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<mods:title id="34E6536ECBFE09ED7F9A2F028D36F9E5">A dryinid wasp in Early Eocene amber from the Paris basin (Hymenoptera: Dryinidae)</mods:title>
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<treatment id="03EFB146FFC5FF9F7813FE3DFC57B36C" ID-DOI="http://doi.org/10.5281/zenodo.5059366" ID-GBIF-Taxon="183233310" ID-Zenodo-Dep="5059366" LSID="urn:lsid:plazi:treatment:03EFB146FFC5FF9F7813FE3DFC57B36C" httpUri="http://treatment.plazi.org/id/03EFB146FFC5FF9F7813FE3DFC57B36C" lastPageId="5" lastPageNumber="36" pageId="1" pageNumber="32">
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<heading id="D0B1B73CFFC5FF9B7813FE3DFD1EB4FA" bold="true" box="[264,653,393,419]" fontSize="11" level="1" pageId="1" pageNumber="32" reason="1">
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<taxonomicName id="4C467BD3FFC5FF9B7813FE3DFDD1B4FA" authority="Peinado &amp; Nel &amp; Waller, 2006" authorityName="Peinado &amp; Nel &amp; Waller" authorityYear="2006" box="[264,578,393,419]" class="Insecta" family="Dryinidae" genus="Pseudodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="parisiensis" status="sp. nov.">
<emphasis id="B932DC42FFC5FF9B7813FE3DFDD1B4FA" bold="true" box="[264,578,393,419]" italics="true" pageId="1" pageNumber="32">Pseudodryinus parisiensis</emphasis>
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<paragraph id="8BF90050FFC5FF9B7813FE18FEE9B49F" blockId="1.[264,653,393,454]" box="[264,378,428,454]" pageId="1" pageNumber="32">
(
<figureCitation id="137D1CD5FFC5FF9B780BFE18FEE1B49F" box="[272,370,428,454]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="1.[264,368,1391,1415]" captionStartId-1="2.[264,367,1051,1075]" captionStartId-2="2.[264,368,1843,1867]" captionStartId-3="3.[264,367,1529,1553]" captionStartId-4="4.[264,368,1301,1325]" captionStartId-5="4.[264,367,1818,1842]" captionStartId-6="5.[264,367,1133,1157]" captionTargetBox-0="[274,1315,864,1371]" captionTargetBox-1="[468,1122,516,1012]" captionTargetBox-2="[269,1317,1182,1836]" captionTargetBox-3="[289,1298,806,1505]" captionTargetBox-4="[300,1286,606,1277]" captionTargetBox-5="[294,1302,1441,1786]" captionTargetBox-6="[466,1121,646,1109]" captionTargetId-0="figure-296@1.[248,1329,849,1388]" captionTargetId-1="figure-111@2.[450,1137,509,1027]" captionTargetId-2="figure-115@2.[269,1317,1180,1837]" captionTargetId-3="figure-308@3.[289,1298,806,1505]" captionTargetId-4="figure-134@4.[300,1286,606,1277]" captionTargetId-5="figure-138@4.[259,1356,1431,1810]" captionTargetId-6="figure-247@5.[466,1121,646,1109]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="2" captionTargetPageId-3="3" captionTargetPageId-4="4" captionTargetPageId-5="4" captionTargetPageId-6="5" captionText-0="FIGURE 1. Pseudodryinus parisiensis sp. n. holotype PA 2378 1/2, drawing of general habitus (scale bar represents 1 mm)." captionText-1="FIGURE 2. Pseudodryinus parisiensis sp. n. holotype PA 2378 1/2, drawing of head, dorsal view (scale bar represents 1 mm)." captionText-2="FIGURE 3. Pseudodryinus parisiensis sp. n. holotype PA 2378 1/2, photograph of fore wing (scale bar represents 0.5 mm)." captionText-3="FIGURE 4. Pseudodryinus parisiensis sp. n. holotype PA 2378 1/2, photograph of fore leg and head, lateral view (scale bar represents 0.5 mm)." captionText-4="FIGURE 5. Pseudodryinus parisiensis sp. n. holotype PA 2378 1/2, photograph of fore leg and head, dorso­lateral view (scale bar represents 0.5 mm)." captionText-5="FIGURE 6. Pseudodryinus parisiensis sp. n. holotype PA 2378 1/2, drawing of fore tarsus (scale bar represents 0.3 mm)." captionText-6="FIGURE 7. Pseudodryinus parisiensis sp. n. holotype PA 2378 1/2, photograph of head with mandibles (arrow), fronto­ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.5058584" figureDoi-1="http://doi.org/10.5281/zenodo.5058586" figureDoi-2="http://doi.org/10.5281/zenodo.5058590" figureDoi-3="http://doi.org/10.5281/zenodo.5058592" figureDoi-4="http://doi.org/10.5281/zenodo.5058594" figureDoi-5="http://doi.org/10.5281/zenodo.5058598" figureDoi-6="http://doi.org/10.5281/zenodo.5058600" httpUri-0="https://zenodo.org/record/5058584/files/figure.png" httpUri-1="https://zenodo.org/record/5058586/files/figure.png" httpUri-2="https://zenodo.org/record/5058590/files/figure.png" httpUri-3="https://zenodo.org/record/5058592/files/figure.png" httpUri-4="https://zenodo.org/record/5058594/files/figure.png" httpUri-5="https://zenodo.org/record/5058598/files/figure.png" httpUri-6="https://zenodo.org/record/5058600/files/figure.png" pageId="1" pageNumber="32">figs 17</figureCitation>
)
</paragraph>
<paragraph id="8BF90050FFC5FF9B7813FE48FB03B677" blockId="1.[264,1322,508,814]" pageId="1" pageNumber="32">Diagnosis: Mandible with four teeth; palpal formula 5/2; maxillary palpomeres long; rudimentary tarsal claw of front leg present; enlarged tarsal claw of front leg with a subapical tooth, as long as fifth tarsal segment, both bearing lamellae; front tarsal segments 2 and 3 with a strong hook; tibial spur formula (1/1/2); ocelli present; occipital carina complete; mesosomal segments not fused; notauli clearly visible on mesoscutum; epicnemium visible; fore wing cells C, R, 1Cu and 2R1 closed, cells 1R1, 1M and 2Cu distally closed by nebulous veins; cells 2Rs, 2M, and 3Cu present but distally opened. The combination of tibial spur (1/1/2) and palpal (5/2) formulae is diagnostic for the new species.</paragraph>
<caption id="DF3950D8FFC5FF9B7813FADBFDA5B0F0" ID-DOI="http://doi.org/10.5281/zenodo.5058584" ID-Zenodo-Dep="5058584" httpUri="https://zenodo.org/record/5058584/files/figure.png" pageId="1" pageNumber="32" startId="1.[264,368,1391,1415]" targetBox="[274,1315,864,1371]" targetPageId="1">
<paragraph id="8BF90050FFC5FF9B7813FADBFDA5B0F0" blockId="1.[264,1322,1391,1449]" pageId="1" pageNumber="32">
<emphasis id="B932DC42FFC5FF9B7813FADBFE1EB0DE" bold="true" box="[264,397,1391,1415]" pageId="1" pageNumber="32">FIGURE 1.</emphasis>
<taxonomicName id="4C467BD3FFC5FF9B788CFADBFD20B0DF" authority="Peinado &amp; Nel &amp; Waller, 2006" authorityName="Peinado &amp; Nel &amp; Waller" authorityYear="2006" box="[407,691,1391,1414]" class="Insecta" family="Dryinidae" genus="Pseudodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="parisiensis" status="sp. nov.">
<emphasis id="B932DC42FFC5FF9B788CFADBFD20B0DF" box="[407,691,1391,1414]" italics="true" pageId="1" pageNumber="32">Pseudodryinus parisiensis</emphasis>
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<taxonomicNameLabel id="A2016139FFC5FF9B7BA6FADBFD6AB0DF" box="[701,761,1391,1414]" pageId="1" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
holotype PA 2378 1/2, drawing of general habitus (scale bar represents 1 mm).
</paragraph>
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<paragraph id="8BF90050FFC5FF987823FA68FEC4B48F" blockId="1.[264,1322,1500,1886]" lastBlockId="2.[264,1323,284,470]" lastPageId="2" lastPageNumber="33" pageId="1" pageNumber="32">
Description. Body and legs brown yellow; body
<quantity id="4CBEADB5FFC5FF9B7A81FA68FC70B0AF" box="[922,995,1500,1526]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="32" unit="mm" value="4.0">4 mm</quantity>
long; head hypognathous, longer than wide; eyes oval, large, occupying nearly the whole length of the head; three ocelli between eyes, making an obtuse equilateral triangle, the median one distinctly larger; occiput short, vertical, and dorsally bordered by a carina; frons occupied by a depression that ends in clypeus; OL = 12; POL = 9; OOL = 30; OPL = 10; TL = 15; antenna with 10 bare segments; scape robust; pedicel and flagellum cylindrical, progressively shortening, with segments in the following proportions 28:10:40:30:22:19:15:14:14:11; antenna much shorter than body, approximately 3.3 times as long as head (length of head measured dorsally from occipital carina behind the ocelli to distal apex of the mandibles): 203:61; maxillary palp with four visible segments but with a distinct angle between second and third visible palpomeres, implying the presence of a not visible basal palpomere, visible palpomeres long and thin, in the following proportions 15:20:20:15; labial palp twosegmented; head broader than pronotum; neck robust; mandibles with three strong teeth and a smaller basal one, third tooth smaller than second and fourth.
</paragraph>
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<paragraph id="8BF90050FFC6FF987813FBAFFDA5B10C" blockId="2.[264,1323,1051,1109]" pageId="2" pageNumber="33">
<emphasis id="B932DC42FFC6FF987813FBAFFE18B16A" bold="true" box="[264,395,1051,1075]" pageId="2" pageNumber="33">FIGURE 2.</emphasis>
<taxonomicName id="4C467BD3FFC6FF987888FBAFFD3EB16B" authority="Peinado &amp; Nel &amp; Waller, 2006" authorityName="Peinado &amp; Nel &amp; Waller" authorityYear="2006" box="[403,685,1051,1074]" class="Insecta" family="Dryinidae" genus="Pseudodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="33" phylum="Arthropoda" rank="species" species="parisiensis" status="sp. nov.">
<emphasis id="B932DC42FFC6FF987888FBAFFD3EB16B" box="[403,685,1051,1074]" italics="true" pageId="2" pageNumber="33">Pseudodryinus parisiensis</emphasis>
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<taxonomicNameLabel id="A2016139FFC6FF987BAFFBAFFD7CB16B" box="[692,751,1051,1074]" pageId="2" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
holotype PA 2378 1/2, drawing of head, dorsal view (scale bar represents 1 mm).
</paragraph>
</caption>
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<paragraph id="8BF90050FFC6FF987813F887FD96B234" blockId="2.[264,1323,1843,1901]" pageId="2" pageNumber="33">
<emphasis id="B932DC42FFC6FF987813F887FE19B212" bold="true" box="[264,394,1843,1867]" pageId="2" pageNumber="33">FIGURE 3.</emphasis>
<taxonomicName id="4C467BD3FFC6FF98788AF880FD39B212" authority="Peinado &amp; Nel &amp; Waller, 2006" authorityName="Peinado &amp; Nel &amp; Waller" authorityYear="2006" box="[401,682,1844,1867]" class="Insecta" family="Dryinidae" genus="Pseudodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="33" phylum="Arthropoda" rank="species" species="parisiensis" status="sp. nov.">
<emphasis id="B932DC42FFC6FF98788AF880FD39B212" box="[401,682,1844,1867]" italics="true" pageId="2" pageNumber="33">Pseudodryinus parisiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2016139FFC6FF987BB4F880FD7BB212" box="[687,744,1844,1867]" pageId="2" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
holotype PA 2378 1/2, photograph of fore wing (scale bar represents 0.5 mm).
</paragraph>
</caption>
<paragraph id="8BF90050FFC7FF997823FEA8FC87B4DF" blockId="3.[264,1324,284,750]" pageId="3" pageNumber="34">
Mesosomal segments delimited by sutures, not fused; mesosoma partly hidden by legs; pronotum apparently bare; notauli complete; propodeum strongly depressed near petiole; epicnemium visible (
<bibRefCitation id="EFD77DA1FFC7FF997B21FED8FD2FB4DF" author="Olmi, M." box="[570,700,364,390]" pageId="3" pageNumber="34" pagination="19 - 62" refId="ref4549" refString="Olmi, M. (1992) Descriptions of new taxa of Dryinidae (Hymenoptera: Chrysidoidea). Frustula Entomologica, (N. S.), 15 (28), 19 - 62." type="journal article" year="1992">Olmi 1992</bibRefCitation>
,
<bibRefCitation id="EFD77DA1FFC7FF997BD3FED8FC94B4DF" author="Olmi, M." box="[712,775,364,390]" pageId="3" pageNumber="34" pagination="57 - 89" refId="ref4587" refString="Olmi, M. (1993) A new generic classification for Thaumatodryininae, Dryininae and Gonatopodinae, with descriptions of new species (Hymenoptera: Dryinidae). Bollettino di Zoologia Agraria e Bachicoltura, (2), 25 (1), 57 - 89." type="journal article" year="1993">1993</bibRefCitation>
).
</paragraph>
<paragraph id="8BF90050FFC7FF997823FE20FB7FB72F" blockId="3.[264,1324,284,750]" pageId="3" pageNumber="34">Fore wing 1.8 mm long; pterostigma narrow; only three cells are closed by tubular veins, i.e. cells C, R, 1Cu and 2R1; cell 1R1 closed but bordered anteriorly and basally by tubular veins, posteriorly and distally by nebulous veins; cells 1M and 2Cu both bordered basally by a tubular vein and by nebulous veins on their three other sides; cells 2Rs, 2M, and 3Cu open with their anterior and posterior sides bordered by nebulous veins; anal margin of wing not visible; wing covered with short and dense bristles on whole surface.</paragraph>
<paragraph id="8BF90050FFC7FF997823FD30FD9BB7B7" blockId="3.[264,1324,284,750]" pageId="3" pageNumber="34">Hind wing narrow not fully visible; four hamuli on anterior edge at ¾ of distance from base to wing apex, with only vein Sc+R present; wing covered with short and dense bristles on whole surface.</paragraph>
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<paragraph id="8BF90050FFC7FF997813FA4DFC9EB36A" blockId="3.[264,1322,1529,1587]" pageId="3" pageNumber="34">
<emphasis id="B932DC42FFC7FF997813FA4DFE1EB348" bold="true" box="[264,397,1529,1553]" pageId="3" pageNumber="34">FIGURE 4.</emphasis>
<taxonomicName id="4C467BD3FFC7FF99788CFA4EFD20B348" authority="Peinado &amp; Nel &amp; Waller, 2006" authorityName="Peinado &amp; Nel &amp; Waller" authorityYear="2006" box="[407,691,1530,1553]" class="Insecta" family="Dryinidae" genus="Pseudodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="parisiensis" status="sp. nov.">
<emphasis id="B932DC42FFC7FF99788CFA4EFD20B348" box="[407,691,1530,1553]" italics="true" pageId="3" pageNumber="34">Pseudodryinus parisiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2016139FFC7FF997BA7FA4DFD6BB349" box="[700,760,1529,1552]" pageId="3" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
holotype PA 2378 1/2, photograph of fore leg and head, lateral view (scale bar represents 0.5 mm).
</paragraph>
</caption>
<paragraph id="8BF90050FFC7FF9E7823F9D3FC87B77F" blockId="3.[264,1323,1639,1905]" lastBlockId="4.[264,1323,284,550]" lastPageId="4" lastPageNumber="35" pageId="3" pageNumber="34">Legs. Front leg very long, with a terminal chela; the two arms of chela of equal length; first arm (enlarged tarsal claw) with a small subapical tooth and at least one row of lamellae, and distinctly shorter than front tibia; apex of enlarged tarsal claw pointed and curved, sheltered in the terminal “cup” of the other branch of chela (fifth tarsomere); this “cup” is nearly circular with a rather large diameter, it is covered with lamellae as is the inner face of the fifth tarsomere; fifth tarsomere with a complete series of long spines on its outer face; arolium present between the two arms of the chela, rather long; second tarsal claw atrophied, very small, half length of arolium; front tarsal segments 2 and 3 both with a strong hook turned upsidedown, both hooks bearing five long bristles at their extremity; fore leg segments in the following proportions: 5:3:12:12:5:1:1.5:6:9 for coxa:trochanter:femur:tibia:tarsomeres 15 respectively; mid leg segments in the following proportions: 12:13:56:60:46:13:13:5:16; hind leg segments in the following proportions: 40:15:75:97:40:20:15:5:12; tibial spur formula 1/1/2; median and posterior legs with an apical pair of claws separated by an arolium.</paragraph>
<caption id="DF3950D8FFC0FF9E7813FAA1FCCDB016" ID-DOI="http://doi.org/10.5281/zenodo.5058594" ID-Zenodo-Dep="5058594" httpUri="https://zenodo.org/record/5058594/files/figure.png" pageId="4" pageNumber="35" startId="4.[264,368,1301,1325]" targetBox="[300,1286,606,1277]" targetPageId="4">
<paragraph id="8BF90050FFC0FF9E7813FAA1FCCDB016" blockId="4.[264,1322,1301,1359]" pageId="4" pageNumber="35">
<emphasis id="B932DC42FFC0FF9E7813FAA1FE1EB074" bold="true" box="[264,397,1301,1325]" pageId="4" pageNumber="35">FIGURE 5.</emphasis>
<taxonomicName id="4C467BD3FFC0FF9E788CFAA1FD20B075" authority="Peinado &amp; Nel &amp; Waller, 2006" authorityName="Peinado &amp; Nel &amp; Waller" authorityYear="2006" box="[407,691,1301,1324]" class="Insecta" family="Dryinidae" genus="Pseudodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="parisiensis" status="sp. nov.">
<emphasis id="B932DC42FFC0FF9E788CFAA1FD20B075" box="[407,691,1301,1324]" italics="true" pageId="4" pageNumber="35">Pseudodryinus parisiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2016139FFC0FF9E7BA7FAA1FD6BB075" box="[700,760,1301,1324]" pageId="4" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
holotype PA 2378 1/2, photograph of fore leg and head, dorso­lateral view (scale bar represents 0.5 mm).
</paragraph>
</caption>
<caption id="DF3950D8FFC0FF9E7813F8AEFD96B20A" ID-DOI="http://doi.org/10.5281/zenodo.5058598" ID-Zenodo-Dep="5058598" httpUri="https://zenodo.org/record/5058598/files/figure.png" pageId="4" pageNumber="35" startId="4.[264,367,1818,1842]" targetBox="[294,1302,1441,1786]" targetPageId="4">
<paragraph id="8BF90050FFC0FF9E7813F8AEFD96B20A" blockId="4.[264,1323,1818,1875]" pageId="4" pageNumber="35">
<emphasis id="B932DC42FFC0FF9E7813F8AEFE18B26B" bold="true" box="[264,395,1818,1842]" pageId="4" pageNumber="35">FIGURE 6.</emphasis>
<taxonomicName id="4C467BD3FFC0FF9E788FF8AEFD3DB268" authority="Peinado &amp; Nel &amp; Waller, 2006" authorityName="Peinado &amp; Nel &amp; Waller" authorityYear="2006" box="[404,686,1818,1841]" class="Insecta" family="Dryinidae" genus="Pseudodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="parisiensis" status="sp. nov.">
<emphasis id="B932DC42FFC0FF9E788FF8AEFD3DB268" box="[404,686,1818,1841]" italics="true" pageId="4" pageNumber="35">Pseudodryinus parisiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2016139FFC0FF9E7BADF8AEFD62B268" box="[694,753,1818,1841]" pageId="4" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
holotype PA 2378 1/2, drawing of fore tarsus (scale bar represents 0.3 mm).
</paragraph>
</caption>
<paragraph id="8BF90050FFC1FF9F7823FEA8FD48B407" blockId="5.[264,1323,284,550]" pageId="5" pageNumber="36">Abdomen with a clearly visible ovipositor; two antepenultimate segments ventrally with four very long bristles (sensillae?).</paragraph>
<paragraph id="8BF90050FFC1FF9F7823FED8FECFB77F" blockId="5.[264,1323,284,550]" pageId="5" pageNumber="36">
<materialsCitation id="3B2E0A0DFFC1FF9F7823FED8FECBB77F" ID-GBIF-Occurrence="3313047301" collectingDate="2378-02-01" collectorName="De Ploeg &amp; Le Quesnoy" country="Canada" location="Canada" pageId="5" pageNumber="36" specimenCount="1" typeStatus="holotype">
Material. Female
<typeStatus id="54FDBEF2FFC1FF9F7B11FED8FDE2B4DF" box="[522,625,364,390]" pageId="5" pageNumber="36" type="holotype">holotype</typeStatus>
specimen PA
<date id="FFF82690FFC1FF9F7A06FED8FC1BB4DF" box="[797,904,364,390]" pageId="5" pageNumber="36" value="2378-02-01">
<collectingDate id="EFBCDF78FFC1FF9F7A06FED8FC1BB4DF" box="[797,904,364,390]" pageId="5" pageNumber="36" value="2378-02-01">2378 1/2</collectingDate>
</date>
, in the same piece of amber with a specimen belonging to the Mantodea, mounted in
<collectingCountry id="F35140C0FFC1FF9F7A45FE20FC25B4F7" box="[862,950,404,430]" name="Canada" pageId="5" pageNumber="36">Canada</collectingCountry>
balsam, in collection
<collectorName id="26B36586FFC1FF9F7DA6FE20FAB8B4F7" box="[1213,1323,404,430]" pageId="5" pageNumber="36">De Ploëg</collectorName>
and Indivision
<taxonomicName id="4C467BD3FFC1FF9F78D8FE08FDBAB48F" box="[451,553,444,470]" family="Langlois" pageId="5" pageNumber="39" rank="family">Langlois</taxonomicName>
Meurine, deposited in Muséum National d'Histoire Naturelle, Paris. Specimens collected in
<collectorName id="26B36586FFC1FF9F7B75FE50FD6CB4A7" box="[622,767,484,510]" pageId="5" pageNumber="36">Le Quesnoy</collectorName>
all bear the letter PA for Paris (meaning Paris Basin)
</materialsCitation>
.
</paragraph>
<caption id="DF3950D8FFC1FF9F7813FBD9FD13B1F1" ID-DOI="http://doi.org/10.5281/zenodo.5058600" ID-Zenodo-Dep="5058600" httpUri="https://zenodo.org/record/5058600/files/figure.png" pageId="5" pageNumber="36" startId="5.[264,367,1133,1157]" targetBox="[466,1121,646,1109]" targetPageId="5">
<paragraph id="8BF90050FFC1FF9F7813FBD9FD13B1F1" blockId="5.[264,1323,1133,1192]" pageId="5" pageNumber="36">
<emphasis id="B932DC42FFC1FF9F7813FBD9FE19B1DC" bold="true" box="[264,394,1133,1157]" pageId="5" pageNumber="36">FIGURE 7.</emphasis>
<taxonomicName id="4C467BD3FFC1FF9F788AFBDAFD3AB1DC" authority="Peinado &amp; Nel &amp; Waller, 2006" authorityName="Peinado &amp; Nel &amp; Waller" authorityYear="2006" box="[401,681,1134,1157]" class="Insecta" family="Dryinidae" genus="Pseudodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="parisiensis" status="sp. nov.">
<emphasis id="B932DC42FFC1FF9F788AFBDAFD3AB1DC" box="[401,681,1134,1157]" italics="true" pageId="5" pageNumber="36">Pseudodryinus parisiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2016139FFC1FF9F7BAAFBDAFD79B1DC" box="[689,746,1134,1157]" pageId="5" pageNumber="36" rank="species">sp. n.</taxonomicNameLabel>
holotype PA 2378 1/2, photograph of head with mandibles (arrow), fronto­ventral view.
</paragraph>
</caption>
<paragraph id="8BF90050FFC1FF9F7823FAB7FA90B044" blockId="5.[264,1322,1283,1589]" box="[312,1283,1283,1309]" pageId="5" pageNumber="36">
Locality deposit. Le Quesnoy, Chevrière, region of Creil, Oise department,
<collectingCountry id="F35140C0FFC1FF9F7DB7FAB7FA93B044" box="[1196,1280,1283,1309]" name="France" pageId="5" pageNumber="36">France</collectingCountry>
.
</paragraph>
<paragraph id="8BF90050FFC1FF9F7823FA9FFD81B0E4" blockId="5.[264,1322,1283,1589]" pageId="5" pageNumber="36">
Geological age. Earliest Eocene, Sparnacian, level MP7 of the mammal fauna of Dormaal. We have demonstrated that the amber is autochthonous and very different from the Baltic amber in age, chemical composition and origin (
<bibRefCitation id="EFD77DA1FFC1FF9F7ABCFACFFBF3B0CC" author="Feugueur, L." box="[935,1120,1403,1429]" pageId="5" pageNumber="36" pagination="1 - 568" refId="ref3590" refString="Feugueur, L. (1963) L'Ypresien du Bassin de Paris. Essai de Monographie stratigraphique. Memoires pour servir a l'explication de la Carte Geologique detaillee de la France, Publication du Ministere de l'Industrie, Paris, 1 - 568." type="book chapter" year="1963">Feugueur 1963</bibRefCitation>
;
<bibRefCitation id="EFD77DA1FFC1FF9F7D6BFACFFED5B0E4" author="De Ploeg, G. &amp; Dutheil, D. &amp; Gheerbrant, E. &amp; Godinot, M. &amp; Jossang, A. &amp; Nel, A. &amp; Paicheler, J. - C. &amp; Pons, D. &amp; Rage, J. - C." pageId="5" pageNumber="36" pagination="108 - 110" refId="ref3405" refString="De Ploeg, G., Dutheil, D., Gheerbrant, E., Godinot, M., Jossang, A., Nel, A., Paicheler, J. - C., Pons, D. &amp; Rage, J. - C. (1998) Un nouveau gisement paleontologique ' Konservat - Lagerstatte' a la base de l'Eocene dans la region de Creil (Oise). In: The Palaeoecene / Eocene boundary in Europe: events and correlations. Symposium Societe Geologique de France, Paris, 19 - 20 janvier 1998, pp. 108 - 110." type="proceedings paper" year="1998">
De Ploëg
<emphasis id="B932DC42FFC1FF9F7DF7FAC8FAB7B0CC" box="[1260,1316,1404,1429]" italics="true" pageId="5" pageNumber="36">et al</emphasis>
. 1998
</bibRefCitation>
;
<bibRefCitation id="EFD77DA1FFC1FF9F7849FA17FD96B0E4" author="Nel, A. &amp; De Ploeg, G. &amp; Dejax, J. &amp; Dutheil, D. &amp; De Franceschi, D. &amp; Gheerbrant, E. &amp; Godinot, M. &amp; Hervet, S. &amp; Menier, J-J. &amp; Auge, M. &amp; Bignot, G. &amp; Cavagnetto, C. &amp; Duffaud, S. &amp; Gaudant, J. &amp; Hua, S. &amp; Jossang, A. &amp; De Lapparent De Broin, F. &amp; Pozzi, J. - P. &amp; Paicheler, J. - C. &amp; Bouchet, F. &amp; Rage, J. - C." box="[338,517,1443,1469]" pageId="5" pageNumber="36" pagination="65 - 72" refId="ref3864" refString="Nel, A., De Ploeg, G., Dejax, J., Dutheil, D., De Franceschi, D., Gheerbrant, E., Godinot, M., Hervet, S., Menier, J-J., Auge, M., Bignot, G., Cavagnetto, C., Duffaud, S., Gaudant, J., Hua, S., Jossang, A., De Lapparent De Broin, F., Pozzi, J. - P., Paicheler, J. - C., Bouchet, F. &amp; Rage, J. - C. (1999) Un gisement sparnacien exceptionnel a plantes, arthropodes et vertebres (Eocene basal, MP 7): Le Quesnoy (Oise, France). Comptes Rendus de l'Academie des Sciences, Sciences de la terre et des planetes, Paris, 329, 65 - 72." type="journal article" year="1999">
Nel
<emphasis id="B932DC42FFC1FF9F789FFA10FE24B0E4" box="[388,439,1444,1469]" italics="true" pageId="5" pageNumber="36">et al</emphasis>
. 1999
</bibRefCitation>
).
</paragraph>
<paragraph id="8BF90050FFC1FF9F7823FA7FFB72B354" blockId="5.[264,1322,1283,1589]" pageId="5" pageNumber="36">
Remarks. The recent
<taxonomicName id="4C467BD3FFC1FF9F7B2EFA7FFD39B0BC" authorityName="Kieffer" authorityYear="1906" box="[565,682,1483,1509]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="family">Dryinidae</taxonomicName>
are parasitoids of
<taxonomicName id="4C467BD3FFC1FF9F7A9CFA7FFB94B0BC" box="[903,1031,1483,1509]" class="Insecta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
: Auchenorrhyncha. Fossil
<taxonomicName id="4C467BD3FFC1FF9F7830FA47FE2EB354" authorityName="Latreille" authorityYear="1825" box="[299,445,1523,1549]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="family">Cicadellidae</taxonomicName>
and Fulgoroidea are rather diverse and frequent in the ëOiseí amber.
</paragraph>
<paragraph id="8BF90050FFC1FF9F7823F9AFFC57B36C" blockId="5.[264,1322,1283,1589]" box="[312,964,1563,1589]" pageId="5" pageNumber="36">Etymology. The species is named after the city of Paris.</paragraph>
</subSubSection>
</treatment>
</document>