFBD332A2409900B1BE58946" box="[309,466,340,366]" name="Malaysia" pageId="17" pageNumber="18">MALAYSIA</collectingCountry>
:
<specimenCount id="9D409DBBFFBD332A24DC900B18508946" box="[480,615,340,366]" pageId="17" pageNumber="18" type="generic">1 specimen</specimenCount>
(
<figureCitation id="137D4AB7FFBD332A2745900B18E78946" box="[633,720,340,366]" captionStart="Fig" captionStartId="19.[227,269,1824,1850]" captionTargetBox="[272,1398,302,1793]" captionTargetId="figure@19.[272,1398,302,1793]" captionTargetPageId="19" captionText="Fig. 12. Pyrops whiteheadi and P. intricatus, in natura, trophobiosis. — A–E. P. whiteheadi. A–B. Tended by cockroaches (Blattodea, Dorylaea, probably D. magna (Schelford, 1909)), Sabah, Danum Valley, 4 Jul. 2013, A. Sorokin. C. Tended by small cockroach (Blattodea, Pseudophyllodromiinae), Sabah, Danum Valley, 15 Dec. 2008, C.C. Lee. D. Sabah, Lower Kinabatangan River, 9 Aug. 2012, N. Messina. E. Tended by an adult gecko, Gehyra mutilata (Wiegmann, 1835) (Squamata: Gekkonidae), Sabah, Sepilok, 6 Jun. 2013, C.C. Lee. — F. P. intricatus tended by cockroach (Blattodea, Dorylaea sp.), Sarawak, Mulu National Park, 15 Oct. 2009, C.C. Lee" figureDoi="http://doi.org/10.5281/zenodo.3785173" httpUri="https://zenodo.org/record/3785173/files/figure.png" pageId="17" pageNumber="18">Fig. 12</figureCitation>
A–B),
<collectingRegion id="498298D0FFBD332A261E900B195A8946" box="[802,877,340,366]" country="Malaysia" name="Sabah" pageId="17" pageNumber="18">Sabah</collectingRegion>
, Danum Valley,
<date id="FFF870F2FFBD332A2109900B1E898946" box="[1077,1214,340,366]" pageId="17" pageNumber="18" value="2013-07-04">4 Jul. 2013</date>
, Anton Sorokin;
<specimenCount id="9D409DBBFFBD332A20B2900B1B6289B9" pageId="17" pageNumber="18" type="generic">1 specimen</specimenCount>
, idem,
<date id="FFF870F2FFBD332A249B9028180889B9" box="[423,575,375,402]" pageId="17" pageNumber="18" value="2013-06-13">13 Jun. 2013</date>
;
<specimenCount id="9D409DBBFFBD332A2776902818E689B9" box="[586,721,375,401]" pageId="17" pageNumber="18" type="generic">1 specimen</specimenCount>
, idem,
<date id="FFF870F2FFBD332A26189028199889B9" box="[804,943,375,401]" pageId="17" pageNumber="18" value="2010-10-03">3 Oct. 2010</date>
, Hok Ping Guek;
<specimenCount id="9D409DBBFFBD332A214090281F3489B9" box="[1148,1283,375,401]" pageId="17" pageNumber="18" type="generic">1 specimen</specimenCount>
, idem, 4 Oct.
</paragraph>
<caption id="DF3906BAFFBD332A25DF968F1FAA8026" ID-DOI="http://doi.org/10.5281/zenodo.3785171" ID-Zenodo-Dep="3785171" httpUri="https://zenodo.org/record/3785171/files/figure.png" pageId="17" pageNumber="18" startId="17.[227,269,2000,2026]" targetBox="[227,1436,465,1962]" targetPageId="17">
<paragraph id="8BF95632FFBD332A25DF968F1FAB8FC2" blockId="17.[227,1436,2000,2027]" box="[227,1436,2000,2027]" pageId="17" pageNumber="18">
<emphasis id="B9328A20FFBD332A25DF968F1B098FC2" bold="true" box="[227,318,2000,2026]" pageId="17" pageNumber="18">Fig. 11.</emphasis>
<taxonomicName id="4C462DB1FFBD332A247A968E1BC08FC2" authorityName="Constant" authorityYear="2015" box="[326,503,2001,2026]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="synavei" status="sp. nov.">
<emphasis id="B9328A20FFBD332A247A968E1BC08FC2" box="[326,503,2001,2026]" italics="true" pageId="17" pageNumber="18">Pyrops synavei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201375BFFBD332A24C3968E186D8FC3" box="[511,602,2001,2027]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
in natura.
<emphasis id="B9328A20FFBD332A27E5968F19278FC2" bold="true" box="[729,784,2000,2026]" pageId="17" pageNumber="18">A–B</emphasis>
. Borneo, Kalimantan, Long Alango, Lalut Birai Sacred
</paragraph>
<paragraph id="8BF95632FFBD332A25DF96AB1FAA8026" blockId="17.[227,1437,2035,2062]" box="[227,1437,2035,2062]" pageId="17" pageNumber="18">
<pageTitle id="CBD98E55FFBD332A25DF96AB1FAA8026" box="[227,1437,2035,2062]" pageId="17" pageNumber="18">
Forest, 2 Sep. 2011, J. Woltering.
<emphasis id="B9328A20FFBD332A275996AC18A98025" bold="true" box="[613,670,2035,2061]" pageId="17" pageNumber="18">C–D</emphasis>
. Borneo, Kalimantan, Lalut Birai Forest, 13 Mar. 2009, D. Forney.
</pageTitle>
</paragraph>
</caption>
<paragraph id="8BF95632FFBE332925DF906E1F7D8AA0" blockId="18.[227,1437,305,649]" pageId="18" pageNumber="19">
2013;
<specimenCount id="9D409DBBFFBE33292412906E1B808963" box="[302,439,305,331]" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
, idem,
<date id="FFF870F2FFBE33292732906E18858963" box="[526,690,305,331]" pageId="18" pageNumber="19" value="2008-08-18">18 Aug. 2008</date>
, Thomas Bridle;
<specimenCount id="9D409DBBFFBE33292642906E1E258963" box="[894,1042,305,331]" pageId="18" pageNumber="19" type="generic">2 specimens</specimenCount>
, idem,
<date id="FFF870F2FFBE33292155906E1F268963" box="[1129,1297,305,331]" pageId="18" pageNumber="19" value="2009-09-27">27 Sept. 2009</date>
, Tim Moss;
<specimenCount id="9D409DBBFFBE332925DF900B1B5B8946" box="[227,364,340,366]" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
, idem,
<date id="FFF870F2FFBE332924F8900B18648946" box="[452,595,340,367]" pageId="18" pageNumber="19" value="2007-06-03">3 Jun. 2007</date>
, Karin Saner;
<specimenCount id="9D409DBBFFBE332927C3900B19BE8946" box="[767,905,340,366]" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
, idem,
<date id="FFF870F2FFBE332926E3900A1E578946" box="[991,1120,340,367]" pageId="18" pageNumber="19" value="2004-08">Aug. 2004</date>
, Dirk Mezger;
<specimenCount id="9D409DBBFFBE3329202A900B1FAB8946" box="[1302,1436,340,366]" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
(
<figureCitation id="137D4AB7FFBE332925D090281B6089BA" box="[236,343,375,402]" captionStart="Fig" captionStartId="19.[227,269,1824,1850]" captionTargetBox="[272,1398,302,1793]" captionTargetId="figure@19.[272,1398,302,1793]" captionTargetPageId="19" captionText="Fig. 12. Pyrops whiteheadi and P. intricatus, in natura, trophobiosis. — A–E. P. whiteheadi. A–B. Tended by cockroaches (Blattodea, Dorylaea, probably D. magna (Schelford, 1909)), Sabah, Danum Valley, 4 Jul. 2013, A. Sorokin. C. Tended by small cockroach (Blattodea, Pseudophyllodromiinae), Sabah, Danum Valley, 15 Dec. 2008, C.C. Lee. D. Sabah, Lower Kinabatangan River, 9 Aug. 2012, N. Messina. E. Tended by an adult gecko, Gehyra mutilata (Wiegmann, 1835) (Squamata: Gekkonidae), Sabah, Sepilok, 6 Jun. 2013, C.C. Lee. — F. P. intricatus tended by cockroach (Blattodea, Dorylaea sp.), Sarawak, Mulu National Park, 15 Oct. 2009, C.C. Lee" figureDoi="http://doi.org/10.5281/zenodo.3785173" httpUri="https://zenodo.org/record/3785173/files/figure.png" pageId="18" pageNumber="19">Fig. 12C</figureCitation>
),
<collectingRegion id="498298D0FFBE3329245190281B8F89B9" box="[365,440,375,401]" country="Malaysia" name="Sabah" pageId="18" pageNumber="19">Sabah</collectingRegion>
, Danum Valley,
<date id="FFF870F2FFBE332927469028192B89BA" box="[634,796,375,402]" pageId="18" pageNumber="19" value="2008-12-15">15 Dec. 2008</date>
, Ch’ien C. Lee;
<specimenCount id="9D409DBBFFBE332926E390271E5289B9" box="[991,1125,375,402]" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
(
<figureCitation id="137D4AB7FFBE3329214A90281ED589BA" box="[1142,1250,375,402]" captionStart="Fig" captionStartId="19.[227,269,1824,1850]" captionTargetBox="[272,1398,302,1793]" captionTargetId="figure@19.[272,1398,302,1793]" captionTargetPageId="19" captionText="Fig. 12. Pyrops whiteheadi and P. intricatus, in natura, trophobiosis. — A–E. P. whiteheadi. A–B. Tended by cockroaches (Blattodea, Dorylaea, probably D. magna (Schelford, 1909)), Sabah, Danum Valley, 4 Jul. 2013, A. Sorokin. C. Tended by small cockroach (Blattodea, Pseudophyllodromiinae), Sabah, Danum Valley, 15 Dec. 2008, C.C. Lee. D. Sabah, Lower Kinabatangan River, 9 Aug. 2012, N. Messina. E. Tended by an adult gecko, Gehyra mutilata (Wiegmann, 1835) (Squamata: Gekkonidae), Sabah, Sepilok, 6 Jun. 2013, C.C. Lee. — F. P. intricatus tended by cockroach (Blattodea, Dorylaea sp.), Sarawak, Mulu National Park, 15 Oct. 2009, C.C. Lee" figureDoi="http://doi.org/10.5281/zenodo.3785173" httpUri="https://zenodo.org/record/3785173/files/figure.png" pageId="18" pageNumber="19">Fig. 12D</figureCitation>
),
<collectingRegion id="498298D0FFBE332921C590281F7389B9" box="[1273,1348,375,401]" country="Malaysia" name="Sabah" pageId="18" pageNumber="19">Sabah</collectingRegion>
, Lower Kinabatangan River,
<date id="FFF870F2FFBE332924E790C4185A899D" box="[475,621,411,437]" pageId="18" pageNumber="19" value="2012-08-09">9 Aug. 2012</date>
, Nicola Messina;
<specimenCount id="9D409DBBFFBE3329260290C419F4899D" box="[830,963,411,437]" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
(
<figureCitation id="137D4AB7FFBE332926EE90C41E0C899D" box="[978,1083,411,437]" captionStart="Fig" captionStartId="19.[227,269,1824,1850]" captionTargetBox="[272,1398,302,1793]" captionTargetId="figure@19.[272,1398,302,1793]" captionTargetPageId="19" captionText="Fig. 12. Pyrops whiteheadi and P. intricatus, in natura, trophobiosis. — A–E. P. whiteheadi. A–B. Tended by cockroaches (Blattodea, Dorylaea, probably D. magna (Schelford, 1909)), Sabah, Danum Valley, 4 Jul. 2013, A. Sorokin. C. Tended by small cockroach (Blattodea, Pseudophyllodromiinae), Sabah, Danum Valley, 15 Dec. 2008, C.C. Lee. D. Sabah, Lower Kinabatangan River, 9 Aug. 2012, N. Messina. E. Tended by an adult gecko, Gehyra mutilata (Wiegmann, 1835) (Squamata: Gekkonidae), Sabah, Sepilok, 6 Jun. 2013, C.C. Lee. — F. P. intricatus tended by cockroach (Blattodea, Dorylaea sp.), Sarawak, Mulu National Park, 15 Oct. 2009, C.C. Lee" figureDoi="http://doi.org/10.5281/zenodo.3785173" httpUri="https://zenodo.org/record/3785173/files/figure.png" pageId="18" pageNumber="19">Fig. 12E</figureCitation>
),
<collectingRegion id="498298D0FFBE3329216C90C41EAC899D" box="[1104,1179,411,437]" country="Malaysia" name="Sabah" pageId="18" pageNumber="19">Sabah</collectingRegion>
, Sepilok,
<date id="FFF870F2FFBE3329203290C41FA0899D" box="[1294,1431,411,437]" pageId="18" pageNumber="19" value="2013-06-06">6 Jun. 2013</date>
, Ch’ien C. Lee;
<specimenCount id="9D409DBBFFBE3329249D90E1180189F0" box="[417,566,446,472]" pageId="18" pageNumber="19" type="generic">2 specimens</specimenCount>
, Keningau,
<date id="FFF870F2FFBE332927F590E1195D89F0" box="[713,874,446,472]" pageId="18" pageNumber="19" value="2011-09-24">24 Sep. 2011</date>
, Khairin Saili;
<specimenCount id="9D409DBBFFBE3329211A90E11E8B89F0" box="[1062,1212,446,472]" pageId="18" pageNumber="19" type="generic">2 specimens</specimenCount>
, Kinabatangan,
<date id="FFF870F2FFBE3329204390E11B6B89D3" pageId="18" pageNumber="19" value="2008-02-14">14 Feb. 2008</date>
, Christophe Maerten;
<specimenCount id="9D409DBBFFBE3329275E90BE18DD89D3" box="[610,746,481,507]" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
, idem,
<date id="FFF870F2FFBE3329267C90BE19EE89D3" box="[832,985,481,508]" pageId="18" pageNumber="19" value="2008-01-21">21 Jan. 2008</date>
, Andrea Ferrari;
<specimenCount id="9D409DBBFFBE3329219D90BE1F1E89D3" box="[1185,1321,481,507]" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
, idem,
<date id="FFF870F2FFBE3329204390BE1B6C8A36" pageId="18" pageNumber="19" value="2009-10-17">17 Oct. 2009</date>
, “Pietra &amp; Paoli”;
<specimenCount id="9D409DBBFFBE33292700935B18F28A36" box="[572,709,516,542]" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
, idem,
<date id="FFF870F2FFBE33292620935B19888A36" box="[796,959,516,543]" pageId="18" pageNumber="19" value="2010-05-24">24 May 2010</date>
, Frank Joas;
<specimenCount id="9D409DBBFFBE33292160935B1ED28A36" box="[1116,1253,516,542]" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
, Kinabatangan, Sukau,
<date id="FFF870F2FFBE3329240793771BEB8A6A" box="[315,476,552,578]" pageId="18" pageNumber="19" value="2011-08-12">12 Aug. 2011</date>
, J.M. Gayman;
<specimenCount id="9D409DBBFFBE332927A4937719178A69" box="[664,800,551,578]" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
, Sandakan,
<date id="FFF870F2FFBE3329269193771E798A6A" box="[941,1102,552,578]" pageId="18" pageNumber="19" value="2013-03-27">27 Mar. 2013</date>
, Gavin Golden;
<specimenCount id="9D409DBBFFBE3329202C93771FAF8A69" box="[1296,1432,551,578]" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
, Sandakan, Sukau,
<date id="FFF870F2FFBE33292483931418538A4D" box="[447,612,587,613]" pageId="18" pageNumber="19" value="2009-08-14">14 Aug. 2009</date>
, Samuel Tan;
<specimenCount id="9D409DBBFFBE33292633931419AF8A4D" box="[783,920,587,613]" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
, Taman Bukit Tawau,
<date id="FFF870F2FFBE3329219993141F7C8A4D" box="[1189,1355,587,613]" pageId="18" pageNumber="19" value="2012-08-17">17 Aug. 2012</date>
, Subki Abdul Hadi;
<specimenCount id="9D409DBBFFBE33292440933118338AA0" box="[380,516,622,648]" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
, Sepilok,
<date id="FFF870F2FFBE332927449331193C8AA0" box="[632,779,622,648]" pageId="18" pageNumber="19" value="2010-08-01">1 Aug. 2010</date>
, J.P. Lawrence;
<specimenCount id="9D409DBBFFBE332926F493311E678AA0" box="[968,1104,622,648]" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
, idem,
<date id="FFF870F2FFBE3329219993311F718AA0" box="[1189,1350,622,649]" pageId="18" pageNumber="19" value="2011-08-15">15 Aug. 2011</date>
.
</paragraph>
<subSubSection id="C35C05B9FFBE332925DF93EF1BCE8C30" pageId="18" pageNumber="19" type="description">
<paragraph id="8BF95632FFBE332925DF93EF18178AE1" blockId="18.[227,544,687,714]" box="[227,544,687,714]" pageId="18" pageNumber="19">
<heading id="D0B1E15EFFBE332925DF93EF18178AE1" bold="true" box="[227,544,687,714]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B9328A20FFBE332925DF93EF18178AE1" bold="true" box="[227,544,687,714]" pageId="18" pageNumber="19">Measurements and ratios</emphasis>
</heading>
</paragraph>
<paragraph id="8BF95632FFBE332925DF93811EE38B34" blockId="18.[227,1436,734,796]" pageId="18" pageNumber="19">
TL:
<specimenCount id="9D409DBBFFBE3329242A93811B1B8AD0" box="[278,300,734,760]" pageId="18" pageNumber="19" type="male">♂</specimenCount>
(n = 2):
<quantity id="4CBEFBD7FFBE332924AC93811BCF8AD0" box="[400,504,734,760]" metricMagnitude="-2" metricUnit="m" metricValue="3.4799999999999995" pageId="18" pageNumber="19" unit="mm" value="34.8">34.8 mm</quantity>
(34.3–35.2);
<specimenCount id="9D409DBBFFBE332927AA9381189B8AD0" box="[662,684,734,760]" pageId="18" pageNumber="19" type="female">♀</specimenCount>
(n = 1): 38.0 mm; TL+process:
<specimenCount id="9D409DBBFFBE3329211E93811E0F8AD0" box="[1058,1080,734,760]" pageId="18" pageNumber="19" type="male">♂</specimenCount>
(n = 2):
<quantity id="4CBEFBD7FFBE332921A193811F328AD0" box="[1181,1285,734,760]" metricMagnitude="-2" metricUnit="m" metricValue="3.9799999999999995" pageId="18" pageNumber="19" unit="mm" value="39.8">39.8 mm</quantity>
(38.7–40.9);
<specimenCount id="9D409DBBFFBE332925DF925D1ACE8B34" box="[227,249,770,796]" pageId="18" pageNumber="19" type="female">♀</specimenCount>
(n = 1):
<quantity id="4CBEFBD7FFBE3329245C925E1BFC8B34" box="[352,459,769,796]" metricMagnitude="-2" metricUnit="m" metricValue="4.22" pageId="18" pageNumber="19" unit="mm" value="42.2">42.2 mm</quantity>
; LTg/BTg = 2.73; BF/BPrH = 3.0; LPr/LF = 2.5; LPr/BPrH = 5.4.
</paragraph>
<paragraph id="8BF95632FFBE332925DF921C1BA18B74" blockId="18.[227,406,834,861]" box="[227,406,834,861]" pageId="18" pageNumber="19">
<heading id="D0B1E15EFFBE332925DF921C1BA18B74" bold="true" box="[227,406,834,861]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B9328A20FFBE332925DF921C1BA18B74" bold="true" box="[227,406,834,861]" pageId="18" pageNumber="19">Male genitalia</emphasis>
</heading>
</paragraph>
<paragraph id="8BF95632FFBE332925DF922D1BCE8C30" blockId="18.[227,1437,881,1048]" pageId="18" pageNumber="19">
Brown with gonostyli paler dorsally and ventrally. Pygofer higher than long, with posterior margin sinuate in lateral view (
<figureCitation id="137D4AB7FFBE332924C892CB18658B86" box="[500,594,916,942]" captionStart="Figs 7–9" captionStartId="15.[227,279,1968,1994]" captionTargetBox="[227,1436,344,1927]" captionTargetId="figure@15.[227,1436,344,1927]" captionTargetPageId="15" captionText="Figs 7–9. Pyrops effusus group, ♂ genitalia. 7. Pyrops effusus. 8. Pyrops gunjii. 9. Pyrops whiteheadi. — A. Lateral view. B. Dorsal view. C. Posteroventral view. An = anal tube, G = gonostylus, Py = pygofer. Scale bar = 1 mm. Photographs by J. Brecko" figureDoi="http://doi.org/10.5281/zenodo.3785167" httpUri="https://zenodo.org/record/3785167/files/figure.png" pageId="18" pageNumber="19">Fig. 9A</figureCitation>
). Anal tube slightly elongate, 1.25 times as long as broad, broader at 4/5 of total length (
<figureCitation id="137D4AB7FFBE332924A092E81BD48BFA" box="[412,483,951,978]" captionStart="Figs 7–9" captionStartId="15.[227,279,1968,1994]" captionTargetBox="[227,1436,344,1927]" captionTargetId="figure@15.[227,1436,344,1927]" captionTargetPageId="15" captionText="Figs 7–9. Pyrops effusus group, ♂ genitalia. 7. Pyrops effusus. 8. Pyrops gunjii. 9. Pyrops whiteheadi. — A. Lateral view. B. Dorsal view. C. Posteroventral view. An = anal tube, G = gonostylus, Py = pygofer. Scale bar = 1 mm. Photographs by J. Brecko" figureDoi="http://doi.org/10.5281/zenodo.3785167" httpUri="https://zenodo.org/record/3785167/files/figure.png" pageId="18" pageNumber="19">Fig. 9</figureCitation>
A–B); lateral margins very slightly sinuate (
<figureCitation id="137D4AB7FFBE332926CD92E81E7A8BFA" box="[1009,1101,951,978]" captionStart="Figs 7–9" captionStartId="15.[227,279,1968,1994]" captionTargetBox="[227,1436,344,1927]" captionTargetId="figure@15.[227,1436,344,1927]" captionTargetPageId="15" captionText="Figs 7–9. Pyrops effusus group, ♂ genitalia. 7. Pyrops effusus. 8. Pyrops gunjii. 9. Pyrops whiteheadi. — A. Lateral view. B. Dorsal view. C. Posteroventral view. An = anal tube, G = gonostylus, Py = pygofer. Scale bar = 1 mm. Photographs by J. Brecko" figureDoi="http://doi.org/10.5281/zenodo.3785167" httpUri="https://zenodo.org/record/3785167/files/figure.png" pageId="18" pageNumber="19">Fig. 9B</figureCitation>
) and apical margin strongly notched in dorsal view (
<figureCitation id="137D4AB7FFBE332924C39284186A8BDD" box="[511,605,987,1013]" captionStart="Figs 7–9" captionStartId="15.[227,279,1968,1994]" captionTargetBox="[227,1436,344,1927]" captionTargetId="figure@15.[227,1436,344,1927]" captionTargetPageId="15" captionText="Figs 7–9. Pyrops effusus group, ♂ genitalia. 7. Pyrops effusus. 8. Pyrops gunjii. 9. Pyrops whiteheadi. — A. Lateral view. B. Dorsal view. C. Posteroventral view. An = anal tube, G = gonostylus, Py = pygofer. Scale bar = 1 mm. Photographs by J. Brecko" figureDoi="http://doi.org/10.5281/zenodo.3785167" httpUri="https://zenodo.org/record/3785167/files/figure.png" pageId="18" pageNumber="19">Fig. 9B</figureCitation>
). Gonostyli (
<figureCitation id="137D4AB7FFBE332927CA928419648BDD" box="[758,851,987,1013]" captionStart="Figs 7–9" captionStartId="15.[227,279,1968,1994]" captionTargetBox="[227,1436,344,1927]" captionTargetId="figure@15.[227,1436,344,1927]" captionTargetPageId="15" captionText="Figs 7–9. Pyrops effusus group, ♂ genitalia. 7. Pyrops effusus. 8. Pyrops gunjii. 9. Pyrops whiteheadi. — A. Lateral view. B. Dorsal view. C. Posteroventral view. An = anal tube, G = gonostylus, Py = pygofer. Scale bar = 1 mm. Photographs by J. Brecko" figureDoi="http://doi.org/10.5281/zenodo.3785167" httpUri="https://zenodo.org/record/3785167/files/figure.png" pageId="18" pageNumber="19">Fig. 9A</figureCitation>
) elongate with posterior margin rounded, slightly narrowing (
<figureCitation id="137D4AB7FFBE3329245792A11BDB8C30" box="[363,492,1022,1048]" captionStart="Figs 7–9" captionStartId="15.[227,279,1968,1994]" captionTargetBox="[227,1436,344,1927]" captionTargetId="figure@15.[227,1436,344,1927]" captionTargetPageId="15" captionText="Figs 7–9. Pyrops effusus group, ♂ genitalia. 7. Pyrops effusus. 8. Pyrops gunjii. 9. Pyrops whiteheadi. — A. Lateral view. B. Dorsal view. C. Posteroventral view. An = anal tube, G = gonostylus, Py = pygofer. Scale bar = 1 mm. Photographs by J. Brecko" figureDoi="http://doi.org/10.5281/zenodo.3785167" httpUri="https://zenodo.org/record/3785167/files/figure.png" pageId="18" pageNumber="19">Fig. 9A, C</figureCitation>
).
</paragraph>
</subSubSection>
</subSubSection>
<subSubSection id="C35C05B9FFBE332925DF956018BA8CA1" box="[227,653,1087,1161]" pageId="18" pageNumber="19" type="distribution">
<paragraph id="8BF95632FFBE332925DF95601B4B8C71" blockId="18.[227,380,1087,1113]" box="[227,380,1087,1113]" pageId="18" pageNumber="19">
<heading id="D0B1E15EFFBE332925DF95601B4B8C71" bold="true" box="[227,380,1087,1113]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B9328A20FFBE332925DF95601B4B8C71" bold="true" box="[227,380,1087,1113]" pageId="18" pageNumber="19">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BF95632FFBE332925DF953118BA8CA1" blockId="18.[227,653,1134,1161]" box="[227,653,1134,1161]" pageId="18" pageNumber="19">
Borneo, known from
<collectingRegion id="498298D0FFBE332924E3953118108CA0" box="[479,551,1134,1160]" country="Malaysia" name="Sabah" pageId="18" pageNumber="19">Sabah</collectingRegion>
(
<figureCitation id="137D4AB7FFBE3329270B953118488CA1" box="[567,639,1134,1161]" captionStart="Fig" captionStartId="14.[227,270,2038,2064]" captionTargetBox="[227,1435,518,1988]" captionTargetId="figure@14.[227,1436,518,1997]" captionTargetPageId="14" captionText="Fig. 6. Pyrops effusus group, distribution map." figureDoi="http://doi.org/10.5281/zenodo.3785165" httpUri="https://zenodo.org/record/3785165/files/figure.png" pageId="18" pageNumber="19">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35C05B9FFBE332F25DF95F019DA8BF1" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" type="biology_ecology">
<subSubSection id="C35C05B9FFBE332925DF95F01FA08D49" pageId="18" pageNumber="19" type="biology_ecology">
<paragraph id="8BF95632FFBE332925DF95F01B758CE1" blockId="18.[227,322,1199,1225]" box="[227,322,1199,1225]" pageId="18" pageNumber="19">
<heading id="D0B1E15EFFBE332925DF95F01B758CE1" bold="true" box="[227,322,1199,1225]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B9328A20FFBE332925DF95F01B758CE1" bold="true" box="[227,322,1199,1225]" pageId="18" pageNumber="19">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="8BF95632FFBE332925DF95811FA08D49" blockId="18.[227,1437,1246,1378]" pageId="18" pageNumber="19">No host tree has been identified for this species, for which records are available from all months of the years, with a higher number of records (8/26) in August. However, we do not conclude that the species is more abundant in August; as most data are from photographs taken by tourists, it is possible that the higher amount of available data is biased by the fact that August is a holiday period in many countries.</paragraph>
</subSubSection>
<subSubSection id="C35C05B9FFBE332F25DF94D619DA8BF1" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" type="discussion">
<paragraph id="8BF95632FFBE332925DF94D619D38D8B" blockId="18.[227,996,1417,1444]" box="[227,996,1417,1444]" pageId="18" pageNumber="19">
<heading id="D0B1E15EFFBE332925DF94D619D38D8B" bold="true" box="[227,996,1417,1444]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B9328A20FFBE332925DF94D619D38D8B" bold="true" box="[227,996,1417,1444]" pageId="18" pageNumber="19">
Notes on trophobiosis with
<taxonomicName id="4C462DB1FFBE3329270B94D619228D8C" baseAuthorityName="Distant" baseAuthorityYear="1889" box="[567,789,1417,1444]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFBE3329270B94D619228D8C" bold="true" box="[567,789,1417,1444]" italics="true" pageId="18" pageNumber="19">Pyrops whiteheadi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C462DB1FFBE3329266F94D619D38D8B" baseAuthorityName="Walker" baseAuthorityYear="1857" box="[851,996,1417,1443]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="intricatus">
<emphasis id="B9328A20FFBE3329266F94D619D38D8B" bold="true" box="[851,996,1417,1443]" italics="true" pageId="18" pageNumber="19">P. intricatus</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF95632FFBE332925DF94E81BFE8EA9" blockId="18.[227,1437,1463,1665]" pageId="18" pageNumber="19">
Trophobiosis with ants commonly occurs in
<taxonomicName id="4C462DB1FFBE332927C494E819408DF9" authorityName="Linnaeus" authorityYear="1758" box="[760,887,1463,1489]" class="Insecta" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
, and ant-attendance involving
<taxonomicName id="4C462DB1FFBE332921DA94E81FAA8DF9" authorityName="Evans" authorityYear="1946" box="[1254,1437,1463,1489]" class="Insecta" infraOrder="Fulgoromorpha" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="4" phylum="Arthropoda" rank="infraOrder">Fulgoromorpha</taxonomicName>
was reviewed by
<bibRefCitation id="EFD72BC3FFBE33292482948418B08DDD" author="Bourgoin" box="[446,647,1499,1525]" firstAuthor="Bourgoin" pageId="18" pageNumber="19" pagination="109 - 124" refId="ref9435" refString="Bourgoin T. 1997. Habitat and ant-attendance in Hemiptera: a phylogenetic test with emphasis on trophobiosis in Fulgoromorpha. In: Grandcolas P. (ed.) The Origin of Biodiversity in Insects: Phylogenetic Tests of Evolutionary Scenarios. Memoires du Museum National d'Histoire Naturelle 173: 109 - 124. Museum National d'Histoire Naturelle, Paris." type="journal article" year="1997">Bourgoin (1997)</bibRefCitation>
who recorded the phenomenon in 5 families of
<taxonomicName id="4C462DB1FFBE332921DA94841FAA8DDD" authorityName="Evans" authorityYear="1946" box="[1254,1437,1499,1525]" class="Insecta" infraOrder="Fulgoromorpha" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="4" phylum="Arthropoda" rank="infraOrder">Fulgoromorpha</taxonomicName>
(
<taxonomicName id="4C462DB1FFBE332925D794A11B648E30" authorityName="Spinola" authorityYear="1839" box="[235,339,1534,1560]" class="Insecta" family="Cixiidae" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="family">Cixiidae</taxonomicName>
,
<taxonomicName id="4C462DB1FFBE3329245C94A11BC18E30" baseAuthorityName="Leach" baseAuthorityYear="1815" box="[352,502,1534,1560]" class="Insecta" family="Delphacidae" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="family">Delphacidae</taxonomicName>
,
<taxonomicName id="4C462DB1FFBE3329273E94A118618E30" authorityName="Spinola" authorityYear="1839" box="[514,598,1534,1560]" class="Insecta" family="Issidae" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="family">Issidae</taxonomicName>
,
<taxonomicName id="4C462DB1FFBE3329275F94A119758E30" authorityName="China &amp; Fennah" authorityYear="1952" box="[611,834,1534,1560]" class="Insecta" family="Hypochthonellidae" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="family">Hypochthonellidae</taxonomicName>
and
<taxonomicName id="4C462DB1FFBE3329264194A11E0B8E31" authorityName="Germar" authorityYear="1821" box="[893,1084,1534,1561]" class="Insecta" family="Tettigometridae" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="family">Tettigometridae</taxonomicName>
). He recognized 4 main
<typeStatus id="54FDE890FFBE3329206294A11FAB8E30" box="[1374,1436,1534,1560]" pageId="18" pageNumber="19">types</typeStatus>
of ant-attendance, from opportunistic attendance by ants to long term attendance where ants collect honeydew directly from the anal opening of the planthopper. At the time, no interaction was known to involve
<taxonomicName id="4C462DB1FFBE3329247E97381BF28EA9" authorityName="Latreille" authorityYear="1807" box="[322,453,1639,1665]" class="Insecta" family="Fulgoridae" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="family">Fulgoridae</taxonomicName>
.
</paragraph>
<paragraph id="8BF95632FFBE332925DF97F619428FBE" blockId="18.[227,1437,1705,1943]" pageId="18" pageNumber="19">
Trophobiotic interactions between
<taxonomicName id="4C462DB1FFBE3329274297F618CA8EEB" authorityName="Latreille" authorityYear="1807" box="[638,765,1705,1731]" class="Insecta" family="Fulgoridae" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="family">Fulgoridae</taxonomicName>
and other animals attracted by the droplets of honeydew produced by the lanternflies were only recently reported by (1)
<bibRefCitation id="EFD72BC3FFBE332926DB97931F2A8ECE" author="Roth &amp; Naskrecki" box="[999,1309,1740,1766]" firstAuthor="Roth" pageId="18" pageNumber="19" pagination="189 - 194" refId="ref10454" refString="Roth L. &amp; Naskrecki P. 2001. Trophobiosis between a blatellid cockroach (Macrophyllodromia spp.) and fulgorids (Enchophora and Copidocephala spp.) in Costa Rica. Journal of Orthoptera Research 10: 189 - 194." type="journal article" year="2001">Roth &amp; Naskrecki (2001)</bibRefCitation>
, involving Neotropical species of the genera
<taxonomicName id="4C462DB1FFBE332927B197B019FD8F21" authority="Stal, 1869" authorityName="Stal" authorityYear="1869" box="[653,970,1775,1802]" class="Insecta" family="Fulgoridae" genus="Copidocephala" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B9328A20FFBE332927B197B019778F21" box="[653,832,1775,1801]" italics="true" pageId="18" pageNumber="19">Copidocephala</emphasis>
Stål, 1869
</taxonomicName>
and
<taxonomicName id="4C462DB1FFBE3329213297B01F618F21" authority="Spinola, 1839" authorityName="Spinola" authorityYear="1839" box="[1038,1366,1775,1801]" class="Insecta" family="Fulgoridae" genus="Enchophora" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B9328A20FFBE3329213297B01EA88F21" box="[1038,1183,1775,1801]" italics="true" pageId="18" pageNumber="19">Enchophora</emphasis>
Spinola, 1839
</taxonomicName>
, with
<taxonomicName id="4C462DB1FFBE332925DF964D1B608F04" box="[227,343,1810,1836]" class="Insecta" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="19" phylum="Arthropoda" rank="order">Blattodea</taxonomicName>
; (2)
<bibRefCitation id="EFD72BC3FFBE332924B6964D18E08F05" author="Naskrecki &amp; Nishida" box="[394,727,1810,1837]" firstAuthor="Naskrecki" pageId="18" pageNumber="19" pagination="2397 - 2402" refId="ref10404" refString="Naskrecki P. &amp; Nishida K. 2007. Novel trophobiotic interactions in lantern bugs (Insecta: Auchenorrhyncha: Fulgoridae). Journal of Natural History 41 (37 - 40): 2397 - 2402. http: // dx. doi. org / 10.1080 / 00222930701633570" type="journal article" year="2007">Naskrecki &amp; Nishida (2007)</bibRefCitation>
, involving Neotropical species of the genera
<taxonomicName id="4C462DB1FFBE332921DE964C1FA28F05" authorityName="Stal" authorityYear="1869" box="[1250,1429,1811,1837]" class="Insecta" family="Fulgoridae" genus="Copidocephala" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B9328A20FFBE332921DE964C1FA28F05" box="[1250,1429,1811,1837]" italics="true" pageId="18" pageNumber="19">Copidocephala</emphasis>
</taxonomicName>
,
<taxonomicName id="4C462DB1FFBE332925DF96691B438F78" authorityName="Spinola" authorityYear="1839" box="[227,372,1846,1872]" class="Insecta" family="Fulgoridae" genus="Enchophora" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B9328A20FFBE332925DF96691B438F78" box="[227,372,1846,1872]" italics="true" pageId="18" pageNumber="19">Enchophora</emphasis>
</taxonomicName>
and
<taxonomicName id="4C462DB1FFBE33292488966918FB8F78" authority="Spinola, 1839" authorityName="Spinola" authorityYear="1839" box="[436,716,1846,1872]" class="Insecta" family="Fulgoridae" genus="Phrictus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B9328A20FFBE33292488966918208F78" box="[436,535,1846,1872]" italics="true" pageId="18" pageNumber="19">Phrictus</emphasis>
Spinola, 1839
</taxonomicName>
, with
<taxonomicName id="4C462DB1FFBE3329262796691E1B8F78" authority=", Lepidoptera" authorityName="Lepidoptera" box="[795,1068,1846,1872]" class="Insecta" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="19" phylum="Arthropoda" rank="order">Blattodea, Lepidoptera</taxonomicName>
and gastropods; (3)
<bibRefCitation id="EFD72BC3FFBE3329201096691BBD8F5B" author="Kemal &amp; Kocak" firstAuthor="Kemal" pageId="18" pageNumber="19" refId="ref9996" refString="Kemal M. &amp; Kocak A. O. 2012. A report on the mutualism between Hemidactylus (Gekkonidae) and" type="book" year="2012">Kemal &amp; Koçak (2012)</bibRefCitation>
between
<taxonomicName id="4C462DB1FFBE332927389605198D8F5C" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[516,954,1881,1908]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="candelaria">
<emphasis id="B9328A20FFBE33292738960518D68F5B" box="[516,737,1881,1907]" italics="true" pageId="18" pageNumber="19">Pyrops candelaria</emphasis>
(Linnaeus, 1758)
</taxonomicName>
and the gecko
<taxonomicName id="4C462DB1FFBE332921BC96061B838FBE" authority="(Schneider, 1792)" baseAuthorityName="Schneider" baseAuthorityYear="1792" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="platyurus">
<emphasis id="B9328A20FFBE332921BC96061FAB8F5B" box="[1152,1436,1881,1907]" italics="true" pageId="18" pageNumber="19">Hemidactylus platyurus</emphasis>
(Schneider, 1792)
</taxonomicName>
(
<taxonomicName id="4C462DB1FFBE332924F99623180A8FBE" box="[453,573,1916,1942]" class="Reptilia" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="order">Squamata</taxonomicName>
:
<taxonomicName id="4C462DB1FFBE33292774962318EF8FBF" authorityName="Gray" authorityYear="1825" box="[584,728,1916,1943]" class="Reptilia" family="Gekkonidae" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="family">Gekkonidae</taxonomicName>
) in
<collectingCountry id="F35116A2FFBE33292638962319478FBE" box="[772,880,1916,1942]" name="Thailand" pageId="18" pageNumber="19">Thailand</collectingCountry>
.
</paragraph>
<paragraph id="8BF95632FFBE332925DF96E21EC78FD3" blockId="18.[227,1436,1981,2043]" pageId="18" pageNumber="19">
Interactions between geckos (5 species belonging to 3 different genera) and another family of
<taxonomicName id="4C462DB1FFBE3329203B96E21FAF8FFF" authorityName="Latreille" authorityYear="1807" box="[1287,1432,1981,2007]" class="Insecta" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="4" phylum="Arthropoda" rank="superFamily" superFamily="Fulgoroidea">Fulgoroidea</taxonomicName>
,
<taxonomicName id="4C462DB1FFBE332925DF96BE1B728FD3" authorityName="Spinola" authorityYear="1839" box="[227,325,2017,2043]" class="Insecta" family="Flatidae" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="family">Flatidae</taxonomicName>
, in
<collectingCountry id="F35116A2FFBE3329245296BE1BC98FD3" box="[366,510,2017,2043]" name="Madagascar" pageId="18" pageNumber="19">Madagascar</collectingCountry>
, were previously reported and analyzed by
<bibRefCitation id="EFD72BC3FFBE332926C096BE1EDB8FD3" author="Folling" box="[1020,1260,2017,2043]" etAl="et al." firstAuthor="Folling" pageId="18" pageNumber="19" pagination="279 - 284" refId="ref9849" refString="Folling M., Knogge C. &amp; Bohme W. 2001. Geckos are milking honeydew-producing planthoppers in Madagascar. Journal of Natural History 35 (2): 279 - 284. http: // dx. doi. org / 10.1080 / 00222930150215378" type="journal article" year="2001">
Fölling
<emphasis id="B9328A20FFBE3329216496BD1EA48FD3" box="[1112,1171,2017,2043]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(2001)
</bibRefCitation>
.
</paragraph>
<caption id="DF3906BAFFBF332825DF967F189C8026" ID-DOI="http://doi.org/10.5281/zenodo.3785173" ID-Zenodo-Dep="3785173" httpUri="https://zenodo.org/record/3785173/files/figure.png" pageId="19" pageNumber="20" startId="19.[227,269,1824,1850]" targetBox="[272,1398,302,1793]" targetPageId="19">
<paragraph id="8BF95632FFBF332825DF967F1FAA8FC2" blockId="19.[227,1437,1824,2027]" pageId="19" pageNumber="20">
<emphasis id="B9328A20FFBF332825DF967F1B0A8F12" bold="true" box="[227,317,1824,1850]" pageId="19" pageNumber="20">Fig. 12.</emphasis>
<taxonomicName id="4C462DB1FFBF3328247F967E182F8F13" baseAuthorityName="Distant" baseAuthorityYear="1889" box="[323,536,1825,1851]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFBF3328247F967E182F8F13" box="[323,536,1825,1851]" italics="true" pageId="19" pageNumber="20">Pyrops whiteheadi</emphasis>
</taxonomicName>
and
<emphasis id="B9328A20FFBF33282773967E18E88F12" box="[591,735,1825,1850]" italics="true" pageId="19" pageNumber="20">
<taxonomicName id="4C462DB1FFBF33282773967E18EC8F12" baseAuthorityName="Walker" baseAuthorityYear="1857" box="[591,731,1825,1850]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="intricatus">P. intricatus</taxonomicName>
,
</emphasis>
in natura, trophobiosis. —
<emphasis id="B9328A20FFBF33282124967F1E628F12" bold="true" box="[1048,1109,1824,1850]" pageId="19" pageNumber="20">A–E.</emphasis>
<taxonomicName id="4C462DB1FFBF33282160967E1EC18F13" baseAuthorityName="Distant" baseAuthorityYear="1889" box="[1116,1270,1825,1851]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFBF33282160967E1EC18F13" box="[1116,1270,1825,1851]" italics="true" pageId="19" pageNumber="20">P. whiteheadi</emphasis>
</taxonomicName>
.
<emphasis id="B9328A20FFBF33282038967F1F0C8F12" bold="true" box="[1284,1339,1824,1850]" pageId="19" pageNumber="20">A–B</emphasis>
. Tended by cockroaches (
<taxonomicName id="4C462DB1FFBF33282496961B182B8F76" box="[426,540,1860,1886]" class="Insecta" kingdom="Animalia" order="Blattodea" pageId="19" pageNumber="20" phylum="Arthropoda" rank="order">Blattodea</taxonomicName>
,
<taxonomicName id="4C462DB1FFBF33282716961B18A18F76" box="[554,662,1860,1886]" class="Insecta" family="Blattidae" genus="Dorylaea" kingdom="Animalia" order="Blattodea" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B9328A20FFBF33282716961B18A18F76" box="[554,662,1860,1886]" italics="true" pageId="19" pageNumber="20">Dorylaea</emphasis>
</taxonomicName>
, probably
<taxonomicName id="4C462DB1FFBF33282624961A1E478F76" authority="(Schelford, 1909))" authorityName="" baseAuthorityName="Schelford" baseAuthorityYear="1909" box="[792,1136,1860,1886]" class="Insecta" family="Blattidae" genus="Dorylaea" kingdom="Animalia" order="Blattodea" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="B9328A20FFBF33282624961A19BB8F76" box="[792,908,1861,1886]" italics="true" pageId="19" pageNumber="20">D. magna</emphasis>
(Schelford, 1909))
</taxonomicName>
, Sabah, Danum Valley, 4 Jul. 2013, A. Sorokin.
<emphasis id="B9328A20FFBF332824DF96381BCF8FA9" bold="true" box="[483,504,1895,1921]" pageId="19" pageNumber="20">C</emphasis>
. Tended by small cockroach (
<taxonomicName id="4C462DB1FFBF33282672963819F78FA9" box="[846,960,1895,1921]" class="Insecta" kingdom="Animalia" order="Blattodea" pageId="19" pageNumber="20" phylum="Arthropoda" rank="order">Blattodea</taxonomicName>
,
<taxonomicName id="4C462DB1FFBF332826F796381ED78FAA" authorityName="Hebard" authorityYear="1929" box="[971,1248,1895,1922]" class="Insecta" family="Ectobiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subFamily" subFamily="Pseudophyllodromiinae">Pseudophyllodromiinae</taxonomicName>
), Sabah, Danum Valley, 15 Dec. 2008, C.C. Lee.
<emphasis id="B9328A20FFBF3328277196D518558F8C" bold="true" box="[589,610,1930,1956]" pageId="19" pageNumber="20">D</emphasis>
. Sabah, Lower Kinabatangan River, 9 Aug. 2012, N. Messina.
<emphasis id="B9328A20FFBF3328201496D51F0C8F8C" bold="true" box="[1320,1339,1930,1956]" pageId="19" pageNumber="20">E</emphasis>
. Tended by an adult gecko,
<taxonomicName id="4C462DB1FFBF332824ED96F219B68FE0" authority="(Wiegmann, 1835)" baseAuthorityName="Wiegmann" baseAuthorityYear="1835" box="[465,897,1965,1992]" class="Reptilia" family="Gekkonidae" genus="Gehyra" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="mutilata">
<emphasis id="B9328A20FFBF332824ED96F218A18FEF" box="[465,662,1965,1991]" italics="true" pageId="19" pageNumber="20">Gehyra mutilata</emphasis>
(Wiegmann, 1835)
</taxonomicName>
(
<taxonomicName id="4C462DB1FFBF332826AA96F21E398FEF" box="[918,1038,1965,1991]" class="Reptilia" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="order">Squamata</taxonomicName>
:
<taxonomicName id="4C462DB1FFBF3328212196F21E878FE0" authorityName="Gray" authorityYear="1825" box="[1053,1200,1965,1992]" class="Reptilia" family="Gekkonidae" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="family">Gekkonidae</taxonomicName>
), Sabah, Sepilok, 6 Jun. 2013, C.C. Lee. —
<emphasis id="B9328A20FFBF3328273C968E18258FC3" bold="true" box="[512,530,2001,2027]" pageId="19" pageNumber="20">F</emphasis>
.
<taxonomicName id="4C462DB1FFBF3328271D968E18998FC2" baseAuthorityName="Walker" baseAuthorityYear="1857" box="[545,686,2001,2026]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="intricatus">
<emphasis id="B9328A20FFBF3328271D968E18998FC2" box="[545,686,2001,2026]" italics="true" pageId="19" pageNumber="20">P. intricatus</emphasis>
</taxonomicName>
tended by cockroach (
<taxonomicName id="4C462DB1FFBF33282684968F1E1A8FC2" box="[952,1069,2000,2026]" class="Insecta" kingdom="Animalia" order="Blattodea" pageId="19" pageNumber="20" phylum="Arthropoda" rank="order">Blattodea</taxonomicName>
,
<taxonomicName id="4C462DB1FFBF33282105968E1E9F8FC3" box="[1081,1192,2001,2027]" class="Insecta" family="Blattidae" genus="Dorylaea" kingdom="Animalia" order="Blattodea" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B9328A20FFBF33282105968E1E9F8FC3" box="[1081,1192,2001,2027]" italics="true" pageId="19" pageNumber="20">Dorylaea</emphasis>
</taxonomicName>
sp.), Sarawak, Mulu
</paragraph>
<paragraph id="8BF95632FFBF332825DF96AB189C8026" blockId="19.[227,683,2035,2062]" box="[227,683,2035,2062]" pageId="19" pageNumber="20">
<pageTitle id="CBD98E55FFBF332825DF96AB189C8026" box="[227,683,2035,2062]" pageId="19" pageNumber="20">National Park, 15 Oct. 2009, C.C. Lee.</pageTitle>
</paragraph>
</caption>
<paragraph id="8BF95632FFB8332F25DF906E1F59899D" blockId="20.[227,1437,305,437]" pageId="20" pageNumber="21">
Similar interactions have been observed and documented between
<taxonomicName id="4C462DB1FFB8332F26C0906D1EE08963" baseAuthorityName="Distant" baseAuthorityYear="1889" box="[1020,1239,305,331]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFB8332F26C0906D1EE08963" box="[1020,1239,305,331]" italics="true" pageId="20" pageNumber="21">Pyrops whiteheadi</emphasis>
</taxonomicName>
and two species of
<taxonomicName id="4C462DB1FFB8332F2438900B1B438946" box="[260,372,340,366]" class="Insecta" kingdom="Animalia" order="Blattodea" pageId="20" pageNumber="21" phylum="Arthropoda" rank="order">Blattodea</taxonomicName>
in
<collectingRegion id="498298D0FFB8332F24A0900B1BD08946" box="[412,487,340,366]" country="Malaysia" name="Sabah" pageId="20" pageNumber="21">Sabah</collectingRegion>
, Danum Valley: one species of the subfamily
<taxonomicName id="4C462DB1FFB8332F2135900B1F178946" authorityName="Hebard" authorityYear="1929" box="[1033,1312,340,366]" class="Insecta" family="Ectobiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="20" pageNumber="21" phylum="Arthropoda" rank="subFamily" subFamily="Pseudophyllodromiinae">Pseudophyllodromiinae</taxonomicName>
(C.C. Lee pers. comm.,
<figureCitation id="137D4AB7FFB8332F24BF90281BDB89BA" box="[387,492,375,402]" captionStart="Fig" captionStartId="19.[227,269,1824,1850]" captionTargetBox="[272,1398,302,1793]" captionTargetId="figure@19.[272,1398,302,1793]" captionTargetPageId="19" captionText="Fig. 12. Pyrops whiteheadi and P. intricatus, in natura, trophobiosis. — A–E. P. whiteheadi. A–B. Tended by cockroaches (Blattodea, Dorylaea, probably D. magna (Schelford, 1909)), Sabah, Danum Valley, 4 Jul. 2013, A. Sorokin. C. Tended by small cockroach (Blattodea, Pseudophyllodromiinae), Sabah, Danum Valley, 15 Dec. 2008, C.C. Lee. D. Sabah, Lower Kinabatangan River, 9 Aug. 2012, N. Messina. E. Tended by an adult gecko, Gehyra mutilata (Wiegmann, 1835) (Squamata: Gekkonidae), Sabah, Sepilok, 6 Jun. 2013, C.C. Lee. — F. P. intricatus tended by cockroach (Blattodea, Dorylaea sp.), Sarawak, Mulu National Park, 15 Oct. 2009, C.C. Lee" figureDoi="http://doi.org/10.5281/zenodo.3785173" httpUri="https://zenodo.org/record/3785173/files/figure.png" pageId="20" pageNumber="21">Fig. 12C</figureCitation>
) and one species of the genus
<taxonomicName id="4C462DB1FFB8332F266A902719F289BA" box="[854,965,376,402]" class="Insecta" family="Blattidae" genus="Dorylaea" kingdom="Animalia" order="Blattodea" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B9328A20FFB8332F266A902719F289BA" box="[854,965,376,402]" italics="true" pageId="20" pageNumber="21">Dorylaea</emphasis>
</taxonomicName>
(A. Sorokin pers. comm.,
<figureCitation id="137D4AB7FFB8332F203C90281F6489BA" box="[1280,1363,375,402]" captionStart="Fig" captionStartId="19.[227,269,1824,1850]" captionTargetBox="[272,1398,302,1793]" captionTargetId="figure@19.[272,1398,302,1793]" captionTargetPageId="19" captionText="Fig. 12. Pyrops whiteheadi and P. intricatus, in natura, trophobiosis. — A–E. P. whiteheadi. A–B. Tended by cockroaches (Blattodea, Dorylaea, probably D. magna (Schelford, 1909)), Sabah, Danum Valley, 4 Jul. 2013, A. Sorokin. C. Tended by small cockroach (Blattodea, Pseudophyllodromiinae), Sabah, Danum Valley, 15 Dec. 2008, C.C. Lee. D. Sabah, Lower Kinabatangan River, 9 Aug. 2012, N. Messina. E. Tended by an adult gecko, Gehyra mutilata (Wiegmann, 1835) (Squamata: Gekkonidae), Sabah, Sepilok, 6 Jun. 2013, C.C. Lee. — F. P. intricatus tended by cockroach (Blattodea, Dorylaea sp.), Sarawak, Mulu National Park, 15 Oct. 2009, C.C. Lee" figureDoi="http://doi.org/10.5281/zenodo.3785173" httpUri="https://zenodo.org/record/3785173/files/figure.png" pageId="20" pageNumber="21">Fig. 12</figureCitation>
A–B), possibly
<taxonomicName id="4C462DB1FFB8332F247090C318A1899D" authority="(Schelford, 1909)" authorityName="" baseAuthorityName="Schelford" baseAuthorityYear="1909" box="[332,662,411,437]" class="Insecta" family="Blattidae" genus="Dorylaea" kingdom="Animalia" order="Blattodea" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="B9328A20FFB8332F247090C31BF7899D" box="[332,448,412,437]" italics="true" pageId="20" pageNumber="21">D. magna</emphasis>
(Schelford, 1909)
</taxonomicName>
(identifications of the
<taxonomicName id="4C462DB1FFB8332F26A390C41E38899D" box="[927,1039,411,437]" class="Insecta" kingdom="Animalia" order="Blattodea" pageId="20" pageNumber="21" phylum="Arthropoda" rank="order">Blattodea</taxonomicName>
by G. Beccaloni,
<date id="FFF870F2FFB8332F21D890C41F55899D" box="[1252,1378,411,437]" pageId="20" pageNumber="21" value="2014-12">Dec. 2014</date>
).
</paragraph>
<paragraph id="8BF95632FFB8332F25DF908318538A48" blockId="20.[227,1436,476,608]" pageId="20" pageNumber="21">
Interactions between one species of gecko (
<taxonomicName id="4C462DB1FFB8332F26389083194B89DE" box="[772,892,476,502]" class="Reptilia" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="order">Squamata</taxonomicName>
:
<taxonomicName id="4C462DB1FFB8332F26B090831E2889DF" authorityName="Gray" authorityYear="1825" box="[908,1055,476,503]" class="Reptilia" family="Gekkonidae" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="family">Gekkonidae</taxonomicName>
),
<taxonomicName id="4C462DB1FFB8332F210490831B108A32" authority="(Wiegmann, 1835)" baseAuthorityName="Wiegmann" baseAuthorityYear="1835" class="Reptilia" family="Gekkonidae" genus="Gehyra" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="mutilata">
<emphasis id="B9328A20FFB8332F210490831EC889DE" box="[1080,1279,476,502]" italics="true" pageId="20" pageNumber="21">Gehyra mutilata</emphasis>
(Wiegmann, 1835)
</taxonomicName>
(identification by O. Pauwels,
<date id="FFF870F2FFB8332F27AB935F19258A32" box="[663,786,512,538]" pageId="20" pageNumber="21" value="2014-12">Dec. 2014</date>
) and
<taxonomicName id="4C462DB1FFB8332F2669935F19C48A31" baseAuthorityName="Distant" baseAuthorityYear="1889" box="[853,1011,511,537]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFB8332F2669935F19C48A31" box="[853,1011,511,537]" italics="true" pageId="20" pageNumber="21">P. whiteheadi</emphasis>
</taxonomicName>
was also observed and documented in
<collectingRegion id="498298D0FFB8332F243E937D1B7A8A14" box="[258,333,546,572]" country="Malaysia" name="Sabah" pageId="20" pageNumber="21">Sabah</collectingRegion>
, Sepilok (C.C. Lee pers. comm.,
<figureCitation id="137D4AB7FFB8332F27EA937D19778A15" box="[726,832,546,573]" captionStart="Fig" captionStartId="19.[227,269,1824,1850]" captionTargetBox="[272,1398,302,1793]" captionTargetId="figure@19.[272,1398,302,1793]" captionTargetPageId="19" captionText="Fig. 12. Pyrops whiteheadi and P. intricatus, in natura, trophobiosis. — A–E. P. whiteheadi. A–B. Tended by cockroaches (Blattodea, Dorylaea, probably D. magna (Schelford, 1909)), Sabah, Danum Valley, 4 Jul. 2013, A. Sorokin. C. Tended by small cockroach (Blattodea, Pseudophyllodromiinae), Sabah, Danum Valley, 15 Dec. 2008, C.C. Lee. D. Sabah, Lower Kinabatangan River, 9 Aug. 2012, N. Messina. E. Tended by an adult gecko, Gehyra mutilata (Wiegmann, 1835) (Squamata: Gekkonidae), Sabah, Sepilok, 6 Jun. 2013, C.C. Lee. — F. P. intricatus tended by cockroach (Blattodea, Dorylaea sp.), Sarawak, Mulu National Park, 15 Oct. 2009, C.C. Lee" figureDoi="http://doi.org/10.5281/zenodo.3785173" httpUri="https://zenodo.org/record/3785173/files/figure.png" pageId="20" pageNumber="21">Fig. 12E</figureCitation>
). Geckos generally are attracted by sugar-rich food (O. Pauwels pers. comm., 2014).
</paragraph>
<paragraph id="8BF95632FFB8332F25DF93D8192D8AC0" blockId="20.[227,1437,647,744]" pageId="20" pageNumber="21">
Although it does not belong to the
<taxonomicName id="4C462DB1FFB8332F274993D818F18A89" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[629,710,647,673]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFB8332F274993D818F18A89" box="[629,710,647,673]" italics="true" pageId="20" pageNumber="21">effusus</emphasis>
</taxonomicName>
group, I should also mention that interactions between
<taxonomicName id="4C462DB1FFB8332F207693D718308AED" authority="(Walker, 1857)" baseAuthorityName="Walker" baseAuthorityYear="1857" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="intricatus">
<emphasis id="B9328A20FFB8332F207693D71B648AEC" italics="true" pageId="20" pageNumber="21">Pyrops intricatus</emphasis>
(Walker, 1857)
</taxonomicName>
and
<taxonomicName id="4C462DB1FFB8332F270293F518998AEC" box="[574,686,682,708]" class="Insecta" kingdom="Animalia" order="Blattodea" pageId="20" pageNumber="21" phylum="Arthropoda" rank="order">Blattodea</taxonomicName>
of the genus
<taxonomicName id="4C462DB1FFB8332F267493F519808AEC" box="[840,951,682,708]" class="Insecta" family="Blattidae" genus="Dorylaea" kingdom="Animalia" order="Blattodea" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B9328A20FFB8332F267493F519808AEC" box="[840,951,682,708]" italics="true" pageId="20" pageNumber="21">Dorylaea</emphasis>
</taxonomicName>
have been documented in
<collectingCountry id="F35116A2FFB8332F21D693F51F658AEC" box="[1258,1362,682,708]" name="Malaysia" pageId="20" pageNumber="21">Sarawak</collectingCountry>
, Mulu National Park (C.C. Lee pers. comm.,
<figureCitation id="137D4AB7FFB8332F27999391193A8AC0" box="[677,781,718,744]" captionStart="Fig" captionStartId="19.[227,269,1824,1850]" captionTargetBox="[272,1398,302,1793]" captionTargetId="figure@19.[272,1398,302,1793]" captionTargetPageId="19" captionText="Fig. 12. Pyrops whiteheadi and P. intricatus, in natura, trophobiosis. — A–E. P. whiteheadi. A–B. Tended by cockroaches (Blattodea, Dorylaea, probably D. magna (Schelford, 1909)), Sabah, Danum Valley, 4 Jul. 2013, A. Sorokin. C. Tended by small cockroach (Blattodea, Pseudophyllodromiinae), Sabah, Danum Valley, 15 Dec. 2008, C.C. Lee. D. Sabah, Lower Kinabatangan River, 9 Aug. 2012, N. Messina. E. Tended by an adult gecko, Gehyra mutilata (Wiegmann, 1835) (Squamata: Gekkonidae), Sabah, Sepilok, 6 Jun. 2013, C.C. Lee. — F. P. intricatus tended by cockroach (Blattodea, Dorylaea sp.), Sarawak, Mulu National Park, 15 Oct. 2009, C.C. Lee" figureDoi="http://doi.org/10.5281/zenodo.3785173" httpUri="https://zenodo.org/record/3785173/files/figure.png" pageId="20" pageNumber="21">Fig. 12F</figureCitation>
).
</paragraph>
<paragraph id="8BF95632FFB8332F25DF925019DA8BF1" blockId="20.[227,1437,783,985]" pageId="20" pageNumber="21">
The unavailabily of such observations for the three other species of the
<taxonomicName id="4C462DB1FFB8332F210492501EBE8B01" baseAuthorityName="Distant" baseAuthorityYear="1891" box="[1080,1161,783,809]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="effusus">
<emphasis id="B9328A20FFB8332F210492501EBE8B01" box="[1080,1161,783,809]" italics="true" pageId="20" pageNumber="21">effusus</emphasis>
</taxonomicName>
group can probably be explained by the fact that
<taxonomicName id="4C462DB1FFB8332F2724926C18808B64" baseAuthorityName="Distant" baseAuthorityYear="1889" box="[536,695,818,844]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B9328A20FFB8332F2724926C18808B64" box="[536,695,818,844]" italics="true" pageId="20" pageNumber="21">P. whiteheadi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C462DB1FFB8332F27CE926C19B78B64" baseAuthorityName="Walker" baseAuthorityYear="1857" box="[754,896,819,844]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="intricatus">
<emphasis id="B9328A20FFB8332F27CE926C19B78B64" box="[754,896,819,844]" italics="true" pageId="20" pageNumber="21">P. intricatus</emphasis>
</taxonomicName>
are present and apparently rather common in several preserved areas (e.g., Danum Valley, Sepilok, Kinabatangan for the first, Mulu for the second) which are extensively visited by naturalists and ecotourists, while the other species are found in more remote regions. All observations reported here come from such sources and it is likely that the same interactions will be documented for the other species in the future.
</paragraph>
</subSubSection>
</subSubSection>
</treatment>
</document>
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